306 lines
29 KiB
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306 lines
29 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.3664830" ID-GBIF-Dataset="dd032848-7430-45d0-85d9-e528f25c83bb" ID-Zenodo-Dep="3664830" checkinTime="1581022300054" checkinUser="plazi" docAuthor="Perrichot, Vincent, Wang, Bo & Barden, Phillip" docDate="2020" docId="03A3850A9F58FFB8FFAEF9310D5F7FC6" docLanguage="en" docName="createceousResearch.109.104.381.pdf" docOrigin="Cretaceous Research 109" docStyle="DocumentStyle{}" docTitle="New remarkable hell ants (Formicidae: Haidomyrmecinae stat. nov.) from mid-Cretaceous amber of northern Myanmar" docType="key" docVersion="5" lastPageId="16" lastPageNumber="17" masterDocId="FF9AFD729F57FFA8FFF6FFD90E4F796E" masterDocTitle="New remarkable hell ants (Formicidae: Haidomyrmecinae stat. nov.) from mid-Cretaceous amber of northern Myanmar" masterLastPageNumber="18" masterPageNumber="1" pageId="15" pageNumber="16" updateTime="1643457689448" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New remarkable hell ants (Formicidae: Haidomyrmecinae stat. nov.) from mid-Cretaceous amber of northern Myanmar</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Perrichot, Vincent</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wang, Bo</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Barden, Phillip</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Cretaceous Research</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2020-01-10</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>109</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>18</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1016/j.cretres.2020.104381</mods:identifier>
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<mods:identifier type="GBIF-Dataset">dd032848-7430-45d0-85d9-e528f25c83bb</mods:identifier>
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<mods:identifier type="Zenodo-Dep">3653072</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3664830" ID-Zenodo-Dep="3664830" LSID="urn:lsid:plazi:treatment:03A3850A9F58FFB8FFAEF9310D5F7FC6" httpUri="http://treatment.plazi.org/id/03A3850A9F58FFB8FFAEF9310D5F7FC6" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
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<subSubSection box="[88,545,1768,1788]" pageId="15" pageNumber="16" type="nomenclature">
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<paragraph blockId="15.[88,545,1768,1788]" box="[88,545,1768,1788]" pageId="15" pageNumber="16">
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<emphasis box="[88,545,1768,1788]" italics="true" pageId="15" pageNumber="16">Key to genera and species of Haidomyrmecinae</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="key">
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<paragraph blockId="15.[89,757,1825,1900]" pageId="15" pageNumber="16">
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1. Clypeus posteriorly with a slightly elevated setose lobe positioned just ventrad or between antennal insertion
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<emphasis box="[114,644,1881,1900]" italics="true" pageId="15" pageNumber="16">………………………………………………………………</emphasis>
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..
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<emphasis box="[654,720,1881,1900]" italics="true" pageId="15" pageNumber="16">………</emphasis>
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... 2
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</paragraph>
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<paragraph blockId="15.[114,757,1909,1984]" pageId="15" pageNumber="16">
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Clypeus posteriorly projecting forward into a cephalic horn arising ventrad, between, or dorsad antennal insertion
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<emphasis box="[114,644,1964,1983]" italics="true" pageId="15" pageNumber="16">………………………………………………………………</emphasis>
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..
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<emphasis box="[654,720,1964,1983]" italics="true" pageId="15" pageNumber="16">………</emphasis>
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... 5
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</paragraph>
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<paragraph blockId="15.[805,1475,178,1959]" pageId="15" pageNumber="16">
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2. Antenna compact, with flagellomeres short (<2 as long as broad); first flagellomere as long as or longer than second one
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<emphasis box="[837,1455,234,253]" italics="true" pageId="15" pageNumber="16">…………………………………………………………………………</emphasis>
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3
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</paragraph>
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<paragraph blockId="15.[805,1475,178,1959]" pageId="15" pageNumber="16">
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Antenna elongate, with flagellomeres long (>2 as long as broad); first flagellomere shorter than second one
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<emphasis italics="true" pageId="15" pageNumber="16">
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………… ………………………………………………
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<taxonomicName authorityName="Dlussky" authorityYear="1996" box="[1229,1366,317,337]" class="Hexapoda" family="Formicidae" genus="Haidomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">Haidomyrmex</taxonomicName>
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</emphasis>
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species: 4
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</paragraph>
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<paragraph blockId="15.[805,1475,178,1959]" pageId="15" pageNumber="16">
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3. All flagellomeres nearly of same length; clypeal setose pad positioned anterior to antennal insertion; ocelli present; two metatibial spurs
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<emphasis italics="true" pageId="15" pageNumber="16">
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<emphasis box="[1001,1376,405,424]" italics="true" pageId="15" pageNumber="16">…………………………………………………………</emphasis>
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…………………
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</emphasis>
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<taxonomicName authority="Perrichot et al., 2008" authorityName="Perrichot" authorityYear="2008" box="[968,1471,432,453]" class="Hexapoda" family="Formicidae" genus="Haidomyrmodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="mammuthus">
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<emphasis box="[968,1256,432,452]" italics="true" pageId="15" pageNumber="16">
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<taxonomicName box="[968,1127,432,452]" genus="Haidomyrmodes" pageId="15" pageNumber="16" rank="genus">Haidomyrmodes</taxonomicName>
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mammuthus
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</emphasis>
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<bibRefCitation author="Perrichot, V. & Lacau, S. & Neraudeau, D. & Nel, A." box="[1261,1471,434,453]" journalOrPublisher="Naturwissenschaften" pageId="15" pageNumber="16" pagination="90" part="95" refId="ref16486" refString="Perrichot, V., Lacau, S., Neraudeau, D., Nel, A., 2008 b. Fossil evidence for the early ant evolution. Naturwissenschaften 95, 85 e 90. https: // doi. org / 10.1007 / s 00114 - 007 - 0301 - 8." title="Fossil evidence for the early ant evolution" type="journal article" year="2008">Perrichot et al., 2008</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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<paragraph blockId="15.[805,1475,178,1959]" pageId="15" pageNumber="16">
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Flagellomere I longest of basal four flagellar articles; clypeal setose pad positioned slightly posterior to antennal insertion; ocelli absent; one metatibial spur
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<emphasis italics="true" pageId="15" pageNumber="16">……………………… …</emphasis>
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..
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<emphasis box="[863,885,545,564]" italics="true" pageId="15" pageNumber="16">…</emphasis>
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..
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<emphasis box="[896,1155,544,564]" italics="true" pageId="15" pageNumber="16">
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……
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<taxonomicName authority="McKellar, Glasier & Engel, 2013" authorityName="McKellar, Glasier & Engel" authorityYear="2013" box="[945,1464,544,564]" class="Hexapoda" family="Formicidae" genus="Haidoterminus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="cippus">
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<taxonomicName box="[945,1089,544,564]" genus="Haidoterminus" pageId="15" pageNumber="16" rank="genus">Haidoterminus</taxonomicName>
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cippus
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<bibRefCitation author="McKellar, R. C. & Glasier, J. R. N. & Engel, M. S." box="[1155,1464,545,564]" journalOrPublisher="The Canadian Entomologist" pageId="15" pageNumber="16" pagination="465" part="145" refId="ref16172" refString="McKellar, R. C., Glasier, J. R. N., Engel, M. S., 2013 a. A new trap-jawed ant (Hymenoptera: Formicidae: Haidomyrmecini) from Canadian Late Cretaceous amber. The Canadian Entomologist 145, 454 e 465. https: // doi. org / 10.4039 / tce. 2013.23." title="A new trap-jawed ant (Hymenoptera: Formicidae: Haidomyrmecini) from Canadian Late Cretaceous amber" type="journal article" year="2013">McKellar, Glasier & Engel, 2013</bibRefCitation>
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</taxonomicName>
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</emphasis>
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.
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</paragraph>
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<paragraph blockId="15.[805,1475,178,1959]" pageId="15" pageNumber="16">
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4. Flagellomere II longest of basal three flagellar articles; frontal triangle moderately elevated; vertex, ventral margin of mandibles, and antennomeres densely setose; mandibles with medioventral blade bearing at least one large apical tooth, with tip of apical portion tapered to acute point, smooth; ocelli reduced to pit-like traces; subpetiolar process present
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<emphasis box="[837,1080,740,759]" italics="true" pageId="15" pageNumber="16">……………………………</emphasis>
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.
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<emphasis box="[1084,1335,739,759]" italics="true" pageId="15" pageNumber="16">
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…
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<taxonomicName authority="Dlussky, 1996" authorityName="Dlussky" authorityYear="1996" box="[1111,1474,739,760]" class="Hexapoda" family="Formicidae" genus="Haidomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="cerberus">
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<taxonomicName authority="Dlussky, 1996" authorityName="Dlussky" authorityYear="1996" box="[1111,1248,739,759]" class="Insecta" family="Formicidae" genus="Haidomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">Haidomyrmex</taxonomicName>
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cerberus
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<bibRefCitation author="Dlussky, G. M." box="[1339,1474,741,760]" journalOrPublisher="Paleontological Journal" pageId="15" pageNumber="16" pagination="454" part="30" refId="ref15710" refString="Dlussky, G. M., 1996. Ants (Hymenoptera: Formicidae) from Burmese amber. Paleontological Journal 30, 449 e 454. Translated from Paleontologicheskii Zhurnal 1996 (3), 83 e 89." title="Ants (Hymenoptera: Formicidae) from Burmese amber" type="journal article" year="1996">Dlussky,1996</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</paragraph>
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<paragraph blockId="15.[805,1475,178,1959]" pageId="15" pageNumber="16">
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Flagellomere II longest; frontal triangle strongly elevated; vertex, ventral margin of mandibles, and antennomeres mostly glabrous; mandibles with medioventral blades asymmetrical, left one bearing three small teeth, right one bearing two larger teeth; tip of apical portion tapered to sharp point, smooth; ocelli and subpetiolar process absent
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<emphasis italics="true" pageId="15" pageNumber="16">
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………………………………… ………………
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<taxonomicName authority="Barden & Grimaldi, 2012" authorityName="Barden & Grimaldi" authorityYear="2012" box="[965,1471,934,955]" class="Hexapoda" family="Formicidae" genus="Haidomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="scimitarus">
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<taxonomicName authority="Dlussky, 1996" authorityName="Dlussky" authorityYear="1996" box="[965,1103,934,954]" class="Insecta" family="Formicidae" genus="Haidomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">Haidomyrmex</taxonomicName>
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scimitarus
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<bibRefCitation author="Barden, P. & Grimaldi, D. A." box="[1218,1471,935,955]" journalOrPublisher="American Museum Novitates" pageId="15" pageNumber="16" pagination="16" part="3755" refId="ref15236" refString="Barden, P., Grimaldi, D. A., 2012. Rediscovery of the bizarre Cretaceous ant Haidomyrmex Dlussky (Hymenoptera: Formicidae), with two new species. American Museum Novitates 3755, 1 e 16. https: // doi. org / 10.1206 / 3755.2." title="Rediscovery of the bizarre Cretaceous ant Haidomyrmex Dlussky (Hymenoptera: Formicidae), with two new species" type="journal article" year="2012">Barden & Grimaldi, 2012</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</paragraph>
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<paragraph blockId="15.[805,1475,178,1959]" pageId="15" pageNumber="16">
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Apical flagellomere longest; frontal triangle feebly elevated; vertex, ventral margin of mandibles, and antennomeres mostly glabrous; mandibles with medioventral blade bearing a single apical tooth, with tip of apical portion tapered to blunt point and minutely serrate; ocelli absent; subpetiolar process a minute tooth
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<emphasis box="[895,1469,1103,1122]" italics="true" pageId="15" pageNumber="16">……………………………………………………………………</emphasis>
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.
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<emphasis box="[831,986,1131,1150]" italics="true" pageId="15" pageNumber="16">…………………</emphasis>
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.
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<emphasis box="[997,1203,1130,1150]" italics="true" pageId="15" pageNumber="16">
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<taxonomicName authorityName="Dlussky" authorityYear="1996" box="[997,1134,1130,1150]" class="Hexapoda" family="Formicidae" genus="Haidomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">Haidomyrmex</taxonomicName>
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zigrasi
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</emphasis>
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..
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<bibRefCitation author="Barden, P. & Grimaldi, D. A." box="[1219,1471,1131,1150]" journalOrPublisher="American Museum Novitates" pageId="15" pageNumber="16" pagination="16" part="3755" refId="ref15236" refString="Barden, P., Grimaldi, D. A., 2012. Rediscovery of the bizarre Cretaceous ant Haidomyrmex Dlussky (Hymenoptera: Formicidae), with two new species. American Museum Novitates 3755, 1 e 16. https: // doi. org / 10.1206 / 3755.2." title="Rediscovery of the bizarre Cretaceous ant Haidomyrmex Dlussky (Hymenoptera: Formicidae), with two new species" type="journal article" year="2012">Barden & Grimaldi, 2012</bibRefCitation>
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.
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</paragraph>
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<paragraph blockId="15.[805,1475,178,1959]" pageId="15" pageNumber="16">
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5. Posterior clypeal projection not furrowed ventrally (
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<emphasis bold="true" box="[1370,1387,1159,1178]" pageId="15" pageNumber="16">¼</emphasis>
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without lateral ridges), forming a pointed or paddle-shaped horn; long, paired trigger setae present, arising from base of clypeal process or ventral surface of apical setose pad; peg-like denticles absent or present only on ventral setose pad; labrum not obviously exposed
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<emphasis box="[920,1428,1298,1317]" italics="true" pageId="15" pageNumber="16">……………………………………………………………</emphasis>
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.
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<emphasis box="[1432,1454,1298,1317]" italics="true" pageId="15" pageNumber="16">…</emphasis>
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6
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</paragraph>
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<paragraph blockId="15.[805,1475,178,1959]" pageId="15" pageNumber="16">
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Posterior clypeal projection distinctly furrowed ventrally, with lateral ridges originating above mandible insertions, and converging posteriorly to form the lateral margins of a horn; paired trigger setae absent; peg-like denticles present along lateral clypeal ridges, or on labrum, or both; labrum exposed ventrally of horn
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<emphasis box="[1013,1455,1466,1485]" italics="true" pageId="15" pageNumber="16">……………………………………………………</emphasis>
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9
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</paragraph>
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<paragraph blockId="15.[805,1475,178,1959]" pageId="15" pageNumber="16">
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6. Clypeal horn a small triangulate cuticular elevation; one pair of trigger setae positioned at base of horn; compound eyes small (
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<collectionCode box="[843,867,1550,1569]" pageId="15" pageNumber="16">OI</collectionCode>
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24); ocelli absent; flagellomeres IIeX compact, barely longer than wide
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<emphasis italics="true" pageId="15" pageNumber="16">
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……………………………………………………………… ………………………
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<taxonomicName authorityName="Perrichot & Wang & Barden" authorityYear="2020" box="[1035,1317,1604,1624]" class="Insecta" family="Formicidae" genus="Protoceratomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="revelatus">Protoceratomyrmex revelatus</taxonomicName>
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</emphasis>
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gen. et. sp. nov.
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</paragraph>
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<paragraph blockId="15.[805,1475,178,1959]" pageId="15" pageNumber="16">
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Clypeal horn paddle-shaped, comprising basal stalk and apical setose pad; compound eyes small to large (
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<collectionCode box="[1294,1318,1661,1680]" pageId="15" pageNumber="16">OI</collectionCode>
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26e39); ocelli present; flagellomeres IIeX elongate, distinctly longer than wide
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<emphasis box="[837,1367,1717,1736]" italics="true" pageId="15" pageNumber="16">………………………………………………………………</emphasis>
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....
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<emphasis box="[1386,1452,1717,1736]" italics="true" pageId="15" pageNumber="16">………</emphasis>
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7
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</paragraph>
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<paragraph blockId="15.[805,1475,178,1959]" pageId="15" pageNumber="16">
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7. Clypeal horn and curved apical portion of mandibles extremely long, reaching high above vertex (HoI 140;
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<collectionCode box="[1282,1325,1773,1792]" country="Malaysia" httpUri="http://grbio.org/cool/nf70-xac8" name="Malaysian Agricultural Research and Development Institute (MARDI)" pageId="15" pageNumber="16" type="Herbarium">MDI</collectionCode>
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130); peg-like denticles present on ventral setose pad; 2 pairs of trigger setae positioned at base of horn's stalk; frontal triangle fused with horn, not distinguishable; gastral constriction between
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<collectionCode box="[953,988,1884,1903]" pageId="15" pageNumber="16">AIII</collectionCode>
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and
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<collectionCode box="[1084,1120,1884,1903]" pageId="15" pageNumber="16">AIV</collectionCode>
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at most faintly impressed
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<emphasis box="[837,1455,1912,1931]" italics="true" pageId="15" pageNumber="16">…………………………………………………………………………</emphasis>
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...
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<emphasis box="[837,1168,1939,1959]" italics="true" pageId="15" pageNumber="16">
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………
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<taxonomicName authority="Perrichot, Wang & Engel, 2016" authorityName="Perrichot, Wang & Engel" authorityYear="2016" box="[908,1474,1939,1959]" class="Hexapoda" family="Formicidae" genus="Ceratomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="ellenbergeri">
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<taxonomicName box="[908,1049,1939,1959]" genus="Ceratomyrmex" pageId="15" pageNumber="16" rank="genus">Ceratomyrmex</taxonomicName>
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ellenbergeri
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<bibRefCitation author="Perrichot, V. & Wang, B. & Engel, M. S." box="[1173,1474,1940,1959]" journalOrPublisher="Current Biology" pageId="15" pageNumber="16" pagination="1472" part="26" refId="ref16608" refString="Perrichot, V., Wang, B., Engel, M. S., 2016. Extreme morphogenesis and ecological specialization among Cretaceous basal ants. Current Biology 26, 1468 e 1472. https: // doi. org / 10.1016 / j. cub. 2016.03.075." title="Extreme morphogenesis and ecological specialization among Cretaceous basal ants" type="journal article" year="2016">Perrichot, Wang & Engel, 2016</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</paragraph>
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<paragraph blockId="16.[117,784,179,1704]" pageId="16" pageNumber="17">
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Clypeal horn and curved apical portion of mandibles distinctly shorter, not reaching above vertex (
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<collectionCode box="[505,550,207,226]" country="Malaysia" httpUri="http://grbio.org/cool/nf70-xac8" name="Malaysian Agricultural Research and Development Institute (MARDI)" pageId="16" pageNumber="17" type="Herbarium">MDI</collectionCode>
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70); peg-like denticles absent; 1 pair of trigger setae positioned on ventral surface of apical setose pad; frontal triangle connecting with horn by cleared cuticular membrane; gastral constriction between
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<collectionCode box="[748,783,290,309]" pageId="16" pageNumber="17">AIII</collectionCode>
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and
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<collectionCode box="[196,232,318,337]" pageId="16" pageNumber="17">AIV</collectionCode>
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deeply impressed
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<emphasis italics="true" pageId="16" pageNumber="17">………………………………………… ……………………………………………</emphasis>
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.
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<taxonomicName authorityName="Barden & Grimaldi" authorityYear="2017" box="[532,676,345,365]" class="Hexapoda" family="Formicidae" genus="Linguamyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
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<emphasis box="[532,676,345,365]" italics="true" pageId="16" pageNumber="17">Linguamyrmex</emphasis>
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</taxonomicName>
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species: 8
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</paragraph>
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<paragraph blockId="16.[117,784,179,1704]" pageId="16" pageNumber="17">
|
|
8 Clypeal horn reduced (HoI 25e35), with stalk almost indistinct and apical setose pad hexagonal, as broad as (worker) or smaller than (gyne) compound eye's length (
|
|
<collectionCode box="[561,591,430,449]" pageId="16" pageNumber="17">EL</collectionCode>
|
|
); mandibles with medioventral blade bearing two teeth and, in worker, surpassing midlength of apical curved portion; compound eyes small in worker (
|
|
<collectionCode box="[352,376,513,532]" pageId="16" pageNumber="17">OI</collectionCode>
|
|
25); propodeum higher than long
|
|
<emphasis box="[145,167,541,560]" italics="true" pageId="16" pageNumber="17">…</emphasis>
|
|
..
|
|
<emphasis box="[177,309,541,560]" italics="true" pageId="16" pageNumber="17">………………</emphasis>
|
|
.
|
|
<emphasis box="[315,623,540,560]" italics="true" pageId="16" pageNumber="17">
|
|
……
|
|
<taxonomicName authorityName="Perrichot & Wang & Barden" authorityYear="2020" box="[365,623,540,560]" class="Hexapoda" family="Formicidae" genus="Linguamyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="brevicornis">
|
|
<taxonomicName box="[365,509,540,560]" genus="Linguamyrmex" pageId="16" pageNumber="17" rank="genus">Linguamyrmex</taxonomicName>
|
|
brevicornis
|
|
</taxonomicName>
|
|
</emphasis>
|
|
gen. et. sp. nov.
|
|
</paragraph>
|
|
<paragraph blockId="16.[117,784,179,1704]" pageId="16" pageNumber="17">
|
|
Clypeal horn conspicuous (HoI 56), with stalk short, not surpassing frontal triangle in dorsal view, and apical setose pad circular, broader than EL in worker; mandibles with medioventral blade smooth and not reaching to midlength of apical curved portion; compound eyes large in worker (
|
|
<collectionCode box="[711,735,681,700]" pageId="16" pageNumber="17">OI</collectionCode>
|
|
39); propodeum longer than high
|
|
<emphasis italics="true" pageId="16" pageNumber="17">……………………………… …………………</emphasis>
|
|
.
|
|
<emphasis box="[305,529,735,755]" italics="true" pageId="16" pageNumber="17">
|
|
…
|
|
<taxonomicName authority="Barden & Grimaldi, 2017" authorityName="Barden & Grimaldi" authorityYear="2017" box="[332,780,735,756]" class="Hexapoda" family="Formicidae" genus="Linguamyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="vladi">
|
|
<taxonomicName box="[332,476,735,755]" genus="Linguamyrmex" pageId="16" pageNumber="17" rank="genus">Linguamyrmex</taxonomicName>
|
|
vladi
|
|
<bibRefCitation author="Barden, P. & Herhold, H. W. & Grimaldi, D. A." box="[534,780,736,756]" journalOrPublisher="Systematic Entomology" pageId="16" pageNumber="17" pagination="846" part="42" refId="ref15337" refString="Barden, P., Herhold, H. W., Grimaldi, D. A., 2017. A new genus of hell ants from the Cretaceous (Hymenoptera: Formicidae: Haidomyrmecini) with a novel head structure. Systematic Entomology 42, 837 e 846. https: // doi. org / 10.1111 / syen. 12253." title="A new genus of hell ants from the Cretaceous (Hymenoptera: Formicidae: Haidomyrmecini) with a novel head structure" type="journal article" year="2017">Barden & Grimaldi, 2017</bibRefCitation>
|
|
</taxonomicName>
|
|
</emphasis>
|
|
.
|
|
</paragraph>
|
|
<paragraph blockId="16.[117,784,179,1704]" pageId="16" pageNumber="17">
|
|
Clypeal horn enormous (HoI 95e105), with apical setose pad circular, nearly as broad as head in dorsal view, and stalk about as long as pad; mandibles with medioventral blade smooth and, in worker, not reaching to midlength of apical curved portion; compound eyes large in worker (
|
|
<collectionCode box="[469,493,876,895]" pageId="16" pageNumber="17">OI</collectionCode>
|
|
44); propodeum longer than high
|
|
<emphasis italics="true" pageId="16" pageNumber="17">…………………………………………………………………… …………………</emphasis>
|
|
..
|
|
<taxonomicName authority="Miao & Wang, 2019" authorityName="Miao & Wang" authorityYear="2019" box="[316,778,930,951]" class="Hexapoda" family="Formicidae" genus="Linguamyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="rhinocerus">
|
|
<emphasis box="[316,570,930,950]" italics="true" pageId="16" pageNumber="17">
|
|
<taxonomicName box="[316,460,930,950]" genus="Linguamyrmex" pageId="16" pageNumber="17" rank="genus">Linguamyrmex</taxonomicName>
|
|
rhinocerus
|
|
</emphasis>
|
|
<bibRefCitation author="Miao, Z. & Wang, M." box="[576,778,932,951]" journalOrPublisher="Journal of Guangxi Normal University" pageId="16" pageNumber="17" pagination="142" part="37" refId="ref16302" refString="Miao, Z., Wang, M., 2019. A new species of hell ants (Hymenoptera: Formicidae: Haidomyrmecini) from the Cretaceous Burmese amber. Journal of Guangxi Normal University 37 (2), 139 e 142. https: // doi. org / 10.16088 / j. issn. 1001 - 6600.2019. 02.017." title="A new species of hell ants (Hymenoptera: Formicidae: Haidomyrmecini) from the Cretaceous Burmese amber" type="journal article" year="2019">Miao & Wang, 2019</bibRefCitation>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
<paragraph blockId="16.[117,784,179,1704]" pageId="16" pageNumber="17">
|
|
9 Body and legs robust; mandibles scythe-shaped, with medioventral triangular blade present; in frontal view, the outline of apical portion of mandibles aligned with clypeal lateral ridges; clypeal horn gradually tapering to rounded tip; genae elongate, the mandibles inserted far from compound eyes
|
|
<emphasis box="[145,719,1099,1118]" italics="true" pageId="16" pageNumber="17">……………………………………………………………………</emphasis>
|
|
..
|
|
<emphasis box="[729,751,1099,1118]" italics="true" pageId="16" pageNumber="17">…</emphasis>
|
|
10
|
|
</paragraph>
|
|
<paragraph blockId="16.[117,784,179,1704]" pageId="16" pageNumber="17">
|
|
Body and legs gracile; mandibles sickle-shaped, with medioventral triangular blade absent; clypeal horn and labrum protruding horizontally, their tip with a deep median notch; apical lateral margins of horn bilobed; genae strongly reduced, the mandibles inserted close to compound eyes
|
|
<emphasis box="[145,322,1266,1285]" italics="true" pageId="16" pageNumber="17">……………………</emphasis>
|
|
.
|
|
<emphasis box="[327,393,1266,1285]" italics="true" pageId="16" pageNumber="17">………</emphasis>
|
|
.
|
|
<taxonomicName authorityName="Perrichot & Wang & Barden" authorityYear="2020" box="[405,619,1265,1285]" class="Insecta" family="Formicidae" genus="Aquilomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="huangi">
|
|
<emphasis box="[405,619,1265,1285]" italics="true" pageId="16" pageNumber="17">Aquilomyrmex huangi</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel box="[626,778,1267,1286]" pageId="16" pageNumber="17">gen. et. sp. nov</taxonomicNameLabel>
|
|
.
|
|
</paragraph>
|
|
<paragraph blockId="16.[117,784,179,1704]" pageId="16" pageNumber="17">
|
|
10 Mandibles with apical curved portion much longer than basal portion and entirely serrate along dorsal margin; with medioventral blade an isosceles tooth not projecting beyond mandibular elbow (labrum and clypeal surface below horn entirely exposed when mandibles closed); ocelli large, antennomeres and compound eyes distinctly elongate
|
|
<emphasis box="[150,624,1460,1481]" italics="true" pageId="16" pageNumber="17">
|
|
……………………………
|
|
<taxonomicName authorityName="Perrichot & Wang & Barden" authorityYear="2020" box="[399,624,1460,1480]" class="Insecta" family="Formicidae" genus="Dhagnathos" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="autokrator">Dhagnathos autokrator</taxonomicName>
|
|
</emphasis>
|
|
<taxonomicNameLabel box="[630,784,1462,1481]" pageId="16" pageNumber="17">gen. et. sp. nov.</taxonomicNameLabel>
|
|
</paragraph>
|
|
<paragraph blockId="16.[117,784,179,1704]" pageId="16" pageNumber="17">
|
|
Mandibles with apical curved portion not distinctly longer than basal portion; with medioventral blade a large triangle gradually tapering to tip of apical portion (labrum and clypeal surface below horn entirely concealed when mandibles closed); dorsal margin of mandibles serrate along distal half of medioventral projection, bidentate apically; ocelli, antennomeres, and compound eyes more reduced
|
|
<emphasis italics="true" pageId="16" pageNumber="17">
|
|
………… ……………………………… ……………………………………
|
|
<taxonomicName authorityName="Perrichot & Wang & Barden" authorityYear="2020" box="[454,623,1684,1704]" class="Insecta" family="Formicidae" genus="Chonidris" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="insolita">Chonidris insolita</taxonomicName>
|
|
</emphasis>
|
|
<taxonomicNameLabel box="[629,780,1685,1704]" pageId="16" pageNumber="17">gen. et. sp. nov</taxonomicNameLabel>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |