treatments-xml/data/03/8F/87/038F87FBFFF9FF8F724BFC7AFA8EF9FC.xml

<document id="DBC9FAD555AA1D027F53F8C3406B03ED" ID-CLB-Dataset="32180" ID-DOI="10.5281/zenodo.886270" ID-GBIF-Dataset="3d6cdea7-79d5-4d75-8ea7-886ce20dd6e3" ID-ISSN="2345-7600" ID-Zenodo-Dep="886270" ID-ZooBank="urn:lsid:zoobank.org:pub:F7024292-9770-4958-845B-EA3BA4B468AE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1504701442361" checkinUser="plazi" docAuthor="Maurice Kottelat" docDate="2017" docId="038F87FBFFF9FF8F724BFC7AFA8EF9FC" docLanguage="en" docName="2017 Schistura colossa and S. klydonion.pdf" docOrigin="Raffles Bulletin Of Zoology 65" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Schistura klydonion Maurice Kottelat 2017, new species" docType="treatment" docVersion="7" lastPageNumber="355" masterDocId="FFB6FF83FFF1FF817346FFD0FFB4FF84" masterDocTitle="Schistura colossa and S. klydonion, two new species of loaches from Bolaven Plateau, southern Laos (Teleostei: Nemacheilidae)" masterLastPageNumber="356" masterPageNumber="341" pageNumber="349" updateTime="1698467968782" updateUser="ExternalLinkService">
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<mods:title id="35793BD73D0C9FFEE5742544AC1FF036">Schistura colossa and S. klydonion, two new species of loaches from Bolaven Plateau, southern Laos (Teleostei: Nemacheilidae)</mods:title>
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<mods:namePart id="023C6FF16DF6D2C8A23974812CD1C7B0">Maurice Kottelat</mods:namePart>
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<mods:title id="4B352D3EAE5338C9CEF39DAE1D7C452B">Raffles Bulletin Of Zoology</mods:title>
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<mods:date id="4E8EB208CC0EBC4F798843E380A32F22">2017</mods:date>
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<heading id="D0D18181FFF9FF89724BFC7AFD36FC46" bold="true" box="[269,642,938,963]" centered="true" fontSize="10" level="2" pageId="8" pageNumber="349" reason="4">
<emphasis id="B952EAFFFFF9FF89724BFC7AFD36FC46" bold="true" box="[269,642,938,963]" pageId="8" pageNumber="349">
<taxonomicName id="4C264D6EFFF9FF89724BFC7AFE59FC46" authority="Maurice Kottelat, 2017" authorityName="Maurice Kottelat" authorityYear="2017" box="[269,493,938,962]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="349" phylum="Chordata" rank="species" species="klydonion" status="sp. nov.">
<emphasis id="B952EAFFFFF9FF89724BFC7AFE59FC46" bold="true" box="[269,493,938,962]" italics="true" pageId="8" pageNumber="349">Schistura klydonion</emphasis>
</taxonomicName>
, 
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<paragraph id="8B9936EDFFF9FF897239FC1BFDA4FC66" blockId="8.[269,642,938,995]" box="[383,528,970,995]" pageId="8" pageNumber="349">
<heading id="D0D18181FFF9FF897239FC1BFDA4FC66" box="[383,528,970,995]" centered="true" fontSize="10" level="3" pageId="8" pageNumber="349" reason="4">
(
<figureCitation id="131D2A68FFF9FF8972C0FC1BFDBCFC66" box="[390,520,970,995]" captionStart-0="Fig. 13" captionStart-1="Fig. 14" captionStart-2="Fig. 15" captionStartId-0="10.[136,171,968,990]" captionStartId-1="11.[113,148,2066,2088]" captionStartId-2="12.[135,170,1748,1770]" captionTargetBox-0="[157,1440,218,903]" captionTargetBox-1="[222,1316,211,2036]" captionTargetBox-2="[142,1442,241,1713]" captionTargetId-0="figure@10.[136,1474,189,951]" captionTargetId-1="figure@11.[222,1341,189,2049]" captionTargetId-2="figure@12.[135,1473,189,1731]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionText-0="Fig. 13. Schistura klydonion, MHNG 2767.085, 75.8 mm SL, holotype; Laos: Xe Namnoy on Bolaven Plateau (right side, reversed)." captionText-1="Fig. 14. Schistura klydonion, paratypes; Laos: Xe Namnoy on Bolaven Plateau; a, CMK 23344, 57.7 mm SL; b, CMK 23344, 61.9 mm SL; and c, CMK 23320, 76.0 mm SL." captionText-2="Fig. 15. Schistura klydonion, paratypes; Laos: Xe Namnoy on Bolaven Plateau; a, CMK 23344, 26.4 mm SL; b, CMK 23344, 27.3 mm SL; c, CMK 23344, 28.1 mm SL; and d, CMK 23320, 29.0 mm SL." httpUri-0="https://zenodo.org/record/886296/files/figure.png" httpUri-1="https://zenodo.org/record/886298/files/figure.png" httpUri-2="https://zenodo.org/record/886300/files/figure.png" pageId="8" pageNumber="349">Figs. 13–15</figureCitation>
)
</heading>
</paragraph>
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<subSubSection id="C33C6566FFF9FF8973CEFBDBFEF7F8C6" pageId="8" pageNumber="349" type="materials_examined">
<paragraph id="8B9936EDFFF9FF8973CEFBDBFD90FB07" blockId="8.[136,775,1034,1155]" pageId="8" pageNumber="349">
<materialsCitation id="3B4E3CB0FFF9FF8973CEFBDBFD90FB07" ID-GBIF-Occurrence="1611915208" collectingDate="2013-01-14" collectionCode="MHNG" collectorName="M. Kottelat &amp; T. Phommavong" country="Laos" county="Bolaven Plateau" elevation="790" latitude="14.878056" location="Xe Namnoy" longLatPrecision="21" longitude="106.58028" municipality="Houay Xoy" pageId="8" pageNumber="349" specimenCode="MHNG 2767.085, 75.8" specimenCount="1" stateProvince="Champasak Province" typeStatus="holotype">
<emphasis id="B952EAFFFFF9FF8973CEFBDBFF42FBA7" bold="true" box="[136,246,1035,1059]" pageId="8" pageNumber="349">
<typeStatus id="549D884FFFF9FF8973CEFBDBFF44FBA7" box="[136,240,1035,1059]" pageId="8" pageNumber="349" type="holotype">Holotype</typeStatus>
.
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<specimenCode id="DB809E96FFF9FF8973BCFBDBFE59FBA6" box="[250,493,1034,1059]" pageId="8" pageNumber="349">
MHNG 2767.085, 
<quantity id="4CDE9B08FFF9FF8972F9FBDAFE59FBA6" box="[447,493,1034,1058]" metricMagnitude="-2" metricUnit="m" metricValue="7.58" pageId="8" pageNumber="349" unit="mm" value="75.8">75.8</quantity>
</specimenCode>
mm SL; 
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: 
<collectingRegion id="49E2F80FFFF9FF8971CAFBDBFF5FFBC7" country="Laos" name="Champasak" pageId="8" pageNumber="349">Champasak Province</collectingRegion>
: 
<collectingCounty id="62F84E61FFF9FF8973BFFBFBFE1BFBC7" box="[249,431,1067,1091]" pageId="8" pageNumber="349">Bolaven Plateau</collectingCounty>
: 
<collectingMunicipality id="6BFDAC97FFF9FF8972FAFBFBFD8AFBC7" box="[444,574,1067,1091]" pageId="8" pageNumber="349">Houay Xoy</collectingMunicipality>
, a tributary of 
<location id="8EF96036FFF9FF8971AEFBFBFF53FBE7" LSID="urn:lsid:plazi:treatment:038F87FBFFF9FF8F724BFC7AFA8EF9FC:8EF96036FFF9FF8971AEFBFBFF53FBE7" country="Laos" county="Bolaven Plateau" latitude="14.878056" longLatPrecision="21" longitude="106.58028" municipality="Houay Xoy" name="Xe Namnoy" pageId="8" pageNumber="349" stateProvince="Champasak Province">Xe Namnoy</location>
, 
<geoCoordinate id="EE12502AFFF9FF8973B2FB99FEC7FBE7" box="[244,371,1097,1123]" direction="north" orientation="latitude" pageId="8" pageNumber="349" precision="15" value="14.878056">14°52′41″N</geoCoordinate>
<geoCoordinate id="EE12502AFFF9FF89723AFB99FDBCFBE7" box="[380,520,1097,1123]" direction="east" orientation="longitude" pageId="8" pageNumber="349" precision="15" value="106.58028">106°34′49″E</geoCoordinate>
; 
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<quantity id="4CDE9B08FFF9FF897153FB9BFDC9FBE7" box="[533,637,1099,1123]" metricMagnitude="2" metricUnit="m" metricValue="7.9" pageId="8" pageNumber="349" unit="m" value="790.0">790 masl</quantity>
</elevation>
; 
<collectorName id="26D3533BFFF9FF8971CFFB9BFCB3FBE7" box="[649,775,1099,1123]" pageId="8" pageNumber="349">M. Kottelat</collectorName>
&amp; 
<collectorName id="26D3533BFFF9FF8973E3FBBBFEE8FB07" box="[165,348,1131,1155]" pageId="8" pageNumber="349">T. Phommavong</collectorName>
, 
<collectingDate id="EFDCE9C5FFF9FF89722FFBBBFD95FB07" box="[361,545,1131,1155]" pageId="8" pageNumber="349" value="2013-01-14">
<date id="FF98102DFFF9FF89722FFBBBFD95FB07" box="[361,545,1131,1155]" pageId="8" pageNumber="349" value="2013-01-14">14 January 2013</date>
</collectingDate>
.
</materialsCitation>
</paragraph>
<paragraph id="8B9936EDFFF9FF8973CEFB7BFE3CF906" blockId="8.[136,775,1195,1667]" pageId="8" pageNumber="349">
<materialsCitation id="3B4E3CB0FFF9FF8973CEFB7BFDEBFB67" ID-GBIF-Occurrence="1611915209" country="Laos" location="Bolaven Plateau" municipality="All" pageId="8" pageNumber="349" specimenCount="1" stateProvince="Champasak Province" typeStatus="paratype">
<emphasis id="B952EAFFFFF9FF8973CEFB7BFEB5FB47" bold="true" box="[136,257,1195,1219]" pageId="8" pageNumber="349">
<typeStatus id="549D884FFFF9FF8973CEFB7BFF48FB47" box="[136,252,1195,1219]" pageId="8" pageNumber="349" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingMunicipality id="6BFDAC97FFF9FF89724CFB7BFE98FB47" box="[266,300,1195,1219]" pageId="8" pageNumber="349">All</collectingMunicipality>
from 
<collectingCountry id="F331767DFFF9FF897235FB7BFE1FFB46" box="[371,427,1195,1218]" name="Laos" pageId="8" pageNumber="349">Laos</collectingCountry>
: 
<collectingRegion id="49E2F80FFFF9FF8972FEFB7BFD15FB47" box="[440,673,1195,1219]" country="Laos" name="Champasak" pageId="8" pageNumber="349">Champasak Province</collectingRegion>
: 
<location id="8EF96036FFF9FF8971E8FB7BFF6FFB67" LSID="urn:lsid:plazi:treatment:038F87FBFFF9FF8F724BFC7AFA8EF9FC:8EF96036FFF9FF8971E8FB7BFF6FFB67" country="Laos" municipality="All" name="Bolaven Plateau" pageId="8" pageNumber="349" stateProvince="Champasak Province">Bolaven Plateau</location>
: CMK 23344, 29, 26.0– 
<quantity id="4CDE9B08FFF9FF89729CFB1AFD83FB66" box="[474,567,1226,1250]" metricMagnitude="-2" metricUnit="m" metricValue="6.65" pageId="8" pageNumber="349" unit="mm" value="66.5">66.5 mm</quantity>
SL
</materialsCitation>
; 
<materialsCitation id="3B4E3CB0FFF9FF89712CFB1BFDE4FA87" ID-GBIF-Occurrence="1611915212" collectingDate="2013-01-14" collectionCode="ZRC" collectorName="M. Kottelat &amp; T. Phommavong" country="Laos" county="Bolaven Plateau" elevation="790" latitude="14.878056" location="Xe Namnoy" longLatPrecision="21" longitude="106.58028" municipality="Houay Xoy" pageId="8" pageNumber="349" specimenCode="ZRC 56223, 5" specimenCount="1" stateProvince="Champasak Province" typeStatus="paratype">
<specimenCode id="DB809E96FFF9FF89712CFB1BFCB6FB66" box="[618,770,1226,1251]" pageId="8" pageNumber="349">ZRC 56223, 5</specimenCode>
, 
<quantity id="4CDE9B08FFF9FF8973CEFB3AFE97FA86" box="[136,291,1258,1282]" metricMagnitude="-2" metricUnit="m" metricValue="4.9" metricValueMax="5.4399999999999995" metricValueMin="4.36" pageId="8" pageNumber="349" unit="mm" value="49.0" valueMax="54.4" valueMin="43.6">43.6–54.4 mm</quantity>
SL; same data as holotype.
</materialsCitation>
— CMK 23320, 25, 25.7–76.0 mm SL; Houay Namkong, a creek tributary of Xe Namnoy; 14°58′14″N 106°33′44″E; 
<quantity id="4CDE9B08FFF9FF897160FAFAFD32FAC7" box="[550,646,1322,1347]" metricMagnitude="2" metricUnit="m" metricValue="7.55" pageId="8" pageNumber="349" unit="m" value="755.0">755 masl</quantity>
; M. Kottelat &amp; T. Phommavong, 
<date id="FF98102DFFF9FF89722BFA9BFD97FAE7" box="[365,547,1355,1379]" pageId="8" pageNumber="349" value="2013-01-12">12 January 2013</date>
. — CMK 23337, 5, 
<quantity id="4CDE9B08FFF9FF8973CEFABAFE98FA06" box="[136,300,1386,1410]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" metricValueMax="5.96" metricValueMin="1.8399999999999999" pageId="8" pageNumber="349" unit="mm" value="39.0" valueMax="59.6" valueMin="18.4">18.4–59.6 mm</quantity>
SL; Houay Xoy, a creek tributary of Xe Namnoy; 14°56′49″N 106°35′02″E; 
<quantity id="4CDE9B08FFF9FF897150FA5BFDC9FA27" box="[534,637,1419,1443]" metricMagnitude="2" metricUnit="m" metricValue="7.709999999999999" pageId="8" pageNumber="349" unit="m" value="771.0">771 masl</quantity>
; M. Kottelat &amp; T. Phommavong, 
<date id="FF98102DFFF9FF89722BFA7BFD97FA47" box="[365,547,1451,1475]" pageId="8" pageNumber="349" value="2013-01-14">14 January 2013</date>
. — CMK 23361, 1, 
<quantity id="4CDE9B08FFF9FF8973CEFA1BFF5DFA66" box="[136,233,1483,1507]" metricMagnitude="-2" metricUnit="m" metricValue="6.859999999999999" pageId="8" pageNumber="349" unit="mm" value="68.6">68.6 mm</quantity>
SL; unnamed creek on road from Tayerkseua to Ban Namtouad; 15°06′32″N 106°35′31″E; 
<quantity id="4CDE9B08FFF9FF897129FA3BFD6DF987" box="[623,729,1515,1539]" metricMagnitude="2" metricUnit="m" metricValue="8.379999999999999" pageId="8" pageNumber="349" unit="m" value="838.0">838 masl</quantity>
; M. Kottelat &amp; T. Phommavong, 
<date id="FF98102DFFF9FF897290F9DAFD27F9A7" box="[470,659,1546,1571]" pageId="8" pageNumber="349" value="2013-01-15">15 January 2013</date>
. — CMK 22363, 1, 51.0 mm SL; Xe Namnoy at bridge downstream of dam site; 15°03′28″N 106°36′10″E; 
<quantity id="4CDE9B08FFF9FF897164F99BFD31F9E7" box="[546,645,1611,1635]" metricMagnitude="2" metricUnit="m" metricValue="7.08" pageId="8" pageNumber="349" unit="m" value="708.0">708 masl</quantity>
; M. Kottelat et al., 
<date id="FF98102DFFF9FF89738BF9BBFE35F907" box="[205,385,1643,1667]" pageId="8" pageNumber="349" value="2011-01-22">22 January 2011</date>
.
