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<document id="42ED9A6F8C6791039D4DFC657F0ADF30" ID-DOI="10.11646/zootaxa.5476.1.24" ID-ISSN="1175-5326" ID-Zenodo-Dep="12681699" ID-ZooBank="880763E8-5AE7-4F9C-AB40-EC5FC386E052" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1720375266466" checkinUser="plazi" docAuthor="Ng, Peter K. L." docDate="2024" docId="03A2321EFFA7FF84FBBD60BCFC9A100D" docLanguage="en" docName="zootaxa.5476.1.24.pdf" docOrigin="Zootaxa 5476 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5476.1.24" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tiwaripotamon nganima Ng, 2024, n. sp." docType="treatment" docVersion="3" lastPageNumber="305" masterDocId="FF9B4A66FFA6FF83FB2A6313FF821620" masterDocTitle="Taxonomic notes on three species of Tiwaripotamon Bott, 1970 (Crustacea: Brachyura: Potamidae) from Vietnam and China, one of which is new to science" masterLastPageNumber="313" masterPageNumber="298" pageNumber="299" updateTime="1720378360316" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="637BE9D4DB710E3A5F100E2280C76C45">Taxonomic notes on three species of Tiwaripotamon Bott, 1970 (Crustacea: Brachyura: Potamidae) from Vietnam and China, one of which is new to science</mods:title>
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<mods:namePart id="7F04E1C0C74C606EACBBEF4612F5FC71">Ng, Peter K. L.</mods:namePart>
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<mods:date id="8A3B6BBC921E33C887F02A81D5B545B3">2024</mods:date>
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<mods:number id="0C28A4D3E9E0F540E77690D6862E4559">2024-07-04</mods:number>
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<mods:number id="141A557FDDBDF50EEB40729D20DD3F00">5476</mods:number>
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<mods:classification id="DFF93442EDA3B2EAFB7180D22EAE362B">journal article</mods:classification>
<mods:identifier id="90DE06D7451465669923EAE71F299F4C" type="DOI">10.11646/zootaxa.5476.1.24</mods:identifier>
<mods:identifier id="24ED6CEED76F3633B4BCBBE934BD5550" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03A2321EFFA7FF84FBBD60BCFC9A100D" ID-DOI="http://doi.org/10.5281/zenodo.12681374" ID-Zenodo-Dep="12681374" LSID="urn:lsid:plazi:treatment:03A2321EFFA7FF84FBBD60BCFC9A100D" httpUri="http://treatment.plazi.org/id/03A2321EFFA7FF84FBBD60BCFC9A100D" lastPageId="7" lastPageNumber="305" pageId="1" pageNumber="299">
<subSubSection id="C311D083FFA7FF82FBBD60BCFD8F15E9" box="[151,525,943,969]" pageId="1" pageNumber="299" type="nomenclature">
<paragraph id="8BB48308FFA7FF82FBBD60BCFD8F15E9" blockId="1.[151,525,943,1005]" box="[151,525,943,969]" pageId="1" pageNumber="299">
<heading id="D0FC3464FFA7FF82FBBD60BCFD8F15E9" bold="true" box="[151,525,943,969]" fontSize="11" level="1" pageId="1" pageNumber="299" reason="1">
<emphasis id="B97F5F1AFFA7FF82FBBD60BCFD8F15E9" bold="true" box="[151,525,943,969]" pageId="1" pageNumber="299">
<taxonomicName id="4C0BF88BFFA7FF82FBBD60BCFE4115E9" authorityName="Ng" authorityYear="2024" box="[151,451,943,969]" class="Malacostraca" family="Potamidae" genus="Tiwaripotamon" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="299" phylum="Arthropoda" rank="species" species="nganima" status="sp. nov.">
<emphasis id="B97F5F1AFFA7FF82FBBD60BCFE4115E9" bold="true" box="[151,451,943,969]" italics="true" pageId="1" pageNumber="299">Tiwaripotamon nganima</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24CE261FFA7FF82FAE060BCFD8F15E9" box="[458,525,943,969]" pageId="1" pageNumber="299" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C311D083FFA7FF82FBBD60C0FE9015CD" box="[151,274,978,1005]" pageId="1" pageNumber="299" type="description">
<paragraph id="8BB48308FFA7FF82FBBD60C0FE9015CD" blockId="1.[151,525,943,1005]" box="[151,274,978,1005]" pageId="1" pageNumber="299">
(
<figureCitation id="13309F8DFFA7FF82FBB560C0FE8B15CD" box="[159,265,978,1005]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="2.[151,250,1899,1923]" captionStartId-1="3.[151,250,1868,1892]" captionStartId-2="4.[151,250,1926,1950]" captionStartId-3="5.[151,250,1308,1332]" captionStartId-4="6.[151,250,1463,1487]" captionTargetBox-0="[151,1435,179,1874]" captionTargetBox-1="[151,1435,309,1845]" captionTargetBox-2="[151,1436,417,1903]" captionTargetBox-3="[151,1436,181,1284]" captionTargetBox-4="[160,1428,189,1432]" captionTargetId-0="figure-15@2.[151,1435,179,1874]" captionTargetId-1="figure-79@3.[151,1435,309,1845]" captionTargetId-2="figure-120@4.[151,1436,417,1903]" captionTargetId-3="figure-25@5.[151,1436,181,1284]" captionTargetId-4="figure-15@6.[151,1436,181,1440]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionText-0="FIGURE 1. Tiwaripotamon nganima n. sp., holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; A, overall dorsal habitus; B, ventral view of habitus; C, frontal view." captionText-1="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." captionText-2="FIGURE 3. Tiwaripotamon nganima n. sp. A–F, holotype male (59.3 × 43.6 mm) (ZRC2022.0810), Vietnam; G, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A, chelae; B, left third maxilliped; C, right fourth ambulatory leg; D, male sternopleonal cavity; E, male posterior thoracic sternites and pleon; F, G, male pleon." captionText-3="FIGURE 4. Tiwaripotamon nganima n. sp. A–E, holotype male (59.3 × 43.6 mm) (ZRC2022.0810), Vietnam; F, G, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam. A, left G1 (ventral view); B, left G1 (dorsal view); C, distal part of left G1 (ventral view); D, distal part of left G1 (dorsal view); E, left G2; F, female anterior thoracic sternites and pleon; G, female sternopleonal cavity and vulvae." captionText-4="FIGURE 5. Tiwaripotamon nganima n. sp. A–E, holotype male (59.3 × 43.6 mm) (ZRC2022.0810), Vietnam; F–H, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A, left G1 (ventral view); B, F, distal part of left G1 (ventral view); C, G, distal part of left G1 (dorsomesial view); D, H, distal part of left G1 (dorsal view); E, left G2. Scales: A, E = 1.0 mm; B–D, F–H = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.12681708" figureDoi-1="http://doi.org/10.5281/zenodo.12681711" figureDoi-2="http://doi.org/10.5281/zenodo.12681715" figureDoi-3="http://doi.org/10.5281/zenodo.12681717" figureDoi-4="http://doi.org/10.5281/zenodo.12681719" httpUri-0="https://zenodo.org/record/12681708/files/figure.png" httpUri-1="https://zenodo.org/record/12681711/files/figure.png" httpUri-2="https://zenodo.org/record/12681715/files/figure.png" httpUri-3="https://zenodo.org/record/12681717/files/figure.png" httpUri-4="https://zenodo.org/record/12681719/files/figure.png" pageId="1" pageNumber="299">Figs. 1–5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C311D083FFA7FF82FBBD6709FAE3125D" pageId="1" pageNumber="299" type="materials_examined">
<paragraph id="8BB48308FFA7FF82FBBD6709FAE3125D" blockId="1.[151,1437,1050,2013]" pageId="1" pageNumber="299">
<emphasis id="B97F5F1AFFA7FF82FBBD6709FE021214" bold="true" box="[151,384,1050,1076]" pageId="1" pageNumber="299">Material examined.</emphasis>
<materialsCitation id="3B638955FFA7FF82FAAC6708FBAE1279" collectingDate="2001-05-24" collectionCode="ZRC" collectorName="Nguyen Van Quang" country="Vietnam" latitude="21.63" location="Huu Lung" longLatPrecision="758" longitude="106.46" pageId="1" pageNumber="299" specimenCode="ZRC 2022.0810" specimenCount="1" stateProvince="Lang Son" typeStatus="holotype">
<typeStatus id="54B03DAAFFA7FF82FAAC6708FE711215" box="[390,499,1051,1077]" pageId="1" pageNumber="299" type="holotype">Holotype</typeStatus>
: male (59.3 ×
<quantity id="4CF32EEDFFA7FF82F9BB6708FD741215" box="[657,758,1051,1077]" metricMagnitude="-2" metricUnit="m" metricValue="4.36" pageId="1" pageNumber="299" unit="mm" value="43.6">43.6 mm</quantity>
) (
<specimenCode id="DBAD2B73FFA7FF82F8266708FC441215" box="[780,966,1051,1077]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="299">ZRC 2022.0810</specimenCode>
), forest between
<location id="8ED4D5D3FFA7FF82FFA86708FB761215" LSID="urn:lsid:plazi:treatment:03A2321EFFA7FF84FBBD60BCFC9A100D:8ED4D5D3FFA7FF82FFA86708FB761215" box="[1154,1268,1051,1077]" country="Vietnam" latitude="21.63" longLatPrecision="758" longitude="106.46" name="Huu Lung" pageId="1" pageNumber="299" stateProvince="Lang Son">Huu Lung</location>
and
<collectingRegion id="49CF4DEAFFA7FF82FE006708FA1A1215" box="[1322,1432,1051,1077]" country="Vietnam" name="Lang Son" pageId="1" pageNumber="299">Lang Son</collectingRegion>
, northern
<collectingCountry id="F31CC398FFA7FF82FBD7672CFEDD1279" box="[253,351,1087,1113]" name="Vietnam" pageId="1" pageNumber="299">Vietnam</collectingCountry>
, ca.
<geoCoordinate id="EE3FE5CFFFA7FF82FABB672CFE721279" box="[401,496,1087,1113]" degrees="21.63" direction="north" orientation="latitude" pageId="1" pageNumber="299" precision="555" value="21.63">21.63°N</geoCoordinate>
<geoCoordinate id="EE3FE5CFFFA7FF82FADD672CFDE11279" box="[503,611,1087,1113]" degrees="106.46" direction="east" orientation="longitude" pageId="1" pageNumber="299" precision="555" value="106.46">106.46°E</geoCoordinate>
, coll.
<collectorName id="26FEE6DEFFA7FF82F982672CFC051279" box="[680,903,1087,1113]" pageId="1" pageNumber="299">Nguyen Van Quang</collectorName>
,
<date id="FFB5A5C8FFA7FF82F8B9672CFBAE1279" box="[915,1068,1087,1113]" pageId="1" pageNumber="299" value="2001-05-24">
<collectingDate id="EFF15C20FFA7FF82F8B9672CFBAE1279" box="[915,1068,1087,1113]" pageId="1" pageNumber="299" value="2001-05-24">24 May 2001</collectingDate>
</date>
</materialsCitation>
.
<typeStatus id="54B03DAAFFA7FF82FF12672CFB2B1279" box="[1080,1193,1087,1113]" pageId="1" pageNumber="299" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D0D4881FFA7FF82FF9E672CFA881279" box="[1204,1290,1087,1113]" count="3" pageId="1" pageNumber="299" type="male">3 males</specimenCount>
(41.5 ×
<quantity id="4CF32EEDFFA7FF82FE41672CFF47125D" metricMagnitude="-2" metricUnit="m" metricValue="3.13" pageId="1" pageNumber="299" unit="mm" value="31.3">31.3 mm</quantity>
, 49.3 ×
<quantity id="4CF32EEDFFA7FF82FA0A6771FE04125D" box="[288,390,1122,1149]" metricMagnitude="-2" metricUnit="m" metricValue="3.52" pageId="1" pageNumber="299" unit="mm" value="35.2">35.2 mm</quantity>
, 54.2 ×
<quantity id="4CF32EEDFFA7FF82FACA6770FDCA125D" box="[480,584,1123,1149]" metricMagnitude="-2" metricUnit="m" metricValue="4.06" pageId="1" pageNumber="299" unit="mm" value="40.6">40.6 mm</quantity>
),
<materialsCitation id="3B638955FFA7FF82F9716770FADF125D" box="[603,1373,1122,1149]" collectingDate="2001-05-24" collectionCode="ZRC" collectorName="Nguyen Van Quang" country="Vietnam" latitude="21.63" location="Huu Lung" longLatPrecision="758" longitude="106.46" pageId="1" pageNumber="299" specimenCode="ZRC 2022.0811" specimenCount="1" specimenCount-female="1" stateProvince="Lang Son" typeStatus="paratype">
<specimenCount id="9D0D4881FFA7FF82F9716770FD3E125D" box="[603,700,1123,1149]" count="1" pageId="1" pageNumber="299" type="female">1 female</specimenCount>
(53.1 ×
<quantity id="4CF32EEDFFA7FF82F8316770FC02125D" box="[795,896,1123,1149]" metricMagnitude="-2" metricUnit="m" metricValue="3.9899999999999998" pageId="1" pageNumber="299" unit="mm" value="39.9">39.9 mm</quantity>
) (
<specimenCode id="DBAD2B73FFA7FF82F8BD6770FBD2125D" box="[919,1104,1123,1149]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="299">ZRC 2022.0811</specimenCode>
), same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C311D083FFA7FF82FBED6795FE7B1071" pageId="1" pageNumber="299" type="diagnosis">
<paragraph id="8BB48308FFA7FF82FBED6795FE7B1071" blockId="1.[151,1437,1050,2013]" pageId="1" pageNumber="299">
<emphasis id="B97F5F1AFFA7FF82FBED6795FEC01280" bold="true" box="[199,322,1158,1184]" pageId="1" pageNumber="299">Diagnosis.</emphasis>
Carapace subquadrate, wider than long (
<figureCitation id="13309F8DFFA7FF82F8326794FCFC1281" box="[792,894,1159,1185]" captionStart="FIGURE 1" captionStartId="2.[151,250,1899,1923]" captionTargetBox="[151,1435,179,1874]" captionTargetId="figure-15@2.[151,1435,179,1874]" captionTargetPageId="2" captionText="FIGURE 1. Tiwaripotamon nganima n. sp., holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; A, overall dorsal habitus; B, ventral view of habitus; C, frontal view." figureDoi="http://doi.org/10.5281/zenodo.12681708" httpUri="https://zenodo.org/record/12681708/files/figure.png" pageId="1" pageNumber="299">Figs. 1A</figureCitation>
,
<figureCitation id="13309F8DFFA7FF82F8A66794FC511281" box="[908,979,1159,1185]" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="1" pageNumber="299">2A–C</figureCitation>
); epigastric cristae very low, postorbital cristae not visible (
<figureCitation id="13309F8DFFA7FF82FA4267B8FE4812E5" box="[360,458,1195,1221]" captionStart="FIGURE 1" captionStartId="2.[151,250,1899,1923]" captionTargetBox="[151,1435,179,1874]" captionTargetId="figure-15@2.[151,1435,179,1874]" captionTargetPageId="2" captionText="FIGURE 1. Tiwaripotamon nganima n. sp., holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; A, overall dorsal habitus; B, ventral view of habitus; C, frontal view." figureDoi="http://doi.org/10.5281/zenodo.12681708" httpUri="https://zenodo.org/record/12681708/files/figure.png" pageId="1" pageNumber="299">Figs. 1A</figureCitation>
,
<figureCitation id="13309F8DFFA7FF82FAFF67B8FD9E12E5" box="[469,540,1195,1221]" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="1" pageNumber="299">2A–C</figureCitation>
); external orbital tooth broadly triangular, low but usually visible, separated from anterolateral margin by indentation or low cleft (
<figureCitation id="13309F8DFFA7FF82F98167DCFCA712C9" box="[683,805,1231,1257]" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="1" pageNumber="299">Fig. 2A–C</figureCitation>
); epibranchial tooth small but visible, rest of anterolateral margin subcristate, gently serrrate (
<figureCitation id="13309F8DFFA7FF82F90C67E0FD20132D" box="[550,674,1267,1293]" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="1" pageNumber="299">Fig. 2A–C</figureCitation>
); suborbital margin convex, smooth, without inner tooth; suborbital region smooth, subhepatic, pterygostomial and sub-branchial regions weakly rugose to smooth (
<figureCitation id="13309F8DFFA7FF82FFCA6604FAC11311" box="[1248,1347,1303,1329]" captionStart="FIGURE 1" captionStartId="2.[151,250,1899,1923]" captionTargetBox="[151,1435,179,1874]" captionTargetId="figure-15@2.[151,1435,179,1874]" captionTargetPageId="2" captionText="FIGURE 1. Tiwaripotamon nganima n. sp., holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; A, overall dorsal habitus; B, ventral view of habitus; C, frontal view." figureDoi="http://doi.org/10.5281/zenodo.12681708" httpUri="https://zenodo.org/record/12681708/files/figure.png" pageId="1" pageNumber="299">Figs. 1C</figureCitation>
,
<figureCitation id="13309F8DFFA7FF82FE786604FF2D1375" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="1" pageNumber="299">2D, E, H</figureCitation>
); posterior margin of epistome with median lobe prominent, triangular to more rounded but distinctly projecting beyond lateral margins (
<figureCitation id="13309F8DFFA7FF82FA8E664CFD871359" box="[420,517,1375,1401]" captionStart="FIGURE 1" captionStartId="2.[151,250,1899,1923]" captionTargetBox="[151,1435,179,1874]" captionTargetId="figure-15@2.[151,1435,179,1874]" captionTargetPageId="2" captionText="FIGURE 1. Tiwaripotamon nganima n. sp., holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; A, overall dorsal habitus; B, ventral view of habitus; C, frontal view." figureDoi="http://doi.org/10.5281/zenodo.12681708" httpUri="https://zenodo.org/record/12681708/files/figure.png" pageId="1" pageNumber="299">Figs. 1C</figureCitation>
,
<figureCitation id="13309F8DFFA7FF82F93B664CFDD81359" box="[529,602,1375,1401]" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="1" pageNumber="299">2D–G</figureCitation>
). Male thoracic sternites prominently punctate (
<figureCitation id="13309F8DFFA7FF82FF46664CFB4F1359" box="[1132,1229,1375,1401]" captionStart="FIGURE 1" captionStartId="2.[151,250,1899,1923]" captionTargetBox="[151,1435,179,1874]" captionTargetId="figure-15@2.[151,1435,179,1874]" captionTargetPageId="2" captionText="FIGURE 1. Tiwaripotamon nganima n. sp., holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; A, overall dorsal habitus; B, ventral view of habitus; C, frontal view." figureDoi="http://doi.org/10.5281/zenodo.12681708" httpUri="https://zenodo.org/record/12681708/files/figure.png" pageId="1" pageNumber="299">Figs. 1B</figureCitation>
,
<figureCitation id="13309F8DFFA7FF82FFF3664CFB7F1359" box="[1241,1277,1375,1401]" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="1" pageNumber="299">2H</figureCitation>
,
<figureCitation id="13309F8DFFA7FF82FE22664CFAAC1359" box="[1288,1326,1375,1401]" captionStart="FIGURE 3" captionStartId="4.[151,250,1926,1950]" captionTargetBox="[151,1436,417,1903]" captionTargetId="figure-120@4.[151,1436,417,1903]" captionTargetPageId="4" captionText="FIGURE 3. Tiwaripotamon nganima n. sp. A–F, holotype male (59.3 × 43.6 mm) (ZRC2022.0810), Vietnam; G, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A, chelae; B, left third maxilliped; C, right fourth ambulatory leg; D, male sternopleonal cavity; E, male posterior thoracic sternites and pleon; F, G, male pleon." figureDoi="http://doi.org/10.5281/zenodo.12681715" httpUri="https://zenodo.org/record/12681715/files/figure.png" pageId="1" pageNumber="299">3D</figureCitation>
); tubercle of male pleonal locking mechanism on posterior third of somite 5 (
<figureCitation id="13309F8DFFA7FF82F8B86690FC7713BD" box="[914,1013,1411,1437]" captionStart="FIGURE 1" captionStartId="2.[151,250,1899,1923]" captionTargetBox="[151,1435,179,1874]" captionTargetId="figure-15@2.[151,1435,179,1874]" captionTargetPageId="2" captionText="FIGURE 1. Tiwaripotamon nganima n. sp., holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; A, overall dorsal habitus; B, ventral view of habitus; C, frontal view." figureDoi="http://doi.org/10.5281/zenodo.12681708" httpUri="https://zenodo.org/record/12681708/files/figure.png" pageId="1" pageNumber="299">Figs. 1B</figureCitation>
,
<figureCitation id="13309F8DFFA7FF82FF296690FBA513BD" box="[1027,1063,1411,1437]" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="1" pageNumber="299">2H</figureCitation>
,
<figureCitation id="13309F8DFFA7FF82FF1E6690FBD813BD" box="[1076,1114,1411,1437]" captionStart="FIGURE 3" captionStartId="4.[151,250,1926,1950]" captionTargetBox="[151,1436,417,1903]" captionTargetId="figure-120@4.[151,1436,417,1903]" captionTargetPageId="4" captionText="FIGURE 3. Tiwaripotamon nganima n. sp. A–F, holotype male (59.3 × 43.6 mm) (ZRC2022.0810), Vietnam; G, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A, chelae; B, left third maxilliped; C, right fourth ambulatory leg; D, male sternopleonal cavity; E, male posterior thoracic sternites and pleon; F, G, male pleon." figureDoi="http://doi.org/10.5281/zenodo.12681715" httpUri="https://zenodo.org/record/12681715/files/figure.png" pageId="1" pageNumber="299">3D</figureCitation>
). Ambulatory legs relatively long (
<figureCitation id="13309F8DFFA7FF82FBF266B4FED913E1" box="[216,347,1447,1473]" captionStart="FIGURE 1" captionStartId="2.[151,250,1899,1923]" captionTargetBox="[151,1435,179,1874]" captionTargetId="figure-15@2.[151,1435,179,1874]" captionTargetPageId="2" captionText="FIGURE 1. Tiwaripotamon nganima n. sp., holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; A, overall dorsal habitus; B, ventral view of habitus; C, frontal view." figureDoi="http://doi.org/10.5281/zenodo.12681708" httpUri="https://zenodo.org/record/12681708/files/figure.png" pageId="1" pageNumber="299">Figs. 1A, B</figureCitation>
,
<figureCitation id="13309F8DFFA7FF82FA4C66B4FE0813E1" box="[358,394,1447,1473]" captionStart="FIGURE 3" captionStartId="4.[151,250,1926,1950]" captionTargetBox="[151,1436,417,1903]" captionTargetId="figure-120@4.[151,1436,417,1903]" captionTargetPageId="4" captionText="FIGURE 3. Tiwaripotamon nganima n. sp. A–F, holotype male (59.3 × 43.6 mm) (ZRC2022.0810), Vietnam; G, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A, chelae; B, left third maxilliped; C, right fourth ambulatory leg; D, male sternopleonal cavity; E, male posterior thoracic sternites and pleon; F, G, male pleon." figureDoi="http://doi.org/10.5281/zenodo.12681715" httpUri="https://zenodo.org/record/12681715/files/figure.png" pageId="1" pageNumber="299">3C</figureCitation>
). Male pleon proportionately more quadrate, somites 3 and 4 relatively less wide (
<figureCitation id="13309F8DFFA7FF82FE3166B4FA1213E1" box="[1307,1424,1447,1473]" captionStart="FIGURE 3" captionStartId="4.[151,250,1926,1950]" captionTargetBox="[151,1436,417,1903]" captionTargetId="figure-120@4.[151,1436,417,1903]" captionTargetPageId="4" captionText="FIGURE 3. Tiwaripotamon nganima n. sp. A–F, holotype male (59.3 × 43.6 mm) (ZRC2022.0810), Vietnam; G, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A, chelae; B, left third maxilliped; C, right fourth ambulatory leg; D, male sternopleonal cavity; E, male posterior thoracic sternites and pleon; F, G, male pleon." figureDoi="http://doi.org/10.5281/zenodo.12681715" httpUri="https://zenodo.org/record/12681715/files/figure.png" pageId="1" pageNumber="299">Fig. 3F, G</figureCitation>
). G1 with subterminal segment strongly curved outwards, outer margin forming deep concavity; terminal segment sharply tapering, distinctly upcurved, outer margin of basal part dilated, forming distinct swelling (
<figureCitation id="13309F8DFFA7FF82FE2566FCFA1A1029" box="[1295,1432,1519,1545]" captionStart="FIGURE 4" captionStartId="5.[151,250,1308,1332]" captionTargetBox="[151,1436,181,1284]" captionTargetId="figure-25@5.[151,1436,181,1284]" captionTargetPageId="5" captionText="FIGURE 4. Tiwaripotamon nganima n. sp. A–E, holotype male (59.3 × 43.6 mm) (ZRC2022.0810), Vietnam; F, G, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam. A, left G1 (ventral view); B, left G1 (dorsal view); C, distal part of left G1 (ventral view); D, distal part of left G1 (dorsal view); E, left G2; F, female anterior thoracic sternites and pleon; G, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.12681717" httpUri="https://zenodo.org/record/12681717/files/figure.png" pageId="1" pageNumber="299">Figs. 4A–D</figureCitation>
,
<figureCitation id="13309F8DFFA7FF82FBBD6501FF63100D" box="[151,225,1554,1581]" captionStart="FIGURE 5" captionStartId="6.[151,250,1463,1487]" captionTargetBox="[160,1428,189,1432]" captionTargetId="figure-15@6.[151,1436,181,1440]" captionTargetPageId="6" captionText="FIGURE 5. Tiwaripotamon nganima n. sp. A–E, holotype male (59.3 × 43.6 mm) (ZRC2022.0810), Vietnam; F–H, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A, left G1 (ventral view); B, F, distal part of left G1 (ventral view); C, G, distal part of left G1 (dorsomesial view); D, H, distal part of left G1 (dorsal view); E, left G2. Scales: A, E = 1.0 mm; B–D, F–H = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681719" httpUri="https://zenodo.org/record/12681719/files/figure.png" pageId="1" pageNumber="299">5A–D</figureCitation>
). Vulva large, ovate and opens obliquely inwards, occupying most of sternite 6, gently pressing onto the suture with sternite 5 (
<figureCitation id="13309F8DFFA7FF82FAB86524FE6F1071" box="[402,493,1591,1617]" captionStart="FIGURE 4" captionStartId="5.[151,250,1308,1332]" captionTargetBox="[151,1436,181,1284]" captionTargetId="figure-25@5.[151,1436,181,1284]" captionTargetPageId="5" captionText="FIGURE 4. Tiwaripotamon nganima n. sp. A–E, holotype male (59.3 × 43.6 mm) (ZRC2022.0810), Vietnam; F, G, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam. A, left G1 (ventral view); B, left G1 (dorsal view); C, distal part of left G1 (ventral view); D, distal part of left G1 (dorsal view); E, left G2; F, female anterior thoracic sternites and pleon; G, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.12681717" httpUri="https://zenodo.org/record/12681717/files/figure.png" pageId="1" pageNumber="299">Fig. 4G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C311D083FFA7FF84FBED6549FE7A178D" lastPageId="7" lastPageNumber="305" pageId="1" pageNumber="299" type="description">
<paragraph id="8BB48308FFA7FF80FBED6549FCF9173D" blockId="1.[151,1437,1050,2013]" lastBlockId="3.[151,1436,151,285]" lastPageId="3" lastPageNumber="301" pageId="1" pageNumber="299">
<emphasis id="B97F5F1AFFA7FF82FBED6549FDA61055" bold="true" box="[199,548,1626,1653]" pageId="1" pageNumber="299">
Description of
<typeStatus id="54B03DAAFFA7FF82FA5C6548FE5F1055" box="[374,477,1627,1653]" pageId="1" pageNumber="299" type="holotype">holotype</typeStatus>
male.
</emphasis>
Carapace subquadrate, wider than long; dorsal surfaces smooth, regions poorly demarcated, cervical grooves barely visible; H-shaped gastro-cardiac depression shallow but discernible (
<figureCitation id="13309F8DFFA7FF82FE1E656CFA1A10B9" box="[1332,1432,1663,1689]" captionStart="FIGURE 1" captionStartId="2.[151,250,1899,1923]" captionTargetBox="[151,1435,179,1874]" captionTargetId="figure-15@2.[151,1435,179,1874]" captionTargetPageId="2" captionText="FIGURE 1. Tiwaripotamon nganima n. sp., holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; A, overall dorsal habitus; B, ventral view of habitus; C, frontal view." figureDoi="http://doi.org/10.5281/zenodo.12681708" httpUri="https://zenodo.org/record/12681708/files/figure.png" pageId="1" pageNumber="299">Figs. 1A</figureCitation>
,
<figureCitation id="13309F8DFFA7FF82FBBD65B0FF3F109D" box="[151,189,1699,1725]" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="1" pageNumber="299">2A</figureCitation>
); dorsal surface (including branchial regions) gently convex, gently inflated (
<figureCitation id="13309F8DFFA7FF82FF2965B0FB00109D" box="[1027,1154,1699,1725]" captionStart="FIGURE 1" captionStartId="2.[151,250,1899,1923]" captionTargetBox="[151,1435,179,1874]" captionTargetId="figure-15@2.[151,1435,179,1874]" captionTargetPageId="2" captionText="FIGURE 1. Tiwaripotamon nganima n. sp., holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; A, overall dorsal habitus; B, ventral view of habitus; C, frontal view." figureDoi="http://doi.org/10.5281/zenodo.12681708" httpUri="https://zenodo.org/record/12681708/files/figure.png" pageId="1" pageNumber="299">Figs. 1A, C</figureCitation>
,
<figureCitation id="13309F8DFFA7FF82FFA165B0FB4D109D" box="[1163,1231,1699,1725]" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="1" pageNumber="299">2A,D</figureCitation>
); epigastric cristae very low, marked by low, short striae; postorbital cristae not visible (
<figureCitation id="13309F8DFFA7FF82F8B165D4FC7D10C1" box="[923,1023,1735,1761]" captionStart="FIGURE 1" captionStartId="2.[151,250,1899,1923]" captionTargetBox="[151,1435,179,1874]" captionTargetId="figure-15@2.[151,1435,179,1874]" captionTargetPageId="2" captionText="FIGURE 1. Tiwaripotamon nganima n. sp., holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; A, overall dorsal habitus; B, ventral view of habitus; C, frontal view." figureDoi="http://doi.org/10.5281/zenodo.12681708" httpUri="https://zenodo.org/record/12681708/files/figure.png" pageId="1" pageNumber="299">Figs. 1A</figureCitation>
,
<figureCitation id="13309F8DFFA7FF82FF2165D4FBB310C1" box="[1035,1073,1735,1761]" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="1" pageNumber="299">2A</figureCitation>
). Frontal margin gently sinuous, lobes not well marked, appears almost entire in dorsal view, level with tips of external orbital tooth in dorsal view; median part distinctly deflexed; contiguous with supraorbital margin (
<figureCitation id="13309F8DFFA7FF82F88F641CFB8B1109" box="[933,1033,1807,1833]" captionStart="FIGURE 1" captionStartId="2.[151,250,1899,1923]" captionTargetBox="[151,1435,179,1874]" captionTargetId="figure-15@2.[151,1435,179,1874]" captionTargetPageId="2" captionText="FIGURE 1. Tiwaripotamon nganima n. sp., holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; A, overall dorsal habitus; B, ventral view of habitus; C, frontal view." figureDoi="http://doi.org/10.5281/zenodo.12681708" httpUri="https://zenodo.org/record/12681708/files/figure.png" pageId="1" pageNumber="299">Figs. 1A</figureCitation>
,
<figureCitation id="13309F8DFFA7FF82FF3E641CFBDE1109" box="[1044,1116,1807,1833]" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="1" pageNumber="299">2A, D</figureCitation>
). Supraorbital margin gently sinuous, without clefts or fissures (
<figureCitation id="13309F8DFFA7FF82F90A6420FD06116D" box="[544,644,1843,1869]" captionStart="FIGURE 1" captionStartId="2.[151,250,1899,1923]" captionTargetBox="[151,1435,179,1874]" captionTargetId="figure-15@2.[151,1435,179,1874]" captionTargetPageId="2" captionText="FIGURE 1. Tiwaripotamon nganima n. sp., holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; A, overall dorsal habitus; B, ventral view of habitus; C, frontal view." figureDoi="http://doi.org/10.5281/zenodo.12681708" httpUri="https://zenodo.org/record/12681708/files/figure.png" pageId="1" pageNumber="299">Figs. 1A</figureCitation>
,
<figureCitation id="13309F8DFFA7FF82F9BA6420FD34116D" box="[656,694,1843,1869]" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="1" pageNumber="299">2A</figureCitation>
). External orbital tooth low, broad, outer margin longer than inner margin; barely demarcated from anterolateral margin by indentation (
<figureCitation id="13309F8DFFA7FF82F8BE6444FC6E1151" box="[916,1004,1879,1905]" captionStart="FIGURE 3" captionStartId="4.[151,250,1926,1950]" captionTargetBox="[151,1436,417,1903]" captionTargetId="figure-120@4.[151,1436,417,1903]" captionTargetPageId="4" captionText="FIGURE 3. Tiwaripotamon nganima n. sp. A–F, holotype male (59.3 × 43.6 mm) (ZRC2022.0810), Vietnam; G, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A, chelae; B, left third maxilliped; C, right fourth ambulatory leg; D, male sternopleonal cavity; E, male posterior thoracic sternites and pleon; F, G, male pleon." figureDoi="http://doi.org/10.5281/zenodo.12681715" httpUri="https://zenodo.org/record/12681715/files/figure.png" pageId="1" pageNumber="299">Fig. 3A</figureCitation>
); epibranchial tooth low hardly visible, rest of anterolateral margin convex, subcristate, serrate; posterolateral margin gently sinuous, converging towards gently convex posterior carapace margin (
<figureCitation id="13309F8DFFA7FF82F95A648CFD571199" box="[624,725,1951,1977]" captionStart="FIGURE 1" captionStartId="2.[151,250,1899,1923]" captionTargetBox="[151,1435,179,1874]" captionTargetId="figure-15@2.[151,1435,179,1874]" captionTargetPageId="2" captionText="FIGURE 1. Tiwaripotamon nganima n. sp., holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; A, overall dorsal habitus; B, ventral view of habitus; C, frontal view." figureDoi="http://doi.org/10.5281/zenodo.12681708" httpUri="https://zenodo.org/record/12681708/files/figure.png" pageId="1" pageNumber="299">Figs. 1A</figureCitation>
,
<figureCitation id="13309F8DFFA7FF82F9CB648CFC851199" box="[737,775,1951,1977]" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="1" pageNumber="299">2A</figureCitation>
). Orbits ovate, large; eyes completely filling space; cornea large, peduncle not elongate (
<figureCitation id="13309F8DFFA7FF82FACE64D0FDC411FD" box="[484,582,1987,2013]" captionStart="FIGURE 1" captionStartId="2.[151,250,1899,1923]" captionTargetBox="[151,1435,179,1874]" captionTargetId="figure-15@2.[151,1435,179,1874]" captionTargetPageId="2" captionText="FIGURE 1. Tiwaripotamon nganima n. sp., holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; A, overall dorsal habitus; B, ventral view of habitus; C, frontal view." figureDoi="http://doi.org/10.5281/zenodo.12681708" httpUri="https://zenodo.org/record/12681708/files/figure.png" pageId="1" pageNumber="299">Figs. 1C</figureCitation>
,
<figureCitation id="13309F8DFFA7FF82F97964D0FD1E11FD" box="[595,668,1987,2013]" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="1" pageNumber="299">2A, D</figureCitation>
). Antennules longitudinally narrow, folding laterally (
<figureCitation id="13309F8DFFA7FF82FFD064D0FADE11FD" box="[1274,1372,1987,2013]" captionStart="FIGURE 1" captionStartId="2.[151,250,1899,1923]" captionTargetBox="[151,1435,179,1874]" captionTargetId="figure-15@2.[151,1435,179,1874]" captionTargetPageId="2" captionText="FIGURE 1. Tiwaripotamon nganima n. sp., holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; A, overall dorsal habitus; B, ventral view of habitus; C, frontal view." figureDoi="http://doi.org/10.5281/zenodo.12681708" httpUri="https://zenodo.org/record/12681708/files/figure.png" pageId="1" pageNumber="299">Figs. 1C</figureCitation>
,
<figureCitation id="13309F8DFFA7FF82FE4364D0FA0D11FD" box="[1385,1423,1987,2013]" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="1" pageNumber="299">2D</figureCitation>
); antennae short, not extending beyond orbit (
<figureCitation id="13309F8DFFA5FF80F98E6384FC891691" box="[676,779,151,177]" captionStart="FIGURE 1" captionStartId="2.[151,250,1899,1923]" captionTargetBox="[151,1435,179,1874]" captionTargetId="figure-15@2.[151,1435,179,1874]" captionTargetPageId="2" captionText="FIGURE 1. Tiwaripotamon nganima n. sp., holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; A, overall dorsal habitus; B, ventral view of habitus; C, frontal view." figureDoi="http://doi.org/10.5281/zenodo.12681708" httpUri="https://zenodo.org/record/12681708/files/figure.png" pageId="3" pageNumber="301">Figs. 1C</figureCitation>
,
<figureCitation id="13309F8DFFA5FF80F8376384FCC11691" box="[797,835,151,177]" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="3" pageNumber="301">2D</figureCitation>
). Epistome longitudinally narrow; posterior margin with broadly triangular median lobe, separated from gently sinuous lateral margins by fissure (
<figureCitation id="13309F8DFFA5FF80FFFD63A8FAB916F5" box="[1239,1339,187,213]" captionStart="FIGURE 1" captionStartId="2.[151,250,1899,1923]" captionTargetBox="[151,1435,179,1874]" captionTargetId="figure-15@2.[151,1435,179,1874]" captionTargetPageId="2" captionText="FIGURE 1. Tiwaripotamon nganima n. sp., holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; A, overall dorsal habitus; B, ventral view of habitus; C, frontal view." figureDoi="http://doi.org/10.5281/zenodo.12681708" httpUri="https://zenodo.org/record/12681708/files/figure.png" pageId="3" pageNumber="301">Figs. 1C</figureCitation>
,
<figureCitation id="13309F8DFFA5FF80FE6063A8FA0D16F5" box="[1354,1423,187,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="3" pageNumber="301">2D, F</figureCitation>
). Suborbital margin convex, smooth, without inner tooth; suborbital region smooth, subhepatic, pterygostomial and sub-branchial regions weakly rugose to smooth (
<figureCitation id="13309F8DFFA5FF80F9926210FC9B173D" box="[696,793,259,285]" captionStart="FIGURE 1" captionStartId="2.[151,250,1899,1923]" captionTargetBox="[151,1435,179,1874]" captionTargetId="figure-15@2.[151,1435,179,1874]" captionTargetPageId="2" captionText="FIGURE 1. Tiwaripotamon nganima n. sp., holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; A, overall dorsal habitus; B, ventral view of habitus; C, frontal view." figureDoi="http://doi.org/10.5281/zenodo.12681708" httpUri="https://zenodo.org/record/12681708/files/figure.png" pageId="3" pageNumber="301">Figs. 1C</figureCitation>
,
<figureCitation id="13309F8DFFA5FF80F80C6210FCEC173D" box="[806,878,259,285]" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="3" pageNumber="301">2D, H</figureCitation>
).
</paragraph>
<caption id="DF74D380FFA4FF81FBBD6478FD221188" ID-DOI="http://doi.org/10.5281/zenodo.12681708" ID-Zenodo-Dep="12681708" httpUri="https://zenodo.org/record/12681708/files/figure.png" pageId="2" pageNumber="300" startId="2.[151,250,1899,1923]" targetBox="[151,1435,179,1874]" targetPageId="2" targetType="figure">
<paragraph id="8BB48308FFA4FF81FBBD6478FD221188" blockId="2.[151,1436,1899,1960]" pageId="2" pageNumber="300">
<emphasis id="B97F5F1AFFA4FF81FBBD6478FE9711A3" bold="true" box="[151,277,1899,1923]" pageId="2" pageNumber="300">FIGURE 1.</emphasis>
<taxonomicName id="4C0BF88BFFA4FF81FA37647FFD9811A3" authorityName="Ng" authorityYear="2024" box="[285,538,1900,1923]" class="Malacostraca" family="Potamidae" genus="Tiwaripotamon" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="300" phylum="Arthropoda" rank="species" species="nganima" status="sp. nov.">
<emphasis id="B97F5F1AFFA4FF81FA37647FFD9811A3" box="[285,538,1900,1923]" italics="true" pageId="2" pageNumber="300">Tiwaripotamon nganima</emphasis>
</taxonomicName>
<emphasis id="B97F5F1AFFA4FF81F908647FFDDF11A4" bold="true" box="[546,605,1900,1924]" pageId="2" pageNumber="300">
<taxonomicNameLabel id="A24CE261FFA4FF81F908647FFDDF11A4" box="[546,605,1900,1924]" pageId="2" pageNumber="300" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; A, overall dorsal habitus; B, ventral view of habitus; C, frontal view.
</paragraph>
</caption>
<caption id="DF74D380FFA5FF80FBBD645FFEDE11F0" ID-DOI="http://doi.org/10.5281/zenodo.12681711" ID-Zenodo-Dep="12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="3" pageNumber="301" startId="3.[151,250,1868,1892]" targetBox="[151,1435,309,1845]" targetPageId="3" targetType="figure">
<paragraph id="8BB48308FFA5FF80FBBD645FFEDE11F0" blockId="3.[151,1437,1868,2000]" pageId="3" pageNumber="301">
<emphasis id="B97F5F1AFFA5FF80FBBD645FFE901144" bold="true" box="[151,274,1868,1892]" pageId="3" pageNumber="301">FIGURE 2.</emphasis>
<taxonomicName id="4C0BF88BFFA5FF80FA3D645EFD921144" authorityName="Ng" authorityYear="2024" box="[279,528,1869,1892]" class="Malacostraca" family="Potamidae" genus="Tiwaripotamon" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="301" phylum="Arthropoda" rank="species" species="nganima" status="sp. nov.">
<emphasis id="B97F5F1AFFA5FF80FA3D645EFD921144" box="[279,528,1869,1892]" italics="true" pageId="3" pageNumber="301">Tiwaripotamon nganima</emphasis>
</taxonomicName>
<emphasis id="B97F5F1AFFA5FF80F93F645FFDCF1144" bold="true" box="[533,589,1868,1892]" pageId="3" pageNumber="301">
<taxonomicNameLabel id="A24CE261FFA5FF80F93F645FFDCF1144" box="[533,589,1868,1892]" pageId="3" pageNumber="301" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon.
</paragraph>
</caption>
<paragraph id="8BB48308FFA2FF87FBED6384FD7D16D9" blockId="4.[151,1436,151,393]" pageId="4" pageNumber="302">
Third maxillipeds relatively short; merus shorter and narrower than ischium, surface slightly rugose; ischium subquadrate with no visible median sulcus; exopod short, curved, reaching to one-third length of merus, with short flagellum shorter than merus width (
<figureCitation id="13309F8DFFA2FF87F91A63CCFD1316D9" box="[560,657,223,249]" captionStart="FIGURE 1" captionStartId="2.[151,250,1899,1923]" captionTargetBox="[151,1435,179,1874]" captionTargetId="figure-15@2.[151,1435,179,1874]" captionTargetPageId="2" captionText="FIGURE 1. Tiwaripotamon nganima n. sp., holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; A, overall dorsal habitus; B, ventral view of habitus; C, frontal view." figureDoi="http://doi.org/10.5281/zenodo.12681708" httpUri="https://zenodo.org/record/12681708/files/figure.png" pageId="4" pageNumber="302">Figs. 1C</figureCitation>
,
<figureCitation id="13309F8DFFA2FF87F9B463CCFD4016D9" box="[670,706,223,249]" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="4" pageNumber="302">2H</figureCitation>
,
<figureCitation id="13309F8DFFA2FF87F9E463CCFD7316D9" box="[718,753,223,249]" captionStart="FIGURE 3" captionStartId="4.[151,250,1926,1950]" captionTargetBox="[151,1436,417,1903]" captionTargetId="figure-120@4.[151,1436,417,1903]" captionTargetPageId="4" captionText="FIGURE 3. Tiwaripotamon nganima n. sp. A–F, holotype male (59.3 × 43.6 mm) (ZRC2022.0810), Vietnam; G, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A, chelae; B, left third maxilliped; C, right fourth ambulatory leg; D, male sternopleonal cavity; E, male posterior thoracic sternites and pleon; F, G, male pleon." figureDoi="http://doi.org/10.5281/zenodo.12681715" httpUri="https://zenodo.org/record/12681715/files/figure.png" pageId="4" pageNumber="302">3B</figureCitation>
).
</paragraph>
<paragraph id="8BB48308FFA2FF87FBED6210FACC17A9" blockId="4.[151,1436,151,393]" pageId="4" pageNumber="302">
Chelipeds elongate; left one prominently larger (
<figureCitation id="13309F8DFFA2FF87F82A6210FC08173D" box="[768,906,259,285]" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="4" pageNumber="302">Figs. 2A–C</figureCitation>
,
<figureCitation id="13309F8DFFA2FF87F8B36210FC3C173D" box="[921,958,259,285]" captionStart="FIGURE 3" captionStartId="4.[151,250,1926,1950]" captionTargetBox="[151,1436,417,1903]" captionTargetId="figure-120@4.[151,1436,417,1903]" captionTargetPageId="4" captionText="FIGURE 3. Tiwaripotamon nganima n. sp. A–F, holotype male (59.3 × 43.6 mm) (ZRC2022.0810), Vietnam; G, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A, chelae; B, left third maxilliped; C, right fourth ambulatory leg; D, male sternopleonal cavity; E, male posterior thoracic sternites and pleon; F, G, male pleon." figureDoi="http://doi.org/10.5281/zenodo.12681715" httpUri="https://zenodo.org/record/12681715/files/figure.png" pageId="4" pageNumber="302">3A</figureCitation>
). Basis-ischium with granules on ventral margin; ventral margins of merus lined with tubercles, dorsal margin uneven, distinctly rugose; carpus with long sharp spine on inner angle, surface slightly rugose; outer surface of chela almost smooth, surface of fingers punctate; fingers longer than palm; fingers gently curved with large cutting teeth evenly spread out (
<figureCitation id="13309F8DFFA2FF87FFA6627CFA9217A9" box="[1164,1296,367,393]" captionStart="FIGURE 1" captionStartId="2.[151,250,1899,1923]" captionTargetBox="[151,1435,179,1874]" captionTargetId="figure-15@2.[151,1435,179,1874]" captionTargetPageId="2" captionText="FIGURE 1. Tiwaripotamon nganima n. sp., holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; A, overall dorsal habitus; B, ventral view of habitus; C, frontal view." figureDoi="http://doi.org/10.5281/zenodo.12681708" httpUri="https://zenodo.org/record/12681708/files/figure.png" pageId="4" pageNumber="302">Figs. 1A–C</figureCitation>
,
<figureCitation id="13309F8DFFA2FF87FE36627CFAC317A9" box="[1308,1345,367,393]" captionStart="FIGURE 3" captionStartId="4.[151,250,1926,1950]" captionTargetBox="[151,1436,417,1903]" captionTargetId="figure-120@4.[151,1436,417,1903]" captionTargetPageId="4" captionText="FIGURE 3. Tiwaripotamon nganima n. sp. A–F, holotype male (59.3 × 43.6 mm) (ZRC2022.0810), Vietnam; G, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A, chelae; B, left third maxilliped; C, right fourth ambulatory leg; D, male sternopleonal cavity; E, male posterior thoracic sternites and pleon; F, G, male pleon." figureDoi="http://doi.org/10.5281/zenodo.12681715" httpUri="https://zenodo.org/record/12681715/files/figure.png" pageId="4" pageNumber="302">3A</figureCitation>
).
</paragraph>
<caption id="DF74D380FFA2FF87FBBD6495FC7511C6" ID-DOI="http://doi.org/10.5281/zenodo.12681715" ID-Zenodo-Dep="12681715" httpUri="https://zenodo.org/record/12681715/files/figure.png" pageId="4" pageNumber="302" startId="4.[151,250,1926,1950]" targetBox="[151,1436,417,1903]" targetPageId="4" targetType="figure">
<paragraph id="8BB48308FFA2FF87FBBD6495FC7511C6" blockId="4.[151,1436,1926,2023]" pageId="4" pageNumber="302">
<emphasis id="B97F5F1AFFA2FF87FBBD6495FE9411BE" bold="true" box="[151,278,1926,1950]" pageId="4" pageNumber="302">FIGURE 3.</emphasis>
<taxonomicName id="4C0BF88BFFA2FF87FA356494FD9E11BE" authorityName="Ng" authorityYear="2024" box="[287,540,1927,1950]" class="Malacostraca" family="Potamidae" genus="Tiwaripotamon" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="302" phylum="Arthropoda" rank="species" species="nganima" status="sp. nov.">
<emphasis id="B97F5F1AFFA2FF87FA356494FD9E11BE" box="[287,540,1927,1950]" italics="true" pageId="4" pageNumber="302">Tiwaripotamon nganima</emphasis>
</taxonomicName>
<emphasis id="B97F5F1AFFA2FF87F90E6494FDE211BF" bold="true" box="[548,608,1927,1951]" pageId="4" pageNumber="302">
<taxonomicNameLabel id="A24CE261FFA2FF87F90E6494FDE211BF" box="[548,608,1927,1951]" pageId="4" pageNumber="302" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
A–F, holotype male (59.3 × 43.6 mm) (ZRC2022.0810), Vietnam; G, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A, chelae; B, left third maxilliped; C, right fourth ambulatory leg; D, male sternopleonal cavity; E, male posterior thoracic sternites and pleon; F, G, male pleon.
</paragraph>
</caption>
<caption id="DF74D380FFA3FF86FBBD660FFE621380" ID-DOI="http://doi.org/10.5281/zenodo.12681717" ID-Zenodo-Dep="12681717" httpUri="https://zenodo.org/record/12681717/files/figure.png" pageId="5" pageNumber="303" startId="5.[151,250,1308,1332]" targetBox="[151,1436,181,1284]" targetPageId="5" targetType="figure">
<paragraph id="8BB48308FFA3FF86FBBD660FFE621380" blockId="5.[151,1436,1308,1440]" pageId="5" pageNumber="303">
<emphasis id="B97F5F1AFFA3FF86FBBD660FFE961314" bold="true" box="[151,276,1308,1332]" pageId="5" pageNumber="303">FIGURE 4.</emphasis>
<taxonomicName id="4C0BF88BFFA3FF86FA31660EFD941314" authorityName="Ng" authorityYear="2024" box="[283,534,1309,1332]" class="Malacostraca" family="Potamidae" genus="Tiwaripotamon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="303" phylum="Arthropoda" rank="species" species="nganima" status="sp. nov.">
<emphasis id="B97F5F1AFFA3FF86FA31660EFD941314" box="[283,534,1309,1332]" italics="true" pageId="5" pageNumber="303">Tiwaripotamon nganima</emphasis>
</taxonomicName>
<emphasis id="B97F5F1AFFA3FF86F937660FFDD51314" bold="true" box="[541,599,1308,1332]" pageId="5" pageNumber="303">
<taxonomicNameLabel id="A24CE261FFA3FF86F937660FFDD51314" box="[541,599,1308,1332]" pageId="5" pageNumber="303" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
A–E, holotype male (59.3 × 43.6 mm) (ZRC2022.0810), Vietnam; F, G, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam. A, left G1 (ventral view); B, left G1 (dorsal view); C, distal part of left G1 (ventral view); D, distal part of left G1 (dorsal view); E, left G2; F, female anterior thoracic sternites and pleon; G, female sternopleonal cavity and vulvae.
</paragraph>
</caption>
<paragraph id="8BB48308FFA3FF86FBED66DDFD831011" blockId="5.[151,1437,1486,2017]" pageId="5" pageNumber="303">
Ambulatory legs relatively long; second leg longest, fourth leg shortest; merus with dorsal margin subcristate, margin uneven but not serrate; carpus with subdorsal ridge; propodus long, with shallow median sulcus; dactylus gently curved (
<figureCitation id="13309F8DFFA3FF86FA6A6505FE461011" box="[320,452,1558,1585]" captionStart="FIGURE 1" captionStartId="2.[151,250,1899,1923]" captionTargetBox="[151,1435,179,1874]" captionTargetId="figure-15@2.[151,1435,179,1874]" captionTargetPageId="2" captionText="FIGURE 1. Tiwaripotamon nganima n. sp., holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; A, overall dorsal habitus; B, ventral view of habitus; C, frontal view." figureDoi="http://doi.org/10.5281/zenodo.12681708" httpUri="https://zenodo.org/record/12681708/files/figure.png" pageId="5" pageNumber="303">Figs. 1A, B</figureCitation>
,
<figureCitation id="13309F8DFFA3FF86FAFA6504FE761011" box="[464,500,1559,1585]" captionStart="FIGURE 3" captionStartId="4.[151,250,1926,1950]" captionTargetBox="[151,1436,417,1903]" captionTargetId="figure-120@4.[151,1436,417,1903]" captionTargetPageId="4" captionText="FIGURE 3. Tiwaripotamon nganima n. sp. A–F, holotype male (59.3 × 43.6 mm) (ZRC2022.0810), Vietnam; G, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A, chelae; B, left third maxilliped; C, right fourth ambulatory leg; D, male sternopleonal cavity; E, male posterior thoracic sternites and pleon; F, G, male pleon." figureDoi="http://doi.org/10.5281/zenodo.12681715" httpUri="https://zenodo.org/record/12681715/files/figure.png" pageId="5" pageNumber="303">3C</figureCitation>
).
</paragraph>
<paragraph id="8BB48308FFA3FF86FBED6529FBD710C5" blockId="5.[151,1437,1486,2017]" pageId="5" pageNumber="303">
Thoracic sternites 1 and 2 completely fused forming triangular plate with convex lateral margins; separated from sternite 3 by straight deep suture; sternites 3 and 4 fused except for shallow lateral groove that is medially interrupted, surface of sternites punctate (
<figureCitation id="13309F8DFFA3FF86F94D6591FD4A10BD" box="[615,712,1666,1693]" captionStart="FIGURE 1" captionStartId="2.[151,250,1899,1923]" captionTargetBox="[151,1435,179,1874]" captionTargetId="figure-15@2.[151,1435,179,1874]" captionTargetPageId="2" captionText="FIGURE 1. Tiwaripotamon nganima n. sp., holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; A, overall dorsal habitus; B, ventral view of habitus; C, frontal view." figureDoi="http://doi.org/10.5281/zenodo.12681708" httpUri="https://zenodo.org/record/12681708/files/figure.png" pageId="5" pageNumber="303">Figs. 1B</figureCitation>
,
<figureCitation id="13309F8DFFA3FF86F9FE6590FD7A10BD" box="[724,760,1667,1693]" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="5" pageNumber="303">2H</figureCitation>
,
<figureCitation id="13309F8DFFA3FF86F82E6590FCA810BD" box="[772,810,1667,1693]" captionStart="FIGURE 3" captionStartId="4.[151,250,1926,1950]" captionTargetBox="[151,1436,417,1903]" captionTargetId="figure-120@4.[151,1436,417,1903]" captionTargetPageId="4" captionText="FIGURE 3. Tiwaripotamon nganima n. sp. A–F, holotype male (59.3 × 43.6 mm) (ZRC2022.0810), Vietnam; G, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A, chelae; B, left third maxilliped; C, right fourth ambulatory leg; D, male sternopleonal cavity; E, male posterior thoracic sternites and pleon; F, G, male pleon." figureDoi="http://doi.org/10.5281/zenodo.12681715" httpUri="https://zenodo.org/record/12681715/files/figure.png" pageId="5" pageNumber="303">3D</figureCitation>
). Somite 8 not visible when pleon closed. Sternopleonal cavity reaching to imaginary line connecting submedian part of coxae of chelipeds, distal part subcristate; tubercle of male pleonal locking mechanism on posterior third of somite 5 (
<figureCitation id="13309F8DFFA3FF86F8AF65D9FC6410C5" box="[901,998,1738,1765]" captionStart="FIGURE 1" captionStartId="2.[151,250,1899,1923]" captionTargetBox="[151,1435,179,1874]" captionTargetId="figure-15@2.[151,1435,179,1874]" captionTargetPageId="2" captionText="FIGURE 1. Tiwaripotamon nganima n. sp., holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; A, overall dorsal habitus; B, ventral view of habitus; C, frontal view." figureDoi="http://doi.org/10.5281/zenodo.12681708" httpUri="https://zenodo.org/record/12681708/files/figure.png" pageId="5" pageNumber="303">Figs. 1B</figureCitation>
,
<figureCitation id="13309F8DFFA3FF86F8D965D8FB9510C5" box="[1011,1047,1739,1765]" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="5" pageNumber="303">2H</figureCitation>
,
<figureCitation id="13309F8DFFA3FF86FF0965D8FBCB10C5" box="[1059,1097,1739,1765]" captionStart="FIGURE 3" captionStartId="4.[151,250,1926,1950]" captionTargetBox="[151,1436,417,1903]" captionTargetId="figure-120@4.[151,1436,417,1903]" captionTargetPageId="4" captionText="FIGURE 3. Tiwaripotamon nganima n. sp. A–F, holotype male (59.3 × 43.6 mm) (ZRC2022.0810), Vietnam; G, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A, chelae; B, left third maxilliped; C, right fourth ambulatory leg; D, male sternopleonal cavity; E, male posterior thoracic sternites and pleon; F, G, male pleon." figureDoi="http://doi.org/10.5281/zenodo.12681715" httpUri="https://zenodo.org/record/12681715/files/figure.png" pageId="5" pageNumber="303">3D</figureCitation>
).
</paragraph>
<paragraph id="8BB48308FFA3FF86FBED65FDFCB01171" blockId="5.[151,1437,1486,2017]" pageId="5" pageNumber="303">
Pleon broadly triangular; somite 1 longitudinally narrow, reaching to edges of coxae of fourth ambulatory legs; somite 2 slightly wider than somite 1; somite 3 widest; somites 4 and 5 trapezoidal; somite 6 sub-rectangular, wide; telson wide, lateral margin gently convex (
<figureCitation id="13309F8DFFA3FF86F95F6425FD5B1171" box="[629,729,1846,1873]" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="5" pageNumber="303">Figs. 2H</figureCitation>
,
<figureCitation id="13309F8DFFA3FF86F9CF6424FCA71171" box="[741,805,1847,1873]" captionStart="FIGURE 3" captionStartId="4.[151,250,1926,1950]" captionTargetBox="[151,1436,417,1903]" captionTargetId="figure-120@4.[151,1436,417,1903]" captionTargetPageId="4" captionText="FIGURE 3. Tiwaripotamon nganima n. sp. A–F, holotype male (59.3 × 43.6 mm) (ZRC2022.0810), Vietnam; G, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A, chelae; B, left third maxilliped; C, right fourth ambulatory leg; D, male sternopleonal cavity; E, male posterior thoracic sternites and pleon; F, G, male pleon." figureDoi="http://doi.org/10.5281/zenodo.12681715" httpUri="https://zenodo.org/record/12681715/files/figure.png" pageId="5" pageNumber="303">3E, F</figureCitation>
).
</paragraph>
<paragraph id="8BB48308FFA3FF86FBED6448FCA811C1" blockId="5.[151,1437,1486,2017]" pageId="5" pageNumber="303">
G1 relatively slender, subterminal segment strongly curved outwards, outer margin forming deep concavity; terminal segment clearly separated from subterminal segment, sharply tapering, distinctly upcurved, outer margin of basal part prominently dilated, forming distinct swelling (
<figureCitation id="13309F8DFFA3FF86F80864B1FC2B119D" box="[802,937,1954,1981]" captionStart="FIGURE 4" captionStartId="5.[151,250,1308,1332]" captionTargetBox="[151,1436,181,1284]" captionTargetId="figure-25@5.[151,1436,181,1284]" captionTargetPageId="5" captionText="FIGURE 4. Tiwaripotamon nganima n. sp. A–E, holotype male (59.3 × 43.6 mm) (ZRC2022.0810), Vietnam; F, G, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam. A, left G1 (ventral view); B, left G1 (dorsal view); C, distal part of left G1 (ventral view); D, distal part of left G1 (dorsal view); E, left G2; F, female anterior thoracic sternites and pleon; G, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.12681717" httpUri="https://zenodo.org/record/12681717/files/figure.png" pageId="5" pageNumber="303">Figs. 4A–D</figureCitation>
,
<figureCitation id="13309F8DFFA3FF86F89F64B1FC7C119D" box="[949,1022,1954,1981]" captionStart="FIGURE 5" captionStartId="6.[151,250,1463,1487]" captionTargetBox="[160,1428,189,1432]" captionTargetId="figure-15@6.[151,1436,181,1440]" captionTargetPageId="6" captionText="FIGURE 5. Tiwaripotamon nganima n. sp. A–E, holotype male (59.3 × 43.6 mm) (ZRC2022.0810), Vietnam; F–H, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A, left G1 (ventral view); B, F, distal part of left G1 (ventral view); C, G, distal part of left G1 (dorsomesial view); D, H, distal part of left G1 (dorsal view); E, left G2. Scales: A, E = 1.0 mm; B–D, F–H = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681719" httpUri="https://zenodo.org/record/12681719/files/figure.png" pageId="5" pageNumber="303">5A–D</figureCitation>
). G2 subequal in length to G1, distal segment two-thirds length of basal segment (
<figureCitation id="13309F8DFFA3FF86F9A464D5FD6D11C1" box="[654,751,1990,2017]" captionStart="FIGURE 4" captionStartId="5.[151,250,1308,1332]" captionTargetBox="[151,1436,181,1284]" captionTargetId="figure-25@5.[151,1436,181,1284]" captionTargetPageId="5" captionText="FIGURE 4. Tiwaripotamon nganima n. sp. A–E, holotype male (59.3 × 43.6 mm) (ZRC2022.0810), Vietnam; F, G, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam. A, left G1 (ventral view); B, left G1 (dorsal view); C, distal part of left G1 (ventral view); D, distal part of left G1 (dorsal view); E, left G2; F, female anterior thoracic sternites and pleon; G, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.12681717" httpUri="https://zenodo.org/record/12681717/files/figure.png" pageId="5" pageNumber="303">Figs. 4E</figureCitation>
,
<figureCitation id="13309F8DFFA3FF86F9D064D5FC9F11C1" box="[762,797,1990,2017]" captionStart="FIGURE 5" captionStartId="6.[151,250,1463,1487]" captionTargetBox="[160,1428,189,1432]" captionTargetId="figure-15@6.[151,1436,181,1440]" captionTargetPageId="6" captionText="FIGURE 5. Tiwaripotamon nganima n. sp. A–E, holotype male (59.3 × 43.6 mm) (ZRC2022.0810), Vietnam; F–H, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A, left G1 (ventral view); B, F, distal part of left G1 (ventral view); C, G, distal part of left G1 (dorsomesial view); D, H, distal part of left G1 (dorsal view); E, left G2. Scales: A, E = 1.0 mm; B–D, F–H = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681719" httpUri="https://zenodo.org/record/12681719/files/figure.png" pageId="5" pageNumber="303">5E</figureCitation>
).
</paragraph>
<caption id="DF74D380FFA0FF85FBBD66A4FE82101B" ID-DOI="http://doi.org/10.5281/zenodo.12681719" ID-Zenodo-Dep="12681719" httpUri="https://zenodo.org/record/12681719/files/figure.png" pageId="6" pageNumber="304" startId="6.[151,250,1463,1487]" targetBox="[160,1428,189,1432]" targetPageId="6" targetType="figure">
<paragraph id="8BB48308FFA0FF85FBBD66A4FE82101B" blockId="6.[151,1436,1463,1595]" pageId="6" pageNumber="304">
<emphasis id="B97F5F1AFFA0FF85FBBD66A4FE9613F0" bold="true" box="[151,276,1463,1488]" pageId="6" pageNumber="304">FIGURE 5.</emphasis>
<taxonomicName id="4C0BF88BFFA0FF85FA3166ABFD9413EF" authorityName="Ng" authorityYear="2024" box="[283,534,1464,1487]" class="Malacostraca" family="Potamidae" genus="Tiwaripotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="304" phylum="Arthropoda" rank="species" species="nganima" status="sp. nov.">
<emphasis id="B97F5F1AFFA0FF85FA3166ABFD9413EF" box="[283,534,1464,1487]" italics="true" pageId="6" pageNumber="304">Tiwaripotamon nganima</emphasis>
</taxonomicName>
<emphasis id="B97F5F1AFFA0FF85F93766ABFDD513F0" bold="true" box="[541,599,1464,1488]" pageId="6" pageNumber="304">
<taxonomicNameLabel id="A24CE261FFA0FF85F93766ABFDD513F0" box="[541,599,1464,1488]" pageId="6" pageNumber="304" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
A–E, holotype male (59.3 × 43.6 mm) (ZRC2022.0810), Vietnam; F–H, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A, left G1 (ventral view); B, F, distal part of left G1 (ventral view); C, G, distal part of left G1 (dorsomesial view); D, H, distal part of left G1 (dorsal view); E, left G2. Scales: A, E = 1.0 mm; B–D, F–H = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BB48308FFA0FF85FBED6579FAB310D0" blockId="6.[151,1437,1641,2028]" pageId="6" pageNumber="304">
<emphasis id="B97F5F1AFFA0FF85FBED6579FE5510A3" bold="true" box="[199,471,1641,1668]" pageId="6" pageNumber="304">Females and variation.</emphasis>
The female specimen agrees with adult males in all aspects except for its more slender and shorter chelipeds. The pleon is ovate and covers almost the entire surface of the thoracic sternum when closed, with the telson semicircular with strongly convex lateral margins (
<figureCitation id="13309F8DFFA0FF85F8BE65A1FC6F10EC" box="[916,1005,1714,1740]" captionStart="FIGURE 4" captionStartId="5.[151,250,1308,1332]" captionTargetBox="[151,1436,181,1284]" captionTargetId="figure-25@5.[151,1436,181,1284]" captionTargetPageId="5" captionText="FIGURE 4. Tiwaripotamon nganima n. sp. A–E, holotype male (59.3 × 43.6 mm) (ZRC2022.0810), Vietnam; F, G, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam. A, left G1 (ventral view); B, left G1 (dorsal view); C, distal part of left G1 (ventral view); D, distal part of left G1 (dorsal view); E, left G2; F, female anterior thoracic sternites and pleon; G, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.12681717" httpUri="https://zenodo.org/record/12681717/files/figure.png" pageId="6" pageNumber="304">Fig. 4F</figureCitation>
). The vulva is large, ovate and opens obliquely inwards, occupying most of sternite 6, gently pressing onto the suture with sternite 5 (
<figureCitation id="13309F8DFFA0FF85FFE065C5FAA710D0" box="[1226,1317,1750,1776]" captionStart="FIGURE 4" captionStartId="5.[151,250,1308,1332]" captionTargetBox="[151,1436,181,1284]" captionTargetId="figure-25@5.[151,1436,181,1284]" captionTargetPageId="5" captionText="FIGURE 4. Tiwaripotamon nganima n. sp. A–E, holotype male (59.3 × 43.6 mm) (ZRC2022.0810), Vietnam; F, G, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam. A, left G1 (ventral view); B, left G1 (dorsal view); C, distal part of left G1 (ventral view); D, distal part of left G1 (dorsal view); E, left G2; F, female anterior thoracic sternites and pleon; G, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.12681717" httpUri="https://zenodo.org/record/12681717/files/figure.png" pageId="6" pageNumber="304">Fig. 4G</figureCitation>
).
</paragraph>
<paragraph id="8BB48308FFA0FF84FBED65E9FC721761" blockId="6.[151,1437,1641,2028]" lastBlockId="7.[151,1437,151,1581]" lastPageId="7" lastPageNumber="305" pageId="6" pageNumber="304">
The series of specimens of
<taxonomicName id="4C0BF88BFFA0FF85FAD565E8FDFF1134" authorityName="Ng" authorityYear="2024" box="[511,637,1787,1812]" class="Malacostraca" family="Potamidae" genus="Tiwaripotamon" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="304" phylum="Arthropoda" rank="species" species="nganima" status="n. sp.">
<emphasis id="B97F5F1AFFA0FF85FAD565E8FDFF1134" box="[511,637,1787,1812]" italics="true" pageId="6" pageNumber="304">T. nganima</emphasis>
</taxonomicName>
<emphasis id="B97F5F1AFFA0FF85F9AC65E9FD441134" bold="true" box="[646,710,1786,1812]" pageId="6" pageNumber="304">
<taxonomicNameLabel id="A24CE261FFA0FF85F9AC65E9FD441134" box="[646,710,1786,1812]" pageId="6" pageNumber="304" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
show that some characters are variable. The median lobe of the posterior margin of the epistome varies in strength, although it is always distinctly projecting beyond the lateral margin; being triangular in the
<typeStatus id="54B03DAAFFA0FF85F92C6451FDEA117C" box="[518,616,1858,1884]" pageId="6" pageNumber="304" type="holotype">holotype</typeStatus>
male (
<figureCitation id="13309F8DFFA0FF85F9966451FC91117C" box="[700,787,1858,1884]" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="6" pageNumber="304">Fig. 2F</figureCitation>
) or with the tip more rounded in others (
<figureCitation id="13309F8DFFA0FF85FFD66451FAD8117C" box="[1276,1370,1858,1884]" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="6" pageNumber="304">Fig. 2G</figureCitation>
). The frontal margin also varies somewhat, being more sinuous and the two lobes more obvious in the female and smaller specimens (
<figureCitation id="13309F8DFFA0FF85FA376499FE191184" box="[285,411,1930,1956]" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="6" pageNumber="304">Fig. 2B, C</figureCitation>
). In addition, large male specimens have a proportionately wider carapace which is more swollen with the dorsal surface more convex (
<figureCitation id="13309F8DFFA0FF85F98F64BDFD7C11E8" box="[677,766,1966,1992]" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="6" pageNumber="304">Fig. 2D</figureCitation>
) (versus surface relatively lower in smaller ones;
<figureCitation id="13309F8DFFA0FF85FE1D64BDFA0D11E8" box="[1335,1423,1966,1992]" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="6" pageNumber="304">Fig. 2E</figureCitation>
); the posterior margin of epistome has the median lobe more prominent, triangular in shape with a sharp tip (
<figureCitation id="13309F8DFFA0FF84FE1564C1FF2A1691" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" lastPageId="7" lastPageNumber="305" pageId="6" pageNumber="304">Fig. 2D, F</figureCitation>
) (versus lower and more rounded in smaller ones;
<figureCitation id="13309F8DFFA1FF84F9DF6384FCF01691" box="[757,882,151,177]" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="7" pageNumber="305">Fig. 2E, G</figureCitation>
); the anterior thoracic sternum, notably sternites 1–3 appear transversely narrower (
<figureCitation id="13309F8DFFA1FF84F90A63A8FDFA16F5" box="[544,632,187,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="7" pageNumber="305">Fig. 2H</figureCitation>
) (versus relatively wider in smaller ones;
<figureCitation id="13309F8DFFA1FF84FF7F63A8FB2616F5" box="[1109,1188,187,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="7" pageNumber="305">Fig. 2I</figureCitation>
); the surfaces of male thoracic sternites 1–4 are distinctly pitted (
<figureCitation id="13309F8DFFA1FF84F95A63CCFD4516D9" box="[624,711,223,249]" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="7" pageNumber="305">Fig. 2H</figureCitation>
) (versus distinctly smoother surface with only scattered pits;
<figureCitation id="13309F8DFFA1FF84FE5A63CCFF30173D" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="7" pageNumber="305">Fig. 2I</figureCitation>
); have a relatively more quadrate male pleon with the lateral margins less divergent (
<figureCitation id="13309F8DFFA1FF84FF436210FB4F173D" box="[1129,1229,259,285]" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="7" pageNumber="305">Figs. 2H</figureCitation>
,
<figureCitation id="13309F8DFFA1FF84FFF36210FB79173D" box="[1241,1275,259,285]" captionStart="FIGURE 3" captionStartId="4.[151,250,1926,1950]" captionTargetBox="[151,1436,417,1903]" captionTargetId="figure-120@4.[151,1436,417,1903]" captionTargetPageId="4" captionText="FIGURE 3. Tiwaripotamon nganima n. sp. A–F, holotype male (59.3 × 43.6 mm) (ZRC2022.0810), Vietnam; G, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A, chelae; B, left third maxilliped; C, right fourth ambulatory leg; D, male sternopleonal cavity; E, male posterior thoracic sternites and pleon; F, G, male pleon." figureDoi="http://doi.org/10.5281/zenodo.12681715" httpUri="https://zenodo.org/record/12681715/files/figure.png" pageId="7" pageNumber="305">3H</figureCitation>
) (versus more triangular with lateral margins more divergent in smaller ones,
<figureCitation id="13309F8DFFA1FF84F8706234FC331761" box="[858,945,295,321]" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="7" pageNumber="305">Figs. 2I</figureCitation>
,
<figureCitation id="13309F8DFFA1FF84F8976234FC611761" box="[957,995,295,321]" captionStart="FIGURE 3" captionStartId="4.[151,250,1926,1950]" captionTargetBox="[151,1436,417,1903]" captionTargetId="figure-120@4.[151,1436,417,1903]" captionTargetPageId="4" captionText="FIGURE 3. Tiwaripotamon nganima n. sp. A–F, holotype male (59.3 × 43.6 mm) (ZRC2022.0810), Vietnam; G, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A, chelae; B, left third maxilliped; C, right fourth ambulatory leg; D, male sternopleonal cavity; E, male posterior thoracic sternites and pleon; F, G, male pleon." figureDoi="http://doi.org/10.5281/zenodo.12681715" httpUri="https://zenodo.org/record/12681715/files/figure.png" pageId="7" pageNumber="305">3G</figureCitation>
).
</paragraph>
<paragraph id="8BB48308FFA1FF84FBED6258FC1B17A9" blockId="7.[151,1437,151,1581]" pageId="7" pageNumber="305">
The diagnostic G1, with the strongly curved subterminal segment and swollen basal part of the terminal segment is constant for large and small specimens (
<figureCitation id="13309F8DFFA1FF84F958627CFD7B17A9" box="[626,761,367,393]" captionStart="FIGURE 4" captionStartId="5.[151,250,1308,1332]" captionTargetBox="[151,1436,181,1284]" captionTargetId="figure-25@5.[151,1436,181,1284]" captionTargetPageId="5" captionText="FIGURE 4. Tiwaripotamon nganima n. sp. A–E, holotype male (59.3 × 43.6 mm) (ZRC2022.0810), Vietnam; F, G, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam. A, left G1 (ventral view); B, left G1 (dorsal view); C, distal part of left G1 (ventral view); D, distal part of left G1 (dorsal view); E, left G2; F, female anterior thoracic sternites and pleon; G, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.12681717" httpUri="https://zenodo.org/record/12681717/files/figure.png" pageId="7" pageNumber="305">Figs. 4A–D</figureCitation>
,
<figureCitation id="13309F8DFFA1FF84F82E627DFC0F17A9" box="[772,909,366,393]" captionStart="FIGURE 5" captionStartId="6.[151,250,1463,1487]" captionTargetBox="[160,1428,189,1432]" captionTargetId="figure-15@6.[151,1436,181,1440]" captionTargetPageId="6" captionText="FIGURE 5. Tiwaripotamon nganima n. sp. A–E, holotype male (59.3 × 43.6 mm) (ZRC2022.0810), Vietnam; F–H, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A, left G1 (ventral view); B, F, distal part of left G1 (ventral view); C, G, distal part of left G1 (dorsomesial view); D, H, distal part of left G1 (dorsal view); E, left G2. Scales: A, E = 1.0 mm; B–D, F–H = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681719" httpUri="https://zenodo.org/record/12681719/files/figure.png" pageId="7" pageNumber="305">5A–D, F–H</figureCitation>
).
</paragraph>
<paragraph id="8BB48308FFA1FF84FBED6281FE7A178D" blockId="7.[151,1437,151,1581]" box="[199,504,402,429]" pageId="7" pageNumber="305">
<emphasis id="B97F5F1AFFA1FF84FBED6281FEEF178C" bold="true" box="[199,365,402,428]" pageId="7" pageNumber="305">Colour in life.</emphasis>
Not known.
</paragraph>
</subSubSection>
<subSubSection id="C311D083FFA1FF84FBED62A4FDBB141D" pageId="7" pageNumber="305" type="etymology">
<paragraph id="8BB48308FFA1FF84FBED62A4FDBB141D" blockId="7.[151,1437,151,1581]" pageId="7" pageNumber="305">
<emphasis id="B97F5F1AFFA1FF84FBED62A4FECC17F1" bold="true" box="[199,334,439,465]" pageId="7" pageNumber="305">Etymology.</emphasis>
The name is derived from an arbitrary combination of the name Ngan with the Latin “
<emphasis id="B97F5F1AFFA1FF84FE6062A4FA1217F0" box="[1354,1424,439,464]" italics="true" pageId="7" pageNumber="305">anima</emphasis>
” for soul; alluding to the kind and generous person Ngan Kee was. She was a good student, a trusted friend and a wonderful colleague; The author will sorely miss her enduring optimism, calming influence and cheeky smile. The name is used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C311D083FFA1FF84FBED6154FC9A100D" pageId="7" pageNumber="305" type="discussion">
<paragraph id="8BB48308FFA1FF84FBED6154FAAB1585" blockId="7.[151,1437,151,1581]" pageId="7" pageNumber="305">
<emphasis id="B97F5F1AFFA1FF84FBED6154FEB91441" bold="true" box="[199,315,583,609]" pageId="7" pageNumber="305">Remarks.</emphasis>
The general carapace features and structure of the G1 of
<taxonomicName id="4C0BF88BFFA1FF84F88C6154FBA31440" authorityName="Ng" authorityYear="2024" box="[934,1057,583,608]" class="Malacostraca" family="Potamidae" genus="Tiwaripotamon" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="305" phylum="Arthropoda" rank="species" species="nganima" status="n. sp.">
<emphasis id="B97F5F1AFFA1FF84F88C6154FBA31440" box="[934,1057,583,608]" italics="true" pageId="7" pageNumber="305">T. nganima</emphasis>
</taxonomicName>
<emphasis id="B97F5F1AFFA1FF84FF0C6154FBE11441" bold="true" box="[1062,1123,583,609]" pageId="7" pageNumber="305">
<taxonomicNameLabel id="A24CE261FFA1FF84FF0C6154FBE11441" box="[1062,1123,583,609]" pageId="7" pageNumber="305" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
most closely resemble those of
<taxonomicName id="4C0BF88BFFA1FF84FB9C6178FE6E14A5" authority="(Balss, 1914)" baseAuthorityName="Balss" baseAuthorityYear="1914" box="[182,492,619,645]" class="Malacostraca" family="Potamidae" genus="Tiwaripotamon" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="305" phylum="Arthropoda" rank="species" species="annamense">
<emphasis id="B97F5F1AFFA1FF84FB9C6178FECD14A4" box="[182,335,619,644]" italics="true" pageId="7" pageNumber="305">T. annamense</emphasis>
(
<bibRefCitation id="EF9AFEF9FFA1FF84FA756178FE6614A5" author="Balss, H." box="[351,484,619,645]" pageId="7" pageNumber="305" pagination="101 - 410" refId="ref5941" refString="Balss, H. (1914) Potamonidenstudien. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 37, 101 - 410. https: // doi. org / 10.5962 / bhl. part. 1071" type="journal article" year="1914">Balss, 1914</bibRefCitation>
)
</taxonomicName>
(Phuc-Son,
<collectingCountry id="F31CC398FFA1FF84F95C6178FD4C14A5" box="[630,718,619,645]" name="Vietnam" pageId="7" pageNumber="305">Annam</collectingCountry>
). Compared to
<taxonomicName id="4C0BF88BFFA1FF84F8526178FB9014A4" baseAuthorityName="Balss" baseAuthorityYear="1914" box="[888,1042,619,644]" class="Malacostraca" family="Potamidae" genus="Tiwaripotamon" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="305" phylum="Arthropoda" rank="species" species="annamense">
<emphasis id="B97F5F1AFFA1FF84F8526178FB9014A4" box="[888,1042,619,644]" italics="true" pageId="7" pageNumber="305">T. annamense</emphasis>
</taxonomicName>
, in
<taxonomicName id="4C0BF88BFFA1FF84FF176178FB3914A4" authorityName="Ng" authorityYear="2024" box="[1085,1211,619,644]" class="Malacostraca" family="Potamidae" genus="Tiwaripotamon" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="305" phylum="Arthropoda" rank="species" species="nganima" status="n. sp.">
<emphasis id="B97F5F1AFFA1FF84FF176178FB3914A4" box="[1085,1211,619,644]" italics="true" pageId="7" pageNumber="305">T. nganima</emphasis>
</taxonomicName>
<emphasis id="B97F5F1AFFA1FF84FFE86178FA8314A5" bold="true" box="[1218,1281,619,645]" pageId="7" pageNumber="305">
<taxonomicNameLabel id="A24CE261FFA1FF84FFE86178FA8314A5" box="[1218,1281,619,645]" pageId="7" pageNumber="305" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, the posterior margin of the epistome has the median lobe clearly defined, projecting anteriorly from the lateral parts, forming sinuous margins (
<figureCitation id="13309F8DFFA1FF84FA7061A0FE4814ED" box="[346,458,691,717]" captionStart="FIGURE 2" captionStartId="3.[151,250,1868,1892]" captionTargetBox="[151,1435,309,1845]" captionTargetId="figure-79@3.[151,1435,309,1845]" captionTargetPageId="3" captionText="FIGURE 2. Tiwaripotamon nganima n. sp.A, D, F, H, holotype male (59.3 × 43.6 mm) (ZRC 2022.0810), Vietnam; B, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam; C, E, G, I, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A–C, dorsal view of carapace; D, E, frontal view of cephalothorax; F, G, epistome; H, I, buccal cavity, anterior posterior thoracic sternites and pleon." figureDoi="http://doi.org/10.5281/zenodo.12681711" httpUri="https://zenodo.org/record/12681711/files/figure.png" pageId="7" pageNumber="305">Fig. 2F, G</figureCitation>
) (versus median lobe low, broadly triangular, with the margin gently contiguous with the lateral parts forming concave margins in
<taxonomicName id="4C0BF88BFFA1FF84F94B61C4FD7814D0" baseAuthorityName="Balss" baseAuthorityYear="1914" box="[609,762,727,752]" class="Malacostraca" family="Potamidae" genus="Tiwaripotamon" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="305" phylum="Arthropoda" rank="species" species="annamense">
<emphasis id="B97F5F1AFFA1FF84F94B61C4FD7814D0" box="[609,762,727,752]" italics="true" pageId="7" pageNumber="305">T. annamense</emphasis>
</taxonomicName>
;
<bibRefCitation id="EF9AFEF9FFA1FF84F82361C4FC3E14D1" author="Ng, P. K. L. &amp; Yeo, D. C. J." box="[777,956,727,753]" pageId="7" pageNumber="305" pagination="275 - 287" refId="ref6723" refString="Ng, P. K. L. &amp; Yeo, D. C. J. (2001) A revision of the genus Tiwaripotamon Bott, 1970 (Decapoda: Brachyura: Potamidae), with a description of a new species. Journal of Crustacean Biology, 21, 275 - 287. https: // doi. org / 10.1163 / 20021975 - 99990123" type="journal article" year="2001">Ng &amp; Yeo 2001</bibRefCitation>
: fig. 1B); the male pleon is proportionately more quadrate, with somites 3 and 4 relatively narrower (
<figureCitation id="13309F8DFFA1FF84F83261E8FC091535" box="[792,907,763,789]" captionStart="FIGURE 3" captionStartId="4.[151,250,1926,1950]" captionTargetBox="[151,1436,417,1903]" captionTargetId="figure-120@4.[151,1436,417,1903]" captionTargetPageId="4" captionText="FIGURE 3. Tiwaripotamon nganima n. sp. A–F, holotype male (59.3 × 43.6 mm) (ZRC2022.0810), Vietnam; G, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A, chelae; B, left third maxilliped; C, right fourth ambulatory leg; D, male sternopleonal cavity; E, male posterior thoracic sternites and pleon; F, G, male pleon." figureDoi="http://doi.org/10.5281/zenodo.12681715" httpUri="https://zenodo.org/record/12681715/files/figure.png" pageId="7" pageNumber="305">Fig. 3F, G</figureCitation>
) (versus pleon wider, especially across somites 3 and
<quantity id="4CF32EEDFFA1FF84FBF4600CFE891519" box="[222,267,799,825]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="7" pageNumber="305" unit="in" value="4.0">4 in</quantity>
<taxonomicName id="4C0BF88BFFA1FF84FA39600CFE2C1518" baseAuthorityName="Balss" baseAuthorityYear="1914" box="[275,430,799,824]" class="Malacostraca" family="Potamidae" genus="Tiwaripotamon" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="305" phylum="Arthropoda" rank="species" species="annamense">
<emphasis id="B97F5F1AFFA1FF84FA39600CFE2C1518" box="[275,430,799,824]" italics="true" pageId="7" pageNumber="305">T. annamense</emphasis>
</taxonomicName>
;
<bibRefCitation id="EF9AFEF9FFA1FF84FA94600CFDF41519" author="Ng, P. K. L. &amp; Yeo, D. C. J." box="[446,630,799,825]" pageId="7" pageNumber="305" pagination="275 - 287" refId="ref6723" refString="Ng, P. K. L. &amp; Yeo, D. C. J. (2001) A revision of the genus Tiwaripotamon Bott, 1970 (Decapoda: Brachyura: Potamidae), with a description of a new species. Journal of Crustacean Biology, 21, 275 - 287. https: // doi. org / 10.1163 / 20021975 - 99990123" type="journal article" year="2001">Ng &amp; Yeo 2001</bibRefCitation>
: fig. 2B); the G1 has the median part of the subterminal segment more strongly curved (
<figureCitation id="13309F8DFFA1FF84FA7E6050FD84157D" box="[340,518,835,861]" captionStart="FIGURE 4" captionStartId="5.[151,250,1308,1332]" captionTargetBox="[151,1436,181,1284]" captionTargetId="figure-25@5.[151,1436,181,1284]" captionTargetPageId="5" captionText="FIGURE 4. Tiwaripotamon nganima n. sp. A–E, holotype male (59.3 × 43.6 mm) (ZRC2022.0810), Vietnam; F, G, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam. A, left G1 (ventral view); B, left G1 (dorsal view); C, distal part of left G1 (ventral view); D, distal part of left G1 (dorsal view); E, left G2; F, female anterior thoracic sternites and pleon; G, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.12681717" httpUri="https://zenodo.org/record/12681717/files/figure.png" pageId="7" pageNumber="305">Figs. 4A, B, 4A</figureCitation>
) (versus gently curved in
<taxonomicName id="4C0BF88BFFA1FF84F80D6050FC42157C" baseAuthorityName="Balss" baseAuthorityYear="1914" box="[807,960,835,860]" class="Malacostraca" family="Potamidae" genus="Tiwaripotamon" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="305" phylum="Arthropoda" rank="species" species="annamense">
<emphasis id="B97F5F1AFFA1FF84F80D6050FC42157C" box="[807,960,835,860]" italics="true" pageId="7" pageNumber="305">T. annamense</emphasis>
</taxonomicName>
;
<bibRefCitation id="EF9AFEF9FFA1FF84F8E46050FB02157D" author="Ng, P. K. L. &amp; Yeo, D. C. J." box="[974,1152,835,861]" pageId="7" pageNumber="305" pagination="275 - 287" refId="ref6723" refString="Ng, P. K. L. &amp; Yeo, D. C. J. (2001) A revision of the genus Tiwaripotamon Bott, 1970 (Decapoda: Brachyura: Potamidae), with a description of a new species. Journal of Crustacean Biology, 21, 275 - 287. https: // doi. org / 10.1163 / 20021975 - 99990123" type="journal article" year="2001">Ng &amp; Yeo 2001</bibRefCitation>
: fig. 2F, H); and the outer surface of the basal part of the terminal segment is prominently swollen, even in smaller specimens (
<figureCitation id="13309F8DFFA1FF84FE3B6074FA1A15A1" box="[1297,1432,871,897]" captionStart="FIGURE 4" captionStartId="5.[151,250,1308,1332]" captionTargetBox="[151,1436,181,1284]" captionTargetId="figure-25@5.[151,1436,181,1284]" captionTargetPageId="5" captionText="FIGURE 4. Tiwaripotamon nganima n. sp. A–E, holotype male (59.3 × 43.6 mm) (ZRC2022.0810), Vietnam; F, G, paratype female (53.1 × 39.9 mm) (ZRC 2022.0811), Vietnam. A, left G1 (ventral view); B, left G1 (dorsal view); C, distal part of left G1 (ventral view); D, distal part of left G1 (dorsal view); E, left G2; F, female anterior thoracic sternites and pleon; G, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.12681717" httpUri="https://zenodo.org/record/12681717/files/figure.png" pageId="7" pageNumber="305">Figs. 4C, D</figureCitation>
,
<figureCitation id="13309F8DFFA1FF84FBBD6099FE9F1585" box="[151,285,906,933]" captionStart="FIGURE 5" captionStartId="6.[151,250,1463,1487]" captionTargetBox="[160,1428,189,1432]" captionTargetId="figure-15@6.[151,1436,181,1440]" captionTargetPageId="6" captionText="FIGURE 5. Tiwaripotamon nganima n. sp. A–E, holotype male (59.3 × 43.6 mm) (ZRC2022.0810), Vietnam; F–H, paratype male (41.5 × 31.3 mm) (ZRC 2022.0811), Vietnam. A, left G1 (ventral view); B, F, distal part of left G1 (ventral view); C, G, distal part of left G1 (dorsomesial view); D, H, distal part of left G1 (dorsal view); E, left G2. Scales: A, E = 1.0 mm; B–D, F–H = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12681719" httpUri="https://zenodo.org/record/12681719/files/figure.png" pageId="7" pageNumber="305">5A–D, F–H</figureCitation>
) (versus surface smooth, without any swelling in
<taxonomicName id="4C0BF88BFFA1FF84F8616098FC661584" baseAuthorityName="Balss" baseAuthorityYear="1914" box="[843,996,907,932]" class="Malacostraca" family="Potamidae" genus="Tiwaripotamon" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="305" phylum="Arthropoda" rank="species" species="annamense">
<emphasis id="B97F5F1AFFA1FF84F8616098FC661584" box="[843,996,907,932]" italics="true" pageId="7" pageNumber="305">T. annamense</emphasis>
</taxonomicName>
;
<bibRefCitation id="EF9AFEF9FFA1FF84F8D96098FB251585" author="Ng, P. K. L. &amp; Yeo, D. C. J." box="[1011,1191,907,933]" pageId="7" pageNumber="305" pagination="275 - 287" refId="ref6723" refString="Ng, P. K. L. &amp; Yeo, D. C. J. (2001) A revision of the genus Tiwaripotamon Bott, 1970 (Decapoda: Brachyura: Potamidae), with a description of a new species. Journal of Crustacean Biology, 21, 275 - 287. https: // doi. org / 10.1163 / 20021975 - 99990123" type="journal article" year="2001">Ng &amp; Yeo 2001</bibRefCitation>
: fig. 2D–I).
</paragraph>
<paragraph id="8BB48308FFA1FF84FBED60BCFC9A100D" blockId="7.[151,1437,151,1581]" pageId="7" pageNumber="305">
The distributions of
<taxonomicName id="4C0BF88BFFA1FF84FA8060BCFDC615E8" baseAuthorityName="Balss" baseAuthorityYear="1914" box="[426,580,943,968]" class="Malacostraca" family="Potamidae" genus="Tiwaripotamon" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="305" phylum="Arthropoda" rank="species" species="annamense">
<emphasis id="B97F5F1AFFA1FF84FA8060BCFDC615E8" box="[426,580,943,968]" italics="true" pageId="7" pageNumber="305">T. annamense</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0BF88BFFA1FF84F95360BCFD7515E8" authorityName="Ng" authorityYear="2024" box="[633,759,943,968]" class="Malacostraca" family="Potamidae" genus="Tiwaripotamon" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="305" phylum="Arthropoda" rank="species" species="nganima" status="n. sp.">
<emphasis id="B97F5F1AFFA1FF84F95360BCFD7515E8" box="[633,759,943,968]" italics="true" pageId="7" pageNumber="305">T. nganima</emphasis>
</taxonomicName>
<emphasis id="B97F5F1AFFA1FF84F9D760BCFCBE15E9" bold="true" box="[765,828,943,969]" pageId="7" pageNumber="305">
<taxonomicNameLabel id="A24CE261FFA1FF84F9D760BCFCBE15E9" box="[765,828,943,969]" pageId="7" pageNumber="305" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
are also
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apart.
<bibRefCitation id="EF9AFEF9FFA1FF84FF6B60BCFA8015E9" author="Ng, P. K. L. &amp; Yeo, D. C. J." box="[1089,1282,943,969]" pageId="7" pageNumber="305" pagination="275 - 287" refId="ref6723" refString="Ng, P. K. L. &amp; Yeo, D. C. J. (2001) A revision of the genus Tiwaripotamon Bott, 1970 (Decapoda: Brachyura: Potamidae), with a description of a new species. Journal of Crustacean Biology, 21, 275 - 287. https: // doi. org / 10.1163 / 20021975 - 99990123" type="journal article" year="2001">Ng &amp; Yeo (2001)</bibRefCitation>
re-examined, redescribed and figured the types of
<taxonomicName id="4C0BF88BFFA1FF84F91060C0FD5715CC" baseAuthorityName="Balss" baseAuthorityYear="1914" box="[570,725,979,1004]" class="Malacostraca" family="Potamidae" genus="Tiwaripotamon" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="305" phylum="Arthropoda" rank="species" species="annamense">
<emphasis id="B97F5F1AFFA1FF84F91060C0FD5715CC" box="[570,725,979,1004]" italics="true" pageId="7" pageNumber="305">T. annamense</emphasis>
</taxonomicName>
and selected a
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male for the species. They noted that the type locality of the species was “Phuc-Son,
<collectingCountry id="F31CC398FFA1FF84F99260E4FC881231" box="[696,778,1015,1041]" name="Vietnam" pageId="7" pageNumber="305">Annam</collectingCountry>
(now in northern
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)” (
<bibRefCitation id="EF9AFEF9FFA1FF84FF4B60E4FAD41231" author="Ng, P. K. L. &amp; Yeo, D. C. J." box="[1121,1366,1015,1041]" pageId="7" pageNumber="305" pagination="275 - 287" refId="ref6723" refString="Ng, P. K. L. &amp; Yeo, D. C. J. (2001) A revision of the genus Tiwaripotamon Bott, 1970 (Decapoda: Brachyura: Potamidae), with a description of a new species. Journal of Crustacean Biology, 21, 275 - 287. https: // doi. org / 10.1163 / 20021975 - 99990123" type="journal article" year="2001">Ng &amp; Yeo 2001: 279</bibRefCitation>
). This is not accurate as the type locality “Phuc-Son,
<collectingCountry id="F31CC398FFA1FF84F9926708FC8F1215" box="[696,781,1051,1077]" name="Vietnam" pageId="7" pageNumber="305">Annam</collectingCountry>
” is actually in central
<collectingCountry id="F31CC398FFA1FF84FF3D6708FBF81215" box="[1047,1146,1051,1077]" name="Vietnam" pageId="7" pageNumber="305">Vietnam</collectingCountry>
.
<bibRefCitation id="EF9AFEF9FFA1FF84FFAD6708FACC1215" author="Balss, H." box="[1159,1358,1051,1077]" pageId="7" pageNumber="305" pagination="101 - 410" refId="ref5941" refString="Balss, H. (1914) Potamonidenstudien. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 37, 101 - 410. https: // doi. org / 10.5962 / bhl. part. 1071" type="journal article" year="1914">Balss (1914: 406</bibRefCitation>
;
<bibRefCitation id="EF9AFEF9FFA1FF84FE766708FF401279" author="Balss, H." pageId="7" pageNumber="305" pagination="143 - 197" refId="ref5983" refString="Balss, H. (1937) Potamoniden (Dekapoda Brachyura) der Philippinen und des Malayischen Archipels. Internationale Revue der gesamten Hydrobiologie und Hydrographie, 34 (3 - 5), 143 - 197. https: // doi. org / 10.1002 / iroh. 19370340111" type="journal article" year="1937">1937: 169</bibRefCitation>
) described
<taxonomicName id="4C0BF88BFFA1FF84FA63672CFDF31278" authorityName="Balss" authorityYear="1914" box="[329,625,1087,1113]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="305" phylum="Arthropoda" rank="species" species="annamensis">
<emphasis id="B97F5F1AFFA1FF84FA63672CFDF31278" box="[329,625,1087,1113]" italics="true" pageId="7" pageNumber="305">Geothelphusa annamensis</emphasis>
</taxonomicName>
on the basis of many specimens collected by the well-known German collector, Hans Fruhstorfer, from Phuc-Son in
<collectingCountry id="F31CC398FFA1FF84F9B26770FD6C125D" box="[664,750,1123,1149]" name="Vietnam" pageId="7" pageNumber="305">Annam</collectingCountry>
. No date was given but
<bibRefCitation id="EF9AFEF9FFA1FF84F8DF6770FB38125D" author="Balss, H." box="[1013,1210,1123,1149]" pageId="7" pageNumber="305" pagination="143 - 197" refId="ref5983" refString="Balss, H. (1937) Potamoniden (Dekapoda Brachyura) der Philippinen und des Malayischen Archipels. Internationale Revue der gesamten Hydrobiologie und Hydrographie, 34 (3 - 5), 143 - 197. https: // doi. org / 10.1002 / iroh. 19370340111" type="journal article" year="1937">Balss (1937: 170)</bibRefCitation>
referred to an insect catalogue which documented Fruhstorfer’s field work, and this indicates that the material was collected between
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(see
<bibRefCitation id="EF9AFEF9FFA1FF84F91867B8FD7612E5" author="Fruhstorfer, H." box="[562,756,1194,1221]" pageId="7" pageNumber="305" pagination="1899 - 1905" refId="ref6622" refString="Fruhstorfer, H. (1905) Tagebuchblatter. Leipzig, Frankestein &amp; Wagner, (i) + 721 pp. [reprint of articles appearing in volumes 16 - 22 (1899 - 1905) of Insekten-Borse]." type="book chapter" year="1905">Fruhstorfer 1905</bibRefCitation>
). The name “Phuc-Son” is well known in the older literature on Vietnamese animals and plants. This location was in “
<collectingCountry id="F31CC398FFA1FF84F83567DCFCFA12C9" box="[799,888,1231,1257]" name="Vietnam" pageId="7" pageNumber="305">Annam</collectingCountry>
”, a French protectorate during colonial times, in what is today central
<collectingCountry id="F31CC398FFA1FF84FAA167E0FE6C132D" box="[395,494,1267,1293]" name="Vietnam" pageId="7" pageNumber="305">Vietnam</collectingCountry>
. “Phuc-Son” is today known as Phýớc Sõn (or Phuoc Son) District, in
<collectingRegion id="49CF4DEAFFA1FF84FE3967E0FF791311" country="Vietnam" name="Quang Nam" pageId="7" pageNumber="305">Quang Nam Province</collectingRegion>
in central
<collectingCountry id="F31CC398FFA1FF84FA5E6604FE551311" box="[372,471,1303,1329]" name="Vietnam" pageId="7" pageNumber="305">Vietnam</collectingCountry>
. This was where the Annamite mountain range is found (today known as Trýong Sõn Mountain, ca.
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<geoCoordinate id="EE3FE5CFFFA1FF84FAB36628FD851375" box="[409,519,1339,1365]" degrees="107.82" direction="east" orientation="longitude" pageId="7" pageNumber="305" precision="555" value="107.82">107.82°E</geoCoordinate>
). In describing a second species,
<taxonomicName id="4C0BF88BFFA1FF84F85B6628FBDE1375" authorityName="Balss" authorityYear="1914" box="[881,1116,1339,1365]" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="305" phylum="Arthropoda" rank="species" species="fruhstorferi">
<emphasis id="B97F5F1AFFA1FF84F85B6628FBDE1375" box="[881,1116,1339,1365]" italics="true" pageId="7" pageNumber="305">Potamon fruhstorferi</emphasis>
</taxonomicName>
(at present in
<taxonomicName id="4C0BF88BFFA1FF84FFDC6628FF561359" authority="Dang" authorityName="Dang" class="Malacostraca" family="Potamidae" genus="Balssipotamon" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="B97F5F1AFFA1FF84FFDC6628FA1E1375" box="[1270,1436,1339,1365]" italics="true" pageId="7" pageNumber="305">Balssipotamon</emphasis>
Đăng
</taxonomicName>
&amp; Hồ, 2008),
<bibRefCitation id="EF9AFEF9FFA1FF84FA55664CFDC81359" author="Balss, H." box="[383,586,1375,1401]" pageId="7" pageNumber="305" pagination="101 - 410" refId="ref5941" refString="Balss, H. (1914) Potamonidenstudien. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 37, 101 - 410. https: // doi. org / 10.5962 / bhl. part. 1071" type="journal article" year="1914">Balss (1914: 403)</bibRefCitation>
noted that it was collected from “
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, Phuc Son,
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westlich vom Hafen Touranne”. Touranne is the old name for what is today the city of
<collectingRegion id="49CF4DEAFFA1FF84F8F76690FBC413BD" box="[989,1094,1411,1437]" country="Vietnam" name="Da Nang" pageId="7" pageNumber="305">Da Nang</collectingRegion>
, a municipality at the edge of
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. Phýớc Sõn District is about
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southwest of
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. This location should not be confused with another two sites also named “Phúc Sõn” in northern
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; one in Tân Yên District,
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(ca.
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) and another in Anh Sõn District,
<collectingRegion id="49CF4DEAFFA1FF84F8AA66FCFBCD1029" box="[896,1103,1519,1545]" country="Vietnam" name="Nghe An" pageId="7" pageNumber="305">Nghe An Province</collectingRegion>
(ca.
<geoCoordinate id="EE3FE5CFFFA1FF84FFAF66FCFB611029" box="[1157,1251,1519,1545]" degrees="18.86" direction="north" orientation="latitude" pageId="7" pageNumber="305" precision="555" value="18.86">18.86°N</geoCoordinate>
<geoCoordinate id="EE3FE5CFFFA1FF84FFC066FDFAD51029" box="[1258,1367,1518,1545]" degrees="105.02" direction="east" orientation="longitude" pageId="7" pageNumber="305" precision="555" value="105.02">105.02°E</geoCoordinate>
), with both areas in what was called “
<collectingCountry id="F31CC398FFA1FF84FADB6500FDC7100D" box="[497,581,1555,1581]" name="Vietnam" pageId="7" pageNumber="305">Tonkin</collectingCountry>
” in colonial times.
</paragraph>
</subSubSection>
</treatment>
</document>
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