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<document id="58C125B6A178EF78C1C0FDF3466A6734" ID-DOI="10.1080/00222933.2019.1646338" ID-GBIF-Dataset="9195a7a3-87bd-40a9-9165-2b2363b953b2" ID-ISSN="1464-5262" ID-Zenodo-Dep="3673704" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582028089786" checkinUser="valdenar" docAuthor="Nguyen, Anh D., Eguchi, Katsuyuki &amp; Hwang, Ui W." docDate="2019" docId="2C5787C93D7F5A61FE4F33D17131FA71" docLanguage="en" docName="JNATHIST.53.21–22.1369–1384.pdf.imf" docOrigin="Journal of Natural History 53 (21)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Hyleoglomeris hoanglien Nguyen ı Katsuyuki Eguchi &amp; Hwang 2019, sp." docType="treatment" docVersion="11" lastPageNumber="1374" masterDocId="D06EFFB13D7D5A67FF8D3612702EFFDB" masterDocTitle="Two new pill millipedes (Diplopoda: Glomerida: Glomeridae) from high mountains of Vietnam" masterLastPageNumber="1384" masterPageNumber="1369" pageNumber="1370" updateTime="1698719133712" updateUser="plazi">
<mods:mods id="76AB8A25D7A58DFF45BD7E697F7B3436" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="F95679E57C63E5608F00DEEE13FED6F9">Two new pill millipedes (Diplopoda: Glomerida: Glomeridae) from high mountains of Vietnam</mods:title>
</mods:titleInfo>
<mods:name id="D8774309602A39A43B278C56D49F08C4" type="personal">
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<mods:namePart id="FB760C04955A3EDFC73074785BA300A1">Nguyen, Anh D.</mods:namePart>
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<mods:namePart id="127AFA1D01AAB29178165DA8EB6327B2">Eguchi, Katsuyuki</mods:namePart>
</mods:name>
<mods:name id="1B85E5852F614D231009AA2AFE02D7AB" type="personal">
<mods:role id="FB36FC5101C6FD7140C287E41DC66B91">
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<mods:namePart id="A4AA135545D6B7F5F74AEC9A976B62E6">Hwang, Ui W.</mods:namePart>
</mods:name>
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<mods:title id="10B7B9F4752C23CBA249BFD8C31BE284">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="8EAB5C5CA21EF03A15EC325154AFB778">
<mods:date id="098025A0EC69811E9F349C1974AE3918">2019</mods:date>
<mods:detail id="3548A5498E1EC01E532EA10A54C0F771" type="pubDate">
<mods:number id="DD274F8EC840D6075A788E99A5244B7A">2019-08-20</mods:number>
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<mods:detail id="D1CF6086BCC6A0D30C9A9147013E91F1" type="volume">
<mods:number id="AC20658C1FB8F057F8FA1F383017319B">53</mods:number>
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<mods:classification id="FFEBE231E02780270EDCBF793323EDBA">journal article</mods:classification>
<mods:identifier id="62AC048C8709E76E599DDE06F240107B" type="DOI">10.1080/00222933.2019.1646338</mods:identifier>
<mods:identifier id="924C953E5824C06C1AA2BFD228D167D3" type="GBIF-Dataset">9195a7a3-87bd-40a9-9165-2b2363b953b2</mods:identifier>
<mods:identifier id="CC6315A58A8602C25B4CF8FE912F9877" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="AB2D89FCE0C9DB75F3D437C3F7F019B1" type="Zenodo-Dep">3673704</mods:identifier>
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<treatment id="2C5787C93D7F5A61FE4F33D17131FA71" ID-DOI="http://doi.org/10.5281/zenodo.3679865" ID-GBIF-Taxon="162162042" ID-Zenodo-Dep="3679865" LSID="urn:lsid:plazi:treatment:2C5787C93D7F5A61FE4F33D17131FA71" httpUri="http://treatment.plazi.org/id/2C5787C93D7F5A61FE4F33D17131FA71" lastPageId="6" lastPageNumber="1374" pageId="2" pageNumber="1370">
<subSubSection id="ECE465543D7F5A65FE4F33D1734FFA07" box="[450,865,1475,1500]" pageId="2" pageNumber="1370" type="nomenclature">
<paragraph id="A44136DF3D7F5A65FE4F33D1734FFA07" blockId="2.[450,865,1475,1535]" box="[450,865,1475,1500]" pageId="2" pageNumber="1370">
<heading id="FF0981B33D7F5A65FE4F33D1734FFA07" box="[450,865,1475,1500]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="1370" reason="2">
<taxonomicName id="63FE4D5C3D7F5A65FE4F33D1730CFA07" authority="Nguyen ı Katsuyuki Eguchi &amp; Hwang, 2019" authorityName="Nguyen ı Katsuyuki Eguchi &amp; Hwang" authorityYear="2019" box="[450,802,1475,1500]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="2" pageNumber="1369" phylum="Arthropoda" rank="species" species="hoanglien" status="sp.">
<emphasis id="968AEACD3D7F5A65FE4F33D172DBFA07" bold="true" box="[450,757,1475,1500]" italics="true" pageId="2" pageNumber="1370">Hyleoglomeris hoanglien</emphasis>
<taxonomicNameLabel id="8DB957B63D7F5A65FD7333D1730CFA07" box="[766,802,1475,1500]" pageId="2" pageNumber="1370" rank="species">