</paragraph>
<paragraph id="8B9936EDFFF9FF8973CEF97AFEF7F8C6" blockId="8.[136,775,1706,1859]" pageId="8" pageNumber="349">
<materialsCitation id="3B4E3CB0FFF9FF8973CEF97AFEF7F8C6" ID-GBIF-Occurrence="1611915200" collectingDate="2013-01-13" collectorName="M. Kottelat &amp; T. Phommavong" country="Laos" elevation="833" latitude="15.027222" location="Xe Namnoy" longLatPrecision="21" longitude="106.60445" municipality="Bolaven Plateau" pageId="8" pageNumber="349" specimenCount="1" stateProvince="Champasak Province" typeStatus="holotype">
<emphasis id="B952EAFFFFF9FF8973CEF97AFE4EF946" bold="true" box="[136,506,1706,1730]" pageId="8" pageNumber="349">
Additional material (non 
<typeStatus id="549D884FFFF9FF8972E9F97AFE59F946" box="[431,493,1706,1730]" pageId="8" pageNumber="349">types</typeStatus>
).
</emphasis>
CMK 24737, 2 (ethanol fxed); 
<collectingCountry id="F331767DFFF9FF89739CF91BFEA0F966" box="[218,276,1739,1762]" name="Laos" pageId="8" pageNumber="349">Laos</collectingCountry>
: 
<collectingRegion id="49E2F80FFFF9FF897267F91BFDA5F967" box="[289,529,1739,1763]" country="Laos" name="Champasak" pageId="8" pageNumber="349">Champasak Province</collectingRegion>
: 
<collectingMunicipality id="6BFDAC97FFF9FF897159F91BFD6EF967" box="[543,730,1739,1763]" pageId="8" pageNumber="349">Bolaven Plateau</collectingMunicipality>
: 
<location id="8EF96036FFF9FF8971AEF91BFF53F887" LSID="urn:lsid:plazi:treatment:038F87FBFFF9FF8F724BFC7AFA8EF9FC:8EF96036FFF9FF8971AEF91BFF53F887" country="Laos" latitude="15.027222" longLatPrecision="21" longitude="106.60445" municipality="Bolaven Plateau" name="Xe Namnoy" pageId="8" pageNumber="349" stateProvince="Champasak Province">Xe Namnoy</location>
, frst rapids downstream of dam site; 
<geoCoordinate id="EE12502AFFF9FF8971CEF93AFCB3F886" box="[648,775,1769,1795]" direction="north" orientation="latitude" pageId="8" pageNumber="349" precision="15" value="15.027222">15°01′38″N</geoCoordinate>
<geoCoordinate id="EE12502AFFF9FF8973CEF8DBFEA3F8A6" box="[136,279,1801,1827]" direction="east" orientation="longitude" pageId="8" pageNumber="349" precision="15" value="106.60445">106°36′16″E</geoCoordinate>
; 
<elevation id="000BD1DEFFF9FF897260F8DBFE24F8A7" box="[294,400,1803,1827]" metricMagnitude="2" metricUnit="m" metricValue="8.33" pageId="8" pageNumber="349" unit="m" value="833.0">
<quantity id="4CDE9B08FFF9FF897260F8DBFE24F8A7" box="[294,400,1803,1827]" metricMagnitude="2" metricUnit="m" metricValue="8.33" pageId="8" pageNumber="349" unit="m" value="833.0">833 masl</quantity>
</elevation>
; 
<collectorName id="26D3533BFFF9FF8972D8F8DBFDABF8A7" box="[414,543,1803,1827]" pageId="8" pageNumber="349">M. Kottelat</collectorName>
&amp; 
<collectorName id="26D3533BFFF9FF89710EF8DBFCB7F8A7" box="[584,771,1803,1827]" pageId="8" pageNumber="349">T. Phommavong</collectorName>
, 
<collectingDate id="EFDCE9C5FFF9FF8973CEF8FBFE88F8C7" box="[136,316,1835,1859]" pageId="8" pageNumber="349" value="2013-01-13">
<date id="FF98102DFFF9FF8973CEF8FBFE88F8C7" box="[136,316,1835,1859]" pageId="8" pageNumber="349" value="2013-01-13">13 January 2013</date>
</collectingDate>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33C6566FFF9FF8973CEF8BAFA06FAE7" pageId="8" pageNumber="349" type="diagnosis">
<paragraph id="8B9936EDFFF9FF8973CEF8BAFBE2FBE0" blockId="8.[136,775,1898,2115]" lastBlockId="8.[835,1475,939,1124]" pageId="8" pageNumber="349">
<emphasis id="B952EAFFFFF9FF8973CEF8BAFF4BF806" bold="true" box="[136,255,1898,1922]" pageId="8" pageNumber="349">Diagnosis.</emphasis>
<taxonomicName id="4C264D6EFFF9FF89724FF8BBFE51F807" authorityName="Maurice Kottelat" authorityYear="2017" box="[265,485,1899,1923]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="8" pageNumber="349" phylum="Chordata" rank="species" species="klydonion">
<emphasis id="B952EAFFFFF9FF89724FF8BBFE51F807" box="[265,485,1899,1923]" italics="true" pageId="8" pageNumber="349">Schistura klydonion</emphasis>
</taxonomicName>
is distinguished from the other species of the genus in Southeast Asia by the following combination of characters: relatively large size (up to at least 
<quantity id="4CDE9B08FFF9FF8973CEF81BFF60F866" box="[136,212,1995,2019]" metricMagnitude="-2" metricUnit="m" metricValue="7.6" pageId="8" pageNumber="349" unit="mm" value="76.0">76 mm</quantity>
SL); body with a midlateral row of 12–21 bars not reaching the dorsal midline, alternating with a middorsal row of saddles or small blotches, and leaving a pale zigzag line between the two rows; lips with a few sparsely set papillae; and black pattern at base of caudal fn made of a vertically elongated blotch, usually with a median constriction at level of lateral line, and a smaller black blotch at base of upper simple and 2–3 posterior procurrent rays, sometimes also a patch of brown pigments on base of lower unbranched and posterior procurrent rays.
</paragraph>
<paragraph id="8B9936EDFFF9FF897005FB5BFA06FAE7" blockId="8.[835,1475,1163,1380]" pageId="8" pageNumber="349">Additional diagnostic characters, not unique to the species are: 8½ branched dorsal-fn rays; no known sexual dimorphism; axillary pelvic lobe rudimentary or small, free; marked dorsal and ventral keels on posterior half of caudal peduncle; depth of caudal peduncle 1.1–1.5 times in its length; pelvic-fn origin about under dorsal-fn origin; 9+8 branched caudal-fn rays; no median notch in upper lip; lateral line complete.</paragraph>
</subSubSection>
<subSubSection id="C33C6566FFF9FF8B7005FA5BFB48F9EA" lastPageId="10" lastPageNumber="351" pageId="8" pageNumber="349" type="description">
<paragraph id="8B9936EDFFF9FF897005FA5BFBDFF827" blockId="8.[835,1475,1419,1955]" pageId="8" pageNumber="349">
<emphasis id="B952EAFFFFF9FF897005FA5BFC6FFA27" bold="true" box="[835,987,1419,1443]" pageId="8" pageNumber="349">Description.</emphasis>
See 
<figureCitation id="131D2A68FFF9FF89775CFA5BFB1CFA27" box="[1050,1192,1419,1443]" captionStart-0="Fig. 13" captionStart-1="Fig. 14" captionStart-2="Fig. 15" captionStartId-0="10.[136,171,968,990]" captionStartId-1="11.[113,148,2066,2088]" captionStartId-2="12.[135,170,1748,1770]" captionTargetBox-0="[157,1440,218,903]" captionTargetBox-1="[222,1316,211,2036]" captionTargetBox-2="[142,1442,241,1713]" captionTargetId-0="figure@10.[136,1474,189,951]" captionTargetId-1="figure@11.[222,1341,189,2049]" captionTargetId-2="figure@12.[135,1473,189,1731]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionText-0="Fig. 13. Schistura klydonion, MHNG 2767.085, 75.8 mm SL, holotype; Laos: Xe Namnoy on Bolaven Plateau (right side, reversed)." captionText-1="Fig. 14. Schistura klydonion, paratypes; Laos: Xe Namnoy on Bolaven Plateau; a, CMK 23344, 57.7 mm SL; b, CMK 23344, 61.9 mm SL; and c, CMK 23320, 76.0 mm SL." captionText-2="Fig. 15. Schistura klydonion, paratypes; Laos: Xe Namnoy on Bolaven Plateau; a, CMK 23344, 26.4 mm SL; b, CMK 23344, 27.3 mm SL; c, CMK 23344, 28.1 mm SL; and d, CMK 23320, 29.0 mm SL." httpUri-0="https://zenodo.org/record/886296/files/figure.png" httpUri-1="https://zenodo.org/record/886298/files/figure.png" httpUri-2="https://zenodo.org/record/886300/files/figure.png" pageId="8" pageNumber="349">Figs. 13–15</figureCitation>
for general appearance and 
<tableCitation id="C6A40356FFF9FF897030FA7BFC7AFA47" box="[886,974,1451,1475]" captionStart="Table 2" captionStartId="9.[113,168,191,213]" captionText="Table 2. Morphometric data of Schistura klydonion (n = 11; MHNG 2767.085, holotype; CMK 23344, 5; CMK 23320, 5). Ranges and means include holotype data." pageId="8" pageNumber="349">Table 2</tableCitation>
for morphometric data of 
<typeStatus id="549D884FFFF9FF897641FA7BFADDFA47" box="[1287,1385,1451,1475]" pageId="8" pageNumber="349" type="holotype">holotype</typeStatus>
and 
<specimenCount id="9D20FD64FFF9FF8976E1FA7BFC01FA60" pageId="8" pageNumber="349" type="generic" typeStatus="paratypes">10 paratypes</specimenCount>
. An elongate nemacheiline with body depth slowly increasing up to slightly in front of dorsal-fn origin. Behind dorsal fn, body depth almost uniform until shortly in front of caudal-fn base, then increasing to caudal-fn base. Dorsal profle continuous between head and body. Head slightly depressed; body slightly compressed anteriorly to compressed posteriorly. Interorbital area fat. In lateral view, eye fushed with or slightly protruding over dorsal profle of head. Cheeks not swollen. Snout pointed but rounded at tip. Depth of caudal peduncle 1.1–1.5 times in its length, deeper posteriorly. Marked dorsal keel on posterior fourth of post-dorsal area (posterior half of caudal peduncle) and ventral keel on posterior half of caudal peduncle. Dorsal keel continuous with upper margin of caudal fn. Largest recorded size 76.0 mm SL.
</paragraph>
<paragraph id="8B9936EDFFF9FF887005F81BFDF6F8DF" blockId="8.[835,1474,1993,2116]" lastBlockId="9.[113,753,1539,1883]" lastPageId="9" lastPageNumber="350" pageId="8" pageNumber="349">Dorsal fn with 4 unbranched and 8½ branched rays; distal margin slightly convex; second branched ray longest. Pectoral fn with 1 unbranched and 9 (7*) or 10 (4) branched rays (including small last ray, usually unbranched), rounded, reaching about halfway to pelvic-fin base; no ray with flamentous extension. Pelvic fn with 1 unbranched and 7 branched rays (including small last ray, usually unbranched); reaching almost to anus; rounded; posterior margin convex; origin at vertical through dorsal-fn origin or below base of unbranched dorsal-fn rays 2–3. Axillary pelvic lobe present, rudimentary to small, free. Anus situated about 1.5–2 eye diameters in front of anal fn, behind posterior extremity of pelvic-fn. Anal fn with 3 unbranched and 5½ branched rays; distal margin convex. Caudal fn with 9+8 branched rays; emarginate, lobes rounded, subequal.</paragraph>
<caption id="DF596665FFF8FF887337FF6FFE3BFF76" pageId="9" pageNumber="350">
<paragraph id="8B9936EDFFF8FF887337FF6FFE3BFF76" blockId="9.[113,1452,191,242]" pageId="9" pageNumber="350">
Table 2. Morphometric data of 
<taxonomicName id="4C264D6EFFF8FF8872E8FF6FFDC5FF50" authorityName="Maurice Kottelat" authorityYear="2017" box="[430,625,191,212]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="9" pageNumber="350" phylum="Chordata" rank="species" species="klydonion">
<emphasis id="B952EAFFFFF8FF8872E8FF6FFDC5FF50" box="[430,625,191,212]" italics="true" pageId="9" pageNumber="350">Schistura klydonion</emphasis>
</taxonomicName>
(n=11; MHNG 2767.085, holotype; CMK 23344, 5; CMK 23320, 5). Ranges and means include holotype data.
</paragraph>
</caption>
<paragraph id="8B9936EDFFF8FF8871F5FEF4FA9DFA37" pageId="9" pageNumber="350">
<table id="F926C44DFFF8007E73C6FEF4FA8EFA30" box="[128,1338,292,1460]" colsContinueFrom="4.[150,1356,292,1460]" gridcols="4" gridrows="35" pageId="9" pageNumber="350">
<tr id="351634AFFFF8007E73C6FEF4FA8EFEBD" box="[128,1338,292,313]" gridrow="0" pageId="9" pageNumber="350" rowspan-0="1">
<th id="76C75DD3FFF8007E71F5FEF4FCA4FEBD" box="[691,784,292,313]" gridcol="1" gridrow="0" pageId="9" pageNumber="350">
<emphasis id="B952EAFFFFF8FF8871F5FEF4FCA4FEBD" bold="true" box="[691,784,292,313]" pageId="9" pageNumber="350">Holotype</emphasis>
</th>
<th id="76C75DD3FFF8007E7085FEF4FB8FFEBD" box="[963,1083,292,313]" gridcol="2" gridrow="0" pageId="9" pageNumber="350">
<emphasis id="B952EAFFFFF8FF887098FEF4FBABFEBD" bold="true" box="[990,1055,292,313]" pageId="9" pageNumber="350">Range</emphasis>
</th>
<th id="76C75DD3FFF8007E77B9FEF4FA8EFEBD" box="[1279,1338,292,313]" gridcol="3" gridrow="0" pageId="9" pageNumber="350">
<emphasis id="B952EAFFFFF8FF8877B9FEF4FA8EFEBD" bold="true" box="[1279,1338,292,313]" pageId="9" pageNumber="350">Mean</emphasis>
</th>
</tr>
<tr id="351634AFFFF8007E73C6FE8AFA8EFEF4" box="[128,1338,346,368]" gridrow="1" pageId="9" pageNumber="350" rowspan-3="1">
<th id="76C75DD3FFF8007E73C6FE8AFE67FEF4" box="[128,467,346,368]" gridcol="0" gridrow="1" pageId="9" pageNumber="350">Standard length (mm)</th>
<td id="76C75DD3FFF8007E71F5FE8AFCA4FEF4" box="[691,784,346,368]" gridcol="1" gridrow="1" pageId="9" pageNumber="350">75.8</td>
<td id="76C75DD3FFF8007E7085FE8AFB8FFEF4" box="[963,1083,346,368]" gridcol="2" gridrow="1" pageId="9" pageNumber="350">52.7–76.0</td>
</tr>
<tr id="351634AFFFF8007E73C6FEAAFA8EFE14" box="[128,1338,378,400]" gridrow="2" pageId="9" pageNumber="350" rowspan-3="1">
<th id="76C75DD3FFF8007E73C6FEAAFE67FE14" box="[128,467,378,400]" gridcol="0" gridrow="2" pageId="9" pageNumber="350">Total length (mm)</th>
<td id="76C75DD3FFF8007E71F5FEAAFCA4FE14" box="[691,784,378,400]" gridcol="1" gridrow="2" pageId="9" pageNumber="350">90.9</td>
<td id="76C75DD3FFF8007E7085FEAAFB8FFE14" box="[963,1083,378,400]" gridcol="2" gridrow="2" pageId="9" pageNumber="350">63.7–91.9</td>
</tr>
<tr id="351634AFFFF8007E73C6FE78FA8EFE3A" box="[128,1338,424,446]" gridrow="3" pageId="9" pageNumber="350" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="76C75DD3FFF8007E73C6FE78FE67FE3A" box="[128,467,424,446]" gridcol="0" gridrow="3" pageId="9" pageNumber="350">
<emphasis id="B952EAFFFFF8FF8873C6FE78FE07FE39" bold="true" box="[128,435,424,446]" pageId="9" pageNumber="350">In percent of standard length</emphasis>
</th>
</tr>
<tr id="351634AFFFF8007E73C6FE18FA8EFE5A" box="[128,1338,456,478]" gridrow="4" pageId="9" pageNumber="350">
<th id="76C75DD3FFF8007E73C6FE18FE67FE5A" box="[128,467,456,478]" gridcol="0" gridrow="4" pageId="9" pageNumber="350">Total length</th>
<td id="76C75DD3FFF8007E71F5FE18FCA4FE5A" box="[691,784,456,478]" gridcol="1" gridrow="4" pageId="9" pageNumber="350">119.9</td>
<td id="76C75DD3FFF8007E7085FE18FB8FFE5A" box="[963,1083,456,478]" gridcol="2" gridrow="4" pageId="9" pageNumber="350">119.4–122.3</td>
<td id="76C75DD3FFF8007E77B9FE18FA8EFE5A" box="[1279,1338,456,478]" gridcol="3" gridrow="4" pageId="9" pageNumber="350">121.3</td>
</tr>
<tr id="351634AFFFF8007E73C6FE38FA8EFE7A" box="[128,1338,488,510]" gridrow="5" pageId="9" pageNumber="350">
<th id="76C75DD3FFF8007E73C6FE38FE67FE7A" box="[128,467,488,510]" gridcol="0" gridrow="5" pageId="9" pageNumber="350">Head length (dorsal)</th>
<td id="76C75DD3FFF8007E71F5FE38FCA4FE7A" box="[691,784,488,510]" gridcol="1" gridrow="5" pageId="9" pageNumber="350">20.8</td>
<td id="76C75DD3FFF8007E7085FE38FB8FFE7A" box="[963,1083,488,510]" gridcol="2" gridrow="5" pageId="9" pageNumber="350">19.9–22.1</td>
<td id="76C75DD3FFF8007E77B9FE38FA8EFE7A" box="[1279,1338,488,510]" gridcol="3" gridrow="5" pageId="9" pageNumber="350">20.8</td>
</tr>
<tr id="351634AFFFF8007E73C6FDD7FA8EFD99" box="[128,1338,519,541]" gridrow="6" pageId="9" pageNumber="350">
<th id="76C75DD3FFF8007E73C6FDD7FE67FD99" box="[128,467,519,541]" gridcol="0" gridrow="6" pageId="9" pageNumber="350">Head length (lateral)</th>
<td id="76C75DD3FFF8007E71F5FDD7FCA4FD99" box="[691,784,519,541]" gridcol="1" gridrow="6" pageId="9" pageNumber="350">25.1</td>
<td id="76C75DD3FFF8007E7085FDD7FB8FFD99" box="[963,1083,519,541]" gridcol="2" gridrow="6" pageId="9" pageNumber="350">23.2–25.1</td>
<td id="76C75DD3FFF8007E77B9FDD7FA8EFD99" box="[1279,1338,519,541]" gridcol="3" gridrow="6" pageId="9" pageNumber="350">24.2</td>
</tr>
<tr id="351634AFFFF8007E73C6FDF7FA8EFDB9" box="[128,1338,551,573]" gridrow="7" pageId="9" pageNumber="350">
<th id="76C75DD3FFF8007E73C6FDF7FE67FDB9" box="[128,467,551,573]" gridcol="0" gridrow="7" pageId="9" pageNumber="350">Predorsal length</th>
<td id="76C75DD3FFF8007E71F5FDF7FCA4FDB9" box="[691,784,551,573]" gridcol="1" gridrow="7" pageId="9" pageNumber="350">54.0</td>
<td id="76C75DD3FFF8007E7085FDF7FB8FFDB9" box="[963,1083,551,573]" gridcol="2" gridrow="7" pageId="9" pageNumber="350">53.3–56.5</td>
<td id="76C75DD3FFF8007E77B9FDF7FA8EFDB9" box="[1279,1338,551,573]" gridcol="3" gridrow="7" pageId="9" pageNumber="350">54.4</td>
</tr>
<tr id="351634AFFFF8007E73C6FD97FA8EFDD9" box="[128,1338,583,605]" gridrow="8" pageId="9" pageNumber="350">
<th id="76C75DD3FFF8007E73C6FD97FE67FDD9" box="[128,467,583,605]" gridcol="0" gridrow="8" pageId="9" pageNumber="350">Prepelvic length</th>
<td id="76C75DD3FFF8007E71F5FD97FCA4FDD9" box="[691,784,583,605]" gridcol="1" gridrow="8" pageId="9" pageNumber="350">50.9</td>
<td id="76C75DD3FFF8007E7085FD97FB8FFDD9" box="[963,1083,583,605]" gridcol="2" gridrow="8" pageId="9" pageNumber="350">50.3–52.2</td>
<td id="76C75DD3FFF8007E77B9FD97FA8EFDD9" box="[1279,1338,583,605]" gridcol="3" gridrow="8" pageId="9" pageNumber="350">51.0</td>
</tr>
<tr id="351634AFFFF8007E73C6FDB7FA8EFDF9" box="[128,1338,615,637]" gridrow="9" pageId="9" pageNumber="350">
<th id="76C75DD3FFF8007E73C6FDB7FE67FDF9" box="[128,467,615,637]" gridcol="0" gridrow="9" pageId="9" pageNumber="350">Pre-anus length</th>
<td id="76C75DD3FFF8007E71F5FDB7FCA4FDF9" box="[691,784,615,637]" gridcol="1" gridrow="9" pageId="9" pageNumber="350">69.8</td>
<td id="76C75DD3FFF8007E7085FDB7FB8FFDF9" box="[963,1083,615,637]" gridcol="2" gridrow="9" pageId="9" pageNumber="350">68.1–71.5</td>
<td id="76C75DD3FFF8007E77B9FDB7FA8EFDF9" box="[1279,1338,615,637]" gridcol="3" gridrow="9" pageId="9" pageNumber="350">69.8</td>
</tr>
<tr id="351634AFFFF8007E73C6FD56FA8EFD18" box="[128,1338,646,668]" gridrow="10" pageId="9" pageNumber="350">
<th id="76C75DD3FFF8007E73C6FD56FE67FD18" box="[128,467,646,668]" gridcol="0" gridrow="10" pageId="9" pageNumber="350">Pre-anal length</th>
<td id="76C75DD3FFF8007E71F5FD56FCA4FD18" box="[691,784,646,668]" gridcol="1" gridrow="10" pageId="9" pageNumber="350">75.9</td>
<td id="76C75DD3FFF8007E7085FD56FB8FFD18" box="[963,1083,646,668]" gridcol="2" gridrow="10" pageId="9" pageNumber="350">75.3–78.2</td>
<td id="76C75DD3FFF8007E77B9FD56FA8EFD18" box="[1279,1338,646,668]" gridcol="3" gridrow="10" pageId="9" pageNumber="350">76.8</td>
</tr>
<tr id="351634AFFFF8007E73C6FD76FA8EFD38" box="[128,1338,678,700]" gridrow="11" pageId="9" pageNumber="350">
<th id="76C75DD3FFF8007E73C6FD76FE67FD38" box="[128,467,678,700]" gridcol="0" gridrow="11" pageId="9" pageNumber="350">Head depth</th>
<td id="76C75DD3FFF8007E71F5FD76FCA4FD38" box="[691,784,678,700]" gridcol="1" gridrow="11" pageId="9" pageNumber="350">11.9</td>
<td id="76C75DD3FFF8007E7085FD76FB8FFD38" box="[963,1083,678,700]" gridcol="2" gridrow="11" pageId="9" pageNumber="350">10.7–12.6</td>
<td id="76C75DD3FFF8007E77B9FD76FA8EFD38" box="[1279,1338,678,700]" gridcol="3" gridrow="11" pageId="9" pageNumber="350">11.7</td>
</tr>
<tr id="351634AFFFF8007E73C6FD16FA8EFD58" box="[128,1338,710,732]" gridrow="12" pageId="9" pageNumber="350">
<th id="76C75DD3FFF8007E73C6FD16FE67FD58" box="[128,467,710,732]" gridcol="0" gridrow="12" pageId="9" pageNumber="350">Body depth at dorsal-fn origin</th>
<td id="76C75DD3FFF8007E71F5FD16FCA4FD58" box="[691,784,710,732]" gridcol="1" gridrow="12" pageId="9" pageNumber="350">17.2</td>
<td id="76C75DD3FFF8007E7085FD16FB8FFD58" box="[963,1083,710,732]" gridcol="2" gridrow="12" pageId="9" pageNumber="350">13.9–17.5</td>
<td id="76C75DD3FFF8007E77B9FD16FA8EFD58" box="[1279,1338,710,732]" gridcol="3" gridrow="12" pageId="9" pageNumber="350">16.2</td>
</tr>
<tr id="351634AFFFF8007E73C6FD36FA8EFD78" box="[128,1338,742,764]" gridrow="13" pageId="9" pageNumber="350">
<th id="76C75DD3FFF8007E73C6FD36FE67FD78" box="[128,467,742,764]" gridcol="0" gridrow="13" pageId="9" pageNumber="350">Depth of caudal peduncle</th>
<td id="76C75DD3FFF8007E71F5FD36FCA4FD78" box="[691,784,742,764]" gridcol="1" gridrow="13" pageId="9" pageNumber="350">13.1</td>
<td id="76C75DD3FFF8007E7085FD36FB8FFD78" box="[963,1083,742,764]" gridcol="2" gridrow="13" pageId="9" pageNumber="350">11.2–13.8</td>
<td id="76C75DD3FFF8007E77B9FD36FA8EFD78" box="[1279,1338,742,764]" gridcol="3" gridrow="13" pageId="9" pageNumber="350">12.6</td>
</tr>
<tr id="351634AFFFF8007E73C6FCD5FA8EFC9F" box="[128,1338,773,795]" gridrow="14" pageId="9" pageNumber="350">
<th id="76C75DD3FFF8007E73C6FCD5FE67FC9F" box="[128,467,773,795]" gridcol="0" gridrow="14" pageId="9" pageNumber="350">Length of caudal peduncle</th>
<td id="76C75DD3FFF8007E71F5FCD5FCA4FC9F" box="[691,784,773,795]" gridcol="1" gridrow="14" pageId="9" pageNumber="350">15.9</td>
<td id="76C75DD3FFF8007E7085FCD5FB8FFC9F" box="[963,1083,773,795]" gridcol="2" gridrow="14" pageId="9" pageNumber="350">15.1–17.6</td>
<td id="76C75DD3FFF8007E77B9FCD5FA8EFC9F" box="[1279,1338,773,795]" gridcol="3" gridrow="14" pageId="9" pageNumber="350">16.5</td>
</tr>
<tr id="351634AFFFF8007E73C6FCF5FA8EFCBF" box="[128,1338,805,827]" gridrow="15" pageId="9" pageNumber="350">
<th id="76C75DD3FFF8007E73C6FCF5FE67FCBF" box="[128,467,805,827]" gridcol="0" gridrow="15" pageId="9" pageNumber="350">Head width</th>
<td id="76C75DD3FFF8007E71F5FCF5FCA4FCBF" box="[691,784,805,827]" gridcol="1" gridrow="15" pageId="9" pageNumber="350">16.1</td>
<td id="76C75DD3FFF8007E7085FCF5FB8FFCBF" box="[963,1083,805,827]" gridcol="2" gridrow="15" pageId="9" pageNumber="350">13.3–16.1</td>
<td id="76C75DD3FFF8007E77B9FCF5FA8EFCBF" box="[1279,1338,805,827]" gridcol="3" gridrow="15" pageId="9" pageNumber="350">14.9</td>
</tr>
<tr id="351634AFFFF8007E73C6FC95FA8EFCDF" box="[128,1338,837,859]" gridrow="16" pageId="9" pageNumber="350">
<th id="76C75DD3FFF8007E73C6FC95FE67FCDF" box="[128,467,837,859]" gridcol="0" gridrow="16" pageId="9" pageNumber="350">Body width at dorsal-fn origin</th>
<td id="76C75DD3FFF8007E71F5FC95FCA4FCDF" box="[691,784,837,859]" gridcol="1" gridrow="16" pageId="9" pageNumber="350">12.7</td>
<td id="76C75DD3FFF8007E7085FC95FB8FFCDF" box="[963,1083,837,859]" gridcol="2" gridrow="16" pageId="9" pageNumber="350">10.8–14.1</td>
<td id="76C75DD3FFF8007E77B9FC95FA8EFCDF" box="[1279,1338,837,859]" gridcol="3" gridrow="16" pageId="9" pageNumber="350">12.5</td>
</tr>
<tr id="351634AFFFF8007E73C6FCB5FA8EFCFF" box="[128,1338,869,891]" gridrow="17" pageId="9" pageNumber="350">
<th id="76C75DD3FFF8007E73C6FCB5FE67FCFF" box="[128,467,869,891]" gridcol="0" gridrow="17" pageId="9" pageNumber="350">Snout length</th>
<td id="76C75DD3FFF8007E71F5FCB5FCA4FCFF" box="[691,784,869,891]" gridcol="1" gridrow="17" pageId="9" pageNumber="350">10.3</td>
<td id="76C75DD3FFF8007E7085FCB5FB8FFCFF" box="[963,1083,869,891]" gridcol="2" gridrow="17" pageId="9" pageNumber="350">8.8–10.7</td>
<td id="76C75DD3FFF8007E77B9FCB5FA8EFCFF" box="[1279,1338,869,891]" gridcol="3" gridrow="17" pageId="9" pageNumber="350">10.0</td>
</tr>
<tr id="351634AFFFF8007E73C6FC54FA8EFC1F" box="[128,1338,900,923]" gridrow="18" pageId="9" pageNumber="350">
<th id="76C75DD3FFF8007E73C6FC54FE67FC1F" box="[128,467,900,923]" gridcol="0" gridrow="18" pageId="9" pageNumber="350">Eye diameter</th>
<td id="76C75DD3FFF8007E71F5FC54FCA4FC1F" box="[691,784,900,923]" gridcol="1" gridrow="18" pageId="9" pageNumber="350">3.8</td>
<td id="76C75DD3FFF8007E7085FC54FB8FFC1F" box="[963,1083,900,923]" gridcol="2" gridrow="18" pageId="9" pageNumber="350">3.6–4.7</td>
<td id="76C75DD3FFF8007E77B9FC54FA8EFC1F" box="[1279,1338,900,923]" gridcol="3" gridrow="18" pageId="9" pageNumber="350">4.3</td>
</tr>
