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<document id="BADEB91704EA80CE18157349D5AA2F21" ID-CLB-Dataset="51205" ID-DOI="10.11646/zootaxa.4061.5.3" ID-GBIF-Dataset="8e4c370b-2a1d-4bc4-a714-16f9f3a1bd65" ID-ISSN="1175-5326" ID-Zenodo-Dep="264294" ID-ZooBank="6EEE4C42-D292-490A-9549-CB3428430920" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459680617424" checkinUser="plazi" docAuthor="Häckel, Martin" docDate="2016" docId="038F87EE01403014FF0BD7BDFA4BFBA8" docLanguage="en" docName="zootaxa.4061.5.3.pdf" docOrigin="Zootaxa 4061 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Craspophorus reflexus" docType="key" docVersion="10" lastPageNumber="522" masterDocId="FFB6FF9601513006FF9CD433FFDDFFEB" masterDocTitle="A contribution to the knowledge of the subfamily Panagaeinae Hope, 1838 from Africa. Part 1. Revision of the Craspedophorus reflexus group (Coleoptera: Carabidae)" masterLastPageNumber="526" masterPageNumber="504" pageNumber="521" updateTime="1698207076058" updateUser="ExternalLinkService">
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<mods:title id="7FC522D75A4A92215C0BD48DD736C4A5">A contribution to the knowledge of the subfamily Panagaeinae Hope, 1838 from Africa. Part 1. Revision of the Craspedophorus reflexus group (Coleoptera: Carabidae)</mods:title>
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<treatment id="038F87EE01403014FF0BD7BDFA4BFBA8" ID-DOI="http://doi.org/10.5281/zenodo.6088147" ID-GBIF-Taxon="119333008" ID-Zenodo-Dep="6088147" LSID="urn:lsid:plazi:treatment:038F87EE01403014FF0BD7BDFA4BFBA8" httpUri="http://treatment.plazi.org/id/038F87EE01403014FF0BD7BDFA4BFBA8" lastPageId="18" lastPageNumber="522" pageId="17" pageNumber="521">
<subSubSection id="C33C657301403017FF0BD7BDFD0EFC43" box="[151,723,910,936]" pageId="17" pageNumber="521" type="nomenclature">
<paragraph id="8B9936F801403017FF0BD7BDFD0EFC43" blockId="17.[151,723,910,936]" box="[151,723,910,936]" pageId="17" pageNumber="521">
<heading id="D0D1819401403017FF0BD7BDFD0EFC43" bold="true" box="[151,723,910,936]" fontSize="11" level="1" pageId="17" pageNumber="521" reason="1">
<emphasis id="B952EAEA01403017FF0BD7BDFD0EFC43" bold="true" box="[151,723,910,936]" pageId="17" pageNumber="521">
Key to species of
<taxonomicName id="4C264D7B01403017FEF1D7BDFD5CFC43" box="[365,641,910,936]" class="Insecta" family="Carabidae" genus="Craspophorus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="521" phylum="Arthropoda" rank="species" species="reflexus">
<emphasis id="B952EAEA01403017FEF1D7BDFD5CFC43" bold="true" box="[365,641,910,936]" italics="true" pageId="17" pageNumber="521">Craspophorus reflexus</emphasis>
</taxonomicName>
group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33C657301403014FF0BD7E6FA4BFBA8" lastPageId="18" lastPageNumber="522" pageId="17" pageNumber="521" type="key">
<keyStep id="30D22E5D01403017FF0BD7E6FA46FBEC" pageId="17" pageNumber="521">
<paragraph id="8B9936F801403017FF0BD7E6FA46FC01" blockId="17.[151,1436,981,2021]" box="[151,1435,981,1002]" pageId="17" pageNumber="521">
<keyLead id="30D795CD01403017FF0BD7E6FA46FC01" box="[151,1435,981,1002]" pageId="17" pageNumber="521">1 Elytra black, unicolor.................................................................................. 2</keyLead>
</paragraph>
<paragraph id="8B9936F801403017FF0BD7C1FA46FBEC" blockId="17.[151,1436,981,2021]" box="[151,1435,1010,1031]" pageId="17" pageNumber="521">
