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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1180.109099" ID-Pensoft-Pub="1313-2970-1180-51" ID-Pensoft-UUID="E89B8C436A615CAD8308264539B52F5A" ID-ZooBank="711DA3A4B9AC49F8AAA40682823DED14" ModsDocID="1313-2970-1180-51" checkinTime="1694791488818" checkinUser="pensoft" docAuthor="Joharchi, Omid, Keum, Seoyoung &amp; Jung, Chuleui" docDate="2023" docId="EA0D6D37B97456BE91378746BB42DFB4" docLanguage="en" docName="ZooKeys 1180: 51-65" docOrigin="ZooKeys 1180" docPubDate="2023-09-15" docSource="http://dx.doi.org/10.3897/zookeys.1180.109099" docTitle="Julolaelaps gigas Joharchi &amp; Keum &amp; Jung 2023, sp. nov." docType="treatment" docUuid="FDD68D96-90DF-4196-9DD5-377DCF03D633" docUuidSource="ZooBank" docVersion="1" id="E89B8C436A615CAD8308264539B52F5A" lastPageNumber="51" masterDocId="E89B8C436A615CAD8308264539B52F5A" masterDocTitle="The first record of the genus Julolaelaps Berlese (Acari, Mesostigmata, Laelapidae) in Republic of Korea and description of a new species on a captive giant African millipede (Spirostreptidae, Archispirostreptus)" masterLastPageNumber="65" masterPageNumber="51" pageNumber="51" updateTime="1694791488818" updateUser="pensoft">
<mods:mods id="4BB8E5404E8BBE9043408B908C68CEA6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A8190F22775679C76C5E117E4D707F2B">
<mods:title id="4A9D16651E28BB6CE2456240FC59C4ED">The first record of the genus Julolaelaps Berlese (Acari, Mesostigmata, Laelapidae) in Republic of Korea and description of a new species on a captive giant African millipede (Spirostreptidae, Archispirostreptus)</mods:title>
</mods:titleInfo>
<mods:name id="B5570C977EA53D31BE06AC1DE181231F" type="personal">
<mods:role id="49EBCD757E3EC0B9EF05372802EB2F05">
<mods:roleTerm id="D6CA195FE359C4273BBE07F95C922D22">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E7EFAF485E0305704586D7A1065CB665">Joharchi, Omid</mods:namePart>
<mods:nameIdentifier id="479C97E5AA4E7DB31FBEB431A0EA1932" type="ORCID">https://orcid.org/0000-0002-2741-4946</mods:nameIdentifier>
<mods:affiliation id="85A3910828D89D31D950EE607AE2FD8F">Agriculture Science and Technology Institute, Andong National University, Andong, Republic of Korea &amp; Institute of Environmental and Agricultural Biology (X-BIO), Tyumen State University, Tyumen, Russia</mods:affiliation>
</mods:name>
<mods:name id="CE24AD73A62E5796AD5A3240C080CEB9" type="personal">
<mods:role id="D2EBCA8BADF2FFB56E6BCA07AF838FF0">
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</mods:role>
<mods:namePart id="243A204B8A0983E9CE002083B926CCB6">Keum, Seoyoung</mods:namePart>
<mods:nameIdentifier id="E4669A6130399321888758F35754BEC7" type="ORCID">https://orcid.org/0000-0002-2219-9423</mods:nameIdentifier>
<mods:affiliation id="B29351EA9C0C23C7A7A2AC4594FA7199">Silkworm and Insect Management Center, Sangju, Republic of Korea</mods:affiliation>
</mods:name>
<mods:name id="2A7B3ADA624BEDCA72319604EB30741A" type="personal">
<mods:role id="26F19A524DD71DDB9034C046DB0DAA99">
<mods:roleTerm id="320FA807A4C7131BA7BE8D19DF059134">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1218AE64B9F24981CF792FE69A37CBAD">Jung, Chuleui</mods:namePart>
<mods:nameIdentifier id="EA5B8A1E95F8E1452232491851FDEDA8" type="ORCID">https://orcid.org/0000-0001-8134-9279</mods:nameIdentifier>
<mods:affiliation id="31C50737F74F32D5ED47279F363F196D">Agriculture Science and Technology Institute, Andong National University, Andong, Republic of Korea</mods:affiliation>
<mods:nameIdentifier id="92FF9E9DA5028623A9E6ED383D73EEA2" type="email">cjung@andong.ac.kr</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="A44084F7140C53562F5DD385614945A4">text</mods:typeOfResource>
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<mods:title id="E09E932C7BE3CF326C8BFF7539838D15">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="DFC7585B29B2C79B3C61A623542C620B">
<mods:date id="F3ABB1F88CB29949B16A07F6DC6E6B48">2023</mods:date>
<mods:detail id="F5DCC75AE9D2A4F271D34359FEA5B4A7" type="pubDate">
<mods:number id="E4F8F9092B27F4A1CF2920A52528C59F">2023-09-15</mods:number>
</mods:detail>
<mods:detail id="74BC8FB8E663D6D2C0BCBF8ABF175F71" type="volume">
<mods:number id="5A3D6CD6DCB2CF8890A95477F8BEC427">1180</mods:number>
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<mods:start id="86E2F84FD93B5C373F6D8872F87A4709">51</mods:start>
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<mods:url id="743495290422302F7CE4F1F8CE17E48E">http://dx.doi.org/10.3897/zookeys.1180.109099</mods:url>
</mods:location>
<mods:classification id="AF1C89592033D42E8E7062695DC5657B">journal article</mods:classification>
<mods:identifier id="393D2899FDD0FBB9DA1A16B8AE1193E5" type="DOI">http://dx.doi.org/10.3897/zookeys.1180.109099</mods:identifier>
<mods:identifier id="61C8DBD9EE095F5AB00627FE86417FB3" type="Pensoft-Pub">1313-2970-1180-51</mods:identifier>
<mods:identifier id="33EF36D050AA137FB02D867011161B46" type="ZooBank">711DA3A4B9AC49F8AAA40682823DED14</mods:identifier>
<mods:identifier id="6673D53ABDBAE9448F017C1E2E9F872D" type="Pensoft-UUID">E89B8C436A615CAD8308264539B52F5A</mods:identifier>
</mods:mods>
<subSection id="AEC7E7EAE685C21BA8E4E4A8908BE623" pageId="0" pageNumber="51" type="results">
<treatment id="EA0D6D37B97456BE91378746BB42DFB4" LSID="urn:lsid:zoobank.org:act:FDD68D96-90DF-4196-9DD5-377DCF03D633" httpUri="http://treatment.plazi.org/id/EA0D6D37B97456BE91378746BB42DFB4" lastPageNumber="51" pageId="0" pageNumber="51">
<subSubSection id="727BB24FEFBF314790A2E663241A2F89" pageId="0" pageNumber="51" type="nomenclature">
<paragraph id="0254EF535A57315BD56448F03D2F6BDC" pageId="0" pageNumber="51">
<taxonomicName id="D1A6420C6B83C17A14D0B6BC77FCC460" LSID="https://zoobank.org/FDD68D96-90DF-4196-9DD5-377DCF03D633" authority="Joharchi &amp; Keum &amp; Jung, 2023" authorityName="Joharchi &amp; Keum &amp; Jung" authorityYear="2023" class="Diplopoda" family="Laelapidae" genus="Julolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Julolaelaps gigas" order="Mesostigmata" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="gigas" status="sp. nov.">Julolaelaps gigas</taxonomicName>
<taxonomicNameLabel id="9EB3E2C0B86560A7BC14F5F5AD40844B" pageId="0" pageNumber="51">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="55DFA3BCD32F2A0CFA598BD8A792EADF" pageId="0" pageNumber="51" type="description">
<paragraph id="F044A60D3CE283BDC6E82EDD992618DC" pageId="0" pageNumber="51">
<figureCitation id="7E088A47544C7E6ED6FC8EFE23597FB7" captionStart="Figures 1–5" captionStartId="F1" captionText="Figures 1 - 5. Julolaelaps gigas sp. nov., female, with symbols for chaetotactic notation of idiosomal setae 1 dorsal idiosoma 2 ventral idiosoma 3 subcapitulum 4 epistome 5 chelicera, lateral view." figureDoi="10.3897/zookeys.1180.109099.figures1-5" httpUri="https://binary.pensoft.net/fig/906333" pageId="0" pageNumber="51">Figs 1-5</figureCitation>
<figureCitation id="8B87C984439CA33B14D23EED6C3BC713" captionStart="Figures 6–9" captionStartId="F2" captionText="Figures 6 - 9. Julolaelaps gigas sp. nov., female, with symbols for chaetotactic notation of legs setae 6 leg I (trochanter-tibia, dorsal aspect) 7 leg II (trochanter-tibia, dorsal aspect) 8 leg III (trochanter-tarsus, dorsal aspect) 9 leg IV (trochanter-tibia, dorsal aspect)." figureDoi="10.3897/zookeys.1180.109099.figures6-9" httpUri="https://binary.pensoft.net/fig/906334" pageId="0" pageNumber="51">, 6-9</figureCitation>
<figureCitation id="FB73ECA15A0C891E2055446BA4C93E90" captionStart="Figures 10, 11" captionStartId="F3" captionText="Figures 10, 11. Julolaelaps gigas sp. nov., male 10 ventral idiosoma 11 chelicera, lateral view." figureDoi="10.3897/zookeys.1180.109099.figures10-11" httpUri="https://binary.pensoft.net/fig/906335" pageId="0" pageNumber="51">, 10, 11</figureCitation>
<figureCitation id="FB2945B89E8923D501B0A02D1949194F" captionStart="Figures 12–16" captionStartId="F4" captionText="Figures 12 - 16. Differential Interference Contrast (DIC) micrographs of Julolaelaps gigas sp. nov., female 12 idiosoma in dorsal view 13 idiosoma in ventral view 14 chelicera in lateral view, male 15 idiosoma in ventral view 16 chelicera in lateral view." figureDoi="10.3897/zookeys.1180.109099.figures12-16" httpUri="https://binary.pensoft.net/fig/906336" pageId="0" pageNumber="51">, 12-16</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="9448FBEA326E0A02E4F5C716EDA74A36" pageId="0" pageNumber="51" type="materials_examined">
<paragraph id="4AA83E32253F86904B0E00327852C587" pageId="0" pageNumber="51">Material examined.</paragraph>
<paragraph id="95E487BF9DA61E78F64BB3D4364B104A" pageId="0" pageNumber="51">
<materialsCitation id="7AE0B7860EC10837985496A3A06AF806" collectingDate="2020-05-19" collectorName="S. Keum, African" country="Republic of Korea" county="Insect Management Center" latitude="36.95" location="Sangju" longLatPrecision="1184" longitude="128.25" municipality="Silkworm" specimenCount="♀" typeStatus="Holotype">
<emphasis id="647EF1DFA3FB4F1A6D841E83E6A4C93D" bold="true" pageId="0" pageNumber="51">
<emphasis id="E2C446C960D2F26C813FC59C0634D44A" italics="true" pageId="0" pageNumber="51">
<typeStatus id="91D5F5454CEAF61D8E237138241CD175">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="9802F6C1F5D5FA8D8036A41D90A0118E" name="South Korea">Republic of Korea</collectingCountry>
•
<specimenCount id="1B9A94C50C5EF5F1878441C5F903E6C9">♀</specimenCount>
;
<location id="528280BAEDA1E761727EFA4A52F85046" LSID="urn:lsid:plazi:treatment:EA0D6D37B97456BE91378746BB42DFB4:528280BAEDA1E761727EFA4A52F85046" country="Republic of Korea" county="Insect Management Center" latitude="36.95" longLatPrecision="1184" longitude="128.25" municipality="Silkworm" name="Sangju">Sangju</location>
,
<collectingMunicipality id="5E0FE359B846C04F4DE1BA8DFFD47C91">Silkworm</collectingMunicipality>
and
<collectingCounty id="F76257ECD389AAB96BDBFCE7FEBD6128">Insect Management Center</collectingCounty>
;
<geoCoordinate id="64880EA9EC77A69BC33366DA27FDC33C" degrees="36" direction="north" minutes="57" orientation="latitude" precision="925" value="36.95">36°57'N</geoCoordinate>
,
<geoCoordinate id="F3860A12ACB825F153D001678DC88006" degrees="128" direction="east" minutes="15" orientation="longitude" precision="925" value="128.25">128°15'E</geoCoordinate>
;
<collectingDate id="27FC2AD6EE15E8CEE465B40F5FEF732B" value="2020-05-19">19 May 2020</collectingDate>
;
<collectorName id="3FF359FC3CB2F80673874D157F4E21CE">S. Keum</collectorName>
leg.; on captive giant
<collectorName id="1F69772E386867A0E6253DE2C53B4993">African</collectorName>
millipede,
<taxonomicName id="5E4730BF966DD2376AAC884543B6CD92" authorityName="Peters" authorityYear="1855" baseAuthorityName="Peters" baseAuthorityYear="1855" class="Diplopoda" family="Spirostreptidae" genus="Archispirostreptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archispirostreptus gigas" order="Spirostreptida" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="gigas">
<emphasis id="6820A0868190183399D5171DD3587FBF" italics="true" pageId="0" pageNumber="51">Archispirostreptus gigas</emphasis>
</taxonomicName>
(
<taxonomicName id="24001A124572E6B0869F204F7FF51092" authorityName="Silvestri" authorityYear="1895" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Spirostreptida" pageId="0" pageNumber="51" phylum="Arthropoda" rank="order">Spirostreptida</taxonomicName>
:
<taxonomicName id="2832FB9928D6690105CF369C0ED04903" class="Diplopoda" family="Spirostreptidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Spirostreptida" pageId="0" pageNumber="51" phylum="Arthropoda" rank="family">Spirostreptidae</taxonomicName>
); NIBRIV 0000905722
</materialsCitation>
.
<materialsCitation id="A8E12882EC258E6468CF9C59A7B5590E" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="S. Keum &amp; African" country="Republic of Korea" county="Insect Management Center" latitude="36.95" location="Republic of Korea" longLatPrecision="1184" longitude="128.25" municipality="Silkworm" specimenCount="4" specimenCount-female="2" specimenCount-male="2" typeStatus="Paratypes">
<emphasis id="CF0BA034D480FD4B813709345EFD157D" bold="true" pageId="0" pageNumber="51">
<emphasis id="F03B5901F27FC2EC62061B606CF39EB6" italics="true" pageId="0" pageNumber="51">
<typeStatus id="CDBAF054A52440013AE35FBB0139CFB9">Paratypes</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="3C4AC270E1487BA79D7DB1620130D75C" name="South Korea">Republic of Korea</collectingCountry>
•
<specimenCount id="AD754FD6F2E4FDBAB2AA64C627E89495" type="female">2♀</specimenCount>
<specimenCount id="A0862E96F1D809FE4B8B989644165E5B" type="male">2♂</specimenCount>
; same data as for holotype; NIBRIV 0000905723, NIBRIV 0000905724, NIBRIV 0000905725, NIBRIV 0000905726
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="7770FAF1A3ACCBAA0BB264155BEC4CCD" pageId="0" pageNumber="51" type="diagnosis">
<paragraph id="D0017E7EF7583552BEC99A0A23494BAC" pageId="0" pageNumber="51">Diagnosis.</paragraph>
<paragraph id="3F6B468278B77174B119D9051781E70F" pageId="0" pageNumber="51">
Dorsal shield smooth, without reticulation, bearing 22 pairs of minute acicular setae, setae
<emphasis id="16286C8120FF55D0C89077F26DD042E8" italics="true" pageId="0" pageNumber="51">Z5</emphasis>
conspicuously longer than other setae on shield, podonotal and opisthonotal parts of dorsal shield slightly hypotrichous, setae in
<emphasis id="C964DC69B7B312562F44F9539C254FEB" italics="true" pageId="0" pageNumber="51">z</emphasis>
- and
<emphasis id="080B8662A5DB5F314813FB37E4479E79" italics="true" pageId="0" pageNumber="51">s/S-series</emphasis>
displaced laterally (on soft cuticle) and some missing (
<emphasis id="F9DB745D57272ECA2BC8C712932CFF62" italics="true" pageId="0" pageNumber="51">z1</emphasis>
, z6,
<emphasis id="3B77C086E24510D83EC20535C00A9739" italics="true" pageId="0" pageNumber="51">J3</emphasis>
); posterior margin of shield somewhat truncated; dorsal shield reduced, not covering entire idiosoma, opisthogastric and lateral soft cuticle with 37 pairs of setae, including 23 pairs of
<emphasis id="D6A040DCD7EB3CD89F8022C0A0AA2634" italics="true" pageId="0" pageNumber="51">r</emphasis>
-
<emphasis id="6A0F3B68C653EB8826932C43DAEECBD7" italics="true" pageId="0" pageNumber="51">R</emphasis>
-
<emphasis id="F09C9DFA352F403770D93A6B95A47AD6" italics="true" pageId="0" pageNumber="51">UR</emphasis>
setae. Sternal shield reduced, bearing only
<emphasis id="1BAD035E9C548411F50811736D0CA6FD" italics="true" pageId="0" pageNumber="51">st1</emphasis>
and
<emphasis id="0111475D1DA3DEB675DE3F3DD9B07400" italics="true" pageId="0" pageNumber="51">st2</emphasis>
, eroded posteriorly,
<emphasis id="F0CB8A1698E808C13A86D73A0A70D185" italics="true" pageId="0" pageNumber="51">st3</emphasis>
and metasternal setae (
<emphasis id="A3684A7006C1AE06570BE68CACEB7DCC" italics="true" pageId="0" pageNumber="51">st4</emphasis>
) inserted on unsclerotized cuticle, epigynal shield drop-shaped, parapodal plates broadly thickened, extending behind coxae IV, endopodal plates III/IV broadly thickened, peritreme relatively long and rather broad, extending to anterior level of coxa II, cheliceral fixed digit with 2-3 teeth. Chaetotaxy of legs normal for free-living
<taxonomicName id="95EDAEBEE5BC56B9F6E3DE14A19FDDC1" authorityName="A.Berlese" authorityYear="1892" class="Arachnida" family="Laelapidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Mesostigmata" pageId="0" pageNumber="51" phylum="Arthropoda" rank="family">Laelapidae</taxonomicName>
, except tibia III with nine setae (two posterolateral setae present) and genu IV with 10 setae (with two posterolateral setae). Male with separate sternogenital and anal shields, chelicerae with edentate digits including a regressed fixed digit and a stylet-like movable digit bordered by a very long spermatodactyl.
</paragraph>
<caption id="205FE6C1C5A47100EA568507FAC4FF5F" doi="10.3897/zookeys.1180.109099.figures1-5" httpUri="https://binary.pensoft.net/fig/906333" pageId="0" pageNumber="51" start="Figures 1–5" startId="F1">
<paragraph id="A91269E417F03C06FC462C1A7C1198DA" pageId="0" pageNumber="51">
<emphasis id="1A2672104319C39AF93B24878187AF42" bold="true" pageId="0" pageNumber="51">Figures 1-5.</emphasis>
<taxonomicName id="CF7D6E0319EFA817831B743D3F459138" authorityName="Joharchi &amp; Keum &amp; Jung" authorityYear="2023" class="Diplopoda" family="Laelapidae" genus="Julolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Julolaelaps gigas" order="Mesostigmata" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="gigas">
<emphasis id="A7D5E64717FBA1CE81C54938ABAC7077" italics="true" pageId="0" pageNumber="51">Julolaelaps gigas</emphasis>
</taxonomicName>
sp. nov., female, with symbols for chaetotactic notation of idiosomal setae
<emphasis id="5FDD75DA24A8530DF5D936417141BD6E" bold="true" pageId="0" pageNumber="51">1</emphasis>
dorsal idiosoma
<emphasis id="AF2083279C9AC92A5D11A2C422D20F8A" bold="true" pageId="0" pageNumber="51">2</emphasis>
ventral idiosoma
<emphasis id="A615ADBE1A2ED1A41DA342B24B39FBC7" bold="true" pageId="0" pageNumber="51">3</emphasis>
subcapitulum
<emphasis id="6A788378427516096EE525416A442D3B" bold="true" pageId="0" pageNumber="51">4</emphasis>
epistome
<emphasis id="526D0FE7F97275E9ED4A9CC4DB5A87F7" bold="true" pageId="0" pageNumber="51">5</emphasis>
chelicera, lateral view.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="C1375A50B8D725BFB7C26202A6F3F2BD" pageId="0" pageNumber="51" type="description">
<paragraph id="E9A8D1584FDDC53CDC55106A66872900" pageId="0" pageNumber="51">Description.</paragraph>
<paragraph id="CF37EF83FB81980EC9E6ACC79C5617EB" pageId="0" pageNumber="51">
<emphasis id="4AF8FAC0ACA1957D6D95B368EAA92663" bold="true" pageId="0" pageNumber="51">Female</emphasis>
(
<emphasis id="8DDC0098BFC2747E5A321DE4A87A0D38" italics="true" pageId="0" pageNumber="51">N</emphasis>
= 3). Figs
<figureCitation id="95F217B96072330DD8A6EBDC6B8F9211" captionStart="Figures 1–5" captionStartId="F1" captionText="Figures 1 - 5. Julolaelaps gigas sp. nov., female, with symbols for chaetotactic notation of idiosomal setae 1 dorsal idiosoma 2 ventral idiosoma 3 subcapitulum 4 epistome 5 chelicera, lateral view." figureDoi="10.3897/zookeys.1180.109099.figures1-5" httpUri="https://binary.pensoft.net/fig/906333" pageId="0" pageNumber="51">1</figureCitation>
-
<figureCitation id="50A17EFF50A2D8F10864FD6F42808CBB" captionStart="Figures 6–9" captionStartId="F2" captionText="Figures 6 - 9. Julolaelaps gigas sp. nov., female, with symbols for chaetotactic notation of legs setae 6 leg I (trochanter-tibia, dorsal aspect) 7 leg II (trochanter-tibia, dorsal aspect) 8 leg III (trochanter-tarsus, dorsal aspect) 9 leg IV (trochanter-tibia, dorsal aspect)." figureDoi="10.3897/zookeys.1180.109099.figures6-9" httpUri="https://binary.pensoft.net/fig/906334" pageId="0" pageNumber="51">9</figureCitation>
,
<figureCitation id="AFFE692FD25B5F3647267945833A1BE0" captionStart="Figures 12–16" captionStartId="F4" captionText="Figures 12 - 16. Differential Interference Contrast (DIC) micrographs of Julolaelaps gigas sp. nov., female 12 idiosoma in dorsal view 13 idiosoma in ventral view 14 chelicera in lateral view, male 15 idiosoma in ventral view 16 chelicera in lateral view." figureDoi="10.3897/zookeys.1180.109099.figures12-16" httpUri="https://binary.pensoft.net/fig/906336" pageId="0" pageNumber="51">12-14</figureCitation>
.
<emphasis id="CDB723F900ABB70EEF988B5A44F1D35A" bold="true" italics="true" pageId="0" pageNumber="51">Dorsal idiosoma</emphasis>
(Figs
<figureCitation id="BB9A0F33E256FD7C98104986C504AF01" captionStart="Figures 1–5" captionStartId="F1" captionText="Figures 1 - 5. Julolaelaps gigas sp. nov., female, with symbols for chaetotactic notation of idiosomal setae 1 dorsal idiosoma 2 ventral idiosoma 3 subcapitulum 4 epistome 5 chelicera, lateral view." figureDoi="10.3897/zookeys.1180.109099.figures1-5" httpUri="https://binary.pensoft.net/fig/906333" pageId="0" pageNumber="51">1</figureCitation>
,
<figureCitation id="C978E4FCD566B96F10E0FF104F52BEB7" captionStart="Figures 12–16" captionStartId="F4" captionText="Figures 12 - 16. Differential Interference Contrast (DIC) micrographs of Julolaelaps gigas sp. nov., female 12 idiosoma in dorsal view 13 idiosoma in ventral view 14 chelicera in lateral view, male 15 idiosoma in ventral view 16 chelicera in lateral view." figureDoi="10.3897/zookeys.1180.109099.figures12-16" httpUri="https://binary.pensoft.net/fig/906336" pageId="0" pageNumber="51">12</figureCitation>
). Length 906-925, width 620-649. Dorsal shield length 687-696, width 391-395, posteriorly somewhat truncate, reduced, not covering entire idiosoma, some dorsal setae left in the soft cuticle lateral to shield, becoming hypertrichous cuticle surrounding dorsal shield, without distinct reticulate ornamentation over whole surface; with 22 pairs of minute acicular setae, 13 pairs of podonotal setae, plus nine pairs of setae on lateral soft cuticle (
<emphasis id="4CB48518B81EC9F323382CC677BDCEF0" italics="true" pageId="0" pageNumber="51">s1</emphasis>
,
<emphasis id="CF3EF297DBE6FC74E962CCC33CC30665" italics="true" pageId="0" pageNumber="51">s2</emphasis>
,
<emphasis id="11FE21D64D358121CAF259DAE5F30320" italics="true" pageId="0" pageNumber="51">s6</emphasis>
,
<emphasis id="284FC67ED8E82F0654B98D43BB528384" italics="true" pageId="0" pageNumber="51">r1-6</emphasis>
), nine pairs of opisthonotal setae, plus 21 pairs of setae on lateral soft cuticle (including
<emphasis id="0DAC1612A7DEE9630D2D535FF03F57B4" italics="true" pageId="0" pageNumber="51">S1-5</emphasis>
and
<emphasis id="EE2E8EE0A5C483F01719F9F904EBBCEE" italics="true" pageId="0" pageNumber="51">R1-7</emphasis>
);
<emphasis id="FED5425AC66FA5521CB2BDA963C83984" italics="true" pageId="0" pageNumber="51">Zx</emphasis>
, unpaired or asymmetrical setae and
<emphasis id="AECD391DA32F2FFC6D4627498A996D70" italics="true" pageId="0" pageNumber="51">z1</emphasis>
,
<emphasis id="2F224640D7EDAB2470411DA1A0F810BF" italics="true" pageId="0" pageNumber="51">z6</emphasis>
,
<emphasis id="CA97069DFE0CFEA3902F2ABDF8858650" italics="true" pageId="0" pageNumber="51">J3</emphasis>
absent, setae similar in length (9-15) and thicknesses; clonal setae
<emphasis id="4B86D834215595DF88EE37889510FE98" italics="true" pageId="0" pageNumber="51">Z5</emphasis>
(26-31) longer than other setae on shield. Muscle insertions prominent as desclerotised circular patches. Dorsal shield with 22 pairs of discernible pore-like structures, including only five pairs of gland openings (
<emphasis id="BA0C4CE3116AA709944EEBA88566236C" italics="true" pageId="0" pageNumber="51">gd1</emphasis>
,
<emphasis id="10EEFB070088ED6448BEF2B587A8AB0D" italics="true" pageId="0" pageNumber="51">gd4</emphasis>
,
<emphasis id="23D895CA769A9E501DCAA3402E47FDE6" italics="true" pageId="0" pageNumber="51">gd6</emphasis>
,
<emphasis id="D606E62ABDD6C24C728EE74B7D094FC2" italics="true" pageId="0" pageNumber="51">gd8</emphasis>
,
<emphasis id="EC6E95019473643DA25DD128BFBC2A6A" italics="true" pageId="0" pageNumber="51">gd9</emphasis>
) and 17 pairs of poroids. Shape, position and relative length and thicknesses of setae shown in Figs
<figureCitation id="3AF0B59E1D313A952C3DFADA2E736CED" captionStart="Figures 1–5" captionStartId="F1" captionText="Figures 1 - 5. Julolaelaps gigas sp. nov., female, with symbols for chaetotactic notation of idiosomal setae 1 dorsal idiosoma 2 ventral idiosoma 3 subcapitulum 4 epistome 5 chelicera, lateral view." figureDoi="10.3897/zookeys.1180.109099.figures1-5" httpUri="https://binary.pensoft.net/fig/906333" pageId="0" pageNumber="51">1</figureCitation>
,
<figureCitation id="A82DA7B1BEB487373200A96FCD773006" captionStart="Figures 12–16" captionStartId="F4" captionText="Figures 12 - 16. Differential Interference Contrast (DIC) micrographs of Julolaelaps gigas sp. nov., female 12 idiosoma in dorsal view 13 idiosoma in ventral view 14 chelicera in lateral view, male 15 idiosoma in ventral view 16 chelicera in lateral view." figureDoi="10.3897/zookeys.1180.109099.figures12-16" httpUri="https://binary.pensoft.net/fig/906336" pageId="0" pageNumber="51">12</figureCitation>
.
