treatments-xml/data/F1/77/8B/F1778B6A5B41FFE963BD03571B66FC32.xml

<document id="F47EC89F3E661B95A85DF6ADEABF8626" ID="10.11646/zootaxa.4243.3.4SLASH10514" ID-CLB-Dataset="36265" ID-DOI="10.11646/zootaxa.4243.3.4" ID-GBIF-Dataset="b4fc5627-7a2f-4c0b-844a-32d26ec39c80" ID-ISSN="1175-5326" ID-Zenodo-Dep="400153" ID-ZooBank="E3A1C72B-EB8F-4EE0-9BF7-99757331E193" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1489655128551" checkinUser="plazi" docAuthor="Santos, Erika, Da Rocha, Clélia M. C., Jr, Edivaldo Gomes &amp; Fontoura, Paulo" docDate="2017" docId="F1778B6A5B41FFE963BD03571B66FC32" docLanguage="en" docName="zootaxa.4243.3.4SLASH10514.pdf" docOrigin="Zootaxa 4243 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Batillipes brasiliensis Santos, Rocha, Jr &amp; Fontoura, 2017, sp. nov." docType="treatment" docVersion="9" lastPageNumber="491" masterDocId="0D4EF3125B43FFE1632A016E1A0BFFDE" masterDocTitle="Three new Batillipes species (Arthrotardigrada: Batillipedidae) from the Brazilian coast" masterLastPageNumber="502" masterPageNumber="483" pageNumber="485" updateTime="1698438041566" updateUser="ExternalLinkService">
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<mods:title id="204B2EF56F48A82CDD67F4175E545CE5">Three new Batillipes species (Arthrotardigrada: Batillipedidae) from the Brazilian coast</mods:title>
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<mods:namePart id="0582C9E85918F18E341DDFFF3DD59B97">Santos, Erika</mods:namePart>
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<mods:namePart id="90A9BD49E2BF1311038EC78E9E37B223">Da Rocha, Clélia M. C.</mods:namePart>
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<mods:namePart id="12E9BCF9AC5F564EF6D01FCEF1119BD6">Jr, Edivaldo Gomes</mods:namePart>
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<mods:namePart id="5B2A34C0D296DE1E2E4F4B02A4694CE0">Fontoura, Paulo</mods:namePart>
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<heading id="22298D105B41FFE363BD03571BF6FD8D" bold="true" box="[151,509,569,595]" fontSize="11" level="1" pageId="2" pageNumber="485" reason="1">
<emphasis id="4BAAE66E5B41FFE363BD03571BF6FD8D" bold="true" box="[151,509,569,595]" pageId="2" pageNumber="485">
<taxonomicName id="BEDE41FF5B41FFE363BD03571B92FD8D" ID-CoL="7b5f7909-db8d-4b31-867e-5275e1f69064" box="[151,409,569,595]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="2" pageNumber="485" phylum="Tardigrada" rank="species" species="brasiliensis" status="sp. nov.">
<emphasis id="4BAAE66E5B41FFE363BD03571B92FD8D" bold="true" box="[151,409,569,595]" italics="true" pageId="2" pageNumber="485">Batillipes brasiliensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="50995B155B41FFE3628A03571BF6FD8D" box="[416,509,569,595]" pageId="2" pageNumber="485" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
</paragraph>
<paragraph id="79613A7C5B41FFE363BD03321B67FDAA" blockId="2.[151,509,569,629]" box="[151,364,604,629]" pageId="2" pageNumber="485">
(
<figureCitation id="E1E526F95B41FFE363B503321B0FFDAA" box="[159,260,604,629]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="3.[151,250,1850,1872]" captionStartId-1="4.[151,250,1691,1713]" captionTargetBox-0="[153,1436,195,1828]" captionTargetBox-1="[153,1432,195,1655]" captionTargetId-0="figure@3.[151,1436,193,1828]" captionTargetId-1="figure@4.[151,1436,193,1669]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 2. Batillipes brasiliensis sp. nov. A. Drawing (ventral view); B. Habitus (ventral view); C. Detail of the head showing cephalic appendages (cm—median cirrus; ci—internal cirrus; cl—lateral cirrus); D. Anterior portion of the body showing clavae (pc—primary clava; sc—secondary clava), first body projection (pr 1) and leg projection of the first leg (lpI). Scale bars: A – B = 50 µm; C – D = 10 µm." captionText-1="FIGURE 3. Batillipes brasiliensis sp. nov. A. Detail of the two first legs of a paratype showing first body projections (pr 1; pr 2), leg projections (lpI; lpII) and first leg spine (pI); B. Posterior portion of the body of a paratype showing the body projection between legs III and IV (pr 4) and sensorial organ of leg IV (pIV); C. Caudal ala (ca) and body projection between legs III and IV (pr 4) of the holotype; D. Toes (t 1 – t 6) of the leg IV of the holotype; E—Detail of the female gonopore (go) and anus (an) of the holotype; F. Detail of the gonopore (go) and anus (an) of the allotypic male. Scale bars = 10 µm." httpUri-0="https://zenodo.org/record/400155/files/figure.png" httpUri-1="https://zenodo.org/record/400156/files/figure.png" pageId="2" pageNumber="485">Figs 2–3</figureCitation>
; Table 1)
</paragraph>
</subSubSection>
<subSubSection id="31C469F75B41FFE363BD03CA1987FC66" pageId="2" pageNumber="485" type="diagnosis">
<paragraph id="79613A7C5B41FFE363BD03CA1987FC66" blockId="2.[151,1437,676,2033]" pageId="2" pageNumber="485">
<emphasis id="4BAAE66E5B41FFE363BD03CA1B01FD63" bold="true" box="[151,266,676,701]" pageId="2" pageNumber="485">Diagnosis</emphasis>
. Small sized 
<taxonomicName id="BEDE41FF5B41FFE3628C03CA1819FD62" box="[422,530,676,700]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="2" pageNumber="485" phylum="Tardigrada" rank="genus">
<emphasis id="4BAAE66E5B41FFE3628C03CA1819FD62" box="[422,530,676,700]" italics="true" pageId="2" pageNumber="485">Batillipes</emphasis>
</taxonomicName>
with tubular undivided primary clavae and papillar secondary clavae. Cephalic cirri with swollen distal tips. Sensorial spines on all legs. Toes 3 and 4 of legs IV with different lengths. Distinct head separated from the body by a neck constriction followed by well-developed lateral processes (“auricles”). Small lateral processes between legs I–III each with a small digit-shaped apex. Processes between legs III and IV fringed with digit-shaped expansions. The caudal apparatus consists of a wide ala-like cuticular expansion covered with detritus. A small semicircular cuticular projection is also present on the coxal region of each leg. Punctate cuticle comprised of small pillars. Rosette-like female gonopore surrounded by a four-valved cuticular structure. Males with circular gonopore with a cuticular crescent-shaped fold.
</paragraph>
</subSubSection>
<subSubSection id="31C469F75B41FFE363ED02AA1B8DFAB6" pageId="2" pageNumber="485" type="materials_examined">
<paragraph id="79613A7C5B41FFE363ED02AA19E1FC03" blockId="2.[151,1437,676,2033]" box="[199,1002,964,989]" pageId="2" pageNumber="485">
<materialsCitation id="C9B630215B41FFE363ED02AA19E1FC03" ID-GBIF-Occurrence="1440156676" box="[199,1002,964,989]" country="Brazil" latitude="-9.8625" location="Gunga Beach" longLatPrecision="21" longitude="-35.904724" pageId="2" pageNumber="485" specimenCount="1" stateProvince="Alagoas" typeStatus="holotype">
<emphasis id="4BAAE66E5B41FFE363ED02AA1B6EFC03" bold="true" box="[199,357,964,989]" pageId="2" pageNumber="485">
<typeStatus id="A66584DE5B41FFE363ED02AA1B09FC03" box="[199,258,964,989]" pageId="2" pageNumber="485">Type</typeStatus>
locality.
</emphasis>
<location id="7C016CA75B41FFE3624602AA180DFC03" LSID="urn:lsid:plazi:treatment:F1778B6A5B41FFE963BD03571B66FC32:7C016CA75B41FFE3624602AA180DFC03" box="[364,518,964,989]" country="Brazil" latitude="-9.8625" longLatPrecision="21" longitude="-35.904724" name="Gunga Beach" pageId="2" pageNumber="485" stateProvince="Alagoas">Gunga Beach</location>
, 
<collectingRegion id="BB1AF49E5B41FFE3613802AA187BFC03" box="[530,624,964,989]" country="Brazil" name="Alagoas" pageId="2" pageNumber="485">Alagoas</collectingRegion>
, 
<collectingCountry id="01C97AEC5B41FFE3615102AA18B4FC03" box="[635,703,964,989]" name="Brazil" pageId="2" pageNumber="485">Brazil</collectingCountry>
(
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, 
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).
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</paragraph>
<paragraph id="79613A7C5B41FFE363ED02891F32FB06" blockId="2.[151,1437,676,2033]" pageId="2" pageNumber="485">
<emphasis id="4BAAE66E5B41FFE363ED02891B71FBDE" bold="true" box="[199,378,999,1024]" pageId="2" pageNumber="485">Type material.</emphasis>
Holotype: adult, female (slide CVII-80) collected at Gunga Beach, mounted in glycerol. Allotype: adult, male (slide TARD/UFRPE 02-11) collected at Amor Beach, mounted in glycerol. Paratypes: 8 females collected at Sossego Beach, mounted in polyvinyl alcohol (slides TARD/UFRPE 
<date id="0D601CBC5B41FFE367BE055E1F35FB96" box="[1172,1342,1071,1096]" pageId="2" pageNumber="485" value="2006-02-03" valueMax="2006-02-02" valueMin="2006-02-03">02-03 to 02-06</date>
and 02- 14); 2 females and one four-toed larva collected at Forte Orange Beach, mounted in polyvinyl alcohol (slides TARD/UFRPE 02-14); 5 specimens collected at Gunga Beach (3 females, 1 male and 1 juvenile), mounted in glycerol (slides TARD/UFRPE 02-08, 02-09 and CVII-81); and 14 specimens collected at Amor Beach (4 females, 4 males and 6 juveniles), mounted in glycerol (slides TARD/UFRPE 
<date id="0D601CBC5B41FFE360B405AE1E4CFB06" box="[926,1095,1216,1240]" pageId="2" pageNumber="485" value="2013-02-10" valueMax="2013-02-02" valueMin="2013-02-10">02-10 to 02-13</date>
, 02-24 and CVII-82).
</paragraph>
<paragraph id="79613A7C5B41FFE363ED058A1B8DFAB6" blockId="2.[151,1437,676,2033]" pageId="2" pageNumber="485">
<materialsCitation id="C9B630215B41FFE363ED058A1B89FAB6" ID-GBIF-Occurrence="1440156678" country="Portugal" county="Department" location="Faculty" municipality="Biology" pageId="2" pageNumber="485" specimenCount="2" stateProvince="Pernambuco" typeStatus="holotype">
<emphasis id="4BAAE66E5B41FFE363ED058A1B80FB23" bold="true" box="[199,395,1252,1277]" pageId="2" pageNumber="485">
<typeStatus id="A66584DE5B41FFE363ED058A1B09FB23" box="[199,258,1252,1277]" pageId="2" pageNumber="485">Type</typeStatus>
repository.
</emphasis>
The 
<typeStatus id="A66584DE5B41FFE362E3058B1BF2FB23" box="[457,505,1253,1277]" pageId="2" pageNumber="485">type</typeStatus>
material (slides CVII-80–82) is deposited in the collection of the 
<collectingCounty id="900042F05B41FFE367DD058B1F70FB22" box="[1271,1403,1253,1276]" pageId="2" pageNumber="485">Department</collectingCounty>
of 
<collectingMunicipality id="9905A0065B41FFE363BD04691AF9FAFE" box="[151,242,1287,1312]" pageId="2" pageNumber="485">Biology</collectingMunicipality>
, 
<location id="7C016CA75B41FFE3622C04691B51FAFE" LSID="urn:lsid:plazi:treatment:F1778B6A5B41FFE963BD03571B66FC32:7C016CA75B41FFE3622C04691B51FAFE" box="[262,346,1287,1312]" country="Portugal" county="Department" municipality="Biology" name="Faculty" pageId="2" pageNumber="485" stateProvince="Pernambuco">Faculty</location>
of 
<location id="7C016CA75B41FFE362BB04691BFDFAFE" LSID="urn:lsid:plazi:treatment:F1778B6A5B41FFE963BD03571B66FC32:7C016CA75B41FFE362BB04691BFDFAFE" box="[401,502,1287,1312]" country="Portugal" county="Department" municipality="Biology" name="Sciences" pageId="2" pageNumber="485" stateProvince="Pernambuco">Sciences</location>
, 
<location id="7C016CA75B41FFE361200469188BFAFE" LSID="urn:lsid:plazi:treatment:F1778B6A5B41FFE963BD03571B66FC32:7C016CA75B41FFE361200469188BFAFE" box="[522,640,1287,1312]" country="Portugal" county="Department" municipality="Biology" name="University" pageId="2" pageNumber="485" stateProvince="Pernambuco">University</location>
of 
<location id="7C016CA75B41FFE3619D046618FCFAC1" LSID="urn:lsid:plazi:treatment:F1778B6A5B41FFE963BD03571B66FC32:7C016CA75B41FFE3619D046618FCFAC1" box="[695,759,1288,1311]" country="Portugal" county="Department" municipality="Biology" name="Porto" pageId="2" pageNumber="485" stateProvince="Pernambuco">Porto</location>
, 
<collectingCountry id="01C97AEC5B41FFE3602004691967FAFE" box="[778,876,1287,1312]" name="Portugal" pageId="2" pageNumber="485">Portugal</collectingCountry>
, and all the other slides in the collection of Tardigrades—UFRPE (
<location id="7C016CA75B41FFE3628D0442182EFA9B" LSID="urn:lsid:plazi:treatment:F1778B6A5B41FFE963BD03571B66FC32:7C016CA75B41FFE3628D0442182EFA9B" box="[423,549,1324,1349]" country="Portugal" county="Department" municipality="Biology" name="Laboratory" pageId="2" pageNumber="485" stateProvince="Pernambuco">Laboratory</location>
of 
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, 
<location id="7C016CA75B41FFE361D90443197CFA9A" LSID="urn:lsid:plazi:treatment:F1778B6A5B41FFE963BD03571B66FC32:7C016CA75B41FFE361D90443197CFA9A" box="[755,887,1325,1348]" country="Portugal" county="Department" municipality="Biology" name="Department" pageId="2" pageNumber="485" stateProvince="Pernambuco">Department</location>
of 
<location id="7C016CA75B41FFE3609904421E05FA9B" LSID="urn:lsid:plazi:treatment:F1778B6A5B41FFE963BD03571B66FC32:7C016CA75B41FFE3609904421E05FA9B" box="[947,1038,1324,1349]" country="Portugal" county="Department" municipality="Biology" name="Biology" pageId="2" pageNumber="485" stateProvince="Pernambuco">Biology</location>
, Universidade Federal Rural de 
<collectingRegion id="BB1AF49E5B41FFE363BD04211B2DFAB6" box="[151,294,1359,1384]" country="Brazil" name="Pernambuco" pageId="2" pageNumber="485">Pernambuco</collectingRegion>
, 
<collectingCountry id="01C97AEC5B41FFE3621B04211B71FAB6" box="[305,378,1359,1384]" name="Brazil" pageId="2" pageNumber="485">Brazil</collectingCountry>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="31C469F75B41FFE363ED041A1887FA26" pageId="2" pageNumber="485" type="etymology">
<paragraph id="79613A7C5B41FFE363ED041A19C0FA53" blockId="2.[151,1437,676,2033]" box="[199,971,1396,1421]" pageId="2" pageNumber="485">
<emphasis id="4BAAE66E5B41FFE363ED041A1B45FA53" bold="true" box="[199,334,1396,1421]" pageId="2" pageNumber="485">Etymology.</emphasis>
The name 
<taxonomicName id="BEDE41FF5B41FFE362E1041A1847FA52" box="[459,588,1396,1420]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="2" pageNumber="485" phylum="Tardigrada" rank="species" species="brasiliensis">
<emphasis id="4BAAE66E5B41FFE362E1041A1847FA52" box="[459,588,1396,1420]" italics="true" pageId="2" pageNumber="485">brasiliensis</emphasis>
</taxonomicName>
refers to the 
<emphasis id="4BAAE66E5B41FFE361F5041A197BFA52" box="[735,880,1396,1420]" italics="true" pageId="2" pageNumber="485">locus typicus</emphasis>
, 
<collectingCountry id="01C97AEC5B41FFE36055041A19CCFA53" box="[895,967,1396,1421]" name="Brazil" pageId="2" pageNumber="485">Brazil</collectingCountry>
.
