291 lines
42 KiB
XML
291 lines
42 KiB
XML
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<mods:title id="A6E569AD08690F672C03672177E88E20">Re-description of the type species of the genera Ganesella Blanford, 1863 and Globotrochus Haas, 1935; with description of a new Ganesella species from Thailand (Eupulmonata, Camaenidae)</mods:title>
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<mods:namePart id="518BD9CA3777DB577E4A1D50FCE33369">Sutcharit, Chirasak</mods:namePart>
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<mods:namePart id="AA057F4755BA7409B1FC7B6823970342">Backeljau, Thierry</mods:namePart>
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<mods:namePart id="88BB121E03004E0627DB8B5CB123D235">Panha, Somsak</mods:namePart>
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<mods:date id="2C3D881610C536B5E0F269B888879C0F">2019</mods:date>
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<mods:number id="D343BF13C5AFE23E8403C4D942C330F9">870</mods:number>
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<mods:identifier id="22CC89A08A4B39136790C75E09CA8C3D" type="DOI">http://dx.doi.org/10.3897/zookeys.870.36970</mods:identifier>
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<treatment id="3DA698469F775B358F4445E647431D68" ID-GBIF-Taxon="159043270" LSID="urn:lsid:plazi:treatment:3DA698469F775B358F4445E647431D68" httpUri="http://treatment.plazi.org/id/3DA698469F775B358F4445E647431D68" lastPageNumber="51" pageId="0" pageNumber="51" scope_family="Camaenidae" scope_superOrder="Eupulmonata">
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<subSubSection id="58F87FC9CF08E0FF09684B62CD434D45" pageId="0" pageNumber="51" type="nomenclature">
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<paragraph id="96F776E67019CC7722B8B054227F24F0" pageId="0" pageNumber="51">
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<taxonomicName id="33826B4E727202F4E6ECF5910F452B32" ID-CoL="7RDCW" LSID="3da69846-9f77-5b35-8f44-45e647431d68" authority="Pfeiffer, 1861" authorityName="Pfeiffer" authorityYear="1861" class="Gastropoda" family="Camaenidae" genus="Ganesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ganesella rhombostoma" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="rhombostoma">Ganesella rhombostoma (Pfeiffer, 1861)</taxonomicName>
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<figureCitation id="4F56AF75156CF25426178C9047D0FA51" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. A, B Ganesella carinella from Keng-khoy, Saraburi (shell width about 15 mm) A live snail and B snails aestivated under loose tree bark, and with white epiphrams attached on substrate C Ganesella rhombostoma from Klong Had, Srakeo (shell height about 15 mm) D Globotrochus onestera from Cuc Phuong, Vietnam (shell width about 15 mm)." figureDoi="10.3897/zookeys.870.36970.figure1" httpUri="https://binary.pensoft.net/fig/323906" pageId="0" pageNumber="51">Figs 1C</figureCitation>
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,
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<figureCitation id="4609A980CE8953CFD7AE6BB46D9F5EC2" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A, B Ganesella carinella A shell from Lopburi CUMZ 4282 and B shell from Tam Brichinda, Chiangmai CUMZ 4165 C-I Ganesella rhombostoma C lectotype NHMUK 20130215 / 1 D paralectotype NHMUK 20130215 / 2 - 3 E shell of " harryleei Thach, 2017 " from Cambodia FMNH 381985 F shell from Klonghad, Srakeo CUMZ 5132 G shell from Chachoengsao CUMZ 5130 H shell from Sirisophon, Cambodia CUMZ 5131 and I shell from Srakaeo CUMZ 4286." figureDoi="10.3897/zookeys.870.36970.figure3" httpUri="https://binary.pensoft.net/fig/323908" pageId="0" pageNumber="51">
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3
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<normalizedToken id="C2F729BC63B9D5C00791D2006B53E502" originalValue="C–I">C-I</normalizedToken>
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</figureCitation>
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,
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<figureCitation id="541F4F97C92386A8C37532A559718355" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A-D Ganesella carinella from Saraburi A general view of genital system B internal structure of penis and vagina C pallial system and pulmonary cavity and D jaw E-G Ganesella rhombostoma from Srakaeo E general view of genital system F internal structure of penis and vagina and G jaw." figureDoi="10.3897/zookeys.870.36970.figure5" httpUri="https://binary.pensoft.net/fig/323910" pageId="0" pageNumber="51">
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5
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<normalizedToken id="5A6BE36791E515F8D75D8E3F6C865C73" originalValue="E–G">E-G</normalizedToken>
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</figureCitation>
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,
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<figureCitation id="D370072EBAC3E7D5FD958D2AB081A8E1" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Radula. A-C Ganesella carinella D-F Ganesella rhombostoma and G-I Globotrochus onestera A, D, G central tooth with the first to the second lateral teeth B, E, H lateral teeth with the tricuspid marginal teeth transition C, F, I outermost marginal teeth. Numbers indicate order of lateral and marginal teeth. Central tooth indicated by ' C'." figureDoi="10.3897/zookeys.870.36970.figure7" httpUri="https://binary.pensoft.net/fig/323912" pageId="0" pageNumber="51">
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7
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<normalizedToken id="76F982F39F1B79E5C0EA22FFC249C351" originalValue="D–F">D-F</normalizedToken>
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</figureCitation>
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,
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<figureCitation id="F196FCDF1B4EFD9CE54CD581254106A4" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Geographic distribution of Ganesella spp. and Globotrochus spp." figureDoi="10.3897/zookeys.870.36970.figure8" httpUri="https://binary.pensoft.net/fig/323913" pageId="0" pageNumber="51">8</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="4F0CFA9FC9EC089FD3EA166CEE540C6C" pageId="0" pageNumber="51" type="reference_group">
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<paragraph id="3976FF13AB1EAD2F09EB295DE3DE57B5" pageId="0" pageNumber="51">
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<taxonomicName id="90CA2979B587363A56ACFEE15964888C" class="Gastropoda" family="Camaenidae" genus="Bulimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bulimus rhombostomus" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="rhombostomus">Bulimus rhombostomus</taxonomicName>
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Pfeiffer, 1861: 194, 195.