<emphasis id="968AEACD3D7F5A65FD7333D1730CFA07" bold="true" box="[766,802,1475,1500]" pageId="2" pageNumber="1370">sp.</emphasis>
</taxonomicNameLabel>
</taxonomicName>
<emphasis id="968AEACD3D7F5A65FCA133D17375FA07" bold="true" box="[812,859,1475,1500]" pageId="2" pageNumber="1370">nov</emphasis>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="ECE465543D7F5A65FDC833F472F0FA24" box="[581,734,1509,1535]" pageId="2" pageNumber="1370" type="description">
<paragraph id="A44136DF3D7F5A65FDC833F472F0FA24" blockId="2.[450,865,1475,1535]" box="[581,734,1509,1535]" pageId="2" pageNumber="1370">
<heading id="FF0981B33D7F5A65FDC833F472F0FA24" box="[581,734,1509,1535]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="1370" reason="5">
(
<figureCitation id="3CC52A5A3D7F5A65FDC033F472F8FA24" box="[589,726,1509,1535]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStartId-0="3.[160,225,919,942]" captionStartId-1="4.[160,225,975,998]" captionStartId-2="5.[160,225,1068,1091]" captionStartId-3="6.[160,225,944,967]" captionTargetBox-0="[164,1152,143,887]" captionTargetBox-1="[164,1152,143,942]" captionTargetBox-2="[263,1052,147,1036]" captionTargetBox-3="[357,957,145,910]" captionTargetId-0="figure@3.[163,1153,142,888]" captionTargetId-1="figure@4.[163,1153,142,943]" captionTargetId-2="figure@5.[258,1058,142,1036]" captionTargetId-3="figure@6.[355,960,142,912]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="Figure 1. Hyleglomeris hoanglien sp. nov., holotype (IEBR-Myr 672H). Whole-body, dorsal view (A), ventral view (B), lateral view (C); collum, anterior view (D)." captionText-1="Figure 2. Hyleoglomeris hoanglien sp. nov., holotype (IEBR-Myr 672H). Ocelli &amp; Tömösváry organ (A); Thoracic shield (B), Antenna (C), Anal shield (D)." captionText-2="Figure 3. Hyleoglomeris hoanglien sp. nov., paratype (IEBR-Myr 672P). Right telopod, posterior view (A), anterior view (B). Syncoxial lobe and horns, posterior view (D). Scale bar = 1 mm." captionText-3="Figure 4. Hyleoglomeris hoanglien sp. nov., paratype (IEBR-Myr 672P). Left telopod, anterior view (A), mesal view (B, C), posterior view (D). Scale bar = 0.3 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3673706" figureDoi-1="http://doi.org/10.5281/zenodo.3673708" figureDoi-2="http://doi.org/10.5281/zenodo.3673710" figureDoi-3="http://doi.org/10.5281/zenodo.3673712" httpUri-0="https://zenodo.org/record/3673706/files/figure.png" httpUri-1="https://zenodo.org/record/3673708/files/figure.png" httpUri-2="https://zenodo.org/record/3673710/files/figure.png" httpUri-3="https://zenodo.org/record/3673712/files/figure.png" pageId="2" pageNumber="1370">
Figures 1
<emphasis id="968AEACD3D7F5A65FD3733F772E6FA25" box="[698,712,1509,1534]" italics="true" pageId="2" pageNumber="1370">–</emphasis>
4
</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="ECE465543D7F5A64FF2D302F7125FB99" lastPageId="3" lastPageNumber="1371" pageId="2" pageNumber="1370" type="materials_examined">
<paragraph id="A44136DF3D7F5A65FF2D302F71AEF98C" blockId="2.[160,1155,1597,1727]" box="[160,384,1597,1623]" pageId="2" pageNumber="1370">
<emphasis id="968AEACD3D7F5A65FF2D302F71AEF98C" bold="true" box="[160,384,1597,1623]" italics="true" pageId="2" pageNumber="1370">Material examined</emphasis>
</paragraph>
<paragraph id="A44136DF3D7F5A65FF2D307272F4F964" blockId="2.[160,1155,1597,1727]" pageId="2" pageNumber="1370">
<materialsCitation id="14963C823D7F5A65FF2D307272F4F964" ID-GBIF-Occurrence="2565952786" collectingDate="2017-01" collectorName="Anh D. Nguyen &amp; K. Eguchi." county="One" elevation="900" location="Pai Hoang Lien National Parki" municipality="Sa Pai Hoang Lien National" pageId="2" pageNumber="1370" specimenCount="1" stateProvince="Lao Cai" typeStatus="holotype">
<emphasis id="968AEACD3D7F5A65FF2D3072713CF9A1" bold="true" box="[160,274,1632,1658]" italics="true" pageId="2" pageNumber="1370">
<typeStatus id="7B45887D3D7F5A65FF2D30727123F9A1" box="[160,269,1632,1658]" pageId="2" pageNumber="1370" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="4D204E533D7F5A65FEAC3070717EF9A1" box="[289,336,1634,1658]" pageId="2" pageNumber="1370">One</collectingCounty>
male (
<emphasis id="968AEACD3D7F5A65FE2D3070727BF9A1" bold="true" box="[416,597,1634,1658]" pageId="2" pageNumber="1370">IEBR-Myr 672H</emphasis>
)ı
<collectingRegion id="663AF83D3D7F5A65FDE63070732AF9A1" box="[619,772,1634,1658]" country="Vietnam" name="Lao Cai" pageId="2" pageNumber="1370">Lao Cai Prov.</collectingRegion>
ı
<collectingMunicipality id="4425ACA53D7F5A65FC833070746AF9A1" box="[782,1092,1634,1658]" pageId="2" pageNumber="1370">
Sa
<location id="A12160043D7F5A65FCBC3070746AF9A1" LSID="urn:lsid:plazi:treatment:2C5787C93D7F5A61FE4F33D17131FA71:A12160043D7F5A65FCBC3070746AF9A1" box="[817,1092,1634,1658]" county="One" municipality="Sa Pai Hoang Lien National" name="Pai Hoang Lien National Parki" pageId="2" pageNumber="1370" stateProvince="Lao Cai">Paı Hoang Lien National</location>
</collectingMunicipality>
Parkı short bamboo forestsı
<location id="A12160043D7F5A65FE2330977266F946" LSID="urn:lsid:plazi:treatment:2C5787C93D7F5A61FE4F33D17131FA71:A12160043D7F5A65FE2330977266F946" box="[430,584,1669,1693]" county="One" municipality="Sa Pai Hoang Lien National" name="Fanxipan Mt." pageId="2" pageNumber="1370" stateProvince="Lao Cai">Fanxipan Mt.</location>
ı 2ı800
<emphasis id="968AEACD3D7F5A65FD1E3096728EF947" box="[659,672,1668,1692]" italics="true" pageId="2" pageNumber="1370">–</emphasis>
2ı
<quantity id="63069B3A3D7F5A65FD3D309772D1F946" box="[688,767,1669,1693]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="1370" unit="m" value="900.0">
<elevation id="2FD3D1EC3D7F5A65FD3D309772D1F946" box="[688,767,1669,1693]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="2" pageNumber="1370" unit="m" value="900.0">900 m</elevation>
</quantity>
alt.ı 103°52
<emphasis id="968AEACD3D7F5A65FC00309673BDF947" bold="true" box="[909,915,1668,1692]" italics="true" pageId="2" pageNumber="1370">ʹ</emphasis>
42
<emphasis id="968AEACD3D7F5A65FC2330967397F947" bold="true" box="[942,953,1668,1692]" italics="true" pageId="2" pageNumber="1370">″</emphasis>
Eı 22°21
<emphasis id="968AEACD3D7F5A65FB9530967430F947" bold="true" box="[1048,1054,1668,1692]" italics="true" pageId="2" pageNumber="1370">ʹ</emphasis>
22
<emphasis id="968AEACD3D7F5A65FBB43096746AF947" bold="true" box="[1081,1092,1668,1692]" italics="true" pageId="2" pageNumber="1370">″</emphasis>
Nı 23 September
<date id="D040101F3D7F5A65FEA930B5714CF964" box="[292,354,1703,1727]" pageId="2" pageNumber="1370" value="2017-01">
<collectingDate id="C004E9F73D7F5A65FEA930B5714CF964" box="[292,354,1703,1727]" pageId="2" pageNumber="1370" value="2017-01">2017ı</collectingDate>
</date>
leg.
<collectorName id="090B53093D7F5A65FE1730B57267F964" box="[410,585,1703,1727]" pageId="2" pageNumber="1370">Anh D. Nguyen</collectorName>
&amp;
<collectorName id="090B53093D7F5A65FDE730B572F4F964" box="[618,730,1703,1727]" pageId="2" pageNumber="1370">K. Eguchi.</collectorName>
</materialsCitation>
</paragraph>
<caption id="F08166573D7E5A64FF2D358572C3FC17" ID-DOI="http://doi.org/10.5281/zenodo.3673706" ID-Zenodo-Dep="3673706" httpUri="https://zenodo.org/record/3673706/files/figure.png" pageId="3" pageNumber="1371" startId="3.[160,225,919,942]" targetBox="[164,1152,143,887]" targetPageId="3">
<paragraph id="A44136DF3D7E5A64FF2D358572C3FC17" blockId="3.[160,1156,919,972]" pageId="3" pageNumber="1371">
<emphasis id="968AEACD3D7E5A64FF2D358570D3FC75" bold="true" box="[160,253,919,942]" pageId="3" pageNumber="1371">Figure 1.</emphasis>
<taxonomicName id="63FE4D5C3D7E5A64FE8B358571C6FC75" authority="Nguyen ı Katsuyuki Eguchi &amp; Hwang, 2019" authorityName="Nguyen ı Katsuyuki Eguchi &amp; Hwang" authorityYear="2019" box="[262,488,919,942]" class="Diplopoda" family="Glomeridae" genus="Hyleglomeris" kingdom="Animalia" order="Glomerida" pageId="3" pageNumber="1371" phylum="Arthropoda" rank="species" species="hoanglien" status="sp. nov.">
<emphasis id="968AEACD3D7E5A64FE8B358571C6FC75" box="[262,488,919,942]" italics="true" pageId="3" pageNumber="1371">Hyleglomeris hoanglien</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DB957B63D7E5A64FE7F3585726AFC75" box="[498,580,919,942]" pageId="3" pageNumber="1371" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (IEBR-Myr 672H). Whole-body, dorsal view (A), ventral view (B), lateral view (C); collum, anterior view (D).
</paragraph>
</caption>
<paragraph id="A44136DF3D7E5A64FF2D32177125FB99" blockId="3.[160,1155,1029,1090]" pageId="3" pageNumber="1371">
<emphasis id="968AEACD3D7E5A64FF2D32177132FBC4" bold="true" box="[160,284,1029,1055]" italics="true" pageId="3" pageNumber="1371">
<typeStatus id="7B45887D3D7E5A64FF2D32177139FBC4" box="[160,279,1029,1055]" pageId="3" pageNumber="1371" type="paratype">Paratypes</typeStatus>
.
</emphasis>
One maleı four females (
<emphasis id="968AEACD3D7E5A64FDEC32157334FBC4" bold="true" box="[609,794,1031,1055]" pageId="3" pageNumber="1371">IEBR-Myr 672P</emphasis>
)ı with the same data as the holotype.
</paragraph>
</subSubSection>
<subSubSection id="ECE465543D7E5A64FF2D326F7325FABC" pageId="3" pageNumber="1371" type="diagnosis">
<paragraph id="A44136DF3D7E5A64FF2D326F713AFB4C" blockId="3.[160,1156,1149,1383]" box="[160,276,1149,1175]" pageId="3" pageNumber="1371">
<emphasis id="968AEACD3D7E5A64FF2D326F713AFB4C" bold="true" box="[160,276,1149,1175]" italics="true" pageId="3" pageNumber="1371">Diagnosis</emphasis>
</paragraph>
<paragraph id="A44136DF3D7E5A64FF2D32B07325FABC" blockId="3.[160,1156,1149,1383]" pageId="3" pageNumber="1371">
A small-sized (
<quantity id="63069B3A3D7E5A64FEB232B071BAFB61" box="[319,404,1186,1210]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="3" pageNumber="1371" unit="mm" value="2.3">2.3 mm</quantity>
wideı and
<quantity id="63069B3A3D7E5A64FD8232B0724DFB61" box="[527,611,1186,1210]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="3" pageNumber="1371" unit="mm" value="5.2">5.2 mm</quantity>
long) and black or blackish-brown body; ocelli 8 + 1; antennal tip with four large sensory cones; thoracic shield with eight transverse striae; telopods short and stout; syncoxital lobe bifid trapeziform; syncoxital horns highly elevatedı directed mesoventrally; tips of syncoxital horns with a long seta and a pointed spine; both prefemoral and femoral trichosteles present; femora condensedı with a ventrally directed process; tibia without modifications.
</paragraph>
</subSubSection>
<subSubSection id="ECE465543D7E5A64FF2D33B070C7F9DA" pageId="3" pageNumber="1371" type="etymology">
<paragraph id="A44136DF3D7E5A64FF2D33B07133FA67" blockId="3.[160,1155,1442,1537]" box="[160,285,1442,1468]" pageId="3" pageNumber="1371">
<emphasis id="968AEACD3D7E5A64FF2D33B07133FA67" bold="true" box="[160,285,1442,1468]" italics="true" pageId="3" pageNumber="1371">Etymology</emphasis>
</paragraph>
<paragraph id="A44136DF3D7E5A64FF2D33D570C7F9DA" blockId="3.[160,1155,1442,1537]" pageId="3" pageNumber="1371">
Named after the localityı
<emphasis id="968AEACD3D7E5A64FE3633D5721AFA04" box="[443,564,1479,1503]" italics="true" pageId="3" pageNumber="1371">‘hoanglien’</emphasis>
noun in appositionı where the
<typeStatus id="7B45887D3D7E5A64FC1B33D573E6FA04" box="[918,968,1479,1503]" pageId="3" pageNumber="1371">type</typeStatus>
specimens were found.
</paragraph>
</subSubSection>
<subSubSection id="ECE465543D7E5A62FF2D302F736DF964" lastPageId="5" lastPageNumber="1373" pageId="3" pageNumber="1371" type="description">
<paragraph id="A44136DF3D7E5A64FF2D302F7108F98C" blockId="3.[160,1155,1597,1692]" box="[160,294,1597,1623]" pageId="3" pageNumber="1371">
<emphasis id="968AEACD3D7E5A64FF2D302F7108F98C" bold="true" box="[160,294,1597,1623]" italics="true" pageId="3" pageNumber="1371">Description</emphasis>
</paragraph>
<paragraph id="A44136DF3D7E5A64FF2D307270D2F947" blockId="3.[160,1155,1597,1692]" pageId="3" pageNumber="1371">
<emphasis id="968AEACD3D7E5A64FF2D307270FAF9A1" bold="true" box="[160,212,1632,1658]" italics="true" pageId="3" pageNumber="1371">Size.</emphasis>
<typeStatus id="7B45887D3D7E5A64FF6830707162F9A1" box="[229,332,1634,1658]" pageId="3" pageNumber="1371" type="holotype">Holotype</typeStatus>
male (
<emphasis id="968AEACD3D7E5A64FE1130707274F9A1" bold="true" box="[412,602,1634,1658]" pageId="3" pageNumber="1371">IEBR-Myr 672 H</emphasis>
). Width of second tergum
<quantity id="63069B3A3D7E5A64FC1F307073C5F9A1" box="[914,1003,1634,1658]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="3" pageNumber="1371" unit="mm" value="2.3">2.3 mm</quantity>
; body length
<quantity id="63069B3A3D7E5A64FF2D309670D7F947" box="[160,249,1668,1692]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="3" pageNumber="1371" unit="mm" value="5.2">5.2 mm</quantity>
.
</paragraph>
<caption id="F08166573D795A63FF2D35DD72ADFBD8" ID-DOI="http://doi.org/10.5281/zenodo.3673708" ID-Zenodo-Dep="3673708" httpUri="https://zenodo.org/record/3673708/files/figure.png" pageId="4" pageNumber="1372" startId="4.[160,225,975,998]" targetBox="[164,1152,143,942]" targetPageId="4">
<paragraph id="A44136DF3D795A63FF2D35DD72ADFBD8" blockId="4.[160,1156,975,1027]" pageId="4" pageNumber="1372">
<emphasis id="968AEACD3D795A63FF2D35DD70D5FC3D" bold="true" box="[160,251,975,998]" pageId="4" pageNumber="1372">Figure 2.</emphasis>
<taxonomicName id="63FE4D5C3D795A63FE8E35DD723DFC3D" authority="Nguyen ı Katsuyuki Eguchi &amp; Hwang, 2019" authorityName="Nguyen ı Katsuyuki Eguchi &amp; Hwang" authorityYear="2019" box="[259,531,975,998]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="4" pageNumber="1369" phylum="Arthropoda" rank="species" species="hoanglien" status="sp.">
<emphasis id="968AEACD3D795A63FE8E35DD71C1FC3D" box="[259,495,975,998]" italics="true" pageId="4" pageNumber="1372">Hyleoglomeris hoanglien</emphasis>
<taxonomicNameLabel id="8DB957B63D795A63FE7B35DD723DFC3D" box="[502,531,975,998]" pageId="4" pageNumber="1372" rank="species">sp.</taxonomicNameLabel>
</taxonomicName>
nov., holotype (IEBR-Myr 672H). Ocelli &amp; Tömösváry organ (A); Thoracic shield (B), Antenna (C), Anal shield (D).
</paragraph>
</caption>
<paragraph id="A44136DF3D795A63FF2D325974AAFB5C" blockId="4.[160,1156,1099,1159]" pageId="4" pageNumber="1372">
<emphasis id="968AEACD3D795A63FF2D325970DBFBBE" bold="true" box="[160,245,1099,1125]" italics="true" pageId="4" pageNumber="1372">Colour.</emphasis>
Generally black or blackish-brown except for a medial brownish-yellow spotı two distolateral marbled brownish-yellow oval spots and light yellowish lateral margins.
</paragraph>
<paragraph id="A44136DF3D795A63FF2D32D171F9FA9E" blockId="4.[160,1155,1219,1349]" pageId="4" pageNumber="1372">
<emphasis id="968AEACD3D795A63FF2D32D170CBFB06" bold="true" box="[160,229,1219,1245]" italics="true" pageId="4" pageNumber="1372">Head.</emphasis>
Ocelli 8 + 1; lenses convexı black contrasting against to brownish background of head. Tömösváry organs transverse ovalı ca. 1.5 X wider than long. Antenna clavate apically; antennomere 6 largeı ca. 2.0
<emphasis id="968AEACD3D795A63FDDE3318724EFAF9" box="[595,608,1290,1314]" italics="true" pageId="4" pageNumber="1372">–</emphasis>
2.5 X longer than wide. Antennal tip with four largeı apical sensory cones.
</paragraph>
<paragraph id="A44136DF3D795A63FF2D339272DCF9FE" blockId="4.[160,1156,1408,1573]" pageId="4" pageNumber="1372">
<emphasis id="968AEACD3D795A63FF2D339270CFFA41" bold="true" box="[160,225,1408,1434]" italics="true" pageId="4" pageNumber="1372">Body.</emphasis>
Collum semicircuları with a very large marbled brownish-black spot centrallyı and two transverse striae; other part of collum black. Thoracic shield (= tergum 2) with a narrow hyposchismı not reaching caudal margin; with 8 clearly distinct transverse striae of which all cross the dorsum. Other terga with 3
<emphasis id="968AEACD3D795A63FD5E33F872CEF9D9" box="[723,736,1514,1538]" italics="true" pageId="4" pageNumber="1372">–</emphasis>
4 shortı vague transverse striae. Anal shield strongly roundedı emarginate mediocaudally.
</paragraph>
<paragraph id="A44136DF3D795A63FF2D3072745BF964" blockId="4.[160,1156,1632,1727]" pageId="4" pageNumber="1372">
<emphasis id="968AEACD3D795A63FF2D307270F5F9A1" bold="true" box="[160,219,1632,1658]" italics="true" pageId="4" pageNumber="1372">Legs.</emphasis>
Leg-pair 17 strongly reducedı 4-segmented; coxite with a rounded outer lobeı with a setiferous tubercle distomesally; podomere 1 with a small distomesal setiferous knob. Leg-pair 18 less strongly reducedı 4-segmentedı with a simple syncoxital notch.
</paragraph>
<caption id="F08166573D785A62FF2D323E742BFBBB" ID-DOI="http://doi.org/10.5281/zenodo.3673710" ID-Zenodo-Dep="3673710" httpUri="https://zenodo.org/record/3673710/files/figure.png" pageId="5" pageNumber="1373" startId="5.[160,225,1068,1091]" targetBox="[263,1052,147,1036]" targetPageId="5">
<paragraph id="A44136DF3D785A62FF2D323E742BFBBB" blockId="5.[160,1155,1068,1120]" pageId="5" pageNumber="1373">
<emphasis id="968AEACD3D785A62FF2D323E70D2FB98" bold="true" box="[160,252,1068,1091]" pageId="5" pageNumber="1373">Figure 3.</emphasis>
<taxonomicName id="63FE4D5C3D785A62FE89323E7238FB98" authority="Nguyen ı Katsuyuki Eguchi &amp; Hwang, 2019" authorityName="Nguyen ı Katsuyuki Eguchi &amp; Hwang" authorityYear="2019" box="[260,534,1068,1091]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="5" pageNumber="1369" phylum="Arthropoda" rank="species" species="hoanglien" status="sp.">
<emphasis id="968AEACD3D785A62FE89323E71DFFB98" box="[260,497,1068,1091]" italics="true" pageId="5" pageNumber="1373">Hyleoglomeris hoanglien</emphasis>
<taxonomicNameLabel id="8DB957B63D785A62FE77323E7238FB98" box="[506,534,1068,1091]" pageId="5" pageNumber="1373" rank="species">sp.</taxonomicNameLabel>
</taxonomicName>
nov., paratype (IEBR-Myr 672P). Right telopod, posterior view (A), anterior view (B). Syncoxial lobe and horns, posterior view (D). Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph id="A44136DF3D785A62FF2D328B7432FA7D" blockId="5.[160,1156,1177,1446]" pageId="5" pageNumber="1373">
<emphasis id="968AEACD3D785A62FF2D328B71E9FB68" bold="true" box="[160,455,1177,1203]" pageId="5" pageNumber="1373">
<emphasis id="968AEACD3D785A62FF2D328B7122FB68" bold="true" box="[160,268,1177,1203]" italics="true" pageId="5" pageNumber="1373">Genitalia</emphasis>
(