</paragraph>
<paragraph id="79613A7C5B41FFE363ED04F91887FA26" blockId="2.[151,1437,676,2033]" pageId="2" pageNumber="485">
<emphasis id="4BAAE66E5B41FFE363ED04F91B88FA6E" bold="true" box="[199,387,1431,1456]" pageId="2" pageNumber="485">Ecological note.</emphasis>
<taxonomicName id="BEDE41FF5B41FFE362A304F71822FA6E" box="[393,553,1432,1456]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="2" pageNumber="485" phylum="Tardigrada" rank="species" species="brasiliensis" status="sp. nov.">
<emphasis id="4BAAE66E5B41FFE362A304F71822FA6E" box="[393,553,1432,1456]" italics="true" pageId="2" pageNumber="485">B. brasiliensis</emphasis>
</taxonomicName>
<emphasis id="4BAAE66E5B41FFE3611A04F91882FA6E" bold="true" box="[560,649,1431,1456]" pageId="2" pageNumber="485">
<taxonomicNameLabel id="50995B155B41FFE3611A04F91882FA6E" box="[560,649,1431,1456]" pageId="2" pageNumber="485" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was found in shallow sublittoral medium to coarse, gravels and quartz sands, in low energy (Sossego Beach) and estuarine beaches (Forte Orange Beach and Gunga Beach) and in reef pools of high energy beaches (Amor Beach).
</paragraph>
</subSubSection>
<subSubSection id="31C469F75B41FFE463ED076A1F2BFBCE" lastPageId="5" lastPageNumber="488" pageId="2" pageNumber="485" type="description">
<paragraph id="79613A7C5B41FFE363ED076A1F3DF876" blockId="2.[151,1437,676,2033]" pageId="2" pageNumber="485">
<emphasis id="4BAAE66E5B41FFE363ED076A181BF9C3" bold="true" box="[199,528,1540,1565]" pageId="2" pageNumber="485">
Description of the 
<typeStatus id="A66584DE5B41FFE36289076A1807F9C3" box="[419,524,1540,1565]" pageId="2" pageNumber="485" type="holotype">holotype</typeStatus>
.
</emphasis>
Female with a body length 124 µm (142 µm including the caudal apparatus) and 45.4 µm wide between the third and fourth pair of legs (
<figureCitation id="E1E526F95B41FFE3606E07491980F99E" box="[836,907,1575,1600]" captionStart="FIGURE 2" captionStartId="3.[151,250,1850,1872]" captionTargetBox="[153,1436,195,1828]" captionTargetId="figure@3.[151,1436,193,1828]" captionTargetPageId="3" captionText="FIGURE 2. Batillipes brasiliensis sp. nov. A. Drawing (ventral view); B. Habitus (ventral view); C. Detail of the head showing cephalic appendages (cm—median cirrus; ci—internal cirrus; cl—lateral cirrus); D. Anterior portion of the body showing clavae (pc—primary clava; sc—secondary clava), first body projection (pr 1) and leg projection of the first leg (lpI). Scale bars: A – B = 50 µm; C – D = 10 µm." httpUri="https://zenodo.org/record/400155/files/figure.png" pageId="2" pageNumber="485">Fig. 2</figureCitation>
A, B). Trapezoid head with eleven cephalic appendages: Internal cirri inserted dorsally on the frontal margin of the head are 18.6 µm long, bearing cirrophores (about 2.2 µm long). External cirri 12.7 µm long, with indistinct cirrophores, inserted more ventrally, near the lateral cirri and primary clavae. The median cirrus, with cirrophore, is 15.5 µm long (
<figureCitation id="E1E526F95B41FFE3677407FA1EAAF972" box="[1118,1185,1684,1709]" captionStart="FIGURE 2" captionStartId="3.[151,250,1850,1872]" captionTargetBox="[153,1436,195,1828]" captionTargetId="figure@3.[151,1436,193,1828]" captionTargetPageId="3" captionText="FIGURE 2. Batillipes brasiliensis sp. nov. A. Drawing (ventral view); B. Habitus (ventral view); C. Detail of the head showing cephalic appendages (cm—median cirrus; ci—internal cirrus; cl—lateral cirrus); D. Anterior portion of the body showing clavae (pc—primary clava; sc—secondary clava), first body projection (pr 1) and leg projection of the first leg (lpI). Scale bars: A – B = 50 µm; C – D = 10 µm." httpUri="https://zenodo.org/record/400155/files/figure.png" pageId="2" pageNumber="485">Fig. 2</figureCitation>
C). Lateral cirri (26.6 µm long) are located dorsally in relation to the unconstricted tubular primary clavae (12.9 µm long). These two appendages share a common pedestal. A van der Land’s organ is present at the base of the primary clava that exhibits a terminal hole (
<figureCitation id="E1E526F95B41FFE3629907911BFCF8C6" box="[435,503,1791,1816]" captionStart="FIGURE 2" captionStartId="3.[151,250,1850,1872]" captionTargetBox="[153,1436,195,1828]" captionTargetId="figure@3.[151,1436,193,1828]" captionTargetPageId="3" captionText="FIGURE 2. Batillipes brasiliensis sp. nov. A. Drawing (ventral view); B. Habitus (ventral view); C. Detail of the head showing cephalic appendages (cm—median cirrus; ci—internal cirrus; cl—lateral cirrus); D. Anterior portion of the body showing clavae (pc—primary clava; sc—secondary clava), first body projection (pr 1) and leg projection of the first leg (lpI). Scale bars: A – B = 50 µm; C – D = 10 µm." httpUri="https://zenodo.org/record/400155/files/figure.png" pageId="2" pageNumber="485">Fig. 2</figureCitation>
D). An indentation is present in the frontal edge of the head between the external cephalic cirrus and the pedestal bearing the primary clava and lateral cirrus. In the frontal edge of the head papillar secondary clavae (
<figureCitation id="E1E526F95B41FFE3624106291BBBF8BE" box="[363,432,1863,1888]" captionStart="FIGURE 2" captionStartId="3.[151,250,1850,1872]" captionTargetBox="[153,1436,195,1828]" captionTargetId="figure@3.[151,1436,193,1828]" captionTargetPageId="3" captionText="FIGURE 2. Batillipes brasiliensis sp. nov. A. Drawing (ventral view); B. Habitus (ventral view); C. Detail of the head showing cephalic appendages (cm—median cirrus; ci—internal cirrus; cl—lateral cirrus); D. Anterior portion of the body showing clavae (pc—primary clava; sc—secondary clava), first body projection (pr 1) and leg projection of the first leg (lpI). Scale bars: A – B = 50 µm; C – D = 10 µm." httpUri="https://zenodo.org/record/400155/files/figure.png" pageId="2" pageNumber="485">Fig. 2</figureCitation>
D) are also well visible (major diameter 2.9 µm). All cephalic cirri, including external cirri, and lateral cirri have swollen tips. Eye spots not recognized. Ovoid pharyngeal bulb 19.7 µm long and 17.7 µm wide. Ventral mouth opening in a protruded triangular cone. Placoids not visible after slide mounting.
</paragraph>
<paragraph id="79613A7C5B41FFE463ED06DB1F17FE9E" blockId="2.[151,1437,676,2033]" lastBlockId="5.[151,1437,151,2013]" lastPageId="5" lastPageNumber="488" pageId="2" pageNumber="485">
Four pairs of ventro-lateral body processes are present: a blunt lateral expansion, 4.1 µm long, between the head and the first pair of legs (
<figureCitation id="E1E526F95B41FFE362C706B61824F82E" box="[493,559,2008,2033]" captionStart="FIGURE 2" captionStartId="3.[151,250,1850,1872]" captionTargetBox="[153,1436,195,1828]" captionTargetId="figure@3.[151,1436,193,1828]" captionTargetPageId="3" captionText="FIGURE 2. Batillipes brasiliensis sp. nov. A. Drawing (ventral view); B. Habitus (ventral view); C. Detail of the head showing cephalic appendages (cm—median cirrus; ci—internal cirrus; cl—lateral cirrus); D. Anterior portion of the body showing clavae (pc—primary clava; sc—secondary clava), first body projection (pr 1) and leg projection of the first leg (lpI). Scale bars: A – B = 50 µm; C – D = 10 µm." httpUri="https://zenodo.org/record/400155/files/figure.png" pageId="2" pageNumber="485">Fig. 2</figureCitation>
D). Lateral processes are also present between all leg pairs. Between the first two pair of legs these blunt processes (6.0, 4.6 µm between legs I–II and II–III respectively) exhibit a thin digitshaped apex (
<figureCitation id="E1E526F95B47FFE5620506191B7AF851" box="[303,369,1911,1936]" captionStart="FIGURE 3" captionStartId="4.[151,250,1691,1713]" captionTargetBox="[153,1432,195,1655]" captionTargetId="figure@4.[151,1436,193,1669]" captionTargetPageId="4" captionText="FIGURE 3. Batillipes brasiliensis sp. nov. A. Detail of the two first legs of a paratype showing first body projections (pr 1; pr 2), leg projections (lpI; lpII) and first leg spine (pI); B. Posterior portion of the body of a paratype showing the body projection between legs III and IV (pr 4) and sensorial organ of leg IV (pIV); C. Caudal ala (ca) and body projection between legs III and IV (pr 4) of the holotype; D. Toes (t 1 – t 6) of the leg IV of the holotype; E—Detail of the female gonopore (go) and anus (an) of the holotype; F. Detail of the gonopore (go) and anus (an) of the allotypic male. Scale bars = 10 µm." httpUri="https://zenodo.org/record/400156/files/figure.png" pageId="4" pageNumber="487">Fig. 3</figureCitation>
A). Between legs III and IV (
<figureCitation id="E1E526F95B47FFE5619706191908F851" box="[701,771,1911,1936]" captionStart="FIGURE 3" captionStartId="4.[151,250,1691,1713]" captionTargetBox="[153,1432,195,1655]" captionTargetId="figure@4.[151,1436,193,1669]" captionTargetPageId="4" captionText="FIGURE 3. Batillipes brasiliensis sp. nov. A. Detail of the two first legs of a paratype showing first body projections (pr 1; pr 2), leg projections (lpI; lpII) and first leg spine (pI); B. Posterior portion of the body of a paratype showing the body projection between legs III and IV (pr 4) and sensorial organ of leg IV (pIV); C. Caudal ala (ca) and body projection between legs III and IV (pr 4) of the holotype; D. Toes (t 1 – t 6) of the leg IV of the holotype; E—Detail of the female gonopore (go) and anus (an) of the holotype; F. Detail of the gonopore (go) and anus (an) of the allotypic male. Scale bars = 10 µm." httpUri="https://zenodo.org/record/400156/files/figure.png" pageId="4" pageNumber="487">Figs 3</figureCitation>
B, C) lateral processes are larger, 5.8 µm long and 10.8 µm wide. In these lateral processes the cuticle is punctated except for the membranous fringed external edge constituted by five digit-shaped expansions (
<figureCitation id="E1E526F95B47FFE561BB06D118DEF809" box="[657,725,1983,2008]" captionStart="FIGURE 2" captionStartId="3.[151,250,1850,1872]" captionTargetBox="[153,1436,195,1828]" captionTargetId="figure@3.[151,1436,193,1828]" captionTargetPageId="3" captionText="FIGURE 2. Batillipes brasiliensis sp. nov. A. Drawing (ventral view); B. Habitus (ventral view); C. Detail of the head showing cephalic appendages (cm—median cirrus; ci—internal cirrus; cl—lateral cirrus); D. Anterior portion of the body showing clavae (pc—primary clava; sc—secondary clava), first body projection (pr 1) and leg projection of the first leg (lpI). Scale bars: A – B = 50 µm; C – D = 10 µm." httpUri="https://zenodo.org/record/400155/files/figure.png" pageId="4" pageNumber="487">Fig. 2</figureCitation>
A, B). The caudal apparatus is an ala-like structure, protruding 17.5 µm and covering almost all the posterior edge of the body. The precise shape of this structure cannot be described because it is covered by detritus (
<figureCitation id="E1E526F95B46FFE4618201D218FEFF0A" box="[680,757,188,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1850,1872]" captionTargetBox="[153,1436,195,1828]" captionTargetId="figure@3.[151,1436,193,1828]" captionTargetPageId="3" captionText="FIGURE 2. Batillipes brasiliensis sp. nov. A. Drawing (ventral view); B. Habitus (ventral view); C. Detail of the head showing cephalic appendages (cm—median cirrus; ci—internal cirrus; cl—lateral cirrus); D. Anterior portion of the body showing clavae (pc—primary clava; sc—secondary clava), first body projection (pr 1) and leg projection of the first leg (lpI). Scale bars: A – B = 50 µm; C – D = 10 µm." httpUri="https://zenodo.org/record/400155/files/figure.png" pageId="5" pageNumber="488">Figs 2</figureCitation>
A, B, 3 C, D). Small semicircular cuticular projections, decreasing in length from the first to the third leg (3.5, 3.3 and 2.1 µm respectively), are also present on the coxal region of legs I–III. This structure is particularly evident in the first leg pair (
<figureCitation id="E1E526F95B46FFE460D4006A1E4DFEC2" box="[1022,1094,260,285]" captionStart="FIGURE 2" captionStartId="3.[151,250,1850,1872]" captionTargetBox="[153,1436,195,1828]" captionTargetId="figure@3.[151,1436,193,1828]" captionTargetPageId="3" captionText="FIGURE 2. Batillipes brasiliensis sp. nov. A. Drawing (ventral view); B. Habitus (ventral view); C. Detail of the head showing cephalic appendages (cm—median cirrus; ci—internal cirrus; cl—lateral cirrus); D. Anterior portion of the body showing clavae (pc—primary clava; sc—secondary clava), first body projection (pr 1) and leg projection of the first leg (lpI). Scale bars: A – B = 50 µm; C – D = 10 µm." httpUri="https://zenodo.org/record/400155/files/figure.png" pageId="5" pageNumber="488">Figs 2</figureCitation>
D, 3 A) and difficult to see in legs III. Lateral processes between legs and cuticular projections on legs often covered by detritus also.
</paragraph>
<caption id="2DA16AF45B40FFE263BD06541832F875" httpUri="https://zenodo.org/record/400155/files/figure.png" pageId="3" pageNumber="486" targetBox="[153,1436,195,1828]" targetPageId="3">
<paragraph id="79613A7C5B40FFE263BD06541832F875" blockId="3.[151,1436,1850,1963]" pageId="3" pageNumber="486">
<emphasis id="4BAAE66E5B40FFE263BD06541B1AF891" bold="true" box="[151,273,1850,1872]" pageId="3" pageNumber="486">FIGURE 2</emphasis>
. 
<taxonomicName id="BEDE41FF5B40FFE2620806541809F88E" box="[290,514,1850,1872]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="3" pageNumber="486" phylum="Tardigrada" rank="species" species="brasiliensis" status="sp. nov.">
<emphasis id="4BAAE66E5B40FFE2620806541809F88E" box="[290,514,1850,1872]" italics="true" pageId="3" pageNumber="486">Batillipes brasiliensis</emphasis>
</taxonomicName>
<emphasis id="4BAAE66E5B40FFE2612706541869F891" bold="true" box="[525,610,1850,1871]" pageId="3" pageNumber="486">
<taxonomicNameLabel id="50995B155B40FFE2612706541869F891" box="[525,610,1850,1871]" pageId="3" pageNumber="486" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A. Drawing (ventral view); B. Habitus (ventral view); C. Detail of the head showing cephalic appendages (cm—median cirrus; ci—internal cirrus; cl—lateral cirrus); D. Anterior portion of the body showing clavae (pc—primary clava; sc—secondary clava), first body projection (pr1) and leg projection of the first leg (lpI). Scale bars: A–B = 50 µm; C–D = 10 µm.
</paragraph>
</caption>
<caption id="2DA16AF45B47FFE563BD07F51F07F8F5" httpUri="https://zenodo.org/record/400156/files/figure.png" pageId="4" pageNumber="487" targetBox="[153,1432,195,1655]" targetPageId="4">
<paragraph id="79613A7C5B47FFE563BD07F51F07F8F5" blockId="4.[151,1436,1691,1835]" pageId="4" pageNumber="487">
<emphasis id="4BAAE66E5B47FFE563BD07F51B05F96E" bold="true" box="[151,270,1691,1713]" pageId="4" pageNumber="487">FIGURE 3</emphasis>
. 