</paragraph>
<paragraph id="880D3699E002EC2C9C14B0F7D944D55C" pageId="0" pageNumber="51">
<emphasis id="1907D35D368EDC92FF564102F940C496" bold="true" italics="true" pageId="0" pageNumber="51">Ventral idiosoma</emphasis>
(Figs
<figureCitation id="A0A378F49D4CAA3420E4FD972A52BBC3" captionStart="Figures 1–5" captionStartId="F1" captionText="Figures 1 - 5. Julolaelaps gigas sp. nov., female, with symbols for chaetotactic notation of idiosomal setae 1 dorsal idiosoma 2 ventral idiosoma 3 subcapitulum 4 epistome 5 chelicera, lateral view." figureDoi="10.3897/zookeys.1180.109099.figures1-5" httpUri="https://binary.pensoft.net/fig/906333" pageId="0" pageNumber="51">2</figureCitation>
,
<figureCitation id="CF4674BFF08E6A7FB0EC60A2A849C1B0" captionStart="Figures 12–16" captionStartId="F4" captionText="Figures 12 - 16. Differential Interference Contrast (DIC) micrographs of Julolaelaps gigas sp. nov., female 12 idiosoma in dorsal view 13 idiosoma in ventral view 14 chelicera in lateral view, male 15 idiosoma in ventral view 16 chelicera in lateral view." figureDoi="10.3897/zookeys.1180.109099.figures12-16" httpUri="https://binary.pensoft.net/fig/906336" pageId="0" pageNumber="51">13</figureCitation>
). Tritosternum with paired pilose laciniae (105-113), short base 30-35
<normalizedToken id="41EEC0075B82F18824EC6EC1791DAC15" originalValue="×">x</normalizedToken>
23-27 wide; presternal platelets absent. Sternal shield reduced (59-63
<normalizedToken id="39969D45ED9DA324F001E92319E510CD" originalValue="×">x</normalizedToken>
134-142 wide), shield with two pairs of smooth sternal setae [
<emphasis id="D0153117AA29BE8D44030E05961EF594" italics="true" pageId="0" pageNumber="51">st1</emphasis>
,
<emphasis id="FE6E69F901F406F45F2BC83EB8A2FB3F" italics="true" pageId="0" pageNumber="51">st2</emphasis>
(20-26)], and two pairs of poroids (
<emphasis id="7E078696343FB3DB4A9EF014BB6D599E" italics="true" pageId="0" pageNumber="51">iv1</emphasis>
,
<emphasis id="1683B30A26522203DFC20B64A936B4F2" italics="true" pageId="0" pageNumber="51">iv2</emphasis>
), eroded posteriorly, surface without reticulate ornamentation, smooth,
<emphasis id="0C728419E1A1DBD30CAB48C3B0EA2BB6" italics="true" pageId="0" pageNumber="51">st3</emphasis>
(30-32) and metasternal setae (
<emphasis id="1A1A3B687A62E0FBA4C4B527166BC855" italics="true" pageId="0" pageNumber="51">st4</emphasis>
, 29-31) inserted on soft integument, metasternal poroids (
<emphasis id="C6D84A070C99908BB67593F968124747" italics="true" pageId="0" pageNumber="51">iv3</emphasis>
) apparently absent (Figs
<figureCitation id="BFEDDE5B18CFB29905D02D3E54632637" captionStart="Figures 1–5" captionStartId="F1" captionText="Figures 1 - 5. Julolaelaps gigas sp. nov., female, with symbols for chaetotactic notation of idiosomal setae 1 dorsal idiosoma 2 ventral idiosoma 3 subcapitulum 4 epistome 5 chelicera, lateral view." figureDoi="10.3897/zookeys.1180.109099.figures1-5" httpUri="https://binary.pensoft.net/fig/906333" pageId="0" pageNumber="51">2</figureCitation>
,
<figureCitation id="8201219CB003E7C77C9E70BAC5820E36" captionStart="Figures 12–16" captionStartId="F4" captionText="Figures 12 - 16. Differential Interference Contrast (DIC) micrographs of Julolaelaps gigas sp. nov., female 12 idiosoma in dorsal view 13 idiosoma in ventral view 14 chelicera in lateral view, male 15 idiosoma in ventral view 16 chelicera in lateral view." figureDoi="10.3897/zookeys.1180.109099.figures12-16" httpUri="https://binary.pensoft.net/fig/906336" pageId="0" pageNumber="51">13</figureCitation>
), shield fused anterolaterally to narrow endopodal strip between coxae I and II, endopodal plates III/IV broadly thickened (Figs
<figureCitation id="96BE488F8C877658585A00473A5F3039" captionStart="Figures 1–5" captionStartId="F1" captionText="Figures 1 - 5. Julolaelaps gigas sp. nov., female, with symbols for chaetotactic notation of idiosomal setae 1 dorsal idiosoma 2 ventral idiosoma 3 subcapitulum 4 epistome 5 chelicera, lateral view." figureDoi="10.3897/zookeys.1180.109099.figures1-5" httpUri="https://binary.pensoft.net/fig/906333" pageId="0" pageNumber="51">2</figureCitation>
,
<figureCitation id="E63DC22E6BD8058F9EEA8EBBAFAC1854" captionStart="Figures 12–16" captionStartId="F4" captionText="Figures 12 - 16. Differential Interference Contrast (DIC) micrographs of Julolaelaps gigas sp. nov., female 12 idiosoma in dorsal view 13 idiosoma in ventral view 14 chelicera in lateral view, male 15 idiosoma in ventral view 16 chelicera in lateral view." figureDoi="10.3897/zookeys.1180.109099.figures12-16" httpUri="https://binary.pensoft.net/fig/906336" pageId="0" pageNumber="51">13</figureCitation>
). Epigynal shield drop-shaped, with an obvious (narrow) neck at level of coxae IV, protruding at level between setae
<emphasis id="20379E55BBB05996EB350404CD06D170" italics="true" pageId="0" pageNumber="51">st5</emphasis>
and
<emphasis id="0D27BDB4F2CE8B22F2CE827E74F1F073" italics="true" pageId="0" pageNumber="51">Jv1</emphasis>
, width (65-70) and length (228-235), anterior and posterior margins rounded, with some irregular longitudinal and oblique lines, otherwise relatively smooth, bearing a pair of simple setae
<emphasis id="ABD47B9EBC42878E3E683C85488E431A" italics="true" pageId="0" pageNumber="51">st5</emphasis>
(20-23) inserted on lateral margins of shield, near level of posterior edge of coxae IV; paragenital poroids
<emphasis id="5BF24970722CDF346EC304D72A1E6BFB" italics="true" pageId="0" pageNumber="51">iv5</emphasis>
located on soft cuticle lateral to shield near seta
<emphasis id="FACE22A5434909221041A74B4BCFAA67" italics="true" pageId="0" pageNumber="51">st5</emphasis>
(Figs
<figureCitation id="C335D7DBC4B54C8A6F5C62E4E24F6A29" captionStart="Figures 1–5" captionStartId="F1" captionText="Figures 1 - 5. Julolaelaps gigas sp. nov., female, with symbols for chaetotactic notation of idiosomal setae 1 dorsal idiosoma 2 ventral idiosoma 3 subcapitulum 4 epistome 5 chelicera, lateral view." figureDoi="10.3897/zookeys.1180.109099.figures1-5" httpUri="https://binary.pensoft.net/fig/906333" pageId="0" pageNumber="51">2</figureCitation>
,
<figureCitation id="F60BEE539FC0718018AA515A2533FC5A" captionStart="Figures 12–16" captionStartId="F4" captionText="Figures 12 - 16. Differential Interference Contrast (DIC) micrographs of Julolaelaps gigas sp. nov., female 12 idiosoma in dorsal view 13 idiosoma in ventral view 14 chelicera in lateral view, male 15 idiosoma in ventral view 16 chelicera in lateral view." figureDoi="10.3897/zookeys.1180.109099.figures12-16" httpUri="https://binary.pensoft.net/fig/906336" pageId="0" pageNumber="51">13</figureCitation>
). Anal shield subtriangular, rounded anteriorly, length 133-138, width 108-114, surface without reticulate ornamentation, smooth, para-anal setae (23-27) shorter than post-anal seta (30-35), cribrum well developed, with 3-4 irregular rows of spicules, each extending from cribrum to near base of post-anal setae; anal opening located at anterior level of shield; pair of glands
<emphasis id="CD3E38BD81BC33E3AF212382E12E07CC" italics="true" pageId="0" pageNumber="51">gv3</emphasis>
inserted on shield lateral margins, at level para-anal setae (Figs
<figureCitation id="FF105D4E721D34918BCB5D4A828FD4B6" captionStart="Figures 1–5" captionStartId="F1" captionText="Figures 1 - 5. Julolaelaps gigas sp. nov., female, with symbols for chaetotactic notation of idiosomal setae 1 dorsal idiosoma 2 ventral idiosoma 3 subcapitulum 4 epistome 5 chelicera, lateral view." figureDoi="10.3897/zookeys.1180.109099.figures1-5" httpUri="https://binary.pensoft.net/fig/906333" pageId="0" pageNumber="51">2</figureCitation>
,
<figureCitation id="23E3347A395FCE342A2B272C28A494AB" captionStart="Figures 12–16" captionStartId="F4" captionText="Figures 12 - 16. Differential Interference Contrast (DIC) micrographs of Julolaelaps gigas sp. nov., female 12 idiosoma in dorsal view 13 idiosoma in ventral view 14 chelicera in lateral view, male 15 idiosoma in ventral view 16 chelicera in lateral view." figureDoi="10.3897/zookeys.1180.109099.figures12-16" httpUri="https://binary.pensoft.net/fig/906336" pageId="0" pageNumber="51">13</figureCitation>
). Soft opisthogastric cuticle surrounding epigynal and anal shields with one pair of suboval metapodal plates (38-43 long
<normalizedToken id="180715F35186B347F6F35F903479F359" originalValue="×">x</normalizedToken>
13-18 wide) and seven pairs of smooth setae (
<emphasis id="A4F94AC69179C448BCE7DE3C04DE2D09" italics="true" pageId="0" pageNumber="51">Jv1-Jv3</emphasis>
,
<emphasis id="76E1D7986EE23C3E668DC75102E0D76C" italics="true" pageId="0" pageNumber="51">Jv5</emphasis>
,
<emphasis id="C67729F0CF5FD1BC7A83ACD078DB20C2" italics="true" pageId="0" pageNumber="51">Zv1-Zv3</emphasis>
), setae more or less same in length (18-24) and thickness (Figs
<figureCitation id="EBB8E371222DE074DA74BA2769CC6D1E" captionStart="Figures 1–5" captionStartId="F1" captionText="Figures 1 - 5. Julolaelaps gigas sp. nov., female, with symbols for chaetotactic notation of idiosomal setae 1 dorsal idiosoma 2 ventral idiosoma 3 subcapitulum 4 epistome 5 chelicera, lateral view." figureDoi="10.3897/zookeys.1180.109099.figures1-5" httpUri="https://binary.pensoft.net/fig/906333" pageId="0" pageNumber="51">2</figureCitation>
,
<figureCitation id="69CD59338B3A4F15FCCB9B08B58E2B60" captionStart="Figures 12–16" captionStartId="F4" captionText="Figures 12 - 16. Differential Interference Contrast (DIC) micrographs of Julolaelaps gigas sp. nov., female 12 idiosoma in dorsal view 13 idiosoma in ventral view 14 chelicera in lateral view, male 15 idiosoma in ventral view 16 chelicera in lateral view." figureDoi="10.3897/zookeys.1180.109099.figures12-16" httpUri="https://binary.pensoft.net/fig/906336" pageId="0" pageNumber="51">13</figureCitation>
). Peritreme relatively long and broad, extending to anterior level of coxa II, peritrematal shield narrow, slightly expanded anteriorly, each shield bearing three discernible pore-like structures, a gland pore
<emphasis id="800B1923F97440366913BFE06CDE514D" italics="true" pageId="0" pageNumber="51">gp2</emphasis>
at level of coxa III, a lyrifissures
<emphasis id="D3D3E21246720F5A7F42AC2361A98DB4" italics="true" pageId="0" pageNumber="51">ip3</emphasis>
and a gland pore
<emphasis id="078653A65BFAE9602F545E66F655DBB1" italics="true" pageId="0" pageNumber="51">gp3</emphasis>
on barely developed poststigmatic section (Figs
<figureCitation id="A7003AC1AA202101DEB2356A20D9C738" captionStart="Figures 1–5" captionStartId="F1" captionText="Figures 1 - 5. Julolaelaps gigas sp. nov., female, with symbols for chaetotactic notation of idiosomal setae 1 dorsal idiosoma 2 ventral idiosoma 3 subcapitulum 4 epistome 5 chelicera, lateral view." figureDoi="10.3897/zookeys.1180.109099.figures1-5" httpUri="https://binary.pensoft.net/fig/906333" pageId="0" pageNumber="51">2</figureCitation>
,
<figureCitation id="AE4AA6270F02192B1949307F90D4C06B" captionStart="Figures 12–16" captionStartId="F4" captionText="Figures 12 - 16. Differential Interference Contrast (DIC) micrographs of Julolaelaps gigas sp. nov., female 12 idiosoma in dorsal view 13 idiosoma in ventral view 14 chelicera in lateral view, male 15 idiosoma in ventral view 16 chelicera in lateral view." figureDoi="10.3897/zookeys.1180.109099.figures12-16" httpUri="https://binary.pensoft.net/fig/906336" pageId="0" pageNumber="51">13</figureCitation>
), anterior part of shield not fused with dorsal shield; parapodal platelets broadly thickened, extending behind coxae IV, bearing gland pore
<emphasis id="1374DE71AF6E405FD45E0CF1E9F45075" italics="true" pageId="0" pageNumber="51">gv2</emphasis>
; exopodal platelets absent (Figs
<figureCitation id="28D5F38925DC2164F9638D650E64DFDA" captionStart="Figures 1–5" captionStartId="F1" captionText="Figures 1 - 5. Julolaelaps gigas sp. nov., female, with symbols for chaetotactic notation of idiosomal setae 1 dorsal idiosoma 2 ventral idiosoma 3 subcapitulum 4 epistome 5 chelicera, lateral view." figureDoi="10.3897/zookeys.1180.109099.figures1-5" httpUri="https://binary.pensoft.net/fig/906333" pageId="0" pageNumber="51">2</figureCitation>
,
<figureCitation id="6109D151DB9AFEEBA77E2B4B870B0386" captionStart="Figures 12–16" captionStartId="F4" captionText="Figures 12 - 16. Differential Interference Contrast (DIC) micrographs of Julolaelaps gigas sp. nov., female 12 idiosoma in dorsal view 13 idiosoma in ventral view 14 chelicera in lateral view, male 15 idiosoma in ventral view 16 chelicera in lateral view." figureDoi="10.3897/zookeys.1180.109099.figures12-16" httpUri="https://binary.pensoft.net/fig/906336" pageId="0" pageNumber="51">13</figureCitation>
).
</paragraph>
<caption id="5C636543C09E7C506DF965368062DFA5" doi="10.3897/zookeys.1180.109099.figures6-9" httpUri="https://binary.pensoft.net/fig/906334" pageId="0" pageNumber="51" start="Figures 6–9" startId="F2">
<paragraph id="FA7CB4215619C8FC1A6C8DDB31D8F303" pageId="0" pageNumber="51">
<emphasis id="7B57DEA300E624E08543B3ADBB97618E" bold="true" pageId="0" pageNumber="51">Figures 6-9.</emphasis>
<taxonomicName id="86C429478284A62D7367B8A87AC8CE50" authorityName="Joharchi &amp; Keum &amp; Jung" authorityYear="2023" class="Diplopoda" family="Laelapidae" genus="Julolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Julolaelaps gigas" order="Mesostigmata" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="gigas">
<emphasis id="700CD4443373CE5635682237F34914D6" italics="true" pageId="0" pageNumber="51">Julolaelaps gigas</emphasis>
</taxonomicName>
sp. nov., female, with symbols for chaetotactic notation of legs setae
<emphasis id="728F12486AC8753D2BD23BCA68B124C3" bold="true" pageId="0" pageNumber="51">6</emphasis>
leg I (trochanter-tibia, dorsal aspect)
<emphasis id="757A8D97F2919CF8A2EEB54DEDAC4742" bold="true" pageId="0" pageNumber="51">7</emphasis>
leg II (trochanter-tibia, dorsal aspect)
<emphasis id="09FC5F4C93E4FF0A252E2A28D744F153" bold="true" pageId="0" pageNumber="51">8</emphasis>
leg III (trochanter-tarsus, dorsal aspect)
<emphasis id="ED40D24B3646B00CF79DCBA0B580723E" bold="true" pageId="0" pageNumber="51">9</emphasis>
leg IV (trochanter-tibia, dorsal aspect).
</paragraph>
</caption>
<paragraph id="012A601F0817AEF0701AE718487F5A72" pageId="0" pageNumber="51">
<emphasis id="211D48DF154E056A248D8089AC1C4038" bold="true" italics="true" pageId="0" pageNumber="51">Gnathosomal structures</emphasis>
(Figs
<figureCitation id="F5E04336388B289FC0B009370BBBBCAA" captionStart="Figures 1–5" captionStartId="F1" captionText="Figures 1 - 5. Julolaelaps gigas sp. nov., female, with symbols for chaetotactic notation of idiosomal setae 1 dorsal idiosoma 2 ventral idiosoma 3 subcapitulum 4 epistome 5 chelicera, lateral view." figureDoi="10.3897/zookeys.1180.109099.figures1-5" httpUri="https://binary.pensoft.net/fig/906333" pageId="0" pageNumber="51">3-5</figureCitation>
,
<figureCitation id="A33CB46AB03455EF7769AF18712EA80D" captionStart="Figures 12–16" captionStartId="F4" captionText="Figures 12 - 16. Differential Interference Contrast (DIC) micrographs of Julolaelaps gigas sp. nov., female 12 idiosoma in dorsal view 13 idiosoma in ventral view 14 chelicera in lateral view, male 15 idiosoma in ventral view 16 chelicera in lateral view." figureDoi="10.3897/zookeys.1180.109099.figures12-16" httpUri="https://binary.pensoft.net/fig/906336" pageId="0" pageNumber="51">14</figureCitation>
). Epistome triangular, with pointed apex and smooth (Fig.
<figureCitation id="3545ADC519D08E7EE6B68124989CCBBF" captionStart="Figures 1–5" captionStartId="F1" captionText="Figures 1 - 5. Julolaelaps gigas sp. nov., female, with symbols for chaetotactic notation of idiosomal setae 1 dorsal idiosoma 2 ventral idiosoma 3 subcapitulum 4 epistome 5 chelicera, lateral view." figureDoi="10.3897/zookeys.1180.109099.figures1-5" httpUri="https://binary.pensoft.net/fig/906333" pageId="0" pageNumber="51">4</figureCitation>
). Hypostomal groove with six transverse rows of denticles, each row with 3-5 denticles, with smooth anterior and posterior transverse lines (Fig.
<figureCitation id="5779C19F7F20079B114056D6E5959C54" captionStart="Figures 1–5" captionStartId="F1" captionText="Figures 1 - 5. Julolaelaps gigas sp. nov., female, with symbols for chaetotactic notation of idiosomal setae 1 dorsal idiosoma 2 ventral idiosoma 3 subcapitulum 4 epistome 5 chelicera, lateral view." figureDoi="10.3897/zookeys.1180.109099.figures1-5" httpUri="https://binary.pensoft.net/fig/906333" pageId="0" pageNumber="51">3</figureCitation>
). Hypostome with four pairs of smooth setae,
<emphasis id="540D86261A5CC4A01CEB59446CE820F4" italics="true" pageId="0" pageNumber="51">pc</emphasis>
(24-26)&gt; h1 = h2 (18-21)&gt;
<emphasis id="0497F0762056A2C1E589860D9699DD39" italics="true" pageId="0" pageNumber="51">h3</emphasis>
(14-16) (Fig.
<figureCitation id="289651D66773BE3531DA1692D1035445" captionStart="Figures 1–5" captionStartId="F1" captionText="Figures 1 - 5. Julolaelaps gigas sp. nov., female, with symbols for chaetotactic notation of idiosomal setae 1 dorsal idiosoma 2 ventral idiosoma 3 subcapitulum 4 epistome 5 chelicera, lateral view." figureDoi="10.3897/zookeys.1180.109099.figures1-5" httpUri="https://binary.pensoft.net/fig/906333" pageId="0" pageNumber="51">3</figureCitation>
). Corniculi robust and horn-like, extending slightly beyond palpfemur. Internal malae with one pair of smooth median projections, flanked by lobes with fimbriate anterior margin; labrum with pilose surface; supralabral process not distinct. Chaetotaxy of palps: trochanter 2, femur 5, genu 6, tibia 14, tarsus 15, all setae smooth; palpfemur with seta
<emphasis id="50DC6036A0AAA9C777017B29DC7ED5C7" italics="true" pageId="0" pageNumber="51">d3</emphasis>
thickened and
<emphasis id="BAFB0A2D7E203B2EC6B32524013AE2F3" italics="true" pageId="0" pageNumber="51">al</emphasis>
paddle-like; palpgenu with
<emphasis id="EC39D6005112DFD1FCB40DF1531E7E96" italics="true" pageId="0" pageNumber="51">al1</emphasis>
stout, blunt,
<emphasis id="97B132895F1A86B48077C4A201DB28D8" italics="true" pageId="0" pageNumber="51">al2</emphasis>
thickened and spatulate; palptarsal apotele two-tined. Fixed digit of chelicera with 2-3 teeth and long pilus dentilis, dorsal cheliceral seta relatively thick and short, arthrodial membrane with a rounded flap and normal filaments; movable digit with two mid-sized teeth (Figs
<figureCitation id="5F906FF46D032DCCF7EDCE9C95921C4B" captionStart="Figures 1–5" captionStartId="F1" captionText="Figures 1 - 5. Julolaelaps gigas sp. nov., female, with symbols for chaetotactic notation of idiosomal setae 1 dorsal idiosoma 2 ventral idiosoma 3 subcapitulum 4 epistome 5 chelicera, lateral view." figureDoi="10.3897/zookeys.1180.109099.figures1-5" httpUri="https://binary.pensoft.net/fig/906333" pageId="0" pageNumber="51">5</figureCitation>
,
<figureCitation id="4DB20DAEC3C909F718E35341E2AA9F58" captionStart="Figures 12–16" captionStartId="F4" captionText="Figures 12 - 16. Differential Interference Contrast (DIC) micrographs of Julolaelaps gigas sp. nov., female 12 idiosoma in dorsal view 13 idiosoma in ventral view 14 chelicera in lateral view, male 15 idiosoma in ventral view 16 chelicera in lateral view." figureDoi="10.3897/zookeys.1180.109099.figures12-16" httpUri="https://binary.pensoft.net/fig/906336" pageId="0" pageNumber="51">14</figureCitation>
).
</paragraph>
<paragraph id="73F9AA06426212445E675DDF71BEE233" pageId="0" pageNumber="51">
<emphasis id="01252752E5747057C8287C2612C2EEDE" bold="true" pageId="0" pageNumber="51">
<emphasis id="3B0E89908C27F911F1C022F154C1A5E7" italics="true" pageId="0" pageNumber="51">Insemination structures</emphasis>
.
</emphasis>
Not seen, apparently unsclerotized.
</paragraph>
<paragraph id="3B87A6EAA79359D0C05D0E3FA5D59FED" pageId="0" pageNumber="51">
<emphasis id="013C048BF0A017D99D34C47DF143110E" bold="true" pageId="0" pageNumber="51">
<emphasis id="626320A1363ED8D1B3BC809BABCF78CE" italics="true" pageId="0" pageNumber="51">Legs</emphasis>
.
</emphasis>
(Figs
<figureCitation id="B314F98CD454AC61F296098D60B4FE2B" captionStart="Figures 6–9" captionStartId="F2" captionText="Figures 6 - 9. Julolaelaps gigas sp. nov., female, with symbols for chaetotactic notation of legs setae 6 leg I (trochanter-tibia, dorsal aspect) 7 leg II (trochanter-tibia, dorsal aspect) 8 leg III (trochanter-tarsus, dorsal aspect) 9 leg IV (trochanter-tibia, dorsal aspect)." figureDoi="10.3897/zookeys.1180.109099.figures6-9" httpUri="https://binary.pensoft.net/fig/906334" pageId="0" pageNumber="51">6-9</figureCitation>
). Legs II and III short (600-650, 625-655), I and IV longer (657-680, 739-755). Chaetotaxy normal for free-living
<taxonomicName id="7D85D98969CB38509E9F6923FBE2D4C8" authorityName="A.Berlese" authorityYear="1892" class="Arachnida" family="Laelapidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Mesostigmata" pageId="0" pageNumber="51" phylum="Arthropoda" rank="family">Laelapidae</taxonomicName>
: Leg I (Fig.
<figureCitation id="35B1B3F88083C7454E446AD913F181EF" captionStart="Figures 6–9" captionStartId="F2" captionText="Figures 6 - 9. Julolaelaps gigas sp. nov., female, with symbols for chaetotactic notation of legs setae 6 leg I (trochanter-tibia, dorsal aspect) 7 leg II (trochanter-tibia, dorsal aspect) 8 leg III (trochanter-tarsus, dorsal aspect) 9 leg IV (trochanter-tibia, dorsal aspect)." figureDoi="10.3897/zookeys.1180.109099.figures6-9" httpUri="https://binary.pensoft.net/fig/906334" pageId="0" pageNumber="51">6</figureCitation>
): coxa 0-0/1, 0/1-0, trochanter 1-0/1, 1/2-1, femur 2-3/1, 2/3-2, genu 2-3/2, 3/1-2, tibia 2-3/2, 3/1-2. Leg II (Fig.
<figureCitation id="B9D503826C92A144F43BC15B8B6FBF29" captionStart="Figures 6–9" captionStartId="F2" captionText="Figures 6 - 9. Julolaelaps gigas sp. nov., female, with symbols for chaetotactic notation of legs setae 6 leg I (trochanter-tibia, dorsal aspect) 7 leg II (trochanter-tibia, dorsal aspect) 8 leg III (trochanter-tarsus, dorsal aspect) 9 leg IV (trochanter-tibia, dorsal aspect)." figureDoi="10.3897/zookeys.1180.109099.figures6-9" httpUri="https://binary.pensoft.net/fig/906334" pageId="0" pageNumber="51">7</figureCitation>
): coxa 0-0/1, 0/1-0, trochanter 1-0/1, 0/2-1, femur 2-3/1, 2/2-1, genu 2-3/1, 2/1-2, tibia 2-2/1, 2/1-2. Leg III (Fig.
<figureCitation id="34E63923FF8AA66B5D54607EED61F28A" captionStart="Figures 6–9" captionStartId="F2" captionText="Figures 6 - 9. Julolaelaps gigas sp. nov., female, with symbols for chaetotactic notation of legs setae 6 leg I (trochanter-tibia, dorsal aspect) 7 leg II (trochanter-tibia, dorsal aspect) 8 leg III (trochanter-tarsus, dorsal aspect) 9 leg IV (trochanter-tibia, dorsal aspect)." figureDoi="10.3897/zookeys.1180.109099.figures6-9" httpUri="https://binary.pensoft.net/fig/906334" pageId="0" pageNumber="51">8</figureCitation>
): coxa 0-0/1, 0/1-0, trochanter 1-1/1, 0/1-1, femur 1-2/1, 1/0-1 (
<emphasis id="56C66602405E3468D3BB9F9132CE1EE1" italics="true" pageId="0" pageNumber="51">ad</emphasis>
thickened and inserted on small tubercles), genu 2-2/1, 2/1-1 (
<emphasis id="511974EACBDED55E9F29C8B6E851CF09" italics="true" pageId="0" pageNumber="51">ad1</emphasis>
and
<emphasis id="6F32926D58311C58A87A61BBB3EF4F70" italics="true" pageId="0" pageNumber="51">pd1</emphasis>
inserted on small tubercles), tibia: 2-1/1, 2/1-2. Leg IV (Fig.
<figureCitation id="2A8098CECC7FFC0932E4B8982E2F3595" captionStart="Figures 6–9" captionStartId="F2" captionText="Figures 6 - 9. Julolaelaps gigas sp. nov., female, with symbols for chaetotactic notation of legs setae 6 leg I (trochanter-tibia, dorsal aspect) 7 leg II (trochanter-tibia, dorsal aspect) 8 leg III (trochanter-tarsus, dorsal aspect) 9 leg IV (trochanter-tibia, dorsal aspect)." figureDoi="10.3897/zookeys.1180.109099.figures6-9" httpUri="https://binary.pensoft.net/fig/906334" pageId="0" pageNumber="51">9</figureCitation>
): coxa 0-0/1, 0/0-0, trochanter 1-1/1, 0/1-1 (
<emphasis id="AFD8003AA39FB478346372C22F7841B8" italics="true" pageId="0" pageNumber="51">ad</emphasis>
thickened), femur 1-2/1, 1/0-1 (
<emphasis id="265B9B2DA6BEAD6AB739D9B62CB8342A" italics="true" pageId="0" pageNumber="51">ad1</emphasis>
longest,
<emphasis id="ACE5D5F2EB4DB4AF5677C26BC0D12508" italics="true" pageId="0" pageNumber="51">ad1</emphasis>
and
<emphasis id="EF3C47A165AA20A5580938980A574A4D" italics="true" pageId="0" pageNumber="51">ad2</emphasis>
inserted on small tubercles), genu 2-2/1, 3/0-2, tibia 2-1/1, 3/1-2. Tarsi II-IV with 18 setae (3-3/2, 3/2-3 +
<emphasis id="68652213505548881EA989EC1796DD7B" italics="true" pageId="0" pageNumber="51">mv</emphasis>
,
<emphasis id="E3E1759F1DA69D74AEBBE8AF34ECFF55" italics="true" pageId="0" pageNumber="51">md</emphasis>
); with some ventral and lateral setae thickened. All pretarsi with well-developed paired claws, rounded pulvilli and normal ambulacral stalk.
</paragraph>
<paragraph id="93B74CA8A84E7026C8F9A07193F00D67" pageId="0" pageNumber="51">
<emphasis id="1D835A44DD2118E2AE5EC2B16BF3DA48" bold="true" pageId="0" pageNumber="51">Male</emphasis>
(
<emphasis id="5501568B26F1C80DF8D644A066CF9EE7" italics="true" pageId="0" pageNumber="51">N</emphasis>
= 2). Figs
<figureCitation id="F6B22A547B71BF5F1485C278E7144846" captionStart="Figures 10, 11" captionStartId="F3" captionText="Figures 10, 11. Julolaelaps gigas sp. nov., male 10 ventral idiosoma 11 chelicera, lateral view." figureDoi="10.3897/zookeys.1180.109099.figures10-11" httpUri="https://binary.pensoft.net/fig/906335" pageId="0" pageNumber="51">10</figureCitation>
,
<figureCitation id="99AFBB67066D14BFCDBF80047709FA6F" captionStart="Figures 10, 11" captionStartId="F3" captionText="Figures 10, 11. Julolaelaps gigas sp. nov., male 10 ventral idiosoma 11 chelicera, lateral view." figureDoi="10.3897/zookeys.1180.109099.figures10-11" httpUri="https://binary.pensoft.net/fig/906335" pageId="0" pageNumber="51">11</figureCitation>
,
<figureCitation id="3BB3817716D3692817F4C9497A113E91" captionStart="Figures 12–16" captionStartId="F4" captionText="Figures 12 - 16. Differential Interference Contrast (DIC) micrographs of Julolaelaps gigas sp. nov., female 12 idiosoma in dorsal view 13 idiosoma in ventral view 14 chelicera in lateral view, male 15 idiosoma in ventral view 16 chelicera in lateral view." figureDoi="10.3897/zookeys.1180.109099.figures12-16" httpUri="https://binary.pensoft.net/fig/906336" pageId="0" pageNumber="51">15</figureCitation>
,
<figureCitation id="3EF40B651F542F2DAA3289770C5F5781" captionStart="Figures 12–16" captionStartId="F4" captionText="Figures 12 - 16. Differential Interference Contrast (DIC) micrographs of Julolaelaps gigas sp. nov., female 12 idiosoma in dorsal view 13 idiosoma in ventral view 14 chelicera in lateral view, male 15 idiosoma in ventral view 16 chelicera in lateral view." figureDoi="10.3897/zookeys.1180.109099.figures12-16" httpUri="https://binary.pensoft.net/fig/906336" pageId="0" pageNumber="51">16</figureCitation>
.
<emphasis id="6D61439D529235FE4E07C992D71CA524" bold="true" pageId="0" pageNumber="51">
<emphasis id="99499191A1342EFCC5D3CB77815ED468" italics="true" pageId="0" pageNumber="51">Dorsal idiosoma</emphasis>
.
</emphasis>
Length 760-768, width 525-530. Dorsal shield length 647-653, width 345-350; ornamentation and chaetotaxy as in female.
</paragraph>
<paragraph id="BF91ED9FA64ACD3CFFE2A1235091D6DD" pageId="0" pageNumber="51">
<emphasis id="23F69BF15CD1EF9CE3DD73787D344211" bold="true" pageId="0" pageNumber="51">
<emphasis id="BBFF8A124F57FF30686226D0BB13C43F" italics="true" pageId="0" pageNumber="51">Ventral idiosoma</emphasis>
.
</emphasis>
(Figs
<figureCitation id="61A13A5551004D37C6DAAF227655D0F7" captionStart="Figures 10, 11" captionStartId="F3" captionText="Figures 10, 11. Julolaelaps gigas sp. nov., male 10 ventral idiosoma 11 chelicera, lateral view." figureDoi="10.3897/zookeys.1180.109099.figures10-11" httpUri="https://binary.pensoft.net/fig/906335" pageId="0" pageNumber="51">10</figureCitation>
,
<figureCitation id="B2B9BF4CB568C4DC93D7308E510B85C6" captionStart="Figures 12–16" captionStartId="F4" captionText="Figures 12 - 16. Differential Interference Contrast (DIC) micrographs of Julolaelaps gigas sp. nov., female 12 idiosoma in dorsal view 13 idiosoma in ventral view 14 chelicera in lateral view, male 15 idiosoma in ventral view 16 chelicera in lateral view." figureDoi="10.3897/zookeys.1180.109099.figures12-16" httpUri="https://binary.pensoft.net/fig/906336" pageId="0" pageNumber="51">15</figureCitation>
). Sternal, genital, endopodal, ventral shields fused into sternogenital shield, 327-333 long from anterior to posterior margins of shield, 175-180 wide at level of
<emphasis id="36D888D66B2FB6733A98FC877F150C1A" italics="true" pageId="0" pageNumber="51">st2</emphasis>
, 160-165 at
<emphasis id="6110AB9E532CDAF9DA799702B8906ACB" italics="true" pageId="0" pageNumber="51">st3</emphasis>
level and 118-123 at
<emphasis id="47ABDB05AC84F001CD68324E9D1CCF19" italics="true" pageId="0" pageNumber="51">st5</emphasis>
level; shield with six pairs of simple sternal setae (
<emphasis id="781184C5743175592EAD7B878CAED206" italics="true" pageId="0" pageNumber="51">st1-5</emphasis>
and
<emphasis id="213D5CDC24AD270D919C24CD2CC22454" italics="true" pageId="0" pageNumber="51">Jv1</emphasis>
) (19-30), and two pairs of poroids; anal shield free, length 117-123, width 93-98, surface without reticulate ornamentation, smooth, post-anal seta (25-28) slightly longer than para-anals (18-23), cribrum well developed, with 3-4 irregular rows of spicules, each extending from cribrum to near base of post-anal setae; anal opening located at anterior level of shield; pair of glands
<emphasis id="EB7B2F8858DAC79AB80155A00B145897" italics="true" pageId="0" pageNumber="51">gv3</emphasis>
inserted on shield lateral margins, at level para-anal setae (Figs
<figureCitation id="3863FE54734AAF7580500D1D114B3822" captionStart="Figures 10, 11" captionStartId="F3" captionText="Figures 10, 11. Julolaelaps gigas sp. nov., male 10 ventral idiosoma 11 chelicera, lateral view." figureDoi="10.3897/zookeys.1180.109099.figures10-11" httpUri="https://binary.pensoft.net/fig/906335" pageId="0" pageNumber="51">10</figureCitation>
,
<figureCitation id="323D44939186602E73FA8CEAC7242CB0" captionStart="Figures 12–16" captionStartId="F4" captionText="Figures 12 - 16. Differential Interference Contrast (DIC) micrographs of Julolaelaps gigas sp. nov., female 12 idiosoma in dorsal view 13 idiosoma in ventral view 14 chelicera in lateral view, male 15 idiosoma in ventral view 16 chelicera in lateral view." figureDoi="10.3897/zookeys.1180.109099.figures12-16" httpUri="https://binary.pensoft.net/fig/906336" pageId="0" pageNumber="51">15</figureCitation>
). Soft opisthogastric and lateral cuticle with 10-12 pairs of setae. Peritremes, peritrematal shields and other ventral structures similar to those in female.
</paragraph>
<paragraph id="0645BEB9C1020ADF566D171EB312CE27" pageId="0" pageNumber="51">
<emphasis id="E17B377CA1CD92B1D596A473836A90C9" bold="true" pageId="0" pageNumber="51">
<emphasis id="CBFEACAA6136159804F8BB740C86E5AE" italics="true" pageId="0" pageNumber="51">Gnathosoma</emphasis>
.
</emphasis>
(Figs
<figureCitation id="4B0F95093CEC6E9D9CA33C979273DC6B" captionStart="Figures 10, 11" captionStartId="F3" captionText="Figures 10, 11. Julolaelaps gigas sp. nov., male 10 ventral idiosoma 11 chelicera, lateral view." figureDoi="10.3897/zookeys.1180.109099.figures10-11" httpUri="https://binary.pensoft.net/fig/906335" pageId="0" pageNumber="51">11</figureCitation>
,
<figureCitation id="C6462FD58D5E76530DA6CA6C471C7A25" captionStart="Figures 12–16" captionStartId="F4" captionText="Figures 12 - 16. Differential Interference Contrast (DIC) micrographs of Julolaelaps gigas sp. nov., female 12 idiosoma in dorsal view 13 idiosoma in ventral view 14 chelicera in lateral view, male 15 idiosoma in ventral view 16 chelicera in lateral view." figureDoi="10.3897/zookeys.1180.109099.figures12-16" httpUri="https://binary.pensoft.net/fig/906336" pageId="0" pageNumber="51">16</figureCitation>
). Chelicerae with edentate digits including a regressed fixed digit and a stylet-like movable digit bordered by a long spermatodactyl, pilus dentilis long. Other gnathosomal structures similar to those in female.
</paragraph>
<paragraph id="08958D98AC51FF4B5DCB703F600B8795" pageId="0" pageNumber="51">
<emphasis id="DA1C7164EB05ADB064AA4CE7A91D8CA3" bold="true" pageId="0" pageNumber="51">
<emphasis id="9BF840CF314BD249746E52A103DE0DAA" italics="true" pageId="0" pageNumber="51">Legs</emphasis>
.
</emphasis>
Chaetotaxy as in female.
</paragraph>
</subSubSection>
<subSubSection id="0379F81BCD6E62A1B0D3A028D556E386" pageId="0" pageNumber="51" type="etymology">
<paragraph id="8E847F7172DB0105573C926EFE7DB4FC" pageId="0" pageNumber="51">Etymology.</paragraph>
<paragraph id="0EFDCD2A63D5C89D02E671A29DB12D27" pageId="0" pageNumber="51">
The name of this species refers to its occurrence on giant African millipede of the species
<taxonomicName id="67AEBEBC455E3CA95D3D7FB1E9B3729D" authorityName="Peters" authorityYear="1855" baseAuthorityName="Peters" baseAuthorityYear="1855" class="Diplopoda" family="Spirostreptidae" genus="Archispirostreptus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archispirostreptus gigas" order="Spirostreptida" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="gigas">
<emphasis id="B77BF86D48B03ACE8893877381C987E8" italics="true" pageId="0" pageNumber="51">Archispirostreptus gigas</emphasis>
</taxonomicName>
(
<taxonomicName id="D3A190D41911E05C074EE0FD2A9B87E2" authorityName="Silvestri" authorityYear="1895" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Spirostreptida" pageId="0" pageNumber="51" phylum="Arthropoda" rank="order">Spirostreptida</taxonomicName>
:
<taxonomicName id="939EBB55050A7AA6BFE0B3CCA990D9ED" class="Diplopoda" family="Spirostreptidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Spirostreptida" pageId="0" pageNumber="51" phylum="Arthropoda" rank="family">Spirostreptidae</taxonomicName>
).
</paragraph>
<caption id="DB7F10064B10BBD54995A6BFF1FF36AD" doi="10.3897/zookeys.1180.109099.figures10-11" httpUri="https://binary.pensoft.net/fig/906335" pageId="0" pageNumber="51" start="Figures 10, 11" startId="F3">
<paragraph id="D5694EE61A0FE351D071ECDC549D3ED0" pageId="0" pageNumber="51">
<emphasis id="35E31B4869A106706D1DEA0226FC82F5" bold="true" pageId="0" pageNumber="51">Figures 10, 11.</emphasis>
<taxonomicName id="FD39C003D3B380D436FA5788A6E1769E" authorityName="Joharchi &amp; Keum &amp; Jung" authorityYear="2023" class="Diplopoda" family="Laelapidae" genus="Julolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Julolaelaps gigas" order="Mesostigmata" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="gigas">
<emphasis id="1EF81C3C3E81CEB311B05941769B0DD1" italics="true" pageId="0" pageNumber="51">Julolaelaps gigas</emphasis>
</taxonomicName>
sp. nov., male
<emphasis id="E37D7C102A7EFE397E7B549F42CBDE48" bold="true" pageId="0" pageNumber="51">10</emphasis>
ventral idiosoma
<emphasis id="B4529F9396399C6DF39C60784B1EDB28" bold="true" pageId="0" pageNumber="51">11</emphasis>
chelicera, lateral view.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="5F6D1002DCFEE3646A11C5C6B1C52866" pageId="0" pageNumber="51" type="differential diagnosis">
<paragraph id="343BE0A88C0FAC3599D1D75309F84DFF" pageId="0" pageNumber="51">Differential diagnosis.</paragraph>
<paragraph id="4D13C34783462F6737AD2EFEDDEDBF04" pageId="0" pageNumber="51">
The female of the new species is unique within
<taxonomicName id="902C8EAB66B038E78A82B934EE191F66" class="Diplopoda" family="Laelapidae" genus="Julolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Julolaelaps" order="Mesostigmata" pageId="0" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="FB41498E3AC91EA2C85CAB51B8976876" italics="true" pageId="0" pageNumber="51">Julolaelaps</emphasis>
</taxonomicName>
because of its conspicuously reduced sternal shield, bearing only two pairs of sternal setae (
<emphasis id="78C46A32DDD108252AC5C34984C9BB34" italics="true" pageId="0" pageNumber="51">st1</emphasis>
,
<emphasis id="F9533A066D3EA88C551AEE0E97FC7DEF" italics="true" pageId="0" pageNumber="51">st2</emphasis>
), eroded posteriorly and endopodal plates III/IV broadly thickened. In other features the new species is closest to
<taxonomicName id="18F6ACA01A1E94660787974B0C69BF7E" lsidName="J. vandaelensis" pageId="0" pageNumber="51" rank="species" species="vandaelensis">
<emphasis id="BF1C5A15AA165BEE6E84A442E3506236" italics="true" pageId="0" pageNumber="51">J. vandaelensis</emphasis>
</taxonomicName>
Maes, 1983 in having a genital shield narrower than anal shield, peritreme almost reaching anterior margin of coxae II and dorsal shield with 22 pairs of setae. It is distinguished from this species by the uniform minute acicular dorsal setae and the reduced dorsal shield, not covering entire idiosoma, leaving an unsclerotised cuticle surrounding the dorsal shield.
</paragraph>
<caption id="BEDF99A28D4B6AB2088449EE6FEEE461" doi="10.3897/zookeys.1180.109099.figures12-16" httpUri="https://binary.pensoft.net/fig/906336" pageId="0" pageNumber="51" start="Figures 12–16" startId="F4">
<paragraph id="BEA62E8F7B09E6CEE0911B60919A7E52" pageId="0" pageNumber="51">
<emphasis id="40C099273E2464A8AF32BD2E4D294E0A" bold="true" pageId="0" pageNumber="51">Figures 12-16.</emphasis>
Differential Interference Contrast (DIC) micrographs of
<taxonomicName id="BA5841FC41EFD254A972A839DDBA7626" authorityName="Joharchi &amp; Keum &amp; Jung" authorityYear="2023" class="Diplopoda" family="Laelapidae" genus="Julolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Julolaelaps gigas" order="Mesostigmata" pageId="0" pageNumber="51" phylum="Arthropoda" rank="species" species="gigas">
<emphasis id="BD6BD77FC24A7B77BDD9AECB4CABA655" italics="true" pageId="0" pageNumber="51">Julolaelaps gigas</emphasis>
</taxonomicName>
sp. nov., female
<emphasis id="6CC34F066FE39806561729E9CB611C0F" bold="true" pageId="0" pageNumber="51">12</emphasis>
idiosoma in dorsal view
<emphasis id="459A71A9F6BF21A09813CA22FA251084" bold="true" pageId="0" pageNumber="51">13</emphasis>
idiosoma in ventral view
<emphasis id="B12B27E875F0D27ADC5C3BABA8640031" bold="true" pageId="0" pageNumber="51">14</emphasis>
chelicera in lateral view, male
<emphasis id="9BFD397E17B295BCC18408EF1565D08D" bold="true" pageId="0" pageNumber="51">15</emphasis>
idiosoma in ventral view
<emphasis id="ABC2FC2D13E91D9A1EF0FE7D85A08901" bold="true" pageId="0" pageNumber="51">16</emphasis>
chelicera in lateral view.
</paragraph>
</caption>
</subSubSection>
</treatment>
</subSection>
</document>
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