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<document id="0187A869F3B7830B0855E55513E224E2" ID-CLB-Dataset="7746" ID-DOI="10.5252/geodiversitas2021v43a5" ID-GBIF-Dataset="e42658a2-0e15-4e90-9bc5-0d00cbcbd781" ID-ISSN="1638-9395" ID-Zenodo-Dep="4605963" ID-ZooBank="urn:lsid:zoobank.org:pub:697FC553-E37B-4EF9-97A4-950E4DEE246C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615828395930" checkinUser="felipe" docAuthor="Zouhri, Samir, Gingerich, Philip D., Khalloufi, Bouziane, Bourdon, Estelle, Adnet, Sylvain, Jouve, Stéphane, Elboudali, Najia, Amane, Ayoub, Rage, Jean-Claude &amp; Tabuce, Rodolphe" docDate="2021" docId="03923C45FF87FF8033B7FA0CFC9717BE" docLanguage="en" docName="Geodiversitas.43.5.121-150.pdf" docOrigin="Geodiversitas 43 (5)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Carcharias Rafinesque 1810" docType="treatment" docVersion="9" lastPageNumber="125" masterDocId="FFAB443DFF81FF883018FFDFFF95166E" masterDocTitle="Middle Eocene vertebrate fauna from the Aridal Formation, Sabkha of Gueran, southwestern Morocco" masterLastPageNumber="150" masterPageNumber="121" pageNumber="125" updateTime="1698933443279" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="5BE9E05E03B1A6FEE3E70B8FAE384C04">Middle Eocene vertebrate fauna from the Aridal Formation, Sabkha of Gueran, southwestern Morocco</mods:title>
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<mods:namePart id="044ABAD748967D165C497A5B6CB70E7B">Khalloufi, Bouziane</mods:namePart>
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<mods:namePart id="DF96928DD6F4888E6B4FEE7B5A6DDD7D">Jouve, Stéphane</mods:namePart>
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<mods:namePart id="510737B292AA8D4B27418AF0859EC7EA">Rage, Jean-Claude</mods:namePart>
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<treatment id="03923C45FF87FF8033B7FA0CFC9717BE" ID-DOI="http://doi.org/10.5281/zenodo.4606613" ID-GBIF-Taxon="180569483" ID-Zenodo-Dep="4606613" LSID="urn:lsid:plazi:treatment:03923C45FF87FF8033B7FA0CFC9717BE" httpUri="http://treatment.plazi.org/id/03923C45FF87FF8033B7FA0CFC9717BE" lastPageId="8" lastPageNumber="125" pageId="6" pageNumber="125">
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<heading id="D0CC3A3FFF87FF8E33B7FA0CFAB81383" box="[943,1325,1490,1517]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="125" reason="2">
<taxonomicName id="4C3BF6D0FF87FF8E33AEFA0DFBB81383" ID-CoL="3HC6" ID-ENA="30500" authorityName="Rafinesque" authorityYear="1810" box="[950,1069,1490,1517]" class="Elasmobranchii" family="Odontaspididae" genus="Carcharias" kingdom="Animalia" order="Lamniformes" pageId="6" pageNumber="125" phylum="Chordata" rank="genus">
<emphasis id="B94F5141FF87FF8E33AEFA0DFBB81383" bold="true" box="[950,1069,1490,1517]" italics="true" pageId="6" pageNumber="125">Carcharias</emphasis>
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<emphasis id="B94F5141FF87FF8E3422FA0DFBEC1383" bold="true" box="[1082,1145,1490,1517]" italics="true" pageId="6" pageNumber="125">koerti</emphasis>
(
<bibRefCitation id="EFAAF0A2FF87FF8E349FFA0CFAB01383" author="STROMER E." box="[1159,1317,1491,1517]" pageId="6" pageNumber="125" pagination="478 - 505" refId="ref25520" refString="STROMER E. 1910. - Reptilien- und Fischreste aus dem marinen Alttertiar von Sudtogo (Westafrica). Zeitschrift der Deutschen Geologischen Gesellschaft 62 (7): 478 - 505." type="journal article" year="1910">Stromer, 1910</bibRefCitation>
)
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<paragraph id="8B848D53FF87FF8E3335F9D3FBBF1050" blockId="6.[813,1455,1548,1598]" pageId="6" pageNumber="125">
MATERIAL. —
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30 broken isolated teeth, figured material included
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Bouj-325 and 326.
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</paragraph>
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<subSubSection id="C321DED8FF87FF8E3335F9ACFB2C1162" pageId="6" pageNumber="125" type="description">
<paragraph id="8B848D53FF87FF8E3335F9ACFC2810E5" blockId="6.[813,1455,1651,1804]" box="[813,957,1651,1677]" pageId="6" pageNumber="125">DESCRIPTION</paragraph>
<paragraph id="8B848D53FF87FF8E3335F94DFB2C1162" blockId="6.[813,1455,1651,1804]" pageId="6" pageNumber="125">
Occurrence of the middle Eocene
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<emphasis id="B94F5141FF87FF8E34BFF94DFA8210C2" box="[1191,1303,1682,1708]" italics="true" pageId="6" pageNumber="125">Carcharias</emphasis>
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<emphasis id="B94F5141FF87FF8E353DF94DFAF510C2" box="[1317,1376,1682,1708]" italics="true" pageId="6" pageNumber="125">koerti</emphasis>
is confirmed in Gueran (
<figureCitation id="130091D6FF87FF8E33E1F96DFBFD10A2" box="[1017,1128,1714,1741]" captionStart="FIG" captionStartId="7.[132,143,1872,1889]" captionTargetBox="[132,1455,215,1848]" captionTargetId="figure-15@7.[132,1455,215,1869]" captionTargetPageId="7" captionText="FIG. 2. — Elasmobranchii Lamniformes teeth remains from Sabkha of Gueran collected around Archaeocete whale carcasses: A-C, Otodus (Carcharocles) cf. sokolowi: A, FSAC Bouj-320 lateral upper tooth in (A1) labial and (A2) lingual views; B, FSAC Bouj-321 anterior lower tooth in labial view; C, FSAC Bouj-322 lateral lower tooth in labial view; D, E, Otodus (Carcharocles) sp.: D, FSAC Bouj-323 antero-lateral tooth in (D1) labial and (D2) lingual views; E, FSAC Bouj-324 lateral tooth (E1) labial and (E2) lingual views; F, G, “Carcharias” koerti; F, FSAC Bouj-325 anterolateral upper tooth in (F1) labial and (F2) lingual views; G, FSAC Bouj-326 anterior tooth of juvenile (G1) labial and (G2) lingual views; H-K, Macrorhizodus praecursor: H, FSAC Bouj-327 anterior lower tooth (H1) labial and (H2) lingual views; I, FSAC Bouj-328 anterior upper tooth in (I1) labial and (I2) lingual views; J, FSAC Bouj-329 lateral upper tooth in (J1) labial and (J2) lingual views; K, FSAC Bouj-330 lateral lower tooth (K1) labial and (K2) lingual views. Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4605967" httpUri="https://zenodo.org/record/4605967/files/figure.png" pageId="6" pageNumber="125">Fig. 2F, G</figureCitation>
). Interestingly, the majority of recovered teeth (e.g.,
<figureCitation id="130091D6FF87FF8E340CF90DFBFD1082" box="[1044,1128,1746,1772]" captionStart="FIG" captionStartId="7.[132,143,1872,1889]" captionTargetBox="[132,1455,215,1848]" captionTargetId="figure-15@7.[132,1455,215,1869]" captionTargetPageId="7" captionText="FIG. 2. — Elasmobranchii Lamniformes teeth remains from Sabkha of Gueran collected around Archaeocete whale carcasses: A-C, Otodus (Carcharocles) cf. sokolowi: A, FSAC Bouj-320 lateral upper tooth in (A1) labial and (A2) lingual views; B, FSAC Bouj-321 anterior lower tooth in labial view; C, FSAC Bouj-322 lateral lower tooth in labial view; D, E, Otodus (Carcharocles) sp.: D, FSAC Bouj-323 antero-lateral tooth in (D1) labial and (D2) lingual views; E, FSAC Bouj-324 lateral tooth (E1) labial and (E2) lingual views; F, G, “Carcharias” koerti; F, FSAC Bouj-325 anterolateral upper tooth in (F1) labial and (F2) lingual views; G, FSAC Bouj-326 anterior tooth of juvenile (G1) labial and (G2) lingual views; H-K, Macrorhizodus praecursor: H, FSAC Bouj-327 anterior lower tooth (H1) labial and (H2) lingual views; I, FSAC Bouj-328 anterior upper tooth in (I1) labial and (I2) lingual views; J, FSAC Bouj-329 lateral upper tooth in (J1) labial and (J2) lingual views; K, FSAC Bouj-330 lateral lower tooth (K1) labial and (K2) lingual views. Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4605967" httpUri="https://zenodo.org/record/4605967/files/figure.png" pageId="6" pageNumber="125">Fig. 2G</figureCitation>
) are relatively small compared to the usual Lutetian representatives.
