384 lines
53 KiB
XML
384 lines
53 KiB
XML
<document id="0766F292EBAA38384424CD181F93CCD2" ID-DOI="10.25674/so95iss1id311" ID-ISSN="2509-9523" ID-Zenodo-Dep="10724236" checkinTime="1709147239332" checkinUser="felipe" docAuthor="Frisch, Johannes & Narakusumo, Raden Pramesa" docDate="2022" docId="03B7C77AFFD1FFE6BC8FFF080C1907D1" docLanguage="en" docName="SoilOrg.95.1.23-73.pdf" docOrigin="Soil Organisms 95 (1)" docSource="http://dx.doi.org/10.25674/so95iss1id311" docStyle="DocumentStyle:EA2ECAC87EC21C07AB805D5B9B4601F2:SoilOrg.2018-.journal_article" docStyleId="EA2ECAC87EC21C07AB805D5B9B4601F2" docStyleName="SoilOrg.2018-.journal_article" docTitle="Scopaeus grandis Frisch 2022, spec. nov." docType="treatment" docVersion="2" lastPageNumber="41" masterDocId="FF8EBF02FFC0FFF4BC5BFFF80D1D040D" masterDocTitle="Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species" masterLastPageNumber="73" masterPageNumber="23" pageNumber="40" updateTime="1709147743548" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods id="7D6C887FBDE34D5BCCF8AA70D6FB35E1" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="629E47D2E446065A2958E077D1FB6241">
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<mods:title id="59265ADA1F006F1B97C86B5AA6D03BE3">Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species</mods:title>
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</mods:titleInfo>
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<mods:name id="8279298941EB246DAA1CD60DFC44F3EA" type="personal">
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<mods:role id="88D515A80E88FC043D84B752B36BD9CC">
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<mods:roleTerm id="C58A9D3A05285A2181D5886A362DF3A5">Author</mods:roleTerm>
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<mods:namePart id="77995D622191E7E279CC5B7C920C0F68">Frisch, Johannes</mods:namePart>
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</mods:name>
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<mods:name id="2C64A16CEFB01057300DFCDAC2D7AB97" type="personal">
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<mods:role id="327842D6A53285CEB72877AA954865A8">
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<mods:roleTerm id="2CADA067F50813F8ECD8CBA9D5A9934E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A2245BBC22372EDFE346DF01130E4B3A">Narakusumo, Raden Pramesa</mods:namePart>
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<mods:affiliation id="B5DC5DD66F7801C2C51D5C731BB86918">MUseUm ZoologicUm Bogoriense, Research Center for Biosystematics and EvolUtion, National Research and Innovation Agency (BRIN), Jl. Raya Jakarta-Bogor KM 46, Cibinong, West Java, 16911, Indonesia</mods:affiliation>
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<mods:typeOfResource id="06EC5C95698D3C5603933B3E7BB007D6">text</mods:typeOfResource>
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<mods:title id="C1ABF6E9D20AAB712CE74DA6F660171D">Soil Organisms</mods:title>
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</mods:titleInfo>
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<mods:part id="E585CEBC13A57757B39877D61C361CF4">
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<mods:date id="F9C62EAFBCB35B90FF84860BCD7D5061">2022</mods:date>
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<mods:number id="1AAF6429F9412491539FC5995F4E99D5">2022-04-01</mods:number>
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<mods:detail id="3496B07B2EFD0F541595BCF3C5156B24" type="volume">
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<mods:number id="EB3B3EC7A92B3AC140DE56565A8D6067">95</mods:number>
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</mods:detail>
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<mods:detail id="03B447B7FE9A66DA71A8BC8A03E92887" type="issue">
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<mods:number id="12E13C52A3E39D195B33794036F41FF2">1</mods:number>
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<mods:start id="D5EA264EC5A02BB302625B321A3DA5FF">23</mods:start>
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<mods:end id="9EBB643D27C3ED376EC61268A025D105">73</mods:end>
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<mods:url id="18F6D409321EE72711E818D73837B9A7">http://dx.doi.org/10.25674/so95iss1id311</mods:url>
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<mods:classification id="7D4031844E8F164D5EE839C05126CF07">journal article</mods:classification>
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<mods:identifier id="AACD198495546B64F878537F5217EEBE" type="DOI">10.25674/so95iss1id311</mods:identifier>
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<mods:identifier id="25305696CCF9EDCBFF1D5E6848AFE700" type="ISSN">2509-9523</mods:identifier>
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<treatment id="03B7C77AFFD1FFE6BC8FFF080C1907D1" ID-DOI="http://doi.org/10.5281/zenodo.10724290" ID-Zenodo-Dep="10724290" LSID="urn:lsid:plazi:treatment:03B7C77AFFD1FFE6BC8FFF080C1907D1" httpUri="http://treatment.plazi.org/id/03B7C77AFFD1FFE6BC8FFF080C1907D1" lastPageId="18" lastPageNumber="41" pageId="17" pageNumber="40">
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<subSubSection id="C30425E7FFD1FFE5BC8FFF080F980507" box="[212,645,239,266]" pageId="17" pageNumber="40" type="nomenclature">
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<paragraph id="8BA1766CFFD1FFE5BC8FFF080F980507" blockId="17.[212,645,239,299]" box="[212,645,239,266]" pageId="17" pageNumber="40">
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<heading id="D0E9C100FFD1FFE5BC8FFF080F980507" box="[212,645,239,266]" fontSize="11" level="2" pageId="17" pageNumber="40" reason="5">
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<taxonomicName id="4C1E0DEFFFD1FFE5BC8FFF080F1E0504" authority="Frisch" authorityName="Frisch" authorityYear="2022" box="[212,515,239,266]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="40" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
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<emphasis id="B96AAA7EFFD1FFE5BC8FFF080F1E0504" bold="true" box="[212,515,239,266]" pageId="17" pageNumber="40">
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<emphasis id="B96AAA7EFFD1FFE5BC8FFF080CB60507" bold="true" box="[212,427,239,266]" italics="true" pageId="17" pageNumber="40">Scopaeus grandis</emphasis>
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Frisch
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</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A2591705FFD1FFE5BE4AFF080F980507" box="[529,645,240,266]" pageId="17" pageNumber="40" rank="species">spec. nov.</taxonomicNameLabel>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C30425E7FFD1FFE5BC8FFEE80F2C0526" box="[212,561,272,299]" pageId="17" pageNumber="40" type="description">
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<paragraph id="8BA1766CFFD1FFE5BC8FFEE80F2C0526" blockId="17.[212,645,239,299]" box="[212,561,272,299]" pageId="17" pageNumber="40">
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(
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<figureCitation id="13256AE9FFD1FFE5BC87FEE80C280526" box="[220,309,272,299]" captionStart="Figures 11–13" captionStartId="7.[189,262,1931,1953]" captionTargetBox="[232,1409,674,1880]" captionTargetId="figure-224@7.[189,1436,660,1912]" captionTargetPageId="7" captionText="Figures 11–13. Habitus of Scopaeus anuliflagellatus, paratype, Jawa Barat, Sukabumi: Cikaniki (11); S. grandis, paratype, Jawa Barat, Sukabumi: Cidahu (12); S. pulcher, paratype, Jawa Barat, Sukabumi: Cikaniki (13)." figureDoi="http://doi.org/10.5281/zenodo.10724244" httpUri="https://zenodo.org/record/10724244/files/figure.png" pageId="17" pageNumber="40">Figs 12</figureCitation>
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,
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<figureCitation id="13256AE9FFD1FFE5BD19FEE90C420526" box="[322,351,273,299]" captionStart="Figures 35–41" captionStartId="16.[151,224,1890,1912]" captionTargetBox="[184,1354,294,1780]" captionTargetId="figure-104@16.[151,1398,258,1871]" captionTargetPageId="16" captionText="Figures 35–41. Aedeagus in lateral view of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (35); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (36); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (37); S. cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (38); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (39); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (40); S. heronifer, holotype, Banten, Lebak: Majasari (41)." figureDoi="http://doi.org/10.5281/zenodo.10724258" httpUri="https://zenodo.org/record/10724258/files/figure.png" pageId="17" pageNumber="40">37</figureCitation>
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,
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<figureCitation id="13256AE9FFD1FFE5BD30FEE90CA30526" box="[363,446,273,299]" captionStart="Figures 80–85" captionStartId="26.[151,224,1884,1906]" captionTargetBox="[223,1322,801,1824]" captionTargetId="figure-325@26.[151,1398,782,1866]" captionTargetPageId="26" captionText="Figures 80–85. Apex of aedeagus in lateral (80, 83), ventral (81, 84), dorsal view (82, 85) of Scopaeus grandis, holotype, Java Barat, Sukabumi: Cidahu (80–82); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (83–85). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median lobe, vl – ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10724276" httpUri="https://zenodo.org/record/10724276/files/figure.png" pageId="17" pageNumber="40">80 –82</figureCitation>
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,
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<figureCitation id="13256AE9FFD1FFE5BD91FEE90CE90526" box="[458,500,273,299]" captionStart="Figures 119–133" captionStartId="38.[151,224,1796,1818]" captionTargetBox="[224,1369,297,1686]" captionTargetId="figure-201@38.[151,1398,236,1777]" captionTargetPageId="38" captionText="Figures 119–133. Abdominal sternite VII (126) and abdominal sternite VIII (119–125, 127– 133) of ♂ of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (119); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (120); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (121); S. jacobsoni, Jawa Barat: Mt Salak (122); S. mixtus, Malaysia, Sabah: Lokan (123); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi: Sirnarasa (124); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi: Cisalimas (125); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (126, 127); S. spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (128); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi:Cidahu (129); S. pulcher, holotype, Jawa Barat, Sukabumi:Cikaniki (130); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (131); S. heronifer, holotype, Banten, Lebak: Majasari (132); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (133)." figureDoi="http://doi.org/10.5281/zenodo.10724304" httpUri="https://zenodo.org/record/10724304/files/figure.png" pageId="17" pageNumber="40">131</figureCitation>
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,
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<figureCitation id="13256AE9FFD1FFE5BE5BFEE80F340526" box="[512,553,272,299]" captionStart="Figures 163–172" captionStartId="42.[151,224,1839,1861]" captionTargetBox="[225,1322,230,1817]" captionTargetId="figure-168@42.[157,1398,210,1820]" captionTargetPageId="42" captionText="Figures 163–172. Sperm pump (163–172) and bursa (165, 167, 170, 172) of ♀ of Scopaeus grandis, paratype, Jawa Barat, Sukabumi: Cidahu (163); S.heronifer, paratype, Jawa Barat, Sukabumi:Sirnarasa (164); S. diversilobatus, paratype, Jawa Barat, Sukabumi, Djampang Kulon (165, with lateral gonocoxal plates IX folded laterally; 166); S. cuspilobatus, paratype, Jawa Barat, Sukabumi: Sirnarasa (167, with lateral gonocoxal plates IX, 168); S. velifer, paratype, Banten, Lebak: Citorek Kidul (169); S. elegantulus, Jawa Barat, Sukabumi: Djampang Kulon (170, with lateral gonocoxal plates IX, 171), S. uncinatus, paratype, Banten, Lebak: Citorek Kidul (172). Long scale bar: 163, 164, 166, 168, 169, 171, 172; short scale bar: 165, 167, 170." figureDoi="http://doi.org/10.5281/zenodo.10724315" httpUri="https://zenodo.org/record/10724315/files/figure.png" pageId="17" pageNumber="40">163</figureCitation>
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)
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C30425E7FFD1FFE5BC8FFEAC0FD106DB" pageId="17" pageNumber="40" type="materials_examined">
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<paragraph id="8BA1766CFFD1FFE5BC8FFEAC0FD106DB" blockId="17.[189,800,340,824]" pageId="17" pageNumber="40">
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<emphasis id="B96AAA7EFFD1FFE5BC8FFEAC0C8F0561" bold="true" box="[212,402,340,364]" pageId="17" pageNumber="40">Type specimens:</emphasis>
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<materialsCitation id="3B767C31FFD1FFE5BDC1FEAC0C9805C2" collectingDate="2015-09-29" collectionCode="MZB" collectorName="Frisch" country="Indonesia" county="Sukabumi" elevation="1210" latitude="-6.7383337" location="Cirasamala River" longLatPrecision="21" longitude="106.71444" municipality="Cidahu" pageId="17" pageNumber="40" specimenCount="1" specimenCount-male="1" stateProvince="Jawa Barat" typeStatus="holotype">
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||
<collectingCountry id="F30936FCFFD1FFE5BDC1FEAC0F1A0561" box="[410,519,340,364]" name="Indonesia" pageId="17" pageNumber="40">Indonesia</collectingCountry>
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:
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<typeStatus id="54A5C8CEFFD1FFE5BE48FEAC0F6A0561" box="[531,631,340,364]" pageId="17" pageNumber="40" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D18BDE5FFD1FFE5BE24FEAD0F8E0561" box="[639,659,341,364]" count="1" pageId="17" pageNumber="40" type="male">♂</specimenCount>
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,
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<collectingRegion id="49DAB88EFFD1FFE5BEFAFEAD0E010561" box="[673,796,341,364]" country="Indonesia" name="Jawa Barat" pageId="17" pageNumber="40">Jawa Barat</collectingRegion>
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,
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<collectingCounty id="62C00EE0FFD1FFE5BCE6FE8D0C330580" box="[189,302,373,397]" pageId="17" pageNumber="40">Sukabumi</collectingCounty>
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, NNW
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<collectingMunicipality id="6BC5EC16FFD1FFE5BDD9FE8D0CC90580" box="[386,468,373,397]" pageId="17" pageNumber="40">Cidahu</collectingMunicipality>
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||
:
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<location id="8EC120B7FFD1FFE5BD85FE8D0F860580" LSID="urn:lsid:plazi:treatment:03B7C77AFFD1FFE6BC8FFF080C1907D1:8EC120B7FFD1FFE5BD85FE8D0F860580" box="[478,667,373,397]" country="Indonesia" county="Sukabumi" latitude="-6.7383337" longLatPrecision="21" longitude="106.71444" municipality="Cidahu" name="Cirasamala River" pageId="17" pageNumber="40" stateProvince="Jawa Barat">Cirasamala River</location>
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(S-slope
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<location id="8EC120B7FFD1FFE5BF5AFE8E0DE105A3" LSID="urn:lsid:plazi:treatment:03B7C77AFFD1FFE6BC8FFF080C1907D1:8EC120B7FFD1FFE5BF5AFE8E0DE105A3" country="Indonesia" county="Sukabumi" latitude="-6.7383337" longLatPrecision="21" longitude="106.71444" municipality="Cidahu" name="Mt Salak" pageId="17" pageNumber="40" stateProvince="Jawa Barat">Mt Salak</location>
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), (
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<geoCoordinate id="EE2A10ABFFD1FFE5BD47FE6E0C8405A3" box="[284,409,406,430]" degrees="06" direction="south" minutes="44" orientation="latitude" pageId="17" pageNumber="40" precision="15" seconds="18" value="-6.7383337">06°44’18’’S</geoCoordinate>
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,
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<geoCoordinate id="EE2A10ABFFD1FFE5BDF1FE6E0F2A05A3" box="[426,567,406,430]" degrees="106" direction="east" minutes="42" orientation="longitude" pageId="17" pageNumber="40" precision="15" seconds="52" value="106.71444">106°42’52’’E</geoCoordinate>
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||
),
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||
<quantity id="4CE6DB89FFD1FFE5BE14FE6E0FB905A0" box="[591,676,406,430]" metricMagnitude="3" metricUnit="m" metricValue="1.21" pageId="17" pageNumber="40" unit="m" value="1210.0">
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||
<elevation id="0033915FFFD1FFE5BE14FE6E0FB905A0" box="[591,676,406,430]" metricMagnitude="3" metricUnit="m" metricValue="1.21" pageId="17" pageNumber="40" unit="m" value="1210.0">1210 m</elevation>
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||
</quantity>
|
||
,
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||
<date id="FFA050ACFFD1FFE5BEEEFE6E0E0105A3" box="[693,796,406,430]" pageId="17" pageNumber="40" value="2015-09-29">
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||
<collectingDate id="EFE4A944FFD1FFE5BEEEFE6E0E0105A3" box="[693,796,406,430]" pageId="17" pageNumber="40" value="2015-09-29">29.9.2015</collectingDate>
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||
</date>
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||
, leg.
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||
<collectorName id="26EB13BAFFD1FFE5BCB0FE4F0C3305C2" box="[235,302,439,463]" pageId="17" pageNumber="40">Frisch</collectorName>
|
||
(
|
||
<collectionCode id="ED0FEEA9FFD1FFE5BD65FE4F0C6005C2" box="[318,381,439,463]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="17" pageNumber="40" type="Museum">MZB</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
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||
<typeStatus id="54A5C8CEFFD1FFE5BDCBFE4F0CE605C2" box="[400,507,439,463]" pageId="17" pageNumber="40" type="paratype">Paratypes</typeStatus>
|
||
(
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||
<specimenCount id="9D18BDE5FFD1FFE5BE51FE4F0FBD05C2" box="[522,672,439,463]" count="12" pageId="17" pageNumber="40" type="generic">12 specimens</specimenCount>
|
||
):
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||
<materialsCitation id="3B767C31FFD1FFE5BEE9FE4F0F0705FD" collectingDate="2015-09-29" collectionCode="MFNB, MZB" collectorName="Frisch" country="Indonesia" county="Sukabumi" elevation="1210" latitude="-6.7383337" location="Cirasamala River" longLatPrecision="21" longitude="106.71444" municipality="Cidahu" pageId="17" pageNumber="40" specimenCount="2" specimenCount-female="2" stateProvince="Jawa Barat" typeStatus="paratype">
|
||
<specimenCount id="9D18BDE5FFD1FFE5BEE9FE4F0FC705C3" box="[690,730,439,463]" count="2" pageId="17" pageNumber="40" type="female">2 ♀</specimenCount>
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||
, same data as holotype (
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<collectionCode id="ED0FEEA9FFD1FFE5BD22FE200CD605FD" box="[377,459,472,496]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="17" pageNumber="40">MFNB</collectionCode>
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||
,
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||
<collectionCode id="ED0FEEA9FFD1FFE5BD8EFE200F0E05FD" box="[469,531,472,496]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="17" pageNumber="40" type="Museum">MZB</collectionCode>
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||
)
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||
</materialsCitation>
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||
.
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||
<materialsCitation id="3B767C31FFD1FFE5BE78FE200C1E065F" collectingDate="2015-03-10" collectionCode="MFNB, MZB" collectorName="Frisch" country="Indonesia" county="Bogor" elevation="990" latitude="-6.7113886" location="Cimuara Herang River" longLatPrecision="21" longitude="106.684166" municipality="Pamijahan" pageId="17" pageNumber="40" specimenCount="2" specimenCount-female="2" stateProvince="Jawa Barat" typeStatus="paratype">
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||
<collectingRegion id="49DAB88EFFD1FFE5BE78FE200F8105E2" box="[547,668,472,495]" country="Indonesia" name="Jawa Barat" pageId="17" pageNumber="40">Jawa Barat</collectingRegion>
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||
:
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<specimenCount id="9D18BDE5FFD1FFE5BEFEFE200FD605E2" box="[677,715,472,496]" count="2" pageId="17" pageNumber="40" type="female">2 ♀</specimenCount>
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||
,
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<collectingCounty id="62C00EE0FFD1FFE5BE8CFE200E0605FD" box="[727,795,472,496]" pageId="17" pageNumber="40">Bogor</collectingCounty>
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||
, SW
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<collectingMunicipality id="6BC5EC16FFD1FFE5BCA9FE000C79061D" box="[242,356,504,528]" pageId="17" pageNumber="40">Pamijahan</collectingMunicipality>
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||
(
|
||
<location id="8EC120B7FFD1FFE5BD23FE000F62061D" LSID="urn:lsid:plazi:treatment:03B7C77AFFD1FFE6BC8FFF080C1907D1:8EC120B7FFD1FFE5BD23FE000F62061D" box="[376,639,504,529]" country="Indonesia" county="Bogor" latitude="-6.7113886" longLatPrecision="21" longitude="106.684166" municipality="Pamijahan" name="Cimuara Herang River" pageId="17" pageNumber="40" stateProvince="Jawa Barat">Cimuara Herang River</location>
|
||
), (
|
||
<geoCoordinate id="EE2A10ABFFD1FFE5BEFBFE000E06061D" box="[672,795,504,528]" degrees="06" direction="south" minutes="42" orientation="latitude" pageId="17" pageNumber="40" precision="15" seconds="41" value="-6.7113886">06°42’41’’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE2A10ABFFD1FFE5BCE6FDE10C55063C" box="[189,328,537,561]" degrees="106" direction="east" minutes="41" orientation="longitude" pageId="17" pageNumber="40" precision="15" seconds="03" value="106.684166">106°41’03’’E</geoCoordinate>
|
||
),
|
||
<quantity id="4CE6DB89FFD1FFE5BD39FDE20CAD063C" box="[354,432,538,562]" metricMagnitude="2" metricUnit="m" metricValue="9.9" pageId="17" pageNumber="40" unit="m" value="990.0">
|
||
<elevation id="0033915FFFD1FFE5BD39FDE20CAD063C" box="[354,432,538,562]" metricMagnitude="2" metricUnit="m" metricValue="9.9" pageId="17" pageNumber="40" unit="m" value="990.0">990 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFA050ACFFD1FFE5BD98FDE10F34063C" box="[451,553,537,561]" pageId="17" pageNumber="40" value="2015-03-10">
|
||
<collectingDate id="EFE4A944FFD1FFE5BD98FDE10F34063C" box="[451,553,537,561]" pageId="17" pageNumber="40" value="2015-03-10">3.10.2015</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26EB13BAFFD1FFE5BE2BFDE10FAE063C" box="[624,691,537,561]" pageId="17" pageNumber="40">Frisch</collectorName>
|
||
(
|
||
<collectionCode id="ED0FEEA9FFD1FFE5BE92FDE20E06063F" box="[713,795,538,562]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="17" pageNumber="40">MFNB</collectionCode>
|
||
,
|
||
<collectionCode id="ED0FEEA9FFD1FFE5BCE6FDC20DE6065F" box="[189,251,570,594]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="17" pageNumber="40" type="Museum">MZB</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B767C31FFD1FFE5BD49FDC20CA70699" collectingDate="2015-02-10" collectionCode="MFNB, MZB" collectorName="Frisch" country="Indonesia" county="Bogor" elevation="1040" latitude="-6.6933336" location="Tapos" longLatPrecision="21" longitude="106.706665" municipality="Tenjolaja" pageId="17" pageNumber="40" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Jawa Barat" typeStatus="paratype">
|
||
<specimenCount id="9D18BDE5FFD1FFE5BD49FDC20C22065F" box="[274,319,570,594]" count="1" pageId="17" pageNumber="40" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D18BDE5FFD1FFE5BD14FDC20C67065F" box="[335,378,570,594]" count="1" pageId="17" pageNumber="40" type="female">1 ♀</specimenCount>
|
||
,
|
||
<collectingCounty id="62C00EE0FFD1FFE5BDD7FDC30CCD065E" box="[396,464,571,595]" pageId="17" pageNumber="40">Bogor</collectingCounty>
|
||
, S
|
||
<collectingMunicipality id="6BC5EC16FFD1FFE5BDA1FDC20F42065F" box="[506,607,570,594]" pageId="17" pageNumber="40">Tenjolaja</collectingMunicipality>
|
||
,
|
||
<location id="8EC120B7FFD1FFE5BE35FDC30FAF065F" LSID="urn:lsid:plazi:treatment:03B7C77AFFD1FFE6BC8FFF080C1907D1:8EC120B7FFD1FFE5BE35FDC30FAF065F" box="[622,690,571,594]" country="Indonesia" county="Bogor" latitude="-6.6933336" longLatPrecision="21" longitude="106.706665" municipality="Tenjolaja" name="Tapos" pageId="17" pageNumber="40" stateProvince="Jawa Barat">Tapos</location>
|
||
:
|
||
<location id="8EC120B7FFD1FFE5BE9BFDC20DE4067E" LSID="urn:lsid:plazi:treatment:03B7C77AFFD1FFE6BC8FFF080C1907D1:8EC120B7FFD1FFE5BE9BFDC20DE4067E" country="Indonesia" county="Bogor" latitude="-6.6933336" longLatPrecision="21" longitude="106.706665" municipality="Tenjolaja" name="Ciampea River" pageId="17" pageNumber="40" stateProvince="Jawa Barat">Ciampea River</location>
|
||
(
|
||
<geoCoordinate id="EE2A10ABFFD1FFE5BD5DFDA30C9E067E" box="[262,387,603,627]" degrees="06" direction="south" minutes="41" orientation="latitude" pageId="17" pageNumber="40" precision="15" seconds="36" value="-6.6933336">06°41’36’’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE2A10ABFFD1FFE5BDD6FDA30F06067E" box="[397,539,603,627]" degrees="106" direction="east" minutes="42" orientation="longitude" pageId="17" pageNumber="40" precision="15" seconds="24" value="106.706665">106°42’24’’E</geoCoordinate>
|
||
),
|
||
<quantity id="4CE6DB89FFD1FFE5BE76FDA30F63067E" box="[557,638,603,627]" metricMagnitude="3" metricUnit="m" metricValue="1.04" pageId="17" pageNumber="40" unit="m" value="1040.0">
|
||
<elevation id="0033915FFFD1FFE5BE76FDA30F63067E" box="[557,638,603,627]" metricMagnitude="3" metricUnit="m" metricValue="1.04" pageId="17" pageNumber="40" unit="m" value="1040.0">1040 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFA050ACFFD1FFE5BED3FDA30FF2067E" box="[648,751,603,627]" pageId="17" pageNumber="40" value="2015-02-10">
|
||
<collectingDate id="EFE4A944FFD1FFE5BED3FDA30FF2067E" box="[648,751,603,627]" pageId="17" pageNumber="40" value="2015-02-10">2.10.2015</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26EB13BAFFD1FFE5BCE6FD840C1D0699" box="[189,256,636,660]" pageId="17" pageNumber="40">Frisch</collectorName>
|
||
(
|
||
<collectionCode id="ED0FEEA9FFD1FFE5BD48FD840C780699" box="[275,357,636,660]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="17" pageNumber="40">MFNB</collectionCode>
|
||
,
|
||
<collectionCode id="ED0FEEA9FFD1FFE5BD2EFD840CAE0699" box="[373,435,636,660]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="17" pageNumber="40" type="Museum">MZB</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B767C31FFD1FFE5BD92FD840F5906B8" collectingDate="1984-06-17" collectionCode="HECO" collectorName="Rougemont" country="Indonesia" location="Batukaru" municipality="Tabanan" pageId="17" pageNumber="40" specimenCount="4" specimenCount-male="4" stateProvince="Bali" typeStatus="paratype">
|
||
<collectingRegion id="49DAB88EFFD1FFE5BD92FD840CE70699" box="[457,506,636,660]" country="Indonesia" name="Bali" pageId="17" pageNumber="40">Bali</collectingRegion>
|
||
:
|
||
<specimenCount id="9D18BDE5FFD1FFE5BE53FD840F280699" box="[520,565,636,660]" count="4" pageId="17" pageNumber="40" type="male">4 ♂</specimenCount>
|
||
,
|
||
<collectingMunicipality id="6BC5EC16FFD1FFE5BE1EFD840FB80699" box="[581,677,636,660]" pageId="17" pageNumber="40">Tabanan</collectingMunicipality>
|
||
:
|
||
<location id="8EC120B7FFD1FFE5BEE8FD840E010699" LSID="urn:lsid:plazi:treatment:03B7C77AFFD1FFE6BC8FFF080C1907D1:8EC120B7FFD1FFE5BEE8FD840E010699" box="[691,796,636,660]" country="Indonesia" municipality="Tabanan" name="Batukaru" pageId="17" pageNumber="40" stateProvince="Bali">Batukaru</location>
|
||
,
|
||
<date id="FFA050ACFFD1FFE5BCE6FD650C3E06B8" box="[189,291,669,693]" pageId="17" pageNumber="40" value="1984-06-17">
|
||
<collectingDate id="EFE4A944FFD1FFE5BCE6FD650C3E06B8" box="[189,291,669,693]" pageId="17" pageNumber="40" value="1984-06-17">17.6.1984</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26EB13BAFFD1FFE5BD3AFD650CC006B8" box="[353,477,669,693]" pageId="17" pageNumber="40">Rougemont</collectorName>
|
||
(
|
||
<collectionCode id="ED0FEEA9FFD1FFE5BDB5FD650F2106B8" box="[494,572,669,693]" pageId="17" pageNumber="40">HECO</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B767C31FFD1FFE5BE0AFD650FD406DB" collectingDate="1991-10-29" collectionCode="MHNG" collectorName="Lobl" country="Indonesia" elevation="600" location="Mt Batukaru" pageId="17" pageNumber="40" specimenCount="2" specimenCount-male="2" stateProvince="Bali" typeStatus="paratype">
|
||
<specimenCount id="9D18BDE5FFD1FFE5BE0AFD650F6106B9" box="[593,636,669,693]" count="2" pageId="17" pageNumber="40" type="male">2 ♂</specimenCount>
|
||
,
|
||
<location id="8EC120B7FFD1FFE5BED1FD650E0106B8" LSID="urn:lsid:plazi:treatment:03B7C77AFFD1FFE6BC8FFF080C1907D1:8EC120B7FFD1FFE5BED1FD650E0106B8" box="[650,796,669,693]" country="Indonesia" name="Mt Batukaru" pageId="17" pageNumber="40" stateProvince="Bali">Mt Batukaru</location>
|
||
,
|
||
<quantity id="4CE6DB89FFD1FFE5BCE6FD460C2206D8" box="[189,319,702,726]" metricMagnitude="2" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="17" pageNumber="40" unit="m" value="600.0" valueMax="700.0" valueMin="500.0">
|
||
<elevation id="0033915FFFD1FFE5BCE6FD460C2206D8" box="[189,319,702,726]" metricMagnitude="2" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="17" pageNumber="40" unit="m" value="600.0" valueMax="700.0" valueMin="500.0">500 – 700 m</elevation>
|
||
</quantity>
|
||
, 28. –
|
||
<date id="FFA050ACFFD1FFE5BD24FD460CED06DB" box="[383,496,702,726]" pageId="17" pageNumber="40" value="1991-10-29">
|
||
<collectingDate id="EFE4A944FFD1FFE5BD24FD460CED06DB" box="[383,496,702,726]" pageId="17" pageNumber="40" value="1991-10-29">29.10.1991</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26EB13BAFFD1FFE5BE73FD460F4706DB" box="[552,602,702,726]" pageId="17" pageNumber="40">Löbl</collectorName>
|
||
(
|
||
<collectionCode id="ED0FEEA9FFD1FFE5BE33FD460FDC06DB" box="[616,705,702,726]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="17" pageNumber="40" type="Museum">MHNG</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30425E7FFD1FFE6BC8FFD260F820715" lastPageId="18" lastPageNumber="41" pageId="17" pageNumber="40" type="description">
|
||
<paragraph id="8BA1766CFFD1FFE5BC8FFD260ED4065F" blockId="17.[189,800,340,824]" lastBlockId="17.[824,1437,241,825]" pageId="17" pageNumber="40">
|
||
<emphasis id="B96AAA7EFFD1FFE5BC8FFD260C7806FB" bold="true" box="[212,357,734,758]" pageId="17" pageNumber="40">Description:</emphasis>
|
||
Habitus and coloring as in
|
||
<figureCitation id="13256AE9FFD1FFE5BE9EFD270E0606FA" box="[709,795,735,759]" captionStart="Figures 11–13" captionStartId="7.[189,262,1931,1953]" captionTargetBox="[232,1409,674,1880]" captionTargetId="figure-224@7.[189,1436,660,1912]" captionTargetPageId="7" captionText="Figures 11–13. Habitus of Scopaeus anuliflagellatus, paratype, Jawa Barat, Sukabumi: Cikaniki (11); S. grandis, paratype, Jawa Barat, Sukabumi: Cidahu (12); S. pulcher, paratype, Jawa Barat, Sukabumi: Cikaniki (13)." figureDoi="http://doi.org/10.5281/zenodo.10724244" httpUri="https://zenodo.org/record/10724244/files/figure.png" pageId="17" pageNumber="40">Fig. 12</figureCitation>
|
||
. Head subcircular. Antennae with penultimate segment about 1.6 times as long as wide. Mesotibia moderately thickened. Forebody surface with fine, dense, setose punctation, without microreticulation, subnitid; abdomen densely punctate and microreticulate, matt. Pubescence of body surface short, decumbent, without conspicuous macrosetae. Body dark brown to black-brown; maxillary palpi medium brown with blackish penultimate segment; antennae with scapus medium brown, segments 2 – 6 blackish, then gradually lighter towards light brown terminal segments; legs medium brown. Total body length
|
||
<quantity id="4CE6DB89FFD1FFE5B80DFDE109FF063C" box="[1110,1250,537,562]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" metricValueMax="4.7" metricValueMin="4.5" pageId="17" pageNumber="40" unit="mm" value="4.6" valueMax="4.7" valueMin="4.5">4.5 – 4.7 mm</quantity>
|
||
; forebody length
|
||
<quantity id="4CE6DB89FFD1FFE5BF63FDC20ED8065F" box="[824,965,570,594]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" metricValueMax="2.7" metricValueMin="2.5" pageId="17" pageNumber="40" unit="mm" value="2.6" valueMax="2.7" valueMin="2.5">2.5 – 2.7 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA1766CFFD1FFE6BF14FDA30F59067E" blockId="17.[824,1437,241,825]" lastBlockId="18.[151,764,241,988]" lastPageId="18" lastPageNumber="41" pageId="17" pageNumber="40">
|
||
Male: Abdominal sternite VII with unmodified posterior margin. Abdominal sternite VIII short, concave in posterior third (
|
||
<figureCitation id="13256AE9FFD1FFE5B85FFD65094306B8" box="[1028,1118,669,693]" captionStart="Figures 119–133" captionStartId="38.[151,224,1796,1818]" captionTargetBox="[224,1369,297,1686]" captionTargetId="figure-201@38.[151,1398,236,1777]" captionTargetPageId="38" captionText="Figures 119–133. Abdominal sternite VII (126) and abdominal sternite VIII (119–125, 127– 133) of ♂ of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (119); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (120); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (121); S. jacobsoni, Jawa Barat: Mt Salak (122); S. mixtus, Malaysia, Sabah: Lokan (123); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi: Sirnarasa (124); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi: Cisalimas (125); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (126, 127); S. spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (128); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi:Cidahu (129); S. pulcher, holotype, Jawa Barat, Sukabumi:Cikaniki (130); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (131); S. heronifer, holotype, Banten, Lebak: Majasari (132); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (133)." figureDoi="http://doi.org/10.5281/zenodo.10724304" httpUri="https://zenodo.org/record/10724304/files/figure.png" pageId="17" pageNumber="40">Fig. 131</figureCitation>
|
||
). Aedeagus (
|
||
<figureCitation id="13256AE9FFD1FFE5B8B6FD65082206B8" box="[1261,1343,669,693]" captionStart="Figures 35–41" captionStartId="16.[151,224,1890,1912]" captionTargetBox="[184,1354,294,1780]" captionTargetId="figure-104@16.[151,1398,258,1871]" captionTargetPageId="16" captionText="Figures 35–41. Aedeagus in lateral view of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (35); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (36); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (37); S. cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (38); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (39); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (40); S. heronifer, holotype, Banten, Lebak: Majasari (41)." figureDoi="http://doi.org/10.5281/zenodo.10724258" httpUri="https://zenodo.org/record/10724258/files/figure.png" pageId="17" pageNumber="40">Figs 37</figureCitation>
|
||
,
|
||
<figureCitation id="13256AE9FFD1FFE5B917FD65088806B8" box="[1356,1429,669,693]" captionStart="Figures 80–85" captionStartId="26.[151,224,1884,1906]" captionTargetBox="[223,1322,801,1824]" captionTargetId="figure-325@26.[151,1398,782,1866]" captionTargetPageId="26" captionText="Figures 80–85. Apex of aedeagus in lateral (80, 83), ventral (81, 84), dorsal view (82, 85) of Scopaeus grandis, holotype, Java Barat, Sukabumi: Cidahu (80–82); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (83–85). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median lobe, vl – ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10724276" httpUri="https://zenodo.org/record/10724276/files/figure.png" pageId="17" pageNumber="40">80 – 82</figureCitation>
|
||
) large, about
|
||
<quantity id="4CE6DB89FFD1FFE5BFE5FD46091206D8" box="[958,1039,702,726]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="17" pageNumber="40" unit="mm" value="1.5">1.5 mm</quantity>
|
||
long; phallobase exceptionally large with small, apicoventrally attached, asymmetrical distal lobes; dextral apical lobe at ventral margin with apical denticle followed by two lobes proximally (
|
||
<figureCitation id="13256AE9FFD1FFE5B94EFCD8087A0734" box="[1301,1383,800,825]" captionStart="Figures 35–41" captionStartId="16.[151,224,1890,1912]" captionTargetBox="[184,1354,294,1780]" captionTargetId="figure-104@16.[151,1398,258,1871]" captionTargetPageId="16" captionText="Figures 35–41. Aedeagus in lateral view of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (35); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (36); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (37); S. cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (38); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (39); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (40); S. heronifer, holotype, Banten, Lebak: Majasari (41)." figureDoi="http://doi.org/10.5281/zenodo.10724258" httpUri="https://zenodo.org/record/10724258/files/figure.png" pageId="17" pageNumber="40">Figs 37</figureCitation>
|
||
,
|
||
<figureCitation id="13256AE9FFD1FFE5B928FCD9088D0734" box="[1395,1424,801,825]" captionStart="Figures 80–85" captionStartId="26.[151,224,1884,1906]" captionTargetBox="[223,1322,801,1824]" captionTargetId="figure-325@26.[151,1398,782,1866]" captionTargetPageId="26" captionText="Figures 80–85. Apex of aedeagus in lateral (80, 83), ventral (81, 84), dorsal view (82, 85) of Scopaeus grandis, holotype, Java Barat, Sukabumi: Cidahu (80–82); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (83–85). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median lobe, vl – ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10724276" httpUri="https://zenodo.org/record/10724276/files/figure.png" pageId="17" pageNumber="40">80</figureCitation>
|
||
), in ventral and dorsal view convex laterally with acute apex (
|
||
<figureCitation id="13256AE9FFD2FFE6BC85FEEA0C7E0527" box="[222,355,274,298]" captionStart="Figures 80–85" captionStartId="26.[151,224,1884,1906]" captionTargetBox="[223,1322,801,1824]" captionTargetId="figure-325@26.[151,1398,782,1866]" captionTargetPageId="26" captionText="Figures 80–85. Apex of aedeagus in lateral (80, 83), ventral (81, 84), dorsal view (82, 85) of Scopaeus grandis, holotype, Java Barat, Sukabumi: Cidahu (80–82); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (83–85). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median lobe, vl – ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10724276" httpUri="https://zenodo.org/record/10724276/files/figure.png" pageId="18" pageNumber="41">Figs 81, 82</figureCitation>
|
||
); sinistral apical lobe, compared to dextral apical lobe, longer with more strongly sclerotized, round end curved to right (
|
||
<figureCitation id="13256AE9FFD2FFE6BD99FEAC0F230561" box="[450,574,340,364]" captionStart="Figures 80–85" captionStartId="26.[151,224,1884,1906]" captionTargetBox="[223,1322,801,1824]" captionTargetId="figure-325@26.[151,1398,782,1866]" captionTargetPageId="26" captionText="Figures 80–85. Apex of aedeagus in lateral (80, 83), ventral (81, 84), dorsal view (82, 85) of Scopaeus grandis, holotype, Java Barat, Sukabumi: Cidahu (80–82); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (83–85). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median lobe, vl – ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10724276" httpUri="https://zenodo.org/record/10724276/files/figure.png" pageId="18" pageNumber="41">Figs 81, 82</figureCitation>
|
||
); flagellum stout, projecting from apical lobes far distad, with narrow winding lying on surface of apical portion of sinistral apical lobe (
|
||
<figureCitation id="13256AE9FFD2FFE6BD40FE4F0C8805C2" box="[283,405,439,463]" captionStart="Figures 80–85" captionStartId="26.[151,224,1884,1906]" captionTargetBox="[223,1322,801,1824]" captionTargetId="figure-325@26.[151,1398,782,1866]" captionTargetPageId="26" captionText="Figures 80–85. Apex of aedeagus in lateral (80, 83), ventral (81, 84), dorsal view (82, 85) of Scopaeus grandis, holotype, Java Barat, Sukabumi: Cidahu (80–82); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (83–85). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median lobe, vl – ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10724276" httpUri="https://zenodo.org/record/10724276/files/figure.png" pageId="18" pageNumber="41">Figs 81, 82</figureCitation>
|
||
); lateral lobes reduced, indicated by groups of setae (
|
||
<figureCitation id="13256AE9FFD2FFE6BDD6FE200F0505FD" box="[397,536,472,496]" captionStart="Figures 80–85" captionStartId="26.[151,224,1884,1906]" captionTargetBox="[223,1322,801,1824]" captionTargetId="figure-325@26.[151,1398,782,1866]" captionTargetPageId="26" captionText="Figures 80–85. Apex of aedeagus in lateral (80, 83), ventral (81, 84), dorsal view (82, 85) of Scopaeus grandis, holotype, Java Barat, Sukabumi: Cidahu (80–82); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (83–85). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median lobe, vl – ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10724276" httpUri="https://zenodo.org/record/10724276/files/figure.png" pageId="18" pageNumber="41">Figs 80 – 82</figureCitation>
|
||
); ventral lobe with strongly emarginate apex, extended asymmetically in proximosinistral and distodextral tooth (
|
||
<figureCitation id="13256AE9FFD2FFE6BE17FDE10F84063F" box="[588,665,537,562]" captionStart="Figures 80–85" captionStartId="26.[151,224,1884,1906]" captionTargetBox="[223,1322,801,1824]" captionTargetId="figure-325@26.[151,1398,782,1866]" captionTargetPageId="26" captionText="Figures 80–85. Apex of aedeagus in lateral (80, 83), ventral (81, 84), dorsal view (82, 85) of Scopaeus grandis, holotype, Java Barat, Sukabumi: Cidahu (80–82); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (83–85). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median lobe, vl – ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10724276" httpUri="https://zenodo.org/record/10724276/files/figure.png" pageId="18" pageNumber="41">Fig. 81</figureCitation>
|
||
); median foramen narrow, with asymmetical, transverse ridge acute-angled projecting distad (
|
||
<figureCitation id="13256AE9FFD2FFE6BDB7FDA30F25067E" box="[492,568,603,627]" captionStart="Figures 80–85" captionStartId="26.[151,224,1884,1906]" captionTargetBox="[223,1322,801,1824]" captionTargetId="figure-325@26.[151,1398,782,1866]" captionTargetPageId="26" captionText="Figures 80–85. Apex of aedeagus in lateral (80, 83), ventral (81, 84), dorsal view (82, 85) of Scopaeus grandis, holotype, Java Barat, Sukabumi: Cidahu (80–82); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (83–85). Abbreviations: al(d) – dextral apical lobe, al(s) – sinistral apical lobe, dl – dorsal lobe, f – flagellum, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median lobe, vl – ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10724276" httpUri="https://zenodo.org/record/10724276/files/figure.png" pageId="18" pageNumber="41">Fig. 81</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF6126E4FFD1FFE5BCE5F8A6088103B5" ID-DOI="http://doi.org/10.5281/zenodo.10724262" ID-Zenodo-Dep="10724262" httpUri="https://zenodo.org/record/10724262/files/figure.png" pageId="17" pageNumber="40" startId="17.[190,263,1886,1908]" targetBox="[263,1341,889,1832]" targetPageId="17" targetType="figure">
|
||
<paragraph id="8BA1766CFFD1FFE5BCE5F8A6088103B5" blockId="17.[189,1437,1886,1976]" pageId="17" pageNumber="40">
|
||
<emphasis id="B96AAA7EFFD1FFE5BCE5F8A60C51037E" bold="true" box="[190,332,1886,1908]" pageId="17" pageNumber="40">Figures 42–49.</emphasis>
|
||
Aedeagus in lateral view of
|
||
<taxonomicName id="4C1E0DEFFFD1FFE5BE01F8A60FF2037E" authorityName="Frisch" authorityYear="2022" box="[602,751,1886,1907]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="40" phylum="Arthropoda" rank="species" species="velifer">
|
||
<emphasis id="B96AAA7EFFD1FFE5BE01F8A60FF2037E" box="[602,751,1886,1907]" italics="true" pageId="17" pageNumber="40">Scopaeus velifer</emphasis>
|
||
</taxonomicName>
|
||
, holotype, Banten, Lebak: Citorek Kidul (
|
||
<emphasis id="B96AAA7EFFD1FFE5B823F8A60992037E" bold="true" box="[1144,1167,1886,1907]" pageId="17" pageNumber="40">42</emphasis>
|
||
);
|
||
<taxonomicName id="4C1E0DEFFFD1FFE5B8FFF8A60838037E" authorityName="Cameron" authorityYear="1930" box="[1188,1317,1886,1907]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="40" phylum="Arthropoda" rank="species" species="elegantulus">
|
||
<emphasis id="B96AAA7EFFD1FFE5B8FFF8A60838037E" box="[1188,1317,1886,1907]" italics="true" pageId="17" pageNumber="40">S. elegantulus</emphasis>
|
||
</taxonomicName>
|
||
, Jawa Barat, Sukabumi, Sirnarasa: Cisarua (
|
||
<emphasis id="B96AAA7EFFD1FFE5BD88F88D0CF70387" bold="true" box="[467,490,1909,1930]" pageId="17" pageNumber="40">43</emphasis>
|
||
);
|
||
<taxonomicName id="4C1E0DEFFFD1FFE5BDA7F88D0F740387" authorityName="Frisch" authorityYear="2022" box="[508,617,1909,1930]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="40" phylum="Arthropoda" rank="species" species="uncinatus">
|
||
<emphasis id="B96AAA7EFFD1FFE5BDA7F88D0F1A0387" box="[508,519,1909,1930]" italics="true" pageId="17" pageNumber="40">S</emphasis>
|
||
.
|
||
<emphasis id="B96AAA7EFFD1FFE5BE4AF88D0F740387" box="[529,617,1909,1930]" italics="true" pageId="17" pageNumber="40">uncinatus</emphasis>
|
||
</taxonomicName>
|
||
, holotype, Banten, Lebak: Citorek Kidul (
|
||
<emphasis id="B96AAA7EFFD1FFE5BFB9F88D0EE40387" bold="true" box="[994,1017,1909,1930]" pageId="17" pageNumber="40">44</emphasis>
|
||
);
|
||
<taxonomicName id="4C1E0DEFFFD1FFE5B851F88D09860387" authorityName="Frisch" authorityYear="2022" box="[1034,1179,1909,1930]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="40" phylum="Arthropoda" rank="species" species="bipenicillatus">
|
||
<emphasis id="B96AAA7EFFD1FFE5B851F88D09080387" box="[1034,1045,1909,1930]" italics="true" pageId="17" pageNumber="40">S</emphasis>
|
||
.
|
||
<emphasis id="B96AAA7EFFD1FFE5B87BF88D09860387" box="[1056,1179,1909,1930]" italics="true" pageId="17" pageNumber="40">bipenicillatus</emphasis>
|
||
</taxonomicName>
|
||
, holotype, Sulawesi Tengah, Sigi: Pagana (
|
||
<emphasis id="B96AAA7EFFD1FFE5BD60F8730C4F03AD" bold="true" box="[315,338,1931,1952]" pageId="17" pageNumber="40">45</emphasis>
|
||
);
|
||
<taxonomicName id="4C1E0DEFFFD1FFE5BD3EF8740CF203AD" authorityName="Frisch" authorityYear="2022" box="[357,495,1931,1953]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="40" phylum="Arthropoda" rank="species" species="bipectenatus">
|
||
<emphasis id="B96AAA7EFFD1FFE5BD3EF8740C6D03AC" box="[357,368,1932,1953]" italics="true" pageId="17" pageNumber="40">S</emphasis>
|
||
.
|
||
<emphasis id="B96AAA7EFFD1FFE5BD27F8730CF203AD" box="[380,495,1931,1952]" italics="true" pageId="17" pageNumber="40">bipectenatus</emphasis>
|
||
</taxonomicName>
|
||
, holotype, Sulawesi Tengah, Poso: Lake Poso (
|
||
<emphasis id="B96AAA7EFFD1FFE5BFC0F8730EAF03AD" bold="true" box="[923,946,1931,1952]" pageId="17" pageNumber="40">46</emphasis>
|
||
);
|
||
<taxonomicName id="4C1E0DEFFFD1FFE5BF9EF874092103AD" authorityName="Frisch" authorityYear="2022" box="[965,1084,1931,1953]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="40" phylum="Arthropoda" rank="species" species="cuspidatus">
|
||
<emphasis id="B96AAA7EFFD1FFE5BF9EF874092103AD" box="[965,1084,1931,1953]" italics="true" pageId="17" pageNumber="40">S. cuspidatus</emphasis>
|
||
</taxonomicName>
|
||
, holotype, Bali, Jembrana: Cekik (
|
||
<emphasis id="B96AAA7EFFD1FFE5B923F873089203AD" bold="true" box="[1400,1423,1931,1952]" pageId="17" pageNumber="40">47</emphasis>
|
||
);
|
||
<taxonomicName id="4C1E0DEFFFD1FFE5BCE6F85A0C7C03B5" authorityName="Frisch" authorityYear="2022" box="[189,353,1954,1976]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="40" phylum="Arthropoda" rank="species" species="crassipunctatus">
|
||
<emphasis id="B96AAA7EFFD1FFE5BCE6F85A0DD503BA" box="[189,200,1954,1975]" italics="true" pageId="17" pageNumber="40">S</emphasis>
|
||
.
|
||
<emphasis id="B96AAA7EFFD1FFE5BC88F85B0C7C03B5" box="[211,353,1955,1976]" italics="true" pageId="17" pageNumber="40">crassipunctatus</emphasis>
|
||
</taxonomicName>
|
||
, holotype, Jawa Barat, Sukabumi, Sirnarasa: Cisarua (
|
||
<emphasis id="B96AAA7EFFD1FFE5BF11F85A0E7C03BA" bold="true" box="[842,865,1954,1975]" pageId="17" pageNumber="40">48</emphasis>
|
||
);
|
||
<taxonomicName id="4C1E0DEFFFD1FFE5BF28F85A091F03BA" authorityName="Frisch" authorityYear="2022" box="[883,1026,1954,1976]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="40" phylum="Arthropoda" rank="species" species="sulawesianus">
|
||
<emphasis id="B96AAA7EFFD1FFE5BF28F85A0E6303BA" box="[883,894,1954,1975]" italics="true" pageId="17" pageNumber="40">S</emphasis>
|
||
.
|
||
<emphasis id="B96AAA7EFFD1FFE5BFD2F85A091F03BA" box="[905,1026,1954,1975]" italics="true" pageId="17" pageNumber="40">sulawesianus</emphasis>
|
||
</taxonomicName>
|
||
, holotype, Sulawesi Selatan: Engrekang (
|
||
<emphasis id="B96AAA7EFFD1FFE5B923F85A089203BA" bold="true" box="[1400,1423,1954,1975]" pageId="17" pageNumber="40">49</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA1766CFFD2FFE6BCF5FD840FC706DB" blockId="18.[151,764,241,988]" pageId="18" pageNumber="41">
|
||
Female: Sperm pump with ends of process segment and apophysis of chamber strongly widened; apophysis adjacent to bursal duct (
|
||
<figureCitation id="13256AE9FFD2FFE6BDC2FD460CEF06DB" box="[409,498,702,726]" captionStart="Figures 163–172" captionStartId="42.[151,224,1839,1861]" captionTargetBox="[225,1322,230,1817]" captionTargetId="figure-168@42.[157,1398,210,1820]" captionTargetPageId="42" captionText="Figures 163–172. Sperm pump (163–172) and bursa (165, 167, 170, 172) of ♀ of Scopaeus grandis, paratype, Jawa Barat, Sukabumi: Cidahu (163); S.heronifer, paratype, Jawa Barat, Sukabumi:Sirnarasa (164); S. diversilobatus, paratype, Jawa Barat, Sukabumi, Djampang Kulon (165, with lateral gonocoxal plates IX folded laterally; 166); S. cuspilobatus, paratype, Jawa Barat, Sukabumi: Sirnarasa (167, with lateral gonocoxal plates IX, 168); S. velifer, paratype, Banten, Lebak: Citorek Kidul (169); S. elegantulus, Jawa Barat, Sukabumi: Djampang Kulon (170, with lateral gonocoxal plates IX, 171), S. uncinatus, paratype, Banten, Lebak: Citorek Kidul (172). Long scale bar: 163, 164, 166, 168, 169, 171, 172; short scale bar: 165, 167, 170." figureDoi="http://doi.org/10.5281/zenodo.10724315" httpUri="https://zenodo.org/record/10724315/files/figure.png" pageId="18" pageNumber="41">Fig. 163</figureCitation>
|
||
); bursa membranous.
|
||
</paragraph>
|
||
<paragraph id="8BA1766CFFD2FFE6BCF5FD270F820715" blockId="18.[151,764,241,988]" pageId="18" pageNumber="41">
|
||
<emphasis id="B96AAA7EFFD2FFE6BCF5FD270C3006FA" bold="true" box="[174,301,735,759]" pageId="18" pageNumber="41">Phylogeny:</emphasis>
|
||
<taxonomicName id="4C1E0DEFFFD2FFE6BD60FD270F1906FA" authorityName="Frisch" authorityYear="2022" box="[315,516,735,759]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="41" phylum="Arthropoda" rank="species" species="grandis">
|
||
<emphasis id="B96AAA7EFFD2FFE6BD60FD270F1906FA" box="[315,516,735,759]" italics="true" pageId="18" pageNumber="41">Scopaeus grandis</emphasis>
|
||
</taxonomicName>
|
||
is a member of the
|
||
<taxonomicName id="4C1E0DEFFFD2FFE6BCCCFCF80C1B0715" authorityName="Oke" authorityYear="1933" box="[151,262,768,792]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="41" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B96AAA7EFFD2FFE6BCCCFCF80C1B0715" box="[151,262,768,792]" italics="true" pageId="18" pageNumber="41">S. gracilis</emphasis>
|
||
</taxonomicName>
|
||
species group (
|
||
<bibRefCitation id="EF8F0B9DFFD2FFE6BDF6FCF80F8E0715" author="Frisch, J. & D. Burckhardt & V. Wolters" box="[429,659,768,792]" pageId="18" pageNumber="41" pagination="27 - 53" refId="ref31920" refString="Frisch, J., D. Burckhardt & V. Wolters (2002): Rove beetles of the subtribe Scopaeina Mulsant & Rey (Coleoptera: Staphylinidae) in the West Palaearctic: Phylogeny, biogeography and species catalogue. - Organisms, Diversity & Evolution 2: 27 - 53." type="journal article" year="2002">Frisch et al. 2002: 39</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30425E7FFD2FFE6BCF5FCD80C6D0754" pageId="18" pageNumber="41" type="distribution">
|
||
<paragraph id="8BA1766CFFD2FFE6BCF5FCD80C6D0754" blockId="18.[151,764,241,988]" pageId="18" pageNumber="41">
|
||
<emphasis id="B96AAA7EFFD2FFE6BCF5FCD80C590735" bold="true" box="[174,324,800,824]" pageId="18" pageNumber="41">Distribution:</emphasis>
|
||
<taxonomicName id="4C1E0DEFFFD2FFE6BD08FCD90F010735" authorityName="Frisch" authorityYear="2022" box="[339,540,800,824]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="41" phylum="Arthropoda" rank="species" species="grandis">
|
||
<emphasis id="B96AAA7EFFD2FFE6BD08FCD90F010735" box="[339,540,800,824]" italics="true" pageId="18" pageNumber="41">Scopaeus grandis</emphasis>
|
||
</taxonomicName>
|
||
is hitherto known from
|
||
<collectingRegion id="49DAB88EFFD2FFE6BC88FCBA0C150754" box="[211,264,834,857]" country="Indonesia" name="Jawa Timur" pageId="18" pageNumber="41">Jawa</collectingRegion>
|
||
and
|
||
<collectingRegion id="49DAB88EFFD2FFE6BD67FCB90C710754" box="[316,364,833,857]" country="Indonesia" name="Bali" pageId="18" pageNumber="41">Bali</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30425E7FFD2FFE6BCF5FC9A0C1907D1" pageId="18" pageNumber="41" type="etymology">
|
||
<paragraph id="8BA1766CFFD2FFE6BCF5FC9A0C1907D1" blockId="18.[151,764,241,988]" pageId="18" pageNumber="41">
|
||
<emphasis id="B96AAA7EFFD2FFE6BCF5FC9A0C290777" bold="true" box="[174,308,866,890]" pageId="18" pageNumber="41">Etymology:</emphasis>
|
||
The epithet
|
||
<taxonomicName id="4C1E0DEFFFD2FFE6BD83FC9A0F360777" authorityName="Frisch" authorityYear="2022" box="[472,555,866,890]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="41" phylum="Arthropoda" rank="species" species="grandis">
|
||
<emphasis id="B96AAA7EFFD2FFE6BD83FC9A0F360777" box="[472,555,866,890]" italics="true" pageId="18" pageNumber="41">grandis</emphasis>
|
||
</taxonomicName>
|
||
(adjective, Latin: ‘big’, ‘great’) refers to the remarkable body size of this new species (
|
||
<figureCitation id="13256AE9FFD2FFE6BD75FC5C0C6107B1" box="[302,380,932,956]" captionStart="Figures 11–13" captionStartId="7.[189,262,1931,1953]" captionTargetBox="[232,1409,674,1880]" captionTargetId="figure-224@7.[189,1436,660,1912]" captionTargetPageId="7" captionText="Figures 11–13. Habitus of Scopaeus anuliflagellatus, paratype, Jawa Barat, Sukabumi: Cikaniki (11); S. grandis, paratype, Jawa Barat, Sukabumi: Cidahu (12); S. pulcher, paratype, Jawa Barat, Sukabumi: Cikaniki (13)." figureDoi="http://doi.org/10.5281/zenodo.10724244" httpUri="https://zenodo.org/record/10724244/files/figure.png" pageId="18" pageNumber="41">Fig. 12</figureCitation>
|
||
) in comparison to the majority of
|
||
<taxonomicName id="4C1E0DEFFFD2FFE6BCCCFC3D0DE007D1" box="[151,253,965,988]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96AAA7EFFD2FFE6BCCCFC3D0DE007D1" box="[151,253,965,988]" italics="true" pageId="18" pageNumber="41">Scopaeus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |