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<document id="7423C92C115316D8FA5CCA386F5ED11F" ID-DOI="10.1080/00222933.2019.1646338" ID-GBIF-Dataset="9195a7a3-87bd-40a9-9165-2b2363b953b2" ID-ISSN="1464-5262" ID-Zenodo-Dep="3673704" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582028089786" checkinUser="valdenar" docAuthor="Nguyen, Anh D., Eguchi, Katsuyuki &amp; Hwang, Ui W." docDate="2019" docId="2C5787C93D705A69FF2D358474AAFD92" docLanguage="en" docName="JNATHIST.53.2122.13691384.pdf.imf" docOrigin="Journal of Natural History 53 (21)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Hyleoglomeris Verhoeffi 1910" docType="key" docVersion="11" lastPageNumber="1382" masterDocId="D06EFFB13D7D5A67FF8D3612702EFFDB" masterDocTitle="Two new pill millipedes (Diplopoda: Glomerida: Glomeridae) from high mountains of Vietnam" masterLastPageNumber="1384" masterPageNumber="1369" pageNumber="1381" updateTime="1698719133712" updateUser="plazi">
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<mods:title id="C686E895AA888594C19009F05FBB0F6C">Two new pill millipedes (Diplopoda: Glomerida: Glomeridae) from high mountains of Vietnam</mods:title>
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<mods:namePart id="7CC7DB7D67E3A9D23C0123EA617374D4">Nguyen, Anh D.</mods:namePart>
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<mods:namePart id="67F358D50E986088D2693DD96A3E4009">Eguchi, Katsuyuki</mods:namePart>
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<mods:namePart id="CE2BC98CF041BC086B03E5A075375385">Hwang, Ui W.</mods:namePart>
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<mods:title id="308F479818CA17F209E2534DAA938A42">Journal of Natural History</mods:title>
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<mods:date id="B136A09D9542AA002C69A9A421A65D93">2019</mods:date>
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<treatment id="2C5787C93D705A69FF2D358474AAFD92" ID-DOI="http://doi.org/10.5281/zenodo.3679859" ID-GBIF-Taxon="162162041" ID-Zenodo-Dep="3679859" LSID="urn:lsid:plazi:treatment:2C5787C93D705A69FF2D358474AAFD92" httpUri="http://treatment.plazi.org/id/2C5787C93D705A69FF2D358474AAFD92" lastPageId="14" lastPageNumber="1382" pageId="13" pageNumber="1381">
<subSubSection id="ECE465543D705A6AFF2D358472E8FC75" box="[160,710,918,942]" pageId="13" pageNumber="1381" type="nomenclature">
<paragraph id="A44136DF3D705A6AFF2D358472E8FC75" blockId="13.[160,710,918,942]" box="[160,710,918,942]" pageId="13" pageNumber="1381">
<heading id="FF0981B33D705A6AFF2D358472E8FC75" box="[160,710,918,942]" pageId="13" pageNumber="1381">
<emphasis id="968AEACD3D705A6AFF2D358472E8FC75" bold="true" box="[160,710,918,942]" pageId="13" pageNumber="1381">
A key to
<taxonomicName id="63FE4D5C3D705A6AFE9D35847199FC75" ID-CoL="7PL2P" authorityName="Verhoeffi" authorityYear="1910" box="[272,439,918,942]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="13" pageNumber="1381" phylum="Arthropoda" rank="genus">
<emphasis id="968AEACD3D705A6AFE9D35847199FC75" bold="true" box="[272,439,918,942]" italics="true" pageId="13" pageNumber="1381">Hyleoglomeris</emphasis>
</taxonomicName>
species from
<collectingCountry id="DCE9764F3D705A6AFDEC358472E8FC75" box="[609,710,918,942]" name="Vietnam" pageId="13" pageNumber="1381">Vietnam</collectingCountry>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="ECE465543D705A69FF2D35CB74AAFD92" lastPageId="14" lastPageNumber="1382" pageId="13" pageNumber="1381" type="key">
<key id="D06F61EE3D705A69FF2D35CB74AAFD92" lastPageId="14" lastPageNumber="1382" pageId="13" pageNumber="1381">
<paragraph id="A44136DF3D705A6AFF2D35CB74ADFBCF" blockId="13.[160,1155,985,1044]" pageId="13" pageNumber="1381">
<keyStep id="1F0A2E7A3D705A6AFF2D35CB74ADFBCF" pageId="13" pageNumber="1381">
<keyLead id="1F0F95EA3D705A6AFF2D35CB74ADFBCF" pageId="13" pageNumber="1381">- Body largely or entirely unpigmented. Troglobites ................................................................... 2 Body well pigmented. Non-troglobites ........................................................................................... 4</keyLead>
</keyStep>
</paragraph>
<paragraph id="A44136DF3D705A6AFF2D322374ADFB6A" blockId="13.[160,1155,1073,1201]" pageId="13" pageNumber="1381">
<keyStep id="1F0A2E7A3D705A6AFF2D322374ADFB6A" pageId="13" pageNumber="1381">
<keyLead id="1F0F95EA3D705A6AFF2D322374ADFB6A" pageId="13" pageNumber="1381">
- Body length ca. 6.0 mmı width of second tergum ca. 3.0 mm. Thoracic shield with six transverse striae. Telopodite with subquadrate syncoxital lobe ............ ............
<taxonomicName id="63FE4D5C3D705A6AFB9F324674ADFBB7" ID-CoL="3NF2L" authority="Body" authorityName="Body" box="[1042,1155,1108,1132]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="13" pageNumber="1381" phylum="Arthropoda" rank="species" species="spelaea">
<emphasis id="968AEACD3D705A6AFB9F324674ADFBB7" box="[1042,1155,1108,1132]" italics="true" pageId="13" pageNumber="1381">H. spelaea</emphasis>
</taxonomicName>
Body length ca. 8.5
<emphasis id="968AEACD3D705A6AFE07326471B9FB55" box="[394,407,1142,1166]" italics="true" pageId="13" pageNumber="1381"></emphasis>
10.0 mmı width of second tergum ca. 4.0
<emphasis id="968AEACD3D705A6AFCE732647359FB55" box="[874,887,1142,1166]" italics="true" pageId="13" pageNumber="1381"></emphasis>
<quantity id="63069B3A3D705A6AFCFA326473E1FB55" box="[887,975,1142,1166]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="13" pageNumber="1381" unit="mm" value="4.1">4.1 mm</quantity>
. Thoracic shield with 7 or more transverse striae. Telopodite with subtrapeziform syncoxital lobe ..... 3
</keyLead>
</keyStep>
</paragraph>
<paragraph id="A44136DF3D705A6AFF2D32DC74ADFA95" blockId="13.[160,1156,1230,1358]" pageId="13" pageNumber="1381">
<keyStep id="1F0A2E7A3D705A6AFF2D32DC74ADFA95" pageId="13" pageNumber="1381">
<keyLead id="1F0F95EA3D705A6AFF2D32DC74ADFA95" pageId="13" pageNumber="1381">
- Syncoxital lobe broadly subtrapeziformı slightly concave. Syncoxital horns without an apical seta ............................................................. .............................................................
<taxonomicName id="63FE4D5C3D705A6AFC6432E374ADFAD2" ID-CoL="6MN37" authority="Syncoxital" authorityName="Syncoxital" box="[1001,1155,1265,1289]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="13" pageNumber="1381" phylum="Arthropoda" rank="species" species="cavernicola">
<emphasis id="968AEACD3D705A6AFC6432E374ADFAD2" box="[1001,1155,1265,1289]" italics="true" pageId="13" pageNumber="1381">H. cavernicola</emphasis>
</taxonomicName>
Syncoxital lobe not broadly subtrapeziformı not concave. Syncoxital horns with an apical seta ............................................................... ...............................................................
<taxonomicName id="63FE4D5C3D705A6AFC71332474ADFA95" ID-CoL="3NF2M" authorityName="Golovatchi Geoffroy and Mauriesi" authorityYear="2006" box="[1020,1155,1334,1358]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="13" pageNumber="1381" phylum="Arthropoda" rank="species" species="speophila">
<emphasis id="968AEACD3D705A6AFC71332474ADFA95" box="[1020,1155,1334,1358]" italics="true" pageId="13" pageNumber="1381">H. speophila</emphasis>
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</paragraph>
<paragraph id="A44136DF3D705A6AFF2D337E74ADFA12" blockId="13.[160,1155,1388,1481]" pageId="13" pageNumber="1381">
<keyStep id="1F0A2E7A3D705A6AFF2D337E74ADFA12" pageId="13" pageNumber="1381">
<keyLead id="1F0F95EA3D705A6AFF2D337E74ADFA12" pageId="13" pageNumber="1381">
- Coloration pattern annulated. Leg 17 3-segmented. Central
<collectingCountry id="DCE9764F3D705A6AFCD1337E7397FA5F" box="[860,953,1388,1412]" name="Vietnam" pageId="13" pageNumber="1381">Vietnam</collectingCountry>
..... .....
<taxonomicName id="63FE4D5C3D705A6AFB87337E74ADFA5F" ID-CoL="6MMR6" authority="Coloration" authorityName="Coloration" box="[1034,1155,1388,1412]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="13" pageNumber="1381" phylum="Arthropoda" rank="species" species="colorata">
<emphasis id="968AEACD3D705A6AFB87337E74ADFA5F" box="[1034,1155,1388,1412]" italics="true" pageId="13" pageNumber="1381">H. colorata</emphasis>
</taxonomicName>
Coloration pattern not annulated. Leg 17 4-segmented. Southern and Northern
<collectingCountry id="DCE9764F3D705A6AFF3C33A37120FA12" box="[177,270,1457,1481]" name="Vietnam" pageId="13" pageNumber="1381">Vietnam</collectingCountry>
.......................................................................................................................................................... 5
</keyLead>
</keyStep>
</paragraph>
<paragraph id="A44136DF3D705A6AFF2D33F474ADF9FA" blockId="13.[160,1155,1510,1569]" pageId="13" pageNumber="1381">
<keyStep id="1F0A2E7A3D705A6AFF2D33F474ADF9FA" pageId="13" pageNumber="1381">
<keyLead id="1F0F95EA3D705A6AFF2D33F474ADF9FA" pageId="13" pageNumber="1381">- Syncoxital lobe subtrapeziformı slightly concave medially .................................................... 6 Syncoxital lobe variable in shape but not subtrapeziform .................................................... 8</keyLead>
</keyStep>
</paragraph>
<paragraph id="A44136DF3D705A6AFF2D302C74ADF965" blockId="13.[160,1159,1598,1726]" pageId="13" pageNumber="1381">
<keyStep id="1F0A2E7A3D705A6AFF2D302C74ADF965" pageId="13" pageNumber="1381">
<keyLead id="1F0F95EA3D705A6AFF2D302C74ADF965" pageId="13" pageNumber="1381">
- Thoracic shield with dark spots. Terga 3
<emphasis id="968AEACD3D705A6AFDFA302C72AAF98D" box="[631,644,1598,1622]" italics="true" pageId="13" pageNumber="1381"></emphasis>
11ı each with 3 rows of large yellow spots .... .................................................................................................................................................
<taxonomicName id="63FE4D5C3D705A6AFC52307374ADF9A2" ID-CoL="3NF2V" authority="Thoracic" authorityName="Thoracic" box="[991,1155,1633,1657]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="13" pageNumber="1381" phylum="Arthropoda" rank="species" species="triangulifera">
<emphasis id="968AEACD3D705A6AFC52307374ADF9A2" box="[991,1155,1633,1657]" italics="true" pageId="13" pageNumber="1381">H. triangulifera</emphasis>
</taxonomicName>
Thoracic shield and terga 3
<emphasis id="968AEACD3D705A6AFE6D309171C3F940" box="[480,493,1667,1691]" italics="true" pageId="13" pageNumber="1381"></emphasis>
11ı each with two large spots strongly contrasting with the background. ................................................................................................................................................. 7
</keyLead>
</keyStep>
</paragraph>
<paragraph id="A44136DF3D735A69FF2D369C74ADFED5" blockId="14.[160,1156,142,270]" pageId="14" pageNumber="1382">
<keyStep id="1F0A2E7A3D735A69FF2D369C74ADFED5" pageId="14" pageNumber="1382">
<keyLead id="1F0F95EA3D735A69FF2D369C74ADFED5" pageId="14" pageNumber="1382">
- Tergal spots blackishı strongly contrasting with yellow-brown background. Thoracic shield with 6
<emphasis id="968AEACD3D735A69FECA36A3717AFF12" box="[327,340,177,201]" italics="true" pageId="14" pageNumber="1382"></emphasis>
7 transverse striaeı of which 3
<emphasis id="968AEACD3D735A69FD2736A37299FF12" box="[682,695,177,201]" italics="true" pageId="14" pageNumber="1382"></emphasis>
4 cross the dorsum ..... .....
<taxonomicName id="63FE4D5C3D735A69FC6836A374AAFF12" ID-CoL="6MN2V" authority="Tergal" authorityName="Tergal" box="[997,1156,177,201]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="14" pageNumber="1382" phylum="Arthropoda" rank="species" species="cattienensis">
<emphasis id="968AEACD3D735A69FC6836A374AAFF12" box="[997,1156,177,201]" italics="true" pageId="14" pageNumber="1382">H. cattienensis</emphasis>
</taxonomicName>
Tergal spots yellow or gray-yellowı strongly contrasting dark or blackish background. Thoracic shield with 9
<emphasis id="968AEACD3D735A69FE2136E47197FED5" box="[428,441,246,270]" italics="true" pageId="14" pageNumber="1382"></emphasis>
10 transverse striaeı of which 7
<emphasis id="968AEACD3D735A69FC9436E47308FED5" box="[793,806,246,270]" italics="true" pageId="14" pageNumber="1382"></emphasis>
8 cross the dorsum
<taxonomicName id="63FE4D5C3D735A69FB9F36E474ADFED5" ID-CoL="3NF2D" authorityName="Attemsi" authorityYear="1938" box="[1042,1155,246,270]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="14" pageNumber="1382" phylum="Arthropoda" rank="species" species="robusta">
<emphasis id="968AEACD3D735A69FB9F36E474ADFED5" box="[1042,1155,246,270]" italics="true" pageId="14" pageNumber="1382">H. robusta</emphasis>
</taxonomicName>
</keyLead>
</keyStep>
</paragraph>
<paragraph id="A44136DF3D735A69FF2D373974ADFE70" blockId="14.[160,1156,299,427]" pageId="14" pageNumber="1382">
<keyStep id="1F0A2E7A3D735A69FF2D373974ADFE70" pageId="14" pageNumber="1382">
<keyLead id="1F0F95EA3D735A69FF2D373974ADFE70" pageId="14" pageNumber="1382">
- Syncoxital lobe highı subquadrate; syncoxital horns directed mesoventradı shorter than syncoxital lobe ....................................................................................................................
<taxonomicName id="63FE4D5C3D735A69FC7C375C74AAFEBD" ID-CoL="6MN2L" authority="Syncoxital" authorityName="Syncoxital" box="[1009,1156,334,358]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="14" pageNumber="1382" phylum="Arthropoda" rank="species" species="fedorenkoi">
<emphasis id="968AEACD3D735A69FC7C375C74AAFEBD" box="[1009,1156,334,358]" italics="true" pageId="14" pageNumber="1382">H. fedorenkoi</emphasis>
</taxonomicName>
Syncoxital lobe not quadrate/subquadrate; syncoxital horns directed mesoventrad or ventradı longer than syncoxital lobe ................................................................................................ 9
</keyLead>
</keyStep>
</paragraph>
<paragraph id="A44136DF3D735A69FF2D37DB74AAFD92" blockId="14.[160,1156,456,585]" pageId="14" pageNumber="1382">
<keyStep id="1F0A2E7A3D735A69FF2D37DB74AAFD92" pageId="14" pageNumber="1382">
<keyLead id="1F0F95EA3D735A69FF2D37DB74AAFD92" pageId="14" pageNumber="1382">
- Syncoxital lobe concave medially to be a bifid trapeziform. Syncoxital horns directed mesoventrad; tip not truncatedı with an apical seta and a spine
<taxonomicName id="63FE4D5C3D735A69FC1537F9740AFDD8" authority="Nguyen ı Katsuyuki Eguchi &amp; Hwang, 2019" authorityName="Nguyen ı Katsuyuki Eguchi &amp; Hwang" authorityYear="2019" box="[920,1060,491,515]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="14" pageNumber="1382" phylum="Arthropoda" rank="species" species="hoanglien" status="sp. nov.">
<emphasis id="968AEACD3D735A69FC1537F9740AFDD8" box="[920,1060,491,515]" italics="true" pageId="14" pageNumber="1382">H. hoanglien</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DB957B63D735A69FBA137F974AAFDD8" box="[1068,1156,491,515]" pageId="14" pageNumber="1382" rank="species">sp. nov.</taxonomicNameLabel>
Syncoxital lobe roundly trapeziform. Syncoxital horns longı directed ventrad; tip truncatedı without apical spine or seta .............................. ..............................
<taxonomicName id="63FE4D5C3D735A69FC293423740DFD92" authority="Nguyen ı Katsuyuki Eguchi &amp; Hwang, 2019" authorityName="Nguyen ı Katsuyuki Eguchi &amp; Hwang" authorityYear="2019" box="[932,1059,561,585]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="14" pageNumber="1382" phylum="Arthropoda" rank="species" species="fanxipan" status="sp. nov.">
<emphasis id="968AEACD3D735A69FC293423740DFD92" box="[932,1059,561,585]" italics="true" pageId="14" pageNumber="1382">H. fanxipan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DB957B63D735A69FBA6342374AAFD92" box="[1067,1156,561,585]" pageId="14" pageNumber="1382" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
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</paragraph>
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