</paragraph>
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<paragraph id="8B848D53FF87FF803335F88EFC9717BE" blockId="6.[812,1455,1842,2028]" lastBlockId="8.[130,775,215,465]" lastPageId="8" lastPageNumber="127" pageId="6" pageNumber="125">
The generic affinity of this large pelagic shark to
<taxonomicName id="4C3BF6D0FF87FF8E352CF88EFA311105" authorityName="Rafinesque" authorityYear="1810" box="[1332,1444,1873,1899]" class="Elasmobranchii" family="Odontaspididae" genus="Carcharias" kingdom="Animalia" order="Lamniformes" pageId="6" pageNumber="125" phylum="Chordata" rank="genus">
<emphasis id="B94F5141FF87FF8E352CF88EFA311105" box="[1332,1444,1873,1899]" italics="true" pageId="6" pageNumber="125">Carcharias</emphasis>
</taxonomicName>
” or “
<taxonomicName id="4C3BF6D0FF87FF8E334CF8AEFB9411E5" authorityName="Cappetta &amp; Nolf" authorityYear="2005" box="[852,1025,1905,1931]" class="Elasmobranchii" family="Odontaspididae" genus="Brachycarcharias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lamniformes" pageId="6" pageNumber="125" phylum="Chordata" rank="genus">
<emphasis id="B94F5141FF87FF8E334CF8AEFB9411E5" box="[852,1025,1905,1931]" italics="true" pageId="6" pageNumber="125">Brachycarcharias</emphasis>
</taxonomicName>
” still remains unclear (e.g.,
<bibRefCitation id="EFAAF0A2FF87FF8E353BF8AEFCF011C5" author="STROUGO A. &amp; CAPPETTA H. &amp; ELNAHAS S." pageId="6" pageNumber="125" pagination="81 - 98" refId="ref25587" refString="STROUGO A., CAPPETTA H. &amp; ELNAHAS S. 2007. - A remarkable Eocene ichthyofauna from the El Gedidia glauconitic sandstone, Bahariya oasis, Egypt, and its stratigraphic implications. MERC Ain Shams University, Earth Science Series 21: 81 - 98." type="book chapter" year="2007">
Strougo
<emphasis id="B94F5141FF87FF8E3566F8ADFA3A11E5" box="[1406,1455,1905,1931]" italics="true" pageId="6" pageNumber="125">et al.</emphasis>
2007
</bibRefCitation>
;
<bibRefCitation id="EFAAF0A2FF87FF8E336BF84EFBF811C5" author="UNDERWOOD C. J. &amp; WARD D. J. &amp; KING C. &amp; ANTAR S. M. &amp; MOUT I. S. &amp; GINGERICH P. D." box="[883,1133,1937,1963]" pageId="6" pageNumber="125" pagination="47 - 66" refId="ref26062" refString="UNDERWOOD C. J., WARD D. J., KING C., ANTAR S. M., ZAL- MOUT I. S. &amp; GINGERICH P. D. 2011. - Shark and ray faunas in the middle and late Eocene of the Fayum area, Egypt. Proceedings of the Geologists' Association 122: 47 - 66. https: // doi. org / 10.1016 / j. pgeola. 20" type="journal article" year="2011">
Underwood
<emphasis id="B94F5141FF87FF8E33E4F84DFBBA11C5" box="[1020,1071,1937,1963]" italics="true" pageId="6" pageNumber="125">et al.</emphasis>
2011
</bibRefCitation>
), it is preferentially known in the Lutetian deposits and is widely distributed around the North and Western African coasts (
<bibRefCitation id="EFAAF0A2FF87FF8E34A6F80EFACC1185" author="STROMER E." box="[1214,1369,2001,2027]" pageId="6" pageNumber="125" pagination="478 - 505" refId="ref25520" refString="STROMER E. 1910. - Reptilien- und Fischreste aus dem marinen Alttertiar von Sudtogo (Westafrica). Zeitschrift der Deutschen Geologischen Gesellschaft 62 (7): 478 - 505." type="journal article" year="1910">Stromer 1910</bibRefCitation>
;
<bibRefCitation id="EFAAF0A2FF87FF803572F80EFF29169C" author="WHITE E. I." lastPageId="8" lastPageNumber="127" pageId="6" pageNumber="125" pagination="319 - 325" refId="ref26333" refString="WHITE E. I. 1955. - Notes on African Tertiary sharks. Colonial Geology and Mineral Resources 5: 319 - 325." type="journal article" year="1955">White 1955</bibRefCitation>
;
<bibRefCitation id="EFAAF0A2FF89FF8030D1FF08FE4F169C" author="DARTEVELLE E. &amp; CASIER E." box="[201,474,215,242]" pageId="8" pageNumber="127" pagination="257 - 568" refId="ref20641" refString="DARTEVELLE E. &amp; CASIER E. 1959. - Les poissons fossiles du Bas- Congo et des regions voisines. Annales du Musee du Congo belge, Serie A 3 (2, 3): 257 - 568." type="journal article" year="1959">Dartevelle &amp; Casier 1959</bibRefCitation>
;
<bibRefCitation id="EFAAF0A2FF89FF8031FFFF08FC95169F" author="CAPPETTA H. &amp; TRAVERSE M." box="[487,768,215,242]" pageId="8" pageNumber="127" pagination="359 - 365" refId="ref20177" refString="CAPPETTA H. &amp; TRAVERSE M. 1988. - Une riche faune de selaciens dans le bassin a phosphate de Kpogame-Hahotoe (Eocene moyen du Togo): Note preliminaire et precisions sur la structure et l'age du gisement. Geobios 21 (3): 359 - 365. https: // doi. org / 10.1016 / S 0016 - 6995 (88) 80058 - 5" type="journal article" year="1988">Cappetta &amp; Traverse 1988</bibRefCitation>
;
<bibRefCitation id="EFAAF0A2FF89FF80309CFF28FEF5177F" author="CAPPETTA H. &amp; PFEIL F. &amp; KITTLER N." box="[132,352,247,273]" pageId="8" pageNumber="127" pagination="81 - 95" refId="ref20247" refString="CAPPETTA H., PFEIL F. &amp; SCHMIDT- KITTLER N. 2000. - New biostratigraphical data on the marine Upper Cretaceous and Palaeogene of Jordan. Newsletters on Stratigraphy 38: 81 - 95. https: // doi. org / 10.1127 / nos / 38 / 2000 / 81" type="journal article" year="2000">
Cappetta
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2000
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;
<bibRefCitation id="EFAAF0A2FF89FF803176FF28FD36177F" author="NOUBHANI A. &amp; CAPPETTA H." box="[366,675,247,273]" pageId="8" pageNumber="127" pagination="1 - 327" refId="ref24148" refString="NOUBHANI A. &amp; CAPPETTA H. 1997. - Les Orectolobiformes, Carcharhiniformes et Myliobatiformes (Elasmobranchii, Neoselachii) des bassins a phosphate du Maroc (Maastrichtien-Lutetien basal). Systematique, biostratigraphie, evolution et dynamique des faunes. Palaeo Ichthyologica 8: 1 - 327." type="journal article" year="1997">Noubhani &amp; Cappetta 1997</bibRefCitation>
;
<bibRefCitation id="EFAAF0A2FF89FF8032A9FF28FF6D175F" author="STROUGO A. &amp; CAPPETTA H. &amp; ELNAHAS S." pageId="8" pageNumber="127" pagination="81 - 98" refId="ref25587" refString="STROUGO A., CAPPETTA H. &amp; ELNAHAS S. 2007. - A remarkable Eocene ichthyofauna from the El Gedidia glauconitic sandstone, Bahariya oasis, Egypt, and its stratigraphic implications. MERC Ain Shams University, Earth Science Series 21: 81 - 98." type="book chapter" year="2007">
Strougo
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2007
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). Rarely recovered in MI (Uppermost LutetianLowermost Bartonian), GA (Lutetian-Bartonian) and lacking in GE A-C (uppermost Bartonian) and the remainder of the succession of Wadi al Hitan,
<collectingCountry id="F32CCDC3FF89FF8031D1FEA8FD9C17FF" box="[457,521,375,401]" name="Egypt" pageId="8" pageNumber="127">Egypt</collectingCountry>
(see
<bibRefCitation id="EFAAF0A2FF89FF80325FFEA8FF2917DF" author="UNDERWOOD C. J. &amp; WARD D. J. &amp; KING C. &amp; ANTAR S. M. &amp; MOUT I. S. &amp; GINGERICH P. D." pageId="8" pageNumber="127" pagination="47 - 66" refId="ref26062" refString="UNDERWOOD C. J., WARD D. J., KING C., ANTAR S. M., ZAL- MOUT I. S. &amp; GINGERICH P. D. 2011. - Shark and ray faunas in the middle and late Eocene of the Fayum area, Egypt. Proceedings of the Geologists' Association 122: 47 - 66. https: // doi. org / 10.1016 / j. pgeola. 20" type="journal article" year="2011">
Underwood
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2011
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), its occurrence in Boujdour area seems to indicate that the age of the deposits is likely older than latest Bartonian.
</paragraph>
</subSubSection>
</treatment>
</document>