treatments-xml/data/03/98/ED/0398ED589D5C352CFEF8F9C1FB00FA8E.xml

<document id="070EF6987E9FEF69CB603A635A833D68" ID-DOI="10.5281/zenodo.174282" ID-GBIF-Dataset="1617eb93-6cef-4e89-8193-431cb0e0f50d" ID-ISSN="1175-5326" ID-Zenodo-Dep="174282" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459794559725" checkinUser="plazi" docAuthor="Brown, Rafe M., Richards, Stephen J., Sukumaran, Jeet &amp; Foufopoulos, Johannes" docDate="2006" docId="0398ED589D5C352CFEF8F9C1FB00FA8E" docLanguage="en" docName="zt01334p068.pdf" docOrigin="Zootaxa 1334" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Platymantis adiastola Brown, Richards, Sukumaran &amp; Foufopoulos, 2006, new species" docType="treatment" docVersion="10" lastPageNumber="63" masterDocId="FFA195209D59353EFFF0FFAFFFF5FF90" masterDocTitle="A new morphologically cryptic species of forest frog (genus Platymantis) from New Britain Island, Bismarck Archipelago" masterLastPageNumber="68" masterPageNumber="45" pageNumber="50" updateTime="1698219521436" updateUser="plazi">
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<mods:title id="5697949F0B16ED86EA9434F99F2B1E17">A new morphologically cryptic species of forest frog (genus Platymantis) from New Britain Island, Bismarck Archipelago</mods:title>
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<mods:namePart id="992AB49D84E815F766FA1CEA2CF727A9">Brown, Rafe M.</mods:namePart>
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<mods:namePart id="8F371A82BBA74D075FA1039185D6F166">Richards, Stephen J.</mods:namePart>
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<mods:namePart id="9408D8F708CD887EC1FEC5B80B66630F">Sukumaran, Jeet</mods:namePart>
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<mods:namePart id="B48118E84C742237606D95D3B514B31C">Foufopoulos, Johannes</mods:namePart>
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<mods:date id="4082706886753A89B90EE4F1CAD229CD">2006</mods:date>
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<treatment id="0398ED589D5C352CFEF8F9C1FB00FA8E" ID-DOI="http://doi.org/10.5281/zenodo.5625771" ID-GBIF-Taxon="119350860" ID-Zenodo-Dep="5625771" LSID="urn:lsid:plazi:treatment:0398ED589D5C352CFEF8F9C1FB00FA8E" httpUri="http://treatment.plazi.org/id/0398ED589D5C352CFEF8F9C1FB00FA8E" lastPageId="18" lastPageNumber="63" pageId="5" pageNumber="50">
<subSubSection id="C32B0FC59D5C353BFEF8F9C1FE74F93B" pageId="5" pageNumber="50" type="nomenclature">
<paragraph id="8B8E5C4E9D5C353BFEF8F9C1FD5FF918" blockId="5.[264,682,1646,1707]" box="[264,682,1646,1672]" pageId="5" pageNumber="50">
<heading id="D0C6EB229D5C353BFEF8F9C1FD5FF918" bold="true" box="[264,682,1646,1672]" fontSize="11" level="1" pageId="5" pageNumber="50" reason="1">
<emphasis id="B945805C9D5C353BFEF8F9C1FD5FF918" bold="true" box="[264,682,1646,1672]" pageId="5" pageNumber="50">
<taxonomicName id="4C3127CD9D5C353BFEF8F9C1FDFEF918" ID-CoL="4JSP3" box="[264,523,1646,1672]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="50" phylum="Chordata" rank="species" species="adiastola" status="sp. nov.">
<emphasis id="B945805C9D5C353BFEF8F9C1FDFEF918" bold="true" box="[264,523,1646,1672]" italics="true" pageId="5" pageNumber="50">Platymantis adiastola</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A2763D279D5C353BFDEBF9C1FD5FF918" box="[539,682,1646,1672]" pageId="5" pageNumber="50" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8E5C4E9D5C353BFEF8F93EFE74F93B" blockId="5.[264,682,1646,1707]" box="[264,385,1681,1707]" pageId="5" pageNumber="50">
(
<figureCitation id="130A40CB9D5C353BFEE0F93EFE8CF93B" box="[272,377,1681,1707]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStartId-0="6.[264,367,1801,1825]" captionStartId-1="7.[264,367,1796,1820]" captionStartId-2="11.[264,367,1529,1553]" captionText-0="FIGURE 4. Dorsal views of P. a d i a s t o l a (A: SAMA R 61908; B: SAMA R 60257), P. s c h m i d t i from Wanui (C: SAMA R 57015; D: SAMA R 57016), and P. schmidti paratypes (E: SAMA 6858; F: SAMA 6862) from Talasea, Willameuz Peninsula. Scale bars = 2 cm." captionText-1="FIGURE 5. Photographs in life of P. a d i a s t o l a (A: paratype SAMA R 61907) and P. schmidti (B: SAMA R 57015)." captionText-2="FIGURE 6. Ventral views of hands and feet of P. a d i a s t o l a (A, B: SAMA R 61908), a recent specimen of P. schmidti from Wanui (C, D: SAMA R 57016), and a paratype of P. schmidti from Talasea (E, F: SAMA R 6862). Scale bars = 2 mm." pageId="5" pageNumber="50">Figs 4–6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32B0FC59D5C3536FEF8F979FADCFE16" lastPageId="8" lastPageNumber="53" pageId="5" pageNumber="50" type="materials_examined">
<paragraph id="8B8E5C4E9D5C353BFEF8F979FB72F8D1" blockId="5.[264,1323,1750,1857]" pageId="5" pageNumber="50">
<emphasis id="B945805C9D5C353BFEF8F979FE8AF960" bold="true" box="[264,383,1750,1776]" pageId="5" pageNumber="50">
<typeStatus id="548AE2EC9D5C353BFEF8F979FE80F960" box="[264,373,1750,1776]" pageId="5" pageNumber="50" type="holotype">
<emphasis id="B945805C9D5C353BFEF8F979FE80F960" bold="true" box="[264,373,1750,1776]" italics="true" pageId="5" pageNumber="50">Holotype</emphasis>
</typeStatus>
:
</emphasis>
<collectionCode id="ED20C48B9D5C353BFE79F978FE2BF961" LSID="urn:lsid:biocol.org:col:34982" box="[393,478,1751,1777]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="5" pageNumber="50">SAMA</collectionCode>
R61906, adult male, collected by S. Richards at Wanui Camp, Wanui River Valley (near Wide Bay), 
<quantity id="4CC9F1AB9D5C353BFD86F950FD34F889" box="[630,705,1791,1817]" metricMagnitude="2" metricUnit="m" metricValue="3.1" pageId="5" pageNumber="50" unit="m" value="310.0">310 m</quantity>
above sea level (
<geoCoordinate id="EE053A899D5C353BFC7DF950FBD4F889" box="[909,1057,1791,1817]" direction="south" orientation="latitude" pageId="5" pageNumber="50" precision="1" value="-5.3606334">05 21.638’S</geoCoordinate>
, 
<geoCoordinate id="EE053A899D5C353BFBDEF950FB28F889" box="[1070,1245,1791,1817]" direction="east" orientation="longitude" pageId="5" pageNumber="50" precision="1" value="152.08777">152° 05.266’E</geoCoordinate>
), New 
<collectingCountry id="F3261CDE9D5C353BFEF8F888FEACF8D1" box="[264,345,1831,1857]" name="United Kingdom" pageId="5" pageNumber="50">Britain</collectingCountry>
Isl., East New 
<collectingCountry id="F3261CDE9D5C353BFDFCF888FDABF8D1" box="[524,606,1831,1857]" name="United Kingdom" pageId="5" pageNumber="50">Britain</collectingCountry>
Province, 
<collectingCountry id="F3261CDE9D5C353BFD2CF888FC34F8D1" box="[732,961,1831,1857]" name="Papua New Guinea" pageId="5" pageNumber="50">Papua New Guinea</collectingCountry>
, 
<date id="FF8F7A8E9D5C353BFC3CF888FB76F8D1" box="[972,1155,1831,1857]" pageId="5" pageNumber="50" value="2000-03-20">20 March 2000</date>
.
</paragraph>
<caption id="DF4E0CC69D5F3538FEF8F8A6FC06F8F6" pageId="6" pageNumber="51">
<paragraph id="8B8E5C4E9D5F3538FEF8F8A6FC06F8F6" blockId="6.[264,1323,1801,1894]" pageId="6" pageNumber="51">
<emphasis id="B945805C9D5F3538FEF8F8A6FE7CF8B1" bold="true" box="[264,393,1801,1825]" pageId="6" pageNumber="51">FIGURE 4.</emphasis>
Dorsal views of 
<emphasis id="B945805C9D5F3538FDCEF8A5FD42F8B1" box="[574,695,1802,1825]" italics="true" pageId="6" pageNumber="51">P. a d i a s t o l a</emphasis>
(A: SAMA R61908; B: SAMA R60257), 
<emphasis id="B945805C9D5F3538FB8CF8A5FB18F8B1" box="[1148,1261,1802,1825]" italics="true" pageId="6" pageNumber="51">P. s c h m i d t i</emphasis>
from Wanui (C: SAMA R57015; D: SAMA R57016), and 
<taxonomicName id="4C3127CD9D5F3538FCAAF883FC27F8D3" box="[858,978,1836,1859]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="51" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B945805C9D5F3538FCAAF883FC27F8D3" box="[858,978,1836,1859]" italics="true" pageId="6" pageNumber="51">P. schmidti</emphasis>
</taxonomicName>
paratypes (E: SAMA 6858; F: SAMA 6862) from Talasea, Willameuz Peninsula. Scale bars = 2 cm.
</paragraph>
</caption>
<caption id="DF4E0CC69D5E3539FEF8F8ABFE35F8AE" pageId="7" pageNumber="52">
<paragraph id="8B8E5C4E9D5E3539FEF8F8ABFE35F8AE" blockId="7.[264,1323,1796,1854]" pageId="7" pageNumber="52">
<emphasis id="B945805C9D5E3539FEF8F8ABFE7EF88C" bold="true" box="[264,395,1796,1820]" pageId="7" pageNumber="52">FIGURE 5.</emphasis>
Photographs in life of 
<emphasis id="B945805C9D5E3539FD79F8ABFCF0F88B" box="[649,773,1796,1819]" italics="true" pageId="7" pageNumber="52">P. a d i a s t o l a</emphasis>
(A: paratype SAMA R61907) and 
<taxonomicName id="4C3127CD9D5E3539FB7AF8ABFAF5F88B" box="[1162,1280,1796,1819]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B945805C9D5E3539FB7AF8ABFAF5F88B" box="[1162,1280,1796,1819]" italics="true" pageId="7" pageNumber="52">P. schmidti</emphasis>
</taxonomicName>
(B: SAMA R57015).
</paragraph>
</caption>
<paragraph id="8B8E5C4E9D513536FEC8FEB3FADCFE16" blockId="8.[264,1323,283,1870]" pageId="8" pageNumber="53">
<emphasis id="B945805C9D513536FEC8FEB3FE4DFEA5" bold="true" box="[312,440,283,310]" pageId="8" pageNumber="53">
<emphasis id="B945805C9D513536FEC8FEB3FE5AFEA6" bold="true" box="[312,431,284,310]" italics="true" pageId="8" pageNumber="53">
<typeStatus id="548AE2EC9D513536FEC8FEB3FE5AFEA6" box="[312,431,284,310]" pageId="8" pageNumber="53" type="paratype">Paratypes</typeStatus>
</emphasis>
:
</emphasis>
<collectionCode id="ED20C48B9D513536FE32FEB3FDE2FEA6" LSID="urn:lsid:biocol.org:col:34982" box="[450,535,284,310]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="8" pageNumber="53">SAMA</collectionCode>
R57014–15, R57040–41, same data as 
<typeStatus id="548AE2EC9D513536FC09FEB3FB95FEA6" box="[1017,1120,284,310]" pageId="8" pageNumber="53" type="holotype">holotype</typeStatus>
except collected 
<date id="FF8F7A8E9D513536FEF8FEEBFE4AFECE" box="[264,447,324,350]" pageId="8" pageNumber="53" value="2000-03-13">13 March 2000</date>
; R60257–8, 
<date id="FF8F7A8E9D513536FDA4FEEBFCFFFECE" box="[596,778,324,350]" pageId="8" pageNumber="53" value="2000-03-15">15 March 2000</date>
; R60259, 
<date id="FF8F7A8E9D513536FC72FEEBFBCCFECE" box="[898,1081,324,350]" pageId="8" pageNumber="53" value="2000-03-18">18 March 2000</date>
; 
<collectionCode id="ED20C48B9D513536FBB5FEEBFB6FFECE" LSID="urn:lsid:biocol.org:col:34982" box="[1093,1178,324,350]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="8" pageNumber="53">SAMA</collectionCode>
R61907, 
<date id="FF8F7A8E9D513536FAFDFEEBFEADFE16" pageId="8" pageNumber="53" value="1919-03">19 March</date>
; 
<collectionCode id="ED20C48B9D513536FE94FEC3FE4CFE16" LSID="urn:lsid:biocol.org:col:34982" box="[356,441,364,390]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="8" pageNumber="53">SAMA</collectionCode>
R61908, 
<collectionCode id="ED20C48B9D513536FDDBFEC3FD8EFE16" box="[555,635,364,390]" pageId="8" pageNumber="53">UPNG</collectionCode>
9974-6, 
<date id="FF8F7A8E9D513536FD14FEC3FC6FFE16" box="[740,922,364,390]" pageId="8" pageNumber="53" value="2000-03-20">20 March 2000</date>
; 
<collectionCode id="ED20C48B9D513536FC56FEC3FC0EFE16" LSID="urn:lsid:biocol.org:col:34982" box="[934,1019,364,390]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="8" pageNumber="53">SAMA</collectionCode>
R61909, 
<date id="FF8F7A8E9D513536FB9EFEC3FAD0FE16" box="[1134,1317,364,390]" pageId="8" pageNumber="53" value="2000-03-22">22 March 2000</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C32B0FC59D513536FEC8FE3BFD41FE6E" pageId="8" pageNumber="53" type="etymology">
<paragraph id="8B8E5C4E9D513536FEC8FE3BFD41FE6E" blockId="8.[264,1323,283,1870]" pageId="8" pageNumber="53">
<emphasis id="B945805C9D513536FEC8FE3BFE37FE3D" bold="true" box="[312,450,403,430]" pageId="8" pageNumber="53">
<emphasis id="B945805C9D513536FEC8FE3BFE4DFE3E" bold="true" box="[312,440,404,430]" italics="true" pageId="8" pageNumber="53">Etymology</emphasis>
:
</emphasis>
The specific epithet is chosen from the Greek 
<emphasis id="B945805C9D513536FBC5FE3BFB5BFE3D" box="[1077,1198,404,429]" italics="true" pageId="8" pageNumber="53">adiastolos</emphasis>
, meaning confused, mixed up, or not separated, in reference to past confusion surrounding the taxonomic status of the new species.
</paragraph>
</subSubSection>
<subSubSection id="C32B0FC59D513536FEC8FDA3FC98FA8E" pageId="8" pageNumber="53" type="diagnosis">
<paragraph id="8B8E5C4E9D513536FEC8FDA3FCFEFC86" blockId="8.[264,1323,283,1870]" pageId="8" pageNumber="53">
<emphasis id="B945805C9D513536FEC8FDA3FE4FFDB5" bold="true" box="[312,442,523,550]" pageId="8" pageNumber="53">
<emphasis id="B945805C9D513536FEC8FDA3FE45FDB6" bold="true" box="[312,432,524,550]" italics="true" pageId="8" pageNumber="53">Diagnosis</emphasis>
:
</emphasis>
<taxonomicName id="4C3127CD9D513536FE3EFDA3FD24FDB5" box="[462,721,524,549]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="adiastola">
<emphasis id="B945805C9D513536FE3EFDA3FD24FDB5" box="[462,721,524,549]" italics="true" pageId="8" pageNumber="53">Platymantis adiastola</emphasis>
</taxonomicName>
is distinguished from congeners by (1) body size (39.5–43.9 for 
<specimenCount id="9D3797C79D513536FE47FD9BFDD7FDDE" box="[439,546,564,590]" pageId="8" pageNumber="53" type="male">11 males</specimenCount>
; 71.7 for one female), (2) narrowly expanded terminal disks of the fingers and toes, (3) skin of dorsum with scattered longitudinal tubercular ridges, including two long (
<quantity id="4CC9F1AB9D513536FDF5FD2BFD9AFD0E" box="[517,623,644,670]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="8" pageNumber="53" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
) medially bowed suprascapular dermal ridges, (4) upper eyelids devoid of conical tubercles, (5) subarticular tubercles of fingers, toes, hands and feet prominent and raised, (6) digits lacking all vestiges of lateral flange and interdigital webbing, and (7) unique advertisement call.
</paragraph>
<paragraph id="8B8E5C4E9D513536FEC8FC8BFC98FA8E" blockId="8.[264,1323,283,1870]" pageId="8" pageNumber="53">
The new species is morphologically nearly identical to 
<taxonomicName id="4C3127CD9D513536FC4BFC8BFBCEFCAD" box="[955,1083,804,829]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B945805C9D513536FC4BFC8BFBCEFCAD" box="[955,1083,804,829]" italics="true" pageId="8" pageNumber="53">P. schmidti</emphasis>
</taxonomicName>
but differs from this species by slight differences in body size and proportions (
<tableCitation id="C6B369F59D513536FC4DFCE3FBE0FCF6" box="[957,1045,844,870]" captionStart="TABLE 1" captionStartId="9.[264,352,283,307]" captionTargetBox="[272,1304,524,1892]" captionTargetPageId="9" captionText="TABLE 1. Morphometric variation (in mm) in adult Platymantis adiastola n. sp. and P. s c h m i d t i (original collections, holotype and paratypes, as well as recent collections). Table entries include mean ± 1 standard deviation (with range shown below), and sample size (n). See text for character abbreviations." httpUri="http://table.plazi.org/id/DF4E0CC69D503537FEF8FEB4FE6AFE34" pageId="8" pageNumber="53" tableUuid="DF4E0CC69D503537FEF8FEB4FE6AFE34">Table 1</tableCitation>
), an apparent tendency towards a more elevated calling perch preference, and a possible greater extent of sexual size dimorphism (
<taxonomicName id="4C3127CD9D513536FE12FC33FD9AFC25" box="[482,623,924,949]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="adiastola">
<emphasis id="B945805C9D513536FE12FC33FD9AFC25" box="[482,623,924,949]" italics="true" pageId="8" pageNumber="53">P. adiastola</emphasis>
</taxonomicName>
male/female SVL = 0.57; 
<taxonomicName id="4C3127CD9D513536FC46FC33FBCCFC25" box="[950,1081,924,949]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B945805C9D513536FC46FC33FBCCFC25" box="[950,1081,924,949]" italics="true" pageId="8" pageNumber="53">P. schmidti</emphasis>
</taxonomicName>
= 0.69). The main distinguishing feature between these two morphologically cryptic forms is the male advertisement call. 
<taxonomicName id="4C3127CD9D513536FE0AFC43FD09FB95" box="[506,764,1004,1029]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="adiastola">
<emphasis id="B945805C9D513536FE0AFC43FD09FB95" box="[506,764,1004,1029]" italics="true" pageId="8" pageNumber="53">Platymantis adiastola</emphasis>
</taxonomicName>
produces a call composed of a series of shrill, high frequency (mean peak dominant frequency 3.0–3.2 kHz), complex, tonal notes whereas 
<taxonomicName id="4C3127CD9D513536FE80FB93FE05FBC5" box="[368,496,1084,1109]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B945805C9D513536FE80FB93FE05FBC5" box="[368,496,1084,1109]" italics="true" pageId="8" pageNumber="53">P. schmidti</emphasis>
</taxonomicName>
calls with a simple train of twin-pulse lower frequency (2.4–2.6 kHz) notes lacking tonal elements. 
<taxonomicName id="4C3127CD9D513536FD98FBCBFC9FFBED" box="[616,874,1124,1149]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="adiastola">
<emphasis id="B945805C9D513536FD98FBCBFC9FFBED" box="[616,874,1124,1149]" italics="true" pageId="8" pageNumber="53">Platymantis adiastola</emphasis>
</taxonomicName>
has a lower mean note repetition rate (4.3 notes/s) than 
<emphasis id="B945805C9D513536FE2BFB23FDA2FB35" box="[475,599,1164,1189]" italics="true" pageId="8" pageNumber="53">P. s c h m i d t i</emphasis>
(12.8 notes/s) and produces fewer mean notes per call (mean = 16.6) than 
<emphasis id="B945805C9D513536FE42FB1BFDC0FB5D" box="[434,565,1204,1229]" italics="true" pageId="8" pageNumber="53">P. s c h m i d t i</emphasis>
(mean = 23.9). Finally, the call of 
<taxonomicName id="4C3127CD9D513536FBE4FB1BFB50FB5D" box="[1044,1189,1204,1229]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="53" phylum="Chordata" rank="species" species="adiastola">
<emphasis id="B945805C9D513536FBE4FB1BFB50FB5D" box="[1044,1189,1204,1229]" italics="true" pageId="8" pageNumber="53">P. adiastola</emphasis>
</taxonomicName>
exhibits a progressively decreasing internote interval across the call, whereas 
<emphasis id="B945805C9D513536FBA4FB73FB22FB65" box="[1108,1239,1244,1269]" italics="true" pageId="8" pageNumber="53">P. s c h m i d t i</emphasis>
has a uniform internote interval throughout the entire call.
</paragraph>
</subSubSection>
<subSubSection id="C32B0FC59D513530FEC8FA83FB13FCAE" lastPageId="14" lastPageNumber="59" pageId="8" pageNumber="53" type="description">
<paragraph id="8B8E5C4E9D513534FEC8FA83FC3BF973" blockId="8.[264,1323,283,1870]" lastBlockId="10.[264,1322,1417,1763]" lastPageId="10" lastPageNumber="55" pageId="8" pageNumber="53">
<emphasis id="B945805C9D513536FEC8FA83FD83FAD6" bold="true" box="[312,630,1324,1350]" pageId="8" pageNumber="53">
<emphasis id="B945805C9D513536FEC8FA83FD99FAD6" bold="true" box="[312,620,1324,1350]" italics="true" pageId="8" pageNumber="53">
Description of 
<typeStatus id="548AE2EC9D513536FDF5FA83FD99FAD6" box="[517,620,1324,1350]" pageId="8" pageNumber="53" type="holotype">holotype</typeStatus>
</emphasis>
:
</emphasis>
A mature, undissected male specimen, in excellent condition; habitus moderately stocky; head distinct, slightly wider in dorsal aspect than body, length 42.6% of SVL; head length 114.2% of head width; snout short, its tip bluntly rounded, protruding slightly beyond lower jaw; snout posteroventrally slanted in lateral aspect; eyes protrude laterally beyond silhouette of head in dorsal aspect, and strongly beyond dorsal surface of head in lateral aspect; labial region flat and smooth, not flared, lips not swollen, not extending past eyes when viewed in dorsal aspect; interorbital region flat; eye diameter 143.4% interorbital distance; pupil horizontally ovoid; canthus rostralis slightly concave; loreal region gently sloping, slightly concave; eye diameter 73.0% of snout length, 126.6% of eye–narial distance; nostrils slightly laterally protuberant; eye narial distance much longer than distance from nostril to tip of snout; internarial region flat; tympanum distinct, its diameter 56.9% of eye diameter; dorsal edge of tympanic annulus in contact with and slightly overlapped by supratympanic fold, the latter extending from dorsoposterior corner of eye, along dorsal edge of tympanum, and 
<emphasis id="B945805C9D503537FE76FE68FAEDFE4C" box="[390,1304,455,476]" italics="true" pageId="9" pageNumber="54">
<taxonomicName id="4C3127CD9D503537FE76FE68FDADFE4C" box="[390,600,455,476]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="adiastola">Platymantis adiastola</taxonomicName>
<taxonomicName id="4C3127CD9D503537FD17FE68FC45FE4C" box="[743,944,455,476]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="schmidti">Platymantis schmidti</taxonomicName>
<taxonomicName id="4C3127CD9D503537FBBFFE68FAEDFE4C" box="[1103,1304,455,476]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="schmidti" status="sp. nov.">Platymantis schmidti</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A2763D279D503537FE76FE4BFE4EFE69" box="[390,443,484,505]" pageId="9" pageNumber="54" rank="species">n. sp.</taxonomicNameLabel>
(original collections) (recent collections) terminating nearly at supraaxillary region; tongue subcircular, with deep posterior notch and narrow anterior attachment; choanae round, minute, at anterolateral edge of palate, widely separated by a distance eight to ten times greater than their diameter, not obscured by palatal shelf; dentigerous process of vomer oblong; vomerine teeth tiny, translucent, numbering four to six; dentigerous process very slightly angled anterolaterally, with closest (posterior) points separated by a distance two times the diameter of one choana, their most distant (anterior) ends separated by distance equal to three times diameter of choanae; openings to vocal sac short slits, slightly posterior to rictus; prominent postrictal tubercle cluster present at posteroventral edge of tympanum.
</paragraph>
<caption id="DF4E0CC69D503537FEF8FEB4FE6AFE34" ID-Table-UUID="DF4E0CC69D503537FEF8FEB4FE6AFE34" httpUri="http://table.plazi.org/id/DF4E0CC69D503537FEF8FEB4FE6AFE34" pageId="9" pageNumber="54" targetBox="[272,1304,524,1892]" targetIsTable="true" targetPageId="9">
<paragraph id="8B8E5C4E9D503537FEF8FEB4FE6AFE34" blockId="9.[264,1323,283,420]" pageId="9" pageNumber="54">
<emphasis id="B945805C9D503537FEF8FEB4FE88FEA3" bold="true" box="[264,381,283,307]" pageId="9" pageNumber="54">TABLE 1.</emphasis>
Morphometric variation (in mm) in adult 
<taxonomicName id="4C3127CD9D503537FCBDFEB3FBCDFEA3" box="[845,1080,284,307]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="adiastola" status="sp. nov.">
<emphasis id="B945805C9D503537FCBDFEB3FBCDFEA3" box="[845,1080,284,307]" italics="true" pageId="9" pageNumber="54">Platymantis adiastola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2763D279D503537FBB1FEB3FB89FEA3" box="[1089,1148,284,307]" pageId="9" pageNumber="54" rank="species">n. sp.</taxonomicNameLabel>
and 
<emphasis id="B945805C9D503537FB44FEB3FAD2FEA3" box="[1204,1319,284,307]" italics="true" pageId="9" pageNumber="54">P. s c h m i d t i</emphasis>
(original collections, holotype and paratypes, as well as recent collections). Table entries include mean ± 1 standard deviation (with range shown below), and sample size (
<emphasis id="B945805C9D503537FBD1FEC8FBDBFEEE" box="[1057,1070,359,382]" italics="true" pageId="9" pageNumber="54">n</emphasis>
). See text for character abbreviations.
</paragraph>
</caption>
<paragraph id="8B8E5C4E9D503537FE76FDA3FB62F8F4" pageId="9" pageNumber="54">
<table id="F931AEEE9D50CAC1FEE0FDA3FAEDF8F4" box="[272,1304,524,1892]" gridcols="8" gridrows="20" pageId="9" pageNumber="54">
<tr id="35015E0C9D50CAC1FEE0FDA3FAEDFDAE" box="[272,1304,524,574]" gridrow="0" pageId="9" pageNumber="54" rowspan-0="1">
<th id="76D037709D50CAC1FE76FDA3FE2AFDAE" box="[390,479,524,574]" gridcol="1" gridrow="0" pageId="9" pageNumber="54">Holotype</th>
<th id="76D037709D50CAC1FE09FDA3FDAFFDAE" box="[505,602,524,574]" gridcol="2" gridrow="0" pageId="9" pageNumber="54">Male (n=10)</th>
<th id="76D037709D50CAC1FD85FDA3FD49FDAE" box="[629,700,524,574]" gridcol="3" gridrow="0" pageId="9" pageNumber="54">Female (n=1)</th>
<th id="76D037709D50CAC1FD17FDA3FCB5FDAE" box="[743,832,524,574]" gridcol="4" gridrow="0" pageId="9" pageNumber="54">Holotype</th>
<th id="76D037709D50CAC1FCA6FDA3FC42FDAE" box="[854,951,524,574]" gridcol="5" gridrow="0" pageId="9" pageNumber="54">Male (n=8)</th>
<th id="76D037709D50CAC1FC21FDA3FBC7FDAE" box="[977,1074,524,574]" gridcol="6" gridrow="0" pageId="9" pageNumber="54">Female (n=8)</th>
<th id="76D037709D50CAC1FBBFFDA3FAEDFDAE" box="[1103,1304,524,574]" gridcol="7" gridrow="0" pageId="9" pageNumber="54">Male (n=7)</th>
</tr>
<tr id="35015E0C9D50CAC1FEE0FDFEFAEDFD14" box="[272,1304,593,644]" gridrow="1" pageId="9" pageNumber="54">
<th id="76D037709D50CAC1FEE0FDFEFE84FD14" box="[272,369,593,644]" gridcol="0" gridrow="1" pageId="9" pageNumber="54">SVL</th>
<td id="76D037709D50CAC1FE76FDFEFE2AFD14" box="[390,479,593,644]" gridcol="1" gridrow="1" pageId="9" pageNumber="54">41.9</td>
<td id="76D037709D50CAC1FE09FDFEFDAFFD14" box="[505,602,593,644]" gridcol="2" gridrow="1" pageId="9" pageNumber="54">41 ± 1.4 39.4–43.9</td>
<td id="76D037709D50CAC1FD85FDFEFD49FD14" box="[629,700,593,644]" gridcol="3" gridrow="1" pageId="9" pageNumber="54">71.7</td>
<td id="76D037709D50CAC1FD17FDFEFCB5FD14" box="[743,832,593,644]" gridcol="4" gridrow="1" pageId="9" pageNumber="54">55.9</td>
<td id="76D037709D50CAC1FCA6FDFEFC42FD14" box="[854,951,593,644]" gridcol="5" gridrow="1" pageId="9" pageNumber="54">34.4 ± 2.1 30.5–36.4</td>
<td id="76D037709D50CAC1FC21FDFEFBC7FD14" box="[977,1074,593,644]" gridcol="6" gridrow="1" pageId="9" pageNumber="54">50.2 ± 5.8 41.1–57.6</td>
<td id="76D037709D50CAC1FBBFFDFEFAEDFD14" box="[1103,1304,593,644]" gridcol="7" gridrow="1" pageId="9" pageNumber="54">39.7 ± 2.3 35.6–42.6</td>
</tr>
<tr id="35015E0C9D50CAC1FEE0FD38FAEDFD59" box="[272,1304,663,713]" gridrow="2" pageId="9" pageNumber="54">
<th id="76D037709D50CAC1FEE0FD38FE84FD59" box="[272,369,663,713]" gridcol="0" gridrow="2" pageId="9" pageNumber="54">ED</th>
<td id="76D037709D50CAC1FE76FD38FE2AFD59" box="[390,479,663,713]" gridcol="1" gridrow="2" pageId="9" pageNumber="54">6.0</td>
<td id="76D037709D50CAC1FE09FD38FDAFFD59" box="[505,602,663,713]" gridcol="2" gridrow="2" pageId="9" pageNumber="54">5.6 ± 0.4 5–6.3</td>
<td id="76D037709D50CAC1FD85FD38FD49FD59" box="[629,700,663,713]" gridcol="3" gridrow="2" pageId="9" pageNumber="54">7.5</td>
<td id="76D037709D50CAC1FD17FD38FCB5FD59" box="[743,832,663,713]" gridcol="4" gridrow="2" pageId="9" pageNumber="54">4.8</td>
<td id="76D037709D50CAC1FCA6FD38FC42FD59" box="[854,951,663,713]" gridcol="5" gridrow="2" pageId="9" pageNumber="54">3.7 ± 0.4 3.1–4.3</td>
<td id="76D037709D50CAC1FC21FD38FBC7FD59" box="[977,1074,663,713]" gridcol="6" gridrow="2" pageId="9" pageNumber="54">5.1 ± 0.5 4.3–5.7</td>
<td id="76D037709D50CAC1FBBFFD38FAEDFD59" box="[1103,1304,663,713]" gridcol="7" gridrow="2" pageId="9" pageNumber="54">3.7 ± 0.5 3.1–4.6</td>
</tr>
<tr id="35015E0C9D50CAC1FEE0FD73FAEDFC9E" box="[272,1304,732,782]" gridrow="3" pageId="9" pageNumber="54">
<th id="76D037709D50CAC1FEE0FD73FE84FC9E" box="[272,369,732,782]" gridcol="0" gridrow="3" pageId="9" pageNumber="54">TD</th>
<td id="76D037709D50CAC1FE76FD73FE2AFC9E" box="[390,479,732,782]" gridcol="1" gridrow="3" pageId="9" pageNumber="54">3.7</td>
<td id="76D037709D50CAC1FE09FD73FDAFFC9E" box="[505,602,732,782]" gridcol="2" gridrow="3" pageId="9" pageNumber="54">3.9 ± 0.3 3.2–4.5</td>
<td id="76D037709D50CAC1FD85FD73FD49FC9E" box="[629,700,732,782]" gridcol="3" gridrow="3" pageId="9" pageNumber="54">5.7</td>
<td id="76D037709D50CAC1FD17FD73FCB5FC9E" box="[743,832,732,782]" gridcol="4" gridrow="3" pageId="9" pageNumber="54">3.8</td>
<td id="76D037709D50CAC1FCA6FD73FC42FC9E" box="[854,951,732,782]" gridcol="5" gridrow="3" pageId="9" pageNumber="54">3.2 ± 0.2 2.9–3.5</td>
<td id="76D037709D50CAC1FC21FD73FBC7FC9E" box="[977,1074,732,782]" gridcol="6" gridrow="3" pageId="9" pageNumber="54">4 ± 0.3 3.6–4.4</td>
<td id="76D037709D50CAC1FBBFFD73FAEDFC9E" box="[1103,1304,732,782]" gridcol="7" gridrow="3" pageId="9" pageNumber="54">2.9 ± 0.2 2.7–3.2</td>
</tr>
<tr id="35015E0C9D50CAC1FEE0FC8EFAEDFCC4" box="[272,1304,801,852]" gridrow="4" pageId="9" pageNumber="54">
<th id="76D037709D50CAC1FEE0FC8EFE84FCC4" box="[272,369,801,852]" gridcol="0" gridrow="4" pageId="9" pageNumber="54">HL</th>
<td id="76D037709D50CAC1FE76FC8EFE2AFCC4" box="[390,479,801,852]" gridcol="1" gridrow="4" pageId="9" pageNumber="54">17.6</td>
<td id="76D037709D50CAC1FE09FC8EFDAFFCC4" box="[505,602,801,852]" gridcol="2" gridrow="4" pageId="9" pageNumber="54">16.9 ± 0.6 16–17.8</td>
<td id="76D037709D50CAC1FD85FC8EFD49FCC4" box="[629,700,801,852]" gridcol="3" gridrow="4" pageId="9" pageNumber="54">27.7</td>
<td id="76D037709D50CAC1FD17FC8EFCB5FCC4" box="[743,832,801,852]" gridcol="4" gridrow="4" pageId="9" pageNumber="54">20.6</td>
<td id="76D037709D50CAC1FCA6FC8EFC42FCC4" box="[854,951,801,852]" gridcol="5" gridrow="4" pageId="9" pageNumber="54">14.3 ± 0.8 13–15.4</td>
<td id="76D037709D50CAC1FC21FC8EFBC7FCC4" box="[977,1074,801,852]" gridcol="6" gridrow="4" pageId="9" pageNumber="54">20 ± 2 16.4–23</td>
<td id="76D037709D50CAC1FBBFFC8EFAEDFCC4" box="[1103,1304,801,852]" gridcol="7" gridrow="4" pageId="9" pageNumber="54">15.9 ± 1.3 14.2–18.1</td>
</tr>
<tr id="35015E0C9D50CAC1FEE0FCC8FAEDFC09" box="[272,1304,871,921]" gridrow="5" pageId="9" pageNumber="54">
<th id="76D037709D50CAC1FEE0FCC8FE84FC09" box="[272,369,871,921]" gridcol="0" gridrow="5" pageId="9" pageNumber="54">SNL</th>
<td id="76D037709D50CAC1FE76FCC8FE2AFC09" box="[390,479,871,921]" gridcol="1" gridrow="5" pageId="9" pageNumber="54">6.9</td>
<td id="76D037709D50CAC1FE09FCC8FDAFFC09" box="[505,602,871,921]" gridcol="2" gridrow="5" pageId="9" pageNumber="54">7.5 ± 0.4 6.9–8.2</td>
<td id="76D037709D50CAC1FD85FCC8FD49FC09" box="[629,700,871,921]" gridcol="3" gridrow="5" pageId="9" pageNumber="54">12.5</td>
<td id="76D037709D50CAC1FD17FCC8FCB5FC09" box="[743,832,871,921]" gridcol="4" gridrow="5" pageId="9" pageNumber="54">8.9</td>
<td id="76D037709D50CAC1FCA6FCC8FC42FC09" box="[854,951,871,921]" gridcol="5" gridrow="5" pageId="9" pageNumber="54">6.6 ± 0.5 5.9–7.5</td>
<td id="76D037709D50CAC1FC21FCC8FBC7FC09" box="[977,1074,871,921]" gridcol="6" gridrow="5" pageId="9" pageNumber="54">9.1 ± 1.1 7.4–10.4</td>
<td id="76D037709D50CAC1FBBFFCC8FAEDFC09" box="[1103,1304,871,921]" gridcol="7" gridrow="5" pageId="9" pageNumber="54">7.0 ± 0.7 6.0–7.9</td>
</tr>
<tr id="35015E0C9D50CAC1FEE0FC03FAEDFC4E" box="[272,1304,940,990]" gridrow="6" pageId="9" pageNumber="54">
<th id="76D037709D50CAC1FEE0FC03FE84FC4E" box="[272,369,940,990]" gridcol="0" gridrow="6" pageId="9" pageNumber="54">IOD</th>
<td id="76D037709D50CAC1FE76FC03FE2AFC4E" box="[390,479,940,990]" gridcol="1" gridrow="6" pageId="9" pageNumber="54">4.7</td>
<td id="76D037709D50CAC1FE09FC03FDAFFC4E" box="[505,602,940,990]" gridcol="2" gridrow="6" pageId="9" pageNumber="54">4.1 ± 0.2 3.9–4.7</td>
<td id="76D037709D50CAC1FD85FC03FD49FC4E" box="[629,700,940,990]" gridcol="3" gridrow="6" pageId="9" pageNumber="54">7.4</td>
<td id="76D037709D50CAC1FD17FC03FCB5FC4E" box="[743,832,940,990]" gridcol="4" gridrow="6" pageId="9" pageNumber="54">4.1</td>
<td id="76D037709D50CAC1FCA6FC03FC42FC4E" box="[854,951,940,990]" gridcol="5" gridrow="6" pageId="9" pageNumber="54">3.3 ± 0.2 3.0–3.5</td>
<td id="76D037709D50CAC1FC21FC03FBC7FC4E" box="[977,1074,940,990]" gridcol="6" gridrow="6" pageId="9" pageNumber="54">4.6 ± 0.6 3.8–5.4</td>
<td id="76D037709D50CAC1FBBFFC03FAEDFC4E" box="[1103,1304,940,990]" gridcol="7" gridrow="6" pageId="9" pageNumber="54">3.1 ± 0.1 3.0–3.2</td>
</tr>
<tr id="35015E0C9D50CAC1FEE0FC5EFAEDFBB4" box="[272,1304,1009,1060]" gridrow="7" pageId="9" pageNumber="54">
<th id="76D037709D50CAC1FEE0FC5EFE84FBB4" box="[272,369,1009,1060]" gridcol="0" gridrow="7" pageId="9" pageNumber="54">HW</th>
<td id="76D037709D50CAC1FE76FC5EFE2AFBB4" box="[390,479,1009,1060]" gridcol="1" gridrow="7" pageId="9" pageNumber="54">16.4</td>
<td id="76D037709D50CAC1FE09FC5EFDAFFBB4" box="[505,602,1009,1060]" gridcol="2" gridrow="7" pageId="9" pageNumber="54">15.4 ± 1.0 14.2–17.5</td>
<td id="76D037709D50CAC1FD85FC5EFD49FBB4" box="[629,700,1009,1060]" gridcol="3" gridrow="7" pageId="9" pageNumber="54">24.7</td>
<td id="76D037709D50CAC1FD17FC5EFCB5FBB4" box="[743,832,1009,1060]" gridcol="4" gridrow="7" pageId="9" pageNumber="54">19.1</td>
<td id="76D037709D50CAC1FCA6FC5EFC42FBB4" box="[854,951,1009,1060]" gridcol="5" gridrow="7" pageId="9" pageNumber="54">12.2 ± 0.7 11.4–13.5</td>
<td id="76D037709D50CAC1FC21FC5EFBC7FBB4" box="[977,1074,1009,1060]" gridcol="6" gridrow="7" pageId="9" pageNumber="54">18.5 ± 2.5 14–22.1</td>
<td id="76D037709D50CAC1FBBFFC5EFAEDFBB4" box="[1103,1304,1009,1060]" gridcol="7" gridrow="7" pageId="9" pageNumber="54">13.9 ± 1 12.3–15.1</td>
</tr>
<tr id="35015E0C9D50CAC1FEE0FB98FAEDFBF9" box="[272,1304,1079,1129]" gridrow="8" pageId="9" pageNumber="54">
<th id="76D037709D50CAC1FEE0FB98FE84FBF9" box="[272,369,1079,1129]" gridcol="0" gridrow="8" pageId="9" pageNumber="54">FA</th>
<td id="76D037709D50CAC1FE76FB98FE2AFBF9" box="[390,479,1079,1129]" gridcol="1" gridrow="8" pageId="9" pageNumber="54">10.4</td>
<td id="76D037709D50CAC1FE09FB98FDAFFBF9" box="[505,602,1079,1129]" gridcol="2" gridrow="8" pageId="9" pageNumber="54">10.1 ± 0.7 8.6–10.8</td>
<td id="76D037709D50CAC1FD85FB98FD49FBF9" box="[629,700,1079,1129]" gridcol="3" gridrow="8" pageId="9" pageNumber="54">16.3</td>
<td id="76D037709D50CAC1FD17FB98FCB5FBF9" box="[743,832,1079,1129]" gridcol="4" gridrow="8" pageId="9" pageNumber="54">10.2</td>
<td id="76D037709D50CAC1FCA6FB98FC42FBF9" box="[854,951,1079,1129]" gridcol="5" gridrow="8" pageId="9" pageNumber="54">7.3 ± 0.9 6.0–8.8</td>
<td id="76D037709D50CAC1FC21FB98FBC7FBF9" box="[977,1074,1079,1129]" gridcol="6" gridrow="8" pageId="9" pageNumber="54">11.1 ± 1.9 8.9–14.2</td>
<td id="76D037709D50CAC1FBBFFB98FAEDFBF9" box="[1103,1304,1079,1129]" gridcol="7" gridrow="8" pageId="9" pageNumber="54">7.3 ± 0.9 6.4–8.8</td>
</tr>
<tr id="35015E0C9D50CAC1FEE0FBD3FAEDFB3E" box="[272,1304,1148,1198]" gridrow="9" pageId="9" pageNumber="54">
<th id="76D037709D50CAC1FEE0FBD3FE84FB3E" box="[272,369,1148,1198]" gridcol="0" gridrow="9" pageId="9" pageNumber="54">TBL</th>
<td id="76D037709D50CAC1FE76FBD3FE2AFB3E" box="[390,479,1148,1198]" gridcol="1" gridrow="9" pageId="9" pageNumber="54">22.5</td>
<td id="76D037709D50CAC1FE09FBD3FDAFFB3E" box="[505,602,1148,1198]" gridcol="2" gridrow="9" pageId="9" pageNumber="54">23.8 ± 0.8 22.5–24.9</td>
<td id="76D037709D50CAC1FD85FBD3FD49FB3E" box="[629,700,1148,1198]" gridcol="3" gridrow="9" pageId="9" pageNumber="54">38.8</td>
<td id="76D037709D50CAC1FD17FBD3FCB5FB3E" box="[743,832,1148,1198]" gridcol="4" gridrow="9" pageId="9" pageNumber="54">33.6</td>
<td id="76D037709D50CAC1FCA6FBD3FC42FB3E" box="[854,951,1148,1198]" gridcol="5" gridrow="9" pageId="9" pageNumber="54">19.4 ± 0.7 18.6–20.5</td>
<td id="76D037709D50CAC1FC21FBD3FBC7FB3E" box="[977,1074,1148,1198]" gridcol="6" gridrow="9" pageId="9" pageNumber="54">28.6 ± 3 25.1–33.6</td>
<td id="76D037709D50CAC1FBBFFBD3FAEDFB3E" box="[1103,1304,1148,1198]" gridcol="7" gridrow="9" pageId="9" pageNumber="54">23.2 ± 1.2 21.3–24.3</td>
</tr>
<tr id="35015E0C9D50CAC1FEE0FB6EFAEDFB64" box="[272,1304,1217,1268]" gridrow="10" pageId="9" pageNumber="54">
<th id="76D037709D50CAC1FEE0FB6EFE84FB64" box="[272,369,1217,1268]" gridcol="0" gridrow="10" pageId="9" pageNumber="54">TSL</th>
<td id="76D037709D50CAC1FE76FB6EFE2AFB64" box="[390,479,1217,1268]" gridcol="1" gridrow="10" pageId="9" pageNumber="54">13.0</td>
<td id="76D037709D50CAC1FE09FB6EFDAFFB64" box="[505,602,1217,1268]" gridcol="2" gridrow="10" pageId="9" pageNumber="54">13.4 ± 0.5 12.6–14</td>
<td id="76D037709D50CAC1FD85FB6EFD49FB64" box="[629,700,1217,1268]" gridcol="3" gridrow="10" pageId="9" pageNumber="54">21.8</td>
<td id="76D037709D50CAC1FD17FB6EFCB5FB64" box="[743,832,1217,1268]" gridcol="4" gridrow="10" pageId="9" pageNumber="54">16.4</td>
<td id="76D037709D50CAC1FCA6FB6EFC42FB64" box="[854,951,1217,1268]" gridcol="5" gridrow="10" pageId="9" pageNumber="54">10.5 ± 0.4 10.0–11.0</td>
<td id="76D037709D50CAC1FC21FB6EFBC7FB64" box="[977,1074,1217,1268]" gridcol="6" gridrow="10" pageId="9" pageNumber="54">15.4 ± 1.4 13.3–17.6</td>
<td id="76D037709D50CAC1FBBFFB6EFAEDFB64" box="[1103,1304,1217,1268]" gridcol="7" gridrow="10" pageId="9" pageNumber="54">11.2 ± 0.8 10.3–12.4</td>
</tr>
<tr id="35015E0C9D50CAC1FEE0FAA8FAEDFAA9" box="[272,1304,1287,1337]" gridrow="11" pageId="9" pageNumber="54">
<th id="76D037709D50CAC1FEE0FAA8FE84FAA9" box="[272,369,1287,1337]" gridcol="0" gridrow="11" pageId="9" pageNumber="54">PL</th>
<td id="76D037709D50CAC1FE76FAA8FE2AFAA9" box="[390,479,1287,1337]" gridcol="1" gridrow="11" pageId="9" pageNumber="54">22.2</td>
<td id="76D037709D50CAC1FE09FAA8FDAFFAA9" box="[505,602,1287,1337]" gridcol="2" gridrow="11" pageId="9" pageNumber="54">22.8 ± 0.6 22–23.9</td>
<td id="76D037709D50CAC1FD85FAA8FD49FAA9" box="[629,700,1287,1337]" gridcol="3" gridrow="11" pageId="9" pageNumber="54">36.5</td>
<td id="76D037709D50CAC1FD17FAA8FCB5FAA9" box="[743,832,1287,1337]" gridcol="4" gridrow="11" pageId="9" pageNumber="54">29.2</td>
<td id="76D037709D50CAC1FCA6FAA8FC42FAA9" box="[854,951,1287,1337]" gridcol="5" gridrow="11" pageId="9" pageNumber="54">17.9 ± 0.6 17.1–19.1</td>
<td id="76D037709D50CAC1FC21FAA8FBC7FAA9" box="[977,1074,1287,1337]" gridcol="6" gridrow="11" pageId="9" pageNumber="54">25.8 ± 2.2 22.7–29.2</td>
<td id="76D037709D50CAC1FBBFFAA8FAEDFAA9" box="[1103,1304,1287,1337]" gridcol="7" gridrow="11" pageId="9" pageNumber="54">20.8 ± 1.5 18.1–22.3</td>
</tr>
<tr id="35015E0C9D50CAC1FEE0FAE3FAEDFAEE" box="[272,1304,1356,1406]" gridrow="12" pageId="9" pageNumber="54">
<th id="76D037709D50CAC1FEE0FAE3FE84FAEE" box="[272,369,1356,1406]" gridcol="0" gridrow="12" pageId="9" pageNumber="54">ML</th>
<td id="76D037709D50CAC1FE76FAE3FE2AFAEE" box="[390,479,1356,1406]" gridcol="1" gridrow="12" pageId="9" pageNumber="54">11.0</td>
<td id="76D037709D50CAC1FE09FAE3FDAFFAEE" box="[505,602,1356,1406]" gridcol="2" gridrow="12" pageId="9" pageNumber="54">10.7 ± 0.5 10.0–11.7</td>
<td id="76D037709D50CAC1FD85FAE3FD49FAEE" box="[629,700,1356,1406]" gridcol="3" gridrow="12" pageId="9" pageNumber="54">17.3</td>
<td id="76D037709D50CAC1FD17FAE3FCB5FAEE" box="[743,832,1356,1406]" gridcol="4" gridrow="12" pageId="9" pageNumber="54">13.9</td>
<td id="76D037709D50CAC1FCA6FAE3FC42FAEE" box="[854,951,1356,1406]" gridcol="5" gridrow="12" pageId="9" pageNumber="54">8.6 ± 0.3 8.1–9.1</td>
<td id="76D037709D50CAC1FC21FAE3FBC7FAEE" box="[977,1074,1356,1406]" gridcol="6" gridrow="12" pageId="9" pageNumber="54">12.5 ± 1.2 10.6–13.9</td>
<td id="76D037709D50CAC1FBBFFAE3FAEDFAEE" box="[1103,1304,1356,1406]" gridcol="7" gridrow="12" pageId="9" pageNumber="54">9.9 ± 1.3 8.1–11.2</td>
</tr>
<tr id="35015E0C9D50CAC1FEE0FA3EFAEDFA54" box="[272,1304,1425,1476]" gridrow="13" pageId="9" pageNumber="54">
<th id="76D037709D50CAC1FEE0FA3EFE84FA54" box="[272,369,1425,1476]" gridcol="0" gridrow="13" pageId="9" pageNumber="54">TL</th>
<td id="76D037709D50CAC1FE76FA3EFE2AFA54" box="[390,479,1425,1476]" gridcol="1" gridrow="13" pageId="9" pageNumber="54">14.8</td>
<td id="76D037709D50CAC1FE09FA3EFDAFFA54" box="[505,602,1425,1476]" gridcol="2" gridrow="13" pageId="9" pageNumber="54">16 ± 0.9 14.8–17.3</td>
<td id="76D037709D50CAC1FD85FA3EFD49FA54" box="[629,700,1425,1476]" gridcol="3" gridrow="13" pageId="9" pageNumber="54">30.0</td>
<td id="76D037709D50CAC1FD17FA3EFCB5FA54" box="[743,832,1425,1476]" gridcol="4" gridrow="13" pageId="9" pageNumber="54">22.6</td>
<td id="76D037709D50CAC1FCA6FA3EFC42FA54" box="[854,951,1425,1476]" gridcol="5" gridrow="13" pageId="9" pageNumber="54">13.6 ± 1.0 12.5–15.4</td>
<td id="76D037709D50CAC1FC21FA3EFBC7FA54" box="[977,1074,1425,1476]" gridcol="6" gridrow="13" pageId="9" pageNumber="54">19.8 ± 2.5 15.8–23.4</td>
<td id="76D037709D50CAC1FBBFFA3EFAEDFA54" box="[1103,1304,1425,1476]" gridcol="7" gridrow="13" pageId="9" pageNumber="54">15.6 ± 2.2 12.4–17.7</td>
</tr>
<tr id="35015E0C9D50CAC1FEE0FA78FAEDF999" box="[272,1304,1495,1545]" gridrow="14" pageId="9" pageNumber="54">
<th id="76D037709D50CAC1FEE0FA78FE84F999" box="[272,369,1495,1545]" gridcol="0" gridrow="14" pageId="9" pageNumber="54">Fin1L</th>
<td id="76D037709D50CAC1FE76FA78FE2AF999" box="[390,479,1495,1545]" gridcol="1" gridrow="14" pageId="9" pageNumber="54">6.1</td>
<td id="76D037709D50CAC1FE09FA78FDAFF999" box="[505,602,1495,1545]" gridcol="2" gridrow="14" pageId="9" pageNumber="54">6 ± 0.5 5.3–6.6</td>
<td id="76D037709D50CAC1FD85FA78FD49F999" box="[629,700,1495,1545]" gridcol="3" gridrow="14" pageId="9" pageNumber="54">10.4</td>
<td id="76D037709D50CAC1FD17FA78FCB5F999" box="[743,832,1495,1545]" gridcol="4" gridrow="14" pageId="9" pageNumber="54">7.4</td>
<td id="76D037709D50CAC1FCA6FA78FC42F999" box="[854,951,1495,1545]" gridcol="5" gridrow="14" pageId="9" pageNumber="54">4.6 ± 0.3 4.2–5.1</td>
<td id="76D037709D50CAC1FC21FA78FBC7F999" box="[977,1074,1495,1545]" gridcol="6" gridrow="14" pageId="9" pageNumber="54">7.4 ± 0.7 6.5–8.5</td>
<td id="76D037709D50CAC1FBBFFA78FAEDF999" box="[1103,1304,1495,1545]" gridcol="7" gridrow="14" pageId="9" pageNumber="54">5.7 ± 0.7 4.4–6.2</td>
</tr>
<tr id="35015E0C9D50CAC1FEE0F9B3FAEDF9DE" box="[272,1304,1564,1614]" gridrow="15" pageId="9" pageNumber="54">
<th id="76D037709D50CAC1FEE0F9B3FE84F9DE" box="[272,369,1564,1614]" gridcol="0" gridrow="15" pageId="9" pageNumber="54">Fin3L</th>
<td id="76D037709D50CAC1FE76F9B3FE2AF9DE" box="[390,479,1564,1614]" gridcol="1" gridrow="15" pageId="9" pageNumber="54">7.6</td>
<td id="76D037709D50CAC1FE09F9B3FDAFF9DE" box="[505,602,1564,1614]" gridcol="2" gridrow="15" pageId="9" pageNumber="54">7.4 ± 0.4 6.9–8.3</td>
<td id="76D037709D50CAC1FD85F9B3FD49F9DE" box="[629,700,1564,1614]" gridcol="3" gridrow="15" pageId="9" pageNumber="54">11.4</td>
<td id="76D037709D50CAC1FD17F9B3FCB5F9DE" box="[743,832,1564,1614]" gridcol="4" gridrow="15" pageId="9" pageNumber="54">9.4</td>
<td id="76D037709D50CAC1FCA6F9B3FC42F9DE" box="[854,951,1564,1614]" gridcol="5" gridrow="15" pageId="9" pageNumber="54">5.9 ± 0.3 5.7–6.4</td>
<td id="76D037709D50CAC1FC21F9B3FBC7F9DE" box="[977,1074,1564,1614]" gridcol="6" gridrow="15" pageId="9" pageNumber="54">8.5 ± 0.9 7.2–9.9</td>
<td id="76D037709D50CAC1FBBFF9B3FAEDF9DE" box="[1103,1304,1564,1614]" gridcol="7" gridrow="15" pageId="9" pageNumber="54">7.1 ± 0.9 5.9–8.1</td>
</tr>
<tr id="35015E0C9D50CAC1FEE0F9CEFAEDF904" box="[272,1304,1633,1684]" gridrow="16" pageId="9" pageNumber="54">
<th id="76D037709D50CAC1FEE0F9CEFE84F904" box="[272,369,1633,1684]" gridcol="0" gridrow="16" pageId="9" pageNumber="54">Fin3DW</th>
<td id="76D037709D50CAC1FE76F9CEFE2AF904" box="[390,479,1633,1684]" gridcol="1" gridrow="16" pageId="9" pageNumber="54">1.5</td>
<td id="76D037709D50CAC1FE09F9CEFDAFF904" box="[505,602,1633,1684]" gridcol="2" gridrow="16" pageId="9" pageNumber="54">1.5 ± 0.1 1.3–1.8</td>
<td id="76D037709D50CAC1FD85F9CEFD49F904" box="[629,700,1633,1684]" gridcol="3" gridrow="16" pageId="9" pageNumber="54">1.9</td>
<td id="76D037709D50CAC1FD17F9CEFCB5F904" box="[743,832,1633,1684]" gridcol="4" gridrow="16" pageId="9" pageNumber="54">1.4</td>
<td id="76D037709D50CAC1FCA6F9CEFC42F904" box="[854,951,1633,1684]" gridcol="5" gridrow="16" pageId="9" pageNumber="54">1.0 ± 0.1 0.9–1.1</td>
<td id="76D037709D50CAC1FC21F9CEFBC7F904" box="[977,1074,1633,1684]" gridcol="6" gridrow="16" pageId="9" pageNumber="54">1.2 ± 0.1 1.1–1.4</td>
<td id="76D037709D50CAC1FBBFF9CEFAEDF904" box="[1103,1304,1633,1684]" gridcol="7" gridrow="16" pageId="9" pageNumber="54">1.1 ± 0.1 1.0–1.3</td>
</tr>
<tr id="35015E0C9D50CAC1FEE0F908FAEDF949" box="[272,1304,1703,1753]" gridrow="17" pageId="9" pageNumber="54">
<th id="76D037709D50CAC1FEE0F908FE84F949" box="[272,369,1703,1753]" gridcol="0" gridrow="17" pageId="9" pageNumber="54">Fin3PPW</th>
<td id="76D037709D50CAC1FE76F908FE2AF949" box="[390,479,1703,1753]" gridcol="1" gridrow="17" pageId="9" pageNumber="54">0.73</td>
<td id="76D037709D50CAC1FE09F908FDAFF949" box="[505,602,1703,1753]" gridcol="2" gridrow="17" pageId="9" pageNumber="54">0.6 ± 0.1 0.6–0.8</td>
<td id="76D037709D50CAC1FD85F908FD49F949" box="[629,700,1703,1753]" gridcol="3" gridrow="17" pageId="9" pageNumber="54">1.07</td>
<td id="76D037709D50CAC1FD17F908FCB5F949" box="[743,832,1703,1753]" gridcol="4" gridrow="17" pageId="9" pageNumber="54">1.01</td>
<td id="76D037709D50CAC1FCA6F908FC42F949" box="[854,951,1703,1753]" gridcol="5" gridrow="17" pageId="9" pageNumber="54">0.7 ± 0.1 0.6–0.9</td>
<td id="76D037709D50CAC1FC21F908FBC7F949" box="[977,1074,1703,1753]" gridcol="6" gridrow="17" pageId="9" pageNumber="54">0.9 ± 0.1 0.8–1</td>
<td id="76D037709D50CAC1FBBFF908FAEDF949" box="[1103,1304,1703,1753]" gridcol="7" gridrow="17" pageId="9" pageNumber="54">0.9 ± 0.1 0.7–1.0</td>
</tr>
<tr id="35015E0C9D50CAC1FEE0F943FAEDF88E" box="[272,1304,1772,1822]" gridrow="18" pageId="9" pageNumber="54">
<th id="76D037709D50CAC1FEE0F943FE84F88E" box="[272,369,1772,1822]" gridcol="0" gridrow="18" pageId="9" pageNumber="54">Toe4DW</th>
<td id="76D037709D50CAC1FE76F943FE2AF88E" box="[390,479,1772,1822]" gridcol="1" gridrow="18" pageId="9" pageNumber="54">1.46</td>
<td id="76D037709D50CAC1FE09F943FDAFF88E" box="[505,602,1772,1822]" gridcol="2" gridrow="18" pageId="9" pageNumber="54">1.6 ± 0.2 1.3–1.8</td>
<td id="76D037709D50CAC1FD85F943FD49F88E" box="[629,700,1772,1822]" gridcol="3" gridrow="18" pageId="9" pageNumber="54">2.27</td>
<td id="76D037709D50CAC1FD17F943FCB5F88E" box="[743,832,1772,1822]" gridcol="4" gridrow="18" pageId="9" pageNumber="54">1.53</td>
<td id="76D037709D50CAC1FCA6F943FC42F88E" box="[854,951,1772,1822]" gridcol="5" gridrow="18" pageId="9" pageNumber="54">1.1 ± 0.1 1–1.2</td>
<td id="76D037709D50CAC1FC21F943FBC7F88E" box="[977,1074,1772,1822]" gridcol="6" gridrow="18" pageId="9" pageNumber="54">1.3 ± 0.2 1.1–1.5</td>
<td id="76D037709D50CAC1FBBFF943FAEDF88E" box="[1103,1304,1772,1822]" gridcol="7" gridrow="18" pageId="9" pageNumber="54">1.2 ± 0.2 1.0–1.5</td>
</tr>
<tr id="35015E0C9D50CAC1FEE0F89EFAEDF8F4" box="[272,1304,1841,1892]" gridrow="19" pageId="9" pageNumber="54">
<th id="76D037709D50CAC1FEE0F89EFE84F8F4" box="[272,369,1841,1892]" gridcol="0" gridrow="19" pageId="9" pageNumber="54">Toe4PPW</th>
<td id="76D037709D50CAC1FE76F89EFE2AF8F4" box="[390,479,1841,1892]" gridcol="1" gridrow="19" pageId="9" pageNumber="54">0.63</td>
<td id="76D037709D50CAC1FE09F89EFDAFF8F4" box="[505,602,1841,1892]" gridcol="2" gridrow="19" pageId="9" pageNumber="54">0.6 ± 0.1 0.5–0.7</td>
<td id="76D037709D50CAC1FD85F89EFD49F8F4" box="[629,700,1841,1892]" gridcol="3" gridrow="19" pageId="9" pageNumber="54">1.21</td>
<td id="76D037709D50CAC1FD17F89EFCB5F8F4" box="[743,832,1841,1892]" gridcol="4" gridrow="19" pageId="9" pageNumber="54">0.91</td>
<td id="76D037709D50CAC1FCA6F89EFC42F8F4" box="[854,951,1841,1892]" gridcol="5" gridrow="19" pageId="9" pageNumber="54">0.7 ± 0.1 0.6–0.9</td>
<td id="76D037709D50CAC1FC21F89EFBC7F8F4" box="[977,1074,1841,1892]" gridcol="6" gridrow="19" pageId="9" pageNumber="54">0.8 ± 0.1 0.7–1</td>
<td id="76D037709D50CAC1FBBFF89EFAEDF8F4" box="[1103,1304,1841,1892]" gridcol="7" gridrow="19" pageId="9" pageNumber="54">0.9 ± 0.1 0.7–1.0</td>
</tr>
</table>
</paragraph>
<caption id="DF4E0CC69D533534FEF8FEB4FD7CFEC5" ID-Table-UUID="DF4E0CC69D533534FEF8FEB4FD7CFEC5" httpUri="http://table.plazi.org/id/DF4E0CC69D533534FEF8FEB4FD7CFEC5" pageId="10" pageNumber="55" targetBox="[272,1263,374,1336]" targetIsTable="true" targetPageId="10">
<paragraph id="8B8E5C4E9D533534FEF8FEB4FD7CFEC5" blockId="10.[264,1323,283,341]" pageId="10" pageNumber="55">
<emphasis id="B945805C9D533534FEF8FEB4FE89FEA3" bold="true" box="[264,380,283,307]" pageId="10" pageNumber="55">TABLE 2.</emphasis>
Loadings for the first four principal components extracted from the correlation matrix. All variables were log transformed.
</paragraph>
</caption>
<paragraph id="8B8E5C4E9D533534FD8EFED9FB1AFAA8" pageId="10" pageNumber="55">
<table id="F931AEEE9D53CAC1FEE0FED9FB1AFAA8" box="[272,1263,374,1336]" gridcols="5" gridrows="23" pageId="10" pageNumber="55">
<tr id="35015E0C9D53CAC1FEE0FED9FB1AFE1D" box="[272,1263,374,397]" gridrow="0" pageId="10" pageNumber="55" rowspan-0="1">
<th id="76D037709D53CAC1FDABFED9FD51FE1D" box="[603,676,374,397]" gridcol="1" gridrow="0" pageId="10" pageNumber="55">I</th>
<th id="76D037709D53CAC1FCEAFED9FC96FE1D" box="[794,867,374,397]" gridcol="2" gridrow="0" pageId="10" pageNumber="55">II</th>
<th id="76D037709D53CAC1FC10FED9FBDCFE1D" box="[992,1065,374,397]" gridcol="3" gridrow="0" pageId="10" pageNumber="55">III</th>
<th id="76D037709D53CAC1FB56FED9FB1AFE1D" box="[1190,1263,374,397]" gridcol="4" gridrow="0" pageId="10" pageNumber="55">IV</th>
</tr>
<tr id="35015E0C9D53CAC1FEE0FE0EFB1AFE28" box="[272,1263,417,440]" gridrow="1" pageId="10" pageNumber="55">
<th id="76D037709D53CAC1FEE0FE0EFE5CFE28" box="[272,425,417,440]" gridcol="0" gridrow="1" pageId="10" pageNumber="55">SVL</th>
<td id="76D037709D53CAC1FDABFE0EFD51FE28" box="[603,676,417,440]" gridcol="1" gridrow="1" pageId="10" pageNumber="55">0.914</td>
<td id="76D037709D53CAC1FCEAFE0EFC96FE28" box="[794,867,417,440]" gridcol="2" gridrow="1" pageId="10" pageNumber="55">0.273</td>
<td id="76D037709D53CAC1FC10FE0EFBDCFE28" box="[992,1065,417,440]" gridcol="3" gridrow="1" pageId="10" pageNumber="55">0.063</td>
<td id="76D037709D53CAC1FB56FE0EFB1AFE28" box="[1190,1263,417,440]" gridcol="4" gridrow="1" pageId="10" pageNumber="55">-0.033</td>
</tr>
<tr id="35015E0C9D53CAC1FEE0FE63FB1AFE73" box="[272,1263,460,483]" gridrow="2" pageId="10" pageNumber="55">
<th id="76D037709D53CAC1FEE0FE63FE5CFE73" box="[272,425,460,483]" gridcol="0" gridrow="2" pageId="10" pageNumber="55">ED</th>
<td id="76D037709D53CAC1FDABFE63FD51FE73" box="[603,676,460,483]" gridcol="1" gridrow="2" pageId="10" pageNumber="55">0.864</td>
<td id="76D037709D53CAC1FCEAFE63FC96FE73" box="[794,867,460,483]" gridcol="2" gridrow="2" pageId="10" pageNumber="55">-0.402</td>
<td id="76D037709D53CAC1FC10FE63FBDCFE73" box="[992,1065,460,483]" gridcol="3" gridrow="2" pageId="10" pageNumber="55">0.116</td>
<td id="76D037709D53CAC1FB56FE63FB1AFE73" box="[1190,1263,460,483]" gridcol="4" gridrow="2" pageId="10" pageNumber="55">-0.056</td>
</tr>
<tr id="35015E0C9D53CAC1FEE0FE59FB1AFD9D" box="[272,1263,502,525]" gridrow="3" pageId="10" pageNumber="55">
<th id="76D037709D53CAC1FEE0FE59FE5CFD9D" box="[272,425,502,525]" gridcol="0" gridrow="3" pageId="10" pageNumber="55">TD</th>
<td id="76D037709D53CAC1FDABFE59FD51FD9D" box="[603,676,502,525]" gridcol="1" gridrow="3" pageId="10" pageNumber="55">0.694</td>
<td id="76D037709D53CAC1FCEAFE59FC96FD9D" box="[794,867,502,525]" gridcol="2" gridrow="3" pageId="10" pageNumber="55">-0.603</td>
<td id="76D037709D53CAC1FC10FE59FBDCFD9D" box="[992,1065,502,525]" gridcol="3" gridrow="3" pageId="10" pageNumber="55">0.117</td>
<td id="76D037709D53CAC1FB56FE59FB1AFD9D" box="[1190,1263,502,525]" gridcol="4" gridrow="3" pageId="10" pageNumber="55">0.240</td>
</tr>
<tr id="35015E0C9D53CAC1FEE0FD8EFB1AFDA8" box="[272,1263,545,568]" gridrow="4" pageId="10" pageNumber="55">
<th id="76D037709D53CAC1FEE0FD8EFE5CFDA8" box="[272,425,545,568]" gridcol="0" gridrow="4" pageId="10" pageNumber="55">HL</th>
<td id="76D037709D53CAC1FDABFD8EFD51FDA8" box="[603,676,545,568]" gridcol="1" gridrow="4" pageId="10" pageNumber="55">0.942</td>
<td id="76D037709D53CAC1FCEAFD8EFC96FDA8" box="[794,867,545,568]" gridcol="2" gridrow="4" pageId="10" pageNumber="55">0.127</td>
<td id="76D037709D53CAC1FC10FD8EFBDCFDA8" box="[992,1065,545,568]" gridcol="3" gridrow="4" pageId="10" pageNumber="55">-0.006</td>
<td id="76D037709D53CAC1FB56FD8EFB1AFDA8" box="[1190,1263,545,568]" gridcol="4" gridrow="4" pageId="10" pageNumber="55">0.035</td>
</tr>
<tr id="35015E0C9D53CAC1FEE0FDE3FB1AFDF3" box="[272,1263,588,611]" gridrow="5" pageId="10" pageNumber="55">
<th id="76D037709D53CAC1FEE0FDE3FE5CFDF3" box="[272,425,588,611]" gridcol="0" gridrow="5" pageId="10" pageNumber="55">SNL</th>
<td id="76D037709D53CAC1FDABFDE3FD51FDF3" box="[603,676,588,611]" gridcol="1" gridrow="5" pageId="10" pageNumber="55">0.836</td>
<td id="76D037709D53CAC1FCEAFDE3FC96FDF3" box="[794,867,588,611]" gridcol="2" gridrow="5" pageId="10" pageNumber="55">0.056</td>
<td id="76D037709D53CAC1FC10FDE3FBDCFDF3" box="[992,1065,588,611]" gridcol="3" gridrow="5" pageId="10" pageNumber="55">-0.194</td>
<td id="76D037709D53CAC1FB56FDE3FB1AFDF3" box="[1190,1263,588,611]" gridcol="4" gridrow="5" pageId="10" pageNumber="55">0.351</td>
</tr>
<tr id="35015E0C9D53CAC1FEE0FDD9FB1AFD1D" box="[272,1263,630,653]" gridrow="6" pageId="10" pageNumber="55">
<th id="76D037709D53CAC1FEE0FDD9FE5CFD1D" box="[272,425,630,653]" gridcol="0" gridrow="6" pageId="10" pageNumber="55">IOD</th>
<td id="76D037709D53CAC1FDABFDD9FD51FD1D" box="[603,676,630,653]" gridcol="1" gridrow="6" pageId="10" pageNumber="55">0.700</td>
<td id="76D037709D53CAC1FCEAFDD9FC96FD1D" box="[794,867,630,653]" gridcol="2" gridrow="6" pageId="10" pageNumber="55">-0.578</td>
<td id="76D037709D53CAC1FC10FDD9FBDCFD1D" box="[992,1065,630,653]" gridcol="3" gridrow="6" pageId="10" pageNumber="55">0.333</td>
<td id="76D037709D53CAC1FB56FDD9FB1AFD1D" box="[1190,1263,630,653]" gridcol="4" gridrow="6" pageId="10" pageNumber="55">-0.154</td>
</tr>
<tr id="35015E0C9D53CAC1FEE0FD0EFB1AFD28" box="[272,1263,673,696]" gridrow="7" pageId="10" pageNumber="55">
<th id="76D037709D53CAC1FEE0FD0EFE5CFD28" box="[272,425,673,696]" gridcol="0" gridrow="7" pageId="10" pageNumber="55">HW</th>
<td id="76D037709D53CAC1FDABFD0EFD51FD28" box="[603,676,673,696]" gridcol="1" gridrow="7" pageId="10" pageNumber="55">0.942</td>
<td id="76D037709D53CAC1FCEAFD0EFC96FD28" box="[794,867,673,696]" gridcol="2" gridrow="7" pageId="10" pageNumber="55">0.009</td>
<td id="76D037709D53CAC1FC10FD0EFBDCFD28" box="[992,1065,673,696]" gridcol="3" gridrow="7" pageId="10" pageNumber="55">-0.005</td>
<td id="76D037709D53CAC1FB56FD0EFB1AFD28" box="[1190,1263,673,696]" gridcol="4" gridrow="7" pageId="10" pageNumber="55">-0.157</td>
</tr>
<tr id="35015E0C9D53CAC1FEE0FD63FB1AFD73" box="[272,1263,716,739]" gridrow="8" pageId="10" pageNumber="55">
<th id="76D037709D53CAC1FEE0FD63FE5CFD73" box="[272,425,716,739]" gridcol="0" gridrow="8" pageId="10" pageNumber="55">FA</th>
<td id="76D037709D53CAC1FDABFD63FD51FD73" box="[603,676,716,739]" gridcol="1" gridrow="8" pageId="10" pageNumber="55">0.811</td>
<td id="76D037709D53CAC1FCEAFD63FC96FD73" box="[794,867,716,739]" gridcol="2" gridrow="8" pageId="10" pageNumber="55">-0.358</td>
<td id="76D037709D53CAC1FC10FD63FBDCFD73" box="[992,1065,716,739]" gridcol="3" gridrow="8" pageId="10" pageNumber="55">0.287</td>
<td id="76D037709D53CAC1FB56FD63FB1AFD73" box="[1190,1263,716,739]" gridcol="4" gridrow="8" pageId="10" pageNumber="55">0.094</td>
</tr>
<tr id="35015E0C9D53CAC1FEE0FD59FB1AFC9D" box="[272,1263,758,781]" gridrow="9" pageId="10" pageNumber="55">
<th id="76D037709D53CAC1FEE0FD59FE5CFC9D" box="[272,425,758,781]" gridcol="0" gridrow="9" pageId="10" pageNumber="55">TBL</th>
<td id="76D037709D53CAC1FDABFD59FD51FC9D" box="[603,676,758,781]" gridcol="1" gridrow="9" pageId="10" pageNumber="55">0.889</td>
<td id="76D037709D53CAC1FCEAFD59FC96FC9D" box="[794,867,758,781]" gridcol="2" gridrow="9" pageId="10" pageNumber="55">0.272</td>
<td id="76D037709D53CAC1FC10FD59FBDCFC9D" box="[992,1065,758,781]" gridcol="3" gridrow="9" pageId="10" pageNumber="55">-0.190</td>
<td id="76D037709D53CAC1FB56FD59FB1AFC9D" box="[1190,1263,758,781]" gridcol="4" gridrow="9" pageId="10" pageNumber="55">-0.122</td>
</tr>
<tr id="35015E0C9D53CAC1FEE0FC8EFB1AFCA8" box="[272,1263,801,824]" gridrow="10" pageId="10" pageNumber="55">
<th id="76D037709D53CAC1FEE0FC8EFE5CFCA8" box="[272,425,801,824]" gridcol="0" gridrow="10" pageId="10" pageNumber="55">TSL</th>
<td id="76D037709D53CAC1FDABFC8EFD51FCA8" box="[603,676,801,824]" gridcol="1" gridrow="10" pageId="10" pageNumber="55">0.945</td>
<td id="76D037709D53CAC1FCEAFC8EFC96FCA8" box="[794,867,801,824]" gridcol="2" gridrow="10" pageId="10" pageNumber="55">-0.237</td>
<td id="76D037709D53CAC1FC10FC8EFBDCFCA8" box="[992,1065,801,824]" gridcol="3" gridrow="10" pageId="10" pageNumber="55">-0.083</td>
<td id="76D037709D53CAC1FB56FC8EFB1AFCA8" box="[1190,1263,801,824]" gridcol="4" gridrow="10" pageId="10" pageNumber="55">0.009</td>
</tr>
<tr id="35015E0C9D53CAC1FEE0FCE3FB1AFCF3" box="[272,1263,844,867]" gridrow="11" pageId="10" pageNumber="55">
<th id="76D037709D53CAC1FEE0FCE3FE5CFCF3" box="[272,425,844,867]" gridcol="0" gridrow="11" pageId="10" pageNumber="55">PL</th>
<td id="76D037709D53CAC1FDABFCE3FD51FCF3" box="[603,676,844,867]" gridcol="1" gridrow="11" pageId="10" pageNumber="55">0.957</td>
<td id="76D037709D53CAC1FCEAFCE3FC96FCF3" box="[794,867,844,867]" gridcol="2" gridrow="11" pageId="10" pageNumber="55">0.088</td>
<td id="76D037709D53CAC1FC10FCE3FBDCFCF3" box="[992,1065,844,867]" gridcol="3" gridrow="11" pageId="10" pageNumber="55">-0.120</td>
<td id="76D037709D53CAC1FB56FCE3FB1AFCF3" box="[1190,1263,844,867]" gridcol="4" gridrow="11" pageId="10" pageNumber="55">-0.080</td>
</tr>
<tr id="35015E0C9D53CAC1FEE0FCD9FB1AFC1D" box="[272,1263,886,909]" gridrow="12" pageId="10" pageNumber="55">
<th id="76D037709D53CAC1FEE0FCD9FE5CFC1D" box="[272,425,886,909]" gridcol="0" gridrow="12" pageId="10" pageNumber="55">ML</th>
<td id="76D037709D53CAC1FDABFCD9FD51FC1D" box="[603,676,886,909]" gridcol="1" gridrow="12" pageId="10" pageNumber="55">0.924</td>
<td id="76D037709D53CAC1FCEAFCD9FC96FC1D" box="[794,867,886,909]" gridcol="2" gridrow="12" pageId="10" pageNumber="55">0.149</td>
<td id="76D037709D53CAC1FC10FCD9FBDCFC1D" box="[992,1065,886,909]" gridcol="3" gridrow="12" pageId="10" pageNumber="55">-0.154</td>
<td id="76D037709D53CAC1FB56FCD9FB1AFC1D" box="[1190,1263,886,909]" gridcol="4" gridrow="12" pageId="10" pageNumber="55">0.009</td>
</tr>
<tr id="35015E0C9D53CAC1FEE0FC0EFB1AFC28" box="[272,1263,929,952]" gridrow="13" pageId="10" pageNumber="55">
<th id="76D037709D53CAC1FEE0FC0EFE5CFC28" box="[272,425,929,952]" gridcol="0" gridrow="13" pageId="10" pageNumber="55">T4L</th>
<td id="76D037709D53CAC1FDABFC0EFD51FC28" box="[603,676,929,952]" gridcol="1" gridrow="13" pageId="10" pageNumber="55">0.792</td>
<td id="76D037709D53CAC1FCEAFC0EFC96FC28" box="[794,867,929,952]" gridcol="2" gridrow="13" pageId="10" pageNumber="55">0.280</td>
<td id="76D037709D53CAC1FC10FC0EFBDCFC28" box="[992,1065,929,952]" gridcol="3" gridrow="13" pageId="10" pageNumber="55">-0.367</td>
<td id="76D037709D53CAC1FB56FC0EFB1AFC28" box="[1190,1263,929,952]" gridcol="4" gridrow="13" pageId="10" pageNumber="55">0.195</td>
</tr>
<tr id="35015E0C9D53CAC1FEE0FC63FB1AFC73" box="[272,1263,972,995]" gridrow="14" pageId="10" pageNumber="55">
<th id="76D037709D53CAC1FEE0FC63FE5CFC73" box="[272,425,972,995]" gridcol="0" gridrow="14" pageId="10" pageNumber="55">F1L</th>
<td id="76D037709D53CAC1FDABFC63FD51FC73" box="[603,676,972,995]" gridcol="1" gridrow="14" pageId="10" pageNumber="55">0.856</td>
<td id="76D037709D53CAC1FCEAFC63FC96FC73" box="[794,867,972,995]" gridcol="2" gridrow="14" pageId="10" pageNumber="55">0.248</td>
<td id="76D037709D53CAC1FC10FC63FBDCFC73" box="[992,1065,972,995]" gridcol="3" gridrow="14" pageId="10" pageNumber="55">-0.119</td>
<td id="76D037709D53CAC1FB56FC63FB1AFC73" box="[1190,1263,972,995]" gridcol="4" gridrow="14" pageId="10" pageNumber="55">-0.284</td>
</tr>
<tr id="35015E0C9D53CAC1FEE0FC59FB1AFB9D" box="[272,1263,1014,1037]" gridrow="15" pageId="10" pageNumber="55">
<th id="76D037709D53CAC1FEE0FC59FE5CFB9D" box="[272,425,1014,1037]" gridcol="0" gridrow="15" pageId="10" pageNumber="55">F3L</th>
<td id="76D037709D53CAC1FDABFC59FD51FB9D" box="[603,676,1014,1037]" gridcol="1" gridrow="15" pageId="10" pageNumber="55">0.896</td>
<td id="76D037709D53CAC1FCEAFC59FC96FB9D" box="[794,867,1014,1037]" gridcol="2" gridrow="15" pageId="10" pageNumber="55">0.287</td>
<td id="76D037709D53CAC1FC10FC59FBDCFB9D" box="[992,1065,1014,1037]" gridcol="3" gridrow="15" pageId="10" pageNumber="55">-0.113</td>
<td id="76D037709D53CAC1FB56FC59FB1AFB9D" box="[1190,1263,1014,1037]" gridcol="4" gridrow="15" pageId="10" pageNumber="55">-0.118</td>
</tr>
<tr id="35015E0C9D53CAC1FEE0FB8EFB1AFBA8" box="[272,1263,1057,1080]" gridrow="16" pageId="10" pageNumber="55">
<th id="76D037709D53CAC1FEE0FB8EFE5CFBA8" box="[272,425,1057,1080]" gridcol="0" gridrow="16" pageId="10" pageNumber="55">F3DW</th>
<td id="76D037709D53CAC1FDABFB8EFD51FBA8" box="[603,676,1057,1080]" gridcol="1" gridrow="16" pageId="10" pageNumber="55">0.902</td>
<td id="76D037709D53CAC1FCEAFB8EFC96FBA8" box="[794,867,1057,1080]" gridcol="2" gridrow="16" pageId="10" pageNumber="55">-0.046</td>
<td id="76D037709D53CAC1FC10FB8EFBDCFBA8" box="[992,1065,1057,1080]" gridcol="3" gridrow="16" pageId="10" pageNumber="55">0.274</td>
<td id="76D037709D53CAC1FB56FB8EFB1AFBA8" box="[1190,1263,1057,1080]" gridcol="4" gridrow="16" pageId="10" pageNumber="55">-0.081</td>
</tr>
<tr id="35015E0C9D53CAC1FEE0FBE3FB1AFBF3" box="[272,1263,1100,1123]" gridrow="17" pageId="10" pageNumber="55">
<th id="76D037709D53CAC1FEE0FBE3FE5CFBF3" box="[272,425,1100,1123]" gridcol="0" gridrow="17" pageId="10" pageNumber="55">F3PPW</th>
<td id="76D037709D53CAC1FDABFBE3FD51FBF3" box="[603,676,1100,1123]" gridcol="1" gridrow="17" pageId="10" pageNumber="55">0.176</td>
<td id="76D037709D53CAC1FCEAFBE3FC96FBF3" box="[794,867,1100,1123]" gridcol="2" gridrow="17" pageId="10" pageNumber="55">0.076</td>
<td id="76D037709D53CAC1FC10FBE3FBDCFBF3" box="[992,1065,1100,1123]" gridcol="3" gridrow="17" pageId="10" pageNumber="55">0.479</td>
<td id="76D037709D53CAC1FB56FBE3FB1AFBF3" box="[1190,1263,1100,1123]" gridcol="4" gridrow="17" pageId="10" pageNumber="55">-0.261</td>
</tr>
<tr id="35015E0C9D53CAC1FEE0FBD9FB1AFB1D" box="[272,1263,1142,1165]" gridrow="18" pageId="10" pageNumber="55">
<th id="76D037709D53CAC1FEE0FBD9FE5CFB1D" box="[272,425,1142,1165]" gridcol="0" gridrow="18" pageId="10" pageNumber="55">T4DW</th>
<td id="76D037709D53CAC1FDABFBD9FD51FB1D" box="[603,676,1142,1165]" gridcol="1" gridrow="18" pageId="10" pageNumber="55">0.865</td>
<td id="76D037709D53CAC1FCEAFBD9FC96FB1D" box="[794,867,1142,1165]" gridcol="2" gridrow="18" pageId="10" pageNumber="55">-0.025</td>
<td id="76D037709D53CAC1FC10FBD9FBDCFB1D" box="[992,1065,1142,1165]" gridcol="3" gridrow="18" pageId="10" pageNumber="55">0.135</td>
<td id="76D037709D53CAC1FB56FBD9FB1AFB1D" box="[1190,1263,1142,1165]" gridcol="4" gridrow="18" pageId="10" pageNumber="55">0.261</td>
</tr>
<tr id="35015E0C9D53CAC1FEE0FB0EFB1AFB28" box="[272,1263,1185,1208]" gridrow="19" pageId="10" pageNumber="55">
<th id="76D037709D53CAC1FEE0FB0EFE5CFB28" box="[272,425,1185,1208]" gridcol="0" gridrow="19" pageId="10" pageNumber="55">T4PPW</th>
<td id="76D037709D53CAC1FDABFB0EFD51FB28" box="[603,676,1185,1208]" gridcol="1" gridrow="19" pageId="10" pageNumber="55">0.047</td>
<td id="76D037709D53CAC1FCEAFB0EFC96FB28" box="[794,867,1185,1208]" gridcol="2" gridrow="19" pageId="10" pageNumber="55">0.074</td>
<td id="76D037709D53CAC1FC10FB0EFBDCFB28" box="[992,1065,1185,1208]" gridcol="3" gridrow="19" pageId="10" pageNumber="55">0.386</td>
<td id="76D037709D53CAC1FB56FB0EFB1AFB28" box="[1190,1263,1185,1208]" gridcol="4" gridrow="19" pageId="10" pageNumber="55">0.462</td>
</tr>
<tr id="35015E0C9D53CAC1FEE0FB63FB1AFB73" box="[272,1263,1228,1251]" gridrow="20" pageId="10" pageNumber="55">
<th id="76D037709D53CAC1FEE0FB63FE5CFB73" box="[272,425,1228,1251]" gridcol="0" gridrow="20" pageId="10" pageNumber="55">Eigenvalues</th>
<td id="76D037709D53CAC1FDABFB63FD51FB73" box="[603,676,1228,1251]" gridcol="1" gridrow="20" pageId="10" pageNumber="55">12.897</td>
<td id="76D037709D53CAC1FCEAFB63FC96FB73" box="[794,867,1228,1251]" gridcol="2" gridrow="20" pageId="10" pageNumber="55">2.605</td>
<td id="76D037709D53CAC1FC10FB63FBDCFB73" box="[992,1065,1228,1251]" gridcol="3" gridrow="20" pageId="10" pageNumber="55">0.977</td>
<td id="76D037709D53CAC1FB56FB63FB1AFB73" box="[1190,1263,1228,1251]" gridcol="4" gridrow="20" pageId="10" pageNumber="55">0.754</td>
</tr>
<tr id="35015E0C9D53CAC1FEE0FB59FB1AFA9D" box="[272,1263,1270,1293]" gridrow="21" pageId="10" pageNumber="55">
<th id="76D037709D53CAC1FEE0FB59FE5CFA9D" box="[272,425,1270,1293]" gridcol="0" gridrow="21" pageId="10" pageNumber="55">% of Variance</th>
<td id="76D037709D53CAC1FDABFB59FD51FA9D" box="[603,676,1270,1293]" gridcol="1" gridrow="21" pageId="10" pageNumber="55">67.878</td>
<td id="76D037709D53CAC1FCEAFB59FC96FA9D" box="[794,867,1270,1293]" gridcol="2" gridrow="21" pageId="10" pageNumber="55">13.709</td>
<td id="76D037709D53CAC1FC10FB59FBDCFA9D" box="[992,1065,1270,1293]" gridcol="3" gridrow="21" pageId="10" pageNumber="55">5.140</td>
<td id="76D037709D53CAC1FB56FB59FB1AFA9D" box="[1190,1263,1270,1293]" gridcol="4" gridrow="21" pageId="10" pageNumber="55">3.968</td>
</tr>
<tr id="35015E0C9D53CAC1FEE0FA8EFB1AFAA8" box="[272,1263,1313,1336]" gridrow="22" pageId="10" pageNumber="55">
<th id="76D037709D53CAC1FEE0FA8EFE5CFAA8" box="[272,425,1313,1336]" gridcol="0" gridrow="22" pageId="10" pageNumber="55">Cumulative %</th>
<td id="76D037709D53CAC1FDABFA8EFD51FAA8" box="[603,676,1313,1336]" gridcol="1" gridrow="22" pageId="10" pageNumber="55">67.878</td>
<td id="76D037709D53CAC1FCEAFA8EFC96FAA8" box="[794,867,1313,1336]" gridcol="2" gridrow="22" pageId="10" pageNumber="55">81.587</td>
<td id="76D037709D53CAC1FC10FA8EFBDCFAA8" box="[992,1065,1313,1336]" gridcol="3" gridrow="22" pageId="10" pageNumber="55">86.728</td>
<td id="76D037709D53CAC1FB56FA8EFB1AFAA8" box="[1190,1263,1313,1336]" gridcol="4" gridrow="22" pageId="10" pageNumber="55">90.696</td>
</tr>
</table>
</paragraph>
<caption id="DF4E0CC69D523535FEF8FA56FCD2F9C5" pageId="11" pageNumber="56">
<paragraph id="8B8E5C4E9D523535FEF8FA56FCD2F9C5" blockId="11.[264,1323,1529,1621]" pageId="11" pageNumber="56">
<emphasis id="B945805C9D523535FEF8FA56FE7BF981" bold="true" box="[264,398,1529,1553]" pageId="11" pageNumber="56">FIGURE 6.</emphasis>
Ventral views of hands and feet of 
<emphasis id="B945805C9D523535FCDCFA56FC5FF980" box="[812,938,1529,1552]" italics="true" pageId="11" pageNumber="56">P. a d i a s t o l a</emphasis>
(A, B: SAMA R61908), a recent specimen of 
<taxonomicName id="4C3127CD9D523535FE65F9B3FDFEF9A3" box="[405,523,1564,1587]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="56" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B945805C9D523535FE65F9B3FDFEF9A3" box="[405,523,1564,1587]" italics="true" pageId="11" pageNumber="56">P. schmidti</emphasis>
</taxonomicName>
from Wanui (C, D: SAMA R57016), and a paratype of 
<taxonomicName id="4C3127CD9D523535FB87F9B3FB1BF9A3" box="[1143,1262,1564,1587]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="56" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B945805C9D523535FB87F9B3FB1BF9A3" box="[1143,1262,1564,1587]" italics="true" pageId="11" pageNumber="56">P. schmidti</emphasis>
</taxonomicName>
from Talasea (E, F: SAMA R6862). Scale bars = 2 mm.
</paragraph>
</caption>
<paragraph id="8B8E5C4E9D523532FEC8F91EFD8DFD56" blockId="11.[264,1323,1713,1899]" lastBlockId="12.[264,1323,284,1870]" lastPageId="12" lastPageNumber="57" pageId="11" pageNumber="56">Skin of dorsal surfaces of body, head, and limbs generally smooth, except for fine longitudinal ridges of variable length on dorsum; a faint pair of medially bowed suprascapular dermal folds present; several other short glandular folds present along dorsolateral edge of trunk and in temporal region; ventral surfaces of trunk, head, throat, and limbs smooth; manus length 45.9% of foot length; digits of manus narrow in appearance and round in cross-section; lateral edges of digits without fleshy dermal flanges; terminal disks moderately expanded and protuberant, with short circumarginal folds on distal tips of digits; minute supraarticlar flaps present above penultimate-ultimate phalangeal articulation; decreasing digital length III, I, II=IV; subarticular tubercles extremely prominent and protuberant, rounded on ventral surfaces; one subarticular tubercle on digits I–II, two tubercles under digits III–IV; supernumerary tubercles very prominent and rounded, present at the base of all digits; minute palmar tubercles preset basal to supernumary tubercles; thenar (inner metacarpal), medial palmar and outer metacarpal tubercle prominent, ovoid, highly convex, and nearly equal in size; slightly raised dermal tubercles present on ventral surfaces of wrists; nuptial pads absent, forearm musculature not hypertrophied.</paragraph>
<paragraph id="8B8E5C4E9D553532FEC8FD7BFD2CFBEE" blockId="12.[264,1323,284,1870]" pageId="12" pageNumber="57">Hindlimbs moderately well developed; foot length 91.9% of tibia length; tibia length 60.3% of snout-vent length; tarsus smooth, lacking folds, flaps, or tubercles; digits of toes moderately expanded and ovoid; slight, narrow lateral dermal flanges present along distal phalanges; circummarginal grooves limited to distal ends of digits; supraarticular cutaneous folds slight; plantar surfaces of pes smooth, with prominently protuberant subarticular tubercles but lacking supernumerary tubercles; subarticular tubercles numbering three under Toe IV, two under Toes III and V, and one each under Toes I and II; decreasing digit length IV, III, V, II, I; outer metatarsal tubercle prominent and subcircular, pointed; inner metatarsal tubercle prominent and elongate, with a sharp venterolateral edge; digits of pes completely free of web; cloacal region smooth and devoid of supracloacaal tubercles or dermal flaps.</paragraph>
<paragraph id="8B8E5C4E9D553532FEC8FB23FC1BFB66" blockId="12.[264,1323,284,1870]" pageId="12" pageNumber="57">
<emphasis id="B945805C9D553532FEC8FB23FD76FB36" bold="true" box="[312,643,1164,1190]" pageId="12" pageNumber="57">
<emphasis id="B945805C9D553532FEC8FB23FD8DFB36" bold="true" box="[312,632,1164,1190]" italics="true" pageId="12" pageNumber="57">
Measurements of 
<typeStatus id="548AE2EC9D553532FDE2FB23FD8DFB36" box="[530,632,1164,1190]" pageId="12" pageNumber="57" type="holotype">holotype</typeStatus>
</emphasis>
:
</emphasis>
SVL 40.6; ED 5.6; TD 3.2; HL 17.3; SNL 7.7; IOD 3.9; HW 15.1; FA 8.6; TBL 24.5; TSL 13.5; PL 22.5; ML 10.3; Toe4L 15.0; Fin1L 5.9; Fin3L 7.2; Fin3DW 1.5; Fin3PPW 0.8; Toe4DW=1.5; Toe4PPW=0.6.
</paragraph>
<paragraph id="8B8E5C4E9D553532FEC8FAABFE61F96E" blockId="12.[264,1323,284,1870]" pageId="12" pageNumber="57">
<emphasis id="B945805C9D553532FEC8FAABFD1BFA8E" bold="true" box="[312,750,1284,1310]" pageId="12" pageNumber="57">
<emphasis id="B945805C9D553532FEC8FAABFD11FA8E" bold="true" box="[312,740,1284,1310]" italics="true" pageId="12" pageNumber="57">
Color of 
<typeStatus id="548AE2EC9D553532FE44FAABFDEFFA8E" box="[436,538,1284,1310]" pageId="12" pageNumber="57" type="holotype">holotype</typeStatus>
in preservative
</emphasis>
:
</emphasis>
Dorsum homogeneous dark brown, blending laterally into light and dark brown marbled flanks with scattered enlarged distinct white spots; dorsal surfaces of head grayish; lateral surfaces of snout and head darker, nearly black with light brown tympanum and distinct white spots on lips; postrictal region heterogeneous, with black and brown blotches and white-tipped postrectal tubercles; dorsal surfaces of limbs brown with darker transverse bars (two on forearm, three on humerus, three on tibia, two on tarsus); dorsal surfaces of hands, feet, and digits dark brown with white bars on phalangeal articulations; inner two digits of hands and feet distinctly lighter than outer digital surfaces; venter pale cream; underside of throat speckled with brown, coalescing into nearly solid brown ventrolateral margins of throat; ventral surfaces of hands dark brown with prominent white subarticular tubercles; ventral surfaces of feet solid dark gray with light gray, velvety digital disks. 
<typeStatus id="548AE2EC9D553532FBDDF913FB6FF946" box="[1069,1178,1724,1750]" pageId="12" pageNumber="57" type="holotype">Holotype</typeStatus>
color in life unrecorded.
</paragraph>
<paragraph id="8B8E5C4E9D553533FEC8F8A3FDAEFCAE" blockId="12.[264,1323,284,1870]" lastBlockId="13.[264,1323,284,1873]" lastPageId="13" lastPageNumber="58" pageId="12" pageNumber="57">
<emphasis id="B945805C9D553532FEC8F8A3FE47F8B6" bold="true" box="[312,434,1804,1830]" pageId="12" pageNumber="57">
<emphasis id="B945805C9D553532FEC8F8A3FE50F8B6" bold="true" box="[312,421,1804,1830]" italics="true" pageId="12" pageNumber="57">Var i at io n</emphasis>
:
</emphasis>
There are three basic color patterns evident in our sample of 
<taxonomicName id="4C3127CD9D553532FB6FF8A3FADFF8B5" box="[1183,1322,1804,1829]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="12" pageNumber="57" phylum="Chordata" rank="species" species="adiastola">
<emphasis id="B945805C9D553532FB6FF8A3FADFF8B5" box="[1183,1322,1804,1829]" italics="true" pageId="12" pageNumber="57">P. adiastola</emphasis>
</taxonomicName>
from the 
<typeStatus id="548AE2EC9D553532FE72F89BFE40F8DE" box="[386,437,1844,1870]" pageId="12" pageNumber="57">type</typeStatus>
locality. The majority (9 of 
<specimenCount id="9D3797C79D553532FCCBF89BFC14F8DE" box="[827,993,1844,1870]" pageId="12" pageNumber="57" type="generic">11 specimens</specimenCount>
) exhibit solid gray dorsal coloration, with faint brown interoribital bars and only scattered flecks of black associated with dorsal tubercular ridges. This largely pigment-free pattern continues on to dorsal surfaces of limbs in a few specimens (SAMA R61907, 61909) but all remaining specimens have distinct transverse brown bars on limbs. The other two major color patterns include a specimen with a single, thick, vertebral yellow stripe (SAMA R61908) and one with two dorsolateral pale lines (SAMA R60259). Lateral surfaces of head with dark bar from tip of snout, through loreal region, encircling orbit, and following supratympanic fold posteroventrally to rictus of jaw. In most specimens (8/11), this dark coloration covers the majority of the lateral portions of the head. In three specimens (SAMA R61907, 61909, 60259), lateral portions of the head are distinctly lighter and darker pigmentation is limited to the canthus rostralis and supratympanic fold. Flank coloration varies from a simple dark (dorsally) to light (ventrally) fade (SAMA R60257, 60259) to a more distinct stratification between dark (above) and pale (below), with numerous moderately distinct pale round spots scattered across flanks.
</paragraph>
<paragraph id="8B8E5C4E9D543533FEC8FCE3FB05FBC6" blockId="13.[264,1323,284,1873]" pageId="13" pageNumber="58">Ventral surfaces of the body are uniform pale cream with throat similarly colored or with a diffuse dark gray ventral surface of the entire head and throat (SAMA R61906, 61908, UPNG 9974-5). In these specimens, the chin and lower lips are distinctly lighter than surrounding throat. Palmar surfaces of the hands vary from very dark gray with white subarticular tubercles (e.g., UPNG 9974) to pale cream throughout their ventral surfaces (SAMA R60257). Plantar surfaces of the feet are uniform light gray (SAMA R60258) to dark gray with lighter subarticular tubercles (SAMA R61906, R61908, UPNG 9974).</paragraph>
<paragraph id="8B8E5C4E9D543533FEC8FBCBFE4EFA76" blockId="13.[264,1323,284,1873]" pageId="13" pageNumber="58">
<emphasis id="B945805C9D543533FEC8FBCBFE20FBEE" bold="true" box="[312,469,1124,1150]" pageId="13" pageNumber="58">
<emphasis id="B945805C9D543533FEC8FBCBFE3EFBEE" bold="true" box="[312,459,1124,1150]" italics="true" pageId="13" pageNumber="58">Color in life</emphasis>
:
</emphasis>
Based on color images of 
<typeStatus id="548AE2EC9D543533FCE1FBCBFC83FBEE" box="[785,886,1124,1150]" pageId="13" pageNumber="58" type="paratype">paratype</typeStatus>
SAMA R61907 before preservation. Dorsal surfaces are khaki to dark tan or brown (e.g. 
<figureCitation id="130A40CB9D543533FC5AFB23FC02FB36" box="[938,1015,1164,1190]" captionStart="FIGURE 5" captionStartId="7.[264,367,1796,1820]" captionText="FIGURE 5. Photographs in life of P. a d i a s t o l a (A: paratype SAMA R 61907) and P. schmidti (B: SAMA R 57015)." pageId="13" pageNumber="58">Fig. 5</figureCitation>
). The dark canthal and supratympanic bars are dark brown, tympanum is light brown, the loreal region is dark brown with gold flecks and the labial region grayish pink with gold bars. The iris is gold above and dark below below pupil and postrictal tubercles are tan with white tips. Bars on limbs are varying shades of brown on a lighter tan or brown background. The dorsal surfaces of the digits are brown with white on spots on digital articulations and the inner two digits on hands and feet and bright white. The throat is white and the venter is white anteriorly, fading to cream or yellow ventrally by groin. Ventral surfaces of limbs are yellow or pink.
</paragraph>
<paragraph id="8B8E5C4E9D543533FEC8FA5BFBAAF946" blockId="13.[264,1323,284,1873]" pageId="13" pageNumber="58">
<emphasis id="B945805C9D543533FEC8FA5BFDF8F99E" bold="true" box="[312,525,1524,1550]" pageId="13" pageNumber="58">
<emphasis id="B945805C9D543533FEC8FA5BFDF6F99E" bold="true" box="[312,515,1524,1550]" italics="true" pageId="13" pageNumber="58">Size dimorphism</emphasis>
:
</emphasis>
The single available female specimen of 
<taxonomicName id="4C3127CD9D543533FBF4FA5BFB65F99D" box="[1028,1168,1524,1549]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="13" pageNumber="58" phylum="Chordata" rank="species" species="adiastola">
<emphasis id="B945805C9D543533FBF4FA5BFB65F99D" box="[1028,1168,1524,1549]" italics="true" pageId="13" pageNumber="58">P. adiastola</emphasis>
</taxonomicName>
has a snoutvent length of 
<quantity id="4CC9F1AB9D543533FE31F9B3FDDAF9A6" box="[449,559,1564,1590]" metricMagnitude="-2" metricUnit="m" metricValue="7.17" pageId="13" pageNumber="58" unit="mm" value="71.7">71.7 mm</quantity>
which is considerably larger than all of the nine available 
<taxonomicName id="4C3127CD9D543533FAE5F9B3FE9EF9CD" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="13" pageNumber="58" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B945805C9D543533FAE5F9B3FE9EF9CD" italics="true" pageId="13" pageNumber="58">P. schmidti</emphasis>
</taxonomicName>
specimens examined by us (Talasea population: SVL 41.1–57.6; 
<tableCitation id="C6B369F59D543533FB8BF9EBFB26F9CE" box="[1147,1235,1604,1630]" captionStart="TABLE 1" captionStartId="9.[264,352,283,307]" captionTargetBox="[272,1304,524,1892]" captionTargetPageId="9" captionText="TABLE 1. Morphometric variation (in mm) in adult Platymantis adiastola n. sp. and P. s c h m i d t i (original collections, holotype and paratypes, as well as recent collections). Table entries include mean ± 1 standard deviation (with range shown below), and sample size (n). See text for character abbreviations." httpUri="http://table.plazi.org/id/DF4E0CC69D503537FEF8FEB4FE6AFE34" pageId="13" pageNumber="58" tableUuid="DF4E0CC69D503537FEF8FEB4FE6AFE34">Table 1</tableCitation>
). This suggests to us the possibility that 
<taxonomicName id="4C3127CD9D543533FD57F9C3FCC1F915" box="[679,820,1644,1669]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="13" pageNumber="58" phylum="Chordata" rank="species" species="adiastola">
<emphasis id="B945805C9D543533FD57F9C3FCC1F915" box="[679,820,1644,1669]" italics="true" pageId="13" pageNumber="58">P. adiastola</emphasis>
</taxonomicName>
may have a greater extent of sexual size dimorphism than 
<taxonomicName id="4C3127CD9D543533FE25F93BFDA0F93D" box="[469,597,1684,1709]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="13" pageNumber="58" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B945805C9D543533FE25F93BFDA0F93D" box="[469,597,1684,1709]" italics="true" pageId="13" pageNumber="58">P. schmidti</emphasis>
</taxonomicName>
(
<emphasis id="B945805C9D543533FD95F93BFD1EF93D" box="[613,747,1684,1709]" italics="true" pageId="13" pageNumber="58">P. a d i a s t o l a</emphasis>
male/female SVL = 0.57; 
<taxonomicName id="4C3127CD9D543533FBD4F93BFB56F93D" box="[1060,1187,1684,1709]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="13" pageNumber="58" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B945805C9D543533FBD4F93BFB56F93D" box="[1060,1187,1684,1709]" italics="true" pageId="13" pageNumber="58">P. schmidti</emphasis>
</taxonomicName>
= 0.69) but larger sample sizes will be required to address this question in the future.
</paragraph>
<paragraph id="8B8E5C4E9D543530FEC8F94BFCB1FDDE" blockId="13.[264,1323,284,1873]" lastBlockId="14.[264,1323,284,1870]" lastPageId="14" lastPageNumber="59" pageId="13" pageNumber="58">
<emphasis id="B945805C9D543533FEC8F94BFD53F96E" bold="true" box="[312,678,1764,1790]" pageId="13" pageNumber="58">
<emphasis id="B945805C9D543533FEC8F94BFD69F96E" bold="true" box="[312,668,1764,1790]" italics="true" pageId="13" pageNumber="58">Ecology and natural history</emphasis>
:
</emphasis>
<taxonomicName id="4C3127CD9D543533FD44F94BFC4FF96D" box="[692,954,1764,1789]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="13" pageNumber="58" phylum="Chordata" rank="species" species="adiastola">
<emphasis id="B945805C9D543533FD44F94BFC4FF96D" box="[692,954,1764,1789]" italics="true" pageId="13" pageNumber="58">Platymantis adiastola</emphasis>
</taxonomicName>
was observed, collected, and recorded at Wanui Camp in primary forest (two hours walking distance from Sampun Village; 
<geoCoordinate id="EE053A899D543533FE9DF898FE66F8D2" box="[365,403,1844,1873]" direction="west" orientation="longitude" pageId="13" pageNumber="58" precision="925" value="-0.083333336">
0 
<geoCoordinate id="EE053A899D543533FE8CF898FE66F8D2" box="[380,403,1844,1873]" direction="south" orientation="latitude" pageId="13" pageNumber="58" precision="1" value="-5.3606334">5o</geoCoordinate>
</geoCoordinate>
21.638'S, 
<geoCoordinate id="EE053A899D543533FDE2F898FD4EF8C1" box="[530,699,1844,1873]" direction="east" orientation="longitude" pageId="13" pageNumber="58" precision="1" value="152.08777">152o 05.266'E</geoCoordinate>
), 
<quantity id="4CC9F1AB9D543533FD20F898FCA3F8C1" box="[720,854,1847,1873]" metricMagnitude="2" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="13" pageNumber="58" unit="m" value="400.0" valueMax="500.0" valueMin="300.0">300–500 m</quantity>
above sea level. This site was sampled for 14 days (
<date id="FF8F7A8E9D573530FE6AFEB3FD8AFEA6" box="[410,639,284,310]" pageId="14" pageNumber="59" value="2000-03-14" valueMax="2000-03-28" valueMin="2000-03-14">14–28 March 2000</date>
). At Wanui the new species was subjectively characterized as 8–10 times more abundant than 
<taxonomicName id="4C3127CD9D573530FD55FEEBFCD0FECD" box="[677,805,324,349]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="14" pageNumber="59" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B945805C9D573530FD55FEEBFCD0FECD" box="[677,805,324,349]" italics="true" pageId="14" pageNumber="59">P. schmidti</emphasis>
</taxonomicName>
. In contrast, 
<taxonomicName id="4C3127CD9D573530FC3AFEEBFBBEFECD" box="[970,1099,324,349]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="14" pageNumber="59" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B945805C9D573530FC3AFEEBFBBEFECD" box="[970,1099,324,349]" italics="true" pageId="14" pageNumber="59">P. schmidti</emphasis>
</taxonomicName>
is clearly the most common form at lower elevations. Vocalizations confirming the sole presence of 
<taxonomicName id="4C3127CD9D573530FAE5FEC3FE9EFE3D" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="14" pageNumber="59" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B945805C9D573530FAE5FEC3FE9EFE3D" italics="true" pageId="14" pageNumber="59">P. schmidti</emphasis>
</taxonomicName>
have been recorded or noted at numerous coastal localities, e.g. Kokopo and the vicinity of Rabaul, (Gazelle Peninsula), Sulu and Silali (foothills of the Nakanai Mountains) and, most significantly, at Talasea, the 
<typeStatus id="548AE2EC9D573530FCACFE4BFC7AFE6E" box="[860,911,484,510]" pageId="14" pageNumber="59">type</typeStatus>
locality of 
<taxonomicName id="4C3127CD9D573530FBE5FE4BFB60FE6D" box="[1045,1173,484,509]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="14" pageNumber="59" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B945805C9D573530FBE5FE4BFB60FE6D" box="[1045,1173,484,509]" italics="true" pageId="14" pageNumber="59">P. schmidti</emphasis>
</taxonomicName>
(SJR and JF pers. obs.). It was also abundant near Sampun Village (~ 
<quantity id="4CC9F1AB9D573530FC43FDA3FC05FDB6" box="[947,1008,524,550]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="59" unit="m" value="50.0">50 m</quantity>
above sea level) and no 
<taxonomicName id="4C3127CD9D573530FAE5FDA3FE80FDDD" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="14" pageNumber="59" phylum="Chordata" rank="species" species="adiastola">
<emphasis id="B945805C9D573530FAE5FDA3FE80FDDD" italics="true" pageId="14" pageNumber="59">P. adiastola</emphasis>
</taxonomicName>
were confirmed from this lowland site.
</paragraph>
<paragraph id="8B8E5C4E9D573530FEC8FDF3FB13FCAE" blockId="14.[264,1323,284,1870]" pageId="14" pageNumber="59">
Where the two species have been observed together, we have noted slight but clear microhabitat differences between the two forms (e.g. Foufopoulos &amp; Richards in press): 
<taxonomicName id="4C3127CD9D573530FAE5FD2BFE80FD55" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="14" pageNumber="59" phylum="Chordata" rank="species" species="adiastola">
<emphasis id="B945805C9D573530FAE5FD2BFE80FD55" italics="true" pageId="14" pageNumber="59">P. adiastola</emphasis>
</taxonomicName>
has been consistently observed calling from elevated perches (leaves and branches) as high as 
<quantity id="4CC9F1AB9D573530FDF2FD7BFDC4FD7E" box="[514,561,724,750]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="59" unit="m" value="2.0">2 m</quantity>
above the ground. In contrast, 
<taxonomicName id="4C3127CD9D573530FC56FD7BFBDDFD7D" box="[934,1064,724,749]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="14" pageNumber="59" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B945805C9D573530FC56FD7BFBDDFD7D" box="[934,1064,724,749]" italics="true" pageId="14" pageNumber="59">P. schmidti</emphasis>
</taxonomicName>
called predominantly from the ground, from low, herbaceous vegetation, or from stumps or fallen logs, consistently less than 
<quantity id="4CC9F1AB9D573530FDF8FC8BFDC0FCAE" box="[520,565,804,830]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="59" unit="m" value="1.0">1 m</quantity>
above the ground (
<bibRefCitation id="EFA021BF9D573530FCE7FC8BFB21FCAD" author="Menzies" box="[791,1236,804,830]" pageId="14" pageNumber="68" refString="Menzies, J. I. (1982 a) The voices of some male Platymantis species of the New Guinea region. British Journal of Herpetology, 6, 1 - 5." type="journal article" year="1982" yearSuffix="a">Menzies 1982a, SJR and JF pers. obs.</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32B0FC59D57352CFEC8FCE3FB00FA8E" lastPageId="18" lastPageNumber="63" pageId="14" pageNumber="59" type="distribution">
<paragraph id="8B8E5C4E9D573530FEC8FCE3FC27FC26" blockId="14.[264,1323,284,1870]" pageId="14" pageNumber="59">
<emphasis id="B945805C9D573530FEC8FCE3FE21FCF6" bold="true" box="[312,468,844,870]" pageId="14" pageNumber="59">
<emphasis id="B945805C9D573530FEC8FCE3FE3EFCF6" bold="true" box="[312,459,844,870]" italics="true" pageId="14" pageNumber="59">Distribution</emphasis>
:
</emphasis>
Confirmed only at two localities (
<figureCitation id="130A40CB9D573530FC9FFCE3FC40FCF6" box="[879,949,844,870]" captionStart="FIGURE 1" captionStartId="1.[264,367,1195,1219]" captionText="FIGURE 1. Map of New Britain Island in relation to the Bismarck Archipelago (inset), showing type locality of P. a d i a s t o l a new species (A: Wanui), the type locality of P. s c h m i d t i (B: Talasea, Willaumez Peninsula), the foothills of the Nakanai Mountains (C: Sulu and Silali Villages), and the East New Britain capital (star: Rabaul / Kokopo)." pageId="14" pageNumber="59">Fig. 1</figureCitation>
): near Sulu and Silali Villages, foothills of the Nakanai Mountains, West New 
<collectingCountry id="F3261CDE9D573530FCCAFCDBFC7EFC1E" box="[826,907,884,910]" name="United Kingdom" pageId="14" pageNumber="59">Britain</collectingCountry>
(
<bibRefCitation id="EFA021BF9D573530FC6CFCDBFB1AFC1E" author="Foufopoulos" box="[924,1263,884,910]" pageId="14" pageNumber="68" refString="Foufopoulos, J. &amp; Brown, R. M. (2004) A new frog of the genus Platymantis (Amphibia; Anura; Ranidae) from New Britain, with a redescription of the poorly-known Platymantis macrosceles. Copeia, 2004, 825 - 841." type="journal article" year="2004">Foufopoulos &amp; Brown 2004</bibRefCitation>
) and Wanui, East New 
<collectingCountry id="F3261CDE9D573530FE2BFC33FDD8FC26" box="[475,557,924,950]" name="United Kingdom" pageId="14" pageNumber="59">Britain</collectingCountry>
(Foufopoulos &amp; Richards in press).
</paragraph>
<paragraph id="8B8E5C4E9D573530FEC8FC6BFBCCFB66" blockId="14.[264,1323,284,1870]" pageId="14" pageNumber="59">
<emphasis id="B945805C9D573530FEC8FC6BFDD8FC4E" bold="true" box="[312,557,964,990]" pageId="14" pageNumber="59">
<emphasis id="B945805C9D573530FEC8FC6BFDD6FC4E" bold="true" box="[312,547,964,990]" italics="true" pageId="14" pageNumber="59">Advertisement calls</emphasis>
:
</emphasis>
The advertisement call of 
<taxonomicName id="4C3127CD9D573530FC80FC6BFC0FFC4D" box="[880,1018,964,989]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="14" pageNumber="59" phylum="Chordata" rank="species" species="adiastola">
<emphasis id="B945805C9D573530FC80FC6BFC0FFC4D" box="[880,1018,964,989]" italics="true" pageId="14" pageNumber="59">P. adiastola</emphasis>
</taxonomicName>
is markedly distinct from that of 
<emphasis id="B945805C9D573530FEADFC43FE2FFB95" box="[349,474,1004,1029]" italics="true" pageId="14" pageNumber="59">P. s c h m i d t i</emphasis>
in both temporal and spectral characteristics. It is, however, similar in general call structure (a rapidly amplitude-modulated train of notes) and so its impression on the human ear is similar. Because the call of the new species is one of the primary characteristics distinguishing the new species, we describe its vocalizations and compare them to those of 
<emphasis id="B945805C9D573530FE23FB23FDA5FB35" box="[467,592,1164,1189]" italics="true" pageId="14" pageNumber="59">P. s c h m i d t i</emphasis>
in some detail, below. The following descriptions are based on 4 vouchered recordings of 
<taxonomicName id="4C3127CD9D573530FD99FB1BFD06FB5D" box="[617,755,1204,1229]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="14" pageNumber="59" phylum="Chordata" rank="species" species="adiastola">
<emphasis id="B945805C9D573530FD99FB1BFD06FB5D" box="[617,755,1204,1229]" italics="true" pageId="14" pageNumber="59">P. adiastola</emphasis>
</taxonomicName>
(2 from the Nakanai Mountains; 2 from Wanui) and 11 of 
<emphasis id="B945805C9D573530FE8DFB73FE0CFB65" box="[381,505,1244,1269]" italics="true" pageId="14" pageNumber="59">P. s c h m i d t i</emphasis>
(9 from the Nakanai Mountains; 2 from Wanui).
</paragraph>
<paragraph id="8B8E5C4E9D573530FEC8FAABFE8AF946" blockId="14.[264,1323,284,1870]" pageId="14" pageNumber="59">
The call of 
<taxonomicName id="4C3127CD9D573530FE31FAABFDB9FA8D" box="[449,588,1284,1309]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="14" pageNumber="59" phylum="Chordata" rank="species" species="adiastola">
<emphasis id="B945805C9D573530FE31FAABFDB9FA8D" box="[449,588,1284,1309]" italics="true" pageId="14" pageNumber="59">P. adiastola</emphasis>
</taxonomicName>
is a series of shrill, high-frequency tonal notes, delivered in a train of variable total duration and note repetition rate. One very distinctive characteristic of the call is the progressive shortening of inter-note intervals throughout the call, such that calls start slowly and gradually shift towards a more rapid note delivery rate towards the end of the note train. The effect on the human ear is a sound akin to that of a marble being dropped and bouncing, faster and faster, until it comes to rest after a final burst of rapid strikes (
<figureCitation id="130A40CB9D573530FE5AFA5BFE04F99E" box="[426,497,1524,1550]" captionStart="FIGURE 7" captionStartId="15.[264,367,1273,1297]" captionText="FIGURE 7. Typical 5.0 s oscillograms from advertisement calls of P. adiastola (A: SAMA R 60258) and P. s c h m i d t i (B: SAMA R 57014). Note decreasing internote interval across the call of P. adiastola (A)." pageId="14" pageNumber="59">Fig. 7</figureCitation>
A). In contrast, the calls of 
<emphasis id="B945805C9D573530FCB0FA5BFC48F99D" box="[832,957,1524,1549]" italics="true" pageId="14" pageNumber="59">P. s c h m i d t i</emphasis>
consist of a much more rapid, evenly-spaced train of dull, lower frequency, stridulating, vibrational notes, like the sound produced by slowly running a finger over the teeth of a comb or running a stick down a length of wood fence (
<figureCitation id="130A40CB9D573530FDE2F9C3FDACF916" box="[530,601,1644,1670]" captionStart="FIGURE 7" captionStartId="15.[264,367,1273,1297]" captionText="FIGURE 7. Typical 5.0 s oscillograms from advertisement calls of P. adiastola (A: SAMA R 60258) and P. s c h m i d t i (B: SAMA R 57014). Note decreasing internote interval across the call of P. adiastola (A)." pageId="14" pageNumber="59">Fig. 7</figureCitation>
B). Menzies stated that the call of 
<emphasis id="B945805C9D573530FC06F9C3FB87F915" box="[1014,1138,1644,1669]" italics="true" pageId="14" pageNumber="59">P. s c h m i d t i</emphasis>
“lacks musical quality and merely sounds like a stick being rubbed against wooden railings” (
<bibRefCitation id="EFA021BF9D573530FB35F93BFE85F946" author="Menzies" pageId="14" pageNumber="68" refString="Menzies, J. I. (1982 a) The voices of some male Platymantis species of the New Guinea region. British Journal of Herpetology, 6, 1 - 5." type="journal article" year="1982" yearSuffix="a">Menzies 1982a: 2</bibRefCitation>
).
</paragraph>
<paragraph id="8B8E5C4E9D573530FEC8F94BFB91F8DE" blockId="14.[264,1323,284,1870]" pageId="14" pageNumber="59">
Individual male mean note repetition rate ([total number of notes – 1]/time from beginning of first note to beginning of last) was 4.0–4.5 (= 4.3 ± 0.4 SD; n = 4) notes/s for 
<taxonomicName id="4C3127CD9D573530FEF8F89BFE64F8DD" box="[264,401,1844,1869]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="14" pageNumber="59" phylum="Chordata" rank="species" species="adiastola">
<emphasis id="B945805C9D573530FEF8F89BFE64F8DD" box="[264,401,1844,1869]" italics="true" pageId="14" pageNumber="59">P. adiastola</emphasis>
</taxonomicName>
and 9.7–15.3 (= 12.8 ± 3.7 SD; n = 9) notes/s for 
<emphasis id="B945805C9D573530FC2DF89BFBACF8DD" box="[989,1113,1844,1869]" italics="true" pageId="14" pageNumber="59">P. s c h m i d t i</emphasis>
.
</paragraph>
<paragraph id="8B8E5C4E9D563531FEC8FEB3FC93FE6E" blockId="15.[264,1323,284,510]" pageId="15" pageNumber="60">
Calls of individual males of 
<taxonomicName id="4C3127CD9D563531FD7FFEB3FCEEFEA5" box="[655,795,284,309]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="15" pageNumber="60" phylum="Chordata" rank="species" species="adiastola">
<emphasis id="B945805C9D563531FD7FFEB3FCEEFEA5" box="[655,795,284,309]" italics="true" pageId="15" pageNumber="60">P. adiastola</emphasis>
</taxonomicName>
contained as few as 9–15 (mean 11.6 ± 3.6 SD) to as many as 11–38 (17.1 ± 8.9 SD) notes per call. The average number of notes per call for our sample of four males was 16.6 ± 3.4 (11.6–17.1; n = 5). Calls of individual male 
<emphasis id="B945805C9D563531FEB8FE3BFE31FE3D" box="[328,452,404,429]" italics="true" pageId="15" pageNumber="60">P. s c h m i d t i</emphasis>
contained as few as 8–15 (mean 13.6 ± 5.2 SD) to as many as 42–55 (45.2 ± 8.1 SD) notes per call. The average number of notes per call for our sample of 
<specimenCount id="9D3797C79D563531FB1DFE13FADDFE45" box="[1261,1320,444,470]" pageId="15" pageNumber="60" type="generic">
13 
<taxonomicName id="4C3127CD9D563531FAE5FE13FADDFE45" box="[1301,1320,444,469]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="15" pageNumber="60" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B945805C9D563531FAE5FE13FADDFE45" box="[1301,1320,444,469]" italics="true" pageId="15" pageNumber="60">P.</emphasis>
</taxonomicName>
</specimenCount>
schmidti recordings was 23.9 ± 8.8 SD (13.6–45.2).
</paragraph>
<caption id="DF4E0CC69D563531FEF8FB56FE4DFAC5" pageId="15" pageNumber="60">
<paragraph id="8B8E5C4E9D563531FEF8FB56FE4DFAC5" blockId="15.[264,1322,1273,1366]" pageId="15" pageNumber="60">
<emphasis id="B945805C9D563531FEF8FB56FE65FA81" bold="true" box="[264,400,1273,1297]" pageId="15" pageNumber="60">FIGURE 7.</emphasis>
Typical 5.0 s oscillograms from advertisement calls of 
<taxonomicName id="4C3127CD9D563531FBECFB56FB6AFA80" box="[1052,1183,1273,1296]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="15" pageNumber="60" phylum="Chordata" rank="species" species="adiastola">
<emphasis id="B945805C9D563531FBECFB56FB6AFA80" box="[1052,1183,1273,1296]" italics="true" pageId="15" pageNumber="60">P. adiastola</emphasis>
</taxonomicName>
(A: SAMA R60258) and 
<emphasis id="B945805C9D563531FE6AFAB3FDFEFAA3" box="[410,523,1308,1331]" italics="true" pageId="15" pageNumber="60">P. s c h m i d t i</emphasis>
(B: SAMA R57014). Note decreasing internote interval across the call of 
<taxonomicName id="4C3127CD9D563531FEF8FA90FE70FAC6" box="[264,389,1343,1366]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="15" pageNumber="60" phylum="Chordata" rank="species" species="adiastola">
<emphasis id="B945805C9D563531FEF8FA90FE70FAC6" box="[264,389,1343,1366]" italics="true" pageId="15" pageNumber="60">P. adiastola</emphasis>
</taxonomicName>
(A).
</paragraph>
</caption>
<paragraph id="8B8E5C4E9D563531FEC8FA1EFED2F8CB" blockId="15.[264,1323,1457,1883]" pageId="15" pageNumber="60">
Individual note structure between the two species was quite different. The shrill, high frequency tonal notes of the call of 
<emphasis id="B945805C9D563531FD2DFA76FC9EFA62" box="[733,875,1497,1522]" italics="true" pageId="15" pageNumber="60">P. a d i a s t o l a</emphasis>
have a relatively complex internal temporal and spectral structure (
<figureCitation id="130A40CB9D563531FD6CF9AEFD16F98B" box="[668,739,1537,1563]" captionStart="FIGURE 8" captionStartId="16.[264,367,1447,1471]" captionText="FIGURE 8. The advertisement call of P. adiastola (paratype SAMA R 60258) recorded at ambient temperature of 25 ° C on 15 March 2000: a 150 ms power spectrum (A: Fast Fourier Transformation; relative amplitude vs. frequency in kHz) and an expanded waveform (B: relative amplitude vs. time in ms) of a single note. A 1500 ms audiospectrogram (C: frequency in kHz vs. time in ms) and oscillogram (D: relative amplitude vs. time in ms) of six typical notes from the call depicted in Fig. 7 A." pageId="15" pageNumber="60">Fig 8</figureCitation>
). Each note contains three distinct frequency components (
<figureCitation id="130A40CB9D563531FE56F986FE1EF9D3" box="[422,491,1577,1603]" captionStart="FIGURE 8" captionStartId="16.[264,367,1447,1471]" captionText="FIGURE 8. The advertisement call of P. adiastola (paratype SAMA R 60258) recorded at ambient temperature of 25 ° C on 15 March 2000: a 150 ms power spectrum (A: Fast Fourier Transformation; relative amplitude vs. frequency in kHz) and an expanded waveform (B: relative amplitude vs. time in ms) of a single note. A 1500 ms audiospectrogram (C: frequency in kHz vs. time in ms) and oscillogram (D: relative amplitude vs. time in ms) of six typical notes from the call depicted in Fig. 7 A." pageId="15" pageNumber="60">Fig. 8</figureCitation>
A, C) and a complex, amplitude modulated internal note structure with pronounced differences in call energy contained in the different amplitude modulated within-note pulses (
<figureCitation id="130A40CB9D563531FE02F9D6FDCCF903" box="[498,569,1657,1683]" captionStart="FIGURE 8" captionStartId="16.[264,367,1447,1471]" captionText="FIGURE 8. The advertisement call of P. adiastola (paratype SAMA R 60258) recorded at ambient temperature of 25 ° C on 15 March 2000: a 150 ms power spectrum (A: Fast Fourier Transformation; relative amplitude vs. frequency in kHz) and an expanded waveform (B: relative amplitude vs. time in ms) of a single note. A 1500 ms audiospectrogram (C: frequency in kHz vs. time in ms) and oscillogram (D: relative amplitude vs. time in ms) of six typical notes from the call depicted in Fig. 7 A." pageId="15" pageNumber="60">Fig. 8</figureCitation>
B). The dull, lower frequency twin-pulse notes of the call of 
<taxonomicName id="4C3127CD9D563531FAE5F9D6FE9EF92A" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="15" pageNumber="60" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B945805C9D563531FAE5F9D6FE9EF92A" italics="true" pageId="15" pageNumber="60">P. schmidti</emphasis>
</taxonomicName>
(
<figureCitation id="130A40CB9D563531FE8DF90EFE31F92B" box="[381,452,1697,1723]" captionStart="FIGURE 9" captionStartId="17.[264,367,1681,1705]" captionText="FIGURE 9. The advertisement call of P. schmidti (SAMA R 57014) recorded at ambient temperature of 28 ° C on 13 March 2000: a 150 ms power spectrum (A: Fast Fourier Transformation; relative amplitude vs. frequency in kHz) and an expanded waveform (B: relative amplitude vs. time in ms) of a single note. A 1500 ms audiospectrogram (C: frequency in kHz vs. time in ms) and oscillogram (D: relative amplitude vs. time in ms) of 12 typical twin-pulse notes from the call depicted in Fig. 7 B." pageId="15" pageNumber="60">Fig. 9</figureCitation>
) are quite different in that each note contains a tightly-spaced twin-pulse structure (
<figureCitation id="130A40CB9D563531FE71F966FE3FF973" box="[385,458,1737,1763]" captionStart="FIGURE 9" captionStartId="17.[264,367,1681,1705]" captionText="FIGURE 9. The advertisement call of P. schmidti (SAMA R 57014) recorded at ambient temperature of 28 ° C on 13 March 2000: a 150 ms power spectrum (A: Fast Fourier Transformation; relative amplitude vs. frequency in kHz) and an expanded waveform (B: relative amplitude vs. time in ms) of a single note. A 1500 ms audiospectrogram (C: frequency in kHz vs. time in ms) and oscillogram (D: relative amplitude vs. time in ms) of 12 typical twin-pulse notes from the call depicted in Fig. 7 B." pageId="15" pageNumber="60">Fig. 9</figureCitation>
B–D), with pulses only slightly differentiated in amplitude (
<figureCitation id="130A40CB9D563531FB6EF966FB1DF973" box="[1182,1256,1737,1763]" captionStart="FIGURE 9" captionStartId="17.[264,367,1681,1705]" captionText="FIGURE 9. The advertisement call of P. schmidti (SAMA R 57014) recorded at ambient temperature of 28 ° C on 13 March 2000: a 150 ms power spectrum (A: Fast Fourier Transformation; relative amplitude vs. frequency in kHz) and an expanded waveform (B: relative amplitude vs. time in ms) of a single note. A 1500 ms audiospectrogram (C: frequency in kHz vs. time in ms) and oscillogram (D: relative amplitude vs. time in ms) of 12 typical twin-pulse notes from the call depicted in Fig. 7 B." pageId="15" pageNumber="60">Fig. 9</figureCitation>
B, D) and both very similar to one another in spectral structure (
<figureCitation id="130A40CB9D563531FC32F95EFBFEF89B" box="[962,1035,1777,1803]" captionStart="FIGURE 9" captionStartId="17.[264,367,1681,1705]" captionText="FIGURE 9. The advertisement call of P. schmidti (SAMA R 57014) recorded at ambient temperature of 28 ° C on 13 March 2000: a 150 ms power spectrum (A: Fast Fourier Transformation; relative amplitude vs. frequency in kHz) and an expanded waveform (B: relative amplitude vs. time in ms) of a single note. A 1500 ms audiospectrogram (C: frequency in kHz vs. time in ms) and oscillogram (D: relative amplitude vs. time in ms) of 12 typical twin-pulse notes from the call depicted in Fig. 7 B." pageId="15" pageNumber="60">Fig. 9</figureCitation>
C). Menzies referred to the call of 
<taxonomicName id="4C3127CD9D563531FE77F8B6FDFDF8A2" box="[391,520,1817,1842]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="15" pageNumber="60" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B945805C9D563531FE77F8B6FDFDF8A2" box="[391,520,1817,1842]" italics="true" pageId="15" pageNumber="60">P. schmidti</emphasis>
</taxonomicName>
as &quot;little more than a succession of double clicks” (
<bibRefCitation id="EFA021BF9D563531FB9EF8B6FEECF8CB" author="Menzies" pageId="15" pageNumber="68" refString="Menzies, J. I. (1982 a) The voices of some male Platymantis species of the New Guinea region. British Journal of Herpetology, 6, 1 - 5." type="journal article" year="1982" yearSuffix="a">Menzies 1982a: 2</bibRefCitation>
).
</paragraph>
<caption id="DF4E0CC69D49352EFEF8FA08FEC5F9FB" pageId="16" pageNumber="61">
<paragraph id="8B8E5C4E9D49352EFEF8FA08FEC5F9FB" blockId="16.[264,1323,1447,1643]" pageId="16" pageNumber="61">
<emphasis id="B945805C9D49352EFEF8FA08FE7FFA2F" bold="true" box="[264,394,1447,1471]" pageId="16" pageNumber="61">FIGURE 8.</emphasis>
The advertisement call of 
<taxonomicName id="4C3127CD9D49352EFD5BFA08FCDCFA2E" box="[683,809,1447,1470]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="61" phylum="Chordata" rank="species" species="adiastola">
<emphasis id="B945805C9D49352EFD5BFA08FCDCFA2E" box="[683,809,1447,1470]" italics="true" pageId="16" pageNumber="61">P. adiastola</emphasis>
</taxonomicName>
(paratype SAMA R60258) recorded at ambient temperature of 25°C on 15 March 2000: a 150 ms power spectrum (A: Fast Fourier Transformation; relative amplitude vs. frequency in kHz) and an expanded waveform (B: relative amplitude vs. time in ms) of a single note. A 1500 ms audiospectrogram (C: frequency in kHz vs. time in ms) and oscillogram (D: relative amplitude vs. time in ms) of six typical notes from the call depicted in Fig. 7A.
</paragraph>
</caption>
<paragraph id="8B8E5C4E9D49352CFEC8F968FD72FE46" blockId="16.[264,1323,1735,1881]" lastBlockId="18.[264,1323,284,1310]" lastPageId="18" lastPageNumber="63" pageId="16" pageNumber="61">
In 
<taxonomicName id="4C3127CD9D49352EFEA9F967FE11F971" box="[345,484,1736,1761]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="61" phylum="Chordata" rank="species" species="adiastola">
<emphasis id="B945805C9D49352EFEA9F967FE11F971" box="[345,484,1736,1761]" italics="true" pageId="16" pageNumber="61">P. adiastola</emphasis>
</taxonomicName>
calls, each note contains a clear fundamental frequency component at 1.0 to 2.2 kHz (mean peak frequency = 1.1–1.3 kHz for four males), an emphasized (=dominant) frequency between 2.5 and 3.6 kHz (mean peak frequency = 3.0–3.2 kHz), and then an additional frequency component between 4.2 and 5.0 kHz (mean peak frequency 4.3–4.8 kHz). The three frequency components are stereotyped, with variation in frequency between males present (and apparently varying inversely proportionately with body size; correlations not shown) but with little or no within-male or within-call frequency modulation or energy shifts. In 
<emphasis id="B945805C9D48352FFCD7FE3BFC44FE3D" box="[807,945,404,429]" italics="true" pageId="17" pageNumber="62">P. s c h m i d t i,</emphasis>
the emphasized (= dominant) frequency lies between 2.2 and 2.9 kHz (peak frequency = 2.4–2.6 kHz for 
<specimenCount id="9D3797C79D48352FFB8FFE13FB19FE46" box="[1151,1260,444,470]" pageId="17" pageNumber="62" type="male">11 males</specimenCount>
), and then minor frequency components include a fundamental between 1.1–1.9 kHz (= 1.5kHz) and weak harmonics between 7.4 and 8.6 kHz. Unlike the new species, 
<taxonomicName id="4C3127CD9D4B352CFBAEFEB3FB2AFEA5" box="[1118,1247,284,309]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="18" pageNumber="63" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B945805C9D4B352CFBAEFEB3FB2AFEA5" box="[1118,1247,284,309]" italics="true" pageId="18" pageNumber="63">P. schmidti</emphasis>
</taxonomicName>
males exhibit a shift in emphasized frequency across the duration of long calls. Thus, calls typically start with the first 6–10 notes at a lower dominant frequency (2.4–2.6 kHz) and males shift the calling energy of subsequent notes to higher dominant frequencies (2.8–2.9 kHz) towards the end of the call.
</paragraph>
<caption id="DF4E0CC69D48352FFEF8F93EFE2EF8C5" pageId="17" pageNumber="62">
<paragraph id="8B8E5C4E9D48352FFEF8F93EFE2EF8C5" blockId="17.[264,1323,1681,1877]" pageId="17" pageNumber="62">
<emphasis id="B945805C9D48352FFEF8F93EFE61F939" bold="true" box="[264,404,1681,1705]" pageId="17" pageNumber="62">FIGURE 9.</emphasis>
The advertisement call of 
<taxonomicName id="4C3127CD9D48352FFD11F93EFCAAF938" box="[737,863,1681,1704]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="17" pageNumber="62" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B945805C9D48352FFD11F93EFCAAF938" box="[737,863,1681,1704]" italics="true" pageId="17" pageNumber="62">P. schmidti</emphasis>
</taxonomicName>
(SAMA R57014) recorded at ambient temperature of 28°C on 13 March 2000: a 150 ms power spectrum (A: Fast Fourier Transformation; relative amplitude vs. frequency in kHz) and an expanded waveform (B: relative amplitude vs. time in ms) of a single note. A 1500 ms audiospectrogram (C: frequency in kHz vs. time in ms) and oscillogram (D: relative amplitude vs. time in ms) of 12 typical twin-pulse notes from the call depicted in Fig. 7B.
</paragraph>
</caption>
<paragraph id="8B8E5C4E9D4B352CFEC8FE4BFB19FC86" blockId="18.[264,1323,284,1310]" pageId="18" pageNumber="63">
Individual notes of 
<taxonomicName id="4C3127CD9D4B352CFDECFE4BFD50FE6D" box="[540,677,484,509]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="18" pageNumber="63" phylum="Chordata" rank="species" species="adiastola">
<emphasis id="B945805C9D4B352CFDECFE4BFD50FE6D" box="[540,677,484,509]" italics="true" pageId="18" pageNumber="63">P. adiastola</emphasis>
</taxonomicName>
begin with 2–4 internal amplitude modulated pulses of 5–10 ms (
<figureCitation id="130A40CB9D4B352CFE8EFDA3FE30FDB6" box="[382,453,524,550]" captionStart="FIGURE 8" captionStartId="16.[264,367,1447,1471]" captionText="FIGURE 8. The advertisement call of P. adiastola (paratype SAMA R 60258) recorded at ambient temperature of 25 ° C on 15 March 2000: a 150 ms power spectrum (A: Fast Fourier Transformation; relative amplitude vs. frequency in kHz) and an expanded waveform (B: relative amplitude vs. time in ms) of a single note. A 1500 ms audiospectrogram (C: frequency in kHz vs. time in ms) and oscillogram (D: relative amplitude vs. time in ms) of six typical notes from the call depicted in Fig. 7 A." pageId="18" pageNumber="63">Fig. 8</figureCitation>
B), followed by the intense tonal note (
<figureCitation id="130A40CB9D4B352CFC65FDA3FC2FFDB6" box="[917,986,524,550]" captionStart="FIGURE 8" captionStartId="16.[264,367,1447,1471]" captionText="FIGURE 8. The advertisement call of P. adiastola (paratype SAMA R 60258) recorded at ambient temperature of 25 ° C on 15 March 2000: a 150 ms power spectrum (A: Fast Fourier Transformation; relative amplitude vs. frequency in kHz) and an expanded waveform (B: relative amplitude vs. time in ms) of a single note. A 1500 ms audiospectrogram (C: frequency in kHz vs. time in ms) and oscillogram (D: relative amplitude vs. time in ms) of six typical notes from the call depicted in Fig. 7 A." pageId="18" pageNumber="63">Fig. 8</figureCitation>
B–D) of 18–25 ms. Average relative intensity (in dB) of each of the preceding pulses is approximately 45–55% of the subsequent tonal portion of each note. Total note duration for individual males ranged from 42.2–46.5 ms (= 44.8 ± 3.8; n=13 notes) to 47.4–53.5 ms (= 50.8 ± 3.1; n=17 notes); mean note duration for our entire sample of recordings was 47.3 ± 4.4 (42.3–56.7; n=4). Inter-note intervals for the same males ranged from 232–255 ms to 245–267 ms; mean inter-note interval for the entire sample was 252.2 ± 26.3 SD ms (242.3–254.7; n=4).
</paragraph>
<paragraph id="8B8E5C4E9D4B352CFEC8FC8BFB00FA8E" blockId="18.[264,1323,284,1310]" pageId="18" pageNumber="63">
Individual notes of 
<taxonomicName id="4C3127CD9D4B352CFDEAFC8BFD6CFCAD" box="[538,665,804,829]" class="Amphibia" family="Ceratobatrachidae" genus="Platymantis" kingdom="Animalia" order="Anura" pageId="18" pageNumber="63" phylum="Chordata" rank="species" species="schmidti">
<emphasis id="B945805C9D4B352CFDEAFC8BFD6CFCAD" box="[538,665,804,829]" italics="true" pageId="18" pageNumber="63">P. schmidti</emphasis>
</taxonomicName>
are paired pulses (
<figureCitation id="130A40CB9D4B352CFC83FC8BFC43FCAE" box="[883,950,804,830]" captionStart="FIGURE 8" captionStartId="16.[264,367,1447,1471]" captionText="FIGURE 8. The advertisement call of P. adiastola (paratype SAMA R 60258) recorded at ambient temperature of 25 ° C on 15 March 2000: a 150 ms power spectrum (A: Fast Fourier Transformation; relative amplitude vs. frequency in kHz) and an expanded waveform (B: relative amplitude vs. time in ms) of a single note. A 1500 ms audiospectrogram (C: frequency in kHz vs. time in ms) and oscillogram (D: relative amplitude vs. time in ms) of six typical notes from the call depicted in Fig. 7 A." pageId="18" pageNumber="63">Fig. 8</figureCitation>
B–D) of similar amplitude (
<figureCitation id="130A40CB9D4B352CFB0BFC8BFEE2FCF6" captionStart="FIGURE 9" captionStartId="17.[264,367,1681,1705]" captionText="FIGURE 9. The advertisement call of P. schmidti (SAMA R 57014) recorded at ambient temperature of 28 ° C on 13 March 2000: a 150 ms power spectrum (A: Fast Fourier Transformation; relative amplitude vs. frequency in kHz) and an expanded waveform (B: relative amplitude vs. time in ms) of a single note. A 1500 ms audiospectrogram (C: frequency in kHz vs. time in ms) and oscillogram (D: relative amplitude vs. time in ms) of 12 typical twin-pulse notes from the call depicted in Fig. 7 B." pageId="18" pageNumber="63">Fig. 9</figureCitation>
D). Both pulses begin with a rapid rise to peak amplitude (2–5 ms), followed by a more gradual decline to ambient levels (
<figureCitation id="130A40CB9D4B352CFD6EFCDBFD10FC1E" box="[670,741,884,910]" captionStart="FIGURE 9" captionStartId="17.[264,367,1681,1705]" captionText="FIGURE 9. The advertisement call of P. schmidti (SAMA R 57014) recorded at ambient temperature of 28 ° C on 13 March 2000: a 150 ms power spectrum (A: Fast Fourier Transformation; relative amplitude vs. frequency in kHz) and an expanded waveform (B: relative amplitude vs. time in ms) of a single note. A 1500 ms audiospectrogram (C: frequency in kHz vs. time in ms) and oscillogram (D: relative amplitude vs. time in ms) of 12 typical twin-pulse notes from the call depicted in Fig. 7 B." pageId="18" pageNumber="63">Fig. 9</figureCitation>
B); in the second pulse, time for amplitude to fall to ambient levels is 2–3 times that of the first pulse. The average relative intensity (in dB) of the first pulses is approximately 75–90% of the second pulse (
<figureCitation id="130A40CB9D4B352CFBE0FC6BFBADFC4E" box="[1040,1112,964,990]" captionStart="FIGURE 9" captionStartId="17.[264,367,1681,1705]" captionText="FIGURE 9. The advertisement call of P. schmidti (SAMA R 57014) recorded at ambient temperature of 28 ° C on 13 March 2000: a 150 ms power spectrum (A: Fast Fourier Transformation; relative amplitude vs. frequency in kHz) and an expanded waveform (B: relative amplitude vs. time in ms) of a single note. A 1500 ms audiospectrogram (C: frequency in kHz vs. time in ms) and oscillogram (D: relative amplitude vs. time in ms) of 12 typical twin-pulse notes from the call depicted in Fig. 7 B." pageId="18" pageNumber="63">Fig. 9</figureCitation>
B, D). Total note duration for individual males ranged from 44.0–47.8 ms (= 45.8 ± 2.9; n=27 notes) to 48.9–61.7 ms (= 56.1 ± 4.7; n=34 notes); mean note duration for our entire sample of recordings was 47.3 ± 4.4 SD (45.8–56.1; n= 
<specimenCount id="9D3797C79D4B352CFCDCFB93FC6CFBC6" box="[812,921,1084,1110]" pageId="18" pageNumber="63" type="male">11 males</specimenCount>
). Duration of the first pulse was consistently briefer (approximately 10–20 ms) than that of the second (25–33 ms). Individual male interpulse intervals ranged from 11.2–14.3 (= 12.4 ± 1.2 SD) to 17.8–22.5 (20.6 ± 2.6 SD and internote intervals varied from 26.3–37.4 (= 32.2 ± 4.6 SD) to 53.2–64.8 (= 58.2 ± 8.3). Mean interpulse interval for the entire sample was 16.7 ± 5.4 SD (12.4–20.6; n=11) and mean internote interval was 46.3 ± 12.9 SD (32.2–58.2; n=11).
</paragraph>
</subSubSection>
</treatment>
</document>