<tr id="351634AFFFF8007E73C6FC74FA8EFC3E" box="[128,1338,932,954]" gridrow="19" pageId="9" pageNumber="350">
<th id="76C75DD3FFF8007E73C6FC74FE67FC3E" box="[128,467,932,954]" gridcol="0" gridrow="19" pageId="9" pageNumber="350">Interorbital width</th>
<td id="76C75DD3FFF8007E71F5FC74FCA4FC3E" box="[691,784,932,954]" gridcol="1" gridrow="19" pageId="9" pageNumber="350">7.2</td>
<td id="76C75DD3FFF8007E7085FC74FB8FFC3E" box="[963,1083,932,954]" gridcol="2" gridrow="19" pageId="9" pageNumber="350">6.3–7.9</td>
<td id="76C75DD3FFF8007E77B9FC74FA8EFC3E" box="[1279,1338,932,954]" gridcol="3" gridrow="19" pageId="9" pageNumber="350">7.1</td>
</tr>
<tr id="351634AFFFF8007E73C6FC14FA8EFC5E" box="[128,1338,964,986]" gridrow="20" pageId="9" pageNumber="350">
<th id="76C75DD3FFF8007E73C6FC14FE67FC5E" box="[128,467,964,986]" gridcol="0" gridrow="20" pageId="9" pageNumber="350">Length of dorsal fn</th>
<td id="76C75DD3FFF8007E71F5FC14FCA4FC5E" box="[691,784,964,986]" gridcol="1" gridrow="20" pageId="9" pageNumber="350">15.7</td>
<td id="76C75DD3FFF8007E7085FC14FB8FFC5E" box="[963,1083,964,986]" gridcol="2" gridrow="20" pageId="9" pageNumber="350">15.7–19.0</td>
<td id="76C75DD3FFF8007E77B9FC14FA8EFC5E" box="[1279,1338,964,986]" gridcol="3" gridrow="20" pageId="9" pageNumber="350">17.6</td>
</tr>
<tr id="351634AFFFF8007E73C6FC34FA8EFC7E" box="[128,1338,996,1018]" gridrow="21" pageId="9" pageNumber="350">
<th id="76C75DD3FFF8007E73C6FC34FE67FC7E" box="[128,467,996,1018]" gridcol="0" gridrow="21" pageId="9" pageNumber="350">Length of upper caudal-fn lobe</th>
<td id="76C75DD3FFF8007E71F5FC34FCA4FC7E" box="[691,784,996,1018]" gridcol="1" gridrow="21" pageId="9" pageNumber="350">19.7</td>
<td id="76C75DD3FFF8007E7085FC34FB8FFC7E" box="[963,1083,996,1018]" gridcol="2" gridrow="21" pageId="9" pageNumber="350">19.7–22.8</td>
<td id="76C75DD3FFF8007E77B9FC34FA8EFC7E" box="[1279,1338,996,1018]" gridcol="3" gridrow="21" pageId="9" pageNumber="350">21.4</td>
</tr>
<tr id="351634AFFFF8007E73C6FBD4FA8EFB9E" box="[128,1338,1028,1050]" gridrow="22" pageId="9" pageNumber="350">
<th id="76C75DD3FFF8007E73C6FBD4FE67FB9E" box="[128,467,1028,1050]" gridcol="0" gridrow="22" pageId="9" pageNumber="350">Length of median caudal-fn rays</th>
<td id="76C75DD3FFF8007E71F5FBD4FCA4FB9E" box="[691,784,1028,1050]" gridcol="1" gridrow="22" pageId="9" pageNumber="350">14.3</td>
<td id="76C75DD3FFF8007E7085FBD4FB8FFB9E" box="[963,1083,1028,1050]" gridcol="2" gridrow="22" pageId="9" pageNumber="350">14.3–19.9</td>
<td id="76C75DD3FFF8007E77B9FBD4FA8EFB9E" box="[1279,1338,1028,1050]" gridcol="3" gridrow="22" pageId="9" pageNumber="350">17.6</td>
</tr>
<tr id="351634AFFFF8007E73C6FBF3FA8EFBBD" box="[128,1338,1059,1081]" gridrow="23" pageId="9" pageNumber="350">
<th id="76C75DD3FFF8007E73C6FBF3FE67FBBD" box="[128,467,1059,1081]" gridcol="0" gridrow="23" pageId="9" pageNumber="350">Length of lower caudal-fn lobe</th>
<td id="76C75DD3FFF8007E71F5FBF3FCA4FBBD" box="[691,784,1059,1081]" gridcol="1" gridrow="23" pageId="9" pageNumber="350">20.0</td>
<td id="76C75DD3FFF8007E7085FBF3FB8FFBBD" box="[963,1083,1059,1081]" gridcol="2" gridrow="23" pageId="9" pageNumber="350">20.0–23.6</td>
<td id="76C75DD3FFF8007E77B9FBF3FA8EFBBD" box="[1279,1338,1059,1081]" gridcol="3" gridrow="23" pageId="9" pageNumber="350">22.0</td>
</tr>
<tr id="351634AFFFF8007E73C6FB93FA8EFBDD" box="[128,1338,1091,1113]" gridrow="24" pageId="9" pageNumber="350">
<th id="76C75DD3FFF8007E73C6FB93FE67FBDD" box="[128,467,1091,1113]" gridcol="0" gridrow="24" pageId="9" pageNumber="350">Length of anal fn</th>
<td id="76C75DD3FFF8007E71F5FB93FCA4FBDD" box="[691,784,1091,1113]" gridcol="1" gridrow="24" pageId="9" pageNumber="350">15.6</td>
<td id="76C75DD3FFF8007E7085FB93FB8FFBDD" box="[963,1083,1091,1113]" gridcol="2" gridrow="24" pageId="9" pageNumber="350">15.4–17.6</td>
<td id="76C75DD3FFF8007E77B9FB93FA8EFBDD" box="[1279,1338,1091,1113]" gridcol="3" gridrow="24" pageId="9" pageNumber="350">16.5</td>
</tr>
<tr id="351634AFFFF8007E73C6FBB3FA8EFBFD" box="[128,1338,1123,1145]" gridrow="25" pageId="9" pageNumber="350">
<th id="76C75DD3FFF8007E73C6FBB3FE67FBFD" box="[128,467,1123,1145]" gridcol="0" gridrow="25" pageId="9" pageNumber="350">Length of pelvic fn</th>
<td id="76C75DD3FFF8007E71F5FBB3FCA4FBFD" box="[691,784,1123,1145]" gridcol="1" gridrow="25" pageId="9" pageNumber="350">15.0</td>
<td id="76C75DD3FFF8007E7085FBB3FB8FFBFD" box="[963,1083,1123,1145]" gridcol="2" gridrow="25" pageId="9" pageNumber="350">15.0–16.8</td>
<td id="76C75DD3FFF8007E77B9FBB3FA8EFBFD" box="[1279,1338,1123,1145]" gridcol="3" gridrow="25" pageId="9" pageNumber="350">15.9</td>
</tr>
<tr id="351634AFFFF8007E73C6FB53FA8EFB1D" box="[128,1338,1155,1177]" gridrow="26" pageId="9" pageNumber="350">
<th id="76C75DD3FFF8007E73C6FB53FE67FB1D" box="[128,467,1155,1177]" gridcol="0" gridrow="26" pageId="9" pageNumber="350">Length of pectoral fn</th>
<td id="76C75DD3FFF8007E71F5FB53FCA4FB1D" box="[691,784,1155,1177]" gridcol="1" gridrow="26" pageId="9" pageNumber="350">16.8</td>
<td id="76C75DD3FFF8007E7085FB53FB8FFB1D" box="[963,1083,1155,1177]" gridcol="2" gridrow="26" pageId="9" pageNumber="350">16.8–18.9</td>
<td id="76C75DD3FFF8007E77B9FB53FA8EFB1D" box="[1279,1338,1155,1177]" gridcol="3" gridrow="26" pageId="9" pageNumber="350">17.8</td>
</tr>
<tr id="351634AFFFF8007E73C6FB61FA8EFB43" box="[128,1338,1201,1223]" gridrow="27" pageId="9" pageNumber="350" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="76C75DD3FFF8007E73C6FB61FE67FB43" box="[128,467,1201,1223]" gridcol="0" gridrow="27" pageId="9" pageNumber="350">
<emphasis id="B952EAFFFFF8FF8873C6FB61FE64FB42" bold="true" box="[128,464,1201,1223]" pageId="9" pageNumber="350">In percent of dorsal head length</emphasis>
</th>
</tr>
<tr id="351634AFFFF8007E73C6FB01FA8EFB63" box="[128,1338,1233,1255]" gridrow="28" pageId="9" pageNumber="350">
<th id="76C75DD3FFF8007E73C6FB01FE67FB63" box="[128,467,1233,1255]" gridcol="0" gridrow="28" pageId="9" pageNumber="350">Snout length</th>
<td id="76C75DD3FFF8007E71F5FB01FCA4FB63" box="[691,784,1233,1255]" gridcol="1" gridrow="28" pageId="9" pageNumber="350">50</td>
<td id="76C75DD3FFF8007E7085FB01FB8FFB63" box="[963,1083,1233,1255]" gridcol="2" gridrow="28" pageId="9" pageNumber="350">44–52</td>
<td id="76C75DD3FFF8007E77B9FB01FA8EFB63" box="[1279,1338,1233,1255]" gridcol="3" gridrow="28" pageId="9" pageNumber="350">48</td>
</tr>
<tr id="351634AFFFF8007E73C6FB21FA8EFA83" box="[128,1338,1265,1287]" gridrow="29" pageId="9" pageNumber="350">
<th id="76C75DD3FFF8007E73C6FB21FE67FA83" box="[128,467,1265,1287]" gridcol="0" gridrow="29" pageId="9" pageNumber="350">Eye diameter</th>
<td id="76C75DD3FFF8007E71F5FB21FCA4FA83" box="[691,784,1265,1287]" gridcol="1" gridrow="29" pageId="9" pageNumber="350">18</td>
<td id="76C75DD3FFF8007E7085FB21FB8FFA83" box="[963,1083,1265,1287]" gridcol="2" gridrow="29" pageId="9" pageNumber="350">18–22</td>
<td id="76C75DD3FFF8007E77B9FB21FA8EFA83" box="[1279,1338,1265,1287]" gridcol="3" gridrow="29" pageId="9" pageNumber="350">20</td>
</tr>
<tr id="351634AFFFF8007E73C6FAC0FA8EFAA2" box="[128,1338,1296,1318]" gridrow="30" pageId="9" pageNumber="350">
<th id="76C75DD3FFF8007E73C6FAC0FE67FAA2" box="[128,467,1296,1318]" gridcol="0" gridrow="30" pageId="9" pageNumber="350">Interorbital width</th>
<td id="76C75DD3FFF8007E71F5FAC0FCA4FAA2" box="[691,784,1296,1318]" gridcol="1" gridrow="30" pageId="9" pageNumber="350">35</td>
<td id="76C75DD3FFF8007E7085FAC0FB8FFAA2" box="[963,1083,1296,1318]" gridcol="2" gridrow="30" pageId="9" pageNumber="350">29–40</td>
<td id="76C75DD3FFF8007E77B9FAC0FA8EFAA2" box="[1279,1338,1296,1318]" gridcol="3" gridrow="30" pageId="9" pageNumber="350">34</td>
</tr>
<tr id="351634AFFFF8007E73C6FAEFFA8EFAD1" box="[128,1338,1343,1365]" gridrow="31" pageId="9" pageNumber="350" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="76C75DD3FFF8007E73C6FAEFFE67FAD1" box="[128,467,1343,1365]" gridcol="0" gridrow="31" pageId="9" pageNumber="350">
<emphasis id="B952EAFFFFF8FF8873C6FAEFFE67FAD0" bold="true" box="[128,467,1343,1365]" pageId="9" pageNumber="350">In percent of lateral head length</emphasis>
</th>
</tr>
<tr id="351634AFFFF8007E73C6FA8EFA8EFAF1" box="[128,1338,1374,1397]" gridrow="32" pageId="9" pageNumber="350">
<th id="76C75DD3FFF8007E73C6FA8EFE67FAF1" box="[128,467,1374,1397]" gridcol="0" gridrow="32" pageId="9" pageNumber="350">Snout length</th>
<td id="76C75DD3FFF8007E71F5FA8EFCA4FAF1" box="[691,784,1374,1397]" gridcol="1" gridrow="32" pageId="9" pageNumber="350">41</td>
<td id="76C75DD3FFF8007E7085FA8EFB8FFAF1" box="[963,1083,1374,1397]" gridcol="2" gridrow="32" pageId="9" pageNumber="350">37–45</td>
<td id="76C75DD3FFF8007E77B9FA8EFA8EFAF1" box="[1279,1338,1374,1397]" gridcol="3" gridrow="32" pageId="9" pageNumber="350">41</td>
</tr>
<tr id="351634AFFFF8007E73C6FAAEFA8EFA10" box="[128,1338,1406,1428]" gridrow="33" pageId="9" pageNumber="350">
<th id="76C75DD3FFF8007E73C6FAAEFE67FA10" box="[128,467,1406,1428]" gridcol="0" gridrow="33" pageId="9" pageNumber="350">Eye diameter</th>
<td id="76C75DD3FFF8007E71F5FAAEFCA4FA10" box="[691,784,1406,1428]" gridcol="1" gridrow="33" pageId="9" pageNumber="350">15</td>
<td id="76C75DD3FFF8007E7085FAAEFB8FFA10" box="[963,1083,1406,1428]" gridcol="2" gridrow="33" pageId="9" pageNumber="350">15–20</td>
<td id="76C75DD3FFF8007E77B9FAAEFA8EFA10" box="[1279,1338,1406,1428]" gridcol="3" gridrow="33" pageId="9" pageNumber="350">18</td>
</tr>
<tr id="351634AFFFF8007E73C6FA4EFA8EFA30" box="[128,1338,1438,1460]" gridrow="34" pageId="9" pageNumber="350">
<th id="76C75DD3FFF8007E73C6FA4EFE67FA30" box="[128,467,1438,1460]" gridcol="0" gridrow="34" pageId="9" pageNumber="350">Interorbital width</th>
<td id="76C75DD3FFF8007E71F5FA4EFCA4FA30" box="[691,784,1438,1460]" gridcol="1" gridrow="34" pageId="9" pageNumber="350">29</td>
<td id="76C75DD3FFF8007E7085FA4EFB8FFA30" box="[963,1083,1438,1460]" gridcol="2" gridrow="34" pageId="9" pageNumber="350">26–33</td>
<td id="76C75DD3FFF8007E77B9FA4EFA8EFA30" box="[1279,1338,1438,1460]" gridcol="3" gridrow="34" pageId="9" pageNumber="350">29</td>
</tr>
</table>
</paragraph>
<paragraph id="8B9936EDFFF8FF887337F853FEE0F79E" blockId="9.[113,752,1923,2075]" pageId="9" pageNumber="350">Body entirely scaled. Scales embedded, more deeply in anterior part. Lateral line complete, with 78–92 pores. Cephalic lateral line system with 6 supraorbital, 4 + 10–12 infraorbital, 9–10 preoperculo-mandibular and 3 supratemporal pores.</paragraph>
<paragraph id="8B9936EDFFF8FF88706BF9D3FA86F87F" blockId="9.[813,1452,1539,2043]" pageId="9" pageNumber="350">
Anterior nostril pierced in front side of a pointed fap-like tube. Posterior nostril adjacent to anterior one, about same size. Mouth strongly arched, gape about 2–2.5 times wider than long (
<figureCitation id="131D2A68FFF8FF8870EDF9B3FBB6F9FF" box="[939,1026,1635,1659]" captionStart="Fig. 16" captionStartId="13.[113,149,609,631]" captionTargetBox="[144,709,189,592]" captionTargetId="figure@13.[113,752,189,592]" captionTargetPageId="13" captionText="Fig. 16. Schistura klydonion, CMK 23320, paratype, 76.0 mm SL; mouth." httpUri="https://zenodo.org/record/886302/files/figure.png" pageId="9" pageNumber="350">Fig. 16</figureCitation>
). Lips thick, with sparsely-set, small papillae. Upper lip without median notch (small notch in one individual), with a few furrows near corner of mouth, edge smooth. crenulated. Processus dentiformis present. Lower lip with narrow median interruption; median part with 2–3 sulci, lateral part with a few shallow ridges. Tip of lower jaw not exposed. A shallow median concavity in lower jaw. Barbels with sparsely set papillae similar to those on lips. Inner rostral barbel reaching corner of mouth; outer one reaching vertical of anterior margin of eye. Maxillary barbel reaching middle of postorbital area. Intestine with a loop immediately behind stomach (
<figureCitation id="131D2A68FFF8FF887780F813FAACF85F" box="[1222,1304,1987,2011]" captionStart="Fig. 17" captionStartId="13.[813,848,962,984]" captionTargetBox="[1041,1301,189,927]" captionTargetId="figure@13.[917,1347,163,978]" captionTargetPageId="13" captionText="Fig. 17. Schistura klydonion, CMK 23344, paratype, 66.5 mm SL; digestive tract. Scale bar = 1 mm." httpUri="https://zenodo.org/record/886304/files/figure.png" pageId="9" pageNumber="350">Fig. 17</figureCitation>
). Air bladder without posterior chamber in abdominal cavity.
