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<document id="28D949C0E7C059999781D04584A17266" ID-CLB-Dataset="31515" ID-DOI="10.11646/zootaxa.4079.3.2" ID-GBIF-Dataset="1e5444f4-9e36-4413-b932-664119da19de" ID-ISSN="1175-5326" ID-Zenodo-Dep="1050797" ID-ZooBank="300B9A6C-F5FF-43A4-A8D2-82C9646F367D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1510811180316" checkinUser="plazi" docAuthor="Liu, Meike, Yang, Xingke &amp; Huang, Min" docDate="2016" docId="C26A87BA7B5E9716FF2F90013C7DB5E3" docLanguage="en" docName="zootaxa.4079.3.2.pdf" docOrigin="Zootaxa 4079 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyllodes quinquemaculatus Liu &amp; Yang &amp; Huang 2016, sp. n." docType="treatment" docVersion="7" lastPageNumber="355" masterDocId="3E53FFC27B56971CFFB8914D3E5CB243" masterDocTitle="One new species and one newly recorded species of Cyllodes Erichson from China (Coleoptera: Nitidulidae: Nitidulinae)" masterLastPageNumber="356" masterPageNumber="345" pageNumber="353" updateTime="1698492281319" updateUser="ExternalLinkService">
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<mods:title id="8F18F423D27B7858384C7375F9EE149B">One new species and one newly recorded species of Cyllodes Erichson from China (Coleoptera: Nitidulidae: Nitidulinae)</mods:title>
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<mods:namePart id="A4E5CA41B95D43E1658E83F7AB17D346">Yang, Xingke</mods:namePart>
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<mods:namePart id="B782E7E4C20BE51DCAB728B81FBE92FA">Huang, Min</mods:namePart>
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<subSubSection id="02D965277B5E9714FF2F90013F72B3C4" pageId="8" pageNumber="353" type="nomenclature">
<paragraph id="4A7C36AC7B5E9714FF2F90013C73B325" blockId="8.[151,559,332,391]" box="[151,559,332,358]" pageId="8" pageNumber="353">
<heading id="113481C07B5E9714FF2F90013C73B325" bold="true" box="[151,559,332,358]" fontSize="11" level="1" pageId="8" pageNumber="353" reason="1">
<emphasis id="78B7EABE7B5E9714FF2F90013C73B325" bold="true" box="[151,559,332,358]" pageId="8" pageNumber="353">
<taxonomicName id="8DC34D2F7B5E9714FF2F90013FBAB325" authority="Liu &amp; Yang &amp; Huang, 2016" authorityName="Liu &amp; Yang &amp; Huang" authorityYear="2016" box="[151,486,332,358]" class="Insecta" family="Nitidulidae" genus="Cyllodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="353" phylum="Arthropoda" rank="species" species="quinquemaculatus" status="sp. nov.">
<emphasis id="78B7EABE7B5E9714FF2F90013FBAB325" bold="true" box="[151,486,332,358]" italics="true" pageId="8" pageNumber="353">Cyllodes quinquemaculatus</emphasis>
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<taxonomicNameLabel id="638457C57B5E9714FE5490013C73B325" box="[492,559,332,358]" pageId="8" pageNumber="353" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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<paragraph id="4A7C36AC7B5E9714FF2F90233F72B3C4" blockId="8.[151,559,332,391]" box="[151,302,366,391]" pageId="8" pageNumber="353">
(
<figureCitation id="D2F82A297B5E9714FF2790233F79B3C4" box="[159,293,366,391]" captionStart-0="FIGURES 17 18" captionStart-1="FIGURES 19 21" captionStart-2="FIGURES 22 24" captionStartId-0="8.[151,264,1845,1868]" captionStartId-1="9.[151,264,1527,1550]" captionStartId-2="10.[151,264,966,989]" captionTargetBox-0="[161,1389,1000,1779]" captionTargetBox-1="[151,1436,193,1506]" captionTargetBox-2="[151,1436,193,945]" captionTargetId-0="figure@8.[151,1436,983,1824]" captionTargetId-1="figure@9.[151,1436,193,1506]" captionTargetId-2="figure@10.[151,1436,193,945]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="FIGURES 17 18. Cyllodes quinquemaculatus sp. n.: 17. Male habitus, dorsal view; 18. Male habitus, ventral view. Scale bar = 0.50 mm." captionText-1="FIGURES 19 21. Cyllodes quinquemaculatus sp. n.: 19. Right antenna; 20. Right mandible; 21. Right maxilla. Scale bar = 0.20 mm." captionText-2="FIGURES 22 24. Cyllodes quinquemaculatus sp. n.: 22. Male tegmen; 23. Male median lobe of aedeagus; 24. Female ovipositor. Scale bar = 0.10 mm." httpUri-0="https://zenodo.org/record/1050812/files/figure.png" httpUri-1="https://zenodo.org/record/1050814/files/figure.png" httpUri-2="https://zenodo.org/record/1050816/files/figure.png" pageId="8" pageNumber="353">Figs. 1724</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="02D965277B5E9716FF2F90FB3F5CB7B3" lastPageId="10" lastPageNumber="355" pageId="8" pageNumber="353" type="description">
<paragraph id="4A7C36AC7B5E9714FF2F90FB3C35B054" blockId="8.[151,1436,438,931]" pageId="8" pageNumber="353">
<emphasis id="78B7EABE7B5E9714FF2F90FB3F75B38C" bold="true" box="[151,297,438,463]" pageId="8" pageNumber="353">Description.</emphasis>
Length
<quantity id="8D3B9B497B5E9714FE3C90FA3F80B38D" box="[388,476,439,463]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="8" pageNumber="353" unit="mm" value="3.7">3.7 mm</quantity>
, width
<quantity id="8D3B9B497B5E9714FD9690FA3CDAB38D" box="[558,646,439,463]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="8" pageNumber="353" unit="mm" value="2.9">2.9 mm</quantity>
. Pronotum reddish-brown with three black spots; elytra black with two reddish-brown spots. Dorsum glossy and shining, surface sculpture consisting of minute punctures with microreticulate interspaces (
<figureCitation id="D2F82A297B5E9714FE6B90B33C07B054" box="[467,603,510,535]" captionStart="FIGURES 17 18" captionStartId="8.[151,264,1845,1868]" captionTargetBox="[161,1389,1000,1779]" captionTargetId="figure@8.[151,1436,983,1824]" captionTargetPageId="8" captionText="FIGURES 17 18. Cyllodes quinquemaculatus sp. n.: 17. Male habitus, dorsal view; 18. Male habitus, ventral view. Scale bar = 0.50 mm." httpUri="https://zenodo.org/record/1050812/files/figure.png" pageId="8" pageNumber="353">Figs. 17, 18</figureCitation>
).
</paragraph>
<paragraph id="4A7C36AC7B5E9714FF7F936F3CEFB1E0" blockId="8.[151,1436,438,931]" pageId="8" pageNumber="353">
Head with anterior half black and posterior half reddish-brown. Surface punctures irregular, circular or crescent shaped, about 1.01.5× eye facet width in diameter, 1.5 puncture diameters apart. Eyes prominent, finely faceted, interfacetal setae absent. Antennal grooves distinctly margined and slightly deepened, slightly convergent posteriorly, the shortest distance between them about 2× width of mentum. Antenna with funicle reddish-brown and 3-segmented club black. Club about 0.8× length of remaining antennomeres combined. Scape 4.0× longer than pedicel. Pedicel small, 0.7× as long as segment 3 (
<figureCitation id="D2F82A297B5E9714FD73939B3D42B0AC" box="[715,798,726,751]" captionStart="FIGURES 19 21" captionStartId="9.[151,264,1527,1550]" captionTargetBox="[151,1436,193,1506]" captionTargetId="figure@9.[151,1436,193,1506]" captionTargetPageId="9" captionText="FIGURES 19 21. Cyllodes quinquemaculatus sp. n.: 19. Right antenna; 20. Right mandible; 21. Right maxilla. Scale bar = 0.20 mm." httpUri="https://zenodo.org/record/1050814/files/figure.png" pageId="8" pageNumber="353">Fig. 19</figureCitation>
). Apical edge of mandibles with four teeth; molar region with large transverse ridges; prostheca with dense setal brush (
<figureCitation id="D2F82A297B5E9714FCC793B73D84B150" box="[895,984,762,787]" captionStart="FIGURES 19 21" captionStartId="9.[151,264,1527,1550]" captionTargetBox="[151,1436,193,1506]" captionTargetId="figure@9.[151,1436,193,1506]" captionTargetPageId="9" captionText="FIGURES 19 21. Cyllodes quinquemaculatus sp. n.: 19. Right antenna; 20. Right mandible; 21. Right maxilla. Scale bar = 0.20 mm." httpUri="https://zenodo.org/record/1050814/files/figure.png" pageId="8" pageNumber="353">Fig. 20</figureCitation>
). Terminal segment of maxillary palpi elongate, nearly 2× longer than remaining palpomeres combined; lacinia with short dense brown setal brush apically and along anterior 1/3 laterally (
<figureCitation id="D2F82A297B5E9714FDC1920F3C8CB118" box="[633,720,834,859]" captionStart="FIGURES 19 21" captionStartId="9.[151,264,1527,1550]" captionTargetBox="[151,1436,193,1506]" captionTargetId="figure@9.[151,1436,193,1506]" captionTargetPageId="9" captionText="FIGURES 19 21. Cyllodes quinquemaculatus sp. n.: 19. Right antenna; 20. Right mandible; 21. Right maxilla. Scale bar = 0.20 mm." httpUri="https://zenodo.org/record/1050814/files/figure.png" pageId="8" pageNumber="353">Fig. 21</figureCitation>
). Terminal segment of labial palpi longer than wide, slightly narrowed apically.
<collectingRegion id="8807F84E7B5E9714FEC9922A3F8CB13D" box="[369,464,871,894]" country="Myanmar" name="Chin" pageId="8" pageNumber="353">Mentum</collectingRegion>
subpentagonal and angulate at medial anterior margin, with only a few faint minute punctures along posteriorly, interspaces smooth.
</paragraph>
<caption id="1EBC66247B5E9714FF2F96783F50B529" httpUri="https://zenodo.org/record/1050812/files/figure.png" pageId="8" pageNumber="353" targetBox="[161,1389,1000,1779]" targetPageId="8">
<paragraph id="4A7C36AC7B5E9714FF2F96783F50B529" blockId="8.[151,1436,1845,1898]" pageId="8" pageNumber="353">
<emphasis id="78B7EABE7B5E9714FF2F96783F08B508" bold="true" box="[151,340,1845,1868]" pageId="8" pageNumber="353">FIGURES 1718.</emphasis>
<taxonomicName id="8DC34D2F7B5E9714FEE3967B3C28B50F" authority="Liu &amp; Yang &amp; Huang, 2016" authorityName="Liu &amp; Yang &amp; Huang" authorityYear="2016" box="[347,628,1846,1868]" class="Insecta" family="Nitidulidae" genus="Cyllodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="353" phylum="Arthropoda" rank="species" species="quinquemaculatus" status="sp. nov.">
<emphasis id="78B7EABE7B5E9714FEE3967B3C28B50F" box="[347,628,1846,1868]" italics="true" pageId="8" pageNumber="353">Cyllodes quinquemaculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="638457C57B5E9714FDC2967A3CE8B50F" box="[634,692,1847,1868]" pageId="8" pageNumber="353" rank="species">
<emphasis id="78B7EABE7B5E9714FDC2967A3CE8B50F" bold="true" box="[634,692,1847,1868]" pageId="8" pageNumber="353">sp. n.</emphasis>
</taxonomicNameLabel>
: 17. Male habitus, dorsal view; 18. Male habitus, ventral view. Scale bar = 0.50 mm.
</paragraph>
</caption>
<caption id="1EBC66247B5F9715FF2F94BA3EA4B46F" httpUri="https://zenodo.org/record/1050814/files/figure.png" pageId="9" pageNumber="354" targetBox="[151,1436,193,1506]" targetPageId="9">
<paragraph id="4A7C36AC7B5F9715FF2F94BA3EA4B46F" blockId="9.[151,1436,1527,1580]" pageId="9" pageNumber="354">
<emphasis id="78B7EABE7B5F9715FF2F94BA3F09B44E" bold="true" box="[151,341,1527,1550]" pageId="9" pageNumber="354">FIGURES 1921.</emphasis>
<taxonomicName id="8DC34D2F7B5F9715FEE594B53C2BB44D" authority="Liu &amp; Yang &amp; Huang, 2016" authorityName="Liu &amp; Yang &amp; Huang" authorityYear="2016" box="[349,631,1528,1550]" class="Insecta" family="Nitidulidae" genus="Cyllodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="354" phylum="Arthropoda" rank="species" species="quinquemaculatus" status="sp. nov.">
<emphasis id="78B7EABE7B5F9715FEE594B53C2BB44D" box="[349,631,1528,1550]" italics="true" pageId="9" pageNumber="354">Cyllodes quinquemaculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="638457C57B5F9715FDC794B53CE6B44E" box="[639,698,1528,1549]" pageId="9" pageNumber="354" rank="species">
<emphasis id="78B7EABE7B5F9715FDC794B53CE6B44E" bold="true" box="[639,698,1528,1549]" pageId="9" pageNumber="354">sp. n.</emphasis>
</taxonomicNameLabel>
: 19. Right antenna; 20. Right mandible; 21. Right maxilla. Scale bar = 0.20 mm.
</paragraph>
</caption>
<paragraph id="4A7C36AC7B5F9715FF7F97183D72B597" blockId="9.[151,1437,1620,2005]" pageId="9" pageNumber="354">Pronotum somewhat transverse (L:W = 1.0:2.4), widest near posterior angles; punctures slightly smaller than those on vertex, surface sculpture smooth and slightly granular laterally and posteriorly. Scutellar shield semicircular. Prosternum with space between coxae and anterior margin about 1.5× width of procoxal process between coxae. Prosternal process slightly expanded posterior to procoxae, with a longitudinal carina. Mesoventrite with punctation similar to that of pronotum, surface sculpturing granular with some faint microreticulation. Mesoventrite distinctly carinate and on more dorsal plane than metaventrite and prosternum; often obscured by prosternal process. Meso-metaventral junction distinctly procurved. Metaventrite overall transverse (W:L = 1.0:3.0); metaventral disc sculpturing granular with small punctures 2.5× their diameter apart. Elytra steeply sloping, covering pygidium completely; some serially punctate striae visible but other punctures distributed randomly; sculpturing granular to smooth, glabrous; punctures similar to those of pronotum while interspaces between them slightly wider than on pronotum.</paragraph>
<caption id="1EBC66247B5C9716FF2F928B3FBEB1B8" httpUri="https://zenodo.org/record/1050816/files/figure.png" pageId="10" pageNumber="355" targetBox="[151,1436,193,945]" targetPageId="10">
<paragraph id="4A7C36AC7B5C9716FF2F928B3FBEB1B8" blockId="10.[151,1436,966,1019]" pageId="10" pageNumber="355">
<emphasis id="78B7EABE7B5C9716FF2F928B3F04B19F" bold="true" box="[151,344,966,989]" pageId="10" pageNumber="355">FIGURES 2224.</emphasis>
<taxonomicName id="8DC34D2F7B5C9716FEDB928B3CDDB19F" authority="Liu &amp; Yang &amp; Huang, 2016" authorityName="Liu &amp; Yang &amp; Huang" authorityYear="2016" box="[355,641,966,988]" class="Insecta" family="Nitidulidae" genus="Cyllodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="355" phylum="Arthropoda" rank="species" species="quinquemaculatus" status="sp. nov.">
<emphasis id="78B7EABE7B5C9716FEDB928B3CDDB19F" box="[355,641,966,988]" italics="true" pageId="10" pageNumber="355">Cyllodes quinquemaculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="638457C57B5C9716FD33928A3C95B19F" box="[651,713,967,988]" pageId="10" pageNumber="355" rank="species">
<emphasis id="78B7EABE7B5C9716FD33928A3C95B19F" bold="true" box="[651,713,967,988]" pageId="10" pageNumber="355">sp. n.</emphasis>
</taxonomicNameLabel>
: 22. Male tegmen; 23. Male median lobe of aedeagus; 24. Female ovipositor. Scale bar = 0.10 mm.
</paragraph>
</caption>
<paragraph id="4A7C36AC7B5C9716FF7F956A3D75B77E" blockId="10.[151,1437,1063,1952]" pageId="10" pageNumber="355">Distance between mesocoxae widest, 0.4× length of antennal club, slightly wider than distance between metacoxae, 3.0× wider than distance between procoxae. Each femora canaliculate for reception of tibiae; widest near middle. Profemur 2.5×, mesofemur 1.5×, and metafemur about 2× width of corresponding tibia. Tibia wide and flat (W:L = 1.0:2.0). Protibia with distinct stout spines along outer edge and two unequal apical spurs, as wide as antennal club. Mesotibia with row of short blunt spines along apical margin and two obvious apical spurs; thick stiff seta-like spines along lateral, apical and distal 2/3 of inner margin. Metatibial armature similar to that of mesotibia, slightly wider than protibia. Protarsus 0.8× width of protibia; mesotarsus as wide as mesotibia; metatarsus slightly narrower than metatibia. Claws simple.</paragraph>
<paragraph id="4A7C36AC7B5C9716FF7F94053F5CB7B3" blockId="10.[151,1437,1063,1952]" pageId="10" pageNumber="355">
Tegmen of male genitalia short and broad (L:W = 1.5:1.0) with apex rounded, setae dense and long around apical margin, with two distinct apico-lateral tufts present (
<figureCitation id="D2F82A297B5C9716FCF894213DCAB7C7" box="[832,918,1388,1413]" captionStart="FIGURES 22 24" captionStartId="10.[151,264,966,989]" captionTargetBox="[151,1436,193,945]" captionTargetId="figure@10.[151,1436,193,945]" captionTargetPageId="10" captionText="FIGURES 22 24. Cyllodes quinquemaculatus sp. n.: 22. Male tegmen; 23. Male median lobe of aedeagus; 24. Female ovipositor. Scale bar = 0.10 mm." httpUri="https://zenodo.org/record/1050816/files/figure.png" pageId="10" pageNumber="355">Fig. 22</figureCitation>
). Median lobe with apex slightly protuberant and with paired elongate fingerlike projections near middle part (
<figureCitation id="D2F82A297B5C9716FCC094C23D91B7EB" box="[888,973,1423,1448]" captionStart="FIGURES 22 24" captionStartId="10.[151,264,966,989]" captionTargetBox="[151,1436,193,945]" captionTargetId="figure@10.[151,1436,193,945]" captionTargetPageId="10" captionText="FIGURES 22 24. Cyllodes quinquemaculatus sp. n.: 22. Male tegmen; 23. Male median lobe of aedeagus; 24. Female ovipositor. Scale bar = 0.10 mm." httpUri="https://zenodo.org/record/1050816/files/figure.png" pageId="10" pageNumber="355">Fig. 23</figureCitation>
). Female genitalia weakly sclerotized, as observed in other
<taxonomicName id="8DC34D2F7B5C9716FEE794F93F9DB78E" authorityName="Erichson" authorityYear="1843" box="[351,449,1460,1485]" class="Insecta" family="Nitidulidae" genus="Cyllodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="355" phylum="Arthropoda" rank="genus">
<emphasis id="78B7EABE7B5C9716FEE794F93F9DB78E" box="[351,449,1460,1485]" italics="true" pageId="10" pageNumber="355">Cyllodes</emphasis>
</taxonomicName>
species, with gonocoxites fused along midline, broadly widened at base (L:W = 1.2:1.0) (
<figureCitation id="D2F82A297B5C9716FF27949A3EA8B7B3" box="[159,244,1495,1520]" captionStart="FIGURES 22 24" captionStartId="10.[151,264,966,989]" captionTargetBox="[151,1436,193,945]" captionTargetId="figure@10.[151,1436,193,945]" captionTargetPageId="10" captionText="FIGURES 22 24. Cyllodes quinquemaculatus sp. n.: 22. Male tegmen; 23. Male median lobe of aedeagus; 24. Female ovipositor. Scale bar = 0.10 mm." httpUri="https://zenodo.org/record/1050816/files/figure.png" pageId="10" pageNumber="355">Fig. 24</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="02D965277B5C9716FF7F94B13B2DB4E7" pageId="10" pageNumber="355" type="diagnosis">
<paragraph id="4A7C36AC7B5C9716FF7F94B13B2DB4E7" blockId="10.[151,1437,1063,1952]" pageId="10" pageNumber="355">
<emphasis id="78B7EABE7B5C9716FF7F94B13F66B456" bold="true" box="[199,314,1532,1557]" pageId="10" pageNumber="355">Diagnosis</emphasis>
. The new species is similar to
<taxonomicName id="8DC34D2F7B5C9716FD3794B13D06B456" baseAuthorityName="Walker" baseAuthorityYear="1859" box="[655,858,1532,1557]" class="Insecta" family="Nitidulidae" genus="Cyllodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="355" phylum="Arthropoda" rank="species" species="bifascies">
<emphasis id="78B7EABE7B5C9716FD3794B13D06B456" box="[655,858,1532,1557]" italics="true" pageId="10" pageNumber="355">Cyllodes bifascies</emphasis>
</taxonomicName>
, but can be distinguished the latter by the following characters: 1) three spots on pronotum and two spots on elytra; 2) apical edge of mandibles present four dentate cuttings; 3) mesoventrite deeply concave for reception of the prosternal process; 4) the ratio of the length of scape and pedicel is 4.0:1.0, larger than other species of
<taxonomicName id="8DC34D2F7B5C9716FD63972A3D61B4C3" authorityName="Erichson" authorityYear="1843" box="[731,829,1639,1664]" class="Insecta" family="Nitidulidae" genus="Cyllodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="355" phylum="Arthropoda" rank="genus">
<emphasis id="78B7EABE7B5C9716FD63972A3D61B4C3" box="[731,829,1639,1664]" italics="true" pageId="10" pageNumber="355">Cyllodes</emphasis>
</taxonomicName>
; 5) male genitalia with tegmen rounded apically and median lobe present two obvious elongate lateral fingerlike projections touched to each other near middle part.
</paragraph>
</subSubSection>
<subSubSection id="02D965277B5C9716FF7F97E23FE4B577" pageId="10" pageNumber="355" type="materials_examined">
<paragraph id="4A7C36AC7B5C9716FF7F97E23FE4B577" blockId="10.[151,1437,1063,1952]" pageId="10" pageNumber="355">
<materialsCitation id="FAAB3CF17B5C9716FF7F97E23F8BB4AF" ID-GBIF-Occurrence="1700783164" collectingDate="1983-05-03" collectionCode="IZAS" country="China" county="Material" elevation="850" location="Jinghong" municipality="Xishuangbanna" pageId="10" pageNumber="355" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="78B7EABE7B5C9716FF7F97E23FF4B48B" bold="true" box="[199,424,1711,1736]" pageId="10" pageNumber="355">
<collectingCounty id="A31D4E207B5C9716FF7F97E23F73B48B" box="[199,303,1711,1736]" pageId="10" pageNumber="355">Material</collectingCounty>
examined
</emphasis>
.
<typeStatus id="9578880E7B5C9716FE0E97E23C7FB48B" box="[438,547,1711,1736]" pageId="10" pageNumber="355" type="holotype">
<emphasis id="78B7EABE7B5C9716FE0E97E23C7FB48B" bold="true" box="[438,547,1711,1736]" pageId="10" pageNumber="355">Holotype</emphasis>
</typeStatus>
.
<specimenCount id="5CC5FD257B5C9716FD8897FC3C1BB48B" box="[560,583,1713,1736]" pageId="10" pageNumber="355" type="male"></specimenCount>
,
<collectingCountry id="32D4763C7B5C9716FDEB97E23CC7B48B" box="[595,667,1711,1736]" name="China" pageId="10" pageNumber="355">China</collectingCountry>
:
<collectingRegion id="8807F84E7B5C9716FD1C97FC3CA1B48B" box="[676,765,1713,1736]" country="China" name="Yunnan" pageId="10" pageNumber="355">Yunnan</collectingRegion>
,
<collectingMunicipality id="AA18ACD67B5C9716FCBF97E23DE4B48B" box="[775,952,1711,1736]" pageId="10" pageNumber="355">Xishuangbanna</collectingMunicipality>
,
<location id="4F1C60777B5C9716FC7A97E23A70B48B" LSID="urn:lsid:plazi:treatment:C26A87BA7B5E9716FF2F90013C7DB5E3:4F1C60777B5C9716FC7A97E23A70B48B" box="[962,1068,1711,1736]" country="China" county="Material" municipality="Xishuangbanna" name="Jinghong" pageId="10" pageNumber="355" stateProvince="Yunnan">Jinghong</location>
,
<location id="4F1C60777B5C9716FB8F97E23AB9B48B" LSID="urn:lsid:plazi:treatment:C26A87BA7B5E9716FF2F90013C7DB5E3:4F1C60777B5C9716FB8F97E23AB9B48B" box="[1079,1253,1711,1736]" country="China" county="Material" municipality="Xishuangbanna" name="Xiaomengyang" pageId="10" pageNumber="355" stateProvince="Yunnan">Xiaomengyang</location>
,
<elevation id="C1EED19F7B5C9716FB5797E23B37B48B" box="[1263,1387,1711,1736]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="10" pageNumber="355" unit="m" value="850.0">
<quantity id="8D3B9B497B5C9716FB5797E23B73B48B" box="[1263,1327,1711,1736]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="10" pageNumber="355" unit="m" value="850.0">850m</quantity>
a.s.l.
</elevation>
,
<collectingDate id="2E39E9847B5C9716FACF97FD3EB1B4AF" pageId="10" pageNumber="355" value="1983-05-03">
<date id="3E7D106C7B5C9716FACF97FD3EB1B4AF" pageId="10" pageNumber="355" value="1983-05-03">03- V-1983</date>
</collectingDate>
, Панфилов (
<collectionCode id="2CD2AE697B5C9716FE3397993F93B4AF" box="[395,463,1748,1772]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="10" pageNumber="355">IZAS</collectionCode>
)
</materialsCitation>
.
<emphasis id="78B7EABE7B5C9716FE5F97993C03B4AE" bold="true" box="[487,607,1748,1773]" pageId="10" pageNumber="355">Paratypes</emphasis>
. (1♂, 1♀).
<materialsCitation id="FAAB3CF17B5C9716FD4997993CC6B553" ID-GBIF-Occurrence="1700783194" collectingDate="1958-08-20" collectionCode="WANG, IZAS" collectorName="Shuyong" country="China" elevation="1065" location="Menga" municipality="Xishuangbanna" pageId="10" pageNumber="355" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<specimenCount id="5CC5FD257B5C9716FD4997993D49B4AF" box="[753,789,1748,1772]" pageId="10" pageNumber="355" type="male">1♂</specimenCount>
,
<collectingCountry id="32D4763C7B5C9716FC9F97993D32B4AE" box="[807,878,1748,1773]" name="China" pageId="10" pageNumber="355">China</collectingCountry>
:
<collectingRegion id="8807F84E7B5C9716FCC697983D8BB4AF" box="[894,983,1749,1772]" country="China" name="Yunnan" pageId="10" pageNumber="355">Yunnan</collectingRegion>
,
<collectingMunicipality id="AA18ACD67B5C9716FC5E97993ACBB4AE" box="[998,1175,1748,1773]" pageId="10" pageNumber="355">Xishuangbanna</collectingMunicipality>
,
<location id="4F1C60777B5C9716FB1097983AA5B4AE" LSID="urn:lsid:plazi:treatment:C26A87BA7B5E9716FF2F90013C7DB5E3:4F1C60777B5C9716FB1097983AA5B4AE" box="[1192,1273,1749,1773]" country="China" municipality="Xishuangbanna" name="Menga" pageId="10" pageNumber="355" stateProvince="Yunnan">Menga</location>
,
<elevation id="C1EED19F7B5C9716FAB097993E90B553" metricMagnitude="3" metricUnit="m" metricValue="1.065" metricValueMax="1.08" metricValueMin="1.05" pageId="10" pageNumber="355" unit="m" value="1065.0" valueMax="1080.0" valueMin="1050.0">
<quantity id="8D3B9B497B5C9716FAB097993BC1B4AE" box="[1288,1437,1748,1773]" metricMagnitude="3" metricUnit="m" metricValue="1.065" metricValueMax="1.08" metricValueMin="1.05" pageId="10" pageNumber="355" unit="m" value="1065.0" valueMax="1080.0" valueMin="1050.0">10501080m</quantity>
a.s.l.
</elevation>
,
<collectingDate id="2E39E9847B5C9716FF6297B53F28B553" box="[218,372,1784,1808]" pageId="10" pageNumber="355" value="1958-08-20">
<date id="3E7D106C7B5C9716FF6297B53F28B553" box="[218,372,1784,1808]" pageId="10" pageNumber="355" value="1958-08-20">20-VIII-1958</date>
</collectingDate>
,
<collectorName id="E736537A7B5C9716FEC797B53FBFB553" box="[383,483,1784,1808]" pageId="10" pageNumber="355">Shuyong</collectorName>
<collectionCode id="2CD2AE697B5C9716FE5397B53C63B553" box="[491,575,1784,1808]" pageId="10" pageNumber="355">WANG</collectionCode>
(
<collectionCode id="2CD2AE697B5C9716FDF797B53CCFB553" box="[591,659,1784,1808]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="10" pageNumber="355">IZAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="FAAB3CF17B5C9716FD1F97B53FE4B577" ID-GBIF-Occurrence="1700783168" collectingDate="1983-04-17" collectionCode="IZAS" collectorName="Fuji PU" country="China" elevation="650" location="Damenglong" municipality="Xishuangbanna" pageId="10" pageNumber="355" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="holotype">
<specimenCount id="5CC5FD257B5C9716FD1F97B53C96B553" box="[679,714,1784,1808]" pageId="10" pageNumber="355" type="female">1♀</specimenCount>
,
<collectingCountry id="32D4763C7B5C9716FD6197B53D7CB553" box="[729,800,1784,1808]" name="China" pageId="10" pageNumber="355">China</collectingCountry>
:
<collectingRegion id="8807F84E7B5C9716FC9397B53DD8B54C" box="[811,900,1784,1807]" country="China" name="Yunnan" pageId="10" pageNumber="355">Yunnan</collectingRegion>
,
<collectingMunicipality id="AA18ACD67B5C9716FC2897B53A1DB553" box="[912,1089,1784,1808]" pageId="10" pageNumber="355">Xishuangbanna</collectingMunicipality>
,
<location id="4F1C60777B5C9716FBF597B53ABDB553" LSID="urn:lsid:plazi:treatment:C26A87BA7B5E9716FF2F90013C7DB5E3:4F1C60777B5C9716FBF597B53ABDB553" box="[1101,1249,1784,1808]" country="China" municipality="Xishuangbanna" name="Damenglong" pageId="10" pageNumber="355" stateProvince="Yunnan">Damenglong</location>
,
<elevation id="C1EED19F7B5C9716FB5497B53B36B553" box="[1260,1386,1784,1808]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="10" pageNumber="355" unit="m" value="650.0">
<quantity id="8D3B9B497B5C9716FB5497B53B70B553" box="[1260,1324,1784,1808]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="10" pageNumber="355" unit="m" value="650.0">650m</quantity>
a.s.l.
</elevation>
,
<collectingDate id="2E39E9847B5C9716FACF97B53EA9B577" pageId="10" pageNumber="355" value="1983-04-17">
<date id="3E7D106C7B5C9716FACF97B53EA9B577" pageId="10" pageNumber="355" value="1983-04-17">17- IV-1983</date>
</collectingDate>
,
<collectorName id="E736537A7B5C9716FEB996513F05B577" box="[257,345,1820,1845]" pageId="10" pageNumber="355">Fuji PU</collectorName>
(
<collectionCode id="2CD2AE697B5C9716FED096513FF0B577" box="[360,428,1820,1844]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="10" pageNumber="355">IZAS</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="02D965277B5C9716FF7F96723C67B53E" pageId="10" pageNumber="355" type="etymology">
<paragraph id="4A7C36AC7B5C9716FF7F96723C67B53E" blockId="10.[151,1437,1063,1952]" pageId="10" pageNumber="355">
<emphasis id="78B7EABE7B5C9716FF7F96723F1AB51B" bold="true" box="[199,326,1855,1880]" pageId="10" pageNumber="355">Etymology</emphasis>
. The specific epithet is derived from the combined Latin words “quinque” and “maculatus” referring to the five spots on its body.
</paragraph>
</subSubSection>
<subSubSection id="02D965277B5C9716FF7F96CA3C7DB5E3" box="[199,545,1927,1952]" pageId="10" pageNumber="355" type="distribution">
<paragraph id="4A7C36AC7B5C9716FF7F96CA3C7DB5E3" blockId="10.[151,1437,1063,1952]" box="[199,545,1927,1952]" pageId="10" pageNumber="355">
<emphasis id="78B7EABE7B5C9716FF7F96CA3F05B5E3" bold="true" box="[199,345,1927,1952]" pageId="10" pageNumber="355">Distribution</emphasis>
.
<collectingCountry id="32D4763C7B5C9716FEDF96CA3FF6B5E3" box="[359,426,1927,1952]" name="China" pageId="10" pageNumber="355">China</collectingCountry>
(
<collectingRegion id="8807F84E7B5C9716FE0196C53C49B5E3" box="[441,533,1928,1952]" country="China" name="Yunnan" pageId="10" pageNumber="355">Yunnan</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>