<figureCitation id="3CC52A5A3D785A62FEAE32887199FB68" box="[291,439,1178,1203]" captionStart-0="Figure 3" captionStart-1="Figure 4" captionStart-2="Figure 5" captionStartId-0="5.[160,225,1068,1091]" captionStartId-1="6.[160,225,944,967]" captionStartId-2="7.[160,225,978,1001]" captionTargetBox-0="[263,1052,147,1036]" captionTargetBox-1="[357,957,145,910]" captionTargetBox-2="[163,1153,142,936]" captionTargetId-0="figure@5.[258,1058,142,1036]" captionTargetId-1="figure@6.[355,960,142,912]" captionTargetId-2="figure@7.[163,1153,142,946]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="Figure 3. Hyleoglomeris hoanglien sp. nov., paratype (IEBR-Myr 672P). Right telopod, posterior view (A), anterior view (B). Syncoxial lobe and horns, posterior view (D). Scale bar = 1 mm." captionText-1="Figure 4. Hyleoglomeris hoanglien sp. nov., paratype (IEBR-Myr 672P). Left telopod, anterior view (A), mesal view (B, C), posterior view (D). Scale bar = 0.3 mm." captionText-2="Figure 5. Hyleoglomeris hoanglien sp. nov., paratype (IEBR-Myr 672P). Tibia of left telopod, ventral view (A); lateral surface (B); syncoxital lobe and horns, anterior view (C); distal part of the syncoxital horn (D)." figureDoi-0="http://doi.org/10.5281/zenodo.3673710" figureDoi-1="http://doi.org/10.5281/zenodo.3673712" figureDoi-2="http://doi.org/10.5281/zenodo.3673714" httpUri-0="https://zenodo.org/record/3673710/files/figure.png" httpUri-1="https://zenodo.org/record/3673712/files/figure.png" httpUri-2="https://zenodo.org/record/3673714/files/figure.png" pageId="5" pageNumber="1373">Figures 3–5</figureCitation>
).
</emphasis>
Telopods shortı stoutı and incrassate. Syncoxital lobe bifid trapeziform. Syncoxital horns highly elevatedı directed mesoventrallyı with numerous setae; tips with a long seta and a pointed spine. Prefemur (
<emphasis id="968AEACD3D785A62FCC832F3735BFB22" bold="true" box="[837,885,1249,1273]" pageId="5" pageNumber="1373">pref</emphasis>
) micropapillate laterally and posterolaterallyı with a longı prominentı finger-shaped trichostele (
<emphasis id="968AEACD3D785A62FC31331173C3FAC0" bold="true" box="[956,1005,1283,1307]" pageId="5" pageNumber="1373">pret</emphasis>
). Femora (
<emphasis id="968AEACD3D785A62FBE833117452FAC0" bold="true" box="[1125,1148,1283,1307]" pageId="5" pageNumber="1373">fe</emphasis>
) condensedı also with a longı prominentı finger-shaped trichostele (
<emphasis id="968AEACD3D785A62FC1A33347399FAE5" bold="true" box="[919,951,1318,1342]" pageId="5" pageNumber="1373">fet</emphasis>
). Femoral process (
<emphasis id="968AEACD3D785A62FF2A335B7091FABA" bold="true" box="[167,191,1353,1377]" pageId="5" pageNumber="1373">fp</emphasis>
) directed ventrally; base broadly enlarged; tip folded; posterior surface densely micropapillate. Tibia (
<emphasis id="968AEACD3D785A62FEC033797173FA58" bold="true" box="[333,349,1387,1411]" pageId="5" pageNumber="1373">ti</emphasis>
) without modificationsı but the tibial process (
<emphasis id="968AEACD3D785A62FCD13379735BFA58" bold="true" box="[860,885,1387,1411]" pageId="5" pageNumber="1373">tp</emphasis>
) directed mesoventrally. Tarsus (
<emphasis id="968AEACD3D785A62FF79339C7125FA7D" bold="true" box="[244,267,1422,1446]" pageId="5" pageNumber="1373">ta</emphasis>
) curvedı bearing an apical seta; inner side also densely micropapillate.
</paragraph>
<paragraph id="A44136DF3D785A62FF2D33C17110F9CB" blockId="5.[160,1155,1491,1552]" pageId="5" pageNumber="1373">
<emphasis id="968AEACD3D785A62FF2D33C1713BFA36" bold="true" box="[160,277,1491,1517]" italics="true" pageId="5" pageNumber="1373">Variation.</emphasis>
Width of second tergum
<quantity id="63069B3A3D785A62FDCD33C772EDFA36" box="[576,707,1493,1517]" metricMagnitude="-3" metricUnit="m" metricValue="2.85" metricValueMax="3.2" metricValueMin="2.5" pageId="5" pageNumber="1373" unit="mm" value="2.85" valueMax="3.2" valueMin="2.5">
2.5
<emphasis id="968AEACD3D785A62FDEC33C77240FA36" box="[609,622,1493,1517]" italics="true" pageId="5" pageNumber="1373">–</emphasis>
3.2 mm
</quantity>
in the female; body length
<quantity id="63069B3A3D785A62FB8D33C774ADFA36" box="[1024,1155,1493,1517]" metricMagnitude="-3" metricUnit="m" metricValue="5.95" metricValueMax="6.7" metricValueMin="5.2" pageId="5" pageNumber="1373" unit="mm" value="5.95" valueMax="6.7" valueMin="5.2">
5.2
<emphasis id="968AEACD3D785A62FBAC33C77400FA36" box="[1057,1070,1493,1517]" italics="true" pageId="5" pageNumber="1373">–</emphasis>
6.7 mm
</quantity>
in the female.
</paragraph>
<paragraph id="A44136DF3D785A62FF2D302F7111F98C" blockId="5.[160,1155,1597,1727]" box="[160,319,1597,1623]" pageId="5" pageNumber="1373">
<emphasis id="968AEACD3D785A62FF2D302F7111F98C" bold="true" box="[160,319,1597,1623]" italics="true" pageId="5" pageNumber="1373">DNA barcode</emphasis>
</paragraph>
<paragraph id="A44136DF3D785A62FF2D3070736DF964" blockId="5.[160,1155,1597,1727]" pageId="5" pageNumber="1373">
COI barcode data (partial) is for the
<typeStatus id="7B45887D3D785A62FDB5307072B2F9A1" box="[568,668,1634,1658]" pageId="5" pageNumber="1373" type="paratype">paratype</typeStatus>
uploaded to GenBank under the accession number MH248038. The new species shares the identity of 87% and 86% with
<taxonomicName id="63FE4D5C3D785A62FF2D30B57184F964" authorityName="Heller" authorityYear="1858" box="[160,426,1703,1727]" class="Diplopoda" family="Glomeridae" genus="Trachysphaera" kingdom="Animalia" order="Glomerida" pageId="5" pageNumber="1373" phylum="Arthropoda" rank="species" species="schmidtii">
<emphasis id="968AEACD3D785A62FF2D30B57184F964" box="[160,426,1703,1727]" italics="true" pageId="5" pageNumber="1373">Trachysphaera schmidtii</emphasis>
</taxonomicName>
and
<taxonomicName id="63FE4D5C3D785A62FE6B30B57286F964" box="[486,680,1703,1727]" class="Diplopoda" family="Glomeridae" genus="Glomeris" kingdom="Animalia" order="Glomerida" pageId="5" pageNumber="1373" phylum="Arthropoda" rank="species" species="connexa">
<emphasis id="968AEACD3D785A62FE6B30B57286F964" box="[486,680,1703,1727]" italics="true" pageId="5" pageNumber="1373">Glomeris connexa</emphasis>
</taxonomicName>
ı respectively.
</paragraph>
</subSubSection>
<caption id="F08166573D7B5A61FF2D35A272C7FC3F" ID-DOI="http://doi.org/10.5281/zenodo.3673712" ID-Zenodo-Dep="3673712" httpUri="https://zenodo.org/record/3673712/files/figure.png" pageId="6" pageNumber="1374" startId="6.[160,225,944,967]" targetBox="[357,957,145,910]" targetPageId="6">
<paragraph id="A44136DF3D7B5A61FF2D35A272C7FC3F" blockId="6.[160,1156,944,996]" pageId="6" pageNumber="1374">
<emphasis id="968AEACD3D7B5A61FF2D35A270D4FC1C" bold="true" box="[160,250,944,967]" pageId="6" pageNumber="1374">Figure 4.</emphasis>
<taxonomicName id="63FE4D5C3D7B5A61FE8F35A2723FFC1C" authority="Nguyen ı Katsuyuki Eguchi &amp; Hwang, 2019" authorityName="Nguyen ı Katsuyuki Eguchi &amp; Hwang" authorityYear="2019" box="[258,529,944,967]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="6" pageNumber="1369" phylum="Arthropoda" rank="species" species="hoanglien" status="sp.">
<emphasis id="968AEACD3D7B5A61FE8F35A271C3FC1C" box="[258,493,944,967]" italics="true" pageId="6" pageNumber="1374">Hyleoglomeris hoanglien</emphasis>
<taxonomicNameLabel id="8DB957B63D7B5A61FE7835A2723FFC1C" box="[501,529,944,967]" pageId="6" pageNumber="1374" rank="species">sp.</taxonomicNameLabel>
</taxonomicName>
nov., paratype (IEBR-Myr 672P). Left telopod, anterior view (A), mesal view (B, C), posterior view (D). Scale bar = 0.3 mm.
</paragraph>
</caption>
<subSubSection id="ECE465543D7B5A61FF2D32017131FA71" pageId="6" pageNumber="1374" type="discussion">
<paragraph id="A44136DF3D7B5A61FF2D32017128FBF6" blockId="6.[160,1156,1043,1450]" box="[160,262,1043,1069]" pageId="6" pageNumber="1374">
<emphasis id="968AEACD3D7B5A61FF2D32017128FBF6" bold="true" box="[160,262,1043,1069]" italics="true" pageId="6" pageNumber="1374">Remarks</emphasis>
</paragraph>
<paragraph id="A44136DF3D7B5A61FF2D32257131FA71" blockId="6.[160,1156,1043,1450]" pageId="6" pageNumber="1374">
The new species is known as the smallest glomerid recorded in
<collectingCountry id="DCE9764F3D7B5A61FCFA322573FBFB94" box="[887,981,1079,1103]" name="Vietnam" pageId="6" pageNumber="1374">Vietnam</collectingCountry>
. The previously smallestı troglobiotic speciesı
<taxonomicName id="63FE4D5C3D7B5A61FE6732487276FBA9" authorityName="Golovatchi Geoffroy and VandenSpiegeli" authorityYear="2013" box="[490,600,1114,1138]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="6" pageNumber="1374" phylum="Arthropoda" rank="species" species="spelaea">
<emphasis id="968AEACD3D7B5A61FE6732487276FBA9" box="[490,600,1114,1138]" italics="true" pageId="6" pageNumber="1374">H. spelaea</emphasis>
</taxonomicName>
ı has a width of the second tergum of ca. 3.0 mm. The new species seems to belong to the
<taxonomicName id="63FE4D5C3D7B5A61FD08326F72FDFB4E" baseAuthorityName="Brolemanni" baseAuthorityYear="1896" box="[645,723,1149,1173]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="6" pageNumber="1374" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="968AEACD3D7B5A61FD08326F72FDFB4E" box="[645,723,1149,1173]" italics="true" pageId="6" pageNumber="1374">sinensis</emphasis>
</taxonomicName>
-group characterised by telopods being incrassate (
<bibRefCitation id="C06F4B2E3D7B5A61FE94328D7226FB6C" author="Golovatch SIi Geoffroy JJi Mauries JP" box="[281,520,1183,1207]" pageId="6" pageNumber="1374" pagination="887 - 915" refId="ref5134" refString="Golovatch SIi Geoffroy JJi Mauries JP. 2006. Review of the millipede genus Hyleoglomeris Verhoeffi 1910 (Diplopodai Glomeridai Glomeridae) i with descriptions of new species from caves in Southeast Asia. Zoosystema. 28 (4): 887 - 915." type="journal article" year="2006">Golovatch et al. 2006</bibRefCitation>
). This group currently consists of two speciesı
<taxonomicName id="63FE4D5C3D7B5A61FB98328D7146FB01" authority="(Brolemanni 1896)" baseAuthorityName="Brolemanni" baseAuthorityYear="1896" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="6" pageNumber="1374" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="968AEACD3D7B5A61FB98328D74ACFB6C" box="[1045,1154,1183,1207]" italics="true" pageId="6" pageNumber="1374">H. sinensis</emphasis>
(Brölemannı 1896)
</taxonomicName>
and
<taxonomicName id="63FE4D5C3D7B5A61FE2932D072D4FB01" authority="Golovatchi 1981" authorityName="Golovatchi" authorityYear="1981" box="[420,762,1218,1242]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="6" pageNumber="1374" phylum="Arthropoda" rank="species" species="emarginata">
<emphasis id="968AEACD3D7B5A61FE2932D07212FB01" box="[420,572,1218,1242]" italics="true" pageId="6" pageNumber="1374">H. emarginata</emphasis>
Golovatchı 1981
</taxonomicName>
ı both from
<collectingCountry id="DCE9764F3D7B5A61FCF232D07391FB01" box="[895,959,1218,1242]" name="China" pageId="6" pageNumber="1374">China</collectingCountry>
. The new species differs from both species in its smaller size (
<quantity id="63069B3A3D7B5A61FDFB32F772E4FB26" box="[630,714,1253,1277]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="6" pageNumber="1374" unit="mm" value="2.3">2.3 mm</quantity>
wideı
<quantity id="63069B3A3D7B5A61FC9C32F7734DFB26" box="[785,867,1253,1277]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="6" pageNumber="1374" unit="mm" value="5.2">5.2 mm</quantity>
long vs. 5.5
<emphasis id="968AEACD3D7B5A61FC6932F673DFFB27" box="[996,1009,1252,1276]" italics="true" pageId="6" pageNumber="1374">–</emphasis>
6.0 mm wideı
<quantity id="63069B3A3D7B5A61FF2D33157130FAC4" box="[160,286,1287,1311]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="9.5" metricValueMin="8.5" pageId="6" pageNumber="1374" unit="mm" value="9.0" valueMax="9.5" valueMin="8.5">
8.5
<emphasis id="968AEACD3D7B5A61FF32331570E2FAC4" box="[191,204,1287,1311]" italics="true" pageId="6" pageNumber="1374">–</emphasis>
9.5 mm
</quantity>
long vs 3.0
<emphasis id="968AEACD3D7B5A61FE1B3315718DFAC4" box="[406,419,1287,1311]" italics="true" pageId="6" pageNumber="1374">–</emphasis>
4.0 mm wideı 6.0
<emphasis id="968AEACD3D7B5A61FDD433157248FAC4" box="[601,614,1287,1311]" italics="true" pageId="6" pageNumber="1374">–</emphasis>
9.0 mm long)ı having 8 + 1 ocelliı and in the shape and direction of syncoxital horns. Syncoxital horns are highly elevatedı directed mesoventrallyı with numerous setae; tips with a long seta and a pointed spine in
<taxonomicName id="63FE4D5C3D7B5A61FCD2335F74AAFABE" authority="Nguyen ı Katsuyuki Eguchi &amp; Hwang, 2019" authorityName="Nguyen ı Katsuyuki Eguchi &amp; Hwang" authorityYear="2019" box="[863,1156,1357,1381]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="6" pageNumber="1369" phylum="Arthropoda" rank="species" species="hoanglien" status="sp.">
<emphasis id="968AEACD3D7B5A61FCD2335F7471FABE" box="[863,1119,1357,1381]" italics="true" pageId="6" pageNumber="1374">Hyleoglomeris hoanglien</emphasis>
<taxonomicNameLabel id="8DB957B63D7B5A61FBE8335F74AAFABE" box="[1125,1156,1357,1381]" pageId="6" pageNumber="1374" rank="species">sp.</taxonomicNameLabel>
</taxonomicName>
nov.; directed mesally and denticulate in
<taxonomicName id="63FE4D5C3D7B5A61FDD8337D7290FA5C" baseAuthorityName="Brolemanni" baseAuthorityYear="1896" box="[597,702,1391,1415]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="6" pageNumber="1374" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="968AEACD3D7B5A61FDD8337D7290FA5C" box="[597,702,1391,1415]" italics="true" pageId="6" pageNumber="1374">H. sinensis</emphasis>
</taxonomicName>
; directed subventrally and lanceolate in
<taxonomicName id="63FE4D5C3D7B5A61FBE0337D7137FA71" authorityName="Golovatchi" authorityYear="1981" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="6" pageNumber="1374" phylum="Arthropoda" rank="species" species="emarginata">
<emphasis id="968AEACD3D7B5A61FBE0337D7137FA71" italics="true" pageId="6" pageNumber="1374">H. emarginata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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