</paragraph>
<paragraph id="8B9936EDFFF8FF88706BF7F3FB72F7BF" blockId="9.[813,1222,2083,2107]" box="[813,1222,2083,2107]" pageId="9" pageNumber="350">
<emphasis id="B952EAFFFFF8FF88706BF7F3FBA7F7BF" bold="true" box="[813,1043,2083,2107]" pageId="9" pageNumber="350">Sexual dimorphism.</emphasis>
None observed.
</paragraph>
<caption id="DF596665FFFBFF8B73CEFC18FA16FC5A" box="[136,1442,968,990]" httpUri="https://zenodo.org/record/886296/files/figure.png" pageId="10" pageNumber="351" targetBox="[157,1440,218,903]" targetPageId="10">
<paragraph id="8B9936EDFFFBFF8B73CEFC18FA16FC5A" blockId="10.[136,1442,968,990]" box="[136,1442,968,990]" pageId="10" pageNumber="351">
Fig. 13. 
<taxonomicName id="4C264D6EFFFBFF8B739AFC18FE29FC59" authorityName="Maurice Kottelat" authorityYear="2017" box="[220,413,968,989]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="351" phylum="Chordata" rank="species" species="klydonion">
<emphasis id="B952EAFFFFFBFF8B739AFC18FE29FC59" box="[220,413,968,989]" italics="true" pageId="10" pageNumber="351">Schistura klydonion</emphasis>
</taxonomicName>
, MHNG 2767.085, 75.8 mm SL, holotype; Laos: Xe Namnoy on Bolaven Plateau (right side, reversed).
</paragraph>
</caption>
<paragraph id="8B9936EDFFFBFF8B73CEFBC6FEC4F90A" blockId="10.[136,776,1046,1678]" pageId="10" pageNumber="351">
<emphasis id="B952EAFFFFFBFF8B73CEFBC6FEADFBAA" bold="true" box="[136,281,1046,1070]" pageId="10" pageNumber="351">Colouration.</emphasis>
About 3 weeks after fxation. Head and body background colour pale greyish brown, throat and belly whitish; except otherwise stated, markings dark grey to black. Head with a few spots on top and interorbital area. Body with 12–21 bars (4–7 predorsal, 2–4 subdorsal, 6–10 postdorsal), regular in most specimens, slightly wider than interspaces. Bars extending on about median ¾ of body depth, reaching downwards to level of pectoral fns, behind pelvic fns reaching close to ventral midline but not continuous with contralaterals. Anterior bars only slightly wider than posterior ones in specimens with numerous bars (for example 
<typeStatus id="549D884FFFFBFF8B7265FAA6FE33FA0A" box="[291,391,1398,1422]" pageId="10" pageNumber="351" type="holotype">holotype</typeStatus>
; 
<figureCitation id="131D2A68FFFBFF8B72D3FAA6FE53FA0A" box="[405,487,1398,1422]" captionStart="Fig. 13" captionStartId="10.[136,171,968,990]" captionTargetBox="[157,1440,218,903]" captionTargetId="figure@10.[136,1474,189,951]" captionTargetPageId="10" captionText="Fig. 13. Schistura klydonion, MHNG 2767.085, 75.8 mm SL, holotype; Laos: Xe Namnoy on Bolaven Plateau (right side, reversed)." httpUri="https://zenodo.org/record/886296/files/figure.png" pageId="10" pageNumber="351">Fig. 13</figureCitation>
) and wider in specimens with few anterior bars (maybe equivalent to 2 bars) (
<figureCitation id="131D2A68FFFBFF8B7197FA46FF10FA4A" captionStart="Fig. 14" captionStartId="11.[113,148,2066,2088]" captionTargetBox="[222,1316,211,2036]" captionTargetId="figure@11.[222,1341,189,2049]" captionTargetPageId="11" captionText="Fig. 14. Schistura klydonion, paratypes; Laos: Xe Namnoy on Bolaven Plateau; a, CMK 23344, 57.7 mm SL; b, CMK 23344, 61.9 mm SL; and c, CMK 23320, 76.0 mm SL." httpUri="https://zenodo.org/record/886298/files/figure.png" pageId="10" pageNumber="351">Figs. 14</figureCitation>
a, 15c). A row of saddles alternating with bars, leaving a narrow pale line zigzagging between row of saddles and row of bars. Saddles regular (especially in specimens with few bars; 
<figureCitation id="131D2A68FFFBFF8B73A9F9C6FE8FF9AA" box="[239,315,1558,1582]" captionStart="Fig. 14" captionStartId="11.[113,148,2066,2088]" captionTargetBox="[222,1316,211,2036]" captionTargetId="figure@11.[222,1341,189,2049]" captionTargetPageId="11" captionText="Fig. 14. Schistura klydonion, paratypes; Laos: Xe Namnoy on Bolaven Plateau; a, CMK 23344, 57.7 mm SL; b, CMK 23344, 61.9 mm SL; and c, CMK 23320, 76.0 mm SL." httpUri="https://zenodo.org/record/886298/files/figure.png" pageId="10" pageNumber="351">Fig. 14</figureCitation>
a), sometimes divided into spots (especially in specimens with many bars in anterior part of body; 
<figureCitation id="131D2A68FFFBFF8B719BF9E6FF11F9EA" captionStart="Fig. 13" captionStartId="10.[136,171,968,990]" captionTargetBox="[157,1440,218,903]" captionTargetId="figure@10.[136,1474,189,951]" captionTargetPageId="10" captionText="Fig. 13. Schistura klydonion, MHNG 2767.085, 75.8 mm SL, holotype; Laos: Xe Namnoy on Bolaven Plateau (right side, reversed)." httpUri="https://zenodo.org/record/886296/files/figure.png" pageId="10" pageNumber="351">Fig. 13</figureCitation>
), sometimes irregular and occasionally connected with some bars (
<figureCitation id="131D2A68FFFBFF8B7240F9A6FEE3F90A" box="[262,343,1654,1678]" captionStart="Fig. 14" captionStartId="11.[113,148,2066,2088]" captionTargetBox="[222,1316,211,2036]" captionTargetId="figure@11.[222,1341,189,2049]" captionTargetPageId="11" captionText="Fig. 14. Schistura klydonion, paratypes; Laos: Xe Namnoy on Bolaven Plateau; a, CMK 23344, 57.7 mm SL; b, CMK 23344, 61.9 mm SL; and c, CMK 23320, 76.0 mm SL." httpUri="https://zenodo.org/record/886298/files/figure.png" pageId="10" pageNumber="351">Fig. 14</figureCitation>
c).
</paragraph>
<paragraph id="8B9936EDFFFBFF8B73CEF966FE34F78A" blockId="10.[136,776,1718,2062]" pageId="10" pageNumber="351">
A conspicuous black blotch on caudal fn base, vertically elongated, occupying about middle third, usually with a constriction at level of lateral line (
<figureCitation id="131D2A68FFFBFF8B716EF926FDCBF88A" box="[552,639,1782,1806]" captionStart="Fig. 13" captionStartId="10.[136,171,968,990]" captionTargetBox="[157,1440,218,903]" captionTargetId="figure@10.[136,1474,189,951]" captionTargetPageId="10" captionText="Fig. 13. Schistura klydonion, MHNG 2767.085, 75.8 mm SL, holotype; Laos: Xe Namnoy on Bolaven Plateau (right side, reversed)." httpUri="https://zenodo.org/record/886296/files/figure.png" pageId="10" pageNumber="351">Fig. 13</figureCitation>
). A smaller black blotch at base of upper simple ray and 2–3 posterior procurrent rays. Both blotches more or less extensively covered by superfcial brown pigments. Sometimes also a patch of superfcial brown pigments on base of lower unbranched ray and posterior procurrent rays (
<figureCitation id="131D2A68FFFBFF8B71C6F846FD64F82A" box="[640,720,1942,1966]" captionStart="Fig. 14" captionStartId="11.[113,148,2066,2088]" captionTargetBox="[222,1316,211,2036]" captionTargetId="figure@11.[222,1341,189,2049]" captionTargetPageId="11" captionText="Fig. 14. Schistura klydonion, paratypes; Laos: Xe Namnoy on Bolaven Plateau; a, CMK 23344, 57.7 mm SL; b, CMK 23344, 61.9 mm SL; and c, CMK 23320, 76.0 mm SL." httpUri="https://zenodo.org/record/886298/files/figure.png" pageId="10" pageNumber="351">Fig. 14</figureCitation>
b). A triangular unpigmented area over procurrent rays and adjacent areas of caudal peduncle and unbranched principal rays. A faint inner axial stripe.
</paragraph>
<paragraph id="8B9936EDFFFBFF8B7005FBC6FB0AFB2A" blockId="10.[835,1474,1046,1198]" pageId="10" pageNumber="351">Dorsal fn hyaline, with blackish pigments along all rays; a black spot at base of simple rays and frst branched ray; an elongated blotch at base of branched rays 2–8, usually dissociated in a subproximal row of diffuse spots on rays; space between these spots hyaline.</paragraph>
<paragraph id="8B9936EDFFFBFF8B7005FB06FBE3FA2A" blockId="10.[835,1474,1238,1454]" pageId="10" pageNumber="351">Caudal fn orange red, with blackish pigments along rays, especially between branches and appearing as two vague, indistinct vertical rows of spots. Anal fin hyaline, with blackish pigments along rays, especially at inner sides of branches. Pelvic fn hyaline, with few blackish pigments along rays. Pectoral fn hyaline, with blackish pigments along dorsal side of rays.</paragraph>
<paragraph id="8B9936EDFFFBFF8B7005FA06FB48F9EA" blockId="10.[835,1475,1494,1646]" pageId="10" pageNumber="351">
In smallest available specimens (less than about 
<quantity id="4CDE9B08FFFBFF8B7635FA06FA76FA69" box="[1395,1474,1494,1518]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="351" unit="mm" value="30.0">30 mm</quantity>
SL; 
<figureCitation id="131D2A68FFFBFF8B7032FA26FC72F98A" box="[884,966,1526,1550]" captionStart="Fig. 15" captionStartId="12.[135,170,1748,1770]" captionTargetBox="[142,1442,241,1713]" captionTargetId="figure@12.[135,1473,189,1731]" captionTargetPageId="12" captionText="Fig. 15. Schistura klydonion, paratypes; Laos: Xe Namnoy on Bolaven Plateau; a, CMK 23344, 26.4 mm SL; b, CMK 23344, 27.3 mm SL; c, CMK 23344, 28.1 mm SL; and d, CMK 23320, 29.0 mm SL." httpUri="https://zenodo.org/record/886300/files/figure.png" pageId="10" pageNumber="351">Fig. 15</figureCitation>
), adult pattern already distinct, although bars shorter, more rounded, sometimes partly fused to form a very irregular midlateral stripe. Saddles irregular, sometimes fused into an irregular middorsal blotch.
</paragraph>
</subSubSection>
<subSubSection id="C33C6566FFFBFF8B7005F946FAF7F869" pageId="10" pageNumber="351" type="biology_ecology">
<paragraph id="8B9936EDFFFBFF8B7005F946FAF7F869" blockId="10.[835,1475,1686,2030]" pageId="10" pageNumber="351">
<emphasis id="B952EAFFFFFBFF8B7005F946FBBDF92A" bold="true" box="[835,1033,1686,1710]" pageId="10" pageNumber="351">Notes on biology.</emphasis>
One female (CMK 23344, 
<quantity id="4CDE9B08FFFBFF8B7675F946FA26F929" box="[1331,1426,1686,1710]" metricMagnitude="-2" metricUnit="m" metricValue="6.65" pageId="10" pageNumber="351" unit="mm" value="66.5">66.5 mm</quantity>
SL) had apparently almost ripe ova, about 1.0 mm diameter; another one (CMK 23320, 69.0 mm SL) had apparently more advanced ova, about 
<quantity id="4CDE9B08FFFBFF8B7734F926FB7CF889" box="[1138,1224,1782,1806]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="10" pageNumber="351" unit="mm" value="1.4">1.4 mm</quantity>
diameter. In both, the ovaries were narrow, elongated, and ova were few and on one or two rows. 
<taxonomicName id="4C264D6EFFFBFF8B70BDF8E6FB7BF8CA" authorityName="Maurice Kottelat" authorityYear="2017" box="[1019,1231,1846,1870]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="351" phylum="Chordata" rank="species" species="klydonion">
<emphasis id="B952EAFFFFFBFF8B70BDF8E6FB7BF8CA" box="[1019,1231,1846,1870]" italics="true" pageId="10" pageNumber="351">Schistura klydonion</emphasis>
</taxonomicName>
was observed in water bodies ranging from small forest streams 
<quantity id="4CDE9B08FFFBFF8B7656F886FA88F8E9" box="[1296,1340,1878,1902]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="351" unit="m" value="2.0">2 m</quantity>
wide to the Xe Namnoy main river about 
<quantity id="4CDE9B08FFFBFF8B77CFF8A6FB74F809" box="[1161,1216,1910,1934]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="351" unit="m" value="30.0">30 m</quantity>
wide, with fast current, riffes and rapids, clear water, over a gravel to rock bottom (
<figureCitation id="131D2A68FFFBFF8B700DF866FC2EF84A" box="[843,922,1974,1998]" captionStart="Fig. 18" captionStartId="13.[813,850,1508,1530]" captionTargetBox="[813,1452,1066,1490]" captionTargetId="figure@13.[813,1452,1066,1490]" captionTargetPageId="13" captionText="Fig. 18. Type locality of Schistura klydonion; Laos: Bolaven Plateau: Houay Xoy; 14 January 2013." httpUri="https://zenodo.org/record/886306/files/figure.png" pageId="10" pageNumber="351">Fig. 18</figureCitation>
). Other species of 
<taxonomicName id="4C264D6EFFFBFF8B7724F866FB73F84A" box="[1122,1223,1974,1998]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="351" phylum="Chordata" rank="genus">
<emphasis id="B952EAFFFFFBFF8B7724F866FB73F84A" box="[1122,1223,1974,1998]" italics="true" pageId="10" pageNumber="351">Schistura</emphasis>
</taxonomicName>
collected together with 
<taxonomicName id="4C264D6EFFFBFF8B7005F806FC7DF86A" authorityName="Maurice Kottelat" authorityYear="2017" box="[835,969,2006,2030]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="351" phylum="Chordata" rank="species" species="klydonion">
<emphasis id="B952EAFFFFFBFF8B7005F806FC7DF86A" box="[835,969,2006,2030]" italics="true" pageId="10" pageNumber="351">S. klydonion</emphasis>
</taxonomicName>
are 
<taxonomicName id="4C264D6EFFFBFF8B70BDF806FBEAF86A" authorityName="Kottelat" authorityYear="2000" box="[1019,1118,2006,2030]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="351" phylum="Chordata" rank="species" species="tizardi">
<emphasis id="B952EAFFFFFBFF8B70BDF806FBEAF86A" box="[1019,1118,2006,2030]" italics="true" pageId="10" pageNumber="351">S. tizardi</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C264D6EFFFBFF8B77D3F806FA88F86A" authorityName="Kottelat" authorityYear="2000" box="[1173,1340,2006,2030]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="351" phylum="Chordata" rank="species" species="bolavenensis">
<emphasis id="B952EAFFFFFBFF8B77D3F806FA88F86A" box="[1173,1340,2006,2030]" italics="true" pageId="10" pageNumber="351">S. bolavenensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="DF596665FFFAFF8A7337F7C2FE5CF7C0" httpUri="https://zenodo.org/record/886298/files/figure.png" pageId="11" pageNumber="352" targetBox="[222,1316,211,2036]" targetPageId="11">
<paragraph id="8B9936EDFFFAFF8A7337F7C2FE5CF7C0" blockId="11.[113,1450,2066,2117]" pageId="11" pageNumber="352">
Fig. 14. 
<taxonomicName id="4C264D6EFFFAFF8A7382F7C2FE32F7A3" authorityName="Maurice Kottelat" authorityYear="2017" box="[196,390,2066,2087]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="352" phylum="Chordata" rank="species" species="klydonion">
<emphasis id="B952EAFFFFFAFF8A7382F7C2FE32F7A3" box="[196,390,2066,2087]" italics="true" pageId="11" pageNumber="352">Schistura klydonion</emphasis>
</taxonomicName>
, paratypes; Laos: Xe Namnoy on Bolaven Plateau; a, CMK 23344, 57.7 mm SL; b, CMK 23344, 61.9 mm SL; and c, CMK 23320, 76.0 mm SL.
</paragraph>
</caption>
<caption id="DF596665FFFDFF8D73C1F904FC9EF882" httpUri="https://zenodo.org/record/886300/files/figure.png" pageId="12" pageNumber="353" targetBox="[142,1442,241,1713]" targetPageId="12">
<paragraph id="8B9936EDFFFDFF8D73C1F904FC9EF882" blockId="12.[135,1473,1748,1799]" pageId="12" pageNumber="353">
Fig. 15. 
<taxonomicName id="4C264D6EFFFDFF8D739DF904FE28F96D" authorityName="Maurice Kottelat" authorityYear="2017" box="[219,412,1748,1769]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="353" phylum="Chordata" rank="species" species="klydonion">
<emphasis id="B952EAFFFFFDFF8D739DF904FE28F96D" box="[219,412,1748,1769]" italics="true" pageId="12" pageNumber="353">Schistura klydonion</emphasis>
</taxonomicName>
, paratypes; Laos: Xe Namnoy on Bolaven Plateau; a, CMK 23344, 26.4 mm SL; b, CMK 23344, 27.3 mm SL; c, CMK 23344, 28.1 mm SL; and d, CMK 23320, 29.0 mm SL.
</paragraph>
</caption>
<subSubSection id="C33C6566FFFDFF8D73CEF8EEFCB3F8F2" pageId="12" pageNumber="353" type="distribution">
<paragraph id="8B9936EDFFFDFF8D73CEF8EEFCB3F8F2" blockId="12.[136,775,1854,1911]" pageId="12" pageNumber="353">
<emphasis id="B952EAFFFFFDFF8D73CEF8EEFEAEF8D2" bold="true" box="[136,282,1854,1878]" pageId="12" pageNumber="353">Distribution.</emphasis>
<taxonomicName id="4C264D6EFFFDFF8D7262F8EEFE4FF8D2" authorityName="Maurice Kottelat" authorityYear="2017" box="[292,507,1854,1878]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="353" phylum="Chordata" rank="species" species="klydonion">
<emphasis id="B952EAFFFFFDFF8D7262F8EEFE4FF8D2" box="[292,507,1854,1878]" italics="true" pageId="12" pageNumber="353">Schistura klydonion</emphasis>
</taxonomicName>
has been observed only in the Xe Namnoy on the Bolaven Plateau, southern 
<collectingCountry id="F331767DFFFDFF8D718DF88FFCB6F8F2" box="[715,770,1887,1910]" name="Laos" pageId="12" pageNumber="353">Laos</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C33C6566FFFDFF8D73CEF84EFDEFF792" pageId="12" pageNumber="353" type="etymology">
<paragraph id="8B9936EDFFFDFF8D73CEF84EFDEFF792" blockId="12.[136,775,1950,2070]" pageId="12" pageNumber="353">
<emphasis id="B952EAFFFFFDFF8D73CEF84EFEBDF832" bold="true" box="[136,265,1950,1974]" pageId="12" pageNumber="353">Etymology.</emphasis>
From the classical Greek 
<emphasis id="B952EAFFFFFDFF8D7150F84EFD31F832" box="[534,645,1950,1974]" italics="true" pageId="12" pageNumber="353">ΚλυδώνΙΟν</emphasis>
(
<taxonomicName id="4C264D6EFFFDFF8D71D7F84EFD4BF832" authorityName="Maurice Kottelat" authorityYear="2017" box="[657,767,1950,1974]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="353" phylum="Chordata" rank="species" species="klydonion">klydonion</taxonomicName>
) meaning small wave, ripple, undulation; allusion to the wavy stripe running along the fank between the row of saddles and the row of bars. A noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C33C6566FFFDFF8F7005F8EEFC57FE53" lastPageId="14" lastPageNumber="355" pageId="12" pageNumber="353" type="discussion">
<paragraph id="8B9936EDFFFDFF8C7005F8EEFE84FBA7" blockId="12.[835,1475,1854,2102]" lastBlockId="13.[113,753,715,1059]" lastPageId="13" lastPageNumber="354" pageId="12" pageNumber="353">
<emphasis id="B952EAFFFFFDFF8D7005F8EEFC06F8D2" bold="true" box="[835,946,1854,1878]" pageId="12" pageNumber="353">Remarks.</emphasis>
The projecting papillae on the lips of 
<taxonomicName id="4C264D6EFFFDFF8D7679F8EFFA76F8D2" authorityName="Maurice Kottelat" authorityYear="2017" box="[1343,1474,1854,1878]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="353" phylum="Chordata" rank="species" species="klydonion">
<emphasis id="B952EAFFFFFDFF8D7679F8EFFA76F8D2" box="[1343,1474,1854,1878]" italics="true" pageId="12" pageNumber="353">S. klydonion</emphasis>
</taxonomicName>
have not been observed or reported in other species of 
<taxonomicName id="4C264D6EFFFDFF8D7005F8AEFC1FF812" box="[835,939,1918,1942]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="353" phylum="Chordata" rank="genus">
<emphasis id="B952EAFFFFFDFF8D7005F8AEFC1FF812" box="[835,939,1918,1942]" italics="true" pageId="12" pageNumber="353">Schistura</emphasis>
</taxonomicName>
. The colour pattern of 
<taxonomicName id="4C264D6EFFFDFF8D77FCF8AFFAF1F812" authorityName="Maurice Kottelat" authorityYear="2017" box="[1210,1349,1918,1942]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="353" phylum="Chordata" rank="species" species="klydonion">
<emphasis id="B952EAFFFFFDFF8D77FCF8AFFAF1F812" box="[1210,1349,1918,1942]" italics="true" pageId="12" pageNumber="353">S. klydonion</emphasis>
</taxonomicName>
made of a midlateral row of bars alternating with a middorsal row of saddles or small blotches, and leaving a pale zigzag line between them is unique among species of 
<taxonomicName id="4C264D6EFFFDFF8D7671F80EFA16F872" box="[1335,1442,2014,2038]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="353" phylum="Chordata" rank="genus">
<emphasis id="B952EAFFFFFDFF8D7671F80EFA16F872" box="[1335,1442,2014,2038]" italics="true" pageId="12" pageNumber="353">Schistura</emphasis>
</taxonomicName>
in Southeast Asia. Some individuals of 
<taxonomicName id="4C264D6EFFFDFF8D779CF82FFAFCF792" authorityName="Maurice Kottelat" authorityYear="2017" box="[1242,1352,2046,2070]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="353" phylum="Chordata" rank="species" species="colossa">
<emphasis id="B952EAFFFFFDFF8D779CF82FFAFCF792" box="[1242,1352,2046,2070]" italics="true" pageId="12" pageNumber="353">S. colossa</emphasis>
</taxonomicName>
(also from Bolaven Plateau, see above) have a colour pattern made of bars that do not reach the dorsal midline, but in this case they are dissociated into an irregular pattern of blotches and do not leave a pale zigzag line. Further, in 
<taxonomicName id="4C264D6EFFFCFF8C717BFCDCFD76FCA7" authorityName="Maurice Kottelat" authorityYear="2017" box="[573,706,779,803]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="354" phylum="Chordata" rank="species" species="klydonion">
<emphasis id="B952EAFFFFFCFF8C717BFCDCFD76FCA7" box="[573,706,779,803]" italics="true" pageId="13" pageNumber="354">S. klydonion</emphasis>
</taxonomicName>
, the pattern of bars and saddles is quite regular on the whole body at all sizes (vs. becoming irregular in posterior part of body in largest specimens in 
<taxonomicName id="4C264D6EFFFCFF8C72A0FCBCFDECFC07" authorityName="Maurice Kottelat" authorityYear="2017" box="[486,600,875,899]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="354" phylum="Chordata" rank="species" species="colossa">
<emphasis id="B952EAFFFFFCFF8C72A0FCBCFDECFC07" box="[486,600,875,899]" italics="true" pageId="13" pageNumber="354">S. colossa</emphasis>
</taxonomicName>
), the head is longer (lateral head length 23.2–25.1% SL, vs. 21.1–23.8), there is no median notch in the upper lip (vs. presence), and the spots at the base of the caudal fn are separated (vs. form a continuous band reaching close to dorsal and ventral midlines).
</paragraph>
<caption id="DF596665FFFCFF8C7337FDB1FF54FD10" httpUri="https://zenodo.org/record/886302/files/figure.png" pageId="13" pageNumber="354" targetBox="[144,709,189,592]" targetPageId="13">
<paragraph id="8B9936EDFFFCFF8C7337FDB1FF54FD10" blockId="13.[113,753,609,660]" pageId="13" pageNumber="354">
Fig. 16. 
<taxonomicName id="4C264D6EFFFCFF8C738CFDB1FE27FDF2" authorityName="Maurice Kottelat" authorityYear="2017" box="[202,403,609,630]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="354" phylum="Chordata" rank="species" species="klydonion">
<emphasis id="B952EAFFFFFCFF8C738CFDB1FE27FDF2" box="[202,403,609,630]" italics="true" pageId="13" pageNumber="354">Schistura klydonion</emphasis>
</taxonomicName>
, CMK 23320, paratype, 76.0 mm SL; mouth.
</paragraph>
</caption>
<caption id="DF596665FFFCFF8C706BFC12FBCFFC71" httpUri="https://zenodo.org/record/886304/files/figure.png" pageId="13" pageNumber="354" targetBox="[1041,1301,189,927]" targetPageId="13">
<paragraph id="8B9936EDFFFCFF8C706BFC12FBCFFC71" blockId="13.[813,1452,962,1013]" pageId="13" pageNumber="354">
Fig. 17. 
<taxonomicName id="4C264D6EFFFCFF8C703BFC12FB89FC53" authorityName="Maurice Kottelat" authorityYear="2017" box="[893,1085,962,983]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="354" phylum="Chordata" rank="species" species="klydonion">
<emphasis id="B952EAFFFFFCFF8C703BFC12FB89FC53" box="[893,1085,962,983]" italics="true" pageId="13" pageNumber="354">Schistura klydonion</emphasis>
</taxonomicName>
, CMK 23344, paratype, 66.5 mm SL; digestive tract. Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph id="8B9936EDFFFCFF8C7337FB9BFE28F827" blockId="13.[113,753,1099,1956]" pageId="13" pageNumber="354">
<taxonomicName id="4C264D6EFFFCFF8C7337FB9BFEFCFBE7" authorityName="Freyhof &amp; Serov" authorityYear="2001" box="[113,328,1099,1123]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="354" phylum="Chordata" rank="species" species="dalatensis">
<emphasis id="B952EAFFFFFCFF8C7337FB9BFEFCFBE7" box="[113,328,1099,1123]" italics="true" pageId="13" pageNumber="354">Schistura dalatensis</emphasis>
</taxonomicName>
from the 
<collectingRegion id="49E2F80FFFFCFF8C72EAFB9CFDA0FBE7" box="[428,532,1099,1124]" country="Vietnam" name="Dong Nai" pageId="13" pageNumber="354">Dong Nai</collectingRegion>
drainage in 
<collectingCountry id="F331767DFFFCFF8C71D2FB9BFD44FBE7" box="[660,752,1099,1123]" name="Vietnam" pageId="13" pageNumber="354">Vietnam</collectingCountry>
has a colour pattern somewhat similar to that of 
<taxonomicName id="4C264D6EFFFCFF8C7121FBBCFD5DFB07" authorityName="Maurice Kottelat" authorityYear="2017" box="[615,745,1131,1155]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="354" phylum="Chordata" rank="species" species="klydonion">
<emphasis id="B952EAFFFFFCFF8C7121FBBCFD5DFB07" box="[615,745,1131,1155]" italics="true" pageId="13" pageNumber="354">S. klydonion</emphasis>
</taxonomicName>
, but it has a midlateral row of irregular blotches instead of the quite regular bars of 
<taxonomicName id="4C264D6EFFFCFF8C723FFB7CFE49FB47" authorityName="Maurice Kottelat" authorityYear="2017" box="[377,509,1195,1219]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="354" phylum="Chordata" rank="species" species="klydonion">
<emphasis id="B952EAFFFFFCFF8C723FFB7CFE49FB47" box="[377,509,1195,1219]" italics="true" pageId="13" pageNumber="354">S. klydonion</emphasis>
</taxonomicName>
and the predorsal area is marbled by irregular spots, saddles and bars (
<bibRefCitation id="EFB74B1CFFFCFF8C713AFB1BFE9BFA87" author="Freyhof J &amp; Serov DV" journalOrPublisher="Ichthyological Exploration of Freshwaters" pageId="13" pageNumber="354" pagination="133 - 191" part="12" refString="Freyhof J &amp; Serov DV (2001) Nemacheiline loaches from Central Vietnam with descriptions of a new genus and 14 new species (Cypriniformes: Balitoridae). Ichthyological Exploration of Freshwaters, 12 (2): 133 - 191." title="Nemacheiline loaches from Central Vietnam with descriptions of a new genus and 14 new species (Cypriniformes: Balitoridae)" type="journal article" year="2001">Freyhof &amp; Serov, 2001: 151</bibRefCitation>
). Besides, in 
<taxonomicName id="4C264D6EFFFCFF8C7285FB3CFDFCFA87" authorityName="Maurice Kottelat" authorityYear="2017" box="[451,584,1259,1283]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="354" phylum="Chordata" rank="species" species="klydonion">
<emphasis id="B952EAFFFFFCFF8C7285FB3CFDFCFA87" box="[451,584,1259,1283]" italics="true" pageId="13" pageNumber="354">S. klydonion</emphasis>
</taxonomicName>
, the dorsal and ventral keels on the caudal peduncle are more developed resulting in the caudal peduncle clearly higher posteriorly (vs. uniform depth; 11.2–13.8% SL, mean 12.6, vs. 9.7–11.5, mean 10.5), the body is deeper anteriorly than behind dorsal fn (vs. depth very uniform from head to caudalfn base; depth at dorsal-fn origin 13.9–17.5% SL, mean 16.2, vs. 12.5–15.3, mean 12.8), lateral line complete (vs. variable, reaching to middle of anal in most specimens). In 
<taxonomicName id="4C264D6EFFFCFF8C7337F9DCFF48F9A7" authorityName="Maurice Kottelat" authorityYear="2017" box="[113,252,1547,1571]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="354" phylum="Chordata" rank="species" species="klydonion">
<emphasis id="B952EAFFFFFCFF8C7337F9DCFF48F9A7" box="[113,252,1547,1571]" italics="true" pageId="13" pageNumber="354">S. klydonion</emphasis>
</taxonomicName>
, there is a vertically elongated black blotch at the middle of caudal-fn base, a blotch at base of upper simple principal ray and sometimes one at the base of the lower one. In 
<taxonomicName id="4C264D6EFFFCFF8C7254F9BCFE2BF907" authorityName="Freyhof &amp; Serov" authorityYear="2001" box="[274,415,1643,1667]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="354" phylum="Chordata" rank="species" species="dalatensis">
<emphasis id="B952EAFFFFFCFF8C7254F9BCFE2BF907" box="[274,415,1643,1667]" italics="true" pageId="13" pageNumber="354">S. dalatensis</emphasis>
</taxonomicName>
the black marks at caudal-fn base are fused into a continuous bar, including also the spot at the base of the lower simple principal ray, which seems to always be present. In 
<taxonomicName id="4C264D6EFFFCFF8C72C6F91CFDB2F967" authorityName="Maurice Kottelat" authorityYear="2017" box="[384,518,1739,1763]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="354" phylum="Chordata" rank="species" species="klydonion">
<emphasis id="B952EAFFFFFCFF8C72C6F91CFDB2F967" box="[384,518,1739,1763]" italics="true" pageId="13" pageNumber="354">S. klydonion</emphasis>
</taxonomicName>
, the juveniles have a midlateral row of elongated blotches and a middorsal row of small saddles, while juveniles 
<taxonomicName id="4C264D6EFFFCFF8C72A2F8DCFDDAF8A7" authorityName="Freyhof &amp; Serov" authorityYear="2001" box="[484,622,1803,1827]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="354" phylum="Chordata" rank="species" species="dalatensis">
<emphasis id="B952EAFFFFFCFF8C72A2F8DCFDDAF8A7" box="[484,622,1803,1827]" italics="true" pageId="13" pageNumber="354">S. dalatensis</emphasis>
</taxonomicName>
have about 10–12 irregular bars, some reaching the dorsal midline, many widened in a blotch at level of lateral line (morphometric data from 
<bibRefCitation id="EFB74B1CFFFCFF8C739DF8BBFE64F807" author="Freyhof J &amp; Serov DV" box="[219,464,1899,1923]" journalOrPublisher="Ichthyological Exploration of Freshwaters" pageId="13" pageNumber="354" pagination="133 - 191" part="12" refString="Freyhof J &amp; Serov DV (2001) Nemacheiline loaches from Central Vietnam with descriptions of a new genus and 14 new species (Cypriniformes: Balitoridae). Ichthyological Exploration of Freshwaters, 12 (2): 133 - 191." title="Nemacheiline loaches from Central Vietnam with descriptions of a new genus and 14 new species (Cypriniformes: Balitoridae)" type="journal article" year="2001">Freyhof &amp; Serov, 2001</bibRefCitation>
; other characters confrmed on 
<typeStatus id="549D884FFFFCFF8C73D3F85CFF4FF820" box="[149,251,1932,1956]" pageId="13" pageNumber="354" type="paratype">paratypes</typeStatus>
CMK 15999).
</paragraph>
<caption id="DF596665FFFCFF8C706BFA34FB1FF992" httpUri="https://zenodo.org/record/886306/files/figure.png" pageId="13" pageNumber="354" targetBox="[813,1452,1066,1490]" targetPageId="13">
<paragraph id="8B9936EDFFFCFF8C706BFA34FB1FF992" blockId="13.[813,1452,1508,1559]" pageId="13" pageNumber="354">
Fig. 18. Type locality of 
<taxonomicName id="4C264D6EFFFCFF8C777CFA34FAB2FA7D" authorityName="Maurice Kottelat" authorityYear="2017" box="[1082,1286,1508,1529]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="354" phylum="Chordata" rank="species" species="klydonion">
<emphasis id="B952EAFFFFFCFF8C777CFA34FAB2FA7D" box="[1082,1286,1508,1529]" italics="true" pageId="13" pageNumber="354">Schistura klydonion</emphasis>
</taxonomicName>
; Laos: Bolaven Plateau: Houay Xoy; 14 January 2013.
</paragraph>
</caption>
<paragraph id="8B9936EDFFFCFF8C7337F81CFA18F922" blockId="13.[113,752,1995,2116]" lastBlockId="13.[813,1452,1614,1702]" pageId="13" pageNumber="354">
Two other species of 
<taxonomicName id="4C264D6EFFFCFF8C7210F81BFE0FF867" box="[342,443,1995,2019]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="354" phylum="Chordata" rank="genus">
<emphasis id="B952EAFFFFFCFF8C7210F81BFE0FF867" box="[342,443,1995,2019]" italics="true" pageId="13" pageNumber="354">Schistura</emphasis>
</taxonomicName>
have been collected together with 
<taxonomicName id="4C264D6EFFFCFF8C73EFF83CFE9AF787" authorityName="Maurice Kottelat" authorityYear="2017" box="[169,302,2027,2051]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="354" phylum="Chordata" rank="species" species="klydonion">
<emphasis id="B952EAFFFFFCFF8C73EFF83CFE9AF787" box="[169,302,2027,2051]" italics="true" pageId="13" pageNumber="354">S. klydonion</emphasis>
</taxonomicName>
: 
<taxonomicName id="4C264D6EFFFCFF8C7278F83CFE14F787" authorityName="Kottelat" authorityYear="2000" box="[318,416,2027,2051]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="354" phylum="Chordata" rank="species" species="tizardi">
<emphasis id="B952EAFFFFFCFF8C7278F83CFE14F787" box="[318,416,2027,2051]" italics="true" pageId="13" pageNumber="354">S. tizardi</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C264D6EFFFCFF8C7291F83CFDC9F787" authorityName="Kottelat" authorityYear="2000" box="[471,637,2027,2051]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="354" phylum="Chordata" rank="species" species="bolavenensis">
<emphasis id="B952EAFFFFFCFF8C7291F83CFDC9F787" box="[471,637,2027,2051]" italics="true" pageId="13" pageNumber="354">S. bolavenensis</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C264D6EFFFCFF8C71CDF83BFF0CF7A7" authorityName="Kottelat" authorityYear="2000" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="354" phylum="Chordata" rank="species" species="tizardi">
<emphasis id="B952EAFFFFFCFF8C71CDF83BFF0CF7A7" italics="true" pageId="13" pageNumber="354">Schistura tizardi</emphasis>
</taxonomicName>
(
<figureCitation id="131D2A68FFFCFF8C738DF7DBFEAAF7A7" box="[203,286,2059,2083]" captionStart="Fig. 16" captionStartId="13.[113,149,609,631]" captionTargetBox="[144,709,189,592]" captionTargetId="figure@13.[113,752,189,592]" captionTargetPageId="13" captionText="Fig. 16. Schistura klydonion, CMK 23320, paratype, 76.0 mm SL; mouth." httpUri="https://zenodo.org/record/886302/files/figure.png" pageId="13" pageNumber="354">Fig. 16</figureCitation>
) has a distinctive appearance, with a fat head, depressed snout, eyes protruding over the dorsal profle, and humped back. Besides, 
<taxonomicName id="4C264D6EFFFCFF8C77F6F99EFA81F9E2" authorityName="Maurice Kottelat" authorityYear="2017" box="[1200,1333,1614,1638]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="354" phylum="Chordata" rank="species" species="klydonion">
<emphasis id="B952EAFFFFFCFF8C77F6F99EFA81F9E2" box="[1200,1333,1614,1638]" italics="true" pageId="13" pageNumber="354">S. klydonion</emphasis>
</taxonomicName>
has 12–21 very contrasted bars restricted to the fank (vs. 7–10, not very contrasted and meeting their contralaterals on the back).
</paragraph>
<paragraph id="8B9936EDFFFCFF8C706BF91EFB46F7C2" blockId="13.[813,1452,1742,2118]" pageId="13" pageNumber="354">
<taxonomicName id="4C264D6EFFFCFF8C706BF91EFBB2F962" authorityName="Maurice Kottelat" authorityYear="2017" box="[813,1030,1742,1766]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="354" phylum="Chordata" rank="species" species="klydonion">
<emphasis id="B952EAFFFFFCFF8C706BF91EFBB2F962" box="[813,1030,1742,1766]" italics="true" pageId="13" pageNumber="354">Schistura klydonion</emphasis>
</taxonomicName>
is distinguished from 
<taxonomicName id="4C264D6EFFFCFF8C7645F91EFA1FF962" authorityName="Kottelat" authorityYear="2000" box="[1283,1451,1742,1766]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="354" phylum="Chordata" rank="species" species="bolavenensis">
<emphasis id="B952EAFFFFFCFF8C7645F91EFA1FF962" box="[1283,1451,1742,1766]" italics="true" pageId="13" pageNumber="354">S. bolavenensis</emphasis>
</taxonomicName>
(
<figureCitation id="131D2A68FFFCFF8C7073F93EFC3CF882" box="[821,904,1774,1798]" captionStart="Fig. 17" captionStartId="13.[813,848,962,984]" captionTargetBox="[1041,1301,189,927]" captionTargetId="figure@13.[917,1347,163,978]" captionTargetPageId="13" captionText="Fig. 17. Schistura klydonion, CMK 23344, paratype, 66.5 mm SL; digestive tract. Scale bar = 1 mm." httpUri="https://zenodo.org/record/886304/files/figure.png" pageId="13" pageNumber="354">Fig. 17</figureCitation>
) by its stouter body (depth 13.9–17.5% SL, vs. 12.8–14.6), a stouter caudal peduncle (depth 11.2–13.8% SL, vs. 9.8–12.0; 1.1–1.5 times in its length, vs. 1.5–1.7), a longer head (lateral head length 23.2–25.1% SL, vs. 19.7– 23.3), no median notch in upper lip (vs. presence), 12–21 bars restricted to the fank (vs. 15–24, meeting their contralateral on back), black pattern at caudal-fn base made of a vertically elongated black blotch at the middle, a small one at the base of the upper simple principal ray and sometimes one at the base of the lower one (vs. blotches fused to form a bar often reaching the dorsal and ventral midlines).
</paragraph>
<paragraph id="8B9936EDFFFFFF8F73CEFF2FFE8FFDB3" blockId="14.[136,775,255,567]" pageId="14" pageNumber="355">
Both 
<taxonomicName id="4C264D6EFFFFFF8F7384FF2FFEF1FE93" authorityName="Maurice Kottelat" authorityYear="2017" box="[194,325,255,279]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="355" phylum="Chordata" rank="species" species="klydonion">
<emphasis id="B952EAFFFFFFFF8F7384FF2FFEF1FE93" box="[194,325,255,279]" italics="true" pageId="14" pageNumber="355">S. klydonion</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C264D6EFFFFFF8F7231FF2FFE55FE93" authorityName="Maurice Kottelat" authorityYear="2017" box="[375,481,255,279]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="355" phylum="Chordata" rank="species" species="colossa">
<emphasis id="B952EAFFFFFFFF8F7231FF2FFE55FE93" box="[375,481,255,279]" italics="true" pageId="14" pageNumber="355">S. colossa</emphasis>
</taxonomicName>
are endemic to the Bolaven Plateau. 
<taxonomicName id="4C264D6EFFFFFF8F73AAFECFFE7AFEB3" authorityName="Maurice Kottelat" authorityYear="2017" box="[236,462,287,311]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="355" phylum="Chordata" rank="species" species="klydonion">
<emphasis id="B952EAFFFFFFFF8F73AAFECFFE7AFEB3" box="[236,462,287,311]" italics="true" pageId="14" pageNumber="355">Schistura klydonion</emphasis>
</taxonomicName>
has been observed only in the Xe Namnoy. Downstream of the waterfalls that border the plateau, the species has not been observed in the Xe Kong foodplain. It was not found in the Houay Makchang Gnai, a tributary of the Xe Namnoy that enters it after a high waterfall. It was also missing in the samples from the Xe Katam, another Xe Namnoy tributaries; the Xe Katam enters the Xe Namnoy on the Xe Kong foodplain, below a high waterfall.
</paragraph>
<paragraph id="8B9936EDFFFFFF8F73CEFD8FFDB5FB93" blockId="14.[136,775,607,1048]" pageId="14" pageNumber="355">
<taxonomicName id="4C264D6EFFFFFF8F73CEFD8FFEE7FDF3" authorityName="Maurice Kottelat" authorityYear="2017" box="[136,339,607,631]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="355" phylum="Chordata" rank="species" species="colossa">
<emphasis id="B952EAFFFFFFFF8F73CEFD8FFEE7FDF3" box="[136,339,607,631]" italics="true" pageId="14" pageNumber="355">Schistura colossa</emphasis>
</taxonomicName>
was collected in the Xe Pian on the Bolaven Plateau. The Xe Pian leaves the plateau on the south through a succession of waterfalls. 
<taxonomicName id="4C264D6EFFFFFF8F7155FD4FFD67FD33" authorityName="Maurice Kottelat" authorityYear="2017" box="[531,723,671,695]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="355" phylum="Chordata" rank="species" species="colossa">
<emphasis id="B952EAFFFFFFFF8F7155FD4FFD67FD33" box="[531,723,671,695]" italics="true" pageId="14" pageNumber="355">Schistura colossa</emphasis>
</taxonomicName>
was not observed downstream of the plateau. A single specimen caught in the Houay Champi and one caught in the Xe Set, both on Bolaven Plateau, cannot be distinguished from the samples from the Xe Pian. The Houay Champi and Xe Set are two tributaries of the Xe Don, which they join after leaving the Bolaven Plateau through waterfalls on the west and on the north, respectively. A single juvenile collected in the Houay Makchang Gnai is tentatively referred to 
<taxonomicName id="4C264D6EFFFFFF8F71B5FC4FFF6DFC53" authorityName="Maurice Kottelat" authorityYear="2017" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="355" phylum="Chordata" rank="species" species="colossa">
<emphasis id="B952EAFFFFFFFF8F71B5FC4FFF6DFC53" italics="true" pageId="14" pageNumber="355">S. colossa</emphasis>
</taxonomicName>
. The Houay Makchang Gnai is a tributary of the Xe Namnoy (see above). 
<taxonomicName id="4C264D6EFFFFFF8F72EDFC0FFDDAFC73" authorityName="Maurice Kottelat" authorityYear="2017" box="[427,622,991,1015]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="355" phylum="Chordata" rank="species" species="colossa">
<emphasis id="B952EAFFFFFFFF8F72EDFC0FFDDAFC73" box="[427,622,991,1015]" italics="true" pageId="14" pageNumber="355">Schistura colossa</emphasis>
</taxonomicName>
has not been observed in the Xe Namnoy itself.
</paragraph>
<paragraph id="8B9936EDFFFFFF8F73CEFBEFFD48F933" blockId="14.[136,775,1087,1719]" pageId="14" pageNumber="355">
Several species present in the Xe Pian are shared with the Houay Makchang Gnai but are missing in the Xe Namnoy (
<taxonomicName id="4C264D6EFFFFFF8F73D7FB50FED6FB13" authorityName="Kottelat" authorityYear="2000" box="[145,354,1151,1175]" class="Actinopterygii" family="Cyprinidae" genus="Poropuntius" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="355" phylum="Chordata" rank="species" species="solitus">
<emphasis id="B952EAFFFFFFFF8F73D7FB50FED6FB13" box="[145,354,1151,1175]" italics="true" pageId="14" pageNumber="355">Poropuntius solitus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C264D6EFFFFFF8F7229FBAFFDEAFB13" box="[367,606,1151,1175]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="355" phylum="Chordata" rank="species" species="forabilis">
<emphasis id="B952EAFFFFFFFF8F7229FBAFFDEAFB13" box="[367,606,1151,1175]" italics="true" pageId="14" pageNumber="355">Glyptothorax forabilis</emphasis>
</taxonomicName>
) and vice versa several species present in the Xe Namnoy have not been observed in the Houay Makchang Gnai (
<taxonomicName id="4C264D6EFFFFFF8F7115FB10FCB4FB53" authorityName="Roberts" authorityYear="1998" box="[595,768,1215,1239]" class="Actinopterygii" family="Cyprinidae" genus="Poropuntius" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="355" phylum="Chordata" rank="species" species="bolovenensis">
<emphasis id="B952EAFFFFFFFF8F7115FB10FCB4FB53" box="[595,768,1215,1239]" italics="true" pageId="14" pageNumber="355">P. bolovenensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C264D6EFFFFFF8F73CEFB30FEF1FB73" authorityName="Kottelat" authorityYear="2000" box="[136,325,1247,1271]" class="Actinopterygii" family="Cyprinidae" genus="Poropuntius" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="355" phylum="Chordata" rank="species" species="lobocheiloides">
<emphasis id="B952EAFFFFFFFF8F73CEFB30FEF1FB73" box="[136,325,1247,1271]" italics="true" pageId="14" pageNumber="355">P. lobocheiloides</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C264D6EFFFFFF8F7213FB0FFE68FB73" authorityName="Maurice Kottelat" authorityYear="2017" box="[341,476,1247,1271]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="355" phylum="Chordata" rank="species" species="klydonion">
<emphasis id="B952EAFFFFFFFF8F7213FB0FFE68FB73" box="[341,476,1247,1271]" italics="true" pageId="14" pageNumber="355">S. klydonion</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C264D6EFFFFFF8F72AAFB30FDC2FB73" box="[492,630,1248,1271]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="355" phylum="Chordata" rank="species" species="porrectus">
<emphasis id="B952EAFFFFFFFF8F72AAFB30FDC2FB73" box="[492,630,1248,1271]" italics="true" pageId="14" pageNumber="355">G. porrectus</emphasis>
</taxonomicName>
). This might be an artefact resulting from an insufficient number of sampling sites in the Houay Makchang Gnai; however, it seems easily explained by topography. The Houay Makchang Gnai fows parallel to the Xe Pian, coming as close as 
<quantity id="4CDE9B08FFFFFF8F71A3FA8FFF1EFA13" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="14" pageNumber="355" unit="km" value="2.5">2.5 km</quantity>
apart (near the sampling site of CMK 23443 and near the type locality of 
<taxonomicName id="4C264D6EFFFFFF8F721FFA4FFE70FA33" authorityName="Maurice Kottelat" authorityYear="2017" box="[345,452,1439,1463]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="355" phylum="Chordata" rank="species" species="colossa">
<emphasis id="B952EAFFFFFFFF8F721FFA4FFE70FA33" box="[345,452,1439,1463]" italics="true" pageId="14" pageNumber="355">S. colossa</emphasis>
</taxonomicName>
, respectively). The altitude of the plateau above the respective shores is about 
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, and the maximum altitude between them about 
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. A third stream (Houay Liang) fows between them, tributary of the Xe Pian, and at one point less than 
<quantity id="4CDE9B08FFFFFF8F7165F9CFFDEDF9B3" box="[547,601,1567,1591]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="355" unit="km" value="1.0">1 km</quantity>
from the Houay Makchang Gnai; there, around 15°04′36″N 100°32′0 1″E, the altitude difference between the respective shores is about 
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. A former connection between the Houay Makchang Gnai and the Xe Pian seems therefore a reasonable hypothesis.
</paragraph>
<paragraph id="8B9936EDFFFFFF8F73CEF90FFD45F873" blockId="14.[136,775,1759,2040]" pageId="14" pageNumber="355">
Although the Houay Makchang Gnai is a tributary of the Xe Namnoy, it reaches it in the gorges descending the plateau and in its last km it descends from about 
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to about 
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, through rapids and waterfalls. It seems a reasonable hypothesis that the connection is relatively recent and that fshes endemic to the Xe Namnoy are not able to ascend the Houay Mackchang Gnai. An exception is possibly 
<taxonomicName id="4C264D6EFFFFFF8F73ACF86FFE25F853" authorityName="Kottelat" authorityYear="2000" box="[234,401,1983,2007]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="355" phylum="Chordata" rank="species" species="bolavenensis">
<emphasis id="B952EAFFFFFFFF8F73ACF86FFE25F853" box="[234,401,1983,2007]" italics="true" pageId="14" pageNumber="355">S. bolavenensis</emphasis>
</taxonomicName>
known from the Xe Namnoy and Houay Makchang Gnai but not observed in the Xe Pian.
</paragraph>
<paragraph id="8B9936EDFFFFFF8F7005FF6FFC57FE53" blockId="14.[835,1475,191,471]" pageId="14" pageNumber="355">
The Xe Katam descends from the plateau through a very high waterfall and enters the Xe Namnoy at the foot of the plateau. Similarly, the topography of the plateau shows that there is no geomorphological obstacle to an earlier connection between the Xe Katam and the Houay Makchang Gnai, and the Xe Pian, which may explain the presence of 
<taxonomicName id="4C264D6EFFFFFF8F7601FEB0FA76FEF3" box="[1351,1474,351,375]" class="Actinopterygii" family="Sisoridae" genus="Glyptothorax" kingdom="Animalia" order="Siluriformes" pageId="14" pageNumber="355" phylum="Chordata" rank="species" species="forabilis">
<emphasis id="B952EAFFFFFFFF8F7601FEB0FA76FEF3" box="[1351,1474,351,375]" italics="true" pageId="14" pageNumber="355">G. forabilis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C264D6EFFFFFF8F7033FE50FC69FE13" authorityName="Kottelat" authorityYear="2000" box="[885,989,383,407]" class="Actinopterygii" family="Cyprinidae" genus="Poropuntius" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="355" phylum="Chordata" rank="species" species="solitus">
<emphasis id="B952EAFFFFFFFF8F7033FE50FC69FE13" box="[885,989,383,407]" italics="true" pageId="14" pageNumber="355">P. solitus</emphasis>
</taxonomicName>
in both rivers and their absence in the Xe Namnoy, and the absence of the Xe Namnoy endemics in the Xe Katam.
</paragraph>
</subSubSection>
<subSubSection id="C33C6566FFFFFF8F7005FE2FFA8EF9FC" pageId="14" pageNumber="355" type="conservation">
<paragraph id="8B9936EDFFFFFF8F7005FE2FFB99FC1C" blockId="14.[835,1475,511,920]" pageId="14" pageNumber="355">
Rivers on the plateau have been impacted by agriculture (especially coffee plantations), deforestation and smallscale mining. With the abrupt drop of about 700 meters, the plateau is of great interest for hydropower development. A complex scheme is under construction that includes two dams diverting waters from the Houay Makchang-Gnai and Xe Pian to a reservoir created on the Xe Namnoy by a third dam, and from there by a penstock to the Xe Kong. These dams result/will result in severe impacts on the aquatic habitats (inundation of rapids, reduction of volume and velocity of discharge below dams, disturbance of annual cycles, turbidity, siltation, genetic exchanges between watersheds, etc.) (see 
<bibRefCitation id="EFB74B1CFFFFFF8F7005FCAFFB95FC1C" author="Kottelat M" box="[835,1057,895,920]" journalOrPublisher="Raffes Bulletin of Zoology, Supplement" pageId="14" pageNumber="355" pagination="1 - 199" part="26" refString="Kottelat M (2012) Conspectus cobitidum: an inventory of the loaches of the world (Teleostei: Cypriniformes: Cobitoidea). Raffes Bulletin of Zoology, Supplement 26: 1 - 199." title="Conspectus cobitidum: an inventory of the loaches of the world (Teleostei: Cypriniformes: Cobitoidea)" type="journal article" year="2012">Kottelat et al., 2012</bibRefCitation>
).
</paragraph>
<paragraph id="8B9936EDFFFFFF8F7005FC10FA75FABC" blockId="14.[835,1474,959,1336]" pageId="14" pageNumber="355">
Two of the known sites of 
<taxonomicName id="4C264D6EFFFFFF8F772CFC10FB62FC53" authorityName="Maurice Kottelat" authorityYear="2017" box="[1130,1238,959,983]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="355" phylum="Chordata" rank="species" species="colossa">
<emphasis id="B952EAFFFFFFFF8F772CFC10FB62FC53" box="[1130,1238,959,983]" italics="true" pageId="14" pageNumber="355">S. colossa</emphasis>
</taxonomicName>
are directly impacted (at Xe Pian dam site and near Houay Makchang Gnai dam site); the remaining three known sites are not impacted by this hydropower scheme, are quite distant and are in two other drainages (Xe Set and Houay Champi). Certainly 
<taxonomicName id="4C264D6EFFFFFF8F76E8FB90FC27FBF3" authorityName="Maurice Kottelat" authorityYear="2017" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="355" phylum="Chordata" rank="species" species="colossa">
<emphasis id="B952EAFFFFFFFF8F76E8FB90FC27FBF3" italics="true" pageId="14" pageNumber="355">S. colossa</emphasis>
</taxonomicName>
is not restricted to these four sites and has a wider distribution in the respective catchments. Still the whole known range is impacted by agriculture and the potential for more hydropower development exists. The decrease in range and habitat quality probably qualifes 
<taxonomicName id="4C264D6EFFFFFF8F7675FB30FA17FB73" authorityName="Maurice Kottelat" authorityYear="2017" box="[1331,1443,1247,1271]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="355" phylum="Chordata" rank="species" species="colossa">
<emphasis id="B952EAFFFFFFFF8F7675FB30FA17FB73" box="[1331,1443,1247,1271]" italics="true" pageId="14" pageNumber="355">S. colossa</emphasis>
</taxonomicName>
to the Near Threatened category under the International Union for Conservation of Nature (IUCN) criteria (
<bibRefCitation id="EFB74B1CFFFFFF8F766FFAF0FA01FABC" author="IUCN [International Union for Conservation of Nature]" bookContentInfo="31 pp." box="[1321,1461,1312,1336]" journalOrPublisher="IUCN, Gland &amp; Cambridge" pageId="14" pageNumber="355" refString="IUCN [International Union for Conservation of Nature] (2001). IUCN Red List Categories and Criteria: Version 3.1. IUCN, Gland &amp; Cambridge, 31 pp." title="IUCN Red List Categories and Criteria: Version 3.1" type="book" year="2001">IUCN, 2001</bibRefCitation>
).
</paragraph>
<paragraph id="8B9936EDFFFFFF8F7005FA8FFA8EF9FC" blockId="14.[835,1475,1375,1656]" pageId="14" pageNumber="355">
<taxonomicName id="4C264D6EFFFFFF8F7005FA8FFBAEFAF3" authorityName="Maurice Kottelat" authorityYear="2017" box="[835,1050,1375,1399]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="355" phylum="Chordata" rank="species" species="klydonion">
<emphasis id="B952EAFFFFFFFF8F7005FA8FFBAEFAF3" box="[835,1050,1375,1399]" italics="true" pageId="14" pageNumber="355">Schistura klydonion</emphasis>
</taxonomicName>
has been observed in the Xe Namnoy mainstream, which will be entirely impacted, and in some of the few headwaters, above the maximum level of the reservoir, presently not much impacted. However, these are small water bodies with limited fow and are not able to support large populations of large adults. The sharp decrease in range, habitat quality and population size probably qualifes 
<taxonomicName id="4C264D6EFFFFFF8F7005F990FC79F9D3" authorityName="Maurice Kottelat" authorityYear="2017" box="[835,973,1599,1623]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="355" phylum="Chordata" rank="species" species="klydonion">
<emphasis id="B952EAFFFFFFFF8F7005F990FC79F9D3" box="[835,973,1599,1623]" italics="true" pageId="14" pageNumber="355">S. klydonion</emphasis>
</taxonomicName>
to the Endangered or Critically Endangered categories under IUCN criteria (
<bibRefCitation id="EFB74B1CFFFFFF8F77E5F9B0FA9BF9FC" author="IUCN [International Union for Conservation of Nature]" bookContentInfo="31 pp." box="[1187,1327,1632,1656]" journalOrPublisher="IUCN, Gland &amp; Cambridge" pageId="14" pageNumber="355" refString="IUCN [International Union for Conservation of Nature] (2001). IUCN Red List Categories and Criteria: Version 3.1. IUCN, Gland &amp; Cambridge, 31 pp." title="IUCN Red List Categories and Criteria: Version 3.1" type="book" year="2001">IUCN, 2001</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>