<keyLead id="30D795CD01403017FF0BD7C1FA46FBEC" box="[151,1435,1010,1031]" pageId="17" pageNumber="521">- Each elytron with at least one orange humeral macula........................................................ 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D22E5D01403017FF0BD03CFA41FB53" pageId="17" pageNumber="521">
<paragraph id="8B9936F801403017FF0BD03CFA41FBB4" blockId="17.[151,1436,981,2021]" pageId="17" pageNumber="521">
<keyLead id="30D795CD01403017FF0BD03CFA41FBB4" pageId="17" pageNumber="521">
2 Elytral striation barely discernible, striae interrupted by large and deep punctures, in apical half punctures nearly quadrate with interspaces reduced to ridges elevated as much as centers of intervals (
<figureCitation id="131D2A7D01403017FC15D01EFC0EFBA9" box="[905,979,1069,1090]" captionStart="FIGURE 27. a. C" captionStartId="14.[151,250,1855,1877]" captionTargetBox="[348,1246,1100,1788]" captionTargetId="figure@14.[340,1247,1095,1795]" captionTargetPageId="14" captionText="FIGURE 27. a. C. ethmoides Alluaud, 1930, HT (male), dorsal habitus; b. microsculpture of elytral humerus, dorsal view; c. left metepisternum outlined in yellow; d. MNHN labels; e. C. impictus (Boheman, 1848), female, Zanzibar, microsculpture of elytral humerus, dorsal view. f. C. ruvumanus sp. n., HT (male), microsculpture of elytral humerus, dorsal view." httpUri="https://zenodo.org/record/264297/files/figure.png" pageId="17" pageNumber="521">Figs 27</figureCitation>
b, c). Elytra punctate more randomly. Albertine Rift (western rim):
<collectingCountry id="F331766801403017FE06D079FE38FBB4" box="[410,485,1098,1119]" name="Uganda" pageId="17" pageNumber="521">Uganda</collectingCountry>
to
<collectingCountry id="F331766801403017FD98D079FD92FBB4" box="[516,591,1098,1119]" name="Malawi" pageId="17" pageNumber="521">Malawi</collectingCountry>
,
<collectingCountry id="F331766801403017FDC5D079FD78FBB4" box="[601,677,1098,1119]" name="Zambia" pageId="17" pageNumber="521">Zambia</collectingCountry>
,
<collectingCountry id="F331766801403017FD32D079FCCAFBB4" box="[686,791,1098,1119]" name="Zimbabwe" pageId="17" pageNumber="521">Zimbabwe</collectingCountry>
, northern and eastern South African Republic............... 3
</keyLead>
</paragraph>
<paragraph id="8B9936F801403017FF0BD054FA41FB53" blockId="17.[151,1436,981,2021]" pageId="17" pageNumber="521">
<keyLead id="30D795CD01403017FF0BD054FA41FB53" pageId="17" pageNumber="521">
- Elytral striae interrupted by rows of smaller and more densely spaced, mostly circular punctures; centers of intervals more elevated than spaces between punctures, which are lodges more deeply. Elytra punctate more evenly (
<figureCitation id="131D2A7D01403017FB39D0B6FB2CFB71" box="[1189,1265,1157,1178]" captionStart="FIGURE 26. C" captionStartId="14.[151,250,971,993]" captionTargetBox="[348,1246,223,909]" captionTargetId="figure@14.[340,1247,211,918]" captionTargetPageId="14" captionText="FIGURE 26. C. ruvumanus sp. n. a. HT (male), dorsal habitus; b. microsculpture of elytral humerus, dorsal view; c. ventrites, ventral view; d. NMWC labels; e. left metepisternum, oblique ventral view; f. left metepisternum outlined in yellow." httpUri="https://zenodo.org/record/264296/files/figure.png" pageId="17" pageNumber="521">Figs 26</figureCitation>
b, 27d). Albertine Rift (eastern rim): Iringa,
<collectingCountry id="F331766801403017FE45D091FDEEFB5C" box="[473,563,1186,1207]" name="Tanzania" pageId="17" pageNumber="521">Tanzania</collectingCountry>
.........................................................