<taxonomicName id="BEDE41FF5B47FFE5623607F51BF0F96F" box="[284,507,1691,1713]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="4" pageNumber="487" phylum="Tardigrada" rank="species" species="brasiliensis" status="sp. nov.">
<emphasis id="4BAAE66E5B47FFE5623607F51BF0F96F" box="[284,507,1691,1713]" italics="true" pageId="4" pageNumber="487">Batillipes brasiliensis</emphasis>
</taxonomicName>
<emphasis id="4BAAE66E5B47FFE5612807F5185EF96E" bold="true" box="[514,597,1691,1712]" pageId="4" pageNumber="487">
<taxonomicNameLabel id="50995B155B47FFE5612807F5185EF96E" box="[514,597,1691,1712]" pageId="4" pageNumber="487" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A. Detail of the two first legs of a paratype showing first body projections (pr1; pr2), leg projections (lpI; lpII) and first leg spine (pI); B. Posterior portion of the body of a paratype showing the body projection between legs III and IV (pr4) and sensorial organ of leg IV (pIV); C. Caudal ala (ca) and body projection between legs III and IV (pr4) of the holotype; D. Toes (t1–t6) of the leg IV of the holotype; E—Detail of the female gonopore (go) and anus (an) of the holotype; F. Detail of the gonopore (go) and anus (an) of the allotypic male. Scale bars = 10 µm.
</paragraph>
</caption>
<paragraph id="79613A7C5B46FFE463ED0022188EFE2A" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="488">
Sensorial spines present on all legs, increasing in length from leg I to leg IV (5.1; 8.2; 9.6, and 12.8 µm long on leg I, II, III and IV respectively). All leg sense organs with swollen tips. The sense organ on leg IV divided into a pronounced cirrophore (5.8 µm long) and a spine-like distal portion (7.0 µm long) with a basal van der Land’s organ (
<figureCitation id="E1E526F95B46FFE463C300D91B25FE0E" box="[233,302,439,464]" captionStart="FIGURE 3" captionStartId="4.[151,250,1691,1713]" captionTargetBox="[153,1432,195,1655]" captionTargetId="figure@4.[151,1436,193,1669]" captionTargetPageId="4" captionText="FIGURE 3. Batillipes brasiliensis sp. nov. A. Detail of the two first legs of a paratype showing first body projections (pr 1; pr 2), leg projections (lpI; lpII) and first leg spine (pI); B. Posterior portion of the body of a paratype showing the body projection between legs III and IV (pr 4) and sensorial organ of leg IV (pIV); C. Caudal ala (ca) and body projection between legs III and IV (pr 4) of the holotype; D. Toes (t 1 – t 6) of the leg IV of the holotype; E—Detail of the female gonopore (go) and anus (an) of the holotype; F. Detail of the gonopore (go) and anus (an) of the allotypic male. Scale bars = 10 µm." httpUri="https://zenodo.org/record/400156/files/figure.png" pageId="5" pageNumber="488">Fig. 3</figureCitation>
B). Sharply pointed and short (13.1 µm long) cirri E with small cirrophores. Cirri E are dorsally placed between lateral processes and leg IV.
</paragraph>
<paragraph id="79613A7C5B46FFE463ED00911819FCE6" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="488">
Telescopic legs with toes of different lengths. On the first three pair of legs (
<figureCitation id="E1E526F95B46FFE4671500911E8FFDC6" box="[1087,1156,511,536]" captionStart="FIGURE 2" captionStartId="3.[151,250,1850,1872]" captionTargetBox="[153,1436,195,1828]" captionTargetId="figure@3.[151,1436,193,1828]" captionTargetPageId="3" captionText="FIGURE 2. Batillipes brasiliensis sp. nov. A. Drawing (ventral view); B. Habitus (ventral view); C. Detail of the head showing cephalic appendages (cm—median cirrus; ci—internal cirrus; cl—lateral cirrus); D. Anterior portion of the body showing clavae (pc—primary clava; sc—secondary clava), first body projection (pr 1) and leg projection of the first leg (lpI). Scale bars: A – B = 50 µm; C – D = 10 µm." httpUri="https://zenodo.org/record/400155/files/figure.png" pageId="5" pageNumber="488">Fig. 2</figureCitation>
B), toe 2 is the shortest (considering toe 1 the most cephalically), toes 3 and 5 are the longest, and toes 1, 3 and 4 are medium sized (in leg II toe 1 is 7.0 µm long, toe 2 is 3.2 µm, toe 3 is 13.0 µm long, toe 4 is 6.2 µm, toe 5 is 14.1 µm, toe 6 is 7.3 µm). On the fourth pair of legs toes (
<figureCitation id="E1E526F95B46FFE462FD03021810FD5A" box="[471,539,620,645]" captionStart="FIGURE 3" captionStartId="4.[151,250,1691,1713]" captionTargetBox="[153,1432,195,1655]" captionTargetId="figure@4.[151,1436,193,1669]" captionTargetPageId="4" captionText="FIGURE 3. Batillipes brasiliensis sp. nov. A. Detail of the two first legs of a paratype showing first body projections (pr 1; pr 2), leg projections (lpI; lpII) and first leg spine (pI); B. Posterior portion of the body of a paratype showing the body projection between legs III and IV (pr 4) and sensorial organ of leg IV (pIV); C. Caudal ala (ca) and body projection between legs III and IV (pr 4) of the holotype; D. Toes (t 1 – t 6) of the leg IV of the holotype; E—Detail of the female gonopore (go) and anus (an) of the holotype; F. Detail of the gonopore (go) and anus (an) of the allotypic male. Scale bars = 10 µm." httpUri="https://zenodo.org/record/400156/files/figure.png" pageId="5" pageNumber="488">Fig. 3</figureCitation>
D) conform the pattern of the D group of species proposed by 
<bibRefCitation id="1D4F478D5B46FFE467D903021B56FD76" author="Kristensen" pageId="5" pageNumber="488" refString="Kristensen, R. M. &amp; Mackness, B. S. (2000) First record of the marine Tardigrade genus Batillipes (Arthrotardigrada: Batillipedidae) from South Australia with a description of a new species. Records of the South Australian Museum, 33 (2), 73 - 87." type="journal article" year="2000">Kristensen and Mackness (2000)</bibRefCitation>
, with toes 3 and 4 of different lengths (6.4 and 10.4 µm long respectively for toes 3 and 4). In those legs toes 1 and 6 are similar in length (respectively 11.3 and 11.0 µm long); toes 2 and especially toes 5 are the longest (16.3 and 20.5 µm long respectively). Toes have the distal stalk enlarged distally (about 2.2 µm wide), ovoid suction discs (3.3 long x 2.8 µm wide measured on toes of leg IV) with conspicuous braces and well developed peripheral thickenings.
</paragraph>
<paragraph id="79613A7C5B46FFE463ED022A1E2AFC7B" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="488">Dorsal cuticle punctated, about 10 pillars/10 µm (each pillar about 1.4 µm high), without evident transverse folds. However, the large amount of detritus covering the dorsal region of the body can hide those lines. Transverse cuticular folds are visible ventrally where the cuticle punctation is more delicate.</paragraph>
<paragraph id="79613A7C5B46FFE463ED02C11F2BFBCE" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="488">
Rosette-shaped gonopore (
<figureCitation id="E1E526F95B46FFE462D202C11836FC16" box="[504,573,943,968]" captionStart="FIGURE 3" captionStartId="4.[151,250,1691,1713]" captionTargetBox="[153,1432,195,1655]" captionTargetId="figure@4.[151,1436,193,1669]" captionTargetPageId="4" captionText="FIGURE 3. Batillipes brasiliensis sp. nov. A. Detail of the two first legs of a paratype showing first body projections (pr 1; pr 2), leg projections (lpI; lpII) and first leg spine (pI); B. Posterior portion of the body of a paratype showing the body projection between legs III and IV (pr 4) and sensorial organ of leg IV (pIV); C. Caudal ala (ca) and body projection between legs III and IV (pr 4) of the holotype; D. Toes (t 1 – t 6) of the leg IV of the holotype; E—Detail of the female gonopore (go) and anus (an) of the holotype; F. Detail of the gonopore (go) and anus (an) of the allotypic male. Scale bars = 10 µm." httpUri="https://zenodo.org/record/400156/files/figure.png" pageId="5" pageNumber="488">Fig. 3</figureCitation>
C) separated from the anus by a weakly defined groove. The gonopore is surrounded by a four-valved cuticular structure. Two larger divergent valves are located posteriorly and two narrow valves are located anteriorly to the gonopore (
<figureCitation id="E1E526F95B46FFE461B3029918D7FBCE" box="[665,732,1015,1040]" captionStart="FIGURE 3" captionStartId="4.[151,250,1691,1713]" captionTargetBox="[153,1432,195,1655]" captionTargetId="figure@4.[151,1436,193,1669]" captionTargetPageId="4" captionText="FIGURE 3. Batillipes brasiliensis sp. nov. A. Detail of the two first legs of a paratype showing first body projections (pr 1; pr 2), leg projections (lpI; lpII) and first leg spine (pI); B. Posterior portion of the body of a paratype showing the body projection between legs III and IV (pr 4) and sensorial organ of leg IV (pIV); C. Caudal ala (ca) and body projection between legs III and IV (pr 4) of the holotype; D. Toes (t 1 – t 6) of the leg IV of the holotype; E—Detail of the female gonopore (go) and anus (an) of the holotype; F. Detail of the gonopore (go) and anus (an) of the allotypic male. Scale bars = 10 µm." httpUri="https://zenodo.org/record/400156/files/figure.png" pageId="5" pageNumber="488">Fig. 3</figureCitation>
E). The anus is 10.9 µm distant from the gonopore.
</paragraph>
</subSubSection>
<subSubSection id="31C469F75B46FFE463ED05721F2AFA43" pageId="5" pageNumber="488" type="discussion">
<paragraph id="79613A7C5B46FFE463ED05721BFEFAEE" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="488">
<emphasis id="4BAAE66E5B46FFE463ED05721B30FBEB" bold="true" box="[199,315,1052,1077]" pageId="5" pageNumber="488">Remarks.</emphasis>
Sexual dimorphism is not evident. Males are similar to females in both qualitative and metric characters (table 1), except for their circular gonopore with a cuticular crescent shaped fold (
<figureCitation id="E1E526F95B46FFE467FD05511F16FB86" box="[1239,1309,1087,1112]" captionStart="FIGURE 3" captionStartId="4.[151,250,1691,1713]" captionTargetBox="[153,1432,195,1655]" captionTargetId="figure@4.[151,1436,193,1669]" captionTargetPageId="4" captionText="FIGURE 3. Batillipes brasiliensis sp. nov. A. Detail of the two first legs of a paratype showing first body projections (pr 1; pr 2), leg projections (lpI; lpII) and first leg spine (pI); B. Posterior portion of the body of a paratype showing the body projection between legs III and IV (pr 4) and sensorial organ of leg IV (pIV); C. Caudal ala (ca) and body projection between legs III and IV (pr 4) of the holotype; D. Toes (t 1 – t 6) of the leg IV of the holotype; E—Detail of the female gonopore (go) and anus (an) of the holotype; F. Detail of the gonopore (go) and anus (an) of the allotypic male. Scale bars = 10 µm." httpUri="https://zenodo.org/record/400156/files/figure.png" pageId="5" pageNumber="488">Fig. 3</figureCitation>
F) located nearer the anus (4.5 µm distant for the anus in the allotypic male). Juveniles, with six toes on each leg but without a visible gonopore, are also similar to adults. The four-toed larva differs from the other juveniles and the adults in having reduced lateral projections without digit-shaped apices; in particular the one located between legs III and IV which lacks the fringed edge. Moreover, the caudal apparatus of this larva is constituted by a conical process without any membranous ala. However, small projections on the coxal region of the three first legs, characteristic of the new species, are present.
</paragraph>
<paragraph id="79613A7C5B46FFE463ED04521F2AFA43" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="488">
On the other hand, among adults, the development of the caudal apparatus (see 
<figureCitation id="E1E526F95B46FFE4671404521E74FA8A" box="[1086,1151,1340,1365]" captionStart="FIGURE 3" captionStartId="4.[151,250,1691,1713]" captionTargetBox="[153,1432,195,1655]" captionTargetId="figure@4.[151,1436,193,1669]" captionTargetPageId="4" captionText="FIGURE 3. Batillipes brasiliensis sp. nov. A. Detail of the two first legs of a paratype showing first body projections (pr 1; pr 2), leg projections (lpI; lpII) and first leg spine (pI); B. Posterior portion of the body of a paratype showing the body projection between legs III and IV (pr 4) and sensorial organ of leg IV (pIV); C. Caudal ala (ca) and body projection between legs III and IV (pr 4) of the holotype; D. Toes (t 1 – t 6) of the leg IV of the holotype; E—Detail of the female gonopore (go) and anus (an) of the holotype; F. Detail of the gonopore (go) and anus (an) of the allotypic male. Scale bars = 10 µm." httpUri="https://zenodo.org/record/400156/files/figure.png" pageId="5" pageNumber="488">Fig. 3</figureCitation>
B, C, F) and the shape of the fourth lateral projection (between legs III and IV), especially in the number (from 2 to 7) of digit-shaped membranous expansions, are very variable (
<figureCitation id="E1E526F95B46FFE461AE04EA18C0FA42" box="[644,715,1412,1437]" captionStart="FIGURE 3" captionStartId="4.[151,250,1691,1713]" captionTargetBox="[153,1432,195,1655]" captionTargetId="figure@4.[151,1436,193,1669]" captionTargetPageId="4" captionText="FIGURE 3. Batillipes brasiliensis sp. nov. A. Detail of the two first legs of a paratype showing first body projections (pr 1; pr 2), leg projections (lpI; lpII) and first leg spine (pI); B. Posterior portion of the body of a paratype showing the body projection between legs III and IV (pr 4) and sensorial organ of leg IV (pIV); C. Caudal ala (ca) and body projection between legs III and IV (pr 4) of the holotype; D. Toes (t 1 – t 6) of the leg IV of the holotype; E—Detail of the female gonopore (go) and anus (an) of the holotype; F. Detail of the gonopore (go) and anus (an) of the allotypic male. Scale bars = 10 µm." httpUri="https://zenodo.org/record/400156/files/figure.png" pageId="5" pageNumber="488">Figs 3</figureCitation>
B, C). Variability in other projections is not relevant.
</paragraph>
</subSubSection>
<subSubSection id="31C469F75B46FFE763ED04C91D2CFD73" lastPageId="6" lastPageNumber="489" pageId="5" pageNumber="488" type="diagnosis">
<paragraph id="79613A7C5B46FFE463ED04C919C0F8DA" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="488">
<emphasis id="4BAAE66E5B46FFE463ED04C91BDAFA1E" bold="true" box="[199,465,1447,1472]" pageId="5" pageNumber="488">Differential diagnosis.</emphasis>
The taxonomy of the genus 
<taxonomicName id="BEDE41FF5B46FFE4600204C61998FA1E" box="[808,915,1448,1472]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="5" pageNumber="488" phylum="Tardigrada" rank="genus">
<emphasis id="4BAAE66E5B46FFE4600204C61998FA1E" box="[808,915,1448,1472]" italics="true" pageId="5" pageNumber="488">Batillipes</emphasis>
</taxonomicName>
is particular problematic due to the reduced number of taxonomic characters traditionally used to distinguish species. Intraspecific variability exhibited by some of those characters and deformations caused by slide mounting are also a source of taxonomic difficulties (
<emphasis id="4BAAE66E5B46FFE466AB049E1F9EF9D6" box="[1409,1429,1520,1544]" italics="true" pageId="5" pageNumber="488">cf</emphasis>
. 
<bibRefCitation id="1D4F478D5B46FFE463BD077A1B34F9F3" author="McKirdy" box="[151,319,1556,1581]" pageId="5" pageNumber="488" refString="McKirdy, D. J. (1975) Batillipes (Heterotardigrada): Comparison of six species from Florida (USA) and a discussion of taxonomic characters within the genus. Memorie dell'Istituto Italiano di Idrobiologia, 32 (Supplement), 177223." type="journal article" year="1975">McKirdy 1975</bibRefCitation>
; 
<bibRefCitation id="1D4F478D5B46FFE46266077A180EF9F2" author="Kristensen" box="[332,517,1556,1581]" pageId="5" pageNumber="488" refString="Kristensen, R. M. (1978) Notes on Marine Heterotardigrades. 1. Description of two Batillipes species, using the electron microscope. Zoologischer Anzeiger, 200, 1 - 17." type="journal article" year="1978">Kristensen 1978</bibRefCitation>
; 
<bibRefCitation id="1D4F478D5B46FFE4613B077B1951F9F2" author="Morone" box="[529,858,1556,1581]" pageId="5" pageNumber="488" refString="Morone De Lucia, R. M., D'Addabo Gallo, M., Grimaldi de Zio, S. (1988) Descrizione di due nuove specie di Batillipedidae (Tardigrada: Heterotardigrada). Cahiers de Biologie Marine, 29, 361 - 373." type="journal article" year="1988">
Morone De Lucia 
<emphasis id="4BAAE66E5B46FFE461CA077B191AF9F2" box="[736,785,1556,1580]" italics="true" pageId="5" pageNumber="488">et al</emphasis>
. 1988
</bibRefCitation>
; Villora Moreno and de 
<bibRefCitation id="1D4F478D5B46FFE46741077A1F49F9F2" author="Zio" box="[1131,1346,1556,1581]" pageId="5" pageNumber="488" refString="Villora-Moreno, S. &amp; de Zio Grimaldi, S. (1993) Redescription and ecology of Batillipes phreaticus Renaud-Debyser, 1959 (Arthrotardigrada, Batillipedidae) in the gulf of Valencia (western Mediterranean). Cahiers de Biologie Marine, 34, 387 - 399." type="journal article" year="1993">Zio Grimaldi 1993</bibRefCitation>
). In this genus, one of the more reliable taxonomic characters is the relative dimension of toes. This indicator was introduced by 
<bibRefCitation id="1D4F478D5B46FFE4621307321BEAF9AA" author="Pollock" box="[313,481,1628,1653]" pageId="5" pageNumber="488" refString="Pollock, L. W. (1970) Batillipes dicrocercus n. sp., Stygarctus granulatus n. sp. and other tardigrade from Woods Hole, Massachussets, U. S. A. Transactions of the American Microscopical Society, 89 (1), 38 - 52." type="journal article" year="1970">Pollock (1970)</bibRefCitation>
and modified by 
<bibRefCitation id="1D4F478D5B46FFE4618207321E12F9AA" author="Kristensen" box="[680,1049,1628,1653]" pageId="5" pageNumber="488" refString="Kristensen, R. M. &amp; Mackness, B. S. (2000) First record of the marine Tardigrade genus Batillipes (Arthrotardigrada: Batillipedidae) from South Australia with a description of a new species. Records of the South Australian Museum, 33 (2), 73 - 87." type="journal article" year="2000">Kristensen and Mackness (2000)</bibRefCitation>
who proposed four patterns of toe lengths on the fourth foot, corresponding to four species groups. 
<taxonomicName id="BEDE41FF5B46FFE4604707EE1E58F946" box="[877,1107,1664,1688]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="5" pageNumber="488" phylum="Tardigrada" rank="species" species="brasiliesis" status="sp. nov.">
<emphasis id="4BAAE66E5B46FFE4604707EE1E58F946" box="[877,1107,1664,1688]" italics="true" pageId="5" pageNumber="488">Batillipes brasiliesis</emphasis>
</taxonomicName>
<emphasis id="4BAAE66E5B46FFE4677007111EBFF946" bold="true" box="[1114,1204,1663,1688]" pageId="5" pageNumber="488">
<taxonomicNameLabel id="50995B155B46FFE4677007111EBFF946" box="[1114,1204,1663,1688]" pageId="5" pageNumber="488" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in having toes 3 and 4 of different lengths and respectively different from toe 1 and toe 2, belongs to the D group of species. Only five species exhibit this toe pattern on leg IV: 
<taxonomicName id="BEDE41FF5B46FFE4615A07A71AD8F8DA" authority="Morone De Lucia, D'Addabbo Gallo and Grimaldi de Zio, 1988" authorityName="Morone De Lucia, D'Addabbo Gallo and Grimaldi de Zio" authorityYear="1988" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="5" pageNumber="488" phylum="Tardigrada" rank="species" species="africanus">
<emphasis id="4BAAE66E5B46FFE4615A07A718F0F93E" box="[624,763,1736,1760]" italics="true" pageId="5" pageNumber="488">B. africanus</emphasis>
Morone De Lucia, D’Addabbo Gallo and Grimaldi de 
<bibRefCitation id="1D4F478D5B46FFE4664407A91AD8F8DA" author="Zio" pageId="5" pageNumber="488" refString="Morone De Lucia, R. M., D'Addabo Gallo, M., Grimaldi de Zio, S. (1988) Descrizione di due nuove specie di Batillipedidae (Tardigrada: Heterotardigrada). Cahiers de Biologie Marine, 29, 361 - 373." type="journal article" year="1988">Zio, 1988</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="BEDE41FF5B46FFE463F707831B48F8DA" box="[221,323,1772,1796]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="5" pageNumber="488" phylum="Tardigrada" rank="species" species="lesteri">
<emphasis id="4BAAE66E5B46FFE463F707831B48F8DA" box="[221,323,1772,1796]" italics="true" pageId="5" pageNumber="488">B. lesteri</emphasis>
</taxonomicName>
; 
<taxonomicName id="BEDE41FF5B46FFE4627B07831BE9F8DA" box="[337,482,1772,1796]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="5" pageNumber="488" phylum="Tardigrada" rank="species" species="tubernatis">
<emphasis id="4BAAE66E5B46FFE4627B07831BE9F8DA" box="[337,482,1772,1796]" italics="true" pageId="5" pageNumber="488">B. tubernatis</emphasis>
</taxonomicName>
; 
<taxonomicName id="BEDE41FF5B46FFE462DB078318F9F8DB" authority="Schulz, 1955" authorityName="Schulz" authorityYear="1955" box="[497,754,1772,1797]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="5" pageNumber="488" phylum="Tardigrada" rank="species" species="similis">
<emphasis id="4BAAE66E5B46FFE462DB07831852F8DA" box="[497,601,1772,1796]" italics="true" pageId="5" pageNumber="488">B. similis</emphasis>
<bibRefCitation id="1D4F478D5B46FFE4614A078218F9F8DB" author="Schulz" box="[608,754,1772,1797]" pageId="5" pageNumber="488" refString="Schulz, E., (1955) Studien an marinen tardigraden. Kieler Meeresforschungen, 11, 74 - 79." type="journal article" year="1955">Schulz, 1955</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="BEDE41FF5B46FFE46003078319CFF8DA" box="[809,964,1772,1796]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="5" pageNumber="488" phylum="Tardigrada" rank="species" species="acuticauda">
<emphasis id="4BAAE66E5B46FFE46003078319CFF8DA" box="[809,964,1772,1796]" italics="true" pageId="5" pageNumber="488">B. acuticauda</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="79613A7C5B46FFE763ED06611E4EFDAC" blockId="5.[151,1437,151,2013]" lastBlockId="6.[119,1895,575,626]" lastPageId="6" lastPageNumber="489" pageId="5" pageNumber="488">
The new species, in having lateral projections, can be clearly distinguished from 
<taxonomicName id="BEDE41FF5B46FFE46787067F1F35F8F6" box="[1197,1342,1808,1832]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="5" pageNumber="488" phylum="Tardigrada" rank="species" species="africanus">
<emphasis id="4BAAE66E5B46FFE46787067F1F35F8F6" box="[1197,1342,1808,1832]" italics="true" pageId="5" pageNumber="488">B. africanus</emphasis>
</taxonomicName>
and 
<emphasis id="4BAAE66E5B46FFE466AE067F1805F892" italics="true" pageId="5" pageNumber="488">
<taxonomicName id="BEDE41FF5B46FFE466AE067F1B07F892" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="5" pageNumber="488" phylum="Tardigrada" rank="species" species="tubernatis">B. tubernatis</taxonomicName>
. 
<taxonomicName id="BEDE41FF5B46FFE46232065A1805F892" box="[280,526,1844,1868]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="5" pageNumber="488" phylum="Tardigrada" rank="species" species="brasiliensis" status="sp. nov.">Batillipes brasiliensis</taxonomicName>
</emphasis>
<emphasis id="4BAAE66E5B46FFE4613D065A1879F893" bold="true" box="[535,626,1844,1869]" pageId="5" pageNumber="488">
<taxonomicNameLabel id="50995B155B46FFE4613D065A1879F893" box="[535,626,1844,1869]" pageId="5" pageNumber="488" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from 
<taxonomicName id="BEDE41FF5B46FFE46026065B197DF892" box="[780,886,1844,1868]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="5" pageNumber="488" phylum="Tardigrada" rank="species" species="similis">
<emphasis id="4BAAE66E5B46FFE46026065B197DF892" box="[780,886,1844,1868]" italics="true" pageId="5" pageNumber="488">B. similis</emphasis>
</taxonomicName>
in the shape of lateral processes and the caudal appendage that, despite variability, are always conical in the last species. In addition, the primary clava of 
<taxonomicName id="BEDE41FF5B46FFE466AE06371AEBF84A" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="5" pageNumber="488" phylum="Tardigrada" rank="species" species="similis">
<emphasis id="4BAAE66E5B46FFE466AE06371AEBF84A" italics="true" pageId="5" pageNumber="488">B. similis</emphasis>
</taxonomicName>
, in having a swollen tip, is drumstick-like and not uniformly tubular as in the new species. On the other hand, the sense organ on leg IV is much longer, almost twice the length, in 
<taxonomicName id="BEDE41FF5B46FFE460CF06CF1E46F866" box="[997,1101,1952,1976]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="5" pageNumber="488" phylum="Tardigrada" rank="species" species="similis">
<emphasis id="4BAAE66E5B46FFE460CF06CF1E46F866" box="[997,1101,1952,1976]" italics="true" pageId="5" pageNumber="488">B. similis</emphasis>
</taxonomicName>
than in the new species; 18% of the body length in 
<taxonomicName id="BEDE41FF5B46FFE4628006AB1819F802" box="[426,530,1988,2012]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="5" pageNumber="488" phylum="Tardigrada" rank="species" species="similis">
<emphasis id="4BAAE66E5B46FFE4628006AB1819F802" box="[426,530,1988,2012]" italics="true" pageId="5" pageNumber="488">B. similis</emphasis>
</taxonomicName>
(mean values, according to Gallo d’Addabbo 
<emphasis id="4BAAE66E5B46FFE4673506AB1E5BF802" box="[1055,1104,1988,2012]" italics="true" pageId="5" pageNumber="488">et al</emphasis>
. 1999) and 10% in the new 
<emphasis id="4BAAE66E5B45FFE7635D03511ADCFD8B" bold="true" box="[119,215,575,597]" pageId="6" pageNumber="489">TABLE]</emphasis>
. Measurements (in µm) of selecteđ morphological structures of specimens of 
<taxonomicName id="BEDE41FF5B45FFE7608303511E7AFD8B" box="[937,1137,575,597]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="6" pageNumber="489" phylum="Tardigrada" rank="species" species="brasiliensis" status="sp. nov.">
<emphasis id="4BAAE66E5B45FFE7608303511E7AFD8B" box="[937,1137,575,597]" italics="true" pageId="6" pageNumber="489">Batillipes brasiliensis</emphasis>
</taxonomicName>
<emphasis id="4BAAE66E5B45FFE7675203511EC8FD8B" bold="true" box="[1144,1219,575,597]" pageId="6" pageNumber="489">
<taxonomicNameLabel id="50995B155B45FFE7675203511EC8FD8B" box="[1144,1219,575,597]" pageId="6" pageNumber="489" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(Holo—Holotype; SD—Stanđarđ đeviation; Range refers to the smallest largest measuređ specimen/structure; N—number of specimens/structures measuređ; Allo—Allotype).
</paragraph>
<paragraph id="79613A7C5B45FFE762F203F91D2CFD73" blockId="6.[119,1831,663,741]" box="[472,1831,663,685]" pageId="6" pageNumber="489">
<emphasis id="4BAAE66E5B45FFE762F203F91D2CFD73" bold="true" box="[472,1831,663,685]" pageId="6" pageNumber="489">FEMALES MALES JUVENILES LARVA</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="31C469F75B45FFE962F203DD1B66FC32" lastPageId="8" lastPageNumber="491" pageId="6" pageNumber="489" type="description">
<paragraph id="79613A7C5B45FFE762F203DD1D18FD17" blockId="6.[119,1831,663,741]" box="[472,1811,691,713]" pageId="6" pageNumber="489">
<emphasis id="4BAAE66E5B45FFE762F203DD1D18FD17" bold="true" box="[472,1811,691,713]" pageId="6" pageNumber="489">adults adults 6-toed 4-toed</emphasis>
</paragraph>
<paragraph id="79613A7C5B45FFE7635D03A11C59FD3B" blockId="6.[119,1831,663,741]" box="[119,1618,719,741]" pageId="6" pageNumber="489">
<emphasis id="4BAAE66E5B45FFE7635D03A11C59FD3B" bold="true" box="[119,1618,719,741]" pageId="6" pageNumber="489">STRUCTURES Holo Mean ± SD (Range); N Allo Mean ± SD (Range); N Mean ± SD (Range); N</emphasis>
</paragraph>
<paragraph id="79613A7C5B45FFE7635D03941CEBF9FD" pageId="6" pageNumber="489">
<table id="0BDEC8DC5B45001E635D03941D6DF9FD" box="[119,1894,762,1571]" gridcols="7" gridrows="20" pageId="6" pageNumber="489">
<tr id="C7EE383E5B45001E635D03941D6DFCCE" box="[119,1894,762,784]" gridrow="0" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D03941B5EFCCE" box="[119,341,762,784]" gridcol="0" gridrow="0" pageId="6" pageNumber="489">Bođy length</th>
<th id="843F51425B45001E62F20394180BFCCE" box="[472,512,762,784]" gridcol="1" gridrow="0" pageId="6" pageNumber="489">142</th>
<th id="843F51425B45001E61610394194AFCCE" box="[587,833,762,784]" gridcol="2" gridrow="0" pageId="6" pageNumber="489">130.3 ±19.3 (91­156); 16</th>
<th id="843F51425B45001E6086039419DFFCCE" box="[940,980,762,784]" gridcol="3" gridrow="0" pageId="6" pageNumber="489">159</th>
<th id="843F51425B45001E673503941F01FCCE" box="[1055,1290,762,784]" gridcol="4" gridrow="0" pageId="6" pageNumber="489">148.3 ±8.8 (142­159); 5</th>
<th id="843F51425B45001E665F03941C6BFCCE" box="[1397,1632,762,784]" gridcol="5" gridrow="0" pageId="6" pageNumber="489">140.1 ±7.8 (130­150); 5</th>
<th id="843F51425B45001E65FC03941D6DFCCE" box="[1750,1894,762,784]" gridcol="6" gridrow="0" pageId="6" pageNumber="489">92</th>
</tr>
<tr id="C7EE383E5B45001E635D024D1D6DFCE7" box="[119,1894,803,825]" gridrow="1" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D024D1B5EFCE7" box="[119,341,803,825]" gridcol="0" gridrow="1" pageId="6" pageNumber="489">Bođy without cauđal ala</th>
<td id="843F51425B45001E62F2024D180BFCE7" box="[472,512,803,825]" gridcol="1" gridrow="1" pageId="6" pageNumber="489">124</td>
<td id="843F51425B45001E6161024D194AFCE7" box="[587,833,803,825]" gridcol="2" gridrow="1" pageId="6" pageNumber="489">121.7 ±17.0 (87­151); 16</td>
<td id="843F51425B45001E6086024D19DFFCE7" box="[940,980,803,825]" gridcol="3" gridrow="1" pageId="6" pageNumber="489">143</td>
<td id="843F51425B45001E6735024D1F01FCE7" box="[1055,1290,803,825]" gridcol="4" gridrow="1" pageId="6" pageNumber="489">134.0 ±7.0 (127­144); 5</td>
<td id="843F51425B45001E665F024D1C6BFCE7" box="[1397,1632,803,825]" gridcol="5" gridrow="1" pageId="6" pageNumber="489">128.2 ±9.0 (116­137); 5</td>
<td id="843F51425B45001E65FC024D1D6DFCE7" box="[1750,1894,803,825]" gridcol="6" gridrow="1" pageId="6" pageNumber="489">87</td>
</tr>
<tr id="C7EE383E5B45001E635D02231D6DFCBD" box="[119,1894,845,867]" gridrow="2" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D02231B5EFCBD" box="[119,341,845,867]" gridcol="0" gridrow="2" pageId="6" pageNumber="489">Bođy wiđth</th>
<td id="843F51425B45001E62F20223180BFCBD" box="[472,512,845,867]" gridcol="1" gridrow="2" pageId="6" pageNumber="489">45.4</td>
<td id="843F51425B45001E61610223194AFCBD" box="[587,833,845,867]" gridcol="2" gridrow="2" pageId="6" pageNumber="489">46.9 ±8.7 (30.5­62.5); 17</td>
<td id="843F51425B45001E6086022319DFFCBD" box="[940,980,845,867]" gridcol="3" gridrow="2" pageId="6" pageNumber="489">40.0</td>
<td id="843F51425B45001E673502231F01FCBD" box="[1055,1290,845,867]" gridcol="4" gridrow="2" pageId="6" pageNumber="489">43.0 ±3.70 (37.6­46.3); 4</td>
<td id="843F51425B45001E665F02231C6BFCBD" box="[1397,1632,845,867]" gridcol="5" gridrow="2" pageId="6" pageNumber="489">48.1 ±7.78 (38.8­57.7); 6</td>
<td id="843F51425B45001E65FC02231D6DFCBD" box="[1750,1894,845,867]" gridcol="6" gridrow="2" pageId="6" pageNumber="489">25.9</td>
</tr>
<tr id="C7EE383E5B45001E635D02181D6DFC52" box="[119,1894,886,908]" gridrow="3" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D02181B5EFC52" box="[119,341,886,908]" gridcol="0" gridrow="3" pageId="6" pageNumber="489">Međian cirrus</th>
<td id="843F51425B45001E62F20218180BFC52" box="[472,512,886,908]" gridcol="1" gridrow="3" pageId="6" pageNumber="489">15.5</td>
<td id="843F51425B45001E61610218194AFC52" box="[587,833,886,908]" gridcol="2" gridrow="3" pageId="6" pageNumber="489">17.4 ±6.0 (11.2­28.8); 7</td>
<td id="843F51425B45001E6086021819DFFC52" box="[940,980,886,908]" gridcol="3" gridrow="3" pageId="6" pageNumber="489">14.0</td>
<td id="843F51425B45001E673502181F01FC52" box="[1055,1290,886,908]" gridcol="4" gridrow="3" pageId="6" pageNumber="489">- (12.4); 1</td>
<td id="843F51425B45001E665F02181C6BFC52" box="[1397,1632,886,908]" gridcol="5" gridrow="3" pageId="6" pageNumber="489">15.2 ±5.25 (10.4­20.8); 3</td>
<td id="843F51425B45001E65FC02181D6DFC52" box="[1750,1894,886,908]" gridcol="6" gridrow="3" pageId="6" pageNumber="489">13.0</td>
</tr>
<tr id="C7EE383E5B45001E635D02CE1D6DFC68" box="[119,1894,928,950]" gridrow="4" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D02CE1B5EFC68" box="[119,341,928,950]" gridcol="0" gridrow="4" pageId="6" pageNumber="489">Internal cirri</th>
<td id="843F51425B45001E62F202CE180BFC68" box="[472,512,928,950]" gridcol="1" gridrow="4" pageId="6" pageNumber="489">18.6</td>
<td id="843F51425B45001E616102CE194AFC68" box="[587,833,928,950]" gridcol="2" gridrow="4" pageId="6" pageNumber="489">18.0 ±2.19 (14.9­22.7); 15</td>
<td id="843F51425B45001E608602CE19DFFC68" box="[940,980,928,950]" gridcol="3" gridrow="4" pageId="6" pageNumber="489">18.5</td>
<td id="843F51425B45001E673502CE1F01FC68" box="[1055,1290,928,950]" gridcol="4" gridrow="4" pageId="6" pageNumber="489">18.1 ±3.61 (13.2­21.0); 5</td>
<td id="843F51425B45001E665F02CE1C6BFC68" box="[1397,1632,928,950]" gridcol="5" gridrow="4" pageId="6" pageNumber="489">15.3 ±3.27 (12.3­18.3); 4</td>
<td id="843F51425B45001E65FC02CE1D6DFC68" box="[1750,1894,928,950]" gridcol="6" gridrow="4" pageId="6" pageNumber="489">11.3</td>
</tr>
<tr id="C7EE383E5B45001E635D02A71D6DFC01" box="[119,1894,969,991]" gridrow="5" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D02A71B5EFC01" box="[119,341,969,991]" gridcol="0" gridrow="5" pageId="6" pageNumber="489">External cirri</th>
<td id="843F51425B45001E62F202A7180BFC01" box="[472,512,969,991]" gridcol="1" gridrow="5" pageId="6" pageNumber="489">12.7</td>
<td id="843F51425B45001E616102A7194AFC01" box="[587,833,969,991]" gridcol="2" gridrow="5" pageId="6" pageNumber="489">13.4 ±1.94 (11.0­17.9); 16</td>
<td id="843F51425B45001E608602A719DFFC01" box="[940,980,969,991]" gridcol="3" gridrow="5" pageId="6" pageNumber="489">8.7</td>
<td id="843F51425B45001E673502A71F01FC01" box="[1055,1290,969,991]" gridcol="4" gridrow="5" pageId="6" pageNumber="489">11.9 ±1.54 (9.9­14.0); 5</td>
<td id="843F51425B45001E665F02A71C6BFC01" box="[1397,1632,969,991]" gridcol="5" gridrow="5" pageId="6" pageNumber="489">11.6 ±1.78 (9.5­13.9); 6</td>
<td id="843F51425B45001E65FC02A71D6DFC01" box="[1750,1894,969,991]" gridcol="6" gridrow="5" pageId="6" pageNumber="489">?</td>
</tr>
<tr id="C7EE383E5B45001E635D029C1D6DFBD6" box="[119,1894,1010,1032]" gridrow="6" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D029C1B5EFBD6" box="[119,341,1010,1032]" gridcol="0" gridrow="6" pageId="6" pageNumber="489">Lateral cirri A</th>
<td id="843F51425B45001E62F2029C180BFBD6" box="[472,512,1010,1032]" gridcol="1" gridrow="6" pageId="6" pageNumber="489">26.6</td>
<td id="843F51425B45001E6161029C194AFBD6" box="[587,833,1010,1032]" gridcol="2" gridrow="6" pageId="6" pageNumber="489">27.7 ±2.15 (24.5­31.5); 16</td>
<td id="843F51425B45001E6086029C19DFFBD6" box="[940,980,1010,1032]" gridcol="3" gridrow="6" pageId="6" pageNumber="489">25.6</td>
<td id="843F51425B45001E6735029C1F01FBD6" box="[1055,1290,1010,1032]" gridcol="4" gridrow="6" pageId="6" pageNumber="489">27.6 ±2.84 (24.4­31.3); 4</td>
<td id="843F51425B45001E665F029C1C6BFBD6" box="[1397,1632,1010,1032]" gridcol="5" gridrow="6" pageId="6" pageNumber="489">22.2 ±6.77 (14.8­29.0); 4</td>
<td id="843F51425B45001E65FC029C1D6DFBD6" box="[1750,1894,1010,1032]" gridcol="6" gridrow="6" pageId="6" pageNumber="489">14.4</td>
</tr>
<tr id="C7EE383E5B45001E635D05721D6DFBEC" box="[119,1894,1052,1074]" gridrow="7" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D05721B5EFBEC" box="[119,341,1052,1074]" gridcol="0" gridrow="7" pageId="6" pageNumber="489">Primary clavae</th>
<td id="843F51425B45001E62F20572180BFBEC" box="[472,512,1052,1074]" gridcol="1" gridrow="7" pageId="6" pageNumber="489">12.9</td>
<td id="843F51425B45001E61610572194AFBEC" box="[587,833,1052,1074]" gridcol="2" gridrow="7" pageId="6" pageNumber="489">12.6 ±1.84 (9.2­15.8); 16</td>
<td id="843F51425B45001E6086057219DFFBEC" box="[940,980,1052,1074]" gridcol="3" gridrow="7" pageId="6" pageNumber="489">13.5</td>
<td id="843F51425B45001E673505721F01FBEC" box="[1055,1290,1052,1074]" gridcol="4" gridrow="7" pageId="6" pageNumber="489">13.1 ±1.0 (12.1­14.1); 4</td>
<td id="843F51425B45001E665F05721C6BFBEC" box="[1397,1632,1052,1074]" gridcol="5" gridrow="7" pageId="6" pageNumber="489">12.4 ±0.71 (11.2­13.0); 5</td>
<td id="843F51425B45001E65FC05721D6DFBEC" box="[1750,1894,1052,1074]" gridcol="6" gridrow="7" pageId="6" pageNumber="489">7.4</td>
</tr>
<tr id="C7EE383E5B45001E635D052B1D6DFB85" box="[119,1894,1093,1115]" gridrow="8" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D052B1B5EFB85" box="[119,341,1093,1115]" gridcol="0" gridrow="8" pageId="6" pageNumber="489">Primary clava / cirrus A</th>
<td id="843F51425B45001E62F2052B180BFB85" box="[472,512,1093,1115]" gridcol="1" gridrow="8" pageId="6" pageNumber="489">0.49</td>
<td id="843F51425B45001E6161052B194AFB85" box="[587,833,1093,1115]" gridcol="2" gridrow="8" pageId="6" pageNumber="489">0.46 ±0.06 (0.35­0.53); 16</td>
<td id="843F51425B45001E6086052B19DFFB85" box="[940,980,1093,1115]" gridcol="3" gridrow="8" pageId="6" pageNumber="489">0.53</td>
<td id="843F51425B45001E6735052B1F01FB85" box="[1055,1290,1093,1115]" gridcol="4" gridrow="8" pageId="6" pageNumber="489">0.48 ±0.06 (0.45­0.57); 4</td>
<td id="843F51425B45001E665F052B1C6BFB85" box="[1397,1632,1093,1115]" gridcol="5" gridrow="8" pageId="6" pageNumber="489">0.65 ±0.15 (0.47­0.76); 3</td>
<td id="843F51425B45001E65FC052B1D6DFB85" box="[1750,1894,1093,1115]" gridcol="6" gridrow="8" pageId="6" pageNumber="489">0.51</td>
</tr>
<tr id="C7EE383E5B45001E635D05011D6DFB5B" box="[119,1894,1135,1157]" gridrow="9" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D05011B5EFB5B" box="[119,341,1135,1157]" gridcol="0" gridrow="9" pageId="6" pageNumber="489">Seconđary clavae</th>
<td id="843F51425B45001E62F20501180BFB5B" box="[472,512,1135,1157]" gridcol="1" gridrow="9" pageId="6" pageNumber="489">2.9</td>
<td id="843F51425B45001E61610501194AFB5B" box="[587,833,1135,1157]" gridcol="2" gridrow="9" pageId="6" pageNumber="489">3.2±0.79 (1.2­4.7); 17</td>
<td id="843F51425B45001E6086050119DFFB5B" box="[940,980,1135,1157]" gridcol="3" gridrow="9" pageId="6" pageNumber="489">?</td>
<td id="843F51425B45001E673505011F01FB5B" box="[1055,1290,1135,1157]" gridcol="4" gridrow="9" pageId="6" pageNumber="489">3.3 ±0.63 (2.6­4.1); 4</td>
<td id="843F51425B45001E665F05011C6BFB5B" box="[1397,1632,1135,1157]" gridcol="5" gridrow="9" pageId="6" pageNumber="489">2.9±0.58 (1.8­3.4); 6</td>
<td id="843F51425B45001E65FC05011D6DFB5B" box="[1750,1894,1135,1157]" gridcol="6" gridrow="9" pageId="6" pageNumber="489">3.3</td>
</tr>
<tr id="C7EE383E5B45001E635D05F61D6DFB70" box="[119,1894,1176,1198]" gridrow="10" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D05F61B5EFB70" box="[119,341,1176,1198]" gridcol="0" gridrow="10" pageId="6" pageNumber="489">Cirri E</th>
<td id="843F51425B45001E62F205F6180BFB70" box="[472,512,1176,1198]" gridcol="1" gridrow="10" pageId="6" pageNumber="489">13.1</td>
<td id="843F51425B45001E616105F6194AFB70" box="[587,833,1176,1198]" gridcol="2" gridrow="10" pageId="6" pageNumber="489">9.8 ±3.02 (7.2­13.1); 3</td>
<td id="843F51425B45001E608605F619DFFB70" box="[940,980,1176,1198]" gridcol="3" gridrow="10" pageId="6" pageNumber="489">7.3</td>
<td id="843F51425B45001E673505F61F01FB70" box="[1055,1290,1176,1198]" gridcol="4" gridrow="10" pageId="6" pageNumber="489">- (8.0); 1</td>
<td id="843F51425B45001E665F05F61C6BFB70" box="[1397,1632,1176,1198]" gridcol="5" gridrow="10" pageId="6" pageNumber="489">9.0 ±2.84 (7.1­12.3); 3</td>
<td id="843F51425B45001E65FC05F61D6DFB70" box="[1750,1894,1176,1198]" gridcol="6" gridrow="10" pageId="6" pageNumber="489">?</td>
</tr>
<tr id="C7EE383E5B45001E635D05AC1D6DFB06" box="[119,1894,1218,1240]" gridrow="11" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D05AC1B5EFB06" box="[119,341,1218,1240]" gridcol="0" gridrow="11" pageId="6" pageNumber="489">Leg I spine</th>
<td id="843F51425B45001E62F205AC180BFB06" box="[472,512,1218,1240]" gridcol="1" gridrow="11" pageId="6" pageNumber="489">5.1</td>
<td id="843F51425B45001E616105AC194AFB06" box="[587,833,1218,1240]" gridcol="2" gridrow="11" pageId="6" pageNumber="489">5.6 ±0.87 (4.0­6.3); 7</td>
<td id="843F51425B45001E608605AC19DFFB06" box="[940,980,1218,1240]" gridcol="3" gridrow="11" pageId="6" pageNumber="489">5.6</td>
<td id="843F51425B45001E673505AC1F01FB06" box="[1055,1290,1218,1240]" gridcol="4" gridrow="11" pageId="6" pageNumber="489">5.4 ±1.10 (4.3­6.5); 3</td>
<td id="843F51425B45001E665F05AC1C6BFB06" box="[1397,1632,1218,1240]" gridcol="5" gridrow="11" pageId="6" pageNumber="489">6.3 ±0.66 (5.7­7.0); 3</td>
<td id="843F51425B45001E65FC05AC1D6DFB06" box="[1750,1894,1218,1240]" gridcol="6" gridrow="11" pageId="6" pageNumber="489">?</td>
</tr>
<tr id="C7EE383E5B45001E635D05851D6DFADF" box="[119,1894,1259,1281]" gridrow="12" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D05851B5EFADF" box="[119,341,1259,1281]" gridcol="0" gridrow="12" pageId="6" pageNumber="489">Leg II spine</th>
<td id="843F51425B45001E62F20585180BFADF" box="[472,512,1259,1281]" gridcol="1" gridrow="12" pageId="6" pageNumber="489">8.2</td>
<td id="843F51425B45001E61610585194AFADF" box="[587,833,1259,1281]" gridcol="2" gridrow="12" pageId="6" pageNumber="489">7.3 ±2.12 (4.6­10.0); 7</td>
<td id="843F51425B45001E6086058519DFFADF" box="[940,980,1259,1281]" gridcol="3" gridrow="12" pageId="6" pageNumber="489">6.3</td>
<td id="843F51425B45001E673505851F01FADF" box="[1055,1290,1259,1281]" gridcol="4" gridrow="12" pageId="6" pageNumber="489">- (10.5­10.6); 2</td>
<td id="843F51425B45001E665F05851C6BFADF" box="[1397,1632,1259,1281]" gridcol="5" gridrow="12" pageId="6" pageNumber="489">- (11.0); 1</td>
<td id="843F51425B45001E65FC05851D6DFADF" box="[1750,1894,1259,1281]" gridcol="6" gridrow="12" pageId="6" pageNumber="489">?</td>
</tr>
<tr id="C7EE383E5B45001E635D047B1D6DFAF5" box="[119,1894,1301,1323]" gridrow="13" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D047B1B5EFAF5" box="[119,341,1301,1323]" gridcol="0" gridrow="13" pageId="6" pageNumber="489">Leg III spine</th>
<td id="843F51425B45001E62F2047B180BFAF5" box="[472,512,1301,1323]" gridcol="1" gridrow="13" pageId="6" pageNumber="489">9.6</td>
<td id="843F51425B45001E6161047B194AFAF5" box="[587,833,1301,1323]" gridcol="2" gridrow="13" pageId="6" pageNumber="489">6.9 ±2.09 (4.6­9.6); 8</td>
<td id="843F51425B45001E6086047B19DFFAF5" box="[940,980,1301,1323]" gridcol="3" gridrow="13" pageId="6" pageNumber="489">8.2</td>
<td id="843F51425B45001E6735047B1F01FAF5" box="[1055,1290,1301,1323]" gridcol="4" gridrow="13" pageId="6" pageNumber="489">- (5.9­9.2); 2</td>
<td id="843F51425B45001E665F047B1C6BFAF5" box="[1397,1632,1301,1323]" gridcol="5" gridrow="13" pageId="6" pageNumber="489">- (8.5­8.7); 2</td>
<td id="843F51425B45001E65FC047B1D6DFAF5" box="[1750,1894,1301,1323]" gridcol="6" gridrow="13" pageId="6" pageNumber="489">?</td>
</tr>
<tr id="C7EE383E5B45001E635D04501D6DFA8A" box="[119,1894,1342,1364]" gridrow="14" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D04501B5EFA8A" box="[119,341,1342,1364]" gridcol="0" gridrow="14" pageId="6" pageNumber="489">Leg IV sense organ</th>
<td id="843F51425B45001E62F20450180BFA8A" box="[472,512,1342,1364]" gridcol="1" gridrow="14" pageId="6" pageNumber="489">12.8</td>
<td id="843F51425B45001E61610450194AFA8A" box="[587,833,1342,1364]" gridcol="2" gridrow="14" pageId="6" pageNumber="489">12.6 ±2.08 (9.3­16.0); 16</td>
<td id="843F51425B45001E6086045019DFFA8A" box="[940,980,1342,1364]" gridcol="3" gridrow="14" pageId="6" pageNumber="489">15.6</td>
<td id="843F51425B45001E673504501F01FA8A" box="[1055,1290,1342,1364]" gridcol="4" gridrow="14" pageId="6" pageNumber="489">13.7 ±1.47 (12.0­15.3); 4</td>
<td id="843F51425B45001E665F04501C6BFA8A" box="[1397,1632,1342,1364]" gridcol="5" gridrow="14" pageId="6" pageNumber="489">13.1 ±2.50 (9.6­15.7); 3</td>
<td id="843F51425B45001E65FC04501D6DFA8A" box="[1750,1894,1342,1364]" gridcol="6" gridrow="14" pageId="6" pageNumber="489">?</td>
</tr>
<tr id="C7EE383E5B45001E635D04091D6DFAA3" box="[119,1894,1383,1405]" gridrow="15" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D04091B5EFAA3" box="[119,341,1383,1405]" gridcol="0" gridrow="15" pageId="6" pageNumber="489">Leg IV base</th>
<td id="843F51425B45001E62F20409180BFAA3" box="[472,512,1383,1405]" gridcol="1" gridrow="15" pageId="6" pageNumber="489">5.8</td>
<td id="843F51425B45001E61610409194AFAA3" box="[587,833,1383,1405]" gridcol="2" gridrow="15" pageId="6" pageNumber="489">6.3 ±0.88 (5.1­7.6); 13</td>
<td id="843F51425B45001E6086040919DFFAA3" box="[940,980,1383,1405]" gridcol="3" gridrow="15" pageId="6" pageNumber="489">7.8</td>
<td id="843F51425B45001E673504091F01FAA3" box="[1055,1290,1383,1405]" gridcol="4" gridrow="15" pageId="6" pageNumber="489">6.2 ±1.15 (4.7­7.1); 4</td>
<td id="843F51425B45001E665F04091C6BFAA3" box="[1397,1632,1383,1405]" gridcol="5" gridrow="15" pageId="6" pageNumber="489">5.9 ±0.66 (5.1­6.7); 4</td>
<td id="843F51425B45001E65FC04091D6DFAA3" box="[1750,1894,1383,1405]" gridcol="6" gridrow="15" pageId="6" pageNumber="489">?</td>
</tr>
<tr id="C7EE383E5B45001E635D04FF1D6DFA79" box="[119,1894,1425,1447]" gridrow="16" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D04FF1B5EFA79" box="[119,341,1425,1447]" gridcol="0" gridrow="16" pageId="6" pageNumber="489">Leg IV spine</th>
<td id="843F51425B45001E62F204FF180BFA79" box="[472,512,1425,1447]" gridcol="1" gridrow="16" pageId="6" pageNumber="489">7.0</td>
<td id="843F51425B45001E616104FF194AFA79" box="[587,833,1425,1447]" gridcol="2" gridrow="16" pageId="6" pageNumber="489">7.2 ±1.61 (5.4­10.9); 11</td>
<td id="843F51425B45001E608604FF19DFFA79" box="[940,980,1425,1447]" gridcol="3" gridrow="16" pageId="6" pageNumber="489">7.8</td>
<td id="843F51425B45001E673504FF1F01FA79" box="[1055,1290,1425,1447]" gridcol="4" gridrow="16" pageId="6" pageNumber="489">6.4 ±1.28 (4.5­7.3); 4</td>
<td id="843F51425B45001E665F04FF1C6BFA79" box="[1397,1632,1425,1447]" gridcol="5" gridrow="16" pageId="6" pageNumber="489">7.6 ±1.70 (5.7­9.5); 5</td>
<td id="843F51425B45001E65FC04FF1D6DFA79" box="[1750,1894,1425,1447]" gridcol="6" gridrow="16" pageId="6" pageNumber="489">?</td>
</tr>
<tr id="C7EE383E5B45001E635D04D41D6DFA0E" box="[119,1894,1466,1488]" gridrow="17" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D04D41B5EFA0E" box="[119,341,1466,1488]" gridcol="0" gridrow="17" pageId="6" pageNumber="489">Leg I projection</th>
<td id="843F51425B45001E62F204D4180BFA0E" box="[472,512,1466,1488]" gridcol="1" gridrow="17" pageId="6" pageNumber="489">3.5</td>
<td id="843F51425B45001E616104D4194AFA0E" box="[587,833,1466,1488]" gridcol="2" gridrow="17" pageId="6" pageNumber="489">3.6 ±0.94 (2.0­5.0); 11</td>
<td id="843F51425B45001E608604D419DFFA0E" box="[940,980,1466,1488]" gridcol="3" gridrow="17" pageId="6" pageNumber="489">3.5</td>
<td id="843F51425B45001E673504D41F01FA0E" box="[1055,1290,1466,1488]" gridcol="4" gridrow="17" pageId="6" pageNumber="489">3.9 ±0.74 (3.1­4.8); 5</td>
<td id="843F51425B45001E665F04D41C6BFA0E" box="[1397,1632,1466,1488]" gridcol="5" gridrow="17" pageId="6" pageNumber="489">3.9 ±0.55 (3.3­4.5); 4</td>
<td id="843F51425B45001E65FC04D41D6DFA0E" box="[1750,1894,1466,1488]" gridcol="6" gridrow="17" pageId="6" pageNumber="489">2.2</td>
</tr>
<tr id="C7EE383E5B45001E635D048A1D6DFA24" box="[119,1894,1508,1530]" gridrow="18" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D048A1B5EFA24" box="[119,341,1508,1530]" gridcol="0" gridrow="18" pageId="6" pageNumber="489">Leg II projection</th>
<td id="843F51425B45001E62F2048A180BFA24" box="[472,512,1508,1530]" gridcol="1" gridrow="18" pageId="6" pageNumber="489">3.3</td>
<td id="843F51425B45001E6161048A194AFA24" box="[587,833,1508,1530]" gridcol="2" gridrow="18" pageId="6" pageNumber="489">3.4 ±0.43 (2.8­4.0); 7</td>
<td id="843F51425B45001E6086048A19DFFA24" box="[940,980,1508,1530]" gridcol="3" gridrow="18" pageId="6" pageNumber="489">5.1</td>
<td id="843F51425B45001E6735048A1F01FA24" box="[1055,1290,1508,1530]" gridcol="4" gridrow="18" pageId="6" pageNumber="489">3.5 ±1.56 (2.0­6.0); 5</td>
<td id="843F51425B45001E665F048A1C6BFA24" box="[1397,1632,1508,1530]" gridcol="5" gridrow="18" pageId="6" pageNumber="489">3.0 ±0.93 (1.8­3.7); 4</td>
<td id="843F51425B45001E65FC048A1D6DFA24" box="[1750,1894,1508,1530]" gridcol="6" gridrow="18" pageId="6" pageNumber="489">?</td>
</tr>
<tr id="C7EE383E5B45001E635D07631D6DF9FD" box="[119,1894,1549,1571]" gridrow="19" pageId="6" pageNumber="489">
<th id="843F51425B45001E635D07631B5EF9FD" box="[119,341,1549,1571]" gridcol="0" gridrow="19" pageId="6" pageNumber="489">Leg III projection</th>
<td id="843F51425B45001E62F20763180BF9FD" box="[472,512,1549,1571]" gridcol="1" gridrow="19" pageId="6" pageNumber="489">2.1</td>
<td id="843F51425B45001E61610763194AF9FD" box="[587,833,1549,1571]" gridcol="2" gridrow="19" pageId="6" pageNumber="489">- (2.2); 1</td>
<td id="843F51425B45001E6086076319DFF9FD" box="[940,980,1549,1571]" gridcol="3" gridrow="19" pageId="6" pageNumber="489">2.4</td>
<td id="843F51425B45001E673507631F01F9FD" box="[1055,1290,1549,1571]" gridcol="4" gridrow="19" pageId="6" pageNumber="489">- (4.4); 2</td>
<td id="843F51425B45001E665F07631C6BF9FD" box="[1397,1632,1549,1571]" gridcol="5" gridrow="19" pageId="6" pageNumber="489">- (1.7­3.3); 2</td>
<td id="843F51425B45001E65FC07631D6DF9FD" box="[1750,1894,1549,1571]" gridcol="6" gridrow="19" pageId="6" pageNumber="489">?</td>
</tr>
</table>
</paragraph>
<paragraph id="79613A7C5B45FFE9656407561F96FF0A" blockId="6.[1614,1894,1592,1614]" lastBlockId="8.[151,1437,151,1005]" lastPageId="8" lastPageNumber="491" pageId="6" pageNumber="489">
<emphasis id="4BAAE66E5B45FFE7656407561D6DF990" box="[1614,1894,1592,1614]" italics="true" pageId="6" pageNumber="489">......continued on the next page</emphasis>
<emphasis id="4BAAE66E5B44FFE6635703CF1AEAFD69" bold="true" box="[125,225,673,695]" pageId="7" pageNumber="490">TABLE].</emphasis>
(Continueđ) species. 
<taxonomicName id="BEDE41FF5B4BFFE963D001F61BE5FF6E" box="[250,494,152,176]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="8" pageNumber="491" phylum="Tardigrada" rank="species" species="acuticauda">
<emphasis id="4BAAE66E5B4BFFE963D001F61BE5FF6E" box="[250,494,152,176]" italics="true" pageId="8" pageNumber="491">Batillipes acuticauda</emphasis>
</taxonomicName>
, in contrary to the new species, has an acute spiky caudal appendage, cephalic appendages sharply pointed, and very different shaped lateral projections that are flattened between legs III and IV.
</paragraph>
<paragraph id="79613A7C5B44FFE66357026A1CEFF9EF" pageId="7" pageNumber="490">
<table id="0BDEC8DC5B44001E635703A21D21F9EF" box="[125,1834,716,1585]" gridcols="7" gridrows="20" pageId="7" pageNumber="490">
<tr id="C7EE383E5B44001E635703A21D21FCC4" box="[125,1834,716,794]" gridrow="0" pageId="7" pageNumber="490">
<th id="843F51425B44001E635703A21B6BFCC4" box="[125,352,716,794]" gridcol="0" gridrow="0" pageId="7" pageNumber="490">STRUCTURES</th>
<th id="843F51425B44001E62F703A2184DFCC4" box="[477,582,716,794]" gridcol="1" gridrow="0" pageId="7" pageNumber="490">FEMALES ađults Holo</th>
<th id="843F51425B44001E617B03A2194DFCC4" box="[593,838,716,794]" gridcol="2" gridrow="0" pageId="7" pageNumber="490">Mean ± SD (Range); N</th>
<th id="843F51425B44001E609B03A219F4FCC4" box="[945,1023,716,794]" gridcol="3" gridrow="0" pageId="7" pageNumber="490">MALES ađults Allo</th>
<th id="843F51425B44001E670E03A21F0FFCC4" box="[1060,1284,716,794]" gridcol="4" gridrow="0" pageId="7" pageNumber="490">Mean ± SD (Range); N</th>
<th id="843F51425B44001E665003A21C6EFCC4" box="[1402,1637,716,794]" gridcol="5" gridrow="0" pageId="7" pageNumber="490">JUVENILES 6-toeđ Mean ± SD (Range); N</th>
<th id="843F51425B44001E65F603A21D21FCC4" box="[1756,1834,716,794]" gridcol="6" gridrow="0" pageId="7" pageNumber="490">LARVA 4-toeđ</th>
</tr>
<tr id="C7EE383E5B44001E6357025E1D21FC98" box="[125,1834,816,838]" gridrow="1" pageId="7" pageNumber="490">
<th id="843F51425B44001E6357025E1B6BFC98" box="[125,352,816,838]" gridcol="0" gridrow="1" pageId="7" pageNumber="490">Bođy projection 1</th>
<td id="843F51425B44001E62F7025E184DFC98" box="[477,582,816,838]" gridcol="1" gridrow="1" pageId="7" pageNumber="490">4.1</td>
<td id="843F51425B44001E617B025E194DFC98" box="[593,838,816,838]" gridcol="2" gridrow="1" pageId="7" pageNumber="490">3.5 ±0.52 (2.8­4.1); 5</td>
<td id="843F51425B44001E609B025E19F4FC98" box="[945,1023,816,838]" gridcol="3" gridrow="1" pageId="7" pageNumber="490">?</td>
<td id="843F51425B44001E670E025E1F0FFC98" box="[1060,1284,816,838]" gridcol="4" gridrow="1" pageId="7" pageNumber="490">- (3.2); 1</td>
<td id="843F51425B44001E6650025E1C6EFC98" box="[1402,1637,816,838]" gridcol="5" gridrow="1" pageId="7" pageNumber="490">- (2.5); 1</td>
<td id="843F51425B44001E65F6025E1D21FC98" box="[1756,1834,816,838]" gridcol="6" gridrow="1" pageId="7" pageNumber="490">1.8</td>
</tr>
<tr id="C7EE383E5B44001E635702371D21FCB1" box="[125,1834,857,879]" gridrow="2" pageId="7" pageNumber="490">
<th id="843F51425B44001E635702371B6BFCB1" box="[125,352,857,879]" gridcol="0" gridrow="2" pageId="7" pageNumber="490">Bođy projection 2</th>
<td id="843F51425B44001E62F70237184DFCB1" box="[477,582,857,879]" gridcol="1" gridrow="2" pageId="7" pageNumber="490">6.0</td>
<td id="843F51425B44001E617B0237194DFCB1" box="[593,838,857,879]" gridcol="2" gridrow="2" pageId="7" pageNumber="490">4.7 ±1.87 (2.7­6.6); 4</td>
<td id="843F51425B44001E609B023719F4FCB1" box="[945,1023,857,879]" gridcol="3" gridrow="2" pageId="7" pageNumber="490">1.9</td>
<td id="843F51425B44001E670E02371F0FFCB1" box="[1060,1284,857,879]" gridcol="4" gridrow="2" pageId="7" pageNumber="490">- (4.8); 1</td>
<td id="843F51425B44001E665002371C6EFCB1" box="[1402,1637,857,879]" gridcol="5" gridrow="2" pageId="7" pageNumber="490">- (3.0); 1</td>
<td id="843F51425B44001E65F602371D21FCB1" box="[1756,1834,857,879]" gridcol="6" gridrow="2" pageId="7" pageNumber="490">2.6</td>
</tr>
<tr id="C7EE383E5B44001E635702ED1D21FC47" box="[125,1834,899,921]" gridrow="3" pageId="7" pageNumber="490">
<th id="843F51425B44001E635702ED1B6BFC47" box="[125,352,899,921]" gridcol="0" gridrow="3" pageId="7" pageNumber="490">Bođy projection 3</th>
<td id="843F51425B44001E62F702ED184DFC47" box="[477,582,899,921]" gridcol="1" gridrow="3" pageId="7" pageNumber="490">4.6</td>
<td id="843F51425B44001E617B02ED194DFC47" box="[593,838,899,921]" gridcol="2" gridrow="3" pageId="7" pageNumber="490">5.1 ±1.51 (2.9­7.7); 10</td>
<td id="843F51425B44001E609B02ED19F4FC47" box="[945,1023,899,921]" gridcol="3" gridrow="3" pageId="7" pageNumber="490">5.6</td>
<td id="843F51425B44001E670E02ED1F0FFC47" box="[1060,1284,899,921]" gridcol="4" gridrow="3" pageId="7" pageNumber="490">5.6 ±0.58 (5.0­6.1); 3</td>
<td id="843F51425B44001E665002ED1C6EFC47" box="[1402,1637,899,921]" gridcol="5" gridrow="3" pageId="7" pageNumber="490">- (3.8); 1</td>
<td id="843F51425B44001E65F602ED1D21FC47" box="[1756,1834,899,921]" gridcol="6" gridrow="3" pageId="7" pageNumber="490">3.1</td>
</tr>
<tr id="C7EE383E5B44001E635702C21D21FC1C" box="[125,1834,940,962]" gridrow="4" pageId="7" pageNumber="490">
<th id="843F51425B44001E635702C21B6BFC1C" box="[125,352,940,962]" gridcol="0" gridrow="4" pageId="7" pageNumber="490">Bođy projection 4 wiđth</th>
<td id="843F51425B44001E62F702C2184DFC1C" box="[477,582,940,962]" gridcol="1" gridrow="4" pageId="7" pageNumber="490">10.8</td>
<td id="843F51425B44001E617B02C2194DFC1C" box="[593,838,940,962]" gridcol="2" gridrow="4" pageId="7" pageNumber="490">11.5 ±3.32 (5.5­18.4); 14</td>
<td id="843F51425B44001E609B02C219F4FC1C" box="[945,1023,940,962]" gridcol="3" gridrow="4" pageId="7" pageNumber="490">17.6</td>
<td id="843F51425B44001E670E02C21F0FFC1C" box="[1060,1284,940,962]" gridcol="4" gridrow="4" pageId="7" pageNumber="490">15.1 ±4.77 (7.5­19.8); 5</td>
<td id="843F51425B44001E665002C21C6EFC1C" box="[1402,1637,940,962]" gridcol="5" gridrow="4" pageId="7" pageNumber="490">11.0 ±4.71 (4.7­16.1); 6</td>
<td id="843F51425B44001E65F602C21D21FC1C" box="[1756,1834,940,962]" gridcol="6" gridrow="4" pageId="7" pageNumber="490">5.1</td>
</tr>
<tr id="C7EE383E5B44001E635702B81D21FC32" box="[125,1834,982,1004]" gridrow="5" pageId="7" pageNumber="490">
<th id="843F51425B44001E635702B81B6BFC32" box="[125,352,982,1004]" gridcol="0" gridrow="5" pageId="7" pageNumber="490">Bođy projection 4 length</th>
<td id="843F51425B44001E62F702B8184DFC32" box="[477,582,982,1004]" gridcol="1" gridrow="5" pageId="7" pageNumber="490">5.8</td>
<td id="843F51425B44001E617B02B8194DFC32" box="[593,838,982,1004]" gridcol="2" gridrow="5" pageId="7" pageNumber="490">9.2 ±3.29 (5.2­17.7); 14</td>
<td id="843F51425B44001E609B02B819F4FC32" box="[945,1023,982,1004]" gridcol="3" gridrow="5" pageId="7" pageNumber="490">10.9</td>
<td id="843F51425B44001E670E02B81F0FFC32" box="[1060,1284,982,1004]" gridcol="4" gridrow="5" pageId="7" pageNumber="490">11.1 ±2.84 (7.4­15.3); 5</td>
<td id="843F51425B44001E665002B81C6EFC32" box="[1402,1637,982,1004]" gridcol="5" gridrow="5" pageId="7" pageNumber="490">9.5 ±3.5 (3.6­14.0); 6</td>
<td id="843F51425B44001E65F602B81D21FC32" box="[1756,1834,982,1004]" gridcol="6" gridrow="5" pageId="7" pageNumber="490">5.3</td>
</tr>
<tr id="C7EE383E5B44001E635702911D21FBCB" box="[125,1834,1023,1045]" gridrow="6" pageId="7" pageNumber="490">
<th id="843F51425B44001E635702911B6BFBCB" box="[125,352,1023,1045]" gridcol="0" gridrow="6" pageId="7" pageNumber="490">Cauđal apparatus</th>
<td id="843F51425B44001E62F70291184DFBCB" box="[477,582,1023,1045]" gridcol="1" gridrow="6" pageId="7" pageNumber="490">17.5</td>
<td id="843F51425B44001E617B0291194DFBCB" box="[593,838,1023,1045]" gridcol="2" gridrow="6" pageId="7" pageNumber="490">10.1 ±4.78 (1.8­19.2); 15</td>
<td id="843F51425B44001E609B029119F4FBCB" box="[945,1023,1023,1045]" gridcol="3" gridrow="6" pageId="7" pageNumber="490">16.3</td>
<td id="843F51425B44001E670E02911F0FFBCB" box="[1060,1284,1023,1045]" gridcol="4" gridrow="6" pageId="7" pageNumber="490">13.2 ±3.14 (9.0­16.2); 5</td>
<td id="843F51425B44001E665002911C6EFBCB" box="[1402,1637,1023,1045]" gridcol="5" gridrow="6" pageId="7" pageNumber="490">12.7 ±4.20 (5.7­17.8); 6</td>
<td id="843F51425B44001E65F602911D21FBCB" box="[1756,1834,1023,1045]" gridcol="6" gridrow="6" pageId="7" pageNumber="490">4.8</td>
</tr>
<tr id="C7EE383E5B44001E635705471D21FBE1" box="[125,1834,1065,1087]" gridrow="7" pageId="7" pageNumber="490">
<th id="843F51425B44001E635705471B6BFBE1" box="[125,352,1065,1087]" gridcol="0" gridrow="7" pageId="7" pageNumber="490">Legs I­III: Toe 1</th>
<td id="843F51425B44001E62F70547184DFBE1" box="[477,582,1065,1087]" gridcol="1" gridrow="7" pageId="7" pageNumber="490">7.0</td>
<td id="843F51425B44001E617B0547194DFBE1" box="[593,838,1065,1087]" gridcol="2" gridrow="7" pageId="7" pageNumber="490">7.4 ±1.44 (5.3­10.1); 11</td>
<td id="843F51425B44001E609B054719F4FBE1" box="[945,1023,1065,1087]" gridcol="3" gridrow="7" pageId="7" pageNumber="490">6.3</td>
<td id="843F51425B44001E670E05471F0FFBE1" box="[1060,1284,1065,1087]" gridcol="4" gridrow="7" pageId="7" pageNumber="490">- (7.3); 1</td>
<td id="843F51425B44001E665005471C6EFBE1" box="[1402,1637,1065,1087]" gridcol="5" gridrow="7" pageId="7" pageNumber="490">- (5.1­10.5); 2</td>
<td id="843F51425B44001E65F605471D21FBE1" box="[1756,1834,1065,1087]" gridcol="6" gridrow="7" pageId="7" pageNumber="490">6.0</td>
</tr>
<tr id="C7EE383E5B44001E6357053C1D21FBB6" box="[125,1834,1106,1128]" gridrow="8" pageId="7" pageNumber="490">
<th id="843F51425B44001E6357053C1B6BFBB6" box="[125,352,1106,1128]" gridcol="0" gridrow="8" pageId="7" pageNumber="490">Toe 2</th>
<td id="843F51425B44001E62F7053C184DFBB6" box="[477,582,1106,1128]" gridcol="1" gridrow="8" pageId="7" pageNumber="490">3.2</td>
<td id="843F51425B44001E617B053C194DFBB6" box="[593,838,1106,1128]" gridcol="2" gridrow="8" pageId="7" pageNumber="490">4.2 ±0.94 (2.5­5.9); 11</td>
<td id="843F51425B44001E609B053C19F4FBB6" box="[945,1023,1106,1128]" gridcol="3" gridrow="8" pageId="7" pageNumber="490">2.4</td>
<td id="843F51425B44001E670E053C1F0FFBB6" box="[1060,1284,1106,1128]" gridcol="4" gridrow="8" pageId="7" pageNumber="490">- (3.8); 1</td>
<td id="843F51425B44001E6650053C1C6EFBB6" box="[1402,1637,1106,1128]" gridcol="5" gridrow="8" pageId="7" pageNumber="490">- (2.6­4.9); 2</td>
<td id="843F51425B44001E65F6053C1D21FBB6" box="[1756,1834,1106,1128]" gridcol="6" gridrow="8" pageId="7" pageNumber="490">-</td>
</tr>
<tr id="C7EE383E5B44001E635705121D21FB4C" box="[125,1834,1148,1170]" gridrow="9" pageId="7" pageNumber="490">
<th id="843F51425B44001E635705121B6BFB4C" box="[125,352,1148,1170]" gridcol="0" gridrow="9" pageId="7" pageNumber="490">Toe 3</th>
<td id="843F51425B44001E62F70512184DFB4C" box="[477,582,1148,1170]" gridcol="1" gridrow="9" pageId="7" pageNumber="490">13.0</td>
<td id="843F51425B44001E617B0512194DFB4C" box="[593,838,1148,1170]" gridcol="2" gridrow="9" pageId="7" pageNumber="490">11.8 ±2.35 (8.8­15.5); 11</td>
<td id="843F51425B44001E609B051219F4FB4C" box="[945,1023,1148,1170]" gridcol="3" gridrow="9" pageId="7" pageNumber="490">13.7</td>
<td id="843F51425B44001E670E05121F0FFB4C" box="[1060,1284,1148,1170]" gridcol="4" gridrow="9" pageId="7" pageNumber="490">- (11.1); 1</td>
<td id="843F51425B44001E665005121C6EFB4C" box="[1402,1637,1148,1170]" gridcol="5" gridrow="9" pageId="7" pageNumber="490">- (9.7­13.0); 2</td>
<td id="843F51425B44001E65F605121D21FB4C" box="[1756,1834,1148,1170]" gridcol="6" gridrow="9" pageId="7" pageNumber="490">7.4</td>
</tr>
<tr id="C7EE383E5B44001E635705CB1D21FB65" box="[125,1834,1189,1211]" gridrow="10" pageId="7" pageNumber="490">
<th id="843F51425B44001E635705CB1B6BFB65" box="[125,352,1189,1211]" gridcol="0" gridrow="10" pageId="7" pageNumber="490">Toe 4</th>
<td id="843F51425B44001E62F705CB184DFB65" box="[477,582,1189,1211]" gridcol="1" gridrow="10" pageId="7" pageNumber="490">6.2</td>
<td id="843F51425B44001E617B05CB194DFB65" box="[593,838,1189,1211]" gridcol="2" gridrow="10" pageId="7" pageNumber="490">6.7 ±1.47 (3.7­8.9); 11</td>
<td id="843F51425B44001E609B05CB19F4FB65" box="[945,1023,1189,1211]" gridcol="3" gridrow="10" pageId="7" pageNumber="490">6.4</td>
<td id="843F51425B44001E670E05CB1F0FFB65" box="[1060,1284,1189,1211]" gridcol="4" gridrow="10" pageId="7" pageNumber="490">- (6.0); 1</td>
<td id="843F51425B44001E665005CB1C6EFB65" box="[1402,1637,1189,1211]" gridcol="5" gridrow="10" pageId="7" pageNumber="490">- (5.2­7.1); 2</td>
<td id="843F51425B44001E65F605CB1D21FB65" box="[1756,1834,1189,1211]" gridcol="6" gridrow="10" pageId="7" pageNumber="490">7.9</td>
</tr>
<tr id="C7EE383E5B44001E635705A11D21FB3B" box="[125,1834,1231,1253]" gridrow="11" pageId="7" pageNumber="490">
<th id="843F51425B44001E635705A11B6BFB3B" box="[125,352,1231,1253]" gridcol="0" gridrow="11" pageId="7" pageNumber="490">Toe 5</th>
<td id="843F51425B44001E62F705A1184DFB3B" box="[477,582,1231,1253]" gridcol="1" gridrow="11" pageId="7" pageNumber="490">14.1</td>
<td id="843F51425B44001E617B05A1194DFB3B" box="[593,838,1231,1253]" gridcol="2" gridrow="11" pageId="7" pageNumber="490">13.1 ±2.05 (9.9­15.7); 10</td>
<td id="843F51425B44001E609B05A119F4FB3B" box="[945,1023,1231,1253]" gridcol="3" gridrow="11" pageId="7" pageNumber="490">16.0</td>
<td id="843F51425B44001E670E05A11F0FFB3B" box="[1060,1284,1231,1253]" gridcol="4" gridrow="11" pageId="7" pageNumber="490">- (15.5); 1</td>
<td id="843F51425B44001E665005A11C6EFB3B" box="[1402,1637,1231,1253]" gridcol="5" gridrow="11" pageId="7" pageNumber="490">- (11.0­14.1); 2</td>
<td id="843F51425B44001E65F605A11D21FB3B" box="[1756,1834,1231,1253]" gridcol="6" gridrow="11" pageId="7" pageNumber="490">-</td>
</tr>
<tr id="C7EE383E5B44001E635705961D21FAD0" box="[125,1834,1272,1294]" gridrow="12" pageId="7" pageNumber="490">
<th id="843F51425B44001E635705961B6BFAD0" box="[125,352,1272,1294]" gridcol="0" gridrow="12" pageId="7" pageNumber="490">Toe 6</th>
<td id="843F51425B44001E62F70596184DFAD0" box="[477,582,1272,1294]" gridcol="1" gridrow="12" pageId="7" pageNumber="490">7.3</td>
<td id="843F51425B44001E617B0596194DFAD0" box="[593,838,1272,1294]" gridcol="2" gridrow="12" pageId="7" pageNumber="490">8.2 ±1.51 (4.8­9.7); 10</td>
<td id="843F51425B44001E609B059619F4FAD0" box="[945,1023,1272,1294]" gridcol="3" gridrow="12" pageId="7" pageNumber="490">8.9</td>
<td id="843F51425B44001E670E05961F0FFAD0" box="[1060,1284,1272,1294]" gridcol="4" gridrow="12" pageId="7" pageNumber="490">- (9.4); 1</td>
<td id="843F51425B44001E665005961C6EFAD0" box="[1402,1637,1272,1294]" gridcol="5" gridrow="12" pageId="7" pageNumber="490">- (6.9­9.2); 2</td>
<td id="843F51425B44001E65F605961D21FAD0" box="[1756,1834,1272,1294]" gridcol="6" gridrow="12" pageId="7" pageNumber="490">13.5</td>
</tr>
<tr id="C7EE383E5B44001E6357044C1D21FAE6" box="[125,1834,1314,1336]" gridrow="13" pageId="7" pageNumber="490">
<th id="843F51425B44001E6357044C1B6BFAE6" box="[125,352,1314,1336]" gridcol="0" gridrow="13" pageId="7" pageNumber="490">Leg IV: Toe 1</th>
<td id="843F51425B44001E62F7044C184DFAE6" box="[477,582,1314,1336]" gridcol="1" gridrow="13" pageId="7" pageNumber="490">11.3</td>
<td id="843F51425B44001E617B044C194DFAE6" box="[593,838,1314,1336]" gridcol="2" gridrow="13" pageId="7" pageNumber="490">11.2 ±0.92 (8.8­12.5); 12</td>
<td id="843F51425B44001E609B044C19F4FAE6" box="[945,1023,1314,1336]" gridcol="3" gridrow="13" pageId="7" pageNumber="490">?</td>
<td id="843F51425B44001E670E044C1F0FFAE6" box="[1060,1284,1314,1336]" gridcol="4" gridrow="13" pageId="7" pageNumber="490">- (9.6­11.8); 2</td>
<td id="843F51425B44001E6650044C1C6EFAE6" box="[1402,1637,1314,1336]" gridcol="5" gridrow="13" pageId="7" pageNumber="490">11.0 ±1.39 (9.3­12.7); 4</td>
<td id="843F51425B44001E65F6044C1D21FAE6" box="[1756,1834,1314,1336]" gridcol="6" gridrow="13" pageId="7" pageNumber="490">7.0</td>
</tr>
<tr id="C7EE383E5B44001E635704221D21FABC" box="[125,1834,1356,1378]" gridrow="14" pageId="7" pageNumber="490">
<th id="843F51425B44001E635704221B6BFABC" box="[125,352,1356,1378]" gridcol="0" gridrow="14" pageId="7" pageNumber="490">Toe 2</th>
<td id="843F51425B44001E62F70422184DFABC" box="[477,582,1356,1378]" gridcol="1" gridrow="14" pageId="7" pageNumber="490">16.3</td>
<td id="843F51425B44001E617B0422194DFABC" box="[593,838,1356,1378]" gridcol="2" gridrow="14" pageId="7" pageNumber="490">16.2 ±1.94 (12.6­18.9); 11</td>
<td id="843F51425B44001E609B042219F4FABC" box="[945,1023,1356,1378]" gridcol="3" gridrow="14" pageId="7" pageNumber="490">?</td>
<td id="843F51425B44001E670E04221F0FFABC" box="[1060,1284,1356,1378]" gridcol="4" gridrow="14" pageId="7" pageNumber="490">- (14.8­17.2); 2</td>
<td id="843F51425B44001E665004221C6EFABC" box="[1402,1637,1356,1378]" gridcol="5" gridrow="14" pageId="7" pageNumber="490">14.7 ±2.07 (11.7­16.2); 4</td>
<td id="843F51425B44001E65F604221D21FABC" box="[1756,1834,1356,1378]" gridcol="6" gridrow="14" pageId="7" pageNumber="490">15.6</td>
</tr>
<tr id="C7EE383E5B44001E6357041B1D21FA55" box="[125,1834,1397,1419]" gridrow="15" pageId="7" pageNumber="490">
<th id="843F51425B44001E6357041B1B6BFA55" box="[125,352,1397,1419]" gridcol="0" gridrow="15" pageId="7" pageNumber="490">Toe 3</th>
<td id="843F51425B44001E62F7041B184DFA55" box="[477,582,1397,1419]" gridcol="1" gridrow="15" pageId="7" pageNumber="490">6.4</td>
<td id="843F51425B44001E617B041B194DFA55" box="[593,838,1397,1419]" gridcol="2" gridrow="15" pageId="7" pageNumber="490">6.5 ±1.10 (3.7­8.1); 12</td>
<td id="843F51425B44001E609B041B19F4FA55" box="[945,1023,1397,1419]" gridcol="3" gridrow="15" pageId="7" pageNumber="490">?</td>
<td id="843F51425B44001E670E041B1F0FFA55" box="[1060,1284,1397,1419]" gridcol="4" gridrow="15" pageId="7" pageNumber="490">- (3.6­5.7); 2</td>
<td id="843F51425B44001E6650041B1C6EFA55" box="[1402,1637,1397,1419]" gridcol="5" gridrow="15" pageId="7" pageNumber="490">5.0 ±1.00 (4.0­6.0); 3</td>
<td id="843F51425B44001E65F6041B1D21FA55" box="[1756,1834,1397,1419]" gridcol="6" gridrow="15" pageId="7" pageNumber="490">-</td>
</tr>
<tr id="C7EE383E5B44001E635704F11D21FA6B" box="[125,1834,1439,1461]" gridrow="16" pageId="7" pageNumber="490">
<th id="843F51425B44001E635704F11B6BFA6B" box="[125,352,1439,1461]" gridcol="0" gridrow="16" pageId="7" pageNumber="490">Toe 4</th>
<td id="843F51425B44001E62F704F1184DFA6B" box="[477,582,1439,1461]" gridcol="1" gridrow="16" pageId="7" pageNumber="490">10.7</td>
<td id="843F51425B44001E617B04F1194DFA6B" box="[593,838,1439,1461]" gridcol="2" gridrow="16" pageId="7" pageNumber="490">10.6 ±0.92 (9.1­11.9); 12</td>
<td id="843F51425B44001E609B04F119F4FA6B" box="[945,1023,1439,1461]" gridcol="3" gridrow="16" pageId="7" pageNumber="490">?</td>
<td id="843F51425B44001E670E04F11F0FFA6B" box="[1060,1284,1439,1461]" gridcol="4" gridrow="16" pageId="7" pageNumber="490">- (7.5­9.3); 2</td>
<td id="843F51425B44001E665004F11C6EFA6B" box="[1402,1637,1439,1461]" gridcol="5" gridrow="16" pageId="7" pageNumber="490">8.8 ±1.96 (7.4­10.2); 3</td>
<td id="843F51425B44001E65F604F11D21FA6B" box="[1756,1834,1439,1461]" gridcol="6" gridrow="16" pageId="7" pageNumber="490">-</td>
</tr>
<tr id="C7EE383E5B44001E635704A61D21FA00" box="[125,1834,1480,1502]" gridrow="17" pageId="7" pageNumber="490">
<th id="843F51425B44001E635704A61B6BFA00" box="[125,352,1480,1502]" gridcol="0" gridrow="17" pageId="7" pageNumber="490">Toe 5</th>
<td id="843F51425B44001E62F704A6184DFA00" box="[477,582,1480,1502]" gridcol="1" gridrow="17" pageId="7" pageNumber="490">20.5</td>
<td id="843F51425B44001E617B04A6194DFA00" box="[593,838,1480,1502]" gridcol="2" gridrow="17" pageId="7" pageNumber="490">19.3 ±2.31 (15.3­22.6); 11</td>
<td id="843F51425B44001E609B04A619F4FA00" box="[945,1023,1480,1502]" gridcol="3" gridrow="17" pageId="7" pageNumber="490">?</td>
<td id="843F51425B44001E670E04A61F0FFA00" box="[1060,1284,1480,1502]" gridcol="4" gridrow="17" pageId="7" pageNumber="490">- (12.8­15.8); 2</td>
<td id="843F51425B44001E665004A61C6EFA00" box="[1402,1637,1480,1502]" gridcol="5" gridrow="17" pageId="7" pageNumber="490">17.9 ±2.73 (14.2­20.7); 4</td>
<td id="843F51425B44001E65F604A61D21FA00" box="[1756,1834,1480,1502]" gridcol="6" gridrow="17" pageId="7" pageNumber="490">16.2</td>
</tr>
<tr id="C7EE383E5B44001E6357049C1D21F9D6" box="[125,1834,1522,1544]" gridrow="18" pageId="7" pageNumber="490">
<th id="843F51425B44001E6357049C1B6BF9D6" box="[125,352,1522,1544]" gridcol="0" gridrow="18" pageId="7" pageNumber="490">Toe 6</th>
<td id="843F51425B44001E62F7049C184DF9D6" box="[477,582,1522,1544]" gridcol="1" gridrow="18" pageId="7" pageNumber="490">11.0</td>
<td id="843F51425B44001E617B049C194DF9D6" box="[593,838,1522,1544]" gridcol="2" gridrow="18" pageId="7" pageNumber="490">10.7 ±1.04 (9.0­12.1); 11</td>
<td id="843F51425B44001E609B049C19F4F9D6" box="[945,1023,1522,1544]" gridcol="3" gridrow="18" pageId="7" pageNumber="490">?</td>
<td id="843F51425B44001E670E049C1F0FF9D6" box="[1060,1284,1522,1544]" gridcol="4" gridrow="18" pageId="7" pageNumber="490">- (8.5­10.8); 2</td>
<td id="843F51425B44001E6650049C1C6EF9D6" box="[1402,1637,1522,1544]" gridcol="5" gridrow="18" pageId="7" pageNumber="490">10.2 ±1.26 (8.5­11.5); 4</td>
<td id="843F51425B44001E65F6049C1D21F9D6" box="[1756,1834,1522,1544]" gridcol="6" gridrow="18" pageId="7" pageNumber="490">?</td>
</tr>
<tr id="C7EE383E5B44001E635707751D21F9EF" box="[125,1834,1563,1585]" gridrow="19" pageId="7" pageNumber="490">
<th id="843F51425B44001E635707751B6BF9EF" box="[125,352,1563,1585]" gridcol="0" gridrow="19" pageId="7" pageNumber="490">Gonopore - Anus</th>
<td id="843F51425B44001E62F70775184DF9EF" box="[477,582,1563,1585]" gridcol="1" gridrow="19" pageId="7" pageNumber="490">10.9</td>
<td id="843F51425B44001E617B0775194DF9EF" box="[593,838,1563,1585]" gridcol="2" gridrow="19" pageId="7" pageNumber="490">8.0 ±2.07 (4.7­11.0); 14</td>
<td id="843F51425B44001E609B077519F4F9EF" box="[945,1023,1563,1585]" gridcol="3" gridrow="19" pageId="7" pageNumber="490">4.5</td>
<td id="843F51425B44001E670E07751F0FF9EF" box="[1060,1284,1563,1585]" gridcol="4" gridrow="19" pageId="7" pageNumber="490">5.5 ±0.49 (5.0­6.1); 5</td>
<td id="843F51425B44001E665007751C6EF9EF" box="[1402,1637,1563,1585]" gridcol="5" gridrow="19" pageId="7" pageNumber="490">-</td>
<td id="843F51425B44001E65F607751D21F9EF" box="[1756,1834,1563,1585]" gridcol="6" gridrow="19" pageId="7" pageNumber="490">-</td>
</tr>
</table>
</paragraph>
<paragraph id="79613A7C5B4BFFE963ED018E1B66FC32" blockId="8.[151,1437,151,1005]" pageId="8" pageNumber="491">
<taxonomicName id="BEDE41FF5B4BFFE963ED018E1BB0FF26" box="[199,443,224,248]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="8" pageNumber="491" phylum="Tardigrada" rank="species" species="brasiliensis" status="sp. nov.">
<emphasis id="4BAAE66E5B4BFFE963ED018E1BB0FF26" box="[199,443,224,248]" italics="true" pageId="8" pageNumber="491">Batillipes brasiliensis</emphasis>
</taxonomicName>
<emphasis id="4BAAE66E5B4BFFE962E801B11817FF26" bold="true" box="[450,540,223,248]" pageId="8" pageNumber="491">
<taxonomicNameLabel id="50995B155B4BFFE962E801B11817FF26" box="[450,540,223,248]" pageId="8" pageNumber="491" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in having swollen external cephalic cirri and the caudal apparatus consisting of an ala-like cuticular expansion covered with detritus is most similar to 
<taxonomicName id="BEDE41FF5B4BFFE96099006B1E12FEC2" box="[947,1049,260,284]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="8" pageNumber="491" phylum="Tardigrada" rank="species" species="lesteri">
<emphasis id="4BAAE66E5B4BFFE96099006B1E12FEC2" box="[947,1049,260,284]" italics="true" pageId="8" pageNumber="491">B. lesteri</emphasis>
</taxonomicName>
. However, 
<taxonomicName id="BEDE41FF5B4BFFE967B1006B1F30FEC2" box="[1179,1339,260,284]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="8" pageNumber="491" phylum="Tardigrada" rank="species" species="brasiliensis" status="sp. nov.">
<emphasis id="4BAAE66E5B4BFFE967B1006B1F30FEC2" box="[1179,1339,260,284]" italics="true" pageId="8" pageNumber="491">B. brasiliensis</emphasis>
</taxonomicName>
<emphasis id="4BAAE66E5B4BFFE96668006A1F97FEC3" bold="true" box="[1346,1436,260,285]" pageId="8" pageNumber="491">
<taxonomicNameLabel id="50995B155B4BFFE96668006A1F97FEC3" box="[1346,1436,260,285]" pageId="8" pageNumber="491" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
exhibits different shaped lateral projections with digit-shaped apices and the one between leg III and IV distally fringed. In addition, external cephalic cirri are horn-shaped in 
<taxonomicName id="BEDE41FF5B4BFFE96078002319BCFEBA" box="[850,951,332,356]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="8" pageNumber="491" phylum="Tardigrada" rank="species" species="lesteri">
<emphasis id="4BAAE66E5B4BFFE96078002319BCFEBA" box="[850,951,332,356]" italics="true" pageId="8" pageNumber="491">B. lesteri</emphasis>
</taxonomicName>
and terminated by a swollen tip in the new species. The two species, that have comparable body lengths, differ in the length of cirri E (the longest cirrus E in the new species is 13.1 µm long while in 
<taxonomicName id="BEDE41FF5B4BFFE9614100FB18DAFE72" box="[619,721,404,428]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="8" pageNumber="491" phylum="Tardigrada" rank="species" species="lesteri">
<emphasis id="4BAAE66E5B4BFFE9614100FB18DAFE72" box="[619,721,404,428]" italics="true" pageId="8" pageNumber="491">B. lesteri</emphasis>
</taxonomicName>
the shortest cirrus E is 19.0 µm long) and leg sense organs that are shorter in the new species. In particular, the sense organ of leg IV is much shorter in the new species (10% of the body length) than in 
<taxonomicName id="BEDE41FF5B4BFFE9629C00B31815FE2A" box="[438,542,476,500]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="8" pageNumber="491" phylum="Tardigrada" rank="species" species="lesteri">
<emphasis id="4BAAE66E5B4BFFE9629C00B31815FE2A" box="[438,542,476,500]" italics="true" pageId="8" pageNumber="491">B. lesteri</emphasis>
</taxonomicName>
where it is comprised of between 20 and 26 % of the body length. Cuticular valved structures associated with the female gonopore are present in both species, but in 
<taxonomicName id="BEDE41FF5B4BFFE96757036F1EE9FDC6" box="[1149,1250,512,536]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="8" pageNumber="491" phylum="Tardigrada" rank="species" species="lesteri">
<emphasis id="4BAAE66E5B4BFFE96757036F1EE9FDC6" box="[1149,1250,512,536]" italics="true" pageId="8" pageNumber="491">B. lesteri</emphasis>
</taxonomicName>
only two valves are present posterior to the gonopore, while there are four in the new species, two anterior and two posterior. Moreover, the new species differs from 
<taxonomicName id="BEDE41FF5B4BFFE96146032718DFFDBE" box="[620,724,584,608]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="8" pageNumber="491" phylum="Tardigrada" rank="species" species="lesteri">
<emphasis id="4BAAE66E5B4BFFE96146032718DFFDBE" box="[620,724,584,608]" italics="true" pageId="8" pageNumber="491">B. lesteri</emphasis>
</taxonomicName>
(and from all 
<taxonomicName id="BEDE41FF5B4BFFE960A9032619E4FDBE" box="[899,1007,584,608]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="8" pageNumber="491" phylum="Tardigrada" rank="genus">
<emphasis id="4BAAE66E5B4BFFE960A9032619E4FDBE" box="[899,1007,584,608]" italics="true" pageId="8" pageNumber="491">Batillipes</emphasis>
</taxonomicName>
species) by a peculiar character: the presence of small semicircular cuticular projections on the coxal region of legs I–III. This character, that cannot be confused with the coxal swelling of the leg articulation, is also present in specimens previously collected in 
<collectingCountry id="01C97AEC5B4BFFE9667203E11F97FD76" box="[1368,1436,655,680]" name="Brazil" pageId="8" pageNumber="491">Brazil</collectingCountry>
and attributed to 
<taxonomicName id="BEDE41FF5B4BFFE9627E03DB18B1FD12" authority="da Rocha et al. 2013" authorityName="da Rocha et al." authorityYear="2013" box="[340,698,692,717]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="8" pageNumber="491" phylum="Tardigrada" rank="species" species="lesteri">
<emphasis id="4BAAE66E5B4BFFE9627E03DB1BB2FD12" box="[340,441,692,716]" italics="true" pageId="8" pageNumber="491">B. lesteri</emphasis>
(da Rocha 
<emphasis id="4BAAE66E5B4BFFE9611D03DB1864FD12" box="[567,623,692,716]" italics="true" pageId="8" pageNumber="491">et al.</emphasis>
2013)
</taxonomicName>
. The re-examination of those specimens, which also have external cephalic cirri with swollen tips, led to the conclusion that they should be attributed to the new species. In those specimens mounted in glycerin, eyes were observed which seem to confirm the presence of these structures in the new species. Eyes are soluble in some mounting media, as the glycerol. The re-examination also confirms the intraspecific variability of the caudal apparatus and lateral projections located between legs III and IV that, together with the large amount of adherent detritus were the reason of misidentification. With this correction, 
<taxonomicName id="BEDE41FF5B4BFFE967D602071F97FC5E" box="[1276,1436,872,896]" class="Heterotardigrada" family="Batillipedidae" genus="Batillipes" kingdom="Animalia" order="Arthrotardigrada" pageId="8" pageNumber="491" phylum="Tardigrada" rank="species" species="brasiliensis" status="sp. nov.">
<emphasis id="4BAAE66E5B4BFFE967D602071F97FC5E" box="[1276,1436,872,896]" italics="true" pageId="8" pageNumber="491">B. brasiliensis</emphasis>
</taxonomicName>
<emphasis id="4BAAE66E5B4BFFE963BD02E21AF8FC7B" bold="true" box="[151,243,908,933]" pageId="8" pageNumber="491">
<taxonomicNameLabel id="50995B155B4BFFE963BD02E21AF8FC7B" box="[151,243,908,933]" pageId="8" pageNumber="491" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is also recorded for Cupe Beach, 
<collectingRegion id="BB1AF49E5B4BFFE961AA02E21907FC7B" box="[640,780,908,933]" country="Brazil" name="Pernambuco" pageId="8" pageNumber="491">Pernambuco</collectingRegion>
State (
<geoCoordinate id="1CEA5CBB5B4BFFE9607502E219A2FC7B" box="[863,937,908,933]" direction="south" orientation="latitude" pageId="8" pageNumber="491" precision="925" value="-8.416667">8°25'S</geoCoordinate>
, 
<geoCoordinate id="1CEA5CBB5B4BFFE9609D02E21E17FC7B" box="[951,1052,908,933]" direction="west" orientation="longitude" pageId="8" pageNumber="491" precision="925" value="-34.916668">34°55'W</geoCoordinate>
), and for the continental shelf, in medium sands at 
<quantity id="BE2697995B4BFFE9624A02C11B90FC19" box="[352,411,943,968]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="491" unit="m" value="35.0">35 m</quantity>
depth, of 
<collectingRegion id="BB1AF49E5B4BFFE9613802DE1858FC16" box="[530,595,944,968]" country="Brazil" name="Ceara" pageId="8" pageNumber="491">Ceará</collectingRegion>
and 
<collectingRegion id="BB1AF49E5B4BFFE961A402C1198BFC19" box="[654,896,943,968]" country="Brazil" name="Rio Grande do Norte" pageId="8" pageNumber="491">Rio Grande do Norte</collectingRegion>
States (4°20'– 
<geoCoordinate id="1CEA5CBB5B4BFFE9670F02C11E65FC16" box="[1061,1134,943,968]" direction="south" orientation="latitude" pageId="8" pageNumber="491" precision="925" value="-4.8333335">4°50'S</geoCoordinate>
; 36°00– 
<geoCoordinate id="1CEA5CBB5B4BFFE967E702C11F3DFC16" box="[1229,1334,943,968]" direction="west" orientation="longitude" pageId="8" pageNumber="491" precision="925" value="-37.25">37°15' W</geoCoordinate>
), (see da Rocha 
<emphasis id="4BAAE66E5B4BFFE963CF02BB1B16FC32" box="[229,285,980,1004]" italics="true" pageId="8" pageNumber="491">et al.</emphasis>
2013).
</paragraph>
</subSubSection>
</treatment>
</document>