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<document id="8319376858A4689ECD55B8DECCFB0870" ID-CLB-Dataset="291999" ID-DOI="10.25674/so95iss1id311" ID-GBIF-Dataset="06b03cc4-e211-4f65-be07-742740b0053c" ID-ISSN="2509-9523" ID-Zenodo-Dep="10724236" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1709147239332" checkinUser="felipe" docAuthor="Frisch, Johannes &amp; Narakusumo, Raden Pramesa" docDate="2022" docId="03B7C77AFFE3FFD0BC8FFD820F1F0796" docLanguage="en" docName="SoilOrg.95.1.23-73.pdf" docOrigin="Soil Organisms 95 (1)" docSource="http://dx.doi.org/10.25674/so95iss1id311" docStyle="DocumentStyle:EA2ECAC87EC21C07AB805D5B9B4601F2.2:SoilOrg.2018-.journal_article" docStyleId="EA2ECAC87EC21C07AB805D5B9B4601F2" docStyleName="SoilOrg.2018-.journal_article" docStyleVersion="2" docTitle="Scopaeus cuspidatus Frisch 2022, spec. nov." docType="treatment" docVersion="4" lastPageNumber="59" masterDocId="FF8EBF02FFC0FFF4BC5BFFF80D1D040D" masterDocTitle="Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species" masterLastPageNumber="73" masterPageNumber="23" pageNumber="58" updateTime="1711467745701" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="58B4211EFCA4AD3EB0C68C0C40AE5B68" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="10AD7852775655ADAF7EAE1D8F643F9C">Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species</mods:title>
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<mods:namePart id="FD2B4765017810681F660CC14843934A">Frisch, Johannes</mods:namePart>
</mods:name>
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<mods:role id="F20C249AF0D0605F10F2E964025BCBDA">
<mods:roleTerm id="36DB51F5F0BB651A0253FE6BAC45322C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8BFC4CFC9DF492348720B7EB8D7FFF41">Narakusumo, Raden Pramesa</mods:namePart>
<mods:affiliation id="8134EC8BA80A3420220007C90528175F">MUseUm ZoologicUm Bogoriense, Research Center for Biosystematics and EvolUtion, National Research and Innovation Agency (BRIN), Jl. Raya Jakarta-Bogor KM 46, Cibinong, West Java, 16911, Indonesia</mods:affiliation>
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<mods:title id="CE45EC176AC12D8E2CE14C9FDBC79245">Soil Organisms</mods:title>
</mods:titleInfo>
<mods:part id="3DFEF5473513CDDFED454C595F5C33CF">
<mods:date id="3EFFA4B8EABA899BC2A2BCD1305C0ABC">2022</mods:date>
<mods:detail id="1DFD7F8811CB9B51F6225CC0DD4FC5B4" type="pubDate">
<mods:number id="44E6FBA8FDEAF14281668519A3E88B7F">2022-04-01</mods:number>
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<mods:number id="BC218E2F94C0992161D248850AC4F9D5">95</mods:number>
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<mods:classification id="193B75F1C982D398DC7074B557400592">journal article</mods:classification>
<mods:identifier id="97303D020E2F3F087659B0F714E67E9A" type="CLB-Dataset">291999</mods:identifier>
<mods:identifier id="8026769BB9FA5D189DFB69703F670D38" type="DOI">10.25674/so95iss1id311</mods:identifier>
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<mods:identifier id="EA69EEAC51AB5914DD409A7741EB134F" type="ISSN">2509-9523</mods:identifier>
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<treatment id="03B7C77AFFE3FFD0BC8FFD820F1F0796" ID-DOI="http://doi.org/10.5281/zenodo.10724327" ID-Zenodo-Dep="10724327" LSID="urn:lsid:plazi:treatment:03B7C77AFFE3FFD0BC8FFD820F1F0796" httpUri="http://treatment.plazi.org/id/03B7C77AFFE3FFD0BC8FFD820F1F0796" lastPageId="36" lastPageNumber="59" pageId="35" pageNumber="58">
<subSubSection id="C30425E7FFE3FFD7BC8FFD820FB60698" box="[212,683,634,661]" pageId="35" pageNumber="58" type="nomenclature">
<paragraph id="8BA1766CFFE3FFD7BC8FFD820FB60698" blockId="35.[212,683,634,694]" box="[212,683,634,661]" pageId="35" pageNumber="58">
<heading id="D0E9C100FFE3FFD7BC8FFD820FB60698" box="[212,683,634,661]" fontSize="11" level="2" pageId="35" pageNumber="58" reason="5">
<taxonomicName id="4C1E0DEFFFE3FFD7BC8FFD820F370699" authority="Frisch" authorityName="Frisch" authorityYear="2022" box="[212,554,634,661]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="58" phylum="Arthropoda" rank="species" species="cuspidatus" status="sp. nov.">
<emphasis id="B96AAA7EFFE3FFD7BC8FFD820F370699" bold="true" box="[212,554,634,661]" pageId="35" pageNumber="58">
<emphasis id="B96AAA7EFFE3FFD7BC8FFD820CCC0698" bold="true" box="[212,465,634,661]" italics="true" pageId="35" pageNumber="58">Scopaeus cuspidatus</emphasis>
Frisch
</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2591705FFE3FFD7BE63FD830FB60698" box="[568,683,635,661]" pageId="35" pageNumber="58" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFE3FFD7BC8FFD630F9D06BB" box="[212,640,667,694]" pageId="35" pageNumber="58" type="description">
<paragraph id="8BA1766CFFE3FFD7BC8FFD630F9D06BB" blockId="35.[212,683,634,694]" box="[212,640,667,694]" pageId="35" pageNumber="58">
(
<figureCitation id="13256AE9FFE3FFD7BC87FD630C2806B8" box="[220,309,667,693]" captionStart="Figures 23–26" captionStartId="14.[151,224,1930,1952]" captionTargetBox="[263,1352,435,1899]" captionTargetId="figure-131@14.[151,1398,422,1911]" captionTargetPageId="14" captionText="Figures 23–26. Habitus of Scopaeus cuspidatus, paratype, Jawa Barat: Mt Salak (23); S. sulawesianus, paratype, Sulawesi Tengah: Tambarana (24); S. niger, paralectotype, Singapore (25); S. borneensis, holotype, Kalimantan Barat: Pontianak (26)." figureDoi="http://doi.org/10.5281/zenodo.10724252" httpUri="https://zenodo.org/record/10724252/files/figure.png" pageId="35" pageNumber="58">Figs 23</figureCitation>
,
<figureCitation id="13256AE9FFE3FFD7BD1AFD630C4306B8" box="[321,350,667,693]" captionStart="Figures 42–49" captionStartId="17.[190,263,1886,1908]" captionTargetBox="[263,1341,889,1832]" captionTargetId="figure-379@17.[189,1436,851,1867]" captionTargetPageId="17" captionText="Figures 42–49. Aedeagus in lateral view of Scopaeus velifer, holotype, Banten, Lebak: Citorek Kidul (42); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (43); S. uncinatus, holotype,Banten, Lebak:Citorek Kidul (44); S. bipenicillatus, holotype,Sulawesi Tengah, Sigi: Pagana (45); S. bipectenatus, holotype, Sulawesi Tengah, Poso: Lake Poso (46); S. cuspidatus, holotype, Bali, Jembrana: Cekik (47); S. crassipunctatus, holotype, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (48); S. sulawesianus, holotype, Sulawesi Selatan: Engrekang (49)." figureDoi="http://doi.org/10.5281/zenodo.10724262" httpUri="https://zenodo.org/record/10724262/files/figure.png" pageId="35" pageNumber="58">47</figureCitation>
,
<figureCitation id="13256AE9FFE3FFD7BD31FD630CCE06BB" box="[362,467,667,694]" captionStart="Figures 113–115" captionStartId="35.[189,262,1908,1930]" captionTargetBox="[261,1322,1006,1857]" captionTargetId="figure-356@35.[189,1436,978,1889]" captionTargetPageId="35" captionText="Figures 113–115. Aedeagus of Scopaeus cuspidatus, holotype, Bali, Jembrana: Cekik in lateral (113), ventral (114), dorsal view (115). Abbreviations: al – apical lobes, dw – dorsodistal window of phallobase, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10724298" httpUri="https://zenodo.org/record/10724298/files/figure.png" pageId="35" pageNumber="58">113 – 115</figureCitation>
, 142, 143, 176)
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFE3FFD7BC8FFD270FF707B1" pageId="35" pageNumber="58" type="materials_examined">
<paragraph id="8BA1766CFFE3FFD7BC8FFD270FF707B1" blockId="35.[189,799,734,956]" pageId="35" pageNumber="58">
<emphasis id="B96AAA7EFFE3FFD7BC8FFD270CBC06FB" bold="true" box="[212,417,734,759]" pageId="35" pageNumber="58">Type specimens:</emphasis>
<materialsCitation id="3B767C31FFE3FFD7BDE3FD270C3F0734" collectionCode="PWCM" collectorName="Wunderle" country="Indonesia" elevation="1994300" location="Cekik" municipality="Jembrana" pageId="35" pageNumber="58" specimenCount="1" specimenCount-male="1" stateProvince="Bali" typeStatus="holotype">
<collectingCountry id="F30936FCFFE3FFD7BDE3FD270F3D06FA" box="[440,544,735,759]" name="Indonesia" pageId="35" pageNumber="58">Indonesia</collectingCountry>
:
<typeStatus id="54A5C8CEFFE3FFD7BE1BFD270FB906FA" box="[576,676,735,759]" pageId="35" pageNumber="58" type="holotype">Holotype</typeStatus>
<specimenCount id="9D18BDE5FFE3FFD7BEE7FD270FCD06FB" box="[700,720,735,758]" count="1" pageId="35" pageNumber="58" type="male">♂</specimenCount>
,
<collectingRegion id="49DAB88EFFE3FFD7BEB7FD270E0606FA" box="[748,795,735,759]" country="Indonesia" name="Bali" pageId="35" pageNumber="58">Bali</collectingRegion>
,
<collectingMunicipality id="6BC5EC16FFE3FFD7BCE6FCF80C340715" box="[189,297,768,792]" pageId="35" pageNumber="58">Jembrana</collectingMunicipality>
:
<location id="8EC120B7FFE3FFD7BD66FCF80C9C0715" LSID="urn:lsid:plazi:treatment:03B7C77AFFE3FFD0BC8FFD820F1F0796:8EC120B7FFE3FFD7BD66FCF80C9C0715" box="[317,385,768,792]" country="Indonesia" municipality="Jembrana" name="Cekik" pageId="35" pageNumber="58" stateProvince="Bali">Cekik</location>
,
<date id="FFA050ACFFE3FFD7BDCCFD070F1C071A" box="[407,513,767,791]" pageId="35" pageNumber="58" value="1994-06-20">
20.6.
<quantity id="4CE6DB89FFE3FFD7BD90FD070F75071A" box="[459,616,767,792]" metricMagnitude="6" metricUnit="m" metricValue="1.9943" pageId="35" pageNumber="58" unit="m" value="1994300.0">
<elevation id="0033915FFFE3FFD7BD90FD070F75071A" box="[459,616,767,792]" metricMagnitude="6" metricUnit="m" metricValue="1.9943" pageId="35" pageNumber="58" unit="m" value="1994300.0">1994, 300 m</elevation>
</quantity>
</date>
, leg.
<collectorName id="26EB13BAFFE3FFD7BEEDFCF80E020715" box="[694,799,768,792]" pageId="35" pageNumber="58">Wunderle</collectorName>
(
<collectionCode id="ED0FEEA9FFE3FFD7BC9FFCD90C060734" box="[196,283,801,825]" pageId="35" pageNumber="58">PWCM</collectionCode>
)
</materialsCitation>
.
<typeStatus id="54A5C8CEFFE3FFD7BD67FCD90CBA0734" box="[316,423,801,825]" pageId="35" pageNumber="58" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D18BDE5FFE3FFD7BD9FFCD90F400735" box="[452,605,800,825]" count="2" pageId="35" pageNumber="58" type="generic">2 specimens</specimenCount>
):
<materialsCitation id="3B767C31FFE3FFD7BE26FCD90FF10777" collectingDate="2017-12-05" collectionCode="MFNB" collectorName="Frisch" country="Indonesia" elevation="430" latitude="-2.1016667" location="Beteleme - Tinompo" longLatPrecision="21" longitude="121.14056" municipality="Morowali" pageId="35" pageNumber="58" specimenCount="1" specimenCount-male="1" stateProvince="Sulawesi Tengah" typeStatus="paratype">
<specimenCount id="9D18BDE5FFE3FFD7BE26FCD90FA90735" box="[637,692,801,825]" count="1" pageId="35" pageNumber="58" type="male">1 ♂</specimenCount>
,
<collectingRegion id="49DAB88EFFE3FFD7BE94FCD80C3D0754" country="Indonesia" name="Sulawesi Tengah" pageId="35" pageNumber="58">Central Sulawesi</collectingRegion>
,
<collectingMunicipality id="6BC5EC16FFE3FFD7BD77FCB90C840754" box="[300,409,833,857]" pageId="35" pageNumber="58">Morowali</collectingMunicipality>
:
<location id="8EC120B7FFE3FFD7BDFFFCB90F950754" LSID="urn:lsid:plazi:treatment:03B7C77AFFE3FFD0BC8FFD820F1F0796:8EC120B7FFE3FFD7BDFFFCB90F950754" box="[420,648,833,857]" country="Indonesia" latitude="-2.1016667" longLatPrecision="21" longitude="121.14056" municipality="Morowali" name="Beteleme - Tinompo" pageId="35" pageNumber="58" stateProvince="Sulawesi Tengah">Beteleme – Tinompo</location>
(
<geoCoordinate id="EE2A10ABFFE3FFD7BEC3FCB90E060754" box="[664,795,833,857]" degrees="02" direction="south" minutes="06" orientation="latitude" pageId="35" pageNumber="58" precision="15" seconds="06" value="-2.1016667">02°06’06’’S</geoCoordinate>
,
<geoCoordinate id="EE2A10ABFFE3FFD7BCE6FC9A0C510777" box="[189,332,866,890]" degrees="121" direction="east" minutes="08" orientation="longitude" pageId="35" pageNumber="58" precision="15" seconds="26" value="121.14056">121°08’26’’E</geoCoordinate>
),
<quantity id="4CE6DB89FFE3FFD7BD07FC9A0C820777" box="[348,415,866,890]" metricMagnitude="2" metricUnit="m" metricValue="4.3" pageId="35" pageNumber="58" unit="m" value="430.0">
<elevation id="0033915FFFE3FFD7BD07FC9A0C820777" box="[348,415,866,890]" metricMagnitude="2" metricUnit="m" metricValue="4.3" pageId="35" pageNumber="58" unit="m" value="430.0">430 m</elevation>
</quantity>
,
<date id="FFA050ACFFE3FFD7BDF3FC9A0F120777" box="[424,527,866,890]" pageId="35" pageNumber="58" value="2017-12-05">
<collectingDate id="EFE4A944FFE3FFD7BDF3FC9A0F120777" box="[424,527,866,890]" pageId="35" pageNumber="58" value="2017-12-05">12.5.2017</collectingDate>
</date>
, leg.
<collectorName id="26EB13BAFFE3FFD7BE18FC9A0F9B0777" box="[579,646,866,890]" pageId="35" pageNumber="58">Frisch</collectorName>
(
<collectionCode id="ED0FEEA9FFE3FFD7BEC9FC9A0FF90777" box="[658,740,866,890]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="35" pageNumber="58">MFNB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B767C31FFE3FFD7BEAFFC9A0FFB07B1" collectingDate="2016-09-13" collectionCode="MZB" collectorName="Frisch" country="Indonesia" elevation="640" latitude="-6.665278" location="Mt Salak" longLatPrecision="21" longitude="106.76" municipality="Bogor" pageId="35" pageNumber="58" specimenCount="1" specimenCount-female="1" stateProvince="Jawa Barat" typeStatus="paratype">
<specimenCount id="9D18BDE5FFE3FFD7BEAFFC9A0E040777" box="[756,793,866,890]" count="1" pageId="35" pageNumber="58" type="female">1 ♀</specimenCount>
,
<collectingRegion id="49DAB88EFFE3FFD7BCE6FC7C0C2B0796" box="[189,310,900,923]" country="Indonesia" name="Jawa Barat" pageId="35" pageNumber="58">Jawa Barat</collectingRegion>
, NO
<collectingMunicipality id="6BC5EC16FFE3FFD7BD36FC7C0CAE0791" box="[365,435,900,924]" pageId="35" pageNumber="58">Bogor</collectingMunicipality>
: NO-slope
<location id="8EC120B7FFE3FFD7BE77FC7C0F900796" LSID="urn:lsid:plazi:treatment:03B7C77AFFE3FFD0BC8FFD820F1F0796:8EC120B7FFE3FFD7BE77FC7C0F900796" box="[556,653,899,923]" country="Indonesia" latitude="-6.665278" longLatPrecision="21" longitude="106.76" municipality="Bogor" name="Mt Salak" pageId="35" pageNumber="58" stateProvince="Jawa Barat">Mt Salak</location>
(
<geoCoordinate id="EE2A10ABFFE3FFD7BEC7FC7B0E070796" box="[668,794,899,923]" degrees="06" direction="south" minutes="39" orientation="latitude" pageId="35" pageNumber="58" precision="15" seconds="55" value="-6.665278">06°39’55’’S</geoCoordinate>
,
<geoCoordinate id="EE2A10ABFFE3FFD7BCE6FC5C0C5707B1" box="[189,330,932,956]" degrees="106" direction="east" minutes="45" orientation="longitude" pageId="35" pageNumber="58" precision="15" seconds="36" value="106.76">106°45’36’’E</geoCoordinate>
),
<quantity id="4CE6DB89FFE3FFD7BD06FC5C0CBE07B6" box="[349,419,932,956]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="35" pageNumber="58" unit="m" value="640.0">
<elevation id="0033915FFFE3FFD7BD06FC5C0CBE07B6" box="[349,419,932,956]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="35" pageNumber="58" unit="m" value="640.0">640 m</elevation>
</quantity>
,
<date id="FFA050ACFFE3FFD7BDF4FC5C0F0B07B1" box="[431,534,932,956]" pageId="35" pageNumber="58" value="2016-09-13">
<collectingDate id="EFE4A944FFE3FFD7BDF4FC5C0F0B07B1" box="[431,534,932,956]" pageId="35" pageNumber="58" value="2016-09-13">13.9.2016</collectingDate>
</date>
, leg.
<collectorName id="26EB13BAFFE3FFD7BE16FC5C0F8D07B1" box="[589,656,932,956]" pageId="35" pageNumber="58">Frisch</collectorName>
(
<collectionCode id="ED0FEEA9FFE3FFD7BEC4FC5C0FC307B1" box="[671,734,932,956]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="35" pageNumber="58" type="Museum">MZB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFE3FFD0BF0AFF090C7106B8" lastPageId="36" lastPageNumber="59" pageId="35" pageNumber="58" type="description">
<paragraph id="8BA1766CFFE3FFD7BF0AFF090812063C" blockId="35.[826,1437,241,956]" pageId="35" pageNumber="58">
<emphasis id="B96AAA7EFFE3FFD7BF0AFF090EC20504" bold="true" box="[849,991,241,265]" pageId="35" pageNumber="58">Description:</emphasis>
Habitus and coloring as in
<figureCitation id="13256AE9FFE3FFD7B919FF0908840507" box="[1346,1433,241,266]" captionStart="Figures 23–26" captionStartId="14.[151,224,1930,1952]" captionTargetBox="[263,1352,435,1899]" captionTargetId="figure-131@14.[151,1398,422,1911]" captionTargetPageId="14" captionText="Figures 23–26. Habitus of Scopaeus cuspidatus, paratype, Jawa Barat: Mt Salak (23); S. sulawesianus, paratype, Sulawesi Tengah: Tambarana (24); S. niger, paralectotype, Singapore (25); S. borneensis, holotype, Kalimantan Barat: Pontianak (26)." figureDoi="http://doi.org/10.5281/zenodo.10724252" httpUri="https://zenodo.org/record/10724252/files/figure.png" pageId="35" pageNumber="58">Fig. 23</figureCitation>
. Head trapezoidal with concave posterior margin. Penultimate antennal segment transverse. Mesotibia moderately thickened. Forebody surface with relatively coarse, spacious, setose punctation; microreticulation absent; body surface shiny. Pubescence of body surface short, decumbent, without conspicuous macrosetae. Forebody unicolorous orange brown; abdomen light brown; appendages yellow-brown. Total body length
<quantity id="4CE6DB89FFE3FFD7BF61FDE20EDB063C" box="[826,966,537,562]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.5" metricValueMin="2.3" pageId="35" pageNumber="58" unit="mm" value="2.4" valueMax="2.5" valueMin="2.3">2.3 – 2.5 mm</quantity>
; forebody length
<quantity id="4CE6DB89FFE3FFD7B8D9FDE20816063C" box="[1154,1291,538,562]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="35" pageNumber="58" unit="mm" value="1.25" valueMax="1.3" valueMin="1.2">1.2 – 1.3 mm</quantity>
.
</paragraph>
<paragraph id="8BA1766CFFE3FFD0BF0AFDC20FD3063F" blockId="35.[826,1437,241,956]" lastBlockId="36.[151,762,241,923]" lastPageId="36" lastPageNumber="59" pageId="35" pageNumber="58">
Male: Posterior margin of abdominal sternite VII extended in narrow, tapered, median lobe (Fig. 142). Abdominal sternite VIII with short, narrow, triangular emargination in posterior eighth and basal margin in median third notably concave, almost reaching subbasal ridge (Fig. 143). Aedeagus (
<figureCitation id="13256AE9FFE3FFD7B8D9FD2709C706FA" box="[1154,1242,735,759]" captionStart="Figures 42–49" captionStartId="17.[190,263,1886,1908]" captionTargetBox="[263,1341,889,1832]" captionTargetId="figure-379@17.[189,1436,851,1867]" captionTargetPageId="17" captionText="Figures 42–49. Aedeagus in lateral view of Scopaeus velifer, holotype, Banten, Lebak: Citorek Kidul (42); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (43); S. uncinatus, holotype,Banten, Lebak:Citorek Kidul (44); S. bipenicillatus, holotype,Sulawesi Tengah, Sigi: Pagana (45); S. bipectenatus, holotype, Sulawesi Tengah, Poso: Lake Poso (46); S. cuspidatus, holotype, Bali, Jembrana: Cekik (47); S. crassipunctatus, holotype, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (48); S. sulawesianus, holotype, Sulawesi Selatan: Engrekang (49)." figureDoi="http://doi.org/10.5281/zenodo.10724262" httpUri="https://zenodo.org/record/10724262/files/figure.png" pageId="35" pageNumber="58">Figs 47</figureCitation>
,
<figureCitation id="13256AE9FFE3FFD7B8B7FD27085606FA" box="[1260,1355,735,759]" captionStart="Figures 113–115" captionStartId="35.[189,262,1908,1930]" captionTargetBox="[261,1322,1006,1857]" captionTargetId="figure-356@35.[189,1436,978,1889]" captionTargetPageId="35" captionText="Figures 113–115. Aedeagus of Scopaeus cuspidatus, holotype, Bali, Jembrana: Cekik in lateral (113), ventral (114), dorsal view (115). Abbreviations: al – apical lobes, dw – dorsodistal window of phallobase, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10724298" httpUri="https://zenodo.org/record/10724298/files/figure.png" pageId="35" pageNumber="58">113 – 115</figureCitation>
) about
<quantity id="4CE6DB89FFE3FFD7BF61FD070E96071A" box="[826,907,767,791]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="35" pageNumber="58" unit="mm" value="0.5">0.5 mm</quantity>
long, strongly sclerotized, thus dark reddish brown; phallobase compact, with large, distad projecting dorsodistal window (
<figureCitation id="13256AE9FFE3FFD7B870FCB909630754" box="[1067,1150,833,857]" captionStart="Figures 42–49" captionStartId="17.[190,263,1886,1908]" captionTargetBox="[263,1341,889,1832]" captionTargetId="figure-379@17.[189,1436,851,1867]" captionTargetPageId="17" captionText="Figures 42–49. Aedeagus in lateral view of Scopaeus velifer, holotype, Banten, Lebak: Citorek Kidul (42); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (43); S. uncinatus, holotype,Banten, Lebak:Citorek Kidul (44); S. bipenicillatus, holotype,Sulawesi Tengah, Sigi: Pagana (45); S. bipectenatus, holotype, Sulawesi Tengah, Poso: Lake Poso (46); S. cuspidatus, holotype, Bali, Jembrana: Cekik (47); S. crassipunctatus, holotype, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (48); S. sulawesianus, holotype, Sulawesi Selatan: Engrekang (49)." figureDoi="http://doi.org/10.5281/zenodo.10724262" httpUri="https://zenodo.org/record/10724262/files/figure.png" pageId="35" pageNumber="58">Fig. 47</figureCitation>
,
<figureCitation id="13256AE9FFE3FFD7B8CBFCB909AA0754" box="[1168,1207,833,857]" captionStart="Figures 113–115" captionStartId="35.[189,262,1908,1930]" captionTargetBox="[261,1322,1006,1857]" captionTargetId="figure-356@35.[189,1436,978,1889]" captionTargetPageId="35" captionText="Figures 113–115. Aedeagus of Scopaeus cuspidatus, holotype, Bali, Jembrana: Cekik in lateral (113), ventral (114), dorsal view (115). Abbreviations: al – apical lobes, dw – dorsodistal window of phallobase, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10724298" httpUri="https://zenodo.org/record/10724298/files/figure.png" pageId="35" pageNumber="58">113</figureCitation>
), strongly narrowed towards long, slender, apicad tapered distal portion of aedeagus with narrow, short apical lobes each with ventroapical widening studded with longitudinal row of about six stout setae followed proximally by weakly sclerotized, broad, longitudinal enlargement with acute ventroproximal end (
<figureCitation id="13256AE9FFE4FFD0BDD1FECB0F370546" box="[394,554,307,331]" captionStart="Figures 113–115" captionStartId="35.[189,262,1908,1930]" captionTargetBox="[261,1322,1006,1857]" captionTargetId="figure-356@35.[189,1436,978,1889]" captionTargetPageId="35" captionText="Figures 113–115. Aedeagus of Scopaeus cuspidatus, holotype, Bali, Jembrana: Cekik in lateral (113), ventral (114), dorsal view (115). Abbreviations: al – apical lobes, dw – dorsodistal window of phallobase, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10724298" httpUri="https://zenodo.org/record/10724298/files/figure.png" pageId="36" pageNumber="59">Figs 113, 114</figureCitation>
); dorsal lobe and flagellum inconspicuous; ventral lobe beginning with wide, triangular, proximal enlargement projecting from phallobase, then weakly sclerotized distad, very thin in ventral view (
<figureCitation id="13256AE9FFE4FFD0BD75FE4F0C9505C2" box="[302,392,439,463]" captionStart="Figures 113–115" captionStartId="35.[189,262,1908,1930]" captionTargetBox="[261,1322,1006,1857]" captionTargetId="figure-356@35.[189,1436,978,1889]" captionTargetPageId="35" captionText="Figures 113–115. Aedeagus of Scopaeus cuspidatus, holotype, Bali, Jembrana: Cekik in lateral (113), ventral (114), dorsal view (115). Abbreviations: al – apical lobes, dw – dorsodistal window of phallobase, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10724298" httpUri="https://zenodo.org/record/10724298/files/figure.png" pageId="36" pageNumber="59">Fig. 114</figureCitation>
), in lateral view straight ventrally and evenly narrowed towards base of apical lobes (
<figureCitation id="13256AE9FFE4FFD0BE91FE200DAF061C" captionStart="Figures 42–49" captionStartId="17.[190,263,1886,1908]" captionTargetBox="[263,1341,889,1832]" captionTargetId="figure-379@17.[189,1436,851,1867]" captionTargetPageId="17" captionText="Figures 42–49. Aedeagus in lateral view of Scopaeus velifer, holotype, Banten, Lebak: Citorek Kidul (42); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (43); S. uncinatus, holotype,Banten, Lebak:Citorek Kidul (44); S. bipenicillatus, holotype,Sulawesi Tengah, Sigi: Pagana (45); S. bipectenatus, holotype, Sulawesi Tengah, Poso: Lake Poso (46); S. cuspidatus, holotype, Bali, Jembrana: Cekik (47); S. crassipunctatus, holotype, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (48); S. sulawesianus, holotype, Sulawesi Selatan: Engrekang (49)." figureDoi="http://doi.org/10.5281/zenodo.10724262" httpUri="https://zenodo.org/record/10724262/files/figure.png" pageId="36" pageNumber="59">Figs 47</figureCitation>
,
<figureCitation id="13256AE9FFE4FFD0BC9EFE010DF1061C" box="[197,236,505,529]" captionStart="Figures 113–115" captionStartId="35.[189,262,1908,1930]" captionTargetBox="[261,1322,1006,1857]" captionTargetId="figure-356@35.[189,1436,978,1889]" captionTargetPageId="35" captionText="Figures 113–115. Aedeagus of Scopaeus cuspidatus, holotype, Bali, Jembrana: Cekik in lateral (113), ventral (114), dorsal view (115). Abbreviations: al – apical lobes, dw – dorsodistal window of phallobase, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10724298" httpUri="https://zenodo.org/record/10724298/files/figure.png" pageId="36" pageNumber="59">113</figureCitation>
); median foramen small, limited distally by ventrally projecting, transverse ridge (
<figureCitation id="13256AE9FFE4FFD0BE68FDE10FDE063F" box="[563,707,537,562]" captionStart="Figures 113–115" captionStartId="35.[189,262,1908,1930]" captionTargetBox="[261,1322,1006,1857]" captionTargetId="figure-356@35.[189,1436,978,1889]" captionTargetPageId="35" captionText="Figures 113–115. Aedeagus of Scopaeus cuspidatus, holotype, Bali, Jembrana: Cekik in lateral (113), ventral (114), dorsal view (115). Abbreviations: al – apical lobes, dw – dorsodistal window of phallobase, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10724298" httpUri="https://zenodo.org/record/10724298/files/figure.png" pageId="36" pageNumber="59">Figs 113, 114</figureCitation>
).
</paragraph>
<caption id="DF6126E4FFE3FFD7BCE6F88C0C4503BA" ID-DOI="http://doi.org/10.5281/zenodo.10724298" ID-Zenodo-Dep="10724298" httpUri="https://zenodo.org/record/10724298/files/figure.png" pageId="35" pageNumber="58" startId="35.[189,262,1908,1930]" targetBox="[261,1322,1006,1857]" targetPageId="35" targetType="figure">
<paragraph id="8BA1766CFFE3FFD7BCE6F88C0C4503BA" blockId="35.[189,1437,1908,1975]" pageId="35" pageNumber="58">
<emphasis id="B96AAA7EFFE3FFD7BCE6F88C0C420384" bold="true" box="[189,351,1908,1930]" pageId="35" pageNumber="58">Figures 113–115.</emphasis>
Aedeagus of
<taxonomicName id="4C1E0DEFFFE3FFD7BDBBF88C0F830384" authorityName="Frisch" authorityYear="2022" box="[480,670,1908,1929]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="58" phylum="Arthropoda" rank="species" species="cuspidatus">
<emphasis id="B96AAA7EFFE3FFD7BDBBF88C0F830384" box="[480,670,1908,1929]" italics="true" pageId="35" pageNumber="58">Scopaeus cuspidatus</emphasis>
</taxonomicName>
, holotype, Bali, Jembrana: Cekik in lateral (
<emphasis id="B96AAA7EFFE3FFD7B867F88C09400384" bold="true" box="[1084,1117,1908,1929]" pageId="35" pageNumber="58">113</emphasis>
), ventral (
<emphasis id="B96AAA7EFFE3FFD7B8E4F88C09FD0384" bold="true" box="[1215,1248,1908,1929]" pageId="35" pageNumber="58">114</emphasis>
), dorsal view (
<emphasis id="B96AAA7EFFE3FFD7B935F88C08920384" bold="true" box="[1390,1423,1908,1929]" pageId="35" pageNumber="58">115</emphasis>
).
<emphasis id="B96AAA7EFFE3FFD7BCE6F8720C5B03AD" bold="true" box="[189,326,1930,1952]" pageId="35" pageNumber="58">Abbreviations</emphasis>
:
<emphasis id="B96AAA7EFFE3FFD7BD0AF8730C7E03AD" bold="true" box="[337,355,1931,1952]" pageId="35" pageNumber="58">al</emphasis>
– apical lobes,
<emphasis id="B96AAA7EFFE3FFD7BDABF8730F1003AD" bold="true" box="[496,525,1931,1952]" pageId="35" pageNumber="58">dw</emphasis>
– dorsodistal window of phallobase,
<emphasis id="B96AAA7EFFE3FFD7BF05F8720E6503AD" bold="true" box="[862,888,1930,1952]" pageId="35" pageNumber="58">mf</emphasis>
– median foramen,
<emphasis id="B96AAA7EFFE3FFD7B875F873095D03AD" bold="true" box="[1070,1088,1931,1952]" pageId="35" pageNumber="58">tr</emphasis>
– transverse ridge of median foramen,
<emphasis id="B96AAA7EFFE3FFD7BCE6F8590DD203BB" bold="true" box="[189,207,1953,1974]" pageId="35" pageNumber="58">vl</emphasis>
– ventral lobe.
</paragraph>
</caption>
<paragraph id="8BA1766CFFE4FFD0BCF5FDC20C7106B8" blockId="36.[151,762,241,923]" pageId="36" pageNumber="59">
Female: Sperm pump with short process segment and long, straight chamber without apophysis, narrowed towards thin, weakly sclerotized bursal duct (
<figureCitation id="13256AE9FFE4FFD0BECAFD840FF30699" box="[657,750,636,660]" captionStart="Figures 173–178" captionStartId="43.[189,262,1885,1907]" captionTargetId="figure-111@43.[189,1436,294,1840]" captionTargetPageId="43" captionText="Figures 173–178. Sperm pump (173–178) and bursa (173) of ♀ of Scopaeus bipenicillatus, paratype, Sulawesi Tengah, Sigi: Pagana (173, with lateral gonocoxal plates IX; 174); S. bipectenatus, paratype, Sulawesi Tengah: Lake Poso (175); S. cuspidatus, paratype, Jawa Barat: Mt Salak (176); S. crassipunctatus, paratype, Jawa Barat, Sukabumi: Sirnarasa (177); S. sulawesianus, paratype, Central Sulawesi, Tambarana (178, with bursal duct). Long scale bar: 174–178; short scale bar: 173." figureDoi="http://doi.org/10.5281/zenodo.10880230" httpUri="https://zenodo.org/record/10880230/files/figure.png" pageId="36" pageNumber="59">Fig. 176</figureCitation>
); bursa membranous.
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFE4FFD0BCF5FD460C1A0715" pageId="36" pageNumber="59" type="distribution">
<paragraph id="8BA1766CFFE4FFD0BCF5FD460C1A0715" blockId="36.[151,762,241,923]" pageId="36" pageNumber="59">
<emphasis id="B96AAA7EFFE4FFD0BCF5FD460C5906DB" bold="true" box="[174,324,702,726]" pageId="36" pageNumber="59">Distribution:</emphasis>
<taxonomicName id="4C1E0DEFFFE4FFD0BD10FD460F3006DB" authorityName="Frisch" authorityYear="2022" box="[331,557,702,726]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="59" phylum="Arthropoda" rank="species" species="cuspidatus">
<emphasis id="B96AAA7EFFE4FFD0BD10FD460F3006DB" box="[331,557,702,726]" italics="true" pageId="36" pageNumber="59">Scopaeus cuspidatus</emphasis>
</taxonomicName>
was found in
<collectingRegion id="49DAB88EFFE4FFD0BE9DFD460FE806DB" box="[710,757,702,726]" country="Indonesia" name="Bali" pageId="36" pageNumber="59">Bali</collectingRegion>
,
<collectingRegion id="49DAB88EFFE4FFD0BCCCFD270DD106FB" box="[151,204,735,758]" country="Indonesia" name="Jawa Timur" pageId="36" pageNumber="59">Jawa</collectingRegion>
and
<collectingRegion id="49DAB88EFFE4FFD0BD5CFD270C7A06FA" box="[263,359,735,759]" country="Indonesia" name="Sulawesi Utara" pageId="36" pageNumber="59">Sulawesi</collectingRegion>
and seems to be widespread across
<collectingCountry id="F30936FCFFE4FFD0BCCCFCF80C1E0715" box="[151,259,768,792]" name="Indonesia" pageId="36" pageNumber="59">Indonesia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFE4FFD0BCF5FCD90F1F0796" pageId="36" pageNumber="59" type="etymology">
<paragraph id="8BA1766CFFE4FFD0BCF5FCD90F1F0796" blockId="36.[151,762,241,923]" pageId="36" pageNumber="59">
<emphasis id="B96AAA7EFFE4FFD0BCF5FCD90C290734" bold="true" box="[174,308,801,825]" pageId="36" pageNumber="59">Etymology:</emphasis>
For this new species the epithet
<taxonomicName id="4C1E0DEFFFE4FFD0BEDEFCD80FE70735" authorityName="Frisch" authorityYear="2022" box="[645,762,800,824]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="59" phylum="Arthropoda" rank="species" species="cuspidatus">
<emphasis id="B96AAA7EFFE4FFD0BEDEFCD80FE70735" box="[645,762,800,824]" italics="true" pageId="36" pageNumber="59">cuspidatus</emphasis>
</taxonomicName>
(adjective, Latin: ‘tapered, pointed’) was chosen in reference of the distinct spine at the posterior margin of abdominal sternite VII (Fig. 142).
</paragraph>
</subSubSection>
</treatment>
</document>
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