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<document id="DA7FFB0064DB9EFCF5D832CD643A7BEA" ID-DOI="10.5281/zenodo.5300975" ID-ISSN="0374-1036" ID-Zenodo-Dep="5300975" ID-ZooBank="9CC5EBA3-8351-4C89-B9B2-0D84C17BBF0B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630098542094" checkinUser="marcus" docAuthor="Clarkson, Bruno, Albertoni, Fabiano Fabian &amp; FIKÁýEK, Martin" docDate="2014" docId="03E4BF23CC72FFD0FE00FC5DFDFDD858" docLanguage="en" docName="ActaEntMusNatPra.2014.54.1.157-194.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 54 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Lachnodacnum Orchymont 1937" docType="treatment" docVersion="4" lastPageNumber="167" masterDocId="FFDDC75BCC71FFDAFF9CFFC1FFE0DE4B" masterDocTitle="Taxonomy and biology of the bromeliad-inhabiting genus Lachnodacnum (Coleoptera: Hydrophilidae: Sphaeridiinae)" masterLastPageNumber="194" masterPageNumber="157" pageNumber="160" updateTime="1699144678491" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="63E9AE9C216ED1DC54E4F1CB49A0DFA5">Taxonomy and biology of the bromeliad-inhabiting genus Lachnodacnum (Coleoptera: Hydrophilidae: Sphaeridiinae)</mods:title>
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<mods:namePart id="1B6CF982D82586E57B725B4E282D83D5">Clarkson, Bruno</mods:namePart>
<mods:affiliation id="538929B67E26E911FF505BD8B34B718F">Programa de Pós-Graduação em Zoologia, Museu Nacional, Laboratório de Entomologia, Departamento de Zoologia, Instituto de Biologia, Universidade Federal do Rio de Janeiro, Caixa Postal 68044, 21941 - 971, Rio de Janeiro, RJ, Brasil; e-mail: brclarkson @ gmail. com</mods:affiliation>
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<mods:namePart id="9D98E1FEC9CCE8F4BD467516116CF035">Albertoni, Fabiano Fabian</mods:namePart>
<mods:affiliation id="EB407AC5FD821BBC733DA684777BB7AB">Programa de Pós-Graduação em Sistemática, taxonomia animal e biodiversidade, Entomologia, Museu de Zoologia da Universidade de São Paulo, Av. Nazaré, 481, Ipiranga, CEP 04263 - 000, São Paulo, SP, Brasil; e-mail: fabianoalbertoni @ gmail. com</mods:affiliation>
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<taxonomicName id="4C4D75B6CC72FFD9FE00FC5DFCF6DDF8" ID-CoL="8X7HN" authority="Orchymont, 1937" authorityName="Orchymont" authorityYear="1937" box="[412,790,924,947]" class="Insecta" family="Hydrophilidae" genus="Lachnodacnum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="B939D227CC72FFD9FE00FC5DFCF6DDF8" bold="true" box="[412,790,924,947]" pageId="3" pageNumber="160">
<emphasis id="B939D227CC72FFD9FE00FC5DFDA9DDF8" bold="true" box="[412,585,924,947]" italics="true" pageId="3" pageNumber="160">Lachnodacnum</emphasis>
<bibRefCitation id="EFDC73C4CC72FFD9FDCCFC5DFCF6DDF8" author="ORCHYMONT A." box="[592,790,924,947]" pageId="3" pageNumber="160" pagination="127" refId="ref18556" refString="ORCHYMONT A. d' 1937: Sphaeridiini bromeliadicoles nouveaux (Coleoptera: Hydrophilidae, Sphaeridiinae). Annals and Magazine of Natural History, Series 10 20: 127 i 140." type="journal article" year="1937">Orchymont, 1937</bibRefCitation>
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<paragraph id="8BF20E35CC72FFD9FFEDFC03FE22DD9E" blockId="3.[113,450,962,981]" box="[113,450,962,981]" pageId="3" pageNumber="160">
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<treatmentCitation id="0AEC2824CC72FFD9FFEDFC03FE5FDD9E" author="ORCHYMONT A." box="[113,447,962,981]" page="134" pageId="3" pageNumber="160" year="1937">
<taxonomicName id="4C4D75B6CC72FFD9FFEDFC03FE5FDD9E" ID-CoL="8X7HN" authority="Orchymont, 1937: 134" authorityName="Orchymont" authorityPageNumber="134" authorityYear="1937" box="[113,447,962,981]" class="Insecta" family="Hydrophilidae" genus="Lachnodacnum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="160" phylum="Arthropoda" rank="genus">
<emphasis id="B939D227CC72FFD9FFEDFC03FF15DD9E" box="[113,245,962,981]" italics="true" pageId="3" pageNumber="160">Lachnodacnum</emphasis>
<bibRefCitation id="EFDC73C4CC72FFD9FF66FC03FE5FDD9E" author="ORCHYMONT A." box="[250,447,962,981]" pageId="3" pageNumber="160" pagination="127" refId="ref18556" refString="ORCHYMONT A. d' 1937: Sphaeridiini bromeliadicoles nouveaux (Coleoptera: Hydrophilidae, Sphaeridiinae). Annals and Magazine of Natural History, Series 10 20: 127 i 140." type="journal article" year="1937">Orchymont, 1937: 134</bibRefCitation>
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.
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<emphasis id="B939D227CC72FFD9FFEDFC2CFEE5DA4F" bold="true" box="[113,261,1005,1028]" pageId="3" pageNumber="160">
<typeStatus id="54F6B097CC72FFD9FFEDFC2CFF49DA4F" box="[113,169,1005,1028]" pageId="3" pageNumber="160">Type</typeStatus>
species.
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<taxonomicName id="4C4D75B6CC72FFD9FE90FC2CFD04DA4F" ID-CoL="3RPTK" authority="Orchymont, 1937" authorityName="Orchymont" authorityYear="1937" box="[268,740,1005,1028]" class="Insecta" family="Hydrophilidae" genus="Lachnodacnum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="160" phylum="Arthropoda" rank="species" species="saundersi">
<emphasis id="B939D227CC72FFD9FE90FC2CFDFFDA4F" box="[268,543,1005,1028]" italics="true" pageId="3" pageNumber="160">Lachnodacnum saundersi</emphasis>
<bibRefCitation id="EFDC73C4CC72FFD9FDB9FC2CFD04DA4F" author="ORCHYMONT A." box="[549,740,1005,1028]" pageId="3" pageNumber="160" pagination="127" refId="ref18556" refString="ORCHYMONT A. d' 1937: Sphaeridiini bromeliadicoles nouveaux (Coleoptera: Hydrophilidae, Sphaeridiinae). Annals and Magazine of Natural History, Series 10 20: 127 i 140." type="journal article" year="1937">Orchymont, 1937</bibRefCitation>
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, by original designation.
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<paragraph id="8BF20E35CC72FFD9FFEDFBCCFCF3DB8F" blockId="3.[113,1090,1005,1604]" pageId="3" pageNumber="160">
<emphasis id="B939D227CC72FFD9FFEDFBCCFF03DA6F" bold="true" box="[113,227,1037,1060]" pageId="3" pageNumber="160">Diagnosis.</emphasis>
Eyes small, rounded in dorsal view (Fig. 3); labial palpomere 2 bearing dense brush of long setae on inner face and distal margin; pronotum weakly convex, anterior corners rounded, posterior corners weakly angulate; hypomeron with large pubescent inner portion and moderately wide marginal glabrous portion (Fig. 26); mesoventrite well-divided from anepisternum 2 except sublaterally, anepisternal suture distinct medially and laterally (Figs 27, 34); preepisternal plate widely attaching and as high as anterior portion of metaventral process, forming a mesometaventral keel, constricted at the area where meso- and metaventrite join each other (
<figureCitation id="137612B0CC72FFD9FC71FB0CFBDDDAAF" box="[1005,1085,1229,1252]" captionStart="Figs 24" captionStartId="8.[150,187,1274,1293]" captionTargetBox="[151,1126,219,1256]" captionTargetId="figure-88@8.[151,1126,219,1257]" captionTargetPageId="8" captionText="Figs 24í28. Adult morphology of Lachnodacnum luederwaldti Orchymont, 1937, SEM micrographs. 24 whole beetle, ventral view; 25 head, ventral view; 26 prothorax in ventral view (prosternum and hypomeron); 27 lateral portion of mesoventrite (arrow indicates the anepisternal suture); 28 antenna." figureDoi="http://doi.org/10.5281/zenodo.5300985" httpUri="https://zenodo.org/record/5300985/files/figure.png" pageId="3" pageNumber="160">Figs 24</figureCitation>
,
<figureCitation id="137612B0CC72FFD9FFEDFB2CFF6BDB4F" box="[113,139,1261,1284]" captionStart="Figs 33" captionStartId="12.[150,187,1115,1134]" captionTargetBox="[177,1100,219,1097]" captionTargetId="figure-259@12.[177,1100,219,1097]" captionTargetPageId="12" captionText="Figs 33í36. Mesoventral elevation of Lachnodacnum Orchymont, 1937, SEM micrographs. In ventral (33í34) and lateral (35í36) views. 33, 35 L. luederwaldti Orchymont, 1937; 34, 36 L. saundersi Orchymont, 1937 (arrow indicates the anepisternal suture)." figureDoi="http://doi.org/10.5281/zenodo.5319472" httpUri="https://zenodo.org/record/5319472/files/figure.png" pageId="3" pageNumber="160">33</figureCitation>
í36); anteromedian pit-like groove large, well developed (Fig. 27); grooves for reception of procoxae not de¿ned (Figs 27, 34); elytron weakly convex, sutural stria very ¿ne but distinct, developed only on elytral apex (Fig. 30); mesofemur pubescent on ventral surface (
<figureCitation id="137612B0CC72FFD9FC79FAECFBD5DB0F" box="[997,1077,1325,1348]" captionStart="Figs 29" captionStartId="9.[113,149,996,1015]" captionTargetBox="[114,1089,219,983]" captionTargetId="figure-186@9.[114,1090,219,983]" captionTargetPageId="9" captionText="Figs 29í32.Adult morphology of Lachnodacnum luederwaldti Orchymont, 1937,SEM micrographs.29 mesofemur; 30 apical portion of elytra; 31 microsculpture of the clypeus; 32 trichobothrium on the elytron." figureDoi="http://doi.org/10.5281/zenodo.5300987" httpUri="https://zenodo.org/record/5300987/files/figure.png" pageId="3" pageNumber="160">Fig. 29</figureCitation>
); metatarsus slightly compressed laterally, metatarsomere 1 about twice as long as metatarsomere 2; aedeagus with rather long and well developed symmetrical phallobase (
<figureCitation id="137612B0CC72FFD9FCE0FAACFC2BDBCF" box="[892,971,1389,1412]" captionStart="Figs 15" captionStartId="6.[150,187,853,872]" captionTargetBox="[239,1036,223,827]" captionTargetId="figure-267@6.[224,1053,212,830]" captionTargetPageId="6" captionText="Figs 15í17. Aedeagus. 15 Lachnodacnum luederwaldti Orchymont, 1937; 16 L. saundersi Orchymont, 1937; 17 Phaenostoma urichi (Scott, 1912)." figureDoi="http://doi.org/10.5281/zenodo.5300981" httpUri="https://zenodo.org/record/5300981/files/figure.png" pageId="3" pageNumber="160">Figs 15</figureCitation>
í16); parameres wide throughout; median lobe wide basally, with a pair of short basal projections, more or less gradually narrowing apicad; gonopore distinct, subapical.
</paragraph>
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<subSubSection id="C3575DBECC72FFD0FFEDFA0CFDFDD858" lastPageId="10" lastPageNumber="167" pageId="3" pageNumber="160" type="description">
<paragraph id="8BF20E35CC72FFD9FFEDFA0CFD27D84F" blockId="3.[113,1090,1005,1604]" pageId="3" pageNumber="160">
<emphasis id="B939D227CC72FFD9FFEDFA0CFEF6DBAF" bold="true" box="[113,278,1485,1508]" pageId="3" pageNumber="160">Redescription.</emphasis>
Body widely oval, weakly convex in lateral view; pronotal and elytral outline continuous in dorsal and lateral views (
<figureCitation id="137612B0CC72FFD9FD8FFA2CFDB6D84F" box="[531,598,1517,1540]" captionStart="Figs 1" captionStartId="4.[151,188,1467,1486]" captionTargetBox="[159,1119,224,1353]" captionTargetId="figure-10@4.[151,1120,215,1362]" captionTargetPageId="4" captionText="Figs 1í10. General habitus of adults and larvae of Lachnodacnum Orchymont, 1937. 1í3 adults of L. luederwaldti Orchymont, 1937 (1 dorsal view, 2 lateral view, 3 dorsal view of the head). 4í8 larva of L. luederwaldti (4 third instar larva in dorsal view; 5 same in ventral view; 6 ¿rst instar in dorsal view; 7 detail of head and thorax of third instar, dorsal view; 8 same, ventral view). 9í10 adults of L. saundersi Orchymont, 1937 (9 dorsal view, 10 lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5300977" httpUri="https://zenodo.org/record/5300977/files/figure.png" pageId="3" pageNumber="160">Figs 1</figureCitation>
í2, 9í10).
</paragraph>
<paragraph id="8BF20E35CC72FFDCFF0CF9CCFDEEDBB9" blockId="3.[113,1090,1005,1604]" lastBlockId="6.[151,1127,955,1586]" lastPageId="6" lastPageNumber="163" pageId="3" pageNumber="160">
<emphasis id="B939D227CC72FFD9FF0CF9CCFF31D86F" bold="true" box="[144,209,1549,1572]" pageId="3" pageNumber="160">
<emphasis id="B939D227CC72FFD9FF0CF9CCFF2AD86F" bold="true" box="[144,202,1549,1572]" italics="true" pageId="3" pageNumber="160">Head</emphasis>
.
</emphasis>
Clypeus wide even anteriorly, only slightly narrowing before eyes, with anteromedian margin truncate (Fig. 3). Frontoclypeal suture conspicuous; lateral portions of frons between anterior margin of eye and frontoclypeal suture lacking punctuation. Eyes small (Fig. 3), not protruding laterad, rounded in dorsal view, surrounded by micropunctures, separated by 7× the width of one eye, distinctly emarginated anteriorly in lateral view. Clypeus and frons with scattered trichobothria. Labrum small, membranous, concealed under clypeus. Maxillary palps with four palpomeres, short and rather stout. Mentum (Fig. 25) 1.4× as wide as long; lateral margins nearly parallel-sided, each side bearing a row of setae [examined in
<taxonomicName id="4C4D75B6CC77FFDCFBCDFB9AFEFEDAD9" authorityName="Orchymont" authorityYear="1937" class="Insecta" family="Hydrophilidae" genus="Lachnodacnum" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="163" phylum="Arthropoda" rank="species" species="luederwaldti">
<emphasis id="B939D227CC77FFDCFBCDFB9AFEFEDAD9" italics="true" pageId="6" pageNumber="163">L. luederwaldti</emphasis>
</taxonomicName>
only, poorly preserved in available specimens of
<taxonomicName id="4C4D75B6CC77FFDCFCB5FBBAFC4DDAD9" authorityName="Orchymont" authorityYear="1937" box="[809,941,1147,1170]" class="Insecta" family="Hydrophilidae" genus="Lachnodacnum" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="163" phylum="Arthropoda" rank="species" species="saundersi">
<emphasis id="B939D227CC77FFDCFCB5FBBAFC4DDAD9" box="[809,941,1147,1170]" italics="true" pageId="6" pageNumber="163">L. saundersi</emphasis>
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]; anterior margin deeply bisinuate; surface of mentum bearing sparsely distributed long setae. Labial palps with three palpomeres: palpomere 2 longest, strongly widened distally, bearing dense brush of long setae on its inner face and distal margin; palpomere 3 much narrower and about 0.6× as long as palpomere 2. Submentum pubescent. Gula slightly constricted between tentorial pits, bearing low longitudinal carina in posterior half [examined only for
<taxonomicName id="4C4D75B6CC77FFDCFC25FADAFBBEDB79" authorityName="Orchymont" authorityYear="1937" box="[953,1118,1307,1330]" class="Insecta" family="Hydrophilidae" genus="Lachnodacnum" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="163" phylum="Arthropoda" rank="species" species="luederwaldti">
<emphasis id="B939D227CC77FFDCFC25FADAFBBEDB79" box="[953,1118,1307,1330]" italics="true" pageId="6" pageNumber="163">L. luederwaldti</emphasis>
</taxonomicName>
] (Fig. 25). Antenna with nine antennomeres (Fig. 28); scapus rather long and thick, its basal portion bent dorsally; pedicel cylindrical; antennomere 3 twice the length of antennomere 4, antennomeres 3í5 narrower than pedicel proximally, slightly widened distally; cupula long and wide, bare, concealing basal portion of antennomere 7; antennal club loosely segmented, depressed dorsoventrally, densely pubescent with few longer and thicker setae on sides and on distal margins of antennomeres.
</paragraph>
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Figs 1í10. General habitus of adults and larvae of
<taxonomicName id="4C4D75B6CC75FFDEFDA1FA7AFCBDDB85" authority="Orchymont, 1937" authorityName="Orchymont" authorityYear="1937" box="[573,861,1467,1486]" class="Insecta" family="Hydrophilidae" genus="Lachnodacnum" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="161" phylum="Arthropoda" rank="genus">
<emphasis id="B939D227CC75FFDEFDA1FA7AFD21DB85" box="[573,705,1467,1486]" italics="true" pageId="4" pageNumber="161">Lachnodacnum</emphasis>
<bibRefCitation id="EFDC73C4CC75FFDEFD59FA7AFCBDDB85" author="ORCHYMONT A." box="[709,861,1467,1486]" pageId="4" pageNumber="161" pagination="127" refId="ref18556" refString="ORCHYMONT A. d' 1937: Sphaeridiini bromeliadicoles nouveaux (Coleoptera: Hydrophilidae, Sphaeridiinae). Annals and Magazine of Natural History, Series 10 20: 127 i 140." type="journal article" year="1937">Orchymont, 1937</bibRefCitation>
</taxonomicName>
. 1í3 adults of
<taxonomicName id="4C4D75B6CC75FFDEFC79FA7AFECFDBA3" authority="Orchymont, 1937" authorityName="Orchymont" authorityYear="1937" class="Insecta" family="Hydrophilidae" genus="Lachnodacnum" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="161" phylum="Arthropoda" rank="species" species="luederwaldti">
<emphasis id="B939D227CC75FFDEFC79FA7AFB86DB85" box="[997,1126,1467,1486]" italics="true" pageId="4" pageNumber="161">L. luederwaldti</emphasis>
<bibRefCitation id="EFDC73C4CC75FFDEFF0BFA14FECFDBA3" author="ORCHYMONT A." box="[151,303,1493,1512]" pageId="4" pageNumber="161" pagination="127" refId="ref18556" refString="ORCHYMONT A. d' 1937: Sphaeridiini bromeliadicoles nouveaux (Coleoptera: Hydrophilidae, Sphaeridiinae). Annals and Magazine of Natural History, Series 10 20: 127 i 140." type="journal article" year="1937">Orchymont, 1937</bibRefCitation>
</taxonomicName>
(1 dorsal view, 2 lateral view, 3 dorsal view of the head). 4í8 larva of
<taxonomicName id="4C4D75B6CC75FFDEFC7FFA14FB86DBA3" authorityName="Orchymont" authorityYear="1937" box="[995,1126,1493,1512]" class="Insecta" family="Hydrophilidae" genus="Lachnodacnum" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="161" phylum="Arthropoda" rank="species" species="luederwaldti">
<emphasis id="B939D227CC75FFDEFC7FFA14FB86DBA3" box="[995,1126,1493,1512]" italics="true" pageId="4" pageNumber="161">L. luederwaldti</emphasis>
</taxonomicName>
(4 third instar larva in dorsal view; 5 same in ventral view; 6 ¿rst instar in dorsal view; 7 detail of head and thorax of third instar, dorsal view; 8 same, ventral view). 9í10 adults of
<taxonomicName id="4C4D75B6CC75FFDEFCC1F9CAFB87D855" authority="Orchymont, 1937" authorityName="Orchymont" authorityYear="1937" box="[861,1127,1547,1566]" class="Insecta" family="Hydrophilidae" genus="Lachnodacnum" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="161" phylum="Arthropoda" rank="species" species="saundersi">
<emphasis id="B939D227CC75FFDEFCC1F9CAFC28D855" box="[861,968,1547,1566]" italics="true" pageId="4" pageNumber="161">L. saundersi</emphasis>
<bibRefCitation id="EFDC73C4CC75FFDEFC52F9CAFB87D855" author="ORCHYMONT A." box="[974,1127,1547,1566]" pageId="4" pageNumber="161" pagination="127" refId="ref18556" refString="ORCHYMONT A. d' 1937: Sphaeridiini bromeliadicoles nouveaux (Coleoptera: Hydrophilidae, Sphaeridiinae). Annals and Magazine of Natural History, Series 10 20: 127 i 140." type="journal article" year="1937">Orchymont, 1937</bibRefCitation>
</taxonomicName>
(9 dorsal view, 10 lateral view).
</paragraph>
</caption>
<caption id="DF325EBDCC74FFDFFFEDFA2BFD2BD879" ID-DOI="http://doi.org/10.5281/zenodo.5300979" ID-Zenodo-Dep="5300979" httpUri="https://zenodo.org/record/5300979/files/figure.png" pageId="5" pageNumber="162" startId="5.[113,150,1514,1533]" targetBox="[119,1083,218,1467]" targetPageId="5">
<paragraph id="8BF20E35CC74FFDFFFEDFA2BFD2BD879" blockId="5.[113,1088,1514,1586]" pageId="5" pageNumber="162">
Figs 11í14. Morphological details of
<taxonomicName id="4C4D75B6CC74FFDFFE33FA2BFD2CDBB6" authority="Orchymont, 1937" authorityName="Orchymont" authorityYear="1937" box="[431,716,1514,1533]" class="Insecta" family="Hydrophilidae" genus="Lachnodacnum" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="162" phylum="Arthropoda" rank="genus">
<emphasis id="B939D227CC74FFDFFE33FA2BFDD3DBB6" box="[431,563,1514,1533]" italics="true" pageId="5" pageNumber="162">Lachnodacnum</emphasis>
<bibRefCitation id="EFDC73C4CC74FFDFFDABFA2BFD2CDBB6" author="ORCHYMONT A." box="[567,716,1514,1533]" pageId="5" pageNumber="162" pagination="127" refId="ref18556" refString="ORCHYMONT A. d' 1937: Sphaeridiini bromeliadicoles nouveaux (Coleoptera: Hydrophilidae, Sphaeridiinae). Annals and Magazine of Natural History, Series 10 20: 127 i 140." type="journal article" year="1937">Orchymont, 1937</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C4D75B6CC74FFDFFD6FFA2BFBE3DBB6" authority="Orchymont, 1937" authorityName="Orchymont" authorityYear="1937" box="[755,1027,1514,1533]" class="Insecta" family="Hydrophilidae" genus="Phaenostoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="162" phylum="Arthropoda" rank="genus">
<emphasis id="B939D227CC74FFDFFD6FFA2BFC87DBB6" box="[755,871,1514,1533]" italics="true" pageId="5" pageNumber="162">Phaenostoma</emphasis>
<bibRefCitation id="EFDC73C4CC74FFDFFCF0FA2BFBE3DBB6" author="ORCHYMONT A." box="[876,1027,1514,1533]" pageId="5" pageNumber="162" pagination="127" refId="ref18556" refString="ORCHYMONT A. d' 1937: Sphaeridiini bromeliadicoles nouveaux (Coleoptera: Hydrophilidae, Sphaeridiinae). Annals and Magazine of Natural History, Series 10 20: 127 i 140." type="journal article" year="1937">Orchymont, 1937</bibRefCitation>
</taxonomicName>
. 11í12 hind wings (11
<taxonomicName id="4C4D75B6CC74FFDFFE88F9C4FD46D853" authority="Orchymont, 1937" authorityName="Orchymont" authorityYear="1937" box="[276,678,1541,1560]" class="Insecta" family="Hydrophilidae" genus="Lachnodacnum" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="162" phylum="Arthropoda" rank="species" species="luederwaldti">
<emphasis id="B939D227CC74FFDFFE88F9C4FDE9D853" box="[276,521,1541,1560]" italics="true" pageId="5" pageNumber="162">Lachnodacnum luederwaldti</emphasis>
<bibRefCitation id="EFDC73C4CC74FFDFFD91F9C4FD46D853" author="ORCHYMONT A." box="[525,678,1541,1560]" pageId="5" pageNumber="162" pagination="127" refId="ref18556" refString="ORCHYMONT A. d' 1937: Sphaeridiini bromeliadicoles nouveaux (Coleoptera: Hydrophilidae, Sphaeridiinae). Annals and Magazine of Natural History, Series 10 20: 127 i 140." type="journal article" year="1937">Orchymont, 1937</bibRefCitation>
</taxonomicName>
; 12
<taxonomicName id="4C4D75B6CC74FFDFFD44F9C4FBE2D853" authority="(Scott, 1912))" authorityName="Clarkson &amp; Albertoni &amp; FIKÁýEK" authorityYear="2014" baseAuthorityName="Scott" baseAuthorityYear="1912" box="[728,1026,1541,1560]" class="Insecta" family="Hydrophilidae" genus="Phaenostoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="162" phylum="Arthropoda" rank="species" species="urichi">
<emphasis id="B939D227CC74FFDFFD44F9C4FC64D853" box="[728,900,1541,1560]" italics="true" pageId="5" pageNumber="162">Phaenostoma urichi</emphasis>
(
<bibRefCitation id="EFDC73C4CC74FFDFFC13F9C4FC14D853" author="SCOTT H." box="[911,1012,1541,1560]" pageId="5" pageNumber="162" pagination="424" refId="ref18616" refString="SCOTT H. 1912: A contribution to the knowledge of the fauna of Bromeliaceae. Annals and Magazine of Natural History, Series 8 10: 424 i 438 + pl. X." type="journal article" year="1912">Scott, 1912</bibRefCitation>
))
</taxonomicName>
. 13í14 detail of spiracular atrium of the third instar larva of
<taxonomicName id="4C4D75B6CC74FFDFFDD8F9DEFD26D879" authorityName="Orchymont" authorityYear="1937" box="[580,710,1567,1586]" class="Insecta" family="Hydrophilidae" genus="Lachnodacnum" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="162" phylum="Arthropoda" rank="species" species="luederwaldti">
<emphasis id="B939D227CC74FFDFFDD8F9DEFD26D879" box="[580,710,1567,1586]" italics="true" pageId="5" pageNumber="162">L. luederwaldti</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF325EBDCC77FFDCFF0AFC94FE0ADDC8" ID-DOI="http://doi.org/10.5281/zenodo.5300981" ID-Zenodo-Dep="5300981" httpUri="https://zenodo.org/record/5300981/files/figure.png" pageId="6" pageNumber="163" startId="6.[150,187,853,872]" targetBox="[239,1036,223,827]" targetPageId="6">
<paragraph id="8BF20E35CC77FFDCFF0AFC94FE0ADDC8" blockId="6.[150,1127,853,899]" pageId="6" pageNumber="163">
Figs 15í17. Aedeagus. 15
<taxonomicName id="4C4D75B6CC77FFDCFE12FC94FCC3DD23" authority="Orchymont, 1937" authorityName="Orchymont" authorityYear="1937" box="[398,803,853,872]" class="Insecta" family="Hydrophilidae" genus="Lachnodacnum" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="163" phylum="Arthropoda" rank="species" species="luederwaldti">
<emphasis id="B939D227CC77FFDCFE12FC94FD64DD23" box="[398,644,853,872]" italics="true" pageId="6" pageNumber="163">Lachnodacnum luederwaldti</emphasis>
<bibRefCitation id="EFDC73C4CC77FFDCFD16FC94FCC3DD23" author="ORCHYMONT A." box="[650,803,853,872]" pageId="6" pageNumber="163" pagination="127" refId="ref18556" refString="ORCHYMONT A. d' 1937: Sphaeridiini bromeliadicoles nouveaux (Coleoptera: Hydrophilidae, Sphaeridiinae). Annals and Magazine of Natural History, Series 10 20: 127 i 140." type="journal article" year="1937">Orchymont, 1937</bibRefCitation>
</taxonomicName>
; 16
<taxonomicName id="4C4D75B6CC77FFDCFCC5FC94FB83DD23" authority="Orchymont, 1937" authorityName="Orchymont" authorityYear="1937" box="[857,1123,853,872]" class="Insecta" family="Hydrophilidae" genus="Lachnodacnum" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="163" phylum="Arthropoda" rank="species" species="saundersi">
<emphasis id="B939D227CC77FFDCFCC5FC94FC23DD23" box="[857,963,853,872]" italics="true" pageId="6" pageNumber="163">L. saundersi</emphasis>
<bibRefCitation id="EFDC73C4CC77FFDCFC55FC94FB83DD23" author="ORCHYMONT A." box="[969,1123,853,872]" pageId="6" pageNumber="163" pagination="127" refId="ref18556" refString="ORCHYMONT A. d' 1937: Sphaeridiini bromeliadicoles nouveaux (Coleoptera: Hydrophilidae, Sphaeridiinae). Annals and Magazine of Natural History, Series 10 20: 127 i 140." type="journal article" year="1937">Orchymont, 1937</bibRefCitation>
</taxonomicName>
; 17
<taxonomicName id="4C4D75B6CC77FFDCFF5EFCB1FE05DDC8" authority="(Scott, 1912)" authorityName="Clarkson &amp; Albertoni &amp; FIKÁýEK" authorityYear="2014" baseAuthorityName="Scott" baseAuthorityYear="1912" box="[194,485,880,899]" class="Insecta" family="Hydrophilidae" genus="Phaenostoma" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="163" phylum="Arthropoda" rank="species" species="urichi">
<emphasis id="B939D227CC77FFDCFF5EFCB1FE8EDDC8" box="[194,366,880,899]" italics="true" pageId="6" pageNumber="163">Phaenostoma urichi</emphasis>
(
<bibRefCitation id="EFDC73C4CC77FFDCFEE6FCB1FE3FDDC8" author="SCOTT H." box="[378,479,880,899]" pageId="6" pageNumber="163" pagination="424" refId="ref18616" refString="SCOTT H. 1912: A contribution to the knowledge of the fauna of Bromeliaceae. Annals and Magazine of Natural History, Series 8 10: 424 i 438 + pl. X." type="journal article" year="1912">Scott, 1912</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BF20E35CC77FFD2FF29FA3AFD0DDBA4" blockId="6.[151,1127,955,1586]" lastBlockId="8.[151,1127,1400,1615]" lastPageId="8" lastPageNumber="165" pageId="6" pageNumber="163">
<emphasis id="B939D227CC77FFDCFF29FA3AFECCD859" bold="true" box="[181,300,1531,1554]" italics="true" pageId="6" pageNumber="163">Prothorax.</emphasis>
Pronotum weakly convex, with a bead on anterior and lateral margins including antero- and posterolateral corners; anterior corners rounded, posterior corners weakly angulate; posterior margin slightly arcuate; pronotal trichobothria present sublaterally. Prosternum not carinate medially, weakly convex (Fig. 26). Procoxal cavities large, open posteriorly, anterolateral aperture of procoxal cavity open (Fig. 26). Hypomeron with large pubescent inner portion and moderately wide marginal glabrous portion.
</paragraph>
<caption id="DF325EBDCC76FFDDFFEDFA7AFF15D818" ID-DOI="http://doi.org/10.5281/zenodo.5300983" ID-Zenodo-Dep="5300983" httpUri="https://zenodo.org/record/5300983/files/figure.png" pageId="7" pageNumber="164" startId="7.[113,150,1467,1486]" targetBox="[113,1088,219,1415]" targetPageId="7">
<paragraph id="8BF20E35CC76FFDDFFEDFA7AFF15D818" blockId="7.[113,1089,1466,1619]" pageId="7" pageNumber="164">
Figs 18í23. Biology of
<taxonomicName id="4C4D75B6CC76FFDDFEDCFA7AFD34DB85" authority="Orchymont, 1937" authorityName="Orchymont" authorityYear="1937" box="[320,724,1467,1486]" class="Insecta" family="Hydrophilidae" genus="Lachnodacnum" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="164" phylum="Arthropoda" rank="species" species="luederwaldti">
<emphasis id="B939D227CC76FFDDFEDCFA7AFDD5DB85" box="[320,565,1467,1486]" italics="true" pageId="7" pageNumber="164">Lachnodacnum luederwaldti</emphasis>
<bibRefCitation id="EFDC73C4CC76FFDDFDA7FA7AFD34DB85" author="ORCHYMONT A." box="[571,724,1467,1486]" pageId="7" pageNumber="164" pagination="127" refId="ref18556" refString="ORCHYMONT A. d' 1937: Sphaeridiini bromeliadicoles nouveaux (Coleoptera: Hydrophilidae, Sphaeridiinae). Annals and Magazine of Natural History, Series 10 20: 127 i 140." type="journal article" year="1937">Orchymont, 1937</bibRefCitation>
</taxonomicName>
. 18 micropool in the rosette of
<taxonomicName id="4C4D75B6CC76FFDDFC6EFA7AFF49DBA3" class="Liliopsida" family="Bromeliaceae" genus="Neoregelia" kingdom="Plantae" order="Poales" pageId="7" pageNumber="164" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B939D227CC76FFDDFC6EFA7AFF68DBA3" italics="true" pageId="7" pageNumber="164">Neoregelia</emphasis>
sp.
</taxonomicName>
, a typical habitat; 19 egg cases attached at the inner face of a bromeliad leaf; 20 third instar larva and pupa in the detritus accumulated in the bromeliad rosette; 21í22 pupa in the broken pupal chamber (21 dorsal view; 22 ventral view); 23 adults staying submerged among bromeliad leaves in the rearing box. Figs 18í19, 23: photo B. Clarkson in Ubatuba Municipality, São Paulo; Figs 20í22: photo by F. F. Albertoni in Florianopolis, Santa Catarina.
</paragraph>
</caption>
<caption id="DF325EBDCC79FFD2FF0AFB3BFCDADB09" ID-DOI="http://doi.org/10.5281/zenodo.5300985" ID-Zenodo-Dep="5300985" httpUri="https://zenodo.org/record/5300985/files/figure.png" pageId="8" pageNumber="165" startId="8.[150,187,1274,1293]" targetBox="[151,1126,219,1256]" targetPageId="8">
<paragraph id="8BF20E35CC79FFD2FF0AFB3BFCDADB09" blockId="8.[150,1127,1273,1346]" pageId="8" pageNumber="165">
Figs 24í28. Adult morphology of
<taxonomicName id="4C4D75B6CC79FFD2FE5CFB3BFCB5DB46" authority="Orchymont, 1937" authorityName="Orchymont" authorityYear="1937" box="[448,853,1274,1293]" class="Insecta" family="Hydrophilidae" genus="Lachnodacnum" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="165" phylum="Arthropoda" rank="species" species="luederwaldti">
<emphasis id="B939D227CC79FFD2FE5CFB3BFD55DB46" box="[448,693,1274,1293]" italics="true" pageId="8" pageNumber="165">Lachnodacnum luederwaldti</emphasis>
<bibRefCitation id="EFDC73C4CC79FFD2FD27FB3BFCB5DB46" author="ORCHYMONT A." box="[699,853,1274,1293]" pageId="8" pageNumber="165" pagination="127" refId="ref18556" refString="ORCHYMONT A. d' 1937: Sphaeridiini bromeliadicoles nouveaux (Coleoptera: Hydrophilidae, Sphaeridiinae). Annals and Magazine of Natural History, Series 10 20: 127 i 140." type="journal article" year="1937">Orchymont, 1937</bibRefCitation>
</taxonomicName>
, SEM micrographs. 24 whole beetle, ventral view; 25 head, ventral view; 26 prothorax in ventral view (prosternum and hypomeron); 27 lateral portion of mesoventrite (arrow indicates the anepisternal suture); 28 antenna.
</paragraph>
</caption>
<paragraph id="8BF20E35CC79FFD3FF29FA39FDAEDB24" blockId="8.[151,1127,1400,1615]" lastBlockId="9.[113,1090,1112,1551]" lastPageId="9" lastPageNumber="166" pageId="8" pageNumber="165">
<emphasis id="B939D227CC79FFD2FF29FA39FEA0D844" bold="true" box="[181,320,1528,1551]" italics="true" pageId="8" pageNumber="165">Mesothorax.</emphasis>
Mesoventrite fused with anepisternum 2 sublaterally, anepisternal suture deeply concave, distinct medially and laterally (Figs 27, 34); epimeron 2 well delimited, divided from anepisternum 2 by a suture, pubescent; anterior collar of mesothorax broad, well demarcated. Mesoventrite highly elevated medially into preepisternal plate; widely attaching metaventral process (
<figureCitation id="137612B0CC78FFD3FEC5FBB9FE4EDAC4" box="[345,430,1144,1167]" captionStart="Figs 24" captionStartId="8.[150,187,1274,1293]" captionTargetBox="[151,1126,219,1256]" captionTargetId="figure-88@8.[151,1126,219,1257]" captionTargetPageId="8" captionText="Figs 24í28. Adult morphology of Lachnodacnum luederwaldti Orchymont, 1937, SEM micrographs. 24 whole beetle, ventral view; 25 head, ventral view; 26 prothorax in ventral view (prosternum and hypomeron); 27 lateral portion of mesoventrite (arrow indicates the anepisternal suture); 28 antenna." figureDoi="http://doi.org/10.5281/zenodo.5300985" httpUri="https://zenodo.org/record/5300985/files/figure.png" pageId="9" pageNumber="166">Figs 24</figureCitation>
,
<figureCitation id="137612B0CC78FFD3FE20FBB9FE37DAC4" box="[444,471,1144,1167]" captionStart="Figs 33" captionStartId="12.[150,187,1115,1134]" captionTargetBox="[177,1100,219,1097]" captionTargetId="figure-259@12.[177,1100,219,1097]" captionTargetPageId="12" captionText="Figs 33í36. Mesoventral elevation of Lachnodacnum Orchymont, 1937, SEM micrographs. In ventral (33í34) and lateral (35í36) views. 33, 35 L. luederwaldti Orchymont, 1937; 34, 36 L. saundersi Orchymont, 1937 (arrow indicates the anepisternal suture)." figureDoi="http://doi.org/10.5281/zenodo.5319472" httpUri="https://zenodo.org/record/5319472/files/figure.png" pageId="9" pageNumber="166">33</figureCitation>
í36); mesoventral process as high as metaventral one in lateral view; anteromedian pit-like groove large, well developed. Grooves for reception of procoxae not de¿ned. Elytron weakly convex, without longitudinal series of larger punctures (
<figureCitation id="137612B0CC78FFD3FFE5FB19FF5ADAA4" box="[121,186,1240,1263]" captionStart="Figs 1" captionStartId="4.[151,188,1467,1486]" captionTargetBox="[159,1119,224,1353]" captionTargetId="figure-10@4.[151,1120,215,1362]" captionTargetPageId="4" captionText="Figs 1í10. General habitus of adults and larvae of Lachnodacnum Orchymont, 1937. 1í3 adults of L. luederwaldti Orchymont, 1937 (1 dorsal view, 2 lateral view, 3 dorsal view of the head). 4í8 larva of L. luederwaldti (4 third instar larva in dorsal view; 5 same in ventral view; 6 ¿rst instar in dorsal view; 7 detail of head and thorax of third instar, dorsal view; 8 same, ventral view). 9í10 adults of L. saundersi Orchymont, 1937 (9 dorsal view, 10 lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5300977" httpUri="https://zenodo.org/record/5300977/files/figure.png" pageId="9" pageNumber="166">Figs 1</figureCitation>
, 9); elytral trichobothria present (Fig. 32); sutural stria very ¿ne but distinct, developed only on elytral apex (Fig. 30). Epipleuron (
<figureCitation id="137612B0CC78FFD3FDDEFB39FD71DB44" box="[578,657,1272,1295]" captionStart="Figs 24" captionStartId="8.[150,187,1274,1293]" captionTargetBox="[151,1126,219,1256]" captionTargetId="figure-88@8.[151,1126,219,1257]" captionTargetPageId="8" captionText="Figs 24í28. Adult morphology of Lachnodacnum luederwaldti Orchymont, 1937, SEM micrographs. 24 whole beetle, ventral view; 25 head, ventral view; 26 prothorax in ventral view (prosternum and hypomeron); 27 lateral portion of mesoventrite (arrow indicates the anepisternal suture); 28 antenna." figureDoi="http://doi.org/10.5281/zenodo.5300985" httpUri="https://zenodo.org/record/5300985/files/figure.png" pageId="9" pageNumber="166">Fig. 24</figureCitation>
) wide and strongly inclined throughout, reaching elytral apex; inner pubescent portion (= epipleuron) very wide anteriorly, strongly narrowing towards the level of posterior coxae. Mesocoxal cavities transverse, narrowly divided medially by meso-metaventral plate.
</paragraph>
<caption id="DF325EBDCC78FFD3FFEDFC25FC2BDA59" ID-DOI="http://doi.org/10.5281/zenodo.5300987" ID-Zenodo-Dep="5300987" httpUri="https://zenodo.org/record/5300987/files/figure.png" pageId="9" pageNumber="166" startId="9.[113,149,996,1015]" targetBox="[114,1089,219,983]" targetPageId="9">
<paragraph id="8BF20E35CC78FFD3FFEDFC25FC2BDA59" blockId="9.[113,1090,996,1042]" pageId="9" pageNumber="166">
Figs 29í32.Adult morphology of
<taxonomicName id="4C4D75B6CC78FFD3FE15FC25FCEFDDBC" authority="Orchymont, 1937" authorityName="Orchymont" authorityYear="1937" box="[393,783,996,1015]" class="Insecta" family="Hydrophilidae" genus="Lachnodacnum" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="166" phylum="Arthropoda" rank="species" species="luederwaldti">
<emphasis id="B939D227CC78FFD3FE15FC25FD97DDBC" box="[393,631,996,1015]" italics="true" pageId="9" pageNumber="166">Lachnodacnum luederwaldti</emphasis>
<bibRefCitation id="EFDC73C4CC78FFD3FDE7FC25FCEFDDBC" author="ORCHYMONT A." box="[635,783,996,1015]" pageId="9" pageNumber="166" pagination="127" refId="ref18556" refString="ORCHYMONT A. d' 1937: Sphaeridiini bromeliadicoles nouveaux (Coleoptera: Hydrophilidae, Sphaeridiinae). Annals and Magazine of Natural History, Series 10 20: 127 i 140." type="journal article" year="1937">Orchymont, 1937</bibRefCitation>
</taxonomicName>
,SEM micrographs.29 mesofemur; 30 apical portion of elytra; 31 microsculpture of the clypeus; 32 trichobothrium on the elytron.
</paragraph>
</caption>
<paragraph id="8BF20E35CC78FFD0FF0CFAB9FE70DCB8" blockId="9.[113,1090,1112,1551]" lastBlockId="10.[151,1127,220,1555]" lastPageId="10" lastPageNumber="167" pageId="9" pageNumber="166">
<emphasis id="B939D227CC78FFD3FF0CFAB9FEF4DBC4" bold="true" box="[144,276,1400,1423]" italics="true" pageId="9" pageNumber="166">Metathorax.</emphasis>
Metaventrite with pubescent surface, slightly elevated into narrow, longitudinal bare plate medially; median plate projecting anteriorly into long and narrow metaventral process joining mesoventral plate to form a meso-metaventral keel (
<figureCitation id="137612B0CC78FFD3FC89FA79FC83DB84" box="[789,867,1464,1487]" captionStart="Figs 24" captionStartId="8.[150,187,1274,1293]" captionTargetBox="[151,1126,219,1256]" captionTargetId="figure-88@8.[151,1126,219,1257]" captionTargetPageId="8" captionText="Figs 24í28. Adult morphology of Lachnodacnum luederwaldti Orchymont, 1937, SEM micrographs. 24 whole beetle, ventral view; 25 head, ventral view; 26 prothorax in ventral view (prosternum and hypomeron); 27 lateral portion of mesoventrite (arrow indicates the anepisternal suture); 28 antenna." figureDoi="http://doi.org/10.5281/zenodo.5300985" httpUri="https://zenodo.org/record/5300985/files/figure.png" pageId="9" pageNumber="166">Fig. 24</figureCitation>
), the keel constricted at the area where meso- and metaventrite join each other. Median bare portion of metaventrite with scattered punctures, punctures densely arranged in anterior half and on posterior margin; punctation of posterior half of the keel sparser than on anterior half, posterior half with weak longitudinal groove. Metepimeron rather large with long posterior projection. Hind wigs (
<figureCitation id="137612B0CC7BFFD0FBA0FF3DFF51DF78" captionStart="Figs 11" captionStartId="5.[113,150,1514,1533]" captionTargetBox="[119,1083,218,1467]" captionTargetId="figure-12@5.[119,1083,218,1483]" captionTargetPageId="5" captionText="Figs 11í14. Morphological details of Lachnodacnum Orchymont, 1937 and Phaenostoma Orchymont, 1937. 11í12 hind wings (11 Lachnodacnum luederwaldti Orchymont, 1937; 12 Phaenostoma urichi (Scott, 1912)). 13í14 detail of spiracular atrium of the third instar larva of L. luederwaldti." figureDoi="http://doi.org/10.5281/zenodo.5300979" httpUri="https://zenodo.org/record/5300979/files/figure.png" pageId="10" pageNumber="167">Fig. 11</figureCitation>
) with R-M loop distally the maximum width of the wing; RA 3+4 not attached to RA basally, forming a opened radial cell; radial cell large, subtrapezoidal, pigmented on anteroapical half; r3 absent; r4 developed, connecting RA 3+4 to R-M loop; apical ¿eld with RA 3 well de¿ned, short, not reaching the margin of the wing apically; RA 4 fused with RP 1, together forming a pigmented area almost reaching wing margin apically; base of RP 1 far of R-M loop; RP 2 forming a pigmented area wide on base, almost reaching R-M loop basally but not reaching edge of the wing apically; RP 3+4 well de¿ned, narrow and long, arising from R-M loop basally, curved posteriad and reaching wing margin apically; medial ¿eld with medial spur rather long, narrow and sharp; MP3+4 present, weakly developed, nearly connected with MP 1+2, not attached to MP 3 and MP 4 +CuA 1; MP 3 and MP 4 +CuA 1 well de¿ned, not connected to basal portions of the veins proximally, reaching wing margin posteriorly; CuA 1 strongly reduced, without connection with MP 3+4, MP 3 and MP 4 +CuA 1; CuA 2 present and well de¿ned, reaching the posterior edge of the wing apically; CuA 3+4 absent and hence wedge cell missing; AA 3 and AA 4 strongly de¿ned and long, nearly reaching posterior margin of the wing; anal ¿eld with venation strongly reduced, composed of a short AP
</paragraph>
<paragraph id="8BF20E35CC7BFFD0FE0CFD2DFE48DCB1" blockId="10.[151,1127,220,1555]" box="[400,424,748,762]" pageId="10" pageNumber="167">3+4</paragraph>
<paragraph id="8BF20E35CC7BFFD0FE32FD1DFD3DDCB8" blockId="10.[151,1127,220,1555]" box="[430,733,732,755]" pageId="10" pageNumber="167">and nearly inconspicuous AP</paragraph>
<paragraph id="8BF20E35CC7BFFD0FD41FD2DFD16DCB1" blockId="10.[151,1127,220,1555]" box="[733,758,748,762]" pageId="10" pageNumber="167">1+2</paragraph>
<paragraph id="8BF20E35CC7BFFD0FD69FD1DFD1CDCB8" blockId="10.[151,1127,220,1555]" box="[757,764,732,755]" pageId="10" pageNumber="167">.</paragraph>
<paragraph id="8BF20E35CC7BFFD0FF29FD3DFC99DB78" blockId="10.[151,1127,220,1555]" pageId="10" pageNumber="167">
<emphasis id="B939D227CC7BFFD0FF29FD3DFF10DD58" bold="true" box="[181,240,764,787]" pageId="10" pageNumber="167">
<emphasis id="B939D227CC7BFFD0FF29FD3DFF08DD58" bold="true" box="[181,232,764,787]" italics="true" pageId="10" pageNumber="167">Legs</emphasis>
.
</emphasis>
Procoxa globular, pubescent; profemur with deep tibial groove delimited by low ventral and high dorsal ridges, ventral surface pubescent except for small bare distal area; protibia cylindrical, bearing series of stout spines and irregularly arranged small spines, distal portion bearing “crown” of smaller stout spines and two large spurs. Mesotrochanter slightly sinuate on posterior margin; ventral surface of mesofemur pubescent in basal 0.75 (
<figureCitation id="137612B0CC7BFFD0FB97FCBDFBBADDD8" box="[1035,1114,892,915]" captionStart="Figs 29" captionStartId="9.[113,149,996,1015]" captionTargetBox="[114,1089,219,983]" captionTargetId="figure-186@9.[114,1090,219,983]" captionTargetPageId="9" captionText="Figs 29í32.Adult morphology of Lachnodacnum luederwaldti Orchymont, 1937,SEM micrographs.29 mesofemur; 30 apical portion of elytra; 31 microsculpture of the clypeus; 32 trichobothrium on the elytron." figureDoi="http://doi.org/10.5281/zenodo.5300987" httpUri="https://zenodo.org/record/5300987/files/figure.png" pageId="10" pageNumber="167">Fig. 29</figureCitation>
), reticulate; tibial groove deep, developed throughout, delimited by high ventral and low dorsal ridges; mesotibia flattened, sparsely covered by short and stout spines, bearing longitudinal series of larger spines and a subapical transverse series of larger spines; distal apex with series of short spines and two long spurs on inner margin. Metatrochanter slightly sinuate on posterior margin, pubescent; metafemur large and wide, ventral surface bare except of sparsely arranged setae on extreme anteroproximal portion, surface reticulate with scattered setiferous punctures; tibial groove well developed, deep, delimited by high ventral and low dorsal ridges. Metatibia flattened, bearing scattered short spines and two series of larger spines on outer margin, and a series of larger spines on inner margin on ventral surface; distal portion with transverse series of moderately long spines and two long spurs on inner margin. Tarsi much shorter than tibiae, each tarsomere bearing a dense brush of long setae ventrally, with longer setae sideward and with a single to few long setae dorsally. Metatarsus slightly compressed laterally, metatarsomere 1 about twice as long as metatarsomere 2; claws small, simply arcuate.
</paragraph>
<paragraph id="8BF20E35CC7BFFD0FF29FAFDFC32DB38" blockId="10.[151,1127,220,1555]" pageId="10" pageNumber="167">
<emphasis id="B939D227CC7BFFD0FF29FAFDFEC6DB18" bold="true" box="[181,294,1340,1363]" pageId="10" pageNumber="167">
<emphasis id="B939D227CC7BFFD0FF29FAFDFEFEDB18" bold="true" box="[181,286,1340,1363]" italics="true" pageId="10" pageNumber="167">Abdomen</emphasis>
.
</emphasis>
With ¿ve ventrites, all ventrites flat, without any longitudinal carina; posterior margin of ventrite 5 without apical notch nor the median group of stout setae.
</paragraph>
<paragraph id="8BF20E35CC7BFFD0FF29FABDFDFDD858" blockId="10.[151,1127,220,1555]" pageId="10" pageNumber="167">
<emphasis id="B939D227CC7BFFD0FF29FABDFEBBDBD8" bold="true" box="[181,347,1404,1427]" italics="true" pageId="10" pageNumber="167">Male genitalia.</emphasis>
Aedeagus (
<figureCitation id="137612B0CC7BFFD0FE4BFABDFDC8DBD8" box="[471,552,1404,1427]" captionStart="Figs 15" captionStartId="6.[150,187,853,872]" captionTargetBox="[239,1036,223,827]" captionTargetId="figure-267@6.[224,1053,212,830]" captionTargetPageId="6" captionText="Figs 15í17. Aedeagus. 15 Lachnodacnum luederwaldti Orchymont, 1937; 16 L. saundersi Orchymont, 1937; 17 Phaenostoma urichi (Scott, 1912)." figureDoi="http://doi.org/10.5281/zenodo.5300981" httpUri="https://zenodo.org/record/5300981/files/figure.png" pageId="10" pageNumber="167">Figs 15</figureCitation>
í16) with rather long and well developed symmetrical phallobase; parameres wide throughout; median lobe wide basally, with a pair of short basal projections, more or less gradually narrowing apicad, gonopore distinct, subapical. Sternite 9 wide basally, bearing rather narrow tongue-shaped median projection, lateral struts arcuate, slightly shorter than median portion.
</paragraph>
</subSubSection>
</treatment>
</document>