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<bibRefCitation id="A4458C3171411C81132763836ECA3DFA" author="Pfeiffer, L" journalOrPublisher="Lipsiae, F. A. Brockhaus" pageId="0" pageNumber="51" refId="B44" refString="Pfeiffer, L, 1868. Monographiae Heliceorum Viventium Supplementum Tertium. . Lipsiae, F.A. Brockhaus" title="Monographiae Heliceorum Viventium Supplementum Tertium." year="1868">Pfeiffer 1868</bibRefCitation>
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: 33.
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</paragraph>
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<paragraph id="C039FB2C58FCEB4175202978B7C7C186" pageId="0" pageNumber="51">
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<taxonomicName id="8AC964A94695907D3B9B9772C397C481" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphidromus rhombostomus" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="rhombostomus">Amphidromus rhombostomus</taxonomicName>
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:
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<bibRefCitation id="1325BCFB29395EC3D0C2DE0D131032F0" author="Pfeiffer, L" journalOrPublisher="Cassellis, Sumptibus Theodori Fischeri" pageId="0" pageNumber="51" refId="B45" refString="Pfeiffer, L, Clessin, S, 1881. Nomenclator Heliceorum Viventium. . Cassellis, Sumptibus Theodori Fischeri" title="Nomenclator Heliceorum Viventium." year="1881">Pfeiffer and Clessin 1881</bibRefCitation>
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: 214.
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<bibRefCitation id="4696074FA0C4066E3273FA24A6E250AD" author="Morlet, L" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="51" pagination="121 - 199" refId="B36" refString="Morlet, L, 1889. Catalogue des coquilles recueillies, par M. Pavie dans le Cambodge et le Royaume de Siam, et description ďespeces nouvelles (1). . Journal de Conchyliologie 37: 121 - 199" title="Catalogue des coquilles recueillies, par M. Pavie dans le Cambodge et le Royaume de Siam, et description ďespeces nouvelles (1)." volume="37" year="1889">Morlet 1889</bibRefCitation>
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: 127.
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<bibRefCitation id="11C495D77D79306B77B8C32B5D277F7B" author="Morlet, L" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="51" pagination="119 - 122" refId="B37" refString="Morlet, L, 1890. Contribution a la faune malacologique du Cambodge et du Siam. . Journal de Conchyliologie 38: 119 - 122" title="Contribution a la faune malacologique du Cambodge et du Siam." volume="38" year="1890">Morlet 1890</bibRefCitation>
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: 121, 122, pl. 3, figs 6, 6a, b.
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<bibRefCitation id="A2167BFE7B0E2A882A55A134693CD713" pageId="0" pageNumber="51" refId="B19">Fischer and Dautzenberg 1904</bibRefCitation>
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||
: 407.
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</paragraph>
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<paragraph id="2D1540570B5F388E84CDC326DA4D559D" pageId="0" pageNumber="51">
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<taxonomicName id="4F039E394F4D1A584A830D26295181E8" class="Gastropoda" family="Camaenidae" genus="Buliminus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Buliminus rhombostomus" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="rhombostomus">Buliminus rhombostomus</taxonomicName>
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:
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<bibRefCitation id="9AA6A7ECF03425C6867E8D029E042477" author="Dautzenberg, P" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="51" pagination="343 - 471" refId="B17" refString="Dautzenberg, P, Fischer, H, 1906. Liste des mollusques recoltes par M.H. Mansuy en Indo-Chine et au Yunnan et description ďespeces nouvelles. . Journal de Conchyliologie 54: 343 - 471" title="Liste des mollusques recoltes par M. H. Mansuy en Indo-Chine et au Yunnan et description ďespeces nouvelles." volume="54" year="1906">Dautzenberg and Fischer 1906</bibRefCitation>
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: 366, 367.
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</paragraph>
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<paragraph id="81D39F4DD39749F6DC3B57154E3373A7" pageId="0" pageNumber="51">
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<taxonomicName id="EEE08F30BCA480E87921A6C6D8FF6CB2" class="Gastropoda" family="Camaenidae" genus="Buliminus" higherTaxonomySource="CoL" infraspecific-rank="var." kingdom="Animalia" lsidName="Buliminus rhombostomus var. pupoidea" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="variety" species="rhombostomus" variety="pupoidea">Buliminus rhombostomus var. pupoidea</taxonomicName>
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<bibRefCitation id="3935F49B6FF3EE0624D91BC6C527D265" author="Dautzenberg, P" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="51" pagination="343 - 471" refId="B17" refString="Dautzenberg, P, Fischer, H, 1906. Liste des mollusques recoltes par M.H. Mansuy en Indo-Chine et au Yunnan et description ďespeces nouvelles. . Journal de Conchyliologie 54: 343 - 471" title="Liste des mollusques recoltes par M. H. Mansuy en Indo-Chine et au Yunnan et description ďespeces nouvelles." volume="54" year="1906">Dautzenberg and Fischer 1906</bibRefCitation>
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: 367. Type locality: Hong-Chon, Cochinchine.
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</paragraph>
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<paragraph id="AF3A54F1C54AFE0ED6450FECEE06A9C6" pageId="0" pageNumber="51">
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<taxonomicName id="8A6028CB1D659B588F864222CD10F6BC" class="Gastropoda" family="Camaenidae" genus="Giardia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Giardia rhombostoma" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="rhombostoma">Giardia rhombostoma</taxonomicName>
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:
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<bibRefCitation id="245B79F21FDD640644FD5A6FBECC1384" author="Schileyko, AA" journalOrPublisher="Ruthenica" pageId="0" pageNumber="51" pagination="1 - 68" refId="B56" refString="Schileyko, AA, 2011. Check-list of land pulmonate molluscs of Vietnam (Gastropoda: Stylommatophora). . Ruthenica 20: 1 - 68" title="Check-list of land pulmonate molluscs of Vietnam (Gastropoda: Stylommatophora)." volume="20" year="2011">Schileyko 2011</bibRefCitation>
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: 46.
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</paragraph>
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<paragraph id="7CED82BC3A53C117D28C0E4FAE3FEFFE" pageId="0" pageNumber="51">
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<taxonomicName id="117824EF6B6D9F9D81AD29B6AB338AD9" class="Gastropoda" family="Camaenidae" genus="Pseudobuliminus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudobuliminus harryleei" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="harryleei">Pseudobuliminus harryleei</taxonomicName>
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Thach, 2017: 54, 55, figs 756-760. Type locality: Suburb of Battambang City, Battambang Province, Northwest Cambodia. New Synonym
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</paragraph>
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<paragraph id="B22E74863BAB4FC26815786749624F1E" pageId="0" pageNumber="51">
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<taxonomicName id="4EA35985FA940C1D18A18F757A366921" class="Gastropoda" family="Camaenidae" genus="Pseudobuliminus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudobuliminus tuongvyae" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="tuongvyae">Pseudobuliminus tuongvyae</taxonomicName>
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Thach, 2017: 56, figs 751-755. Type locality: Ha Tien, Kien Gaing Province, Southwest Vietnam. New Synonym
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</paragraph>
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<paragraph id="054EF9B716ED9D64DB00578A3D9AE11E" pageId="0" pageNumber="51">
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<taxonomicName id="1EB31913B39212B207A250F97A99B5F3" class="Gastropoda" family="Camaenidae" genus="Pseudobuliminus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudobuliminus huberi" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="huberi">Pseudobuliminus huberi</taxonomicName>
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Thach, 2017: 55, figs 759-760. Type locality: 20 km of Kampong Trach District, Kampot Province, Northwest Cambodia. New Synonym
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</paragraph>
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</subSubSection>
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<subSubSection id="D217540C5E12344905E50C928AB6D64B" pageId="0" pageNumber="51" type="type locality">
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<paragraph id="BFDCFA88FC618E43296656D1CF16A95D" pageId="0" pageNumber="51">Type locality.</paragraph>
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<paragraph id="32CF343B7CD46D1BD79AE90FCE05AF06" pageId="0" pageNumber="51">Camboja [Cambodia].</paragraph>
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</subSubSection>
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<subSubSection id="82D294FEB82D86D4FE28BF316FB2200C" pageId="0" pageNumber="51" type="materials_examined">
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<paragraph id="FDD3AC4664F2FD248F901E0D633B2C7A" pageId="0" pageNumber="51">Material examined.</paragraph>
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<paragraph id="ED4ECFB6CCC2BA9041BC1461A8AB31F0" pageId="0" pageNumber="51">
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<emphasis id="2F0D6CA94AF7FB1F9466DD6220CD9388" bold="true" pageId="0" pageNumber="51">Type specimens.</emphasis>
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Three syntypes of
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<taxonomicName id="751F83FA63AF222D2C74B527CB736250" class="Gastropoda" family="Camaenidae" genus="Bulimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bulimus rhombostomus" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="rhombostomus">
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<emphasis id="6D77FF06296DA45257229660DC622F2E" italics="true" pageId="0" pageNumber="51">Bulimus rhombostomus</emphasis>
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</taxonomicName>
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Pfeiffer, 1861 in H. Cuming collection, the shell that best matches with the original description is designated here as the lectotype NHMUK 20130215/1 (
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<figureCitation id="90BC80D0DCBEFA01847F148111B23CA0" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A, B Ganesella carinella A shell from Lopburi CUMZ 4282 and B shell from Tam Brichinda, Chiangmai CUMZ 4165 C-I Ganesella rhombostoma C lectotype NHMUK 20130215 / 1 D paralectotype NHMUK 20130215 / 2 - 3 E shell of " harryleei Thach, 2017 " from Cambodia FMNH 381985 F shell from Klonghad, Srakeo CUMZ 5132 G shell from Chachoengsao CUMZ 5130 H shell from Sirisophon, Cambodia CUMZ 5131 and I shell from Srakaeo CUMZ 4286." figureDoi="10.3897/zookeys.870.36970.figure3" httpUri="https://binary.pensoft.net/fig/323908" pageId="0" pageNumber="51">Fig. 3C</figureCitation>
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, height 16.7 mm, width 11.5 mm) to stabilize the name; the other two shells from the same lot become the paralectotypes NHMUK 20130215/2-3 (2 shells;
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<figureCitation id="82E22D298B8A97F08B5ECA27C86D4167" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A, B Ganesella carinella A shell from Lopburi CUMZ 4282 and B shell from Tam Brichinda, Chiangmai CUMZ 4165 C-I Ganesella rhombostoma C lectotype NHMUK 20130215 / 1 D paralectotype NHMUK 20130215 / 2 - 3 E shell of " harryleei Thach, 2017 " from Cambodia FMNH 381985 F shell from Klonghad, Srakeo CUMZ 5132 G shell from Chachoengsao CUMZ 5130 H shell from Sirisophon, Cambodia CUMZ 5131 and I shell from Srakaeo CUMZ 4286." figureDoi="10.3897/zookeys.870.36970.figure3" httpUri="https://binary.pensoft.net/fig/323908" pageId="0" pageNumber="51">Fig. 3D</figureCitation>
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, height 18.4 mm, width 13.2 mm).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9A4FD0467B00B1C9203632802910CF75" pageId="0" pageNumber="51" type="materials_examined">
|
||
<paragraph id="D0692F598BB5380274E70E65FC2FE76D" pageId="0" pageNumber="51">Additional material.</paragraph>
|
||
<paragraph id="B570B4171D3AB5ECF38439925F71744A" pageId="0" pageNumber="51">
|
||
<emphasis id="43B3D63D5842D319A1F95ABECA251E09" bold="true" pageId="0" pageNumber="51">VIETNAM</emphasis>
|
||
: Ha Tien, Kien Gaing Province, Southwest Vietnam: holotype of
|
||
<taxonomicName id="C472DA3C7299DDC2B331939E0E211467" class="Gastropoda" family="Camaenidae" genus="Pseudobuliminus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudobuliminus tuongvyae" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="tuongvyae">
|
||
<emphasis id="020D6547A526A5CFE0FBE7BC02C044AD" italics="true" pageId="0" pageNumber="51">Pseudobuliminus tuongvyae</emphasis>
|
||
</taxonomicName>
|
||
Thach, 2017 MNHN-IM-2000-33203.
|
||
<emphasis id="EDA92A650BC12484740C5569D4D9CFA0" bold="true" pageId="0" pageNumber="51">CAMBODIA</emphasis>
|
||
: Suburb of Battambang City, Battambang Province, Northwest Cambodia: holotype of
|
||
<taxonomicName id="426F8C25B2B87C37CCB37951062CB044" class="Gastropoda" family="Camaenidae" genus="Pseudobuliminus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudobuliminus harryleei" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="harryleei">
|
||
<emphasis id="DC6CB8D0B699597AAB998A6488B987BF" italics="true" pageId="0" pageNumber="51">Pseudobuliminus harryleei</emphasis>
|
||
</taxonomicName>
|
||
Thach, 2017 FMNH 381985 (
|
||
<figureCitation id="1DBFF42A034D9C122F6B2EB97634BE2B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A, B Ganesella carinella A shell from Lopburi CUMZ 4282 and B shell from Tam Brichinda, Chiangmai CUMZ 4165 C-I Ganesella rhombostoma C lectotype NHMUK 20130215 / 1 D paralectotype NHMUK 20130215 / 2 - 3 E shell of " harryleei Thach, 2017 " from Cambodia FMNH 381985 F shell from Klonghad, Srakeo CUMZ 5132 G shell from Chachoengsao CUMZ 5130 H shell from Sirisophon, Cambodia CUMZ 5131 and I shell from Srakaeo CUMZ 4286." figureDoi="10.3897/zookeys.870.36970.figure3" httpUri="https://binary.pensoft.net/fig/323908" pageId="0" pageNumber="51">Fig. 3E</figureCitation>
|
||
). Wat Thammaban Khiri, Sirisophon Town, Banteay Meanchey Province (
|
||
<geoCoordinate id="13D7DC103C4DF14F704BDAB41395BE32" degrees="13" direction="north" minutes="37" orientation="latitude" precision="1" seconds="58.1" value="13.632806">13°37'58.1"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="77E2522AD1B1E1BC205A34DD9B79713C" degrees="102" direction="east" minutes="56" orientation="longitude" precision="1" seconds="38.0" value="102.94389">102°56'38.0"E</geoCoordinate>
|
||
): CUMZ 5131 (
|
||
<figureCitation id="A7A93B19B06482A589663EB1008E24FE" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A, B Ganesella carinella A shell from Lopburi CUMZ 4282 and B shell from Tam Brichinda, Chiangmai CUMZ 4165 C-I Ganesella rhombostoma C lectotype NHMUK 20130215 / 1 D paralectotype NHMUK 20130215 / 2 - 3 E shell of " harryleei Thach, 2017 " from Cambodia FMNH 381985 F shell from Klonghad, Srakeo CUMZ 5132 G shell from Chachoengsao CUMZ 5130 H shell from Sirisophon, Cambodia CUMZ 5131 and I shell from Srakaeo CUMZ 4286." figureDoi="10.3897/zookeys.870.36970.figure3" httpUri="https://binary.pensoft.net/fig/323908" pageId="0" pageNumber="51">Fig. 3H</figureCitation>
|
||
).
|
||
<emphasis id="CD8408301B2DCCB9564188E843BAE442" bold="true" pageId="0" pageNumber="51">THAILAND</emphasis>
|
||
: Tam Leoum, Klonghad District, Srakaeo Province: CUMZ 4286 (
|
||
<figureCitation id="1F627D4F4685B3D685DEF0282C2C230F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A, B Ganesella carinella A shell from Lopburi CUMZ 4282 and B shell from Tam Brichinda, Chiangmai CUMZ 4165 C-I Ganesella rhombostoma C lectotype NHMUK 20130215 / 1 D paralectotype NHMUK 20130215 / 2 - 3 E shell of " harryleei Thach, 2017 " from Cambodia FMNH 381985 F shell from Klonghad, Srakeo CUMZ 5132 G shell from Chachoengsao CUMZ 5130 H shell from Sirisophon, Cambodia CUMZ 5131 and I shell from Srakaeo CUMZ 4286." figureDoi="10.3897/zookeys.870.36970.figure3" httpUri="https://binary.pensoft.net/fig/323908" pageId="0" pageNumber="51">Fig. 3I</figureCitation>
|
||
). Tam Pha Pheung, Klonghad District, Srakaeo Province: CUMZ 5124. Tam Srithong, Klonghad District, Srakaeo Province: CUMZ 4070, 5118, 5132 (
|
||
<figureCitation id="BD49B5BEBAAEE3ED14BD67F3BCFF7C9C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A, B Ganesella carinella A shell from Lopburi CUMZ 4282 and B shell from Tam Brichinda, Chiangmai CUMZ 4165 C-I Ganesella rhombostoma C lectotype NHMUK 20130215 / 1 D paralectotype NHMUK 20130215 / 2 - 3 E shell of " harryleei Thach, 2017 " from Cambodia FMNH 381985 F shell from Klonghad, Srakeo CUMZ 5132 G shell from Chachoengsao CUMZ 5130 H shell from Sirisophon, Cambodia CUMZ 5131 and I shell from Srakaeo CUMZ 4286." figureDoi="10.3897/zookeys.870.36970.figure3" httpUri="https://binary.pensoft.net/fig/323908" pageId="0" pageNumber="51">Fig. 3F</figureCitation>
|
||
). Tam Phet Phothong, Klonghad District, Srakaeo Province: CUMZ 4600, 5119. Khoa Phlapphueng Thong, Wang Somboon District, Srakaeo Province: CUMZ 4069. Wat Khao Maka, Muang District, Srakaeo Province: CUMZ 4071, 4598, 4599, 5130 (
|
||
<figureCitation id="66E26E33E99B18E39C70ECC0160D0082" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A, B Ganesella carinella A shell from Lopburi CUMZ 4282 and B shell from Tam Brichinda, Chiangmai CUMZ 4165 C-I Ganesella rhombostoma C lectotype NHMUK 20130215 / 1 D paralectotype NHMUK 20130215 / 2 - 3 E shell of " harryleei Thach, 2017 " from Cambodia FMNH 381985 F shell from Klonghad, Srakeo CUMZ 5132 G shell from Chachoengsao CUMZ 5130 H shell from Sirisophon, Cambodia CUMZ 5131 and I shell from Srakaeo CUMZ 4286." figureDoi="10.3897/zookeys.870.36970.figure3" httpUri="https://binary.pensoft.net/fig/323908" pageId="0" pageNumber="51">Fig. 3G</figureCitation>
|
||
). Tam Kaeo Sawanbandan, Pong Namron District, Chanthaburi Province: CUMZ 4285.Tam Rad, Tha Takiep District, Chachoengsao Province: CUMZ 5117.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7BDC6A4A3EF0E34963DBBFD39F05C4A4" pageId="0" pageNumber="51" type="description">
|
||
<paragraph id="4BFA674E4ACEE9F30482A835DED4F06A" pageId="0" pageNumber="51">Description.</paragraph>
|
||
<paragraph id="9C873C68512B9B86DE985D106CB1CD5E" pageId="0" pageNumber="51">
|
||
<emphasis id="109A841E6F3B78D334CDF7FCC589842C" bold="true" pageId="0" pageNumber="51">Shell.</emphasis>
|
||
Shell small, sinistral, thin and trochoid. Apex acute with blackish colour; embryonic shell smooth. Whorls 5-6, increasing regularly, convex; suture wide and shallow. Shell surface smooth or with fine growth lines. Last whorl large, well rounded, keeled near aperture; with or without brownish spiral band. Shell monochrome white, yellow to light brownish and translucent. Aperture semi-ovate, open subventrally; lip expanded and whitish; parietal callus transparent. Umbilicus rimate; columella wide and whitish.
|
||
</paragraph>
|
||
<paragraph id="F177807D9961AE70CF1EF0C6E5517027" pageId="0" pageNumber="51">
|
||
<emphasis id="0C996921692542BA111B82AB81D75FA6" bold="true" pageId="0" pageNumber="51">Genital organs.</emphasis>
|
||
Atrium (at) short (
|
||
<emphasis id="D093ACD41AA2AB5CE32356D7AA254E05" italics="true" pageId="0" pageNumber="51">N</emphasis>
|
||
= 10). Penis (p) cylindrical, long and may be as long as the vagina. Epiphallus (e) and flagellum (fl) each about half as long as the penis. Vas deferens (vd) long and narrow, extending from free oviduct and connected to distal part of epiphallus. Penial retractor muscle (pr) long and slightly thickened (
|
||
<figureCitation id="51AF6D61EF9B7650BD9D213FADF2C475" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A-D Ganesella carinella from Saraburi A general view of genital system B internal structure of penis and vagina C pallial system and pulmonary cavity and D jaw E-G Ganesella rhombostoma from Srakaeo E general view of genital system F internal structure of penis and vagina and G jaw." figureDoi="10.3897/zookeys.870.36970.figure5" httpUri="https://binary.pensoft.net/fig/323910" pageId="0" pageNumber="51">Fig. 5E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="1DEABB9E18FD5BA584686660EAF3F3D4" pageId="0" pageNumber="51">
|
||
Penial verge (pv) small, conical, and with smooth surface. Penial wall ribbed, forming a series of irregular, smooth longitudinal pilasters (pp) that encircle penial verge (
|
||
<figureCitation id="52623CB376EB30CCB46C4E44ED1E1AD7" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A-D Ganesella carinella from Saraburi A general view of genital system B internal structure of penis and vagina C pallial system and pulmonary cavity and D jaw E-G Ganesella rhombostoma from Srakaeo E general view of genital system F internal structure of penis and vagina and G jaw." figureDoi="10.3897/zookeys.870.36970.figure5" httpUri="https://binary.pensoft.net/fig/323910" pageId="0" pageNumber="51">Fig. 5F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="720E70358A7E9AFA54378252E1F11412" pageId="0" pageNumber="51">
|
||
Vagina (v) large, cylindrical about as long as penis. Gametolytic duct (gd) short, cylindrical, gradually tapering towards gametolytic sac (gs). Free oviduct (fo) short, about half as long as vagina; oviduct (ov) enlarged with curled lobules. Albumen gland (ag) curved ligulate. Hermaphroditic duct (hd) convoluted and connected to hermaphrodic gland (hg) (
|
||
<figureCitation id="955BFEDD817949A4E46FC5F64AB8F339" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A-D Ganesella carinella from Saraburi A general view of genital system B internal structure of penis and vagina C pallial system and pulmonary cavity and D jaw E-G Ganesella rhombostoma from Srakaeo E general view of genital system F internal structure of penis and vagina and G jaw." figureDoi="10.3897/zookeys.870.36970.figure5" httpUri="https://binary.pensoft.net/fig/323910" pageId="0" pageNumber="51">Fig. 5E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D5CAA9180FAA1407BD70BF095A64074C" pageId="0" pageNumber="51">
|
||
Vaginal wall with several smooth, longitudinal pilasters; vaginal wall itself with strong longitudinal ridges through the vaginal chamber (
|
||
<figureCitation id="BD8350327321F4035C0AA8FEBD47967C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A-D Ganesella carinella from Saraburi A general view of genital system B internal structure of penis and vagina C pallial system and pulmonary cavity and D jaw E-G Ganesella rhombostoma from Srakaeo E general view of genital system F internal structure of penis and vagina and G jaw." figureDoi="10.3897/zookeys.870.36970.figure5" httpUri="https://binary.pensoft.net/fig/323910" pageId="0" pageNumber="51">Fig. 5F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="102577B63867C2896A31E5EF56EE8433" pageId="0" pageNumber="51">
|
||
<emphasis id="D5540524EC86D2F44001F5C5E9672945" bold="true" pageId="0" pageNumber="51">Radula.</emphasis>
|
||
Teeth arranged in nearly straight rows, each row containing 58 (29-(6-9)-1-(7-9)-28) teeth. Central tooth triangular, symmetric monocuspid with dull cusp. Lateral teeth asymmetric, teeth no. 1-6 monocuspid and no. 7-10 bicuspid with very small ectocone (
|
||
<figureCitation id="A67729BD615CC46F5C8896B7D401EF96" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Radula. A-C Ganesella carinella D-F Ganesella rhombostoma and G-I Globotrochus onestera A, D, G central tooth with the first to the second lateral teeth B, E, H lateral teeth with the tricuspid marginal teeth transition C, F, I outermost marginal teeth. Numbers indicate order of lateral and marginal teeth. Central tooth indicated by ' C'." figureDoi="10.3897/zookeys.870.36970.figure7" httpUri="https://binary.pensoft.net/fig/323912" pageId="0" pageNumber="51">Fig. 7D, E</figureCitation>
|
||
). Marginal teeth start from teeth no. 9-10, tricuspid, endocone very small to absent, mesocone large triangular, and ectocone very small with pointed cusp. Outermost marginal teeth (near radula edge) tricuspid, endocone small, mesocone large with curved cusp, and ectocone with one, two or more pointed cusps (
|
||
<figureCitation id="47030022ED630BB833AA116C0DD8D1FF" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Radula. A-C Ganesella carinella D-F Ganesella rhombostoma and G-I Globotrochus onestera A, D, G central tooth with the first to the second lateral teeth B, E, H lateral teeth with the tricuspid marginal teeth transition C, F, I outermost marginal teeth. Numbers indicate order of lateral and marginal teeth. Central tooth indicated by ' C'." figureDoi="10.3897/zookeys.870.36970.figure7" httpUri="https://binary.pensoft.net/fig/323912" pageId="0" pageNumber="51">Fig. 7E, F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D88D198E7DCCFE4F15C0510FA76B6A50" pageId="0" pageNumber="51">
|
||
Jaw crescent, with anteriorly convex cutting margin. Vertical ribs thin, and variable in number and size (
|
||
<figureCitation id="70F897422DC6FE45F3E09F8F4331EEB9" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. A-D Ganesella carinella from Saraburi A general view of genital system B internal structure of penis and vagina C pallial system and pulmonary cavity and D jaw E-G Ganesella rhombostoma from Srakaeo E general view of genital system F internal structure of penis and vagina and G jaw." figureDoi="10.3897/zookeys.870.36970.figure5" httpUri="https://binary.pensoft.net/fig/323910" pageId="0" pageNumber="51">Fig. 5G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EEB7E90525CBCA6CFDCA6A330E1F2F28" pageId="0" pageNumber="51" type="distribution">
|
||
<paragraph id="32FCF04075CCD68D563A5B13846D7F98" pageId="0" pageNumber="51">Distribution.</paragraph>
|
||
<paragraph id="D818EE98F90D438E702340B618F7BA8F" pageId="0" pageNumber="51">
|
||
This species was formerly known from its type locality in Cambodia (
|
||
<bibRefCitation id="34E3477517096C0DC90D6B8B0FA7900C" author="Morlet, L" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="51" pagination="121 - 199" refId="B36" refString="Morlet, L, 1889. Catalogue des coquilles recueillies, par M. Pavie dans le Cambodge et le Royaume de Siam, et description ďespeces nouvelles (1). . Journal de Conchyliologie 37: 121 - 199" title="Catalogue des coquilles recueillies, par M. Pavie dans le Cambodge et le Royaume de Siam, et description ďespeces nouvelles (1)." volume="37" year="1889">Morlet 1889</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="54D3BED010089C6DE2E52E2B31B76456" author="Morlet, L" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="51" pagination="119 - 122" refId="B37" refString="Morlet, L, 1890. Contribution a la faune malacologique du Cambodge et du Siam. . Journal de Conchyliologie 38: 119 - 122" title="Contribution a la faune malacologique du Cambodge et du Siam." volume="38" year="1890">1890</bibRefCitation>
|
||
) and some inaccurate localities recorded from Vietnam (
|
||
<bibRefCitation id="E02243A81FFE1145400C9BC5DBAA3BAD" author="Schileyko, AA" journalOrPublisher="Ruthenica" pageId="0" pageNumber="51" pagination="1 - 68" refId="B56" refString="Schileyko, AA, 2011. Check-list of land pulmonate molluscs of Vietnam (Gastropoda: Stylommatophora). . Ruthenica 20: 1 - 68" title="Check-list of land pulmonate molluscs of Vietnam (Gastropoda: Stylommatophora)." volume="20" year="2011">Schileyko 2011</bibRefCitation>
|
||
). In Thailand, it is known from several localities in Chachoengsao, Srakeo and Chanthaburi Provinces.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9055E0DEEDA14E6BD05E05409129FB52" pageId="0" pageNumber="51" type="remarks">
|
||
<paragraph id="7FB6A82786E349892D7A6FBE009706D1" pageId="0" pageNumber="51">Remarks.</paragraph>
|
||
<paragraph id="1BAA8AC3673A9D5FD1C423820F535EE7" pageId="0" pageNumber="51">
|
||
<taxonomicName id="6224EB310146E8CA0576A912BB6ED83D" class="Gastropoda" family="Camaenidae" genus="Ganesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ganesella rhombostoma" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="rhombostoma">
|
||
<emphasis id="E2F0FC386758BD00AADB9E5ADF040D73" italics="true" pageId="0" pageNumber="51">Ganesella rhombostoma</emphasis>
|
||
</taxonomicName>
|
||
has long been overlooked and its taxonomic status has been unclear. The species has been erroneously assigned to
|
||
<taxonomicName id="4D5709B7D6294A3635AD184BDBCE5E35" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphidromus" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="genus">
|
||
<emphasis id="D14EBC358E5106B28496CDEBA21580D0" italics="true" pageId="0" pageNumber="51">Amphidromus</emphasis>
|
||
</taxonomicName>
|
||
Albers, 1850 and
|
||
<taxonomicName id="908E40D20973DE479F1AA4BB89B26BD7" class="Gastropoda" family="Camaenidae" genus="Giardia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Giardia" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="genus">
|
||
<emphasis id="5A7F8E88486013B3873B8D2F29BD3FAB" italics="true" pageId="0" pageNumber="51">Giardia</emphasis>
|
||
</taxonomicName>
|
||
Ancey, 1907 (see
|
||
<bibRefCitation id="9E7B74A8B3FB0D0DAB01ABE68FB75C62" author="Dautzenberg, P" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="51" pagination="343 - 471" refId="B17" refString="Dautzenberg, P, Fischer, H, 1906. Liste des mollusques recoltes par M.H. Mansuy en Indo-Chine et au Yunnan et description ďespeces nouvelles. . Journal de Conchyliologie 54: 343 - 471" title="Liste des mollusques recoltes par M. H. Mansuy en Indo-Chine et au Yunnan et description ďespeces nouvelles." volume="54" year="1906">Dautzenberg and Fischer 1906</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="B994C71BA8E507AFE18E670B609EEDB3" author="Schileyko, AA" journalOrPublisher="Ruthenica" pageId="0" pageNumber="51" pagination="1 - 68" refId="B56" refString="Schileyko, AA, 2011. Check-list of land pulmonate molluscs of Vietnam (Gastropoda: Stylommatophora). . Ruthenica 20: 1 - 68" title="Check-list of land pulmonate molluscs of Vietnam (Gastropoda: Stylommatophora)." volume="20" year="2011">Schileyko 2011</bibRefCitation>
|
||
). Yet, its trochoid shell and sub-ventrally opening aperture show that it belongs to neither of these genera, since
|
||
<taxonomicName id="5A687511E27652DC93012741B5D58640" class="Gastropoda" family="Camaenidae" genus="Giardia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Giardia" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="genus">
|
||
<emphasis id="4DC5FC7CE04B7642942E59A69178868A" italics="true" pageId="0" pageNumber="51">Giardia</emphasis>
|
||
</taxonomicName>
|
||
(type species
|
||
<taxonomicName id="AB264233CB4E3C5F2E7C2E941E530B99" class="Gastropoda" family="Camaenidae" genus="Bulimus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bulimus siamensis" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="siamensis">
|
||
<emphasis id="BAC5BC806BD76B8F5F91B0B33F3E2CCC" italics="true" pageId="0" pageNumber="51">Bulimus siamensis</emphasis>
|
||
</taxonomicName>
|
||
Redfield, 1853) and
|
||
<taxonomicName id="A25A81D98EC4CB37D78ADFEDDE0DE226" class="Gastropoda" family="Camaenidae" genus="Amphidromus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphidromus" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="genus">
|
||
<emphasis id="7931AC7AB29FBD9FF3ED325E347379C6" italics="true" pageId="0" pageNumber="51">Amphidromus</emphasis>
|
||
</taxonomicName>
|
||
(type species
|
||
<taxonomicName id="90F9F71327B0D6A8B6EE34FFE11FF41A" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helix perversus" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="species" species="perversus">
|
||
<emphasis id="146D653F4E304BEF6FC449CF5E3A749D" italics="true" pageId="0" pageNumber="51">Helix perversus</emphasis>
|
||
</taxonomicName>
|
||
Linnaeus, 1758) have an elongate, ovate shell, a non-deflected last whorl, an ovate and laterally opening aperture, and a narrowly opened umbilicus (see also
|
||
<bibRefCitation id="89B66F9650F8B9EDCE5E48F4E1A89E03" pageId="0" pageNumber="51" refId="B54">Schileyko (2003)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="625B0CB0883243530136FF765277AB6E" DOI="https://doi.org/10.1093/mollus/eyi044" author="Sutcharit, C" journalOrPublisher="Journal of Molluscan Studies" pageId="0" pageNumber="51" pagination="1 - 30" refId="B60" refString="Sutcharit, C, Panha, S, 2006. Taxonomic review of the tree snail Amphidromus Albers, 1850 (Pulmonata: Camaenidae) in Thailand and adjacent areas: subgenusAmphidromus. . Journal of Molluscan Studies 72: 1 - 30" title="Taxonomic review of the tree snail Amphidromus Albers, 1850 (Pulmonata: Camaenidae) in Thailand and adjacent areas: subgenusAmphidromus." url="https://doi.org/10.1093/mollus/eyi044" volume="72" year="2006">Sutcharit and Panha (2006)</bibRefCitation>
|
||
for further comparisons). Moreover, the anatomical evidence presented here suggests that this species rather belongs to
|
||
<taxonomicName id="15E977BBFC94842CC10E851082A2211B" class="Gastropoda" family="Camaenidae" genus="Ganesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ganesella" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="genus">
|
||
<emphasis id="3B591FCAB7CE068CCD29B19EB1C3380E" italics="true" pageId="0" pageNumber="51">Ganesella</emphasis>
|
||
</taxonomicName>
|
||
. However, the relationships between
|
||
<taxonomicName id="60F8FB24694FE5352AE2683D1F70D9B9" class="Gastropoda" family="Camaenidae" genus="Ganesella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ganesella" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="genus">
|
||
<emphasis id="E9CCCD1F4AADF68A6B7FD4C3294FD619" italics="true" pageId="0" pageNumber="51">Ganesella</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="AD13D7AEE2B2D4D8334FF3382E6570A2" class="Gastropoda" family="Camaenidae" genus="Giardia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Giardia" order="Stylommatophora" pageId="0" pageNumber="51" phylum="Mollusca" rank="genus">
|
||
<emphasis id="8D4904ECD63A16DD52239F5773E6C7DE" italics="true" pageId="0" pageNumber="51">Giardia</emphasis>
|
||
</taxonomicName>
|
||
needs further investigation by molecular analysis.
|
||
</paragraph>
|
||
<paragraph id="89A778486A37E8B7083758AADED13976" pageId="0" pageNumber="51">
|
||
Specimens from isolated limestone outcrops at Khao Maka, Chacheongsao are smaller and have a more ovate shell without a brownish spiral band on of the last whorl (
|
||
<figureCitation id="6B52439C6C7A88C0D80BF2F83757690C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A, B Ganesella carinella A shell from Lopburi CUMZ 4282 and B shell from Tam Brichinda, Chiangmai CUMZ 4165 C-I Ganesella rhombostoma C lectotype NHMUK 20130215 / 1 D paralectotype NHMUK 20130215 / 2 - 3 E shell of " harryleei Thach, 2017 " from Cambodia FMNH 381985 F shell from Klonghad, Srakeo CUMZ 5132 G shell from Chachoengsao CUMZ 5130 H shell from Sirisophon, Cambodia CUMZ 5131 and I shell from Srakaeo CUMZ 4286." figureDoi="10.3897/zookeys.870.36970.figure3" httpUri="https://binary.pensoft.net/fig/323908" pageId="0" pageNumber="51">Fig. 3G</figureCitation>
|
||
). However, anatomically they are indistinguishable from typical
|
||
<taxonomicName id="690D3900E9DFEEEF1AD36DA0F5F389B6" lsidName="G. rhombostoma" pageId="0" pageNumber="51" rank="species" species="rhombostoma">
|
||
<emphasis id="3D604C3A218498CF84B614AAD3A8C644" italics="true" pageId="0" pageNumber="51">G. rhombostoma</emphasis>
|
||
</taxonomicName>
|
||
and, therefore, they are considered conspecific.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |