574 lines
93 KiB
XML
574 lines
93 KiB
XML
<document id="FBDC93F5383B5993BA980F70786C7C42" ID-DOI="10.11646/zootaxa.3743.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5271036" ID-ZooBank="1F36D59E-7737-4D50-8436-8584CF150DB2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629943368999" checkinUser="felipe" docAuthor="YAMAZAKI, KAZUNORI, YAMAZAKI, MIDORI & UESHIMA, REI" docDate="2013" docId="AF33878FFFCE6869FF1FDE1DE925A1A8" docLanguage="en" docName="zt03743p071.pdf" docOrigin="Zootaxa 3743 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.3743.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hungerfordia papilio subsp. stenoptera K. Yamazaki and Ueshima 2013, subsp. nov." docType="treatment" docVersion="3" lastPageNumber="65" masterDocId="530AFFF7FFF26829FF88D813ED35A557" masterDocTitle="<p> <strong> Systematic review of diplommatinid land snails (Caenogastropoda, Diplommatinidae) endemic to the Palau Islands. (1) Generic classification and revision of <em> Hungerfordia </ em> </ strong> <br /> <strong> species with highly developed axial ribs </ strong> </ p>" masterLastPageNumber="71" masterPageNumber="1" pageNumber="61" updateTime="1699130124248" updateUser="plazi">
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<mods:title id="A274FBA54D34F454CE5099723D00DA08"><p> <strong> Systematic review of diplommatinid land snails (Caenogastropoda, Diplommatinidae) endemic to the Palau Islands. (1) Generic classification and revision of <em> Hungerfordia </ em> </ strong> <br /> <strong> species with highly developed axial ribs </ strong> </ p></mods:title>
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<mods:roleTerm id="1003AD520F4EB7B207FA7E7BA9E18C60">Author</mods:roleTerm>
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<mods:namePart id="02F9A658BF3D630B9F98F56E3B83E1DC">YAMAZAKI, KAZUNORI</mods:namePart>
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<mods:roleTerm id="888275586FB1E04896800765028C7A1F">Author</mods:roleTerm>
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<mods:namePart id="CA42456D46877F1E2309E0609F02E3C1">YAMAZAKI, MIDORI</mods:namePart>
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<mods:roleTerm id="2EE70D48A1AF923C31F86DB5914579BC">Author</mods:roleTerm>
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<mods:namePart id="416C057910CD7F4E2FE2413934E45E7C">UESHIMA, REI</mods:namePart>
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<mods:typeOfResource id="14A5E201BB3747F678F31500C2198CDC">text</mods:typeOfResource>
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<mods:title id="19C74E76AAAB81ADAE0C7EE2C1318E9F">Zootaxa</mods:title>
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<mods:part id="74BE134A4E97BB8382FCFCADE10EC70E">
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<mods:date id="A8555E98B36128E2AF5585988FBA955B">2013</mods:date>
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<mods:number id="2D261F46C1F227ECD103D1E24C903DFB">2013-12-02</mods:number>
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<mods:number id="9A5410CD10FCEFA45E1E8C6F1B3E9CDD">3743</mods:number>
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<mods:url id="746BAACD9F7ADF06BB2535EBD5308415">http://dx.doi.org/10.11646/zootaxa.3743.1.1</mods:url>
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<mods:classification id="CC2B665FA6A42CFAFCDEDBCD1B9DEEC1">journal article</mods:classification>
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<mods:identifier id="D7BF19EF5902252E9AE9D4A66E1C90C3" type="DOI">10.11646/zootaxa.3743.1.1</mods:identifier>
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<treatment id="AF33878FFFCE6869FF1FDE1DE925A1A8" LSID="urn:lsid:plazi:treatment:AF33878FFFCE6869FF1FDE1DE925A1A8" httpUri="http://treatment.plazi.org/id/AF33878FFFCE6869FF1FDE1DE925A1A8" lastPageId="64" lastPageNumber="65" pageId="60" pageNumber="61">
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<subSubSection id="6F806512FFCE6815FF1FDE1DE937A37E" box="[151,1026,1550,1577]" pageId="60" pageNumber="61" type="nomenclature">
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<paragraph id="27253699FFCE6815FF1FDE1DE937A37E" blockId="60.[151,1026,1550,1610]" box="[151,1026,1550,1577]" pageId="60" pageNumber="61">
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<heading id="7C6D81F5FFCE6815FF1FDE1DE937A37E" bold="true" box="[151,1026,1550,1577]" fontSize="11" level="2" pageId="60" pageNumber="61" reason="4">
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<emphasis id="15EEEA8BFFCE6815FF1FDE1DE937A37E" bold="true" box="[151,1026,1550,1577]" pageId="60" pageNumber="61">
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<taxonomicName id="E09A4D1AFFCE6815FF1FDE1DEE44A37E" ID-CoL="7TWG3" authority="K. Yamazaki and Ueshima" authorityName="K. Yamazaki and Ueshima" authorityYear="2013" box="[151,881,1550,1577]" class="Gastropoda" family="Diplommatinidae" genus="Hungerfordia" kingdom="Animalia" order="Architaenioglossa" pageId="60" pageNumber="61" phylum="Mollusca" rank="subSpecies" species="papilio" status="subsp. nov." subSpecies="stenoptera">
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<emphasis id="15EEEA8BFFCE6815FF1FDE1DEF28A37F" bold="true" box="[151,541,1550,1576]" italics="true" pageId="60" pageNumber="61">Hungerfordia papilio stenoptera</emphasis>
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K. Yamazaki and Ueshima
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</taxonomicName>
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<taxonomicNameLabel id="0EDD57F0FFCE6815FCF0DE1CE937A37E" box="[888,1026,1551,1577]" pageId="60" pageNumber="61" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="6F806512FFCE6815FF1FDE22EF5AA3D8" pageId="60" pageNumber="61" type="description">
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<paragraph id="27253699FFCE6815FF1FDE22EC5CA31D" blockId="60.[151,1026,1550,1610]" box="[151,361,1585,1610]" pageId="60" pageNumber="61">
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(
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<figureCitation id="BFA12A1CFFCE6815FF17DE22EC36A31D" box="[159,259,1585,1610]" captionStart="FIGURE 9" captionStartId="11.[151,250,1790,1812]" captionTargetBox="[274,1314,195,1747]" captionTargetId="figure-167@11.[273,1315,193,1748]" captionTargetPageId="11" captionText="FIGURE 9. Radulae of Hungerfordia species. (A) H. pelewensis Beddome, 1889, B0283; (B) H. triplochilus sp. nov., topotypic paratype, B0158; (C) H. expansilabris sp. nov., topotypic paratype, B1299; (D) H. nudicollum sp. nov., topotypic paratype, B0364; (E) H. echinata echinata sp. et subsp. nov., paratype, B1084; (F) H. echinata tubulispina subsp. nov., topotypic paratype, B0169; (G) H. elegantissima sp. nov., topotypic paratype, B0375; (H) H. goniobasis goniobasis sp. et subsp. nov., topotypic paratype, B0235; (I) H. goniobasis dmasechensis subsp. nov., topotypic paratype, B0579; (J) H. lamellata (Crosse, 1866), B0157; (K) H. subalata sp. nov., paratype, B0021; (L) H. pteropurpuroides sp. nov., topotypic paratype, B1116; (M) H. alata (Crosse, 1866), B1041; (N) H. papilio papilio sp. et subsp. nov., paratype, B0374; (O) H. papilio stenoptera subsp. nov., paratype, B0761. Scale bars, 10 µm. All specimens, UMUTZ-MG." figureDoi="http://doi.org/10.5281/zenodo.5271050" httpUri="https://zenodo.org/record/5271050/files/figure.png" pageId="60" pageNumber="61">Figs. 9O</figureCitation>
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,
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<figureCitation id="BFA12A1CFFCE6815FE86DE22EC18A31D" box="[270,301,1585,1610]" captionStart="FIGURE 28" captionStartId="62.[151,249,1859,1881]" captionTargetBox="[151,1436,193,1837]" captionTargetId="figure-128@62.[151,1436,193,1837]" captionTargetPageId="62" captionText="FIGURE 28. Hungerfordia papilio stenoptera subsp. nov. (A1–A4) A topotypic paratype (B0223a); (B) columella (dorsal view), (C) axial ribs above the aperture, and (D) sculpture on the last 2–4 whorls of topotypic paratypes (B0223a, b); (E) a topotypic paratype after removal of the winged ribs, B0223c; (F) a paratype from Loc. Ik3-1, B0212; (G1–G4) holotype, B0223T; (H) a paratype from Loc. Mc4-1, B0334; (I) a paratype from Loc. Mc2-1, B0236; (J1, J2) outer surface, and (J3, J4) inner surface of operculum of a topotypic paratype, B0223. Scale bars 1mm (A–I), 0.5 mm (J). All specimens, UMUTZ-MG." figureDoi="http://doi.org/10.5281/zenodo.5271074" httpUri="https://zenodo.org/record/5271074/files/figure.png" pageId="60" pageNumber="61">28</figureCitation>
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,
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<figureCitation id="BFA12A1CFFCE6815FEB1DE22EC54A31D" box="[313,353,1585,1610]" captionStart="FIGURE 29" captionStartId="63.[151,250,1751,1773]" captionTargetBox="[163,1423,151,1726]" captionTargetId="figure-184@63.[163,1423,149,1726]" captionTargetPageId="63" captionText="FIGURE 29. Shell size and shape variation among Hungerfordia species with highly developed ribs. (A) H. triplochilus sp. nov., holotype; (B) H. expansilabris sp. nov., holotype; (C) H. nudicollum sp. nov., holotype; (D) H. echinata echinata sp. et subsp. nov., holotype; (E) H. echinata tubulispina subsp. nov., holotype; (F1) H. lamellata (Crosse, 1866), B0199a; (F2) H. lamellata (Crosse, 1866), B0282b; (G) H. alata (Crosse, 1866), B0446; (H) H. pteropurpuroides sp. nov., holotype; (I1) H. subalata sp. nov., paratype (B0655); (I2) H. subalata sp. nov., holotype; (J) H. papilio stenoptera subsp. nov., holotype; (K) H. papilio papilio sp. et subsp. nov., holotype; (L) H. goniobasis dmasechensis subsp. nov., holotype; (M) H. goniobasis goniobasis sp. et subsp. nov., holotype; (N) H. elegantissima sp. nov., holotype; (O1) H. pelewensis Beddome, 1889, B0542; (O2) H. pelewensis Beddome, 1889, B0383. Scale bar, 1mm. All specimens, UMUTZ-MG." figureDoi="http://doi.org/10.5281/zenodo.5271076" httpUri="https://zenodo.org/record/5271076/files/figure.png" pageId="60" pageNumber="61">29J</figureCitation>
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)
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</paragraph>
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||
<paragraph id="27253699FFCE6815FF1FDE6AEF5AA3D8" blockId="60.[151,623,1657,1679]" box="[151,623,1657,1679]" pageId="60" pageNumber="61">
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||
<taxonomicName id="E09A4D1AFFCE6815FF1FDE6AEC17A3D8" authorityName="YAMAZAKI & YAMAZAKI & UESHIMA" authorityYear="2013" box="[151,290,1657,1679]" class="Gastropoda" family="Diplommatinidae" genus="Hungerfordia" kingdom="Animalia" order="Architaenioglossa" pageId="60" pageNumber="61" phylum="Mollusca" rank="genus">
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<emphasis id="15EEEA8BFFCE6815FF1FDE6AEC17A3D8" box="[151,290,1657,1679]" italics="true" pageId="60" pageNumber="61">Hungerfordia</emphasis>
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</taxonomicName>
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U:
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<bibRefCitation id="430B4B68FFCE6815FECFDE6AECECA3D8" author="Rundell, R. J." box="[327,473,1657,1679]" pageId="60" pageNumber="61" pagination="3401 - 3412" refId="ref49187" refString="Rundell, R. J. (2008) Cryptic diversity, molecular phylogeny and biogeography of the rock- and leaf-dwelling land snails of Belau (Republic of Palau, Oceania). Philosophical Transactions of the Royal Society B, 363, 3401 - 3412. http: // dx. doi. org / 10.1098 / rstb. 2008.0110" type="journal article" year="2008">Rundell, 2008</bibRefCitation>
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, 3403, fig. 3C.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="6F806512FFCE6814FF1FDEAFEFA4A417" lastPageId="61" lastPageNumber="62" pageId="60" pageNumber="61" type="materials_examined">
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<paragraph id="27253699FFCE6815FF1FDEAFE8A9A2A2" blockId="60.[151,1437,1724,2037]" pageId="60" pageNumber="61">
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||
<emphasis id="15EEEA8BFFCE6815FF1FDEAFECBEA382" bold="true" box="[151,395,1724,1749]" pageId="60" pageNumber="61">Materials examined.</emphasis>
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<materialsCitation id="97F23CC4FFCE6815FE1BDEAFE912A3AE" collectingDate="2005-06-29" collectionCode="R, T" collectorName="Li el Kim & KAZUNORI YAMAZAKI" country="Palau" location="East of Li" municipality="Mecherchar island" pageId="60" pageNumber="61" specimenCount="1" typeStatus="holotype">
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<emphasis id="15EEEA8BFFCE6815FE1BDEAFEF34A382" bold="true" box="[403,513,1724,1749]" italics="true" pageId="60" pageNumber="61">
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||
<typeStatus id="F821883BFFCE6815FE1BDEAFECCBA382" box="[403,510,1724,1749]" pageId="60" pageNumber="61" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<location id="22456042FFCE6815FD81DEAEEF4DA382" LSID="urn:lsid:plazi:treatment:AF33878FFFCE6869FF1FDE1DE925A1A8:22456042FFCE6815FD81DEAEEF4DA382" box="[521,632,1724,1749]" country="Palau" municipality="Mecherchar island" name="East of Li" pageId="60" pageNumber="61">
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East of
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<collectorName id="8A6F534FFFCE6815FDD7DEAFEF4DA382" box="[607,632,1724,1749]" pageId="60" pageNumber="61">Li</collectorName>
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</location>
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el Kim, northern coast of central part of
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<collectingMunicipality id="C741ACE3FFCE6815FBCDDEAFE820A382" box="[1093,1301,1724,1749]" pageId="60" pageNumber="61">Mecherchar island</collectingMunicipality>
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(Loc. Mc1- 1),
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<collectingCountry id="5F8D7609FFCE6815FF35DEF3EDCAA3AE" box="[189,255,1760,1785]" name="Palau" pageId="60" pageNumber="61">Palau</collectingCountry>
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||
. Collected by
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<collectionCode id="418BAE5CFFCE6815FE28DEF2EC83A3AF" box="[416,438,1761,1784]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="60" pageNumber="61">R</collectionCode>
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||
. U. and
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||
<collectorName id="8A6F534FFFCE6815FD9CDEF2EF7AA3AF" box="[532,591,1761,1784]" name="KAZUNORI YAMAZAKI" pageId="60" pageNumber="61">K. Y.</collectorName>
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||
on
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||
<date id="53241059FFCE6815FDF2DEF2EE2EA3AE" box="[634,795,1760,1785]" pageId="60" pageNumber="61" value="2005-06-29">
|
||
<collectingDate id="4360E9B1FFCE6815FDF2DEF2EE2EA3AE" box="[634,795,1760,1785]" pageId="60" pageNumber="61" value="2005-06-29">June 29, 2005</collectingDate>
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||
</date>
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||
. UMUTZ-MG-B0223
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||
<collectionCode id="418BAE5CFFCE6815FB9EDEF3E912A3AE" box="[1046,1063,1760,1785]" name="Tavera, Department of Geology and Geophysics" pageId="60" pageNumber="61">T</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
<emphasis id="15EEEA8BFFCE6815FBBADEF3EC74A24A" bold="true" pageId="60" pageNumber="61">
|
||
<emphasis id="15EEEA8BFFCE6815FBBADEF3E99FA3A0" bold="true" box="[1074,1194,1760,1783]" italics="true" pageId="60" pageNumber="61">
|
||
<typeStatus id="F821883BFFCE6815FBBADEF3E993A3A0" box="[1074,1190,1760,1783]" pageId="60" pageNumber="61" type="paratype">Paratypes</typeStatus>
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||
.
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</emphasis>
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Mecherchar and its satellite islets:
|
||
</emphasis>
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||
Loc. Mc1-1,
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||
<specimenCount id="319CFD10FFCE6815FE6CDF17EFB4A24A" box="[484,641,1796,1821]" pageId="60" pageNumber="61" type="generic">57 specimens</specimenCount>
|
||
,
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<date id="53241059FFCE6815FD07DF17EE21A24A" box="[655,788,1796,1821]" pageId="60" pageNumber="61" value="2005-06-29">29/VI/2005</date>
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||
, UMUTZ-MG-B0223; Loc. Mc2-1,
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||
<specimenCount id="319CFD10FFCE6815FB32DF17E87FA24A" box="[1210,1354,1796,1821]" pageId="60" pageNumber="61" type="generic">6 specimens</specimenCount>
|
||
,
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||
<date id="53241059FFCE6815FAD0DF17EDE7A216" pageId="60" pageNumber="61" value="2005-07-01">1/VII/ 2005</date>
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||
, UMUTZ-MG-B0236; Loc. Mc4-1,
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||
<specimenCount id="319CFD10FFCE6815FD1CDF3BEE02A216" box="[660,823,1832,1857]" pageId="60" pageNumber="61" type="generic">63 specimens</specimenCount>
|
||
,
|
||
<date id="53241059FFCE6815FCC5DF3BEEF8A216" box="[845,973,1832,1857]" pageId="60" pageNumber="61" value="2006-07-09">9/VII/2006</date>
|
||
, UMUTZ-MG-B0334; Loc. Mc5-1,
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||
<specimenCount id="319CFD10FFCE6815FA06DF3BEC31A232" pageId="60" pageNumber="61" type="generic">1 specimen</specimenCount>
|
||
,
|
||
<date id="53241059FFCE6815FE99DF5FECA0A232" box="[273,405,1868,1893]" pageId="60" pageNumber="61" value="2006-11-15">15/XI/2006</date>
|
||
, UMUTZ-MG-B0556; Loc. Mc5-2,
|
||
<specimenCount id="319CFD10FFCE6815FCB3DF5EEEFFA232" box="[827,970,1868,1893]" pageId="60" pageNumber="61" type="generic">7 specimens</specimenCount>
|
||
,
|
||
<date id="53241059FFCE6815FC50DF5FE969A232" box="[984,1116,1868,1893]" pageId="60" pageNumber="61" value="2006-11-16">16/XI/2006</date>
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||
, UMUTZ-MG-B0583; Loc. Mc5-3,
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<specimenCount id="319CFD10FFCE6815FF7CDF63ECA7A2DE" box="[244,402,1904,1929]" pageId="60" pageNumber="61" type="generic">52 specimens</specimenCount>
|
||
,
|
||
<date id="53241059FFCE6815FE29DF63EF10A2DE" box="[417,549,1904,1929]" pageId="60" pageNumber="61" value="2006-11-17">17/XI/2006</date>
|
||
, UMUTZ-MG-B0630; Loc. Mc6-1,
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<specimenCount id="319CFD10FFCE6815FC47DF63E96AA2DE" box="[975,1119,1904,1929]" pageId="60" pageNumber="61" type="generic">2 specimens</specimenCount>
|
||
,
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||
<date id="53241059FFCE6815FBE7DF63E9DCA2DE" box="[1135,1257,1904,1929]" pageId="60" pageNumber="61" value="2007-10-23">23/X/2007</date>
|
||
, UMUTZ-MG- B0761; Loc. Mc6-2,
|
||
<specimenCount id="319CFD10FFCE6815FE1BDF87EF07A2FA" box="[403,562,1940,1965]" pageId="60" pageNumber="61" type="generic">22 specimens</specimenCount>
|
||
,
|
||
<date id="53241059FFCE6815FDCBDF87EF88A2FA" box="[579,701,1940,1965]" pageId="60" pageNumber="61" value="2007-10-25">25/X/2007</date>
|
||
, UMUTZ-MG-B0822; Loc. Mc6-3,
|
||
<specimenCount id="319CFD10FFCE6815FBE5DF87E839A2FA" box="[1133,1292,1940,1965]" pageId="60" pageNumber="61" type="generic">13 specimens</specimenCount>
|
||
,
|
||
<date id="53241059FFCE6815FA95DF87E8ADA2FA" box="[1309,1432,1940,1965]" pageId="60" pageNumber="61" value="2007-10-25">25/X/2007</date>
|
||
, UMUTZ-MG-B0836; Loc. Mc6-4,
|
||
<specimenCount id="319CFD10FFCE6815FDA6DFABEFF9A286" box="[558,716,1976,2001]" pageId="60" pageNumber="61" type="generic">13 specimens</specimenCount>
|
||
,
|
||
<date id="53241059FFCE6815FD52DFABEE61A286" box="[730,852,1976,2001]" pageId="60" pageNumber="61" value="2007-10-25">25/X/2007</date>
|
||
, UMUTZ-MG-B0849; Loc. Mc6-5,
|
||
<specimenCount id="319CFD10FFCE6815FB72DFABE8ADA286" box="[1274,1432,1976,2001]" pageId="60" pageNumber="61" type="generic">10 specimens</specimenCount>
|
||
,
|
||
<date id="53241059FFCE6815FF1FDFCFEC27A2A2" box="[151,274,2012,2037]" pageId="60" pageNumber="61" value="2007-10-26">26/X/2007</date>
|
||
, UMUTZ-MG-B0869; Loc. Mc8-1,
|
||
<specimenCount id="319CFD10FFCE6815FD3FDFCFEE61A2A2" box="[695,852,2012,2037]" pageId="60" pageNumber="61" type="generic">27 specimens</specimenCount>
|
||
,
|
||
<date id="53241059FFCE6815FCEADFCFEEEEA2A2" box="[866,987,2012,2037]" pageId="60" pageNumber="61" value="2011-02-26">26/II/2011</date>
|
||
, UMUTZ-MG-B1202; Loc. Mc8-2, 49
|
||
</paragraph>
|
||
<paragraph id="27253699FFCF6814FF1FD884EFA4A417" blockId="61.[151,1437,151,2023]" pageId="61" pageNumber="62">
|
||
specimens,
|
||
<date id="53241059FFCF6814FE94D884ECA1A5E7" box="[284,404,151,176]" pageId="61" pageNumber="62" value="2011-02-28">28/II/2011</date>
|
||
, UMUTZ-MG-B1253; Loc. Mc8-3,
|
||
<specimenCount id="319CFD10FFCF6814FCB2D884EEE3A5E7" box="[826,982,151,176]" pageId="61" pageNumber="62" type="generic">11 specimens</specimenCount>
|
||
,
|
||
<date id="53241059FFCF6814FC6CD884E969A5E7" box="[996,1116,151,176]" pageId="61" pageNumber="62" value="2011-02-28">28/II/2011</date>
|
||
, UMUTZ-MG-B1261; Loc. Mc8-4,
|
||
<specimenCount id="319CFD10FFCF6814FF7CD8AFECA7A582" box="[244,402,188,213]" pageId="61" pageNumber="62" type="generic">32 specimens</specimenCount>
|
||
,
|
||
<date id="53241059FFCF6814FE29D8AFEF2CA582" box="[417,537,188,213]" pageId="61" pageNumber="62" value="2011-02-28">28/II/2011</date>
|
||
, UMUTZ-MG-B1272; Loc. Mc8-5,
|
||
<specimenCount id="319CFD10FFCF6814FC4CD8AFE956A582" box="[964,1123,188,213]" pageId="61" pageNumber="62" type="generic">36 specimens</specimenCount>
|
||
,
|
||
<date id="53241059FFCF6814FBF9D8AFE9DCA582" box="[1137,1257,188,213]" pageId="61" pageNumber="62" value="2011-02-28">28/II/2011</date>
|
||
, UMUTZ-MG- B1284; Loc. Mc3-1,
|
||
<specimenCount id="319CFD10FFCF6814FE0AD8CCEF28A5AF" box="[386,541,223,248]" pageId="61" pageNumber="62" type="generic">33 specimens</specimenCount>
|
||
,
|
||
<date id="53241059FFCF6814FDA0D8CCEF99A5AF" box="[552,684,223,248]" pageId="61" pageNumber="62" value="2005-06-28">28/VI/2005</date>
|
||
, UMUTZ-MG-B0193.
|
||
<emphasis id="15EEEA8BFFCF6814FC3BD8CCE909A5AF" bold="true" box="[947,1084,223,248]" pageId="61" pageNumber="62">Euidelchol:</emphasis>
|
||
Loc. Ik3-1,
|
||
<specimenCount id="319CFD10FFCF6814FB4FD8CCE858A5AF" box="[1223,1389,223,248]" pageId="61" pageNumber="62" type="generic">111 specimens</specimenCount>
|
||
,
|
||
<date id="53241059FFCF6814FAF0D8CCEDC2A44A" pageId="61" pageNumber="62" value="2005-06-29">29/ VI/2005</date>
|
||
, UMUTZ-MG-B0212; Loc. Ik6-1,
|
||
<specimenCount id="319CFD10FFCF6814FD2BD917EE70A44A" box="[675,837,260,285]" pageId="61" pageNumber="62" type="generic">24 specimens</specimenCount>
|
||
,
|
||
<date id="53241059FFCF6814FCD1D917EEE6A44A" box="[857,979,260,285]" pageId="61" pageNumber="62" value="2007-10-25">25/X/2007</date>
|
||
, UMUTZ-MG-B0806; Loc. Ik8-2,
|
||
<specimenCount id="319CFD10FFCF6814FA08D917EC3AA417" pageId="61" pageNumber="62" type="generic">36 specimens</specimenCount>
|
||
,
|
||
<date id="53241059FFCF6814FE91D934ECA7A417" box="[281,402,295,320]" pageId="61" pageNumber="62" value="2011-02-26">26/II/2011</date>
|
||
, UMUTZ-MG-B1193.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6F806512FFCF6814FF4DD95FEE76A4A5" pageId="61" pageNumber="62" type="diagnosis">
|
||
<paragraph id="27253699FFCF6814FF4DD95FEE76A4A5" blockId="61.[151,1437,151,2023]" pageId="61" pageNumber="62">
|
||
<emphasis id="15EEEA8BFFCF6814FF4DD95FEC0AA432" bold="true" box="[197,319,332,357]" pageId="61" pageNumber="62">Diagnosis.</emphasis>
|
||
Shell generally similar to
|
||
<taxonomicName id="E09A4D1AFFCF6814FD0ED95EEE69A433" authority="papilio" authorityName="papilio K. Yamazaki and Ueshima" authorityYear="2013" box="[646,860,332,356]" class="Gastropoda" family="Diplommatinidae" genus="Hungerfordia" kingdom="Animalia" order="Architaenioglossa" pageId="61" pageNumber="62" phylum="Mollusca" rank="subSpecies" species="papilio" subSpecies="papilio">
|
||
<emphasis id="15EEEA8BFFCF6814FD0ED95EEE69A433" box="[646,860,332,356]" italics="true" pageId="61" pageNumber="62">H. papilio papilio</emphasis>
|
||
</taxonomicName>
|
||
but differs from the nominal subspecies by the following features: axial ribs
|
||
<emphasis id="15EEEA8BFFCF6814FE6FD97CEF24A4D0" box="[487,529,367,391]" italics="true" pageId="61" pageNumber="62">less</emphasis>
|
||
synchronized with those on the previous whorls; wing-like projections of axial ribs on the last 2–3 whorls narrower than the nominal subspecies, clearly separated from those on the previous whorls, more strongly folded at the periphery; angular margin of the inner peristome more strongly protruded. Distribution restricted to Mecherchar and Euidelchol islands.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6F806512FFCF6869FF4DD9EEEEBBA74D" lastPageId="64" lastPageNumber="65" pageId="61" pageNumber="62" type="description">
|
||
<paragraph id="27253699FFCF6814FF4DD9EEEEB2A2B0" blockId="61.[151,1437,151,2023]" pageId="61" pageNumber="62">
|
||
<emphasis id="15EEEA8BFFCF6814FF4DD9EEECA9A741" bold="true" box="[197,412,509,534]" pageId="61" pageNumber="62">
|
||
Description.
|
||
<emphasis id="15EEEA8BFFCF6814FEE9D9EEECA9A741" bold="true" box="[353,412,509,534]" italics="true" pageId="61" pageNumber="62">Shell</emphasis>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="BFA12A1CFFCF6814FE38D9EEEF0BA741" box="[432,574,509,534]" captionStart="FIGURE 28" captionStartId="62.[151,249,1859,1881]" captionTargetBox="[151,1436,193,1837]" captionTargetId="figure-128@62.[151,1436,193,1837]" captionTargetPageId="62" captionText="FIGURE 28. Hungerfordia papilio stenoptera subsp. nov. (A1–A4) A topotypic paratype (B0223a); (B) columella (dorsal view), (C) axial ribs above the aperture, and (D) sculpture on the last 2–4 whorls of topotypic paratypes (B0223a, b); (E) a topotypic paratype after removal of the winged ribs, B0223c; (F) a paratype from Loc. Ik3-1, B0212; (G1–G4) holotype, B0223T; (H) a paratype from Loc. Mc4-1, B0334; (I) a paratype from Loc. Mc2-1, B0236; (J1, J2) outer surface, and (J3, J4) inner surface of operculum of a topotypic paratype, B0223. Scale bars 1mm (A–I), 0.5 mm (J). All specimens, UMUTZ-MG." figureDoi="http://doi.org/10.5281/zenodo.5271074" httpUri="https://zenodo.org/record/5271074/files/figure.png" pageId="61" pageNumber="62">Figs. 28A–I</figureCitation>
|
||
,
|
||
<figureCitation id="BFA12A1CFFCF6814FDC5D9EEEF4CA741" box="[589,633,509,534]" captionStart="FIGURE 29" captionStartId="63.[151,250,1751,1773]" captionTargetBox="[163,1423,151,1726]" captionTargetId="figure-184@63.[163,1423,149,1726]" captionTargetPageId="63" captionText="FIGURE 29. Shell size and shape variation among Hungerfordia species with highly developed ribs. (A) H. triplochilus sp. nov., holotype; (B) H. expansilabris sp. nov., holotype; (C) H. nudicollum sp. nov., holotype; (D) H. echinata echinata sp. et subsp. nov., holotype; (E) H. echinata tubulispina subsp. nov., holotype; (F1) H. lamellata (Crosse, 1866), B0199a; (F2) H. lamellata (Crosse, 1866), B0282b; (G) H. alata (Crosse, 1866), B0446; (H) H. pteropurpuroides sp. nov., holotype; (I1) H. subalata sp. nov., paratype (B0655); (I2) H. subalata sp. nov., holotype; (J) H. papilio stenoptera subsp. nov., holotype; (K) H. papilio papilio sp. et subsp. nov., holotype; (L) H. goniobasis dmasechensis subsp. nov., holotype; (M) H. goniobasis goniobasis sp. et subsp. nov., holotype; (N) H. elegantissima sp. nov., holotype; (O1) H. pelewensis Beddome, 1889, B0542; (O2) H. pelewensis Beddome, 1889, B0383. Scale bar, 1mm. All specimens, UMUTZ-MG." figureDoi="http://doi.org/10.5281/zenodo.5271076" httpUri="https://zenodo.org/record/5271076/files/figure.png" pageId="61" pageNumber="62">29J</figureCitation>
|
||
). Shell sinistral, large for
|
||
<taxonomicName id="E09A4D1AFFCF6814FC26D9EEE992A741" box="[942,1191,509,534]" class="Gastropoda" family="Diplommatinidae" genus="Hungerfordia" kingdom="Animalia" order="Architaenioglossa" pageId="61" pageNumber="62" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="15EEEA8BFFCF6814FC26D9EEE972A742" box="[942,1095,509,533]" italics="true" pageId="61" pageNumber="62">Hungerfordia</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
, slender conical, last whorl widest when removing the protruded ribs and peristomes. Shell color semi-transparent throughout the whorls, upper whorls occasionally yellow-colored with the color becoming deeper towards the apex. Whorls 7–8, convex; last 1/16–1/8 whorls (including interspace between the outer and inner peristomes) expanded upward and ascending onto the penultimate whorl in left lateral view. Suture impressed. Constriction located above the parietal/ columellar junction of the aperture, associated with abrupt reduction of the wing-like ribs, internally without palatal plica except a low axial-crest caused by the constriction. Tuba 7/8 whorls. Protoconch 1+1/4 to 1.5 whorls, smooth on the apical whorl, weakly and finely punctated on the lower whorl. Apical septum absent. Axial ribs developed on the teleoconch, highly developed as wings on most whorls: ribs undeveloped on the first 1/4 whorls of the teleoconch; ribs on the following 1/4–1/3 whorls of initial teleoconch low, moderately or widely spaced, not folded; ribs on the subsequent 2–3 whorls (upper whorls higher than the antepenultimate whorl) widely and highly protruded as wings, not folded or very weakly folded at the periphery, very widely spaced at almost a constant interval of 7 ribs in a whorl, almost synchronized with those on the previous whorls, but occasionally with a few non-synchronized ribs; wing-like projections of the ribs very high, very wide, thin, fragile, arising at or slightly below the suture, almost flat, protruded upward, mostly overlapped with and attaching to those on the previous whorls, with fine and weak spiral sculptures on the outer surface; ribs on the penultimate and antepenultimate whorls also highly and widely protruded as wings, very widely spaced (8–11 ribs in a whorl),
|
||
<emphasis id="15EEEA8BFFCF6814FB43DC26E819A11A" box="[1227,1324,1077,1101]" italics="true" pageId="61" pageNumber="62">partially</emphasis>
|
||
or
|
||
<emphasis id="15EEEA8BFFCF6814FADCDC26E8A9A11A" box="[1364,1436,1077,1101]" italics="true" pageId="61" pageNumber="62">hardly</emphasis>
|
||
synchronized with those on the previous whorls, weakly folded at the periphery; wing-like projections of the ribs narrower than and clearly separated from those on the previous whorls, curved upward, rather stout, arising at short distance below or just below the suture, upper margin weakly folded at the periphery over the lower margin, protruded rather upward; ribs above the aperture (ribs on the last 1 to the last 3/4 whorls around the constriction) abruptly becoming very closely spaced and very low, without wing-like projection, fine; ribs on the last 3/4 whorls highly developed as wings again, very widely spaced; wing-like projections of the ribs arising at longer distance below the suture than those on upper whorls (near periphery of the whorl), protruded laterally or rather downward, upper margin more prominently folded or curled at the periphery over the lower margin. Spiral sculptures present on the teleoconch, fine, very weak, frequently reduced on lower whorls, rather prominent on the outer surface of the wing-like ribs, sinuous in high magnification. Interspace between the wing-like ribs almost smooth on most whorls, with weak growth wrinkles on lower whorls; upper growth wrinkles developed below the suture on the last 5/8 whorls, fine, very low; lower growth wrinkles occasionally present
|
||
<emphasis id="15EEEA8BFFCF6814FC5BDDCDE923A0A1" box="[979,1046,1502,1526]" italics="true" pageId="61" pageNumber="62">above</emphasis>
|
||
the suture of the last whorl, very weak, short, almost vertical. Base convex, without basal keel. Axial protrusions of shell wall (along the ribs) strongly developed on the last 3/4 whorls, hidden by the wing-like projections, abapertural and adapertural sides equally wide and concave in umbilical view. Umbilicus clearly opened in most populations, but very narrow in a few populations, width of 4–9 % of the shell diameter (including the expanded ribs and outer peristome). Aperture hardly or slightly tilted downward against the coiling axis, hardly protruded; columellar axis within aperture almost vertical. Columellar tooth absent. Peristome double, expanded: outer peristome developed, widely expanded beyond the inner one at the palatal and basal sides, often with rounded palatal edge, weakly reflexed outward, smooth and polished on the inner surface; inner peristome pentagonal circular or oval, widely expanded, moderately protruded from the outer one, upper palatal (angular) margin more or less protruded forward in umbilical view and weakly depressed in front view, palatal margin frequently concave just below the angular protrusion in left lateral view, basal margin occasionally concave in umbilical view, parietal margin located below the middle level of the upper body whorl. Interspace between the inner and outer peristomes of moderate width, smooth on the outer surface; uppermost margin ascending onto the penultimate whorl toward the aperture in left lateral view; base expanded toward the aperture in left lateral view.
|
||
</paragraph>
|
||
<caption id="73E56611FFCC6817FF1FDF50E8BCA28C" ID-DOI="http://doi.org/10.5281/zenodo.5271074" ID-Zenodo-Dep="5271074" httpUri="https://zenodo.org/record/5271074/files/figure.png" pageId="62" pageNumber="63" startId="62.[151,249,1859,1881]" targetBox="[151,1436,193,1837]" targetPageId="62">
|
||
<paragraph id="27253699FFCC6817FF1FDF50E8BCA28C" blockId="62.[151,1436,1858,2011]" pageId="62" pageNumber="63">
|
||
<emphasis id="15EEEA8BFFCC6817FF1FDF50EE33A20E" bold="true" box="[151,774,1858,1881]" pageId="62" pageNumber="63">
|
||
FIGURE 28.
|
||
<taxonomicName id="E09A4D1AFFCC6817FEA4DF51EFB6A20F" authority="YAMAZAKI & YAMAZAKI & UESHIMA, 2013" authorityName="YAMAZAKI & YAMAZAKI & UESHIMA" authorityYear="2013" box="[300,643,1858,1880]" class="Gastropoda" family="Diplommatinidae" genus="Hungerfordia" kingdom="Animalia" order="Architaenioglossa" pageId="62" pageNumber="63" phylum="Mollusca" rank="subSpecies" species="papilio" status="subsp. nov." subSpecies="stenoptera">
|
||
<emphasis id="15EEEA8BFFCC6817FEA4DF51EFB6A20F" bold="true" box="[300,643,1858,1880]" italics="true" pageId="62" pageNumber="63">Hungerfordia papilio stenoptera</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="0EDD57F0FFCC6817FD04DF50EE33A20E" box="[652,774,1859,1881]" pageId="62" pageNumber="63" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(A
|
||
<subScript id="BB1E34DCFFCC6817FCA2DF43EE04A20B" attach="left" box="[810,817,1872,1884]" fontSize="5" pageId="62" pageNumber="63">1</subScript>
|
||
–A
|
||
<subScript id="BB1E34DCFFCC6817FCD8DF43EE62A20B" attach="left" box="[848,855,1872,1884]" fontSize="5" pageId="62" pageNumber="63">4</subScript>
|
||
) A topotypic paratype (B0223a); (B) columella (dorsal view), (C) axial ribs above the aperture, and (D) sculpture on the last 2–4 whorls of topotypic paratypes (B0223a, b); (E) a topotypic paratype after removal of the winged ribs, B0223c; (F) a paratype from Loc. Ik3-1, B0212; (G
|
||
<subScript id="BB1E34DCFFCC6817FB75DF83E831A2CB" attach="left" box="[1277,1284,1936,1948]" fontSize="5" pageId="62" pageNumber="63">1</subScript>
|
||
–G
|
||
<subScript id="BB1E34DCFFCC6817FAABDF83E81FA2CB" attach="left" box="[1315,1322,1936,1948]" fontSize="5" pageId="62" pageNumber="63">4</subScript>
|
||
) holotype, B0223T; (H) a paratype from Loc. Mc4-1, B0334; (I) a paratype from Loc. Mc2-1, B0236; (J
|
||
<subScript id="BB1E34DCFFCC6817FBECDFA2E95EA2EA" attach="left" box="[1124,1131,1969,1981]" fontSize="5" pageId="62" pageNumber="63">1</subScript>
|
||
, J
|
||
<subScript id="BB1E34DCFFCC6817FB0BDFA2E9BFA2EA" attach="left" box="[1155,1162,1969,1981]" fontSize="5" pageId="62" pageNumber="63">2</subScript>
|
||
) outer surface, and (J
|
||
<subScript id="BB1E34DCFFCC6817FAE5DFA2E841A2EA" attach="left" box="[1389,1396,1969,1981]" fontSize="5" pageId="62" pageNumber="63">3</subScript>
|
||
, J
|
||
<subScript id="BB1E34DCFFCC6817FA04DFA2E8A6A2EA" attach="left" box="[1420,1427,1969,1981]" fontSize="5" pageId="62" pageNumber="63">4</subScript>
|
||
) inner surface of operculum of a topotypic paratype, B0223. Scale bars 1mm (A–I), 0.5 mm (J). All specimens, UMUTZ-MG.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="73E56611FFCD6816FF1FDEC4E918A299" ID-DOI="http://doi.org/10.5281/zenodo.5271076" ID-Zenodo-Dep="5271076" httpUri="https://zenodo.org/record/5271076/files/figure.png" pageId="63" pageNumber="64" startId="63.[151,250,1751,1773]" targetBox="[163,1423,151,1726]" targetPageId="63">
|
||
<paragraph id="27253699FFCD6816FF1FDEC4E918A299" blockId="63.[151,1437,1751,2002]" pageId="63" pageNumber="64">
|
||
<emphasis id="15EEEA8BFFCD6816FF1FDEC4E9E0A3BA" bold="true" box="[151,1237,1751,1773]" pageId="63" pageNumber="64">
|
||
FIGURE 29. Shell size and shape variation among
|
||
<taxonomicName id="E09A4D1AFFCD6816FD4CDEC4EE92A3BA" box="[708,935,1751,1773]" class="Gastropoda" family="Diplommatinidae" genus="Hungerfordia" kingdom="Animalia" order="Architaenioglossa" pageId="63" pageNumber="64" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="15EEEA8BFFCD6816FD4CDEC4EE60A3BA" bold="true" box="[708,853,1751,1773]" italics="true" pageId="63" pageNumber="64">Hungerfordia</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
with highly developed ribs.
|
||
</emphasis>
|
||
(A)
|
||
<taxonomicName id="E09A4D1AFFCD6816FA8FDECBE8A9A3BA" authority="YAMAZAKI & YAMAZAKI & UESHIMA, 2013" authorityName="YAMAZAKI & YAMAZAKI & UESHIMA" authorityYear="2013" box="[1287,1436,1751,1773]" class="Gastropoda" family="Diplommatinidae" genus="Hungerfordia" kingdom="Animalia" order="Architaenioglossa" pageId="63" pageNumber="64" phylum="Mollusca" rank="species" species="triplochilus" status="sp. nov.">
|
||
<emphasis id="15EEEA8BFFCD6816FA8FDECBE8A9A3BA" box="[1287,1436,1751,1773]" italics="true" pageId="63" pageNumber="64">H. triplochilus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="15EEEA8BFFCD6816FF1FDEE5EDD2A25B" bold="true" box="[151,231,1782,1804]" pageId="63" pageNumber="64">
|
||
<taxonomicNameLabel id="0EDD57F0FFCD6816FF1FDEE5EDD2A25B" box="[151,231,1782,1804]" pageId="63" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype; (B)
|
||
<taxonomicName id="E09A4D1AFFCD6816FE0BDEE4EF1EA25B" authority="YAMAZAKI & YAMAZAKI & UESHIMA, 2013" authorityName="YAMAZAKI & YAMAZAKI & UESHIMA" authorityYear="2013" box="[387,555,1782,1804]" class="Gastropoda" family="Diplommatinidae" genus="Hungerfordia" kingdom="Animalia" order="Architaenioglossa" pageId="63" pageNumber="64" phylum="Mollusca" rank="species" species="expansilabris" status="sp. nov.">
|
||
<emphasis id="15EEEA8BFFCD6816FE0BDEE4EF1EA25B" box="[387,555,1782,1804]" italics="true" pageId="63" pageNumber="64">H. expansilabris</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="15EEEA8BFFCD6816FDB9DEE5EFB7A25B" bold="true" box="[561,642,1782,1804]" pageId="63" pageNumber="64">
|
||
<taxonomicNameLabel id="0EDD57F0FFCD6816FDB9DEE5EFB7A25B" box="[561,642,1782,1804]" pageId="63" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype; (C)
|
||
<taxonomicName id="E09A4D1AFFCD6816FC96DEE4EE9AA25B" authority="YAMAZAKI & YAMAZAKI & UESHIMA, 2013" authorityName="YAMAZAKI & YAMAZAKI & UESHIMA" authorityYear="2013" box="[798,943,1782,1804]" class="Gastropoda" family="Diplommatinidae" genus="Hungerfordia" kingdom="Animalia" order="Architaenioglossa" pageId="63" pageNumber="64" phylum="Mollusca" rank="species" species="nudicollum" status="sp. nov.">
|
||
<emphasis id="15EEEA8BFFCD6816FC96DEE4EE9AA25B" box="[798,943,1782,1804]" italics="true" pageId="63" pageNumber="64">H. nudicollum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="15EEEA8BFFCD6816FC3DDEE5E933A25B" bold="true" box="[949,1030,1782,1804]" pageId="63" pageNumber="64">
|
||
<taxonomicNameLabel id="0EDD57F0FFCD6816FC3DDEE5E933A25B" box="[949,1030,1782,1804]" pageId="63" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype; (D)
|
||
<taxonomicName id="E09A4D1AFFCD6816FB2BDEE4E842A25B" authority="YAMAZAKI & YAMAZAKI & UESHIMA, 2013" authorityName="echinata K. Yamazaki and Ueshima" authorityYear="2013" box="[1187,1399,1782,1804]" class="Gastropoda" family="Diplommatinidae" genus="Hungerfordia" kingdom="Animalia" order="Architaenioglossa" pageId="63" pageNumber="64" phylum="Mollusca" rank="subSpecies" species="echinata" status="sp. et subsp. nov." subSpecies="echinata">
|
||
<emphasis id="15EEEA8BFFCD6816FB2BDEE4E842A25B" box="[1187,1399,1782,1804]" italics="true" pageId="63" pageNumber="64">H. echinata echinata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="15EEEA8BFFCD6816FAF5DEE5EC1DA27D" bold="true" pageId="63" pageNumber="64">
|
||
<taxonomicNameLabel id="0EDD57F0FFCD6816FAF5DEE5EC1DA27D" pageId="63" pageNumber="64" rank="subSpecies">sp. et subsp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype; (E)
|
||
<taxonomicName id="E09A4D1AFFCD6816FE4CDF06EF86A27D" authorityName="YAMAZAKI & YAMAZAKI & UESHIMA" authorityYear="2013" box="[452,691,1812,1834]" class="Gastropoda" family="Diplommatinidae" genus="Hungerfordia" kingdom="Animalia" order="Architaenioglossa" pageId="63" pageNumber="64" phylum="Mollusca" rank="subSpecies" species="echinata" subSpecies="tubulispina">
|
||
<emphasis id="15EEEA8BFFCD6816FE4CDF06EF86A27D" box="[452,691,1812,1834]" italics="true" pageId="63" pageNumber="64">H. echinata tubulispina</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="15EEEA8BFFCD6816FD32DF07EE04A27D" bold="true" box="[698,817,1812,1834]" pageId="63" pageNumber="64">subsp. nov.</emphasis>
|
||
, holotype; (F
|
||
<subScript id="BB1E34DCFFCD6816FC33DF31EEF7A279" attach="left" box="[955,962,1826,1838]" fontSize="5" pageId="63" pageNumber="64">1</subScript>
|
||
)
|
||
<taxonomicName id="E09A4D1AFFCD6816FC59DF06E9D9A27D" authority="(Crosse, 1866)" authorityName="YAMAZAKI & YAMAZAKI & UESHIMA" authorityYear="2013" baseAuthorityName="Crosse" baseAuthorityYear="1866" box="[977,1260,1812,1834]" class="Gastropoda" family="Diplommatinidae" genus="Hungerfordia" kingdom="Animalia" order="Architaenioglossa" pageId="63" pageNumber="64" phylum="Mollusca" rank="species" species="lamellata">
|
||
<emphasis id="15EEEA8BFFCD6816FC59DF06E964A27D" box="[977,1105,1812,1834]" italics="true" pageId="63" pageNumber="64">H. lamellata</emphasis>
|
||
(
|
||
<bibRefCitation id="430B4B68FFCD6816FBD6DF07E9D0A27D" author="Crosse, H." box="[1118,1253,1812,1834]" pageId="63" pageNumber="64" pagination="346 - 350" refId="ref48511" refString="Crosse, H. (1866) Note sur les Mollusques opercules terrestres des iles Pelew ou Palaos. Journal de Conchyliologie, 14, 346 - 350." type="journal article" year="1866">Crosse, 1866</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, B0199a; (F
|
||
<subScript id="BB1E34DCFFCD6816FAE5DF31E841A279" attach="right" box="[1389,1396,1826,1838]" fontSize="5" pageId="63" pageNumber="64">2</subScript>
|
||
)
|
||
<taxonomicName id="E09A4D1AFFCD6816FA0BDF06ECADA21B" authority="(Crosse, 1866)" authorityName="YAMAZAKI & YAMAZAKI & UESHIMA" authorityYear="2013" baseAuthorityName="Crosse" baseAuthorityYear="1866" class="Gastropoda" family="Diplommatinidae" genus="Hungerfordia" kingdom="Animalia" order="Architaenioglossa" pageId="63" pageNumber="64" phylum="Mollusca" rank="species" species="lamellata">
|
||
<emphasis id="15EEEA8BFFCD6816FA0BDF06EDC2A21B" italics="true" pageId="63" pageNumber="64">H. lamellata</emphasis>
|
||
(
|
||
<bibRefCitation id="430B4B68FFCD6816FE8EDF25ECA4A21B" author="Crosse, H." box="[262,401,1846,1868]" pageId="63" pageNumber="64" pagination="346 - 350" refId="ref48511" refString="Crosse, H. (1866) Note sur les Mollusques opercules terrestres des iles Pelew ou Palaos. Journal de Conchyliologie, 14, 346 - 350." type="journal article" year="1866">Crosse, 1866</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, B0282b; (G)
|
||
<taxonomicName id="E09A4D1AFFCD6816FDBADF24EE1CA21B" authority="(Crosse, 1866)" authorityName="YAMAZAKI & YAMAZAKI & UESHIMA" authorityYear="2013" baseAuthorityName="Crosse" baseAuthorityYear="1866" box="[562,809,1846,1868]" class="Gastropoda" family="Diplommatinidae" genus="Hungerfordia" kingdom="Animalia" order="Architaenioglossa" pageId="63" pageNumber="64" phylum="Mollusca" rank="species" species="alata">
|
||
<emphasis id="15EEEA8BFFCD6816FDBADF24EFB2A21B" box="[562,647,1846,1868]" italics="true" pageId="63" pageNumber="64">H. alata</emphasis>
|
||
(
|
||
<bibRefCitation id="430B4B68FFCD6816FD1FDF25EE17A21B" author="Crosse, H." box="[663,802,1846,1868]" pageId="63" pageNumber="64" pagination="346 - 350" refId="ref48511" refString="Crosse, H. (1866) Note sur les Mollusques opercules terrestres des iles Pelew ou Palaos. Journal de Conchyliologie, 14, 346 - 350." type="journal article" year="1866">Crosse, 1866</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, B0446; (H)
|
||
<taxonomicName id="E09A4D1AFFCD6816FC3EDF24E9B3A21B" authority="YAMAZAKI & YAMAZAKI & UESHIMA, 2013" authorityName="YAMAZAKI & YAMAZAKI & UESHIMA" authorityYear="2013" box="[950,1158,1846,1868]" class="Gastropoda" family="Diplommatinidae" genus="Hungerfordia" kingdom="Animalia" order="Architaenioglossa" pageId="63" pageNumber="64" phylum="Mollusca" rank="species" species="pteropurpuroides" status="sp. nov.">
|
||
<emphasis id="15EEEA8BFFCD6816FC3EDF24E9B3A21B" box="[950,1158,1846,1868]" italics="true" pageId="63" pageNumber="64">H. pteropurpuroides</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="15EEEA8BFFCD6816FB07DF25E9D7A21B" bold="true" box="[1167,1250,1846,1868]" pageId="63" pageNumber="64">
|
||
<taxonomicNameLabel id="0EDD57F0FFCD6816FB07DF25E9D7A21B" box="[1167,1250,1846,1868]" pageId="63" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype; (I
|
||
<subScript id="BB1E34DCFFCD6816FAE3DF50E847A218" attach="left" box="[1387,1394,1859,1871]" fontSize="5" pageId="63" pageNumber="64">1</subScript>
|
||
)
|
||
<taxonomicName id="E09A4D1AFFCD6816FA0BDF24EDDBA23A" authority="YAMAZAKI & YAMAZAKI & UESHIMA, 2013" authorityName="YAMAZAKI & YAMAZAKI & UESHIMA" authorityYear="2013" class="Gastropoda" family="Diplommatinidae" genus="Hungerfordia" kingdom="Animalia" order="Architaenioglossa" pageId="63" pageNumber="64" phylum="Mollusca" rank="species" species="subalata" status="sp. nov.">
|
||
<emphasis id="15EEEA8BFFCD6816FA0BDF24EDDBA23A" italics="true" pageId="63" pageNumber="64">H. subalata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="15EEEA8BFFCD6816FF7CDF44EC71A23A" bold="true" box="[244,324,1879,1901]" pageId="63" pageNumber="64">
|
||
<taxonomicNameLabel id="0EDD57F0FFCD6816FF7CDF44EC71A23A" box="[244,324,1879,1901]" pageId="63" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype (B0655); (I
|
||
<subScript id="BB1E34DCFFCD6816FDA8DF76EF12A226" attach="left" box="[544,551,1893,1905]" fontSize="5" pageId="63" pageNumber="64">2</subScript>
|
||
)
|
||
<taxonomicName id="E09A4D1AFFCD6816FDBDDF4BEF9EA23A" authority="YAMAZAKI & YAMAZAKI & UESHIMA, 2013" authorityName="YAMAZAKI & YAMAZAKI & UESHIMA" authorityYear="2013" box="[565,683,1879,1901]" class="Gastropoda" family="Diplommatinidae" genus="Hungerfordia" kingdom="Animalia" order="Architaenioglossa" pageId="63" pageNumber="64" phylum="Mollusca" rank="species" species="subalata" status="sp. nov.">
|
||
<emphasis id="15EEEA8BFFCD6816FDBDDF4BEF9EA23A" box="[565,683,1879,1901]" italics="true" pageId="63" pageNumber="64">H. subalata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="15EEEA8BFFCD6816FD39DF44EE34A23A" bold="true" box="[689,769,1879,1901]" pageId="63" pageNumber="64">
|
||
<taxonomicNameLabel id="0EDD57F0FFCD6816FD39DF44EE34A23A" box="[689,769,1879,1901]" pageId="63" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype; (J)
|
||
<taxonomicName id="E09A4D1AFFCD6816FC1CDF4BE959A23A" authorityName="YAMAZAKI & YAMAZAKI & UESHIMA" authorityYear="2013" box="[916,1132,1879,1901]" class="Gastropoda" family="Diplommatinidae" genus="Hungerfordia" kingdom="Animalia" order="Architaenioglossa" pageId="63" pageNumber="64" phylum="Mollusca" rank="subSpecies" species="papilio" subSpecies="stenoptera">
|
||
<emphasis id="15EEEA8BFFCD6816FC1CDF4BE959A23A" box="[916,1132,1879,1901]" italics="true" pageId="63" pageNumber="64">H. papilio stenoptera</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="15EEEA8BFFCD6816FBFADF44E9DDA23A" bold="true" box="[1138,1256,1879,1901]" pageId="63" pageNumber="64">subsp. nov.</emphasis>
|
||
, holotype; (K)
|
||
<taxonomicName id="E09A4D1AFFCD6816FA0BDF4BEC07A2D9" authority="YAMAZAKI & YAMAZAKI & UESHIMA, 2013" authorityName="papilio K. Yamazaki and Ueshima" authorityYear="2013" class="Gastropoda" family="Diplommatinidae" genus="Hungerfordia" kingdom="Animalia" order="Architaenioglossa" pageId="63" pageNumber="64" phylum="Mollusca" rank="subSpecies" species="papilio" status="sp. et subsp. nov." subSpecies="papilio">
|
||
<emphasis id="15EEEA8BFFCD6816FA0BDF4BEC07A2D9" italics="true" pageId="63" pageNