treatments-xml/data/03/A2/12/03A212449A6B3246FF42F41BFD42EC75.xml

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<mods:title id="EAC757DD5DB7D856D92FBF9775FFDDCD">Two new species of Leptarma (Decapoda, Brachyura, Sesarmidae) from the Western Indian Ocean with notes on their phylogeny</mods:title>
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<mods:namePart id="D0BF33988962D66644842B802F28F85C">Shahdadi, Adnan</mods:namePart>
<mods:affiliation id="5B15B7BF7AFCE786C6D0F575C10AC2A2">Department of Marine Biology, Faculty of Marine Sciences and Technology, University of Hormozgan, Bandar Abbas, Iran. adnan 1361 @ gmail. com; https: // orcid. org / 0000 - 0001 - 5131 - 7700</mods:affiliation>
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<mods:namePart id="7F94E588A3DDE54F4A3E3ECC134AE5A5">Schubart, Christoph D.</mods:namePart>
<mods:affiliation id="38508B5F8642FA3E3E901EB2767C4BE1">Zoology &amp; Evolution, University of Regensburg, 93040 Regensburg, Germany.</mods:affiliation>
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<heading id="D0FC143E9A6B324EFF42F41BFE37E901" bold="true" box="[151,500,1053,1079]" fontSize="11" level="1" pageId="6" pageNumber="445" reason="1">
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<taxonomicName id="4C0BD8D19A6B324EFF42F41BFE69E901" ID-CoL="84GM8" authority="Shahdadi &amp; Schubart, 2020" authorityName="Shahdadi &amp; Schubart" authorityYear="2020" box="[151,426,1053,1079]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="445" phylum="Arthropoda" rank="species" species="reunionense" status="sp. nov.">
<emphasis id="B97F7F409A6B324EFF42F41BFE69E901" bold="true" box="[151,426,1053,1079]" italics="true" pageId="6" pageNumber="445">Leptarma reunionense</emphasis>
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<taxonomicNameLabel id="A24CC23B9A6B324EFE64F41BFE37E901" box="[433,500,1053,1079]" pageId="6" pageNumber="445" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="8BB4A3529A6B324EFF42F447FE18E96D" blockId="6.[151,968,1053,1151]" box="[151,475,1089,1115]" pageId="6" pageNumber="445">
(
<figureCitation id="1330BFD79A6B324EFF4AF447FEC8E96D" box="[159,267,1089,1115]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStartId-0="7.[151,250,1980,2005]" captionStartId-1="8.[151,250,1769,1794]" captionStartId-2="9.[151,250,1508,1533]" captionTargetBox-0="[167,1419,611,1937]" captionTargetBox-1="[180,1408,181,1743]" captionTargetBox-2="[153,1434,181,1482]" captionTargetId-0="figure@7.[149,1437,597,1955]" captionTargetId-1="figure@8.[180,1408,181,1743]" captionTargetId-2="figure@9.[153,1434,181,1482]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionText-0="FIGURE 4. L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356). A, dorsal habitus; B, ventral habitus." captionText-1="FIGURE 5. L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356). A, frontal view; B, right G1, dorsal view (arrows pointing to broken part); C, right chela, outer view; D, chelar dactylar tubercles, outer view; E, right chela, dorsal view; F, chelar dactylar tubercles, dorsal view; G, pleon; H, pleonal somites 1–3." captionText-2="FIGURE 6. L. reunionense n. sp. paratype, male (11.6×9.8) (UF12354). A, dorsal habitus; B–E, right G1; B, dorsal view; C, apical corneous part, dorsal view (arrows pointing to aperture); D, apical corneous part, ventral view; E, front view of apical corneous part." figureDoi-0="http://doi.org/10.5281/zenodo.3954585" figureDoi-1="http://doi.org/10.5281/zenodo.3954587" figureDoi-2="http://doi.org/10.5281/zenodo.3954589" httpUri-0="https://zenodo.org/record/3954585/files/figure.png" httpUri-1="https://zenodo.org/record/3954587/files/figure.png" httpUri-2="https://zenodo.org/record/3954589/files/figure.png" pageId="6" pageNumber="445">
Figs. 4– 
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, 
<figureCitation id="1330BFD79A6B324EFEC2F447FEFBE96D" box="[279,312,1089,1115]" captionStart="FIGURE 7" captionStartId="10.[151,250,1866,1891]" captionTargetBox="[151,1431,190,1835]" captionTargetId="figure@10.[150,1436,181,1841]" captionTargetPageId="10" captionText="FIGURE 7. Comparisons of carapace and ambulatory legs, dorsal views. A, L. dispersum n. sp. holotype, male (12.2×10.1) (MNHN-IU-2012-660); B, L. lenzii, male (11.8×9.9) (ZRC 2018.1373); C, L. gazi, paratype, male (15.5×13.1) (MZUF 3671); D, L. leptosoma, male (19.0×17.5) (MZUF 4978); E, L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356); F, L. liho, holotype, male (13.8×12.5) (SMF 36266); G, L. moluccense, lectotype, male (14.6×13.6) (RMNH.CRUS.D 102590); H, L. paucitorum, holotype, male (19.9×18.8) (MZB.cru. 2243)." figureDoi="http://doi.org/10.5281/zenodo.3954593" httpUri="https://zenodo.org/record/3954593/files/figure.png" pageId="6" pageNumber="445">7E</figureCitation>
, 
<figureCitation id="1330BFD79A6B324EFE91F447FEABE96D" box="[324,360,1089,1115]" captionStart="FIGURE 8" captionStartId="11.[151,250,1705,1730]" captionTargetBox="[157,1430,489,1681]" captionTargetId="figure@11.[157,1430,489,1681]" captionTargetPageId="11" captionText="FIGURE 8. Comparisons of male chelar dactylus and tuberculation patterns, outer views. A, L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356); B, L. liho, holotype, male (13.8×12.5) (SMF 36266); C, L. jamelense, holotype, male (11.5×10.0) (USNM 45917); D, L. moluccense, lectotype, male (14.6×13.6) (RMNH.CRUS.D 102590); L. paucitorum, holotype, male (19.9×18.8) (MZB.cru. 2243)." figureDoi="http://doi.org/10.5281/zenodo.3954595" httpUri="https://zenodo.org/record/3954595/files/figure.png" pageId="6" pageNumber="445">8A</figureCitation>
, 
<figureCitation id="1330BFD79A6B324EFEA1F447FE5AE96D" box="[372,409,1089,1115]" captionStart="FIGURE 9" captionStartId="12.[151,250,1898,1923]" captionTargetBox="[408,1179,181,1873]" captionTargetId="figure@12.[408,1179,181,1873]" captionTargetPageId="12" captionText="FIGURE 9. Comparisons of male chelar dactylus and tuberculation patterns, dorsal views. A, L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356); B, L. liho, holotype, male (13.8×12.5) (SMF 36266); C, L. jamelense, holotype, male (11.5×10.0) (USNM 45917); D, L. moluccense, lectotype, male (14.6×13.6) (RMNH.CRUS.D 102590); L. paucitorum, holotype, male (19.9×18.8) (MZB.cru. 2243)." figureDoi="http://doi.org/10.5281/zenodo.3954597" httpUri="https://zenodo.org/record/3954597/files/figure.png" pageId="6" pageNumber="445">9A</figureCitation>
, 
<figureCitation id="1330BFD79A6B324EFE71F447FE10E96D" box="[420,467,1089,1115]" captionStart="FIGURE 10" captionStartId="13.[151,250,1672,1697]" captionTargetBox="[155,1432,181,1646]" captionTargetId="figure@13.[155,1432,181,1646]" captionTargetPageId="13" captionText="FIGURE 10. Comparisons of G1, apical part, dorsal views. A, L. dispersum n. sp. holotype, male (12.2×10.1) (MNHN-IU- 2012-660); B, L. lenzii, male (11.8×9.9) (ZRC 2018.1373); C, L. gazi, paratype, male (15.5×13.1) (MZUF 3671); D, L. leptosoma, male (19.0×17.5) (MZUF 4978); E, L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356); F, L. liho, holotype, male (13.8×12.5) (SMF 36266); G, L. jamelense, holotype, male (11.5×10.0) (USNM 45917); H, L. moluccense, lectotype, male (14.6×13.6) (RMNH.CRUS.D 102590); I, L. paucitorum, holotype, male (19.9×18.8) (MZB.cru. 2243)." figureDoi="http://doi.org/10.5281/zenodo.3954599" httpUri="https://zenodo.org/record/3954599/files/figure.png" pageId="6" pageNumber="445">10E</figureCitation>
)
</paragraph>
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urn:lsid:zoobank.org:act: 
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<paragraph id="8BB4A3529A6B324EFF42F4ACFB54E9F7" blockId="6.[151,1175,1192,1217]" box="[151,1175,1192,1217]" pageId="6" pageNumber="445">
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<taxonomicName id="4C0BD8D19A6B324EFF42F4ACFEBDE9F7" ID-CoL="4DBZJ" baseAuthorityName="Latreille" baseAuthorityYear="1803" box="[151,382,1193,1217]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="445" phylum="Arthropoda" rank="species" species="plicatum">
<emphasis id="B97F7F409A6B324EFF42F4ACFEBDE9F7" box="[151,382,1193,1217]" italics="true" pageId="6" pageNumber="445">Parasesarma plicatum</emphasis>
</taxonomicName>
— 
<treatmentCitation id="0AAA85439A6B324EFE42F4AEFD96E9F7" author="Poupin, J." box="[407,597,1192,1217]" page="132" pageId="6" pageNumber="445" year="2009">
<bibRefCitation id="EF9ADEA39A6B324EFE42F4AEFD96E9F7" author="Poupin, J." box="[407,597,1192,1217]" pageId="6" pageNumber="445" refId="ref8954" refString="Poupin, J. (2009) Crustaces de la Reunion, Decapodes et Stomatopodes. IRD Editions, Marseille, 140 pp." type="book" year="2009">Poupin (2009: 132</bibRefCitation>
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, Fig. p. 36); not 
<taxonomicName id="4C0BD8D19A6B324EFD2BF4ACFB50E9F7" ID-CoL="4DBZJ" authority="(Latreille, 1803)" baseAuthorityName="Latreille" baseAuthorityYear="1803" box="[766,1171,1192,1217]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="445" phylum="Arthropoda" rank="species" species="plicatum">
<emphasis id="B97F7F409A6B324EFD2BF4ACFC27E9F7" box="[766,996,1193,1217]" italics="true" pageId="6" pageNumber="445">Parasesarma plicatum</emphasis>
(Latreille, 1803)
</taxonomicName>
.
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</paragraph>
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<emphasis id="B97F7F409A6B324EFF42F4E9FEBFE83C" bold="true" box="[151,380,1263,1290]" pageId="6" pageNumber="445">Material examined</emphasis>
. 
<materialsCitation id="3B63A90F9A6B324EFE59F4E9FC00E83F" ID-GBIF-Occurrence="2816753303" box="[396,963,1263,1290]" collectionCode="UF" country="Réunion" location="Reunion" pageId="6" pageNumber="445" specimenCode="UF12356" specimenCount="1" typeStatus="holotype">
<typeStatus id="54B01DF09A6B324EFE59F4E9FE3AE83F" box="[396,505,1263,1289]" pageId="6" pageNumber="445" type="holotype">Holotype</typeStatus>
: male (11.8×9.92) (
<specimenCode id="DBAD0B299A6B324EFD0EF4E9FC89E83C" box="[731,842,1263,1290]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="6" pageNumber="445" type="Museum">UF12356</specimenCode>
), 
<collectingCountry id="F31CE3C29A6B324EFCB5F4E9FC00E83F" box="[864,963,1263,1289]" name="Réunion" pageId="6" pageNumber="445">Réunion</collectingCountry>
</materialsCitation>
. 
<materialsCitation id="3B63A90F9A6B324EFC1AF4F6FE86E81B" ID-GBIF-Occurrence="2816753301" collectionCode="UF" country="Reunion Island" location="Reunion Island" pageId="6" pageNumber="445" specimenCode="UF12354" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="54B01DF09A6B324EFC1AF4F6FBF6E83C" box="[975,1077,1264,1290]" pageId="6" pageNumber="445" type="paratype">Paratype</typeStatus>
: 
<specimenCount id="9D0D68DB9A6B324EFB97F4F6FB53E83F" box="[1090,1168,1263,1290]" pageId="6" pageNumber="445" type="male">1 male</specimenCount>
(11.6×9.8) (
<specimenCode id="DBAD0B299A6B324EFAF4F4E9FA53E83C" box="[1313,1424,1263,1290]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="6" pageNumber="445" type="Museum">UF12354</specimenCode>
), 
<collectingCountry id="F31CE3C29A6B324EFF42F515FE86E81B" box="[151,325,1299,1325]" name="Reunion Island" pageId="6" pageNumber="445">Réunion Island</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C311F0D99A6B324EFF12F531FD24EB7B" pageId="6" pageNumber="445" type="diagnosis">
<paragraph id="8BB4A3529A6B324EFF12F531FD24EB7B" blockId="6.[151,1436,1263,2010]" pageId="6" pageNumber="445">
<emphasis id="B97F7F409A6B324EFF12F531FEF9E864" bold="true" box="[199,314,1335,1362]" pageId="6" pageNumber="445">Diagnosis</emphasis>
. Carapace subrectangular, slightly broader than long, front deflexed, almost straight in dorsal view, median post-frontal lobes wider than lateral ones, lateral carapace margin slightly concave. Eye with short and stout eyestalk, wider than long, corneal part slightly wider. Male chela large, robust, upper surface of palm with two oblique pectinated crests, dorsal surface of dactylus bearing 9 or 10 low, rounded, asymmetric tubercles, with longer distal slope, proximal ones positioned at middle of dorsal face of dactylus with oblique keel. Ambulatory legs relatively long, third pair longest, total length (basis-dactylus) ca. 1.8 times carapace width; male pleon triangular, somite 2 medially longer than lateral edges. G1 stout, straight; apical corneous process long, tip truncate, bent at an angle of about 65° to vertical axis, aperture terminal.
</paragraph>
</subSubSection>
<subSubSection id="C311F0D99A6B3241FF12F651FD5BEAC3" lastPageId="9" lastPageNumber="448" pageId="6" pageNumber="445" type="description">
<paragraph id="8BB4A3529A6B324EFF12F651FC9EEA7C" blockId="6.[151,1436,1263,2010]" pageId="6" pageNumber="445">
<emphasis id="B97F7F409A6B324EFF12F651FE92EB44" bold="true" box="[199,337,1623,1650]" pageId="6" pageNumber="445">Description</emphasis>
. Carapace subrectangular, slightly broader than long (cw/cl = 1.17), greatest width between exorbital angles; carapace surface smooth; front ca. 0.55 times carapace width, deflexed, almost straight in dorsal view; post-frontal lobes prominent, median lobes wider than lateral ones, separated by deep furrow; dorsal carapace regions well indicated, gastric region demarcated, cardiac region separated from intestinal region, lateral branchial ridges prominent; upper orbital border smooth, lower orbital border finely granulate; anterolateral margin with sharp exorbital angle, lateral margin slightly concave, edged with row of short setae. Eye with eyestalk short, stout, wider than long, corneal part slightly wider (
<figureCitation id="1330BFD79A6B324EFD5EF729FD2CEA7C" box="[651,751,1839,1866]" captionStart="FIGURE 4" captionStartId="7.[151,250,1980,2005]" captionTargetBox="[167,1419,611,1937]" captionTargetId="figure@7.[149,1437,597,1955]" captionTargetPageId="7" captionText="FIGURE 4. L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356). A, dorsal habitus; B, ventral habitus." figureDoi="http://doi.org/10.5281/zenodo.3954585" httpUri="https://zenodo.org/record/3954585/files/figure.png" pageId="6" pageNumber="445">Figs. 4A</figureCitation>
, 
<figureCitation id="1330BFD79A6B324EFD2FF729FCDCEA7C" box="[762,799,1839,1866]" captionStart="FIGURE 5" captionStartId="8.[151,250,1769,1794]" captionTargetBox="[180,1408,181,1743]" captionTargetId="figure@8.[180,1408,181,1743]" captionTargetPageId="8" captionText="FIGURE 5. L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356). A, frontal view; B, right G1, dorsal view (arrows pointing to broken part); C, right chela, outer view; D, chelar dactylar tubercles, outer view; E, right chela, dorsal view; F, chelar dactylar tubercles, dorsal view; G, pleon; H, pleonal somites 1–3." figureDoi="http://doi.org/10.5281/zenodo.3954587" httpUri="https://zenodo.org/record/3954587/files/figure.png" pageId="6" pageNumber="445">5A</figureCitation>
, 
<figureCitation id="1330BFD79A6B324EFCFEF729FC93EA7C" box="[811,848,1839,1866]" captionStart="FIGURE 6" captionStartId="9.[151,250,1508,1533]" captionTargetBox="[153,1434,181,1482]" captionTargetId="figure@9.[153,1434,181,1482]" captionTargetPageId="9" captionText="FIGURE 6. L. reunionense n. sp. paratype, male (11.6×9.8) (UF12354). A, dorsal habitus; B–E, right G1; B, dorsal view; C, apical corneous part, dorsal view (arrows pointing to aperture); D, apical corneous part, ventral view; E, front view of apical corneous part." figureDoi="http://doi.org/10.5281/zenodo.3954589" httpUri="https://zenodo.org/record/3954589/files/figure.png" pageId="6" pageNumber="445">6A</figureCitation>
).
</paragraph>
<paragraph id="8BB4A3529A6B324FFF12F755FE62EC99" blockId="6.[151,1436,1263,2010]" lastBlockId="7.[151,1437,153,576]" lastPageId="7" lastPageNumber="446" pageId="6" pageNumber="445">
Male chela large (palm length/cw ca. 0.68) (
<figureCitation id="1330BFD79A6B324EFD6DF755FCD9EA58" box="[696,794,1875,1902]" captionStart="FIGURE 4" captionStartId="7.[151,250,1980,2005]" captionTargetBox="[167,1419,611,1937]" captionTargetId="figure@7.[149,1437,597,1955]" captionTargetPageId="7" captionText="FIGURE 4. L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356). A, dorsal habitus; B, ventral habitus." figureDoi="http://doi.org/10.5281/zenodo.3954585" httpUri="https://zenodo.org/record/3954585/files/figure.png" pageId="6" pageNumber="445">Figs. 4B</figureCitation>
, 
<figureCitation id="1330BFD79A6B324EFCF2F755FC88EA58" box="[807,843,1875,1902]" captionStart="FIGURE 5" captionStartId="8.[151,250,1769,1794]" captionTargetBox="[180,1408,181,1743]" captionTargetId="figure@8.[180,1408,181,1743]" captionTargetPageId="8" captionText="FIGURE 5. L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356). A, frontal view; B, right G1, dorsal view (arrows pointing to broken part); C, right chela, outer view; D, chelar dactylar tubercles, outer view; E, right chela, dorsal view; F, chelar dactylar tubercles, dorsal view; G, pleon; H, pleonal somites 1–3." figureDoi="http://doi.org/10.5281/zenodo.3954587" httpUri="https://zenodo.org/record/3954587/files/figure.png" pageId="6" pageNumber="445">5A</figureCitation>
), robust (palm width/length ca. 0.61) (
<figureCitation id="1330BFD79A6B324EFB29F755FA9DEA58" box="[1276,1374,1875,1902]" captionStart="FIGURE 4" captionStartId="7.[151,250,1980,2005]" captionTargetBox="[167,1419,611,1937]" captionTargetId="figure@7.[149,1437,597,1955]" captionTargetPageId="7" captionText="FIGURE 4. L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356). A, dorsal habitus; B, ventral habitus." figureDoi="http://doi.org/10.5281/zenodo.3954585" httpUri="https://zenodo.org/record/3954585/files/figure.png" pageId="6" pageNumber="445">Figs. 4B</figureCitation>
, 
<figureCitation id="1330BFD79A6B324EFABFF755FA4DEA58" box="[1386,1422,1875,1902]" captionStart="FIGURE 5" captionStartId="8.[151,250,1769,1794]" captionTargetBox="[180,1408,181,1743]" captionTargetId="figure@8.[180,1408,181,1743]" captionTargetPageId="8" captionText="FIGURE 5. L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356). A, frontal view; B, right G1, dorsal view (arrows pointing to broken part); C, right chela, outer view; D, chelar dactylar tubercles, outer view; E, right chela, dorsal view; F, chelar dactylar tubercles, dorsal view; G, pleon; H, pleonal somites 1–3." figureDoi="http://doi.org/10.5281/zenodo.3954587" httpUri="https://zenodo.org/record/3954587/files/figure.png" pageId="6" pageNumber="445">5C</figureCitation>
); merus with granulate dorsal border, no distinct subdistal spine, ventral border granulate, anterior border granulate, with distinct subdistal spine, accompanied by several spines distally, inner face smooth with longitudinal row of setae, anterior face smooth, outer face with rows of fine granules; upper surface of palm with 2 oblique pectinated crests, distal (primary) crest composed of 13-16 teeth; secondary crest well developed, with 6-8 teeth, both crests terminating at inner end in 1-2 granule(s) and several granules on outer side (
<figureCitation id="1330BFD79A6A324FFBD1F0BBFB98EDE1" box="[1028,1115,188,215]" captionStart="FIGURE 5" captionStartId="8.[151,250,1769,1794]" captionTargetBox="[180,1408,181,1743]" captionTargetId="figure@8.[180,1408,181,1743]" captionTargetPageId="8" captionText="FIGURE 5. L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356). A, frontal view; B, right G1, dorsal view (arrows pointing to broken part); C, right chela, outer view; D, chelar dactylar tubercles, outer view; E, right chela, dorsal view; F, chelar dactylar tubercles, dorsal view; G, pleon; H, pleonal somites 1–3." figureDoi="http://doi.org/10.5281/zenodo.3954587" httpUri="https://zenodo.org/record/3954587/files/figure.png" pageId="7" pageNumber="446">Fig. 5E</figureCitation>
); outer surface of palm with fine granules (rows of fine granules on the proximal half) except for smooth, punctate fixed finger (
<figureCitation id="1330BFD79A6A324FFAD8F0E7FAA5EDCD" box="[1293,1382,224,251]" captionStart="FIGURE 5" captionStartId="8.[151,250,1769,1794]" captionTargetBox="[180,1408,181,1743]" captionTargetId="figure@8.[180,1408,181,1743]" captionTargetPageId="8" captionText="FIGURE 5. L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356). A, frontal view; B, right G1, dorsal view (arrows pointing to broken part); C, right chela, outer view; D, chelar dactylar tubercles, outer view; E, right chela, dorsal view; F, chelar dactylar tubercles, dorsal view; G, pleon; H, pleonal somites 1–3." figureDoi="http://doi.org/10.5281/zenodo.3954587" httpUri="https://zenodo.org/record/3954587/files/figure.png" pageId="7" pageNumber="446">Fig. 5C</figureCitation>
); inner surface of palm granular except area facing carpus and distal half of finger; no vertical ridge, dactylus slightly curved downward and inward, dorsal surface bearing 9 or 10 low but distinct, rounded tubercles, distinct to tip, tubercles asymmetric with distal slope longer distal slope, proximal ones positioned at middle of dorsal face of dactylus with oblique keel; small pointed granules on inner side, cutting edge of both fingers with a series of variably sized teeth (
<figureCitation id="1330BFD79A6A324FFEC8F193FE9CEC99" box="[285,351,404,431]" captionStart="FIGURE 5" captionStartId="8.[151,250,1769,1794]" captionTargetBox="[180,1408,181,1743]" captionTargetId="figure@8.[180,1408,181,1743]" captionTargetPageId="8" captionText="FIGURE 5. L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356). A, frontal view; B, right G1, dorsal view (arrows pointing to broken part); C, right chela, outer view; D, chelar dactylar tubercles, outer view; E, right chela, dorsal view; F, chelar dactylar tubercles, dorsal view; G, pleon; H, pleonal somites 1–3." figureDoi="http://doi.org/10.5281/zenodo.3954587" httpUri="https://zenodo.org/record/3954587/files/figure.png" pageId="7" pageNumber="446">Fig. 5</figureCitation>
C–F).
</paragraph>
<paragraph id="8BB4A3529A6A324FFF12F1BFFE4FEF76" blockId="7.[151,1437,153,576]" pageId="7" pageNumber="446">
Ambulatory legs flattened dorso-ventrally, relatively long, third pair longest, total length (basis-dactylus) ca. 1.8 times carapace width, tufts of long setae scattered along prpodi and dactyli, merus with anterior margin crenulated, ca. 2.26 times as long as wide, propodus ca. 2.8 times as long as wide, dactylus ca. 0.8 times length of propodus (
<figureCitation id="1330BFD79A6A324FFF3CF223FE8EEF09" box="[233,333,549,575]" captionStart="FIGURE 4" captionStartId="7.[151,250,1980,2005]" captionTargetBox="[167,1419,611,1937]" captionTargetId="figure@7.[149,1437,597,1955]" captionTargetPageId="7" captionText="FIGURE 4. L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356). A, dorsal habitus; B, ventral habitus." figureDoi="http://doi.org/10.5281/zenodo.3954585" httpUri="https://zenodo.org/record/3954585/files/figure.png" pageId="7" pageNumber="446">Figs. 4A</figureCitation>
, 
<figureCitation id="1330BFD79A6A324FFE8CF223FEBCEF76" box="[345,383,549,576]" captionStart="FIGURE 6" captionStartId="9.[151,250,1508,1533]" captionTargetBox="[153,1434,181,1482]" captionTargetId="figure@9.[153,1434,181,1482]" captionTargetPageId="9" captionText="FIGURE 6. L. reunionense n. sp. paratype, male (11.6×9.8) (UF12354). A, dorsal habitus; B–E, right G1; B, dorsal view; C, apical corneous part, dorsal view (arrows pointing to aperture); D, apical corneous part, ventral view; E, front view of apical corneous part." figureDoi="http://doi.org/10.5281/zenodo.3954589" httpUri="https://zenodo.org/record/3954589/files/figure.png" pageId="7" pageNumber="446">6A</figureCitation>
).
</paragraph>
<caption id="DF74F3DA9A6A324FFF42F7BAFB3CEAE3" ID-DOI="http://doi.org/10.5281/zenodo.3954585" ID-Zenodo-Dep="3954585" box="[151,1279,1980,2005]" httpUri="https://zenodo.org/record/3954585/files/figure.png" pageId="7" pageNumber="446" startId="7.[151,250,1980,2005]" targetBox="[167,1419,611,1937]" targetPageId="7">
<paragraph id="8BB4A3529A6A324FFF42F7BAFB3CEAE3" blockId="7.[151,1279,1980,2005]" box="[151,1279,1980,2005]" pageId="7" pageNumber="446">
<emphasis id="B97F7F409A6A324FFF42F7BAFED0EAE3" bold="true" box="[151,275,1980,2005]" pageId="7" pageNumber="446">FIGURE 4.</emphasis>
<taxonomicName id="4C0BD8D19A6A324FFECCF7B8FE6CEAE3" authority="Shahdadi &amp; Schubart, 2020" authorityName="Shahdadi &amp; Schubart" authorityYear="2020" box="[281,431,1982,2005]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="446" phylum="Arthropoda" rank="species" species="reunionense" status="sp. nov.">
<emphasis id="B97F7F409A6A324FFECCF7B8FE6CEAE3" box="[281,431,1982,2005]" italics="true" pageId="7" pageNumber="446">L. reunionense</emphasis>
</taxonomicName>
<emphasis id="B97F7F409A6A324FFE60F7BBFE2CEAE3" bold="true" box="[437,495,1981,2005]" pageId="7" pageNumber="446">
<taxonomicNameLabel id="A24CC23B9A6A324FFE60F7BBFE2CEAE3" box="[437,495,1981,2005]" pageId="7" pageNumber="446" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
holotype, male (11.8×9.92) (UF12356). A, dorsal habitus; B, ventral habitus.
</paragraph>
</caption>
<caption id="DF74F3DA9A653240FF42F6EFFCECEA7C" ID-DOI="http://doi.org/10.5281/zenodo.3954587" ID-Zenodo-Dep="3954587" httpUri="https://zenodo.org/record/3954587/files/figure.png" pageId="8" pageNumber="447" startId="8.[151,250,1769,1794]" targetBox="[180,1408,181,1743]" targetPageId="8">
<paragraph id="8BB4A3529A653240FF42F6EFFCECEA7C" blockId="8.[151,1436,1769,1866]" pageId="8" pageNumber="447">
<emphasis id="B97F7F409A653240FF42F6EFFED6EA34" bold="true" box="[151,277,1769,1794]" pageId="8" pageNumber="447">FIGURE 5.</emphasis>
<taxonomicName id="4C0BD8D19A653240FEC8F6ECFE76EA37" authority="Shahdadi &amp; Schubart, 2020" authorityName="Shahdadi &amp; Schubart" authorityYear="2020" box="[285,437,1770,1793]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="447" phylum="Arthropoda" rank="species" species="reunionense" status="sp. nov.">
<emphasis id="B97F7F409A653240FEC8F6ECFE76EA37" box="[285,437,1770,1793]" italics="true" pageId="8" pageNumber="447">L. reunionense</emphasis>
</taxonomicName>
<emphasis id="B97F7F409A653240FE68F6ECFE3BEA34" bold="true" box="[445,504,1770,1794]" pageId="8" pageNumber="447">
<taxonomicNameLabel id="A24CC23B9A653240FE68F6ECFE3BEA34" box="[445,504,1770,1794]" pageId="8" pageNumber="447" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
holotype, male (11.8×9.92) (UF12356). A, frontal view; B, right G1, dorsal view (arrows pointing to broken part); C, right chela, outer view; D, chelar dactylar tubercles, outer view; E, right chela, dorsal view; F, chelar dactylar tubercles, dorsal view; G, pleon; H, pleonal somites 1–3.
</paragraph>
</caption>
<caption id="DF74F3DA9A643241FF42F5E2FEEBEB72" ID-DOI="http://doi.org/10.5281/zenodo.3954589" ID-Zenodo-Dep="3954589" httpUri="https://zenodo.org/record/3954589/files/figure.png" pageId="9" pageNumber="448" startId="9.[151,250,1508,1533]" targetBox="[153,1434,181,1482]" targetPageId="9">
<paragraph id="8BB4A3529A643241FF42F5E2FEEBEB72" blockId="9.[151,1436,1508,1604]" pageId="9" pageNumber="448">
<emphasis id="B97F7F409A643241FF42F5E2FED7E8CB" bold="true" box="[151,276,1508,1533]" pageId="9" pageNumber="448">FIGURE 6.</emphasis>
<taxonomicName id="4C0BD8D19A643241FECEF5E0FE71E8CB" authority="Shahdadi &amp; Schubart, 2020" authorityName="Shahdadi &amp; Schubart" authorityYear="2020" box="[283,434,1510,1533]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="448" phylum="Arthropoda" rank="species" species="reunionense" status="sp. nov.">
<emphasis id="B97F7F409A643241FECEF5E0FE71E8CB" box="[283,434,1510,1533]" italics="true" pageId="9" pageNumber="448">L. reunionense</emphasis>
</taxonomicName>
<emphasis id="B97F7F409A643241FE6CF5E3FE30E8CB" bold="true" box="[441,499,1509,1533]" pageId="9" pageNumber="448">
<taxonomicNameLabel id="A24CC23B9A643241FE6CF5E3FE30E8CB" box="[441,499,1509,1533]" pageId="9" pageNumber="448" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
paratype, male (11.6×9.8) (UF12354). A, dorsal habitus; B–E, right G1; B, dorsal view; C, apical corneous part, dorsal view (arrows pointing to aperture); D, apical corneous part, ventral view; E, front view of apical corneous part.
</paragraph>
</caption>
<paragraph id="8BB4A3529A643241FF12F691FE59EBCF" blockId="9.[151,1436,1686,2038]" pageId="9" pageNumber="448">
Male pleon triangular (
<figureCitation id="1330BFD79A643241FE1EF691FDEDEB87" box="[459,558,1687,1713]" captionStart="FIGURE 4" captionStartId="7.[151,250,1980,2005]" captionTargetBox="[167,1419,611,1937]" captionTargetId="figure@7.[149,1437,597,1955]" captionTargetPageId="7" captionText="FIGURE 4. L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356). A, dorsal habitus; B, ventral habitus." figureDoi="http://doi.org/10.5281/zenodo.3954585" httpUri="https://zenodo.org/record/3954585/files/figure.png" pageId="9" pageNumber="448">Figs. 4B</figureCitation>
, 
<figureCitation id="1330BFD79A643241FDEEF690FDA3EB87" box="[571,608,1686,1713]" captionStart="FIGURE 5" captionStartId="8.[151,250,1769,1794]" captionTargetBox="[180,1408,181,1743]" captionTargetId="figure@8.[180,1408,181,1743]" captionTargetPageId="8" captionText="FIGURE 5. L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356). A, frontal view; B, right G1, dorsal view (arrows pointing to broken part); C, right chela, outer view; D, chelar dactylar tubercles, outer view; E, right chela, dorsal view; F, chelar dactylar tubercles, dorsal view; G, pleon; H, pleonal somites 1–3." figureDoi="http://doi.org/10.5281/zenodo.3954587" httpUri="https://zenodo.org/record/3954587/files/figure.png" pageId="9" pageNumber="448">5G</figureCitation>
), with telson almost as long as basal width, as long as somite 6; somite 6 longer than others; somites 4 and 5 trapezoidal, somite 3 widest, laterally convex, somite 2 medially longer than lateral edges (
<figureCitation id="1330BFD79A643241FEE6F6D9FE4DEBCF" box="[307,398,1758,1785]" captionStart="FIGURE 5" captionStartId="8.[151,250,1769,1794]" captionTargetBox="[180,1408,181,1743]" captionTargetId="figure@8.[180,1408,181,1743]" captionTargetPageId="8" captionText="FIGURE 5. L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356). A, frontal view; B, right G1, dorsal view (arrows pointing to broken part); C, right chela, outer view; D, chelar dactylar tubercles, outer view; E, right chela, dorsal view; F, chelar dactylar tubercles, dorsal view; G, pleon; H, pleonal somites 1–3." figureDoi="http://doi.org/10.5281/zenodo.3954587" httpUri="https://zenodo.org/record/3954587/files/figure.png" pageId="9" pageNumber="448">Fig. 5H</figureCitation>
).
</paragraph>
<paragraph id="8BB4A3529A643241FF12F705FCAFEA74" blockId="9.[151,1436,1686,2038]" pageId="9" pageNumber="448">
G1 stout, rather long, straight; apical corneous process long, tip truncate, bent at angle of about 65° to vertical axis, arched in cross section, aperture terminal (
<figureCitation id="1330BFD79A643241FD7BF721FCD3EA77" box="[686,784,1830,1857]" captionStart="FIGURE 5" captionStartId="8.[151,250,1769,1794]" captionTargetBox="[180,1408,181,1743]" captionTargetId="figure@8.[180,1408,181,1743]" captionTargetPageId="8" captionText="FIGURE 5. L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356). A, frontal view; B, right G1, dorsal view (arrows pointing to broken part); C, right chela, outer view; D, chelar dactylar tubercles, outer view; E, right chela, dorsal view; F, chelar dactylar tubercles, dorsal view; G, pleon; H, pleonal somites 1–3." figureDoi="http://doi.org/10.5281/zenodo.3954587" httpUri="https://zenodo.org/record/3954587/files/figure.png" pageId="9" pageNumber="448">Figs. 5B</figureCitation>
, 
<figureCitation id="1330BFD79A643241FCC9F721FCE9EA74" box="[796,810,1831,1858]" captionStart="FIGURE 6" captionStartId="9.[151,250,1508,1533]" captionTargetBox="[153,1434,181,1482]" captionTargetId="figure@9.[153,1434,181,1482]" captionTargetPageId="9" captionText="FIGURE 6. L. reunionense n. sp. paratype, male (11.6×9.8) (UF12354). A, dorsal habitus; B–E, right G1; B, dorsal view; C, apical corneous part, dorsal view (arrows pointing to aperture); D, apical corneous part, ventral view; E, front view of apical corneous part." figureDoi="http://doi.org/10.5281/zenodo.3954589" httpUri="https://zenodo.org/record/3954589/files/figure.png" pageId="9" pageNumber="448">6</figureCitation>
B–E).
</paragraph>
<paragraph id="8BB4A3529A643241FF12F74DFD02EA53" blockId="9.[151,1436,1686,2038]" box="[199,705,1867,1894]" pageId="9" pageNumber="448">
<emphasis id="B97F7F409A643241FF12F74DFEDDEA53" bold="true" box="[199,286,1867,1893]" pageId="9" pageNumber="448">Female</emphasis>
. No female specimen was examined.
</paragraph>
<paragraph id="8BB4A3529A643241FF12F768FA8EEA9B" blockId="9.[151,1436,1686,2038]" pageId="9" pageNumber="448">
<emphasis id="B97F7F409A643241FF12F768FEC8EABF" bold="true" box="[199,267,1902,1929]" pageId="9" pageNumber="448">Color</emphasis>
. The specimens preserved in ethanol do not show any coloration. According to the photo in 
<bibRefCitation id="EF9ADEA39A643241FA98F769FF18EA9B" author="Poupin, J." pageId="9" pageNumber="448" refId="ref8954" refString="Poupin, J. (2009) Crustaces de la Reunion, Decapodes et Stomatopodes. IRD Editions, Marseille, 140 pp." type="book" year="2009">Poupin (2009</bibRefCitation>
, p. 36), the carapace and pereiopods are light brown with black clouds, chela brighter, fingers yellow.
</paragraph>
<paragraph id="8BB4A3529A643241FF12F7B0FD5BEAC3" blockId="9.[151,1436,1686,2038]" pageId="9" pageNumber="448">
<emphasis id="B97F7F409A643241FF12F7B0FE2FEAE7" bold="true" box="[199,492,1974,2001]" pageId="9" pageNumber="448">Habitat and distribution</emphasis>
. No information to be found on label. According to 
<bibRefCitation id="EF9ADEA39A643241FBE1F7B1FAC3EAE7" author="Poupin, J." box="[1076,1280,1974,2001]" pageId="9" pageNumber="448" refId="ref8954" refString="Poupin, J. (2009) Crustaces de la Reunion, Decapodes et Stomatopodes. IRD Editions, Marseille, 140 pp." type="book" year="2009">Poupin (2009: 35)</bibRefCitation>
the crab lives in mangrove and muddy swamps of estuaries.
</paragraph>
</subSubSection>
<caption id="DF74F3DA9A673242FF42F74CFD2AEAC5" ID-DOI="http://doi.org/10.5281/zenodo.3954593" ID-Zenodo-Dep="3954593" httpUri="https://zenodo.org/record/3954593/files/figure.png" pageId="10" pageNumber="449" startId="10.[151,250,1866,1891]" targetBox="[151,1431,190,1835]" targetPageId="10">
<paragraph id="8BB4A3529A673242FF42F74CFD2AEAC5" blockId="10.[151,1437,1866,2035]" pageId="10" pageNumber="449">
<emphasis id="B97F7F409A673242FF42F74CFED6EA55" bold="true" box="[151,277,1866,1891]" pageId="10" pageNumber="449">FIGURE 7.</emphasis>
Comparisons of carapace and ambulatory legs, dorsal views. A, 
<taxonomicName id="4C0BD8D19A673242FC60F74AFBF9EA55" authority="Shahdadi &amp; Schubart, 2020" authorityName="Shahdadi &amp; Schubart" authorityYear="2020" box="[949,1082,1867,1891]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="449" phylum="Arthropoda" rank="species" species="dispersum" status="sp. nov.">
<emphasis id="B97F7F409A673242FC60F74AFBF9EA55" box="[949,1082,1867,1891]" italics="true" pageId="10" pageNumber="449">L. dispersum</emphasis>
</taxonomicName>
<emphasis id="B97F7F409A673242FB97F74DFBBEEA55" bold="true" box="[1090,1149,1867,1891]" pageId="10" pageNumber="449">
<taxonomicNameLabel id="A24CC23B9A673242FB97F74DFBBEEA55" box="[1090,1149,1867,1891]" pageId="10" pageNumber="449" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
holotype, male (12.2×10.1) (MNHN-IU-2012-660); B, 
<taxonomicName id="4C0BD8D19A673242FE7BF776FDC3EAB1" baseAuthorityName="De Man" baseAuthorityYear="1895" box="[430,512,1903,1927]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="449" phylum="Arthropoda" rank="species" species="lenzii">
<emphasis id="B97F7F409A673242FE7BF776FDC3EAB1" box="[430,512,1903,1927]" italics="true" pageId="10" pageNumber="449">L. lenzii</emphasis>
</taxonomicName>
, male (11.8×9.9) (ZRC 2018.1373); C, 
<taxonomicName id="4C0BD8D19A673242FC42F776FC1FEAB1" box="[919,988,1904,1927]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="449" phylum="Arthropoda" rank="species" species="gazi">
<emphasis id="B97F7F409A673242FC42F776FC1FEAB1" box="[919,988,1904,1927]" italics="true" pageId="10" pageNumber="449">L. gazi</emphasis>
</taxonomicName>
, paratype, male (15.5×13.1) (MZUF 3671); D, 
<taxonomicName id="4C0BD8D19A673242FF62F792FEF8EA9D" baseAuthorityName="Hilgendorf" baseAuthorityYear="1869" box="[183,315,1939,1963]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="449" phylum="Arthropoda" rank="species" species="leptosoma">
<emphasis id="B97F7F409A673242FF62F792FEF8EA9D" box="[183,315,1939,1963]" italics="true" pageId="10" pageNumber="449">L. leptosoma</emphasis>
</taxonomicName>
, male (19.0×17.5) (MZUF 4978); E, 
<taxonomicName id="4C0BD8D19A673242FD68F792FC97EA9D" authority="Shahdadi &amp; Schubart, 2020" authorityName="Shahdadi &amp; Schubart" authorityYear="2020" box="[701,852,1940,1963]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="449" phylum="Arthropoda" rank="species" species="reunionense" status="sp. nov.">
<emphasis id="B97F7F409A673242FD68F792FC97EA9D" box="[701,852,1940,1963]" italics="true" pageId="10" pageNumber="449">L. reunionense</emphasis>
</taxonomicName>
<emphasis id="B97F7F409A673242FC8EF795FC55EA9D" bold="true" box="[859,918,1939,1963]" pageId="10" pageNumber="449">
<taxonomicNameLabel id="A24CC23B9A673242FC8EF795FC55EA9D" box="[859,918,1939,1963]" pageId="10" pageNumber="449" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
holotype, male (11.8×9.92) (UF12356); F, 
<taxonomicName id="4C0BD8D19A673242FA86F792FA56EA9D" baseAuthorityName="Koller, Liu &amp; Schubart" baseAuthorityYear="2010" box="[1363,1429,1939,1963]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="449" phylum="Arthropoda" rank="species" species="liho">
<emphasis id="B97F7F409A673242FA86F792FA56EA9D" box="[1363,1429,1939,1963]" italics="true" pageId="10" pageNumber="449">L. liho</emphasis>
</taxonomicName>
, holotype, male (13.8×12.5) (SMF 36266); G, 
<taxonomicName id="4C0BD8D19A673242FDADF7BEFCCFEAF9" baseAuthorityName="De Man" baseAuthorityYear="1892" box="[632,780,1975,1999]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="449" phylum="Arthropoda" rank="species" species="moluccense">
<emphasis id="B97F7F409A673242FDADF7BEFCCFEAF9" box="[632,780,1975,1999]" italics="true" pageId="10" pageNumber="449">L. moluccense</emphasis>
</taxonomicName>
, lectotype, male (14.6×13.6) (RMNH.CRUS.D 102590); H, 
<taxonomicName id="4C0BD8D19A673242FA5DF7BEFECFEAC5" baseAuthorityName="Rahayu &amp; Ng" baseAuthorityYear="2009" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="449" phylum="Arthropoda" rank="species" species="paucitorum">
<emphasis id="B97F7F409A673242FA5DF7BEFECFEAC5" italics="true" pageId="10" pageNumber="449">L. paucitorum</emphasis>
</taxonomicName>
, holotype, male (19.9×18.8) (MZB.cru. 2243).
</paragraph>
</caption>
<subSubSection id="C311F0D99A663243FF12F09FFA50ED85" box="[199,1427,153,180]" pageId="11" pageNumber="450" type="etymology">
<paragraph id="8BB4A3529A663243FF12F09FFA50ED85" blockId="11.[151,1437,153,467]" box="[199,1427,153,180]" pageId="11" pageNumber="450">
<emphasis id="B97F7F409A663243FF12F09FFE8BED85" bold="true" box="[199,328,153,179]" pageId="11" pageNumber="450">Etymology</emphasis>
. The name is derived from the region of occurrence (
<typeStatus id="54B01DF09A663243FC4BF09FFC13ED85" box="[926,976,153,179]" pageId="11" pageNumber="450">type</typeStatus>
locality) of the species, 
<collectingCountry id="F31CE3C29A663243FB34F09FFA4CED85" box="[1249,1423,153,179]" name="Reunion Island" pageId="11" pageNumber="450">Réunion Island</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C311F0D99A663246FF12F0BBFD42EC75" lastPageId="14" lastPageNumber="453" pageId="11" pageNumber="450" type="discussion">
<paragraph id="8BB4A3529A663243FF12F0BBFC44ECE5" blockId="11.[151,1437,153,467]" pageId="11" pageNumber="450">
<emphasis id="B97F7F409A663243FF12F0BBFEF7EDE1" bold="true" box="[199,308,189,215]" pageId="11" pageNumber="450">Remarks</emphasis>
. Among other 
<taxonomicName id="4C0BD8D19A663243FE08F0B8FD89EDE1" authorityName="Shahdadi, Fratini &amp; Schubart" authorityYear="2020" box="[477,586,190,215]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="450" phylum="Arthropoda" rank="genus">
<emphasis id="B97F7F409A663243FE08F0B8FD89EDE1" box="[477,586,190,215]" italics="true" pageId="11" pageNumber="450">Leptarma</emphasis>
</taxonomicName>
, the new species resembles most 
<taxonomicName id="4C0BD8D19A663243FC13F0B8FBD3EDE1" baseAuthorityName="Koller, Liu &amp; Schubart" baseAuthorityYear="2010" box="[966,1040,189,215]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="450" phylum="Arthropoda" rank="species" species="liho">
<emphasis id="B97F7F409A663243FC13F0B8FBD3EDE1" box="[966,1040,189,215]" italics="true" pageId="11" pageNumber="450">L. liho</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0BD8D19A663243FB92F0B8FB16EDE1" baseAuthorityName="Rathbun" baseAuthorityYear="1914" box="[1095,1237,189,215]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="450" phylum="Arthropoda" rank="species" species="jamelense">
<emphasis id="B97F7F409A663243FB92F0B8FB16EDE1" box="[1095,1237,189,215]" italics="true" pageId="11" pageNumber="450">L. jamelense</emphasis>
</taxonomicName>
, both from South 
<collectingCountry id="F31CE3C29A663243FF42F0E7FEC7EDCD" box="[151,260,225,251]" name="China" pageId="11" pageNumber="450">East Asia</collectingCountry>
, in general appearance (
<figureCitation id="1330BFD79A663243FDCDF0E7FD4EEDCD" box="[536,653,225,251]" captionStart="FIGURE 7" captionStartId="10.[151,250,1866,1891]" captionTargetBox="[151,1431,190,1835]" captionTargetId="figure@10.[150,1436,181,1841]" captionTargetPageId="10" captionText="FIGURE 7. Comparisons of carapace and ambulatory legs, dorsal views. A, L. dispersum n. sp. holotype, male (12.2×10.1) (MNHN-IU-2012-660); B, L. lenzii, male (11.8×9.9) (ZRC 2018.1373); C, L. gazi, paratype, male (15.5×13.1) (MZUF 3671); D, L. leptosoma, male (19.0×17.5) (MZUF 4978); E, L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356); F, L. liho, holotype, male (13.8×12.5) (SMF 36266); G, L. moluccense, lectotype, male (14.6×13.6) (RMNH.CRUS.D 102590); H, L. paucitorum, holotype, male (19.9×18.8) (MZB.cru. 2243)." figureDoi="http://doi.org/10.5281/zenodo.3954593" httpUri="https://zenodo.org/record/3954593/files/figure.png" pageId="11" pageNumber="450">Fig. 7E, F</figureCitation>
), chelar (
<figureCitation id="1330BFD79A663243FD2DF0E7FC86EDCD" box="[760,837,225,251]" captionStart="FIGURE 8" captionStartId="11.[151,250,1705,1730]" captionTargetBox="[157,1430,489,1681]" captionTargetId="figure@11.[157,1430,489,1681]" captionTargetPageId="11" captionText="FIGURE 8. Comparisons of male chelar dactylus and tuberculation patterns, outer views. A, L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356); B, L. liho, holotype, male (13.8×12.5) (SMF 36266); C, L. jamelense, holotype, male (11.5×10.0) (USNM 45917); D, L. moluccense, lectotype, male (14.6×13.6) (RMNH.CRUS.D 102590); L. paucitorum, holotype, male (19.9×18.8) (MZB.cru. 2243)." figureDoi="http://doi.org/10.5281/zenodo.3954595" httpUri="https://zenodo.org/record/3954595/files/figure.png" pageId="11" pageNumber="450">Figs. 8</figureCitation>
A–C, 9A–C) and G1 morphology (
<figureCitation id="1330BFD79A663243FB01F0E7FAE7EDCD" box="[1236,1316,225,251]" captionStart="FIGURE 10" captionStartId="13.[151,250,1672,1697]" captionTargetBox="[155,1432,181,1646]" captionTargetId="figure@13.[155,1432,181,1646]" captionTargetPageId="13" captionText="FIGURE 10. Comparisons of G1, apical part, dorsal views. A, L. dispersum n. sp. holotype, male (12.2×10.1) (MNHN-IU- 2012-660); B, L. lenzii, male (11.8×9.9) (ZRC 2018.1373); C, L. gazi, paratype, male (15.5×13.1) (MZUF 3671); D, L. leptosoma, male (19.0×17.5) (MZUF 4978); E, L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356); F, L. liho, holotype, male (13.8×12.5) (SMF 36266); G, L. jamelense, holotype, male (11.5×10.0) (USNM 45917); H, L. moluccense, lectotype, male (14.6×13.6) (RMNH.CRUS.D 102590); I, L. paucitorum, holotype, male (19.9×18.8) (MZB.cru. 2243)." figureDoi="http://doi.org/10.5281/zenodo.3954599" httpUri="https://zenodo.org/record/3954599/files/figure.png" pageId="11" pageNumber="450">Fig. 10</figureCitation>
E–G). The new species, however, has a wider carapace compared to the other two species, i.e. cw/cl ca. 
<quantity id="4CF30EB79A663243FB73F103FB25EC29" box="[1190,1254,261,287]" metricMagnitude="-2" metricUnit="m" metricValue="3.048" pageId="11" pageNumber="450" unit="in" value="1.2">1.2 in</quantity>
the new species vs. 
<quantity id="4CF30EB79A663243FF6AF12FFEC2EC75" box="[191,257,297,323]" metricMagnitude="-2" metricUnit="m" metricValue="2.794" pageId="11" pageNumber="450" unit="in" value="1.1">1.1 in</quantity>
<taxonomicName id="4C0BD8D19A663243FEDCF12CFE97EC75" baseAuthorityName="Koller, Liu &amp; Schubart" baseAuthorityYear="2010" box="[265,340,297,323]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="450" phylum="Arthropoda" rank="species" species="liho">
<emphasis id="B97F7F409A663243FEDCF12CFE97EC75" box="[265,340,297,323]" italics="true" pageId="11" pageNumber="450">L. liho</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0BD8D19A663243FE59F12CFDD8EC75" baseAuthorityName="Rathbun" baseAuthorityYear="1914" box="[396,539,297,323]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="450" phylum="Arthropoda" rank="species" species="jamelense">
<emphasis id="B97F7F409A663243FE59F12CFDD8EC75" box="[396,539,297,323]" italics="true" pageId="11" pageNumber="450">L. jamelense</emphasis>
</taxonomicName>
. In dorsal view, the front is more sinuous in 
<taxonomicName id="4C0BD8D19A663243FBC8F12CFBABEC75" baseAuthorityName="Koller, Liu &amp; Schubart" baseAuthorityYear="2010" box="[1053,1128,297,323]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="450" phylum="Arthropoda" rank="species" species="liho">
<emphasis id="B97F7F409A663243FBC8F12CFBABEC75" box="[1053,1128,297,323]" italics="true" pageId="11" pageNumber="450">L. liho</emphasis>
</taxonomicName>
(
<figureCitation id="1330BFD79A663243FBADF12FFB0FEC75" box="[1144,1228,297,323]" captionStart="FIGURE 7" captionStartId="10.[151,250,1866,1891]" captionTargetBox="[151,1431,190,1835]" captionTargetId="figure@10.[150,1436,181,1841]" captionTargetPageId="10" captionText="FIGURE 7. Comparisons of carapace and ambulatory legs, dorsal views. A, L. dispersum n. sp. holotype, male (12.2×10.1) (MNHN-IU-2012-660); B, L. lenzii, male (11.8×9.9) (ZRC 2018.1373); C, L. gazi, paratype, male (15.5×13.1) (MZUF 3671); D, L. leptosoma, male (19.0×17.5) (MZUF 4978); E, L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356); F, L. liho, holotype, male (13.8×12.5) (SMF 36266); G, L. moluccense, lectotype, male (14.6×13.6) (RMNH.CRUS.D 102590); H, L. paucitorum, holotype, male (19.9×18.8) (MZB.cru. 2243)." figureDoi="http://doi.org/10.5281/zenodo.3954593" httpUri="https://zenodo.org/record/3954593/files/figure.png" pageId="11" pageNumber="450">Fig. 7F</figureCitation>
) and 
<taxonomicName id="4C0BD8D19A663243FAD8F12CFA5FEC75" baseAuthorityName="Rathbun" baseAuthorityYear="1914" box="[1293,1436,297,323]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="450" phylum="Arthropoda" rank="species" species="jamelense">
<emphasis id="B97F7F409A663243FAD8F12CFA5FEC75" box="[1293,1436,297,323]" italics="true" pageId="11" pageNumber="450">L. jamelense</emphasis>
</taxonomicName>
compared to the almost straight front of the new species (
<figureCitation id="1330BFD79A663243FCF5F14BFCACEC5E" box="[800,879,333,360]" captionStart="FIGURE 7" captionStartId="10.[151,250,1866,1891]" captionTargetBox="[151,1431,190,1835]" captionTargetId="figure@10.[150,1436,181,1841]" captionTargetPageId="10" captionText="FIGURE 7. Comparisons of carapace and ambulatory legs, dorsal views. A, L. dispersum n. sp. holotype, male (12.2×10.1) (MNHN-IU-2012-660); B, L. lenzii, male (11.8×9.9) (ZRC 2018.1373); C, L. gazi, paratype, male (15.5×13.1) (MZUF 3671); D, L. leptosoma, male (19.0×17.5) (MZUF 4978); E, L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356); F, L. liho, holotype, male (13.8×12.5) (SMF 36266); G, L. moluccense, lectotype, male (14.6×13.6) (RMNH.CRUS.D 102590); H, L. paucitorum, holotype, male (19.9×18.8) (MZB.cru. 2243)." figureDoi="http://doi.org/10.5281/zenodo.3954593" httpUri="https://zenodo.org/record/3954593/files/figure.png" pageId="11" pageNumber="450">Fig.7E</figureCitation>
). The new species also differs from the two other species in chelar dactylar tubercles (9-10 rounded tubercles in the new species (
<figureCitation id="1330BFD79A663243FBF6F177FBBFECBD" box="[1059,1148,369,395]" captionStart="FIGURE 9" captionStartId="12.[151,250,1898,1923]" captionTargetBox="[408,1179,181,1873]" captionTargetId="figure@12.[408,1179,181,1873]" captionTargetPageId="12" captionText="FIGURE 9. Comparisons of male chelar dactylus and tuberculation patterns, dorsal views. A, L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356); B, L. liho, holotype, male (13.8×12.5) (SMF 36266); C, L. jamelense, holotype, male (11.5×10.0) (USNM 45917); D, L. moluccense, lectotype, male (14.6×13.6) (RMNH.CRUS.D 102590); L. paucitorum, holotype, male (19.9×18.8) (MZB.cru. 2243)." figureDoi="http://doi.org/10.5281/zenodo.3954597" httpUri="https://zenodo.org/record/3954597/files/figure.png" pageId="11" pageNumber="450">Fig. 9A</figureCitation>
) vs. 12-14 oval tubercles in 
<taxonomicName id="4C0BD8D19A663243FF61F190FF3DEC99" baseAuthorityName="Koller, Liu &amp; Schubart" baseAuthorityYear="2010" box="[180,254,405,431]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="450" phylum="Arthropoda" rank="species" species="liho">
<emphasis id="B97F7F409A663243FF61F190FF3DEC99" box="[180,254,405,431]" italics="true" pageId="11" pageNumber="450">L. liho</emphasis>
</taxonomicName>
(
<figureCitation id="1330BFD79A663243FED8F193FEA0EC99" box="[269,355,405,431]" captionStart="FIGURE 9" captionStartId="12.[151,250,1898,1923]" captionTargetBox="[408,1179,181,1873]" captionTargetId="figure@12.[408,1179,181,1873]" captionTargetPageId="12" captionText="FIGURE 9. Comparisons of male chelar dactylus and tuberculation patterns, dorsal views. A, L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356); B, L. liho, holotype, male (13.8×12.5) (SMF 36266); C, L. jamelense, holotype, male (11.5×10.0) (USNM 45917); D, L. moluccense, lectotype, male (14.6×13.6) (RMNH.CRUS.D 102590); L. paucitorum, holotype, male (19.9×18.8) (MZB.cru. 2243)." figureDoi="http://doi.org/10.5281/zenodo.3954597" httpUri="https://zenodo.org/record/3954597/files/figure.png" pageId="11" pageNumber="450">Fig. 9B</figureCitation>
) and 
<taxonomicName id="4C0BD8D19A663243FE77F190FDF3EC99" baseAuthorityName="Rathbun" baseAuthorityYear="1914" box="[418,560,405,431]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="450" phylum="Arthropoda" rank="species" species="jamelense">
<emphasis id="B97F7F409A663243FE77F190FDF3EC99" box="[418,560,405,431]" italics="true" pageId="11" pageNumber="450">L. jamelense</emphasis>
</taxonomicName>
(
<figureCitation id="1330BFD79A663243FDEAF193FD55EC99" box="[575,662,405,431]" captionStart="FIGURE 9" captionStartId="12.[151,250,1898,1923]" captionTargetBox="[408,1179,181,1873]" captionTargetId="figure@12.[408,1179,181,1873]" captionTargetPageId="12" captionText="FIGURE 9. Comparisons of male chelar dactylus and tuberculation patterns, dorsal views. A, L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356); B, L. liho, holotype, male (13.8×12.5) (SMF 36266); C, L. jamelense, holotype, male (11.5×10.0) (USNM 45917); D, L. moluccense, lectotype, male (14.6×13.6) (RMNH.CRUS.D 102590); L. paucitorum, holotype, male (19.9×18.8) (MZB.cru. 2243)." figureDoi="http://doi.org/10.5281/zenodo.3954597" httpUri="https://zenodo.org/record/3954597/files/figure.png" pageId="11" pageNumber="450">Fig. 9C</figureCitation>
)). The new species also has G1s with slightly longer apical corneous processes, compared to the other two similar species (
<figureCitation id="1330BFD79A663243FD27F1BFFC81ECE5" box="[754,834,441,467]" captionStart="FIGURE 10" captionStartId="13.[151,250,1672,1697]" captionTargetBox="[155,1432,181,1646]" captionTargetId="figure@13.[155,1432,181,1646]" captionTargetPageId="13" captionText="FIGURE 10. Comparisons of G1, apical part, dorsal views. A, L. dispersum n. sp. holotype, male (12.2×10.1) (MNHN-IU- 2012-660); B, L. lenzii, male (11.8×9.9) (ZRC 2018.1373); C, L. gazi, paratype, male (15.5×13.1) (MZUF 3671); D, L. leptosoma, male (19.0×17.5) (MZUF 4978); E, L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356); F, L. liho, holotype, male (13.8×12.5) (SMF 36266); G, L. jamelense, holotype, male (11.5×10.0) (USNM 45917); H, L. moluccense, lectotype, male (14.6×13.6) (RMNH.CRUS.D 102590); I, L. paucitorum, holotype, male (19.9×18.8) (MZB.cru. 2243)." figureDoi="http://doi.org/10.5281/zenodo.3954599" httpUri="https://zenodo.org/record/3954599/files/figure.png" pageId="11" pageNumber="450">Fig. 10</figureCitation>
E–G).
</paragraph>
<caption id="DF74F3DA9A663243FF42F6AFFE09EA18" ID-DOI="http://doi.org/10.5281/zenodo.3954595" ID-Zenodo-Dep="3954595" httpUri="https://zenodo.org/record/3954595/files/figure.png" pageId="11" pageNumber="450" startId="11.[151,250,1705,1730]" targetBox="[157,1430,489,1681]" targetPageId="11">
<paragraph id="8BB4A3529A663243FF42F6AFFE09EA18" blockId="11.[151,1436,1705,1838]" pageId="11" pageNumber="450">
<emphasis id="B97F7F409A663243FF42F6AFFED7EBF4" bold="true" box="[151,276,1705,1730]" pageId="11" pageNumber="450">FIGURE 8.</emphasis>
Comparisons of male chelar dactylus and tuberculation patterns, outer views. A, 
<taxonomicName id="4C0BD8D19A663243FB89F6ADFB37EBF4" authority="Shahdadi &amp; Schubart, 2020" authorityName="Shahdadi &amp; Schubart" authorityYear="2020" box="[1116,1268,1707,1730]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="450" phylum="Arthropoda" rank="species" species="reunionense" status="sp. nov.">
<emphasis id="B97F7F409A663243FB89F6ADFB37EBF4" box="[1116,1268,1707,1730]" italics="true" pageId="11" pageNumber="450">L. reunionense</emphasis>
</taxonomicName>
<emphasis id="B97F7F409A663243FB2EF6ACFAF5EBF4" bold="true" box="[1275,1334,1706,1730]" pageId="11" pageNumber="450">
<taxonomicNameLabel id="A24CC23B9A663243FB2EF6ACFAF5EBF4" box="[1275,1334,1706,1730]" pageId="11" pageNumber="450" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
holotype, male (11.8×9.92) (UF12356); B, 
<taxonomicName id="4C0BD8D19A663243FE30F6C9FDE5EBD0" baseAuthorityName="Koller, Liu &amp; Schubart" baseAuthorityYear="2010" box="[485,550,1742,1766]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="450" phylum="Arthropoda" rank="species" species="liho">
<emphasis id="B97F7F409A663243FE30F6C9FDE5EBD0" box="[485,550,1742,1766]" italics="true" pageId="11" pageNumber="450">L. liho</emphasis>
</taxonomicName>
, holotype, male (13.8×12.5) (SMF 36266); C, 
<taxonomicName id="4C0BD8D19A663243FC2BF6C9FBBEEBD0" baseAuthorityName="Rathbun" baseAuthorityYear="1914" box="[1022,1149,1742,1766]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="450" phylum="Arthropoda" rank="species" species="jamelense">
<emphasis id="B97F7F409A663243FC2BF6C9FBBEEBD0" box="[1022,1149,1742,1766]" italics="true" pageId="11" pageNumber="450">L. jamelense</emphasis>
</taxonomicName>
, holotype, male (11.5×10.0) (USNM 45917); D, 
<taxonomicName id="4C0BD8D19A663243FEB8F6F5FDC2EA3C" baseAuthorityName="De Man" baseAuthorityYear="1892" box="[365,513,1778,1802]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="450" phylum="Arthropoda" rank="species" species="moluccense">
<emphasis id="B97F7F409A663243FEB8F6F5FDC2EA3C" box="[365,513,1778,1802]" italics="true" pageId="11" pageNumber="450">L. moluccense</emphasis>
</taxonomicName>
, lectotype, male (14.6×13.6) (RMNH.CRUS.D 102590); 
<taxonomicName id="4C0BD8D19A663243FB8AF6F5FB31EA3C" baseAuthorityName="Rahayu &amp; Ng" baseAuthorityYear="2009" box="[1119,1266,1779,1802]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="450" phylum="Arthropoda" rank="species" species="paucitorum">
<emphasis id="B97F7F409A663243FB8AF6F5FB31EA3C" box="[1119,1266,1779,1802]" italics="true" pageId="11" pageNumber="450">L. paucitorum</emphasis>
</taxonomicName>
, holotype, male (19.9×18.8) (MZB.cru. 2243).
</paragraph>
</caption>
<caption id="DF74F3DA9A613244FF42F76CFE09EAD9" ID-DOI="http://doi.org/10.5281/zenodo.3954597" ID-Zenodo-Dep="3954597" httpUri="https://zenodo.org/record/3954597/files/figure.png" pageId="12" pageNumber="451" startId="12.[151,250,1898,1923]" targetBox="[408,1179,181,1873]" targetPageId="12">
<paragraph id="8BB4A3529A613244FF42F76CFE09EAD9" blockId="12.[151,1437,1898,2031]" pageId="12" pageNumber="451">
<emphasis id="B97F7F409A613244FF42F76CFED7EAB5" bold="true" box="[151,276,1898,1923]" pageId="12" pageNumber="451">FIGURE 9.</emphasis>
Comparisons of male chelar dactylus and tuberculation patterns, dorsal views. A, 
<taxonomicName id="4C0BD8D19A613244FB8AF76DFB36EAB4" authority="Shahdadi &amp; Schubart, 2020" authorityName="Shahdadi &amp; Schubart" authorityYear="2020" box="[1119,1269,1899,1922]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="451" phylum="Arthropoda" rank="species" species="reunionense" status="sp. nov.">
<emphasis id="B97F7F409A613244FB8AF76DFB36EAB4" box="[1119,1269,1899,1922]" italics="true" pageId="12" pageNumber="451">L. reunionense</emphasis>
</taxonomicName>
<emphasis id="B97F7F409A613244FB29F76DFAF5EAB5" bold="true" box="[1276,1334,1899,1923]" pageId="12" pageNumber="451">
<taxonomicNameLabel id="A24CC23B9A613244FB29F76DFAF5EAB5" box="[1276,1334,1899,1923]" pageId="12" pageNumber="451" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
holotype, male (11.8×9.92) (UF12356); B, 
<taxonomicName id="4C0BD8D19A613244FE30F789FDE5EA91" baseAuthorityName="Koller, Liu &amp; Schubart" baseAuthorityYear="2010" box="[485,550,1935,1959]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="451" phylum="Arthropoda" rank="species" species="liho">
<emphasis id="B97F7F409A613244FE30F789FDE5EA91" box="[485,550,1935,1959]" italics="true" pageId="12" pageNumber="451">L. liho</emphasis>
</taxonomicName>
, holotype, male (13.8×12.5) (SMF 36266); C, 
<taxonomicName id="4C0BD8D19A613244FC2BF789FBBEEA91" baseAuthorityName="Rathbun" baseAuthorityYear="1914" box="[1022,1149,1935,1959]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="451" phylum="Arthropoda" rank="species" species="jamelense">
<emphasis id="B97F7F409A613244FC2BF789FBBEEA91" box="[1022,1149,1935,1959]" italics="true" pageId="12" pageNumber="451">L. jamelense</emphasis>
</taxonomicName>
, holotype, male (11.5×10.0) (USNM 45917); D, 
<taxonomicName id="4C0BD8D19A613244FEB8F7B5FDC2EAFD" baseAuthorityName="De Man" baseAuthorityYear="1892" box="[365,513,1971,1995]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="451" phylum="Arthropoda" rank="species" species="moluccense">
<emphasis id="B97F7F409A613244FEB8F7B5FDC2EAFD" box="[365,513,1971,1995]" italics="true" pageId="12" pageNumber="451">L. moluccense</emphasis>
</taxonomicName>
, lectotype, male (14.6×13.6) (RMNH.CRUS.D 102590); 
<taxonomicName id="4C0BD8D19A613244FB8AF7B5FB31EAFC" baseAuthorityName="Rahayu &amp; Ng" baseAuthorityYear="2009" box="[1119,1266,1971,1994]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="451" phylum="Arthropoda" rank="species" species="paucitorum">
<emphasis id="B97F7F409A613244FB8AF7B5FB31EAFC" box="[1119,1266,1971,1994]" italics="true" pageId="12" pageNumber="451">L. paucitorum</emphasis>
</taxonomicName>
, holotype, male (19.9×18.8) (MZB.cru. 2243).
</paragraph>
</caption>
<caption id="DF74F3DA9A603245FF42F68EFB77EA07" ID-DOI="http://doi.org/10.5281/zenodo.3954599" ID-Zenodo-Dep="3954599" httpUri="https://zenodo.org/record/3954599/files/figure.png" pageId="13" pageNumber="452" startId="13.[151,250,1672,1697]" targetBox="[155,1432,181,1646]" targetPageId="13">
<paragraph id="8BB4A3529A603245FF42F68EFB77EA07" blockId="13.[151,1437,1672,1841]" pageId="13" pageNumber="452">
<emphasis id="B97F7F409A603245FF42F68EFEE2EB97" bold="true" box="[151,289,1672,1697]" pageId="13" pageNumber="452">FIGURE 10.</emphasis>
Comparisons of G1, apical part, dorsal views. A, 
<taxonomicName id="4C0BD8D19A603245FCFEF68FFC6CEB96" authority="Shahdadi &amp; Schubart, 2020" authorityName="Shahdadi &amp; Schubart" authorityYear="2020" box="[811,943,1672,1696]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="452" phylum="Arthropoda" rank="species" species="dispersum" status="sp. nov.">
<emphasis id="B97F7F409A603245FCFEF68FFC6CEB96" box="[811,943,1672,1696]" italics="true" pageId="13" pageNumber="452">L. dispersum</emphasis>
</taxonomicName>
<emphasis id="B97F7F409A603245FC6DF68FFC30EB97" bold="true" box="[952,1011,1673,1697]" pageId="13" pageNumber="452">
<taxonomicNameLabel id="A24CC23B9A603245FC6DF68FFC30EB97" box="[952,1011,1673,1697]" pageId="13" pageNumber="452" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
holotype, male (12.2×10.1) (MNHN-IU- 2012-660); B, 
<taxonomicName id="4C0BD8D19A603245FEF8F6ABFE43EBF2" baseAuthorityName="De Man" baseAuthorityYear="1895" box="[301,384,1708,1732]" class="Malacostraca" family="Sesarmidae" genus="Parasesarma" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="452" phylum="Arthropoda" rank="species" species="lenzii">
<emphasis id="B97F7F409A603245FEF8F6ABFE43EBF2" box="[301,384,1708,1732]" italics="true" pageId="13" pageNumber="452">L. lenzii</emphasis>
</taxonomicName>
, male (11.8×9.9) (ZRC 2018.1373); C, 
<taxonomicName id="4C0BD8D19A603245FCCDF6ABFC9EEBF2" box="[792,861,1709,1732]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="452" phylum="Arthropoda" rank="species" species="gazi">
<emphasis id="B97F7F409A603245FCCDF6ABFC9EEBF2" box="[792,861,1709,1732]" italics="true" pageId="13" pageNumber="452">L. gazi</emphasis>
</taxonomicName>
, paratype, male (15.5×13.1) (MZUF 3671); D, 
<taxonomicName id="4C0BD8D19A603245FA93F6ABFF0FEBDE" baseAuthorityName="Hilgendorf" baseAuthorityYear="1869" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="452" phylum="Arthropoda" rank="species" species="leptosoma">
<emphasis id="B97F7F409A603245FA93F6ABFF0FEBDE" italics="true" pageId="13" pageNumber="452">L. leptosoma</emphasis>
</taxonomicName>
, male (19.0×17.5) (MZUF 4978); E, 
<taxonomicName id="4C0BD8D19A603245FD87F6D7FD2AEBDE" authority="Shahdadi &amp; Schubart, 2020" authorityName="Shahdadi &amp; Schubart" authorityYear="2020" box="[594,745,1745,1768]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="452" phylum="Arthropoda" rank="species" species="reunionense" status="sp. nov.">
<emphasis id="B97F7F409A603245FD87F6D7FD2AEBDE" box="[594,745,1745,1768]" italics="true" pageId="13" pageNumber="452">L. reunionense</emphasis>
</taxonomicName>
<emphasis id="B97F7F409A603245FD24F6D7FCE8EBDF" bold="true" box="[753,811,1745,1769]" pageId="13" pageNumber="452">
<taxonomicNameLabel id="A24CC23B9A603245FD24F6D7FCE8EBDF" box="[753,811,1745,1769]" pageId="13" pageNumber="452" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
holotype, male (11.8×9.92) (UF12356); F, 
<taxonomicName id="4C0BD8D19A603245FB3EF6D7FAECEBDE" baseAuthorityName="Koller, Liu &amp; Schubart" baseAuthorityYear="2010" box="[1259,1327,1744,1768]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="452" phylum="Arthropoda" rank="species" species="liho">
<emphasis id="B97F7F409A603245FB3EF6D7FAECEBDE" box="[1259,1327,1744,1768]" italics="true" pageId="13" pageNumber="452">L. liho</emphasis>
</taxonomicName>
, holotype, male (13.8×12.5) (SMF 36266); G, 
<taxonomicName id="4C0BD8D19A603245FE28F6F3FDBFEA3A" baseAuthorityName="Rathbun" baseAuthorityYear="1914" box="[509,636,1780,1804]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="452" phylum="Arthropoda" rank="species" species="jamelense">
<emphasis id="B97F7F409A603245FE28F6F3FDBFEA3A" box="[509,636,1780,1804]" italics="true" pageId="13" pageNumber="452">L. jamelense</emphasis>
</taxonomicName>
, holotype, male (11.5×10.0) (USNM 45917); H, 
<taxonomicName id="4C0BD8D19A603245FBB2F6F3FB34EA3A" baseAuthorityName="De Man" baseAuthorityYear="1892" box="[1127,1271,1780,1804]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="452" phylum="Arthropoda" rank="species" species="moluccense">
<emphasis id="B97F7F409A603245FBB2F6F3FB34EA3A" box="[1127,1271,1780,1804]" italics="true" pageId="13" pageNumber="452">L. moluccense</emphasis>
</taxonomicName>
, lectotype, male (14.6×13.6) (RMNH.CRUS.D 102590); I, 
<taxonomicName id="4C0BD8D19A603245FD9DF71FFD1BEA06" baseAuthorityName="Rahayu &amp; Ng" baseAuthorityYear="2009" box="[584,728,1817,1840]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="452" phylum="Arthropoda" rank="species" species="paucitorum">
<emphasis id="B97F7F409A603245FD9DF71FFD1BEA06" box="[584,728,1817,1840]" italics="true" pageId="13" pageNumber="452">L. paucitorum</emphasis>
</taxonomicName>
, holotype, male (19.9×18.8) (MZB.cru. 2243).
</paragraph>
</caption>
<paragraph id="8BB4A3529A603246FF12F785FA48EDE1" blockId="13.[151,1436,1923,2021]" lastBlockId="14.[151,1437,153,323]" lastPageId="14" lastPageNumber="453" pageId="13" pageNumber="452">
The new species shows phylogenetic association (
<figureCitation id="1330BFD79A603245FD21F785FC89EAAB" box="[756,842,1923,1949]" captionStart="FIGURE 1" captionStartId="2.[151,250,1798,1823]" captionTargetBox="[220,1376,684,1749]" captionTargetId="figure@2.[189,1398,669,1773]" captionTargetPageId="2" captionText="FIGURE 1. A, Phylogenetic consensus tree constructed with Maximum Likelihood (ML) and Bayesian Inference (BI) of the COX1 gene, focusing on Leptarma species; four species of Parasesarma were added as outgroups (only confidence values higher than 50% for ML (bootstrap value, bt) and 90% for BI (posterior probabilities, pp) are shown in the trees as bt/pp; B, C, maximum parsimony haplotype network of COX1 gene constructed with PopART for specimens of B, L. dispersum n. sp. and allied species; C, L. reunionense n. sp. and allied species." figureDoi="http://doi.org/10.5281/zenodo.3954579" httpUri="https://zenodo.org/record/3954579/files/figure.png" pageId="13" pageNumber="452">Fig. 1A</figureCitation>
) and some morphological resemblance to the species pair 
<taxonomicName id="4C0BD8D19A603245FF1CF7AEFEA9EAF7" baseAuthorityName="De Man" baseAuthorityYear="1892" box="[201,362,1959,1985]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="452" phylum="Arthropoda" rank="species" species="moluccense">
<emphasis id="B97F7F409A603245FF1CF7AEFEA9EAF7" box="[201,362,1959,1985]" italics="true" pageId="13" pageNumber="452">L. moluccense</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0BD8D19A603245FE4AF7AEFDFEEAF7" baseAuthorityName="Rahayu &amp; Ng" baseAuthorityYear="2009" box="[415,573,1960,1985]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="452" phylum="Arthropoda" rank="species" species="paucitorum">
<emphasis id="B97F7F409A603245FE4AF7AEFDFEEAF7" box="[415,573,1960,1985]" italics="true" pageId="13" pageNumber="452">L. paucitorum</emphasis>
</taxonomicName>
. The new species, however, has a wider carapace compared to these two species (
<figureCitation id="1330BFD79A603245FF06F7CDFE93EAD3" box="[211,336,1995,2021]" captionStart="FIGURE 7" captionStartId="10.[151,250,1866,1891]" captionTargetBox="[151,1431,190,1835]" captionTargetId="figure@10.[150,1436,181,1841]" captionTargetPageId="10" captionText="FIGURE 7. Comparisons of carapace and ambulatory legs, dorsal views. A, L. dispersum n. sp. holotype, male (12.2×10.1) (MNHN-IU-2012-660); B, L. lenzii, male (11.8×9.9) (ZRC 2018.1373); C, L. gazi, paratype, male (15.5×13.1) (MZUF 3671); D, L. leptosoma, male (19.0×17.5) (MZUF 4978); E, L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356); F, L. liho, holotype, male (13.8×12.5) (SMF 36266); G, L. moluccense, lectotype, male (14.6×13.6) (RMNH.CRUS.D 102590); H, L. paucitorum, holotype, male (19.9×18.8) (MZB.cru. 2243)." figureDoi="http://doi.org/10.5281/zenodo.3954593" httpUri="https://zenodo.org/record/3954593/files/figure.png" pageId="13" pageNumber="452">Fig. 7G, H</figureCitation>
), i.e. cw/cl ca. 
<quantity id="4CF30EB79A603245FE29F7CDFDFEEAD3" box="[508,573,1995,2021]" metricMagnitude="-2" metricUnit="m" metricValue="3.048" pageId="13" pageNumber="452" unit="in" value="1.2">1.2 in</quantity>
the new species, while it is less than 
<quantity id="4CF30EB79A603245FC3DF7CDFBEAEAD3" box="[1000,1065,1995,2021]" metricMagnitude="-2" metricUnit="m" metricValue="2.794" pageId="13" pageNumber="452" unit="in" value="1.1">1.1 in</quantity>
the other two species. They also differ in chelar dactylar tuberculation and G1 morphology. The new species has more tubercles (9-10) than 
<taxonomicName id="4C0BD8D19A633246FA86F09CFF34EDE1" baseAuthorityName="De Man" baseAuthorityYear="1892" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="453" phylum="Arthropoda" rank="species" species="moluccense">
<emphasis id="B97F7F409A633246FA86F09CFF34EDE1" italics="true" pageId="14" pageNumber="453">L. moluccense</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0BD8D19A633246FEFBF0B8FE0EEDE1" baseAuthorityName="Rahayu &amp; Ng" baseAuthorityYear="2009" box="[302,461,190,215]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="453" phylum="Arthropoda" rank="species" species="paucitorum">
<emphasis id="B97F7F409A633246FEFBF0B8FE0EEDE1" box="[302,461,190,215]" italics="true" pageId="14" pageNumber="453">L. paucitorum</emphasis>
</taxonomicName>
(6-8) (
<figureCitation id="1330BFD79A633246FDCEF0BBFD5EEDE1" box="[539,669,189,215]" captionStart="FIGURE 8" captionStartId="11.[151,250,1705,1730]" captionTargetBox="[157,1430,489,1681]" captionTargetId="figure@11.[157,1430,489,1681]" captionTargetPageId="11" captionText="FIGURE 8. Comparisons of male chelar dactylus and tuberculation patterns, outer views. A, L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356); B, L. liho, holotype, male (13.8×12.5) (SMF 36266); C, L. jamelense, holotype, male (11.5×10.0) (USNM 45917); D, L. moluccense, lectotype, male (14.6×13.6) (RMNH.CRUS.D 102590); L. paucitorum, holotype, male (19.9×18.8) (MZB.cru. 2243)." figureDoi="http://doi.org/10.5281/zenodo.3954595" httpUri="https://zenodo.org/record/3954595/files/figure.png" pageId="14" pageNumber="453">Figs. 8D, E</figureCitation>
, 
<figureCitation id="1330BFD79A633246FD7FF0BBFD2FEDE1" box="[682,748,189,215]" captionStart="FIGURE 9" captionStartId="12.[151,250,1898,1923]" captionTargetBox="[408,1179,181,1873]" captionTargetId="figure@12.[408,1179,181,1873]" captionTargetPageId="12" captionText="FIGURE 9. Comparisons of male chelar dactylus and tuberculation patterns, dorsal views. A, L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356); B, L. liho, holotype, male (13.8×12.5) (SMF 36266); C, L. jamelense, holotype, male (11.5×10.0) (USNM 45917); D, L. moluccense, lectotype, male (14.6×13.6) (RMNH.CRUS.D 102590); L. paucitorum, holotype, male (19.9×18.8) (MZB.cru. 2243)." figureDoi="http://doi.org/10.5281/zenodo.3954597" httpUri="https://zenodo.org/record/3954597/files/figure.png" pageId="14" pageNumber="453">9D, E</figureCitation>
) and G1s with longer apical corneous process (
<figureCitation id="1330BFD79A633246FB2AF0BBFABEEDE1" box="[1279,1405,189,215]" captionStart="FIGURE 10" captionStartId="13.[151,250,1672,1697]" captionTargetBox="[155,1432,181,1646]" captionTargetId="figure@13.[155,1432,181,1646]" captionTargetPageId="13" captionText="FIGURE 10. Comparisons of G1, apical part, dorsal views. A, L. dispersum n. sp. holotype, male (12.2×10.1) (MNHN-IU- 2012-660); B, L. lenzii, male (11.8×9.9) (ZRC 2018.1373); C, L. gazi, paratype, male (15.5×13.1) (MZUF 3671); D, L. leptosoma, male (19.0×17.5) (MZUF 4978); E, L. reunionense n. sp. holotype, male (11.8×9.92) (UF12356); F, L. liho, holotype, male (13.8×12.5) (SMF 36266); G, L. jamelense, holotype, male (11.5×10.0) (USNM 45917); H, L. moluccense, lectotype, male (14.6×13.6) (RMNH.CRUS.D 102590); I, L. paucitorum, holotype, male (19.9×18.8) (MZB.cru. 2243)." figureDoi="http://doi.org/10.5281/zenodo.3954599" httpUri="https://zenodo.org/record/3954599/files/figure.png" pageId="14" pageNumber="453">Fig. 10H, I</figureCitation>
).
</paragraph>
<paragraph id="8BB4A3529A633246FF12F0E4FD42EC75" blockId="14.[151,1437,153,323]" pageId="14" pageNumber="453">
<taxonomicName id="4C0BD8D19A633246FF12F0E4FE01EDCD" authority="Shahdadi &amp; Schubart, 2020" authorityName="Shahdadi &amp; Schubart" authorityYear="2020" box="[199,450,226,251]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="453" phylum="Arthropoda" rank="species" species="reunionense" status="sp. nov.">
<emphasis id="B97F7F409A633246FF12F0E4FE01EDCD" box="[199,450,226,251]" italics="true" pageId="14" pageNumber="453">Leptarma reunionense</emphasis>
</taxonomicName>
<emphasis id="B97F7F409A633246FE1DF0E7FDC5EDCD" bold="true" box="[456,518,225,251]" pageId="14" pageNumber="453">
<taxonomicNameLabel id="A24CC23B9A633246FE1DF0E7FDC5EDCD" box="[456,518,225,251]" pageId="14" pageNumber="453" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
also resembles 
<taxonomicName id="4C0BD8D19A633246FD60F0E4FCFAEDCD" baseAuthorityName="Li, Ng &amp; Shih" baseAuthorityYear="2019" box="[693,825,225,251]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="453" phylum="Arthropoda" rank="species" species="aurifrons">
<emphasis id="B97F7F409A633246FD60F0E4FCFAEDCD" box="[693,825,225,251]" italics="true" pageId="14" pageNumber="453">L. aurifrons</emphasis>
</taxonomicName>
in general appearance as well as in chelar morphology. 
<taxonomicName id="4C0BD8D19A633246FF42F100FEFFEC29" authority="Shahdadi &amp; Schubart, 2020" authorityName="Shahdadi &amp; Schubart" authorityYear="2020" box="[151,316,262,287]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="453" phylum="Arthropoda" rank="species" species="reunionense" status="sp. nov.">
<emphasis id="B97F7F409A633246FF42F100FEFFEC29" box="[151,316,262,287]" italics="true" pageId="14" pageNumber="453">L. reunionense</emphasis>
</taxonomicName>
<emphasis id="B97F7F409A633246FE97F103FE42EC29" bold="true" box="[322,385,261,287]" pageId="14" pageNumber="453">
<taxonomicNameLabel id="A24CC23B9A633246FE97F103FE42EC29" box="[322,385,261,287]" pageId="14" pageNumber="453" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, however, differs from 
<taxonomicName id="4C0BD8D19A633246FD52F100FCC8EC29" baseAuthorityName="Li, Ng &amp; Shih" baseAuthorityYear="2019" box="[647,779,261,287]" class="Malacostraca" family="Sesarmidae" genus="Leptarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="453" phylum="Arthropoda" rank="species" species="aurifrons">
<emphasis id="B97F7F409A633246FD52F100FCC8EC29" box="[647,779,261,287]" italics="true" pageId="14" pageNumber="453">L. aurifrons</emphasis>
</taxonomicName>
in G1 morphology, which has longer and more bent corneous process (see 
<bibRefCitation id="EF9ADEA39A633246FE81F12FFE35EC75" author="Li, J. - J. &amp; Shih, H. T. &amp; Ng, P. K. L." box="[340,502,297,323]" pageId="14" pageNumber="453" pagination="40" refId="ref8602" refString="Li, J. - J., Shih, H. T. &amp; Ng, P. K. L. (2019) Three new species and two new records of Parasesarma De Man, 1895 (Crustacea: Brachyura: Sesarmidae) from Taiwan and the Philippines from morphological and molecular evidence. Zoological Studies, 58, 40." type="journal article" year="2019">
Li 
<emphasis id="B97F7F409A633246FEA1F12CFE66EC75" box="[372,421,297,323]" italics="true" pageId="14" pageNumber="453">et al</emphasis>
., 2019
</bibRefCitation>
: 
<figureCitation id="1330BFD79A633246FDD4F12FFDB7EC75" box="[513,628,297,323]" captionStart="FIGURE 3" captionStartId="5.[151,250,1685,1710]" captionTargetBox="[152,1435,345,1660]" captionTargetId="figure@5.[152,1435,345,1660]" captionTargetPageId="5" captionText="FIGURE 3. L. dispersum n. sp. holotype, male (12.2×10.1) (MNHN-IU-2012-660).A, frontal view; B, right chela, outer view; C, right chela, dorsal view; D, right chela, palm pectinated crest; E, pleon; F, pleonal somites 1–3; G–J, G1; right G, dorsal view; H, apical corneous part, dorsal view (arrows pointing to aperture); I, apical corneous part, ventral view; J, front view of apical corneous part." figureDoi="http://doi.org/10.5281/zenodo.3954583" httpUri="https://zenodo.org/record/3954583/files/figure.png" pageId="14" pageNumber="453">Fig. 3E, F</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>