<taxonomicName id="4C264D7B01403017FB44D091FA82FB5C" ID-CoL="Z5YY" box="[1240,1375,1186,1207]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="521" phylum="Arthropoda" rank="species" species="ruvumanus" status="sp. nov.">
<emphasis id="B952EAEA01403017FB44D091FA82FB5C" box="[1240,1375,1186,1207]" italics="true" pageId="17" pageNumber="521">C. ruvumanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A261579101403017FAFAD091FA41FB53" box="[1382,1436,1186,1208]" pageId="17" pageNumber="521" rank="species">
<emphasis id="B952EAEA01403017FAFAD091FA41FB53" bold="true" box="[1382,1436,1186,1208]" pageId="17" pageNumber="521">sp. n.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D22E5D01403017FF0BD0F1FA41FA4F" pageId="17" pageNumber="521">
<paragraph id="8B9936F801403017FF0BD0F1FA4BFAC4" blockId="17.[151,1436,981,2021]" pageId="17" pageNumber="521">
<keyLead id="30D795CD01403017FF0BD0F1FA4BFAC4" pageId="17" pageNumber="521">
3 Antennae with three basal antennomeres black; forelegs with 4th and 5th tarsores brownish. Pronotum with lateral margins moderately sinuate posteriad near the hind angles, this presenting very small but sharp dent; disc less coarsely punctured. Elytra with striae more deeply impressed, less coarsely punctured. Tarsi covered by ferruginous setae. Somewhat smaller species (
<quantity id="4CDE9B1D01403017FF7BD129FE88FAC4" box="[231,341,1306,1327]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.3" metricValueMin="2.2" pageId="17" pageNumber="521" unit="mm" value="22.5" valueMax="23.0" valueMin="22.0">2223 mm</quantity>
). Albertine Rift,
<collectingCountry id="F331766801403017FE6AD129FD9CFAC4" box="[502,577,1306,1327]" name="Uganda" pageId="17" pageNumber="521">Uganda</collectingCountry>
to
<collectingCountry id="F331766801403017FDC3D129FD4DFAC4" box="[607,656,1306,1327]" name="Democratic Republic of the Congo" pageId="17" pageNumber="521">DRC</collectingCountry>
(Kivu)....................................
<taxonomicName id="4C264D7B01403017FB15D129FA4BFAC4" ID-CoL="6BC4T" authority="Alluaud, 1930" authorityName="Alluaud" authorityYear="1930" box="[1161,1430,1306,1327]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="521" phylum="Arthropoda" rank="species" species="ethmoides">
<emphasis id="B952EAEA01403017FB15D129FADBFAC4" box="[1161,1286,1306,1327]" italics="true" pageId="17" pageNumber="521">C. ethmoides</emphasis>
<bibRefCitation id="EFB74B0901403017FA90D129FA4BFAC4" author="Alluaud" box="[1292,1430,1306,1327]" pageId="17" pageNumber="525" refString="Alluaud, C. (1930) Note sur les Panageides dAfrique tropicale. Afra, Cahiers d'Entomologie, 1, 1 - 9." type="journal article" year="1930">Alluaud, 1930</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9936F801403017FF0BD104FA41FA4F" blockId="17.[151,1436,981,2021]" pageId="17" pageNumber="521">
<keyLead id="30D795CD01403017FF0BD104FA41FA4F" pageId="17" pageNumber="521">
- Antennae black throughout. Pronotum with lateral margins very weakly sinuate posteriad near the hind angles, which are moderately indented and obtuse; disc more coarsely punctured. Elytra with striae less deeply impressed, coarsely punctured. Tarsi covered by black setae. Somewhat larger species (
<quantity id="4CDE9B1D01403017FD23D141FCF6FA6C" box="[703,811,1394,1415]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" metricValueMax="2.4" metricValueMin="2.3" pageId="17" pageNumber="521" unit="mm" value="23.5" valueMax="24.0" valueMin="23.0">2324 mm</quantity>
).
<collectingCountry id="F331766801403017FCA7D141FC78FA6C" box="[827,933,1394,1415]" name="Democratic Republic of the Congo" pageId="17" pageNumber="521">DR Congo</collectingCountry>
, eastern
<collectingCountry id="F331766801403017FC64D141FBE2FA6C" box="[1016,1087,1394,1415]" name="Angola" pageId="17" pageNumber="521">Angola</collectingCountry>
to
<collectingCountry id="F331766801403017FBC0D141FB75FA6C" box="[1116,1192,1394,1415]" name="Malawi" pageId="17" pageNumber="521">Malawi</collectingCountry>
,
<collectingCountry id="F331766801403017FB2DD141FB20FA6C" box="[1201,1277,1394,1415]" name="Zambia" pageId="17" pageNumber="521">Zambia</collectingCountry>
,
<collectingCountry id="F331766801403017FA9BD141FAADFA6C" box="[1287,1392,1394,1415]" name="Zimbabwe" pageId="17" pageNumber="521">Zimbabwe</collectingCountry>
, and
<collectingCountry id="F331766801403017FF7DD1BCFE81FA4F" box="[225,348,1423,1444]" name="South Africa" pageId="17" pageNumber="521">South Africa</collectingCountry>
..................................................................
<taxonomicName id="4C264D7B01403017FB1DD1BCFA41FA4F" ID-CoL="Z5VY" authority="Boheman, 1838" authorityName="Boheman" authorityYear="1838" box="[1153,1436,1423,1444]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="521" phylum="Arthropoda" rank="species" species="impictus">
<emphasis id="B952EAEA01403017FB1DD1BCFB33FA4F" box="[1153,1262,1423,1444]" italics="true" pageId="17" pageNumber="521">C. impictus</emphasis>
(Boheman, 1838)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D22E5D01403017FF0BD19CFA46F9F5" pageId="17" pageNumber="521">
<paragraph id="8B9936F801403017FF0BD19CFA41FA09" blockId="17.[151,1436,981,2021]" pageId="17" pageNumber="521">
<keyLead id="30D795CD01403017FF0BD19CFA41FA09" pageId="17" pageNumber="521">
4 Each elytron ornamented by a single yellow macula in basal half, extending from 2nd to 8th interval; apical macula always absents. West of Central Africa (Nana-Mambéré)................................................
<taxonomicName id="4C264D7B01403017FB63D1FFFA82FA0A" ID-CoL="Z5SW" box="[1279,1375,1484,1505]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="521" phylum="Arthropoda" rank="species" species="arnosti" status="sp. nov.">
<emphasis id="B952EAEA01403017FB63D1FFFA82FA0A" box="[1279,1375,1484,1505]" italics="true" pageId="17" pageNumber="521">C. arnosti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A261579101403017FAFAD1FFFA41FA09" box="[1382,1436,1484,1506]" pageId="17" pageNumber="521" rank="species">
<emphasis id="B952EAEA01403017FAFAD1FFFA41FA09" bold="true" box="[1382,1436,1484,1506]" pageId="17" pageNumber="521">sp. n.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8B9936F801403017FF0BD1D9FA46F9F5" blockId="17.[151,1436,981,2021]" pageId="17" pageNumber="521">
<keyLead id="30D795CD01403017FF0BD1D9FA46F9F5" pageId="17" pageNumber="521">- Each elytron ornamented by two orange maculae, one in basal half and other in apical half, both encompassing spots on different intervals; if macula reduced, then spots always present at least on 6th and 4th intervals............................. 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D22E5D01403017FF0BD214FA41F8A2" pageId="17" pageNumber="521">
<paragraph id="8B9936F801403017FF0BD214FA4BF91A" blockId="17.[151,1436,981,2021]" pageId="17" pageNumber="521">
<keyLead id="30D795CD01403017FF0BD214FA4BF91A" pageId="17" pageNumber="521">
5 Terminal labial palpomere more dilated (securiform) than in other species of this group. Pronotal and elytral base parallels nearly throughout length (at least 9/10 of length), curves slightly forward only near hind angles. Pronotum punctate more sparsely and finely, disc strongly convex. Elytra with four orange maculae, each markedly larger than in other species because of its great width as well as length; humeral macula extending from 2nd interval to margin, preapical macula extending from 3rd to 8th interval. All macular spots very long, enlarging maculae, coloration resembling that in
<taxonomicName id="4C264D7B01403017FBC5D292FB00F95D" box="[1113,1245,1697,1718]" class="Insecta" family="Carabidae" genus="Craspophorus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="521" phylum="Arthropoda" rank="species" species="bifasciatus">
<emphasis id="B952EAEA01403017FBC5D292FB00F95D" box="[1113,1245,1697,1718]" italics="true" pageId="17" pageNumber="521">C. bifasciatus</emphasis>
</taxonomicName>
(Laporte de Castelnau, 1835) from Oriental Region, or
<taxonomicName id="4C264D7B01403017FDDED28DFCCBF938" ID-CoL="4C97P" box="[578,790,1726,1747]" class="Insecta" family="Carabidae" genus="Panagaeus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="521" phylum="Arthropoda" rank="species" species="cruxmajor">
<emphasis id="B952EAEA01403017FDDED28DFCCBF938" box="[578,790,1726,1747]" italics="true" pageId="17" pageNumber="521">Panagaeus cruxmajor</emphasis>
</taxonomicName>
(L., 1758) from Palearctic Region....................................................................................................
<taxonomicName id="4C264D7B01403017FBC3D2EFFA4BF91A" ID-CoL="Z5WM" authority="Burgeon, 1930" authorityName="Burgeon" authorityYear="1930" box="[1119,1430,1756,1777]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="521" phylum="Arthropoda" rank="species" species="latemaculatus">
<emphasis id="B952EAEA01403017FBC3D2EFFADCF91A" box="[1119,1281,1756,1777]" italics="true" pageId="17" pageNumber="521">C. latemaculatus</emphasis>
<bibRefCitation id="EFB74B0901403017FA94D2EFFA4BF91A" author="Burgeon" box="[1288,1430,1756,1777]" pageId="17" pageNumber="526" refString="Burgeon, L. (1930) Les Panagaeini du Musee du Congo. Revue de Zoologie et de Botanique Africaines, 19, 151 - 166." type="journal article" year="1930">Burgeon, 1930</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9936F801403017FF0BD2CAFA41F8A2" blockId="17.[151,1436,981,2021]" pageId="17" pageNumber="521">
<keyLead id="30D795CD01403017FF0BD2CAFA41F8A2" pageId="17" pageNumber="521">- Terminal labial palpomere more-or-less elongated, kidney-shaped. Pronotal and elytral base parallels only medially (ca. 3/5 of length), near hind angles curves forward in obligue fashion. Pronotum usually unevenly densely and coarsely punctate, disc only slightly convex. Elytra with maculae markedly smaller, because macular spots are much shorter................... 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D22E5D01403017FF0BD362FA46F80E" pageId="17" pageNumber="521">
<paragraph id="8B9936F801403017FF0BD362FA46F86D" blockId="17.[151,1436,981,2021]" pageId="17" pageNumber="521">
<keyLead id="30D795CD01403017FF0BD362FA46F86D" pageId="17" pageNumber="521">6 Elytral orange maculae strongly reduced, both humeral and preapical, humeral reduced to 25 macular spots, two elongated, located on 6th and 8th interval, one shorter on 4th interval; spots on 5th and 7th intervals markedly shorter or absent.......... 7</keyLead>
</paragraph>
<paragraph id="8B9936F801403017FF0BD3A3FA46F80E" blockId="17.[151,1436,981,2021]" pageId="17" pageNumber="521">
<keyLead id="30D795CD01403017FF0BD3A3FA46F80E" pageId="17" pageNumber="521">- Elytral orange maculae less reduced, humeral macula expanded, forming fascia, extending from 4th to 8th interval, with moreor-less serrate margins, often with shorter spot on 3rd interval and rarely also on 2nd interval, spot on 4th interval longer, almost as long as spot on 6th interval............................................................................ 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D22E5D01433014FF0BD4A9FA46FEA0" pageId="18" pageNumber="522">
<paragraph id="8B9936F801433014FF0BD4A9FA4BFEE0" blockId="18.[151,1437,151,1091]" pageId="18" pageNumber="522">
<keyLead id="30D795CD01433014FF0BD4A9FA4BFEE0" pageId="18" pageNumber="522">
7 Pronotum narrow, pronotal to elytral width ratio ≥ 1.5. Humeral macula comprised of three elongated spots on intervals 4th, 6th and 8th. Preapical macula also comprised of three spots, each with different length, located on same intervals as in humeral macula, in addition with a small yellow spot on 7th interval. Spot on 6th interval twice as long as spots on intervals 4th and 8th.
<collectingCountry id="F331766801433014FF7DD4C5FE91FEE0" box="[225,332,246,267]" name="Democratic Republic of the Congo" pageId="18" pageNumber="522">DR Congo</collectingCountry>
: Orientale Province.......................................................
<taxonomicName id="4C264D7B01433014FB03D4C5FA4BFEE0" ID-CoL="Z5ZF" authority="Alluaud, 1930" authorityName="Alluaud" authorityYear="1930" box="[1183,1430,246,267]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="522" phylum="Arthropoda" rank="species" species="stanleyi">
<emphasis id="B952EAEA01433014FB03D4C5FADBFEE0" box="[1183,1286,246,267]" italics="true" pageId="18" pageNumber="522">C. stanleyi</emphasis>
<bibRefCitation id="EFB74B0901433014FA90D4C5FA4BFEE0" author="Alluaud" box="[1292,1430,246,267]" pageId="18" pageNumber="525" refString="Alluaud, C. (1930) Note sur les Panageides dAfrique tropicale. Afra, Cahiers d'Entomologie, 1, 1 - 9." type="journal article" year="1930">Alluaud, 1930</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9936F801433014FF0BD525FA46FEA0" blockId="18.[151,1437,151,1091]" pageId="18" pageNumber="522">
<keyLead id="30D795CD01433014FF0BD525FA46FEA0" pageId="18" pageNumber="522">- Pronotum wider, pronotal to elytral width ratio &lt;1.5. Preapical macula with spot on 6th interval not more than one-half longer than spots on 4th and 8th intervals......................................................................... 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D22E5D01433014FF0BD560FA46FDE9" pageId="18" pageNumber="522">
<paragraph id="8B9936F801433014FF0BD560FA4BFE2E" blockId="18.[151,1437,151,1091]" pageId="18" pageNumber="522">
<keyLead id="30D795CD01433014FF0BD560FA4BFE2E" pageId="18" pageNumber="522">
8 Pronotum smaller, markedly narrower than elytra. Elytral striae densely and coarsely punctured. Humeral macula compact, extending from 4th to 8th interval, but macular spots short (at most twice longer than wide), present on each interval (4th, 5th, 6th, 7th, 8th), mostly without absence on odd-numbered intervals. Somewhat smaller species (
<quantity id="4CDE9B1D01433014FBB2D5A0FBABFE43" box="[1070,1142,403,424]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="18" pageNumber="522" unit="mm" value="21.0">21 mm</quantity>
).
<collectingCountry id="F331766801433014FB1AD5A0FB07FE43" box="[1158,1242,403,424]" name="Ethiopia" pageId="18" pageNumber="522">Ethiopia</collectingCountry>
,
<collectingCountry id="F331766801433014FB7FD5A0FA80FE43" box="[1251,1373,403,424]" name="South Sudan" pageId="18" pageNumber="522">South Sudan</collectingCountry>
.....................................................................................
<taxonomicName id="4C264D7B01433014FB31D583FA4BFE2E" ID-CoL="Z5TK" authority="Alluaud, 1930" authorityName="Alluaud" authorityYear="1930" box="[1197,1430,432,453]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="522" phylum="Arthropoda" rank="species" species="bozasi">
<emphasis id="B952EAEA01433014FB31D583FADBFE2E" box="[1197,1286,432,453]" italics="true" pageId="18" pageNumber="522">C. bozasi</emphasis>
<bibRefCitation id="EFB74B0901433014FA90D583FA4BFE2E" author="Alluaud" box="[1292,1430,432,453]" pageId="18" pageNumber="525" refString="Alluaud, C. (1930) Note sur les Panageides dAfrique tropicale. Afra, Cahiers d'Entomologie, 1, 1 - 9." type="journal article" year="1930">Alluaud, 1930</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9936F801433014FF0BD5FEFA46FDE9" blockId="18.[151,1437,151,1091]" pageId="18" pageNumber="522">
<keyLead id="30D795CD01433014FF0BD5FEFA46FDE9" pageId="18" pageNumber="522">
- Pronotum larger, almost as wide as elytra. Elytral striae punctured more finely or less densely. Humeral macula interrupted because spot on 5th interval is almost absent, spot on 7th interval is residual, rarely absent also. Larger species (
<quantity id="4CDE9B1D01433014FB63D5DEFAB4FDE9" box="[1279,1385,493,514]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" metricValueMax="2.7" metricValueMin="2.2" pageId="18" pageNumber="522" unit="mm" value="24.5" valueMax="27.0" valueMin="22.0">2227 mm</quantity>
).. 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D22E5D01433014FF0BD638FA46FD32" pageId="18" pageNumber="522">
<paragraph id="8B9936F801433014FF0BD638FA41FD97" blockId="18.[151,1437,151,1091]" pageId="18" pageNumber="522">
<keyLead id="30D795CD01433014FF0BD638FA41FD97" pageId="18" pageNumber="522">
9 Preapical macula markedly oblique posterolaterally, toward suture displaced anteriorly by about three-fifths; macular spots longer than those in humeral macula, located on same intervals (4th, 6th, 8th), spot on 7th interval narrower, shorter and placed more posteriorly than others, and one small spot on 5th interval at level of posterior margins of both contiguous spots. Large species (
<quantity id="4CDE9B1D01433014FEAFD654FEA6FD97" box="[307,379,615,636]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="18" pageNumber="522" unit="mm" value="22.0">22 mm</quantity>
), but somewhat smaller than following one.
<collectingCountry id="F331766801433014FCB8D654FC35FD97" box="[804,1000,615,636]" name="Republic of the Congo" pageId="18" pageNumber="522">Congo (Brazzaville)</collectingCountry>
...............
<taxonomicName id="4C264D7B01433014FB02D654FA41FD97" ID-CoL="Z5WW" authority="Alluaud, 1932" authorityName="Alluaud" authorityYear="1932" box="[1182,1436,615,636]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="522" phylum="Arthropoda" rank="species" species="lebaudyi">
<emphasis id="B952EAEA01433014FB02D654FAD0FD97" box="[1182,1293,615,636]" italics="true" pageId="18" pageNumber="522">C. lebaudyi</emphasis>
<bibRefCitation id="EFB74B0901433014FA8FD654FA41FD97" author="Alluaud" box="[1299,1436,615,636]" pageId="18" pageNumber="525" refString="Alluaud, C. (1932) III. Carabiques d Afrique tropicale. Afra, Cahiers d'Entomologie, 5, 9 - 17." type="journal article" year="1932">Alluaud, 1932</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9936F801433014FF0BD6B4FA46FD32" blockId="18.[151,1437,151,1091]" pageId="18" pageNumber="522">
<keyLead id="30D795CD01433014FF0BD6B4FA46FD32" pageId="18" pageNumber="522">
- Preapical macula transverse. Macular spots as long as those of humeral macula, spot on 4th interval approximately as far from base as that on 7th interval. Larger species (
<quantity id="4CDE9B1D01433014FDE0D694FD35FD57" box="[636,744,679,700]" metricMagnitude="-2" metricUnit="m" metricValue="2.55" metricValueMax="2.8" metricValueMin="2.3" pageId="18" pageNumber="522" unit="mm" value="25.5" valueMax="28.0" valueMin="23.0">2328 mm</quantity>
). Southern
<collectingCountry id="F331766801433014FCC9D694FC74FD57" box="[853,937,679,700]" name="Ethiopia" pageId="18" pageNumber="522">Ethiopia</collectingCountry>
,
<collectingCountry id="F331766801433014FC2ED694FC2EFD57" box="[946,1011,679,700]" name="Kenya" pageId="18" pageNumber="522">Kenya</collectingCountry>
,
<collectingCountry id="F331766801433014FC60D694FB88FD57" box="[1020,1109,679,700]" name="Tanzania" pageId="18" pageNumber="522">Tanzania</collectingCountry>
,
<collectingCountry id="F331766801433014FBC2D694FB6EFD57" box="[1118,1203,679,700]" name="Tanzania" pageId="18" pageNumber="522">Zanzibar</collectingCountry>
...............................................................................................
<taxonomicName id="4C264D7B01433014FBEAD6F7FA46FD32" ID-CoL="6BC46" authority="Harold, 1879" authorityName="Harold" authorityYear="1879" box="[1142,1435,708,729]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="522" phylum="Arthropoda" rank="species" species="carbonarius">
<emphasis id="B952EAEA01433014FBEAD6F7FADAFD32" box="[1142,1287,708,729]" italics="true" pageId="18" pageNumber="522">C. carbonarius</emphasis>
(
<bibRefCitation id="EFB74B0901433014FA8FD6F7FA49FD32" author="Harold" box="[1299,1428,708,729]" pageId="18" pageNumber="526" refString="Harold, E. von (1879) Bericht uber die von den Herren A. von Homeyer und P. Pogge in Angola und im Lunda-Reiche gesammelten Kafer. Coleopterologische Hefte, 16, 1 - 221." type="journal article" year="1879">Harold, 1879</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D22E5D01433014FF0BD6D1FA46FC65" pageId="18" pageNumber="522">
<paragraph id="8B9936F801433014FF0BD6D1FA4BFCBA" blockId="18.[151,1437,151,1091]" pageId="18" pageNumber="522">
<keyLead id="30D795CD01433014FF0BD6D1FA4BFCBA" pageId="18" pageNumber="522">
10 Pronotum more sinuate behind midlength, with distinct, almost rectangular hind angles. Elytra nearly glabrous, only lateral interstices with a few hairs; striae deep and punctured, intervals punctured densely and not too deeply. Both yellow elytral fasciae with spot on 5th interval much shorter than on others, but always present.
<collectingCountry id="F331766801433014FC57D72CFBEBFCDF" box="[971,1078,799,820]" name="Democratic Republic of the Congo" pageId="18" pageNumber="522">DR Congo</collectingCountry>
: Orientale Prov....................................................................................................
<taxonomicName id="4C264D7B01433014FB21D70FFA4BFCBA" ID-CoL="Z5TG" authority="Bates, 1886" authorityName="Bates" authorityYear="1886" box="[1213,1430,828,849]" class="Insecta" family="Carabidae" genus="Craspedophorus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="522" phylum="Arthropoda" rank="species" species="bonnyi">
<emphasis id="B952EAEA01433014FB21D70FFAC2FCBA" box="[1213,1311,828,849]" italics="true" pageId="18" pageNumber="522">C. bonnyi</emphasis>
Bates, 1886
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9936F801433014FF0BD76FFA46FC65" blockId="18.[151,1437,151,1091]" pageId="18" pageNumber="522">
<keyLead id="30D795CD01433014FF0BD76FFA46FC65" pageId="18" pageNumber="522">- Pronotum less sinuate behind midlength. Surface coarsely sculptured and more densely setose. Humeral fascia with spot on 5th interval only slightly shorter than others................................................................... 11</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D22E5D01433014FF0BD7A5FA4BFBA8" pageId="18" pageNumber="522">
<paragraph id="8B9936F801433014FF0BD7A5FA41FC02" blockId="18.[151,1437,151,1091]" pageId="18" pageNumber="522">
<keyLead id="30D795CD01433014FF0BD7A5FA41FC02" pageId="18" pageNumber="522">
11 Pronotum narrower, pronotal to elytral width ratio ≥ 1.5. Humeral macula with posterior margin strongly serrate; preapical macula comprised of three small triangular or ovoid spots covering intervals 4th, 6th and 8th. Somewhat smaller species,
<quantity id="4CDE9B1D01433014FACFD785FA45FC20" box="[1363,1432,950,971]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="18" pageNumber="522" unit="mm" value="22.0">22 mm</quantity>
. Democratic
<collectingCountry id="F331766801433014FECBD7E7FE47FC02" box="[343,410,980,1001]" name="Republic of the Congo" pageId="18" pageNumber="522">Congo</collectingCountry>
: Bandundu Province.................................................
<taxonomicName id="4C264D7B01433014FB30D7E7FA41FC02" authority="Kolbe, 1889" authorityName="Kolbe" authorityYear="1889" box="[1196,1436,980,1001]" class="Insecta" family="Carabidae" genus="Craspdophorus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="522" phylum="Arthropoda" rank="species" species="buettneri">
<emphasis id="B952EAEA01433014FB30D7E7FAC3FC02" box="[1196,1310,980,1001]" italics="true" pageId="18" pageNumber="522">C. buettneri</emphasis>
<bibRefCitation id="EFB74B0901433014FAB9D7E7FA41FC02" author="Kolbe" box="[1317,1436,980,1001]" pageId="18" pageNumber="526" refString="Kolbe, H. J. (1889) Bericht uber die von Herrn Dr. R. Buttner im Gebiete des unteren Quango und Kongo gesammelten Coleopteren. Entomologische Zeitung, 1889 (4 - 6), 113 - 133." type="journal article" year="1889">Kolbe, 1889</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B9936F801433014FF0BD7C2FA4BFBA8" blockId="18.[151,1437,151,1091]" pageId="18" pageNumber="522">
<keyLead id="30D795CD01433014FF0BD7C2FA4BFBA8" pageId="18" pageNumber="522">
- Pronotum wider, pronotal to elytral width ratio &lt;1.5. Humeral macula with posterior margin weakly serrate; preapical macula extending from 4th to 8th interval, with spot on 6th interval prolonged more anteriorly than spot on 7th interval. Larger species,
<quantity id="4CDE9B1D01433014FF7DD01DFE91FBA8" box="[225,332,1070,1091]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" metricValueMax="2.9" metricValueMin="2.3" pageId="18" pageNumber="522" unit="mm" value="26.0" valueMax="29.0" valueMin="23.0">2329 mm</quantity>
. West Africa:
<collectingCountry id="F331766801433014FE4FD01DFDC2FBA8" box="[467,543,1070,1091]" name="Senegal" pageId="18" pageNumber="522">Senegal</collectingCountry>
to
<collectingCountry id="F331766801433014FDA2D01DFD6EFBA8" box="[574,691,1070,1091]" name="Ivory Coast" pageId="18" pageNumber="522">Ivory Coast</collectingCountry>
.......................................
<taxonomicName id="4C264D7B01433014FB18D01DFA4BFBA8" authority="Fabricius, 1781" authorityName="Fabricius" authorityYear="1781" box="[1156,1430,1070,1091]" class="Insecta" family="Carabidae" genus="Craspophorus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="522" phylum="Arthropoda" rank="species" species="reflexus">
<emphasis id="B952EAEA01433014FB18D01DFB36FBA8" box="[1156,1259,1070,1091]" italics="true" pageId="18" pageNumber="522">C. reflexus</emphasis>
(
<bibRefCitation id="EFB74B0901433014FB6BD01DFA52FBA8" author="Fabricius" box="[1271,1423,1070,1091]" pageId="18" pageNumber="526" refString="Fabricius, J. C. (1781) Species Insectorum inhibentes eorum differenties specificas, synonyma, auctorum, loca natalia, metamorphosim adiectis observationibus, descriptionibus. Tomus I. C. E. Bohn, Hamburg et Kilonii, vii + 522 pp." type="book" year="1781">Fabricius, 1781</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>