241 lines
42 KiB
XML
241 lines
42 KiB
XML
<document id="9C0B764E91FAABE98883CA377483C34B" ID-DOI="10.25674/so95iss1id311" ID-ISSN="2509-9523" ID-Zenodo-Dep="10724236" checkinTime="1709147239332" checkinUser="felipe" docAuthor="Frisch, Johannes & Narakusumo, Raden Pramesa" docDate="2022" docId="03B7C77AFFC6FFF2BF72FE6C0E760240" docLanguage="en" docName="SoilOrg.95.1.23-73.pdf" docOrigin="Soil Organisms 95 (1)" docSource="http://dx.doi.org/10.25674/so95iss1id311" docStyle="DocumentStyle:EA2ECAC87EC21C07AB805D5B9B4601F2:SoilOrg.2018-.journal_article" docStyleId="EA2ECAC87EC21C07AB805D5B9B4601F2" docStyleName="SoilOrg.2018-.journal_article" docTitle="Scopaeus riedeli Frisch 2022, spec. nov." docType="treatment" docVersion="2" lastPageNumber="29" masterDocId="FF8EBF02FFC0FFF4BC5BFFF80D1D040D" masterDocTitle="Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species" masterLastPageNumber="73" masterPageNumber="23" pageNumber="29" updateTime="1709147743548" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="420A08B796D42A5BA89F79C38165ED5B">Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species</mods:title>
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<mods:name id="6CD26C94125A076D95AC5E52BDBE8688" type="personal">
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<mods:roleTerm id="B2C01522B31F04CC1D1939DE20DBFDB4">Author</mods:roleTerm>
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<mods:namePart id="E8B86811C74512C7DCFF9374FB2CB11D">Frisch, Johannes</mods:namePart>
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<mods:roleTerm id="3890278EE820F3100D968860D182BBB2">Author</mods:roleTerm>
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<mods:namePart id="DFD958405503A96976F81C2001FBB00E">Narakusumo, Raden Pramesa</mods:namePart>
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<mods:affiliation id="395AE68E6E5205373A30AA3D7CD7BEC4">MUseUm ZoologicUm Bogoriense, Research Center for Biosystematics and EvolUtion, National Research and Innovation Agency (BRIN), Jl. Raya Jakarta-Bogor KM 46, Cibinong, West Java, 16911, Indonesia</mods:affiliation>
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<mods:title id="970116484D08583799C4C192A4A0671E">Soil Organisms</mods:title>
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<mods:part id="8AE3330FF43CD8114707612B1856FFB8">
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<mods:date id="A95CFC8BCE17D45F6AEED728B0661D44">2022</mods:date>
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<mods:number id="AE915804DF042E808C845AEBEA82B68E">2022-04-01</mods:number>
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<mods:number id="BBF8E2256674E8CF973EA16EB555EE7D">95</mods:number>
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<mods:url id="812255FE8DEA49198E5E69AA8E308DA0">http://dx.doi.org/10.25674/so95iss1id311</mods:url>
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<mods:classification id="A97714F3CA8F91BE5E8205600F957DB5">journal article</mods:classification>
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<mods:identifier id="D260002E873C503F7C6A6C7CC602FD6B" type="DOI">10.25674/so95iss1id311</mods:identifier>
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<treatment id="03B7C77AFFC6FFF2BF72FE6C0E760240" ID-DOI="http://doi.org/10.5281/zenodo.10724254" ID-Zenodo-Dep="10724254" LSID="urn:lsid:plazi:treatment:03B7C77AFFC6FFF2BF72FE6C0E760240" httpUri="http://treatment.plazi.org/id/03B7C77AFFC6FFF2BF72FE6C0E760240" lastPageNumber="29" pageId="6" pageNumber="29">
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<subSubSection id="C30425E7FFC6FFF2BF72FE6C09D405A2" box="[809,1225,404,431]" pageId="6" pageNumber="29" type="nomenclature">
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<paragraph id="8BA1766CFFC6FFF2BF72FE6C09D405A2" blockId="6.[809,1225,404,464]" box="[809,1225,404,431]" pageId="6" pageNumber="29">
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<heading id="D0E9C100FFC6FFF2BF72FE6C09D405A2" box="[809,1225,404,431]" fontSize="11" level="2" pageId="6" pageNumber="29" reason="5">
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<taxonomicName id="4C1E0DEFFFC6FFF2BF72FE6C095B05A3" authority="Frisch" authorityName="Frisch" authorityYear="2022" box="[809,1094,404,431]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="29" phylum="Arthropoda" rank="species" species="riedeli" status="sp. nov.">
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<emphasis id="B96AAA7EFFC6FFF2BF72FE6C095B05A3" bold="true" box="[809,1094,404,431]" pageId="6" pageNumber="29">
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<emphasis id="B96AAA7EFFC6FFF2BF72FE6C0EF205A2" bold="true" box="[809,1007,404,431]" italics="true" pageId="6" pageNumber="29">Scopaeus riedeli</emphasis>
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Frisch
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</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A2591705FFC6FFF2B80EFE6D09D405A2" box="[1109,1225,405,431]" pageId="6" pageNumber="29" rank="species">spec. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C30425E7FFC6FFF2BF72FE4D095605C2" box="[809,1099,437,464]" pageId="6" pageNumber="29" type="description">
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<paragraph id="8BA1766CFFC6FFF2BF72FE4D095605C2" blockId="6.[809,1225,404,464]" box="[809,1099,437,464]" pageId="6" pageNumber="29">
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(
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<figureCitation id="13256AE9FFC6FFF2BF6AFE4D0E6005C2" box="[817,893,437,463]" captionStart="Figures 1–7" captionStartId="4.[151,224,1886,1908]" captionTargetBox="[224,1327,277,1777]" captionTargetId="figure-15@4.[151,1398,256,1864]" captionTargetPageId="4" captionText="Figures 1–7. Habitus of Scopaeus javanus, Jawa Barat, Sukabumi: Kiara Dua (1); S. javanus, paralectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (2); S. batukaruensis, paratype, Bali, Tabanan: Batukaru (3); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (4); S. jacobsoni, Jawa Barat: Mt Salak (5); S. mixtus, Malaysia, Sabah: Lokan (6); S. halimunsalakensis, paratype, Jawa Barat, Sukabumi: Sirnaresmi (7)." figureDoi="http://doi.org/10.5281/zenodo.10724238" httpUri="https://zenodo.org/record/10724238/files/figure.png" pageId="6" pageNumber="29">Figs 4</figureCitation>
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,
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<figureCitation id="13256AE9FFC6FFF2BFD1FE4D0EB705C2" box="[906,938,437,463]" captionStart="Figures 27–34" captionStartId="15.[189,262,1865,1887]" captionTargetBox="[263,1347,308,1774]" captionTargetId="figure-115@15.[189,1436,278,1846]" captionTargetPageId="15" captionText="Figures 27–34. Aedeagus in lateral view of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (27); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (28); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (29); S. jacobsoni, holotype, Sumatra: Padang (30); S. mixtus, holotype, Malaysia, Sabah: Sandakan (31); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi:Sirnarasa (32); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi:Cisalimas (33); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (34)." figureDoi="http://doi.org/10.5281/zenodo.10724256" httpUri="https://zenodo.org/record/10724256/files/figure.png" pageId="6" pageNumber="29">29</figureCitation>
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,
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<figureCitation id="13256AE9FFC6FFF2BFEDFE4D091405DD" box="[950,1033,437,464]" captionStart="Figures 56–61" captionStartId="21.[189,262,1841,1863]" captionTargetBox="[261,1359,384,1727]" captionTargetId="figure-167@21.[189,1436,329,1822]" captionTargetPageId="21" captionText="Figures 56–61. Apex of aedeagus in lateral (56, 59), ventral (57, 60), dorsal view (58, 61) of Scopaeus riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (56–58); S. jacobsoni, holotype, Sumatra: Padang (59–61). Abbreviations: al – apical lobes, dl – dorsal lobe, ll – lateral lobes, mf – median foramen, s – groups of setae of lateral lobes, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: ventral margin of apical lobes (I); apex of dorsal lobe (II); apicolateral margins of apical lobes (III); ventral lobes of dorsal lobe (IV); apical extension of dextral, ventral spine of dorsal lobe (V); apical extension of sinistral, ventral spine of dorsal lobe (VI); unpaired lobe of aedeagus (VII)." figureDoi="http://doi.org/10.5281/zenodo.10724267" httpUri="https://zenodo.org/record/10724267/files/figure.png" pageId="6" pageNumber="29">56 –58</figureCitation>
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,
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<figureCitation id="13256AE9FFC6FFF2B84EFE4D095F05C2" box="[1045,1090,437,463]" captionStart="Figures 119–133" captionStartId="38.[151,224,1796,1818]" captionTargetBox="[224,1369,297,1686]" captionTargetId="figure-201@38.[151,1398,236,1777]" captionTargetPageId="38" captionText="Figures 119–133. Abdominal sternite VII (126) and abdominal sternite VIII (119–125, 127– 133) of ♂ of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (119); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (120); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (121); S. jacobsoni, Jawa Barat: Mt Salak (122); S. mixtus, Malaysia, Sabah: Lokan (123); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi: Sirnarasa (124); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi: Cisalimas (125); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (126, 127); S. spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (128); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi:Cidahu (129); S. pulcher, holotype, Jawa Barat, Sukabumi:Cikaniki (130); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (131); S. heronifer, holotype, Banten, Lebak: Majasari (132); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (133)." figureDoi="http://doi.org/10.5281/zenodo.10724304" httpUri="https://zenodo.org/record/10724304/files/figure.png" pageId="6" pageNumber="29">121</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C30425E7FFC6FFF2BF72FE010EAC067E" pageId="6" pageNumber="29" type="materials_examined">
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<paragraph id="8BA1766CFFC6FFF2BF72FE010EAC067E" blockId="6.[786,1399,504,1613]" pageId="6" pageNumber="29">
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<emphasis id="B96AAA7EFFC6FFF2BF72FE010EF8061D" bold="true" box="[809,997,504,529]" pageId="6" pageNumber="29">Type specimen:</emphasis>
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<materialsCitation id="3B767C31FFC6FFF2BFADFE000EAC067E" collectingDate="2006-08-15" collectionCode="MZB" collectorName="SO Kalianda & Riedel" country="Indonesia" elevation="1170" latitude="-5.7776" location="Rajabasa Mts" longLatPrecision="1" longitude="105.62463" municipality="South" pageId="6" pageNumber="29" specimenCount="1" specimenCount-male="1" stateProvince="Sumatera Utara" typeStatus="holotype">
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<typeStatus id="54A5C8CEFFC6FFF2BFADFE000944061D" box="[1014,1113,504,528]" pageId="6" pageNumber="29" type="holotype">Holotype</typeStatus>
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||
<specimenCount id="9D18BDE5FFC6FFF2B830FE01099C061D" box="[1131,1153,505,528]" count="1" pageId="6" pageNumber="29" type="male">♂</specimenCount>
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,
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||
<collectingCountry id="F30936FCFFC6FFF2B8CCFE00081C061D" box="[1175,1281,504,528]" name="Indonesia" pageId="6" pageNumber="29">Indonesia</collectingCountry>
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||
,
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||
<collectingRegion id="49DAB88EFFC6FFF2B94CFE01086E061C" box="[1303,1395,505,529]" country="Indonesia" name="Sumatera Utara" pageId="6" pageNumber="29">Sumatra</collectingRegion>
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||
,
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<collectingRegion id="49DAB88EFFC6FFF2BF49FDE20E66063F" box="[786,891,538,562]" country="Indonesia" name="Lampung" pageId="6" pageNumber="29">Lampung</collectingRegion>
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,
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<collectingMunicipality id="6BC5EC16FFC6FFF2BFD3FDE10EDB063C" box="[904,966,537,561]" pageId="6" pageNumber="29">South</collectingMunicipality>
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<collectingRegion id="49DAB88EFFC6FFF2BF8BFDE20924063F" box="[976,1081,538,562]" country="Indonesia" name="Lampung" pageId="6" pageNumber="29">Lampung</collectingRegion>
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,
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<collectorName id="26EB13BAFFC6FFF2B81CFDE209CA063C" box="[1095,1239,537,562]" pageId="6" pageNumber="29">SO Kalianda</collectorName>
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:
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<location id="8EC120B7FFC6FFF2B8B8FDE1086B063C" LSID="urn:lsid:plazi:treatment:03B7C77AFFC6FFF2BF72FE6C0E760240:8EC120B7FFC6FFF2B8B8FDE1086B063C" box="[1251,1398,537,561]" country="Indonesia" latitude="-5.7776" longLatPrecision="1" longitude="105.62463" municipality="South" name="Rajabasa Mts" pageId="6" pageNumber="29" stateProvince="Sumatera Utara">Rajabasa Mts</location>
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(
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<geoCoordinate id="EE2A10ABFFC6FFF2BF41FDC20EB9065F" box="[794,932,570,594]" degrees="05" direction="south" minutes="46.656" orientation="latitude" pageId="6" pageNumber="29" precision="1" value="-5.7776">05°46.656’S</geoCoordinate>
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,
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<geoCoordinate id="EE2A10ABFFC6FFF2BFEEFDC20953065F" box="[949,1102,570,594]" degrees="105" direction="east" minutes="37.478" orientation="longitude" pageId="6" pageNumber="29" precision="1" value="105.62463">105°37.478’E</geoCoordinate>
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),
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<quantity id="4CE6DB89FFC6FFF2B83CFDC209A2065F" box="[1127,1215,570,594]" metricMagnitude="3" metricUnit="m" metricValue="1.17" pageId="6" pageNumber="29" unit="m" value="1170.0">
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<elevation id="0033915FFFC6FFF2B83CFDC209A2065F" box="[1127,1215,570,594]" metricMagnitude="3" metricUnit="m" metricValue="1.17" pageId="6" pageNumber="29" unit="m" value="1170.0">1170 m</elevation>
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</quantity>
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,
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<date id="FFA050ACFFC6FFF2B88AFDC20823065F" box="[1233,1342,570,594]" pageId="6" pageNumber="29" value="2006-08-15">
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<collectingDate id="EFE4A944FFC6FFF2B88AFDC20823065F" box="[1233,1342,570,594]" pageId="6" pageNumber="29" value="2006-08-15">15.8.2006</collectingDate>
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</date>
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, leg.
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<collectorName id="26EB13BAFFC6FFF2BF49FDA30E45067E" box="[786,856,603,627]" pageId="6" pageNumber="29">Riedel</collectorName>
|
||
(
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<collectionCode id="ED0FEEA9FFC6FFF2BF3DFDA30EB8067E" box="[870,933,603,627]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="6" pageNumber="29" type="Museum">MZB</collectionCode>
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||
).
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</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="C30425E7FFC6FFF2BF72FD840EF40032" pageId="6" pageNumber="29" type="description">
|
||
<paragraph id="8BA1766CFFC6FFF2BF72FD840966071A" blockId="6.[786,1399,504,1613]" pageId="6" pageNumber="29">
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||
<emphasis id="B96AAA7EFFC6FFF2BF72FD840EAB0699" bold="true" box="[809,950,636,660]" pageId="6" pageNumber="29">Description:</emphasis>
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Habitus and coloring (
|
||
<figureCitation id="13256AE9FFC6FFF2B897FD8408090699" box="[1228,1300,636,660]" captionStart="Figures 1–7" captionStartId="4.[151,224,1886,1908]" captionTargetBox="[224,1327,277,1777]" captionTargetId="figure-15@4.[151,1398,256,1864]" captionTargetPageId="4" captionText="Figures 1–7. Habitus of Scopaeus javanus, Jawa Barat, Sukabumi: Kiara Dua (1); S. javanus, paralectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (2); S. batukaruensis, paratype, Bali, Tabanan: Batukaru (3); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (4); S. jacobsoni, Jawa Barat: Mt Salak (5); S. mixtus, Malaysia, Sabah: Lokan (6); S. halimunsalakensis, paratype, Jawa Barat, Sukabumi: Sirnaresmi (7)." figureDoi="http://doi.org/10.5281/zenodo.10724238" httpUri="https://zenodo.org/record/10724238/files/figure.png" pageId="6" pageNumber="29">Fig. 4</figureCitation>
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) like in brachypterous, dark colored
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<taxonomicName id="4C1E0DEFFFC6FFF2B83CFD65082C06B8" authorityName="Cameron" authorityYear="1936" box="[1127,1329,669,693]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="29" phylum="Arthropoda" rank="species" species="javanus">
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||
<emphasis id="B96AAA7EFFC6FFF2B83CFD65082C06B8" box="[1127,1329,669,693]" italics="true" pageId="6" pageNumber="29">Scopaeus javanus</emphasis>
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</taxonomicName>
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(
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<figureCitation id="13256AE9FFC6FFF2B910FD650E0306DB" captionStart="Figures 1–7" captionStartId="4.[151,224,1886,1908]" captionTargetBox="[224,1327,277,1777]" captionTargetId="figure-15@4.[151,1398,256,1864]" captionTargetPageId="4" captionText="Figures 1–7. Habitus of Scopaeus javanus, Jawa Barat, Sukabumi: Kiara Dua (1); S. javanus, paralectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (2); S. batukaruensis, paratype, Bali, Tabanan: Batukaru (3); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (4); S. jacobsoni, Jawa Barat: Mt Salak (5); S. mixtus, Malaysia, Sabah: Lokan (6); S. halimunsalakensis, paratype, Jawa Barat, Sukabumi: Sirnaresmi (7)." figureDoi="http://doi.org/10.5281/zenodo.10724238" httpUri="https://zenodo.org/record/10724238/files/figure.png" pageId="6" pageNumber="29">Fig. 1</figureCitation>
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) and
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<taxonomicName id="4C1E0DEFFFC6FFF2BF34FD46093406DB" authorityName="Frisch" authorityYear="2022" box="[879,1065,702,726]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="29" phylum="Arthropoda" rank="species" species="batukaruensis">
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<emphasis id="B96AAA7EFFC6FFF2BF34FD46093406DB" box="[879,1065,702,726]" italics="true" pageId="6" pageNumber="29">S. batukaruensis</emphasis>
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</taxonomicName>
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(
|
||
<figureCitation id="13256AE9FFC6FFF2B81AFD46099606DB" box="[1089,1163,702,726]" captionStart="Figures 1–7" captionStartId="4.[151,224,1886,1908]" captionTargetBox="[224,1327,277,1777]" captionTargetId="figure-15@4.[151,1398,256,1864]" captionTargetPageId="4" captionText="Figures 1–7. Habitus of Scopaeus javanus, Jawa Barat, Sukabumi: Kiara Dua (1); S. javanus, paralectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (2); S. batukaruensis, paratype, Bali, Tabanan: Batukaru (3); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (4); S. jacobsoni, Jawa Barat: Mt Salak (5); S. mixtus, Malaysia, Sabah: Lokan (6); S. halimunsalakensis, paratype, Jawa Barat, Sukabumi: Sirnaresmi (7)." figureDoi="http://doi.org/10.5281/zenodo.10724238" httpUri="https://zenodo.org/record/10724238/files/figure.png" pageId="6" pageNumber="29">Fig. 3</figureCitation>
|
||
).
|
||
<typeStatus id="54A5C8CEFFC6FFF2B8F2FD46081106DB" box="[1193,1292,702,726]" pageId="6" pageNumber="29" type="holotype">Holotype</typeStatus>
|
||
(
|
||
<figureCitation id="13256AE9FFC6FFF2B97EFD46087306DB" box="[1317,1390,702,726]" captionStart="Figures 1–7" captionStartId="4.[151,224,1886,1908]" captionTargetBox="[224,1327,277,1777]" captionTargetId="figure-15@4.[151,1398,256,1864]" captionTargetPageId="4" captionText="Figures 1–7. Habitus of Scopaeus javanus, Jawa Barat, Sukabumi: Kiara Dua (1); S. javanus, paralectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (2); S. batukaruensis, paratype, Bali, Tabanan: Batukaru (3); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (4); S. jacobsoni, Jawa Barat: Mt Salak (5); S. mixtus, Malaysia, Sabah: Lokan (6); S. halimunsalakensis, paratype, Jawa Barat, Sukabumi: Sirnaresmi (7)." figureDoi="http://doi.org/10.5281/zenodo.10724238" httpUri="https://zenodo.org/record/10724238/files/figure.png" pageId="6" pageNumber="29">Fig. 4</figureCitation>
|
||
) brachypterous, without palisade fringe. Total body length
|
||
<quantity id="4CE6DB89FFC6FFF2BF49FCF80E7A071A" box="[786,871,768,792]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="6" pageNumber="29" unit="mm" value="2.9">2.9 mm</quantity>
|
||
; forebody length
|
||
<quantity id="4CE6DB89FFC6FFF2B879FD07096A071A" box="[1058,1143,767,791]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="29" unit="mm" value="1.6">1.6 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA1766CFFC6FFF2BF72FCD809B00012" blockId="6.[786,1399,504,1613]" pageId="6" pageNumber="29">
|
||
Male: Abdominal sternites VII and VIII (
|
||
<figureCitation id="13256AE9FFC6FFF2B8AAFCD808520734" box="[1265,1359,800,825]" captionStart="Figures 119–133" captionStartId="38.[151,224,1796,1818]" captionTargetBox="[224,1369,297,1686]" captionTargetId="figure-201@38.[151,1398,236,1777]" captionTargetPageId="38" captionText="Figures 119–133. Abdominal sternite VII (126) and abdominal sternite VIII (119–125, 127– 133) of ♂ of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (119); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (120); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (121); S. jacobsoni, Jawa Barat: Mt Salak (122); S. mixtus, Malaysia, Sabah: Lokan (123); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi: Sirnarasa (124); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi: Cisalimas (125); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (126, 127); S. spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (128); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi:Cidahu (129); S. pulcher, holotype, Jawa Barat, Sukabumi:Cikaniki (130); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (131); S. heronifer, holotype, Banten, Lebak: Majasari (132); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (133)." figureDoi="http://doi.org/10.5281/zenodo.10724304" httpUri="https://zenodo.org/record/10724304/files/figure.png" pageId="6" pageNumber="29">Fig. 121</figureCitation>
|
||
) as in
|
||
<taxonomicName id="4C1E0DEFFFC6FFF2BF74FCBA0E820754" authorityName="Cameron" authorityYear="1936" box="[815,927,834,857]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="29" phylum="Arthropoda" rank="species" species="javanus">
|
||
<emphasis id="B96AAA7EFFC6FFF2BF74FCBA0E820754" box="[815,927,834,857]" italics="true" pageId="6" pageNumber="29">S. javanus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13256AE9FFC6FFF2BFF4FCB909170754" box="[943,1034,833,857]" captionStart="Figures 119–133" captionStartId="38.[151,224,1796,1818]" captionTargetBox="[224,1369,297,1686]" captionTargetId="figure-201@38.[151,1398,236,1777]" captionTargetPageId="38" captionText="Figures 119–133. Abdominal sternite VII (126) and abdominal sternite VIII (119–125, 127– 133) of ♂ of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (119); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (120); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (121); S. jacobsoni, Jawa Barat: Mt Salak (122); S. mixtus, Malaysia, Sabah: Lokan (123); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi: Sirnarasa (124); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi: Cisalimas (125); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (126, 127); S. spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (128); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi:Cidahu (129); S. pulcher, holotype, Jawa Barat, Sukabumi:Cikaniki (130); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (131); S. heronifer, holotype, Banten, Lebak: Majasari (132); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (133)." figureDoi="http://doi.org/10.5281/zenodo.10724304" httpUri="https://zenodo.org/record/10724304/files/figure.png" pageId="6" pageNumber="29">Fig. 119</figureCitation>
|
||
) and
|
||
<taxonomicName id="4C1E0DEFFFC6FFF2B812FCBA09E10754" authorityName="Frisch" authorityYear="2022" box="[1097,1276,833,857]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="29" phylum="Arthropoda" rank="species" species="batukaruensis">
|
||
<emphasis id="B96AAA7EFFC6FFF2B812FCBA09E10754" box="[1097,1276,833,857]" italics="true" pageId="6" pageNumber="29">S. batukaruensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13256AE9FFC6FFF2B950FCB908760754" box="[1291,1387,833,857]" captionStart="Figures 119–133" captionStartId="38.[151,224,1796,1818]" captionTargetBox="[224,1369,297,1686]" captionTargetId="figure-201@38.[151,1398,236,1777]" captionTargetPageId="38" captionText="Figures 119–133. Abdominal sternite VII (126) and abdominal sternite VIII (119–125, 127– 133) of ♂ of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (119); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (120); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (121); S. jacobsoni, Jawa Barat: Mt Salak (122); S. mixtus, Malaysia, Sabah: Lokan (123); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi: Sirnarasa (124); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi: Cisalimas (125); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (126, 127); S. spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (128); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi:Cidahu (129); S. pulcher, holotype, Jawa Barat, Sukabumi:Cikaniki (130); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (131); S. heronifer, holotype, Banten, Lebak: Majasari (132); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (133)." figureDoi="http://doi.org/10.5281/zenodo.10724304" httpUri="https://zenodo.org/record/10724304/files/figure.png" pageId="6" pageNumber="29">Fig. 120</figureCitation>
|
||
). Aedeagus (
|
||
<figureCitation id="13256AE9FFC6FFF2BFD6FC9A0EFE0777" box="[909,995,866,890]" captionStart="Figures 27–34" captionStartId="15.[189,262,1865,1887]" captionTargetBox="[263,1347,308,1774]" captionTargetId="figure-115@15.[189,1436,278,1846]" captionTargetPageId="15" captionText="Figures 27–34. Aedeagus in lateral view of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (27); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (28); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (29); S. jacobsoni, holotype, Sumatra: Padang (30); S. mixtus, holotype, Malaysia, Sabah: Sandakan (31); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi:Sirnarasa (32); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi:Cisalimas (33); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (34)." figureDoi="http://doi.org/10.5281/zenodo.10724256" httpUri="https://zenodo.org/record/10724256/files/figure.png" pageId="6" pageNumber="29">Figs 29</figureCitation>
|
||
,
|
||
<figureCitation id="13256AE9FFC6FFF2BFAAFC9A09210777" box="[1009,1084,866,890]" captionStart="Figures 56–61" captionStartId="21.[189,262,1841,1863]" captionTargetBox="[261,1359,384,1727]" captionTargetId="figure-167@21.[189,1436,329,1822]" captionTargetPageId="21" captionText="Figures 56–61. Apex of aedeagus in lateral (56, 59), ventral (57, 60), dorsal view (58, 61) of Scopaeus riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (56–58); S. jacobsoni, holotype, Sumatra: Padang (59–61). Abbreviations: al – apical lobes, dl – dorsal lobe, ll – lateral lobes, mf – median foramen, s – groups of setae of lateral lobes, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: ventral margin of apical lobes (I); apex of dorsal lobe (II); apicolateral margins of apical lobes (III); ventral lobes of dorsal lobe (IV); apical extension of dextral, ventral spine of dorsal lobe (V); apical extension of sinistral, ventral spine of dorsal lobe (VI); unpaired lobe of aedeagus (VII)." figureDoi="http://doi.org/10.5281/zenodo.10724267" httpUri="https://zenodo.org/record/10724267/files/figure.png" pageId="6" pageNumber="29">56 – 58</figureCitation>
|
||
)
|
||
<quantity id="4CE6DB89FFC6FFF2B816FC9A09830777" box="[1101,1182,866,890]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="29" unit="mm" value="0.7">0.7 mm</quantity>
|
||
long, distinguished from that of
|
||
<taxonomicName id="4C1E0DEFFFC6FFF2BFC1FC7B09150796" authorityName="Cameron" authorityYear="1936" box="[922,1032,899,923]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="29" phylum="Arthropoda" rank="species" species="javanus">
|
||
<emphasis id="B96AAA7EFFC6FFF2BFC1FC7B09150796" box="[922,1032,899,923]" italics="true" pageId="6" pageNumber="29">S. javanus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1E0DEFFFC6FFF2B865FC7B09F20796" authorityName="Frisch" authorityYear="2022" box="[1086,1263,899,923]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="29" phylum="Arthropoda" rank="species" species="batukaruensis">
|
||
<emphasis id="B96AAA7EFFC6FFF2B865FC7B09F20796" box="[1086,1263,899,923]" italics="true" pageId="6" pageNumber="29">S. batukaruensis</emphasis>
|
||
</taxonomicName>
|
||
by ventrally inemarginate (
|
||
<figureCitation id="13256AE9FFC6FFF2BFEBFC5C091907B1" box="[944,1028,932,956]" captionStart="Figures 56–61" captionStartId="21.[189,262,1841,1863]" captionTargetBox="[261,1359,384,1727]" captionTargetId="figure-167@21.[189,1436,329,1822]" captionTargetPageId="21" captionText="Figures 56–61. Apex of aedeagus in lateral (56, 59), ventral (57, 60), dorsal view (58, 61) of Scopaeus riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (56–58); S. jacobsoni, holotype, Sumatra: Padang (59–61). Abbreviations: al – apical lobes, dl – dorsal lobe, ll – lateral lobes, mf – median foramen, s – groups of setae of lateral lobes, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: ventral margin of apical lobes (I); apex of dorsal lobe (II); apicolateral margins of apical lobes (III); ventral lobes of dorsal lobe (IV); apical extension of dextral, ventral spine of dorsal lobe (V); apical extension of sinistral, ventral spine of dorsal lobe (VI); unpaired lobe of aedeagus (VII)." figureDoi="http://doi.org/10.5281/zenodo.10724267" httpUri="https://zenodo.org/record/10724267/files/figure.png" pageId="6" pageNumber="29">Fig. 56</figureCitation>
|
||
: arrow I), apicolaterally widened apical lobes (
|
||
<figureCitation id="13256AE9FFC6FFF2BFFEFC3D0EEA07D0" box="[933,1015,965,989]" captionStart="Figures 56–61" captionStartId="21.[189,262,1841,1863]" captionTargetBox="[261,1359,384,1727]" captionTargetId="figure-167@21.[189,1436,329,1822]" captionTargetPageId="21" captionText="Figures 56–61. Apex of aedeagus in lateral (56, 59), ventral (57, 60), dorsal view (58, 61) of Scopaeus riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (56–58); S. jacobsoni, holotype, Sumatra: Padang (59–61). Abbreviations: al – apical lobes, dl – dorsal lobe, ll – lateral lobes, mf – median foramen, s – groups of setae of lateral lobes, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: ventral margin of apical lobes (I); apex of dorsal lobe (II); apicolateral margins of apical lobes (III); ventral lobes of dorsal lobe (IV); apical extension of dextral, ventral spine of dorsal lobe (V); apical extension of sinistral, ventral spine of dorsal lobe (VI); unpaired lobe of aedeagus (VII)." figureDoi="http://doi.org/10.5281/zenodo.10724267" httpUri="https://zenodo.org/record/10724267/files/figure.png" pageId="6" pageNumber="29">Fig. 57</figureCitation>
|
||
: arrow
|
||
<collectionCode id="ED0FEEA9FFC6FFF2B817FC3D097407D1" box="[1100,1129,965,988]" pageId="6" pageNumber="29">III</collectionCode>
|
||
,
|
||
<figureCitation id="13256AE9FFC6FFF2B82CFC3D09D407D0" box="[1143,1225,965,989]" captionStart="Figures 56–61" captionStartId="21.[189,262,1841,1863]" captionTargetBox="[261,1359,384,1727]" captionTargetId="figure-167@21.[189,1436,329,1822]" captionTargetPageId="21" captionText="Figures 56–61. Apex of aedeagus in lateral (56, 59), ventral (57, 60), dorsal view (58, 61) of Scopaeus riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (56–58); S. jacobsoni, holotype, Sumatra: Padang (59–61). Abbreviations: al – apical lobes, dl – dorsal lobe, ll – lateral lobes, mf – median foramen, s – groups of setae of lateral lobes, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: ventral margin of apical lobes (I); apex of dorsal lobe (II); apicolateral margins of apical lobes (III); ventral lobes of dorsal lobe (IV); apical extension of dextral, ventral spine of dorsal lobe (V); apical extension of sinistral, ventral spine of dorsal lobe (VI); unpaired lobe of aedeagus (VII)." figureDoi="http://doi.org/10.5281/zenodo.10724267" httpUri="https://zenodo.org/record/10724267/files/figure.png" pageId="6" pageNumber="29">Fig. 58</figureCitation>
|
||
); different from aedeagus of
|
||
<taxonomicName id="4C1E0DEFFFC6FFF2BFCCFC1E091807F0" authorityName="Cameron" authorityYear="1936" box="[919,1029,998,1021]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="29" phylum="Arthropoda" rank="species" species="javanus">
|
||
<emphasis id="B96AAA7EFFC6FFF2BFCCFC1E091807F0" box="[919,1029,998,1021]" italics="true" pageId="6" pageNumber="29">S. javanus</emphasis>
|
||
</taxonomicName>
|
||
moreover by narrow, not lobiform apex of dorsal lobe (
|
||
<figureCitation id="13256AE9FFC6FFF2BFB7FBFF09210013" box="[1004,1084,1030,1055]" captionStart="Figures 56–61" captionStartId="21.[189,262,1841,1863]" captionTargetBox="[261,1359,384,1727]" captionTargetId="figure-167@21.[189,1436,329,1822]" captionTargetPageId="21" captionText="Figures 56–61. Apex of aedeagus in lateral (56, 59), ventral (57, 60), dorsal view (58, 61) of Scopaeus riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (56–58); S. jacobsoni, holotype, Sumatra: Padang (59–61). Abbreviations: al – apical lobes, dl – dorsal lobe, ll – lateral lobes, mf – median foramen, s – groups of setae of lateral lobes, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: ventral margin of apical lobes (I); apex of dorsal lobe (II); apicolateral margins of apical lobes (III); ventral lobes of dorsal lobe (IV); apical extension of dextral, ventral spine of dorsal lobe (V); apical extension of sinistral, ventral spine of dorsal lobe (VI); unpaired lobe of aedeagus (VII)." figureDoi="http://doi.org/10.5281/zenodo.10724267" httpUri="https://zenodo.org/record/10724267/files/figure.png" pageId="6" pageNumber="29">Fig. 56</figureCitation>
|
||
: arrow
|
||
<collectionCode id="ED0FEEA9FFC6FFF2B8D7FBFF09BC0012" box="[1164,1185,1031,1055]" pageId="6" pageNumber="29">II</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA1766CFFC6FFF2BF72FBDF0EF40032" blockId="6.[786,1399,504,1613]" box="[809,1001,1063,1087]" pageId="6" pageNumber="29">Female unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30425E7FFC6FFF2BF72FBB00ED9008C" pageId="6" pageNumber="29" type="distribution">
|
||
<paragraph id="8BA1766CFFC6FFF2BF72FBB00ED9008C" blockId="6.[786,1399,504,1613]" pageId="6" pageNumber="29">
|
||
<emphasis id="B96AAA7EFFC6FFF2BF72FBB00EA0006D" bold="true" box="[809,957,1096,1120]" pageId="6" pageNumber="29">Distribution:</emphasis>
|
||
<taxonomicName id="4C1E0DEFFFC6FFF2BF98FBB1096E006D" authorityName="Frisch" authorityYear="2022" box="[963,1139,1096,1120]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="29" phylum="Arthropoda" rank="species" species="riedeli">
|
||
<emphasis id="B96AAA7EFFC6FFF2BF98FBB1096E006D" box="[963,1139,1096,1120]" italics="true" pageId="6" pageNumber="29">Scopaeus riedeli</emphasis>
|
||
</taxonomicName>
|
||
is known from the south of
|
||
<collectingRegion id="49DAB88EFFC6FFF2BF74FB910E95008C" box="[815,904,1129,1153]" country="Indonesia" name="Sumatera Utara" pageId="6" pageNumber="29">Sumatra</collectingRegion>
|
||
only.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30425E7FFC6FFF2BF72FB720EFF014A" pageId="6" pageNumber="29" type="etymology">
|
||
<paragraph id="8BA1766CFFC6FFF2BF72FB720EFF014A" blockId="6.[786,1399,504,1613]" pageId="6" pageNumber="29">
|
||
<emphasis id="B96AAA7EFFC6FFF2BF72FB720EB000AF" bold="true" box="[809,941,1162,1186]" pageId="6" pageNumber="29">Etymology:</emphasis>
|
||
The epithet
|
||
<taxonomicName id="4C1E0DEFFFC6FFF2B806FB7209BE00AF" authorityName="Frisch" authorityYear="2022" box="[1117,1187,1162,1186]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="29" phylum="Arthropoda" rank="species" species="riedeli">
|
||
<emphasis id="B96AAA7EFFC6FFF2B806FB7209BE00AF" box="[1117,1187,1162,1186]" italics="true" pageId="6" pageNumber="29">riedeli</emphasis>
|
||
</taxonomicName>
|
||
(Latinized noun, derived from the surname ‘Riedel’, genitive, singular) is chosen in honor of Alexander Riedel, specialist of Indonesian
|
||
<taxonomicName id="4C1E0DEFFFC6FFF2BFC3FB1509320108" box="[920,1071,1261,1285]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="29" phylum="Arthropoda" rank="family">Curculionidae</taxonomicName>
|
||
at Staatliches Museum für Naturkunde Karlsruhe,
|
||
<collectingCountry id="F30936FCFFC6FFF2B854FAF6096E012B" box="[1039,1139,1294,1318]" name="Germany" pageId="6" pageNumber="29">Germany</collectingCountry>
|
||
, who collected this new species in Sumatra.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30425E7FFC6FFF2BF72FAB70E760240" pageId="6" pageNumber="29" type="discussion">
|
||
<paragraph id="8BA1766CFFC6FFF2BF72FAB70E760240" blockId="6.[786,1399,504,1613]" pageId="6" pageNumber="29">
|
||
<emphasis id="B96AAA7EFFC6FFF2BF72FAB70EB9016A" bold="true" box="[809,932,1359,1383]" pageId="6" pageNumber="29">Comment:</emphasis>
|
||
<taxonomicName id="4C1E0DEFFFC6FFF2BFE0FAA8098D016A" authorityName="Cameron" authorityYear="1936" box="[955,1168,1360,1383]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="29" phylum="Arthropoda" rank="species" species="javanus">
|
||
<emphasis id="B96AAA7EFFC6FFF2BFE0FAA8098D016A" box="[955,1168,1360,1383]" italics="true" pageId="6" pageNumber="29">Scopaeus javanus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1E0DEFFFC6FFF2B8F5FAA8086B016A" authorityName="Frisch" authorityYear="2022" box="[1198,1398,1359,1383]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="29" phylum="Arthropoda" rank="species" species="batukaruensis">
|
||
<emphasis id="B96AAA7EFFC6FFF2B8F5FAA8086B016A" box="[1198,1398,1359,1383]" italics="true" pageId="6" pageNumber="29">S. batukaruensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1E0DEFFFC6FFF2BF13FA890EA90185" authorityName="Frisch" authorityYear="2022" box="[840,948,1392,1416]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="29" phylum="Arthropoda" rank="species" species="riedeli">
|
||
<emphasis id="B96AAA7EFFC6FFF2BF13FA890EA90185" box="[840,948,1392,1416]" italics="true" pageId="6" pageNumber="29">S. riedeli</emphasis>
|
||
</taxonomicName>
|
||
form a species group with allopatric distribution in the Sunda Islands. The species are linked by the similar habitus, wing-dimorphism (
|
||
<figureCitation id="13256AE9FFC6FFF2B95AFA4A087201C7" box="[1281,1391,1458,1482]" captionStart="Figures 1–7" captionStartId="4.[151,224,1886,1908]" captionTargetBox="[224,1327,277,1777]" captionTargetId="figure-15@4.[151,1398,256,1864]" captionTargetPageId="4" captionText="Figures 1–7. Habitus of Scopaeus javanus, Jawa Barat, Sukabumi: Kiara Dua (1); S. javanus, paralectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (2); S. batukaruensis, paratype, Bali, Tabanan: Batukaru (3); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (4); S. jacobsoni, Jawa Barat: Mt Salak (5); S. mixtus, Malaysia, Sabah: Lokan (6); S. halimunsalakensis, paratype, Jawa Barat, Sukabumi: Sirnaresmi (7)." figureDoi="http://doi.org/10.5281/zenodo.10724238" httpUri="https://zenodo.org/record/10724238/files/figure.png" pageId="6" pageNumber="29">Figs 1 – 4</figureCitation>
|
||
) and the characteristic shape of the aedeagus with a strongly ventrad curved, from the apical lobes strongly projecting, distally extended dorsal lobe (
|
||
<figureCitation id="13256AE9FFC6FFF2B8B7F9ED086F0220" box="[1260,1394,1557,1581]" captionStart="Figures 27–34" captionStartId="15.[189,262,1865,1887]" captionTargetBox="[263,1347,308,1774]" captionTargetId="figure-115@15.[189,1436,278,1846]" captionTargetPageId="15" captionText="Figures 27–34. Aedeagus in lateral view of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (27); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (28); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (29); S. jacobsoni, holotype, Sumatra: Padang (30); S. mixtus, holotype, Malaysia, Sabah: Sandakan (31); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi:Sirnarasa (32); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi:Cisalimas (33); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (34)." figureDoi="http://doi.org/10.5281/zenodo.10724256" httpUri="https://zenodo.org/record/10724256/files/figure.png" pageId="6" pageNumber="29">Figs 27 – 29</figureCitation>
|
||
,
|
||
<figureCitation id="13256AE9FFC6FFF2BF49F9CD0E420240" box="[786,863,1589,1613]" captionStart-0="Figures 50–55" captionStart-1="Figures 56–61" captionStartId-0="20.[150,223,1886,1908]" captionStartId-1="21.[189,262,1841,1863]" captionTargetBox-0="[222,1327,582,1831]" captionTargetBox-1="[261,1359,384,1727]" captionTargetId-0="figure-271@20.[151,1398,568,1866]" captionTargetId-1="figure-167@21.[189,1436,329,1822]" captionTargetPageId-0="20" captionTargetPageId-1="21" captionText-0="Figures 50–55. Apex of aedeagus in lateral (50, 53), ventral (51, 54), dorsal view (52, 55) of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (50–52); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (53–55). Abbreviations: al – apical lobes, dl – dorsal lobe, ll – lateral lobes, mf – median foramen, tr – transverse ridge of median foramen. Arrows: apex of dorsal lobe (I, IV), apicoventral emargination of apical lobes (III), apicolateral margin of apical lobes (II, V)." captionText-1="Figures 56–61. Apex of aedeagus in lateral (56, 59), ventral (57, 60), dorsal view (58, 61) of Scopaeus riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (56–58); S. jacobsoni, holotype, Sumatra: Padang (59–61). Abbreviations: al – apical lobes, dl – dorsal lobe, ll – lateral lobes, mf – median foramen, s – groups of setae of lateral lobes, tr – transverse ridge of median foramen, vl – ventral lobe. Arrows: ventral margin of apical lobes (I); apex of dorsal lobe (II); apicolateral margins of apical lobes (III); ventral lobes of dorsal lobe (IV); apical extension of dextral, ventral spine of dorsal lobe (V); apical extension of sinistral, ventral spine of dorsal lobe (VI); unpaired lobe of aedeagus (VII)." figureDoi-0="http://doi.org/10.5281/zenodo.10724264" figureDoi-1="http://doi.org/10.5281/zenodo.10724267" httpUri-0="https://zenodo.org/record/10724264/files/figure.png" httpUri-1="https://zenodo.org/record/10724267/files/figure.png" pageId="6" pageNumber="29">50 – 58</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |