150 lines
16 KiB
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150 lines
16 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.335.5567" ID-GBIF-Dataset="f0c6f528-5c7c-4e61-87c3-dbf43b9350ef" ID-PMC="PMC3800789" ID-Pensoft-Pub="1313-2970-335-1" ID-PubMed="24146563" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-335-1" ModsDocOrigin="ZooKeys 335" ModsDocTitle="Shallow water marine gammaridean amphipods of Pulau Tioman, Malaysia, with the description of a new species" checkinTime="1451246890703" checkinUser="pensoft" docAuthor="Azman, B. A. R. & Othman, B. H. R." docDate="2013" docId="67A002947C6414D0C20E28F316D014ED" docLanguage="en" docName="ZooKeys 335: 1-31" docOrigin="ZooKeys 335" docSource="http://dx.doi.org/10.3897/zookeys.335.5567" docTitle="Ericthonius pugnax Dana 1853" docType="treatment" docVersion="3" lastPageNumber="9" masterDocId="6B111C28A76E1D660F77FF9FFFCFFFCB" masterDocTitle="Shallow water marine gammaridean amphipods of Pulau Tioman, Malaysia, with the description of a new species" masterLastPageNumber="31" masterPageNumber="1" pageNumber="9" updateTime="1668156506126" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Shallow water marine gammaridean amphipods of Pulau Tioman, Malaysia, with the description of a new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Azman, B. A. R.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Othman, B. H. R.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="volume">
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<mods:number>335</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>31</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.335.5567</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.335.5567</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-335-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152048371" LSID="urn:lsid:plazi:treatment:67A002947C6414D0C20E28F316D014ED" httpUri="http://treatment.plazi.org/id/67A002947C6414D0C20E28F316D014ED" lastPageNumber="9" pageId="8" pageNumber="9">
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<subSubSection pageId="8" pageNumber="9" type="nomenclature">
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<paragraph pageId="8" pageNumber="9">
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<taxonomicName LSID="http://species-id.net/wiki/Ericthonius_pugnax" authority="Dana, 1853" authorityName="Dana" authorityYear="1853" class="Malacostraca" family="Ischyroceridae" genus="Ericthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ericthonius pugnax" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="pugnax">Ericthonius pugnax (Dana, 1853)</taxonomicName>
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Figure 6
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="9" type="reference_group">
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<paragraph pageId="8" pageNumber="9">
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<taxonomicName class="Malacostraca" family="Ischyroceridae" genus="Pyctilus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pyctilus macrodactylus" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="macrodactylus">Pyctilus macrodactylus</taxonomicName>
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Synonymy: Dana, 1853: 974.
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</paragraph>
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<paragraph pageId="8" pageNumber="9">
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<taxonomicName class="Malacostraca" family="Ischyroceridae" genus="Ericthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ericthonius macrodactylus" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="macrodactylus">Ericthonius macrodactylus</taxonomicName>
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Stebbing, 1906: 672;
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<bibRefCitation author="Walker, AO" journalOrPublisher="Report of the Government of Ceylon Pearl Oyster Fishery of the Gulf of Manaar" pageId="21" pageNumber="22" pagination="229 - 300" title="Report on Amphipoda collected by Professor Herdman, at Ceylon, in 1902." volume="17" year="1904">Walker 1904</bibRefCitation>
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: 292, fig. 48.
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</paragraph>
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<paragraph pageId="8" pageNumber="9">
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<taxonomicName class="Malacostraca" family="Ischyroceridae" genus="Ericthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ericthonius pugnax" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="pugnax">Ericthonius pugnax</taxonomicName>
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Stebbing, 1906: 672;
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<bibRefCitation pageId="8" pageNumber="9">Pirlot 1938</bibRefCitation>
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: 352;
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<bibRefCitation author="Hurley, DE" journalOrPublisher="Transactions of the Royal Society of New Zealand" pageId="18" pageNumber="19" pagination="431 - 460" volume="82" year="1954">Hurley 1954</bibRefCitation>
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: 445, figs 40-61;
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<bibRefCitation author="Nagata, K" journalOrPublisher="Publications of the Seto Marine Biological Laboratory" pageId="19" pageNumber="20" pagination="163 - 182" title="Preliminary notes on benthic gammaridean Amphipoda from the Zostera region of Mihara Bay, Seto Inland Sea, Japan." volume="8" year="1960">Nagata 1960</bibRefCitation>
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: 179, pl. 179, figs 99-102;
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<bibRefCitation author="Nagata, K" journalOrPublisher="Publication of Seto Marine Biological Laboratory" pageId="19" pageNumber="20" pagination="291 - 322" title="Studies on marine Gammaridean Amphipoda of the Seto Inland Sea III." volume="13" year="1965 b">Nagata 1965b</bibRefCitation>
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: 320, fig. 40;
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<bibRefCitation pageId="8" pageNumber="9">Nayar 1967</bibRefCitation>
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:162;
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<bibRefCitation author="Ledoyer, M" journalOrPublisher="Recueil des Travaux de la Station Marine Endoume, Fascicule hors serie Supplement" pageId="18" pageNumber="19" pagination="179 - 182" title="Amphipodes tubicoles des feuilles des herbiers de phanerogames marines de la region de Tulear (Madagascar)." volume="9" year="1969">Ledoyer 1969</bibRefCitation>
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: 179, fig. 1;
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<bibRefCitation author="Hirayama, A" journalOrPublisher="Publication of Seto Marine Biological Laboratory" pageId="17" pageNumber="18" pagination="1 - 53" title="Taxonomic studies on the shallow water Gammaridean Amphipoda of west Kyushu, Japan. IV. Dexaminidae (Guernea), Eophiliantidae, Eusoridae, Haustoriidae, Hyalidae and Ischyroceridae." volume="30" year="1985">Hirayama 1985</bibRefCitation>
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: 52,
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<bibRefCitation author="Ledoyer, M" journalOrPublisher="Memorie del Museo Civico di Storia Naturale di Verona" pageId="18" pageNumber="19" pagination="45 - 84" title="Les Oedicerotidae (Crustacea Amphipoda) De La Mer Mediterranee." volume="9" year="1982 a">Ledoyer 1982a</bibRefCitation>
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: 628, fig. 239;
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<bibRefCitation author="Moore, PG" journalOrPublisher="Journal of Natural History" pageId="19" pageNumber="20" pagination="705 - 732" title="Taxonomic observations on the genera Xenocheria Haswell and Ericthonius Milne-Edwards (Crustacea: Amphipoda) from Australian coastal waters." url="10.1080/00222938800770461" volume="22" year="1988">Moore 1988</bibRefCitation>
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: 727-730, fig. 14;
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<bibRefCitation author="Kim, W" journalOrPublisher="Korean J. Zool." pageId="18" pageNumber="19" pagination="232 - 252" title="The Marine Amphipod Crustaceans of Ulreung Island, Korea: Part 1." volume="34" year="1991">Kim and Kim 1991</bibRefCitation>
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: 246-247, figs 13-14.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="9" type="material">
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<paragraph pageId="8" pageNumber="9">Material.</paragraph>
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<paragraph pageId="8" pageNumber="9">
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2 specimens, TIO 5, Monkey Bay, Pulau Tioman, 2
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="2.8258333">2°49'33"N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="15" value="104.1625">104°9'45"E</geoCoordinate>
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, coral rubble, Azman, Josim, 22 August 1996; 86 specimens, TIO 15, Renggis, Pulau Tioman,
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="2.8097222">2°48'35"N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="15" value="104.134995">104°8'6"E</geoCoordinate>
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, seagrass, Azman, Rayida, 15 July 1999, UKM I.D. 5712-5719; 2 specimens, TIO 33, Tomok, Pulau Tioman,
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="2.7938888">2°47'38"N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="15" value="104.12112">104°7'16"E</geoCoordinate>
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, live corals (
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<taxonomicName class="Anthozoa" family="Poritidae" genus="Porites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Porites" order="Scleractinia" pageId="8" pageNumber="9" phylum="Cnidaria" rank="genus">Porites</taxonomicName>
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sp.,
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<taxonomicName class="Anthozoa" family="Acroporidae" genus="Montipora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Montipora" order="Scleractinia" pageId="8" pageNumber="9" phylum="Cnidaria" rank="genus">Montipora</taxonomicName>
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sp.,
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<taxonomicName class="Anthozoa" family="Acroporidae" genus="Acropora" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acropora" order="Scleractinia" pageId="8" pageNumber="9" phylum="Cnidaria" rank="genus">Acropora</taxonomicName>
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sp.), Azman, B.A.R, Kee, A.A., Zuhaimi, S., Maekawa, T., Okashita, T., 22 March 2004.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="9" type="remarks">
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<paragraph pageId="8" pageNumber="9">Remarks.</paragraph>
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<paragraph pageId="8" pageNumber="9">
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The Pulau Tioman specimens agree well with figures of
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<taxonomicName class="Malacostraca" family="Ischyroceridae" genus="Ericthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ericthonius pugnax" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="pugnax">Ericthonius pugnax</taxonomicName>
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from
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<bibRefCitation author="Walker, AO" journalOrPublisher="Report of the Government of Ceylon Pearl Oyster Fishery of the Gulf of Manaar" pageId="21" pageNumber="22" pagination="229 - 300" title="Report on Amphipoda collected by Professor Herdman, at Ceylon, in 1902." volume="17" year="1904">Walker (1904)</bibRefCitation>
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,
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<bibRefCitation author="Nagata, K" journalOrPublisher="Publications of the Seto Marine Biological Laboratory" pageId="19" pageNumber="20" pagination="163 - 182" title="Preliminary notes on benthic gammaridean Amphipoda from the Zostera region of Mihara Bay, Seto Inland Sea, Japan." volume="8" year="1960">Nagata (1960</bibRefCitation>
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, 1965c),
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<bibRefCitation author="Ledoyer, M" journalOrPublisher="Centre National de la Recherche Scientifique-Paris" pageId="19" pageNumber="20" title="Faune de Madagascar. Crustaces Amphipodes Gammariens; Families des Haustoriidae a' Vitjazianidae." volume="59" year="1982 c">Ledoyer (1982c)</bibRefCitation>
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and
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<bibRefCitation author="Moore, PG" journalOrPublisher="Journal of Natural History" pageId="19" pageNumber="20" pagination="705 - 732" title="Taxonomic observations on the genera Xenocheria Haswell and Ericthonius Milne-Edwards (Crustacea: Amphipoda) from Australian coastal waters." url="10.1080/00222938800770461" volume="22" year="1988">Moore (1988)</bibRefCitation>
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. This species seems to have a wide-ranging distribution as far as the Arabian Sea. From closer examination of the specimens, the males not only have distally bidentate carpal spur of gnathopod 2 but also a mixture of single dentate carpal spur (see also
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<bibRefCitation author="Nagata, K" journalOrPublisher="Publications of the Seto Marine Biological Laboratory" pageId="19" pageNumber="20" pagination="163 - 182" title="Preliminary notes on benthic gammaridean Amphipoda from the Zostera region of Mihara Bay, Seto Inland Sea, Japan." volume="8" year="1960">Nagata 1960</bibRefCitation>
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;
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<bibRefCitation author="Ledoyer, M" journalOrPublisher="Recueil des Travaux de la Station Marine Endoume, Fascicule hors serie Supplement" pageId="18" pageNumber="19" pagination="179 - 182" title="Amphipodes tubicoles des feuilles des herbiers de phanerogames marines de la region de Tulear (Madagascar)." volume="9" year="1969">Ledoyer 1969</bibRefCitation>
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,
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<bibRefCitation author="Ledoyer, M" journalOrPublisher="Mexico" pageId="19" pageNumber="20" pagination="171 - 200" title="Faune mobile des herbiers de phanerogames marines (Holodule et Thalassia) de la lagune de terminus (Mexique, Campeche). II. Les Gammariens (Crustacea). An. Inst. Cienc. del Mar y Limnol. Univ. Nal. Auton." volume="13" year="1986">1986</bibRefCitation>
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). As mentioned by
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<bibRefCitation author="Moore, PG" journalOrPublisher="Journal of Natural History" pageId="19" pageNumber="20" pagination="705 - 732" title="Taxonomic observations on the genera Xenocheria Haswell and Ericthonius Milne-Edwards (Crustacea: Amphipoda) from Australian coastal waters." url="10.1080/00222938800770461" volume="22" year="1988">Moore (1988)</bibRefCitation>
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the carpal bidentate spur of ganthopod 2 will eventually be lost in hyperadult males. Unfortunately the stipulation over hyper adult morphological changes has brought several descriptions of new species that actually represented a single species (i.e
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<taxonomicName class="Malacostraca" family="Ischyroceridae" genus="Ericthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ericthonius macrodactylus" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="macrodactylus">Ericthonius macrodactylus</taxonomicName>
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). Prior to this,
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<bibRefCitation author="Ledoyer, M" journalOrPublisher="Recueil des Travaux de la Station Marine Endoume, Fascicule hors serie Supplement" pageId="18" pageNumber="19" pagination="179 - 182" title="Amphipodes tubicoles des feuilles des herbiers de phanerogames marines de la region de Tulear (Madagascar)." volume="9" year="1969">Ledoyer (1969)</bibRefCitation>
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synonymized
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<taxonomicName class="Malacostraca" family="Ischyroceridae" genus="Ericthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ericthonius macrodactylus" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="macrodactylus">Ericthonius macrodactylus</taxonomicName>
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as a hyperadult form of
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<taxonomicName class="Malacostraca" family="Ischyroceridae" genus="Ericthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ericthonius pugnax" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="pugnax">Ericthonius pugnax</taxonomicName>
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. Examination of the Pulau Tioman specimens confirms the presence of this hyper adult morphology: that the basis of pereopod 5 possesses a strongly developed posterodistal lobe. The Pulau Tioman specimens are undoubtedly assigned to
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<taxonomicName class="Malacostraca" family="Ischyroceridae" genus="Ericthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ericthonius pugnax" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="pugnax">Ericthonius pugnax</taxonomicName>
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and constitute the first record of this species for Peninsular Malaysia.
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</paragraph>
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<caption pageId="8" pageNumber="9">
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<paragraph pageId="8" pageNumber="9">
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Figure 6.
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<taxonomicName class="Malacostraca" family="Ischyroceridae" genus="Ericthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ericthonius pugnax" order="Amphipoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="pugnax">Ericthonius pugnax</taxonomicName>
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(Dana, 1853), male, (UKMMZ-1136), 3.8 mm. Pulau Tioman, South China Sea. Scale for G2 ♂ represents 0.6 mm; A1, A2 HD and ABD scales = 0.5 mm, G1 ♂ scale = 0.25 mm; P5 scale = 0.2 mm.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |