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212 lines
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<document id="824944513DDB6B97B72550B1F6C479D2" ID-CLB-Dataset="7746" ID-DOI="10.5252/geodiversitas2021v43a5" ID-GBIF-Dataset="e42658a2-0e15-4e90-9bc5-0d00cbcbd781" ID-ISSN="1638-9395" ID-Zenodo-Dep="4605963" ID-ZooBank="urn:lsid:zoobank.org:pub:697FC553-E37B-4EF9-97A4-950E4DEE246C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615828395930" checkinUser="felipe" docAuthor="Zouhri, Samir, Gingerich, Philip D., Khalloufi, Bouziane, Bourdon, Estelle, Adnet, Sylvain, Jouve, Stéphane, Elboudali, Najia, Amane, Ayoub, Rage, Jean-Claude & Tabuce, Rodolphe" docDate="2021" docId="03923C45FF8DFF843147FEA8FD9E1023" docLanguage="en" docName="Geodiversitas.43.5.121-150.pdf" docOrigin="Geodiversitas 43 (5)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Cylindracanthus sp. Leidy 1856" docType="treatment" docVersion="6" lastPageNumber="131" masterDocId="FFAB443DFF81FF883018FFDFFF95166E" masterDocTitle="Middle Eocene vertebrate fauna from the Aridal Formation, Sabkha of Gueran, southwestern Morocco" masterLastPageNumber="150" masterPageNumber="121" pageNumber="131" updateTime="1698933443279" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:titleInfo id="8983C36C096EE60696D7334323C76D98">
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<mods:title id="A71BB66C77ED82BCD099CD58A5062FF3">Middle Eocene vertebrate fauna from the Aridal Formation, Sabkha of Gueran, southwestern Morocco</mods:title>
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<mods:namePart id="EF6B5151E4725B7CABCF6249590EB18C">Zouhri, Samir</mods:namePart>
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<mods:namePart id="CB3D53EF46154CB6DFDDA200F000D644">Gingerich, Philip D.</mods:namePart>
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<mods:namePart id="3E9810DEDD6F038AB85D5948A0FB8890">Khalloufi, Bouziane</mods:namePart>
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<mods:namePart id="6A5F17E3AF3270977ECC7F6125563054">Bourdon, Estelle</mods:namePart>
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<mods:namePart id="BB53392E3349B20D5036EB62FF5DEC1E">Adnet, Sylvain</mods:namePart>
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<mods:namePart id="6624519F98EA6EC30FDC629AA6044B51">Jouve, Stéphane</mods:namePart>
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<mods:namePart id="B07760061FB62240F6E17FF415A9095B">Elboudali, Najia</mods:namePart>
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<mods:namePart id="86F0636705D2D9D840F4F45845C949E5">Amane, Ayoub</mods:namePart>
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<mods:namePart id="E03E82398AFFE2B575F7A00F03BAD607">Rage, Jean-Claude</mods:namePart>
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<mods:namePart id="6D24FAAD88B4E8EED0AD051FB26ADA78">Tabuce, Rodolphe</mods:namePart>
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<mods:typeOfResource id="DA280F1290E747CC9B0DBA148E2319F6">text</mods:typeOfResource>
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<mods:title id="15825BEA1F3C0797426392B56F10A6C0">Geodiversitas</mods:title>
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<mods:part id="9D4E5339F258A6931C2BA64282909EEB">
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<mods:date id="7F9B19D42DA8748724904199BB96BD0C">2021</mods:date>
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<mods:number id="C2618853F7CA49D3939649821AF0ED6D">2021-03-11</mods:number>
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<mods:detail id="4E148F0ADF5C7B77688A77092A46C88B" type="volume">
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<mods:number id="A20C86AF8195A1B97D64F5762F33FAFD">43</mods:number>
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<mods:detail id="3C0C6E3A57AF53569EF0D9D8FCC93373" type="issue">
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<mods:number id="F4FC00C89B971CE821A4391DCA0F053F">5</mods:number>
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<mods:start id="2AA93E7089D5DE3E3746638970D1B730">121</mods:start>
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<mods:end id="333D89F1553BF86B87DAD97AB7F8FCF5">150</mods:end>
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<mods:identifier id="A81E1BDB287DD87056F398457EE2231C" type="CLB-Dataset">7746</mods:identifier>
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<mods:identifier id="819B077E0FECA2A020EE228CDC254E24" type="DOI">10.5252/geodiversitas2021v43a5</mods:identifier>
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<mods:identifier id="B5FF33757489E99C08A799681970D821" type="GBIF-Dataset">e42658a2-0e15-4e90-9bc5-0d00cbcbd781</mods:identifier>
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<mods:identifier id="0E83B452678C58C30F0E2AEDDB41C69B" type="ISSN">1638-9395</mods:identifier>
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<treatment id="03923C45FF8DFF843147FEA8FD9E1023" ID-DOI="http://doi.org/10.5281/zenodo.4606623" ID-GBIF-Taxon="180569454" ID-Zenodo-Dep="4606623" LSID="urn:lsid:plazi:treatment:03923C45FF8DFF843147FEA8FD9E1023" httpUri="http://treatment.plazi.org/id/03923C45FF8DFF843147FEA8FD9E1023" lastPageNumber="131" pageId="12" pageNumber="131">
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<subSubSection id="C321DED8FF8DFF843147FEA8FDB817FF" box="[351,557,375,401]" pageId="12" pageNumber="131" type="nomenclature">
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<paragraph id="8B848D53FF8DFF843147FEA8FDB817FF" blockId="12.[351,557,375,401]" box="[351,557,375,401]" pageId="12" pageNumber="131">
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<heading id="D0CC3A3FFF8DFF843147FEA8FDB817FF" box="[351,557,375,401]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="131" reason="2">
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<taxonomicName id="4C3BF6D0FF8DFF843147FEA8FD9D17FF" authorityName="Leidy" authorityYear="1856" box="[351,520,375,401]" class="Actinopterygii" family="Scombridae" genus="Cylindracanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="131" phylum="Chordata" rank="species" species="sp.">
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<emphasis id="B94F5141FF8DFF843147FEA8FD9D17FF" box="[351,520,375,401]" italics="true" pageId="12" pageNumber="131">Cylindracanthus</emphasis>
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</taxonomicName>
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sp.
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C321DED8FF8DFF84309CFE17FD981794" pageId="12" pageNumber="131" type="materials_examined">
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<paragraph id="8B848D53FF8DFF84309CFE17FD981794" blockId="12.[131,775,456,506]" pageId="12" pageNumber="131">
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EXAMINED MATERIAL. —
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<materialsCitation id="3B53870EFF8DFF84316FFE17FD981794" ID-GBIF-Occurrence="3053068313" pageId="12" pageNumber="131">
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FSAC Bouj-141, 356 (
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<figureCitation id="130091D6FF8DFF843249FE17FD0D178E" box="[593,664,456,480]" captionStart="FIG" captionStartId="14.[132,143,1781,1798]" captionTargetBox="[132,1455,215,1723]" captionTargetId="figure-26@14.[698,1267,779,1347]" captionTargetPageId="14" captionText="FIG. 5. — Actinopterygii and Testudines from the Sabkha of Gueran: A-G, Actinopterygii: Cylindracanthus sp., FSAC Bouj-356, rostrum in (A1) lateral and (A2) transversal views; B-E, Percomorpha indet.: B, FSAC Bouj-369,vertebra in (B1) lateral and (B2) transversal views; C, FSAC Bouj-363, vertebra in (C1) lateral and (C2) transversal views; D, FSAC Bouj-370, basioccipital in left lateral view; E, FSAC Bouj-371, fin spine in anterior view; F, Scombridae indet., FSAC Bouj-358, incomplete jawbone in labial view; G, Siluriformes indet., FSAC Bouj-372, right pectoral spine in anterior view; H-P, Testudines: H-L, Cheloniidae indet., FSAC Bouj-109,fragmentary costal in (H1) lateral transection, (H2) ventral, (H3) dorsal and (H4) medial transection views; I, FSAC Bouj-352,fragmentary lateral process of right hyoplastron in (I1) ventral and (I2) dorsal views; J, FSAC Bouj-351, subcomplete right hypoplastron in (J1) ventral and (J2) dorsal views; K, FSAC Bouj- 353, fragment of dermal plate, unlocated on the carapace, in dorsal view; L, FSAC Bouj-354, fragment of dermal plate, unlocated on the shell in (L1, L2) both faces; M, Dermochelyidae indet., FSAC Bouj-350, lateral pubic process of right pubis in (M1) ventral and (M2) dorsal views; N,? Podocnemidoidea indet., FSAC Bouj-196, fragment of right hypoplastron in (N1) dorsal and (N2) ventral views; O, Indeterminata (? Podocnemidoidea indet.), FSAC Bouj-95, fragment of dermal plate of carapace in (O1) dorsal and (O2) ventral views; P, Gueran turtles, pieces of the four fragments of Figure 5I, J, L and M, as preserved in view of conjunction in situ. Scale bars: 20 mm." figureDoi="http://doi.org/10.5281/zenodo.4605975" httpUri="https://zenodo.org/record/4605975/files/figure.png" pageId="12" pageNumber="131">Fig. 5A</figureCitation>
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), 357, 358, 359, 360, 361, 362, fragments of rostra.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C321DED8FF8DFF84309CFDE8FEB41561" pageId="12" pageNumber="131" type="description">
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<paragraph id="8B848D53FF8DFF84309CFDE8FE801421" blockId="12.[132,775,567,783]" box="[132,277,567,593]" pageId="12" pageNumber="131">DESCRIPTION</paragraph>
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<paragraph id="8B848D53FF8DFF84309CFD89FEB41561" blockId="12.[132,775,567,783]" pageId="12" pageNumber="131">The rostra are incomplete and cylindrical. The external surface is crossed by sub-parallel longitudinal ridges, sometimes convergent. The cross-section is circular, with a notched circumference owing to the longitudinal ridges. A unique median canal lies in the center of the cross-section.</paragraph>
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</subSubSection>
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<subSubSection id="C321DED8FF8DFF84309CFCE9FD9E1023" pageId="12" pageNumber="131" type="discussion">
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<paragraph id="8B848D53FF8DFF84309CFCE9FF791520" blockId="12.[131,775,822,1613]" box="[132,236,822,848]" pageId="12" pageNumber="131">REMARKS</paragraph>
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<paragraph id="8B848D53FF8DFF84309CFC8AFD9E1023" blockId="12.[131,775,822,1613]" pageId="12" pageNumber="131">
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These remains are very common in Boujdour. The notched circular cross-section showing a canal and the fluted external surface correspond to the rostrum of
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<taxonomicName id="4C3BF6D0FF8DFF843201FC4AFD5C15C1" authorityName="Leidy" authorityYear="1856" box="[537,713,917,943]" class="Actinopterygii" family="Scombridae" genus="Cylindracanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="131" phylum="Chordata" rank="genus">
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<emphasis id="B94F5141FF8DFF843201FC4AFD5C15C1" box="[537,713,917,943]" italics="true" pageId="12" pageNumber="131">Cylindracanthus</emphasis>
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</taxonomicName>
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. This genus is only known by these peculiar rostra, sometimes showing two rows of minute teeth. They are retrieved in various localities from the Cretaceous to Eocene (and possibly Miocene and Pliocene, see
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<bibRefCitation id="EFAAF0A2FF8DFF8431ECFBCBFD1E1240" author="SCHULTZ O." box="[500,651,1044,1070]" pageId="12" pageNumber="131" pagination="95 - 202" refId="ref25320" refString="SCHULTZ O. 1987. - Taxonomische Neugruppierung der Uberfamilie Xiphioidea (Pisces, Osteichthyes). Annalen des Naturhistorischen Museums in Wien. Serie A fur Mineralogie und Petrographie, Geolo - gie und Palaontologie, Anthropologie und Prahistorie 89: 95 - 202." type="journal article" year="1987">Schultz 1987</bibRefCitation>
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) in Africa, Asia, Europe, and North and South America (
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<bibRefCitation id="EFAAF0A2FF8DFF8432A8FBEBFF541200" author="SCHULTZ O." pageId="12" pageNumber="131" pagination="95 - 202" refId="ref25320" refString="SCHULTZ O. 1987. - Taxonomische Neugruppierung der Uberfamilie Xiphioidea (Pisces, Osteichthyes). Annalen des Naturhistorischen Museums in Wien. Serie A fur Mineralogie und Petrographie, Geolo - gie und Palaontologie, Anthropologie und Prahistorie 89: 95 - 202." type="journal article" year="1987">Schultz 1987</bibRefCitation>
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;
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<bibRefCitation id="EFAAF0A2FF8DFF8430CCFB8BFE311200" author="GALLO V. & FIGUEIREDO F. J. DE & CARVALHO M. S. S. DE" box="[212,420,1108,1134]" pageId="12" pageNumber="131" pagination="175 - 206" refId="ref20985" refString="GALLO V., FIGUEIREDO F. J. DE & CARVALHO M. S. S. DE 2012. - Sintese da paleoictiofauna marinha das bacias costeiras do Brasil e comentarios sobre a paleoictiofauna cronocorrelata da margem oeste da Africa, in GALLO V., SILVA H. M. A., BRITO P. M. M., FIGUEIREDO F. J. DE (eds), Paleontologia de Vertebrados: Relacao entre America do Sul e Africa. Interciencia, Rio de Janeiro: 175 - 206." type="book chapter" year="2012">
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Gallo
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<emphasis id="B94F5141FF8DFF843139FB8AFEC91200" box="[289,348,1108,1134]" italics="true" pageId="12" pageNumber="131">et al.</emphasis>
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2012
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</bibRefCitation>
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;
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<bibRefCitation id="EFAAF0A2FF8DFF8431AFFB8BFDE31200" author="AVERIANOV A." box="[439,630,1108,1134]" pageId="12" pageNumber="131" pagination="1 - 107" refId="ref19225" refString="AVERIANOV A. 2014. - Review of taxonomy, geographic distribution, and paleoenvironments of Azhdarchidae (Pterosauria). ZooKeys 432 (7914): 1 - 107. https: // doi. org / 10.3897 / zookeys. 432.7913" type="journal article" year="2014">Averianov 2014</bibRefCitation>
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;
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<bibRefCitation id="EFAAF0A2FF8DFF843291FB8BFE9612E0" author="GRANDSTAFF B. S. & PELLEGRINI R. A. & PARRIS D. C. & CLEMENTS D." pageId="12" pageNumber="131" pagination="130 - 143" refId="ref21365" refString="GRANDSTAFF B. S., PELLEGRINI R. A., PARRIS D. C. & CLEMENTS D. 2017. - Thin section microscopy of the fossil fish Cylindracanthus. Proceedings of the South Dakota Academy of Science 96: 130 - 143." type="journal article" year="2017">
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Grandstaff
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<emphasis id="B94F5141FF8DFF84309CFBAAFF2B12E0" box="[132,190,1140,1166]" italics="true" pageId="12" pageNumber="131">et al.</emphasis>
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2017
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</bibRefCitation>
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). Putative isolated vertebrae have also been reported but without anatomical connection with the rostrum, (
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<bibRefCitation id="EFAAF0A2FF8DFF8430E1FB6BFE0512A0" author="LERICHE M." box="[249,400,1204,1230]" pageId="12" pageNumber="131" pagination="231 - 363" refId="ref22806" refString="LERICHE M. 1910. - Les Poissons oligocenes de la Belgique. Memoires du Musee royal d'Histoire naturelle de Belgique 5: 231 - 363." type="journal article" year="1910">Leriche 1910</bibRefCitation>
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;
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<bibRefCitation id="EFAAF0A2FF8DFF843187FB6BFDBC12A0" author="WHITE E. I." box="[415,553,1204,1230]" pageId="12" pageNumber="131" pagination="1 - 82" refId="ref26308" refString="WHITE E. I. 1926. - Eocene fishes from Nigeria. Bulletin of the Nigerian Geological Survey 10: 1 - 82." type="journal article" year="1926">White 1926</bibRefCitation>
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). The phylogenetic relationships of
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<taxonomicName id="4C3BF6D0FF8DFF84315CFB0BFE6E1280" authorityName="Leidy" authorityYear="1856" box="[324,507,1236,1262]" class="Actinopterygii" family="Scombridae" genus="Cylindracanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="131" phylum="Chordata" rank="genus">
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<emphasis id="B94F5141FF8DFF84315CFB0BFE6E1280" box="[324,507,1236,1262]" italics="true" pageId="12" pageNumber="131">Cylindracanthus</emphasis>
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</taxonomicName>
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are still discussed and affinities with chimaeroids, billfishes, dercetids, acipenseriforms and beloniforms have been proposed (
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<bibRefCitation id="EFAAF0A2FF8DFF8432AAFACCFF551323" author="SCHULTZ O." pageId="12" pageNumber="131" pagination="95 - 202" refId="ref25320" refString="SCHULTZ O. 1987. - Taxonomische Neugruppierung der Uberfamilie Xiphioidea (Pisces, Osteichthyes). Annalen des Naturhistorischen Museums in Wien. Serie A fur Mineralogie und Petrographie, Geolo - gie und Palaontologie, Anthropologie und Prahistorie 89: 95 - 202." type="journal article" year="1987">Schultz 1987</bibRefCitation>
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;
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<bibRefCitation id="EFAAF0A2FF8DFF8430CAFAEBFEFE1323" author="WEEMS & R. E" box="[210,363,1331,1358]" pageId="12" pageNumber="131" pagination="53 - 99" refId="ref26186" refString="WEEMS R. E. 1999. - Part 4. Actinopterygian fishes from the Fisher / Sullivan site, in WEEMS R. E. & GRIMSLEY G. J. (eds), Early Eocene Vertebrates and Plants from the Fisher / Sullivan Site (Nanjemoy Formation) Stafford County, Virginia. Pub. 152. Virginia Division of Mineral Resources, Charlottesville: 53 - 99." type="book chapter" year="1999">Weems 1999</bibRefCitation>
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;
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<bibRefCitation id="EFAAF0A2FF8DFF843165FAECFDDC1323" author="PARRIS D. C. & GRANDSTAFF B. S. & BELL G. L. J." box="[381,585,1331,1357]" pageId="12" pageNumber="131" pagination="161 - 172" refId="ref24555" refString="PARRIS D. C., GRANDSTAFF B. S. & BELL G. L. J. 2001. - Reassessment of the affinities of the extinct genus Cylindracanthus (Osteichthyes). Proceedings of the South Dakota Academy of Science 80: 161 - 172." type="journal article" year="2001">
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Parris
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<emphasis id="B94F5141FF8DFF8431D1FAEBFD961323" box="[457,515,1331,1357]" italics="true" pageId="12" pageNumber="131">et al.</emphasis>
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2001
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;
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<bibRefCitation id="EFAAF0A2FF8DFF843243FAECFD6A1323" author="MONSCH K. A." box="[603,767,1331,1357]" pageId="12" pageNumber="131" pagination="445 - 489" refId="ref23857" refString="MONSCH K. A. 2004. - Revision of the scombroid fishes from the Cenozoic of England. Transactions of the Royal Society of Edinburgh Earth Sciences 95: 445 - 489. https: // doi. org / 10.1017 / S 0263593300001164" type="journal article" year="2004">Monsch 2004</bibRefCitation>
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;
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<bibRefCitation id="EFAAF0A2FF8DFF84309CFA8CFEA21303" author="FRIEDMAN M." box="[132,311,1363,1389]" pageId="12" pageNumber="131" pagination="113 - 142" refId="ref20915" refString="FRIEDMAN M. 2012. - Ray-finned fishes (Osteichthyes, Actinopterygii) from the type Maastrichtian, the Netherlands and Belgium, in JAGT J. W. M., DONOVAN S. K., JAGT- YAZYKOVA E. A. (eds), Fossils of the type Maastrichtian (Part 1). Scripta Geologica (Special Issue) 8: 113 - 142." type="journal article" year="2012">Friedman 2012</bibRefCitation>
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;
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<bibRefCitation id="EFAAF0A2FF8DFF843151FA8CFDA61303" author="BONDE N. & LEAL M. E. C." box="[329,563,1363,1389]" pageId="12" pageNumber="131" pagination="50 - 54" refId="ref19817" refString="BONDE N. & LEAL M. E. C. 2017. - Danian teleosteans of the North Atlantic Region compared with the Maastrichtian. Research & Knowledge 3 (2): 50 - 54. https: // doi. org / 10.14456 / randk. 2017.26" type="journal article" year="2017">Bonde & Leal 2017</bibRefCitation>
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) while
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<bibRefCitation id="EFAAF0A2FF8DFF843288FA8CFE8013E0" author="GRANDSTAFF B. S. & PELLEGRINI R. A. & PARRIS D. C. & CLEMENTS D." pageId="12" pageNumber="131" pagination="130 - 143" refId="ref21365" refString="GRANDSTAFF B. S., PELLEGRINI R. A., PARRIS D. C. & CLEMENTS D. 2017. - Thin section microscopy of the fossil fish Cylindracanthus. Proceedings of the South Dakota Academy of Science 96: 130 - 143." type="journal article" year="2017">
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Grandstaff
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<emphasis id="B94F5141FF8DFF84309CFAABFF2813E3" box="[132,189,1395,1421]" italics="true" pageId="12" pageNumber="131">et al.</emphasis>
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(2017)
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</bibRefCitation>
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excluded structural resemblances with the billfish
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<taxonomicName id="4C3BF6D0FF8DFF8430CFFA4CFE7913C3" authority="Lacepede, 1802" authorityName="Lacepede" authorityYear="1802" box="[215,492,1427,1453]" class="Actinopterygii" family="Istiophoridae" genus="Makaira" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="131" phylum="Chordata" rank="genus">
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<emphasis id="B94F5141FF8DFF8430CFFA4CFEA313C3" box="[215,310,1427,1453]" italics="true" pageId="12" pageNumber="131">Makaira</emphasis>
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Lacepède, 1802
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</taxonomicName>
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and the paddlefish
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<taxonomicName id="4C3BF6D0FF8DFF8432DBFA4CFEF213A3" authority="Lacepede, 1797" authorityName="Lacepede" authorityYear="1797" class="Actinopterygii" family="Polyodontidae" genus="Polyodon" kingdom="Animalia" order="Acipenseriformes" pageId="12" pageNumber="131" phylum="Chordata" rank="genus">
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<emphasis id="B94F5141FF8DFF8432DBFA4CFF3913A3" italics="true" pageId="12" pageNumber="131">Polyodon</emphasis>
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Lacepède, 1797
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</taxonomicName>
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by analysing thin sections. In North Africa,
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<taxonomicName id="4C3BF6D0FF8DFF8430CEFA0CFE1C1383" authorityName="Leidy" authorityYear="1856" box="[214,393,1491,1517]" class="Actinopterygii" family="Scombridae" genus="Cylindracanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="131" phylum="Chordata" rank="genus">
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<emphasis id="B94F5141FF8DFF8430CEFA0CFE1C1383" box="[214,393,1491,1517]" italics="true" pageId="12" pageNumber="131">Cylindracanthus</emphasis>
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</taxonomicName>
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occurs in the Ypresian beds of the Phosphate basins of
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|
<collectingCountry id="F32CCDC3FF8DFF84316EFA2CFE481063" box="[374,477,1523,1549]" name="Morocco" pageId="12" pageNumber="131">Morocco</collectingCountry>
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and
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<collectingCountry id="F32CCDC3FF8DFF843204FA2CFDF81063" box="[540,621,1523,1549]" name="Algeria" pageId="12" pageNumber="131">Algeria</collectingCountry>
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(
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<bibRefCitation id="EFAAF0A2FF8DFF84329BFA2CFF2A1043" author="ARAMBOURG C." pageId="12" pageNumber="131" pagination="1 - 372" refId="ref19190" refString="ARAMBOURG C. 1952. - Les Vertebres fossiles des gisements de phosphates (Maroc, Algerie, Tunisie). Notes et Memoires du Service Geologique du Maroc 92: 1 - 372." type="journal article" year="1952">Arambourg 1952</bibRefCitation>
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;
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<bibRefCitation id="EFAAF0A2FF8DFF8430D6F9CDFE2F1043" author="KHALLOUFI B. & BRITO P. M. & CAVIN L. & DUTHEIL D. B." box="[206,442,1554,1581]" pageId="12" pageNumber="131" pagination="167 - 248" refId="ref22321" refString="KHALLOUFI B., BRITO P. M. M, CAVIN L. & DUTHEIL D. B. 2017. - Revue des ichthyofaunes mesozoiques et cenozoiques marocaines, in ZOUHRI S. (ed.), Paleontologie des vertebres du Maroc: etat des connaissances. Memoires de la Societe geologique de France, nouvelle serie, Paris 180: 167 - 248." type="journal article" year="2017">
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Khalloufi
|
|
<emphasis id="B94F5141FF8DFF843159F9CCFEE21042" box="[321,375,1554,1580]" italics="true" pageId="12" pageNumber="131">et al.</emphasis>
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2017
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|
</bibRefCitation>
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|
), and in the Priabonian beds of Ad-Dakhla (
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<bibRefCitation id="EFAAF0A2FF8DFF84312BF9EDFE6F1023" author="ADNET S. & CAPPETTA H. & TABUCE R." box="[307,506,1586,1613]" pageId="12" pageNumber="131" pagination="860 - 870" refId="ref18939" refString="ADNET S., CAPPETTA H. & TABUCE R. 2010. - A middle-late Eocene vertebrate fauna (marine fish and mammals) from southwestern Morocco preliminary report: age and palaeobiogeographical implications. Geological Magazine 147: 860 - 870. https: // doi. org / 10.1017 / s 0016756810000348" type="journal article" year="2010">
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Adnet
|
|
<emphasis id="B94F5141FF8DFF843198F9ECFE221022" box="[384,439,1586,1612]" italics="true" pageId="12" pageNumber="131">et al.</emphasis>
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2010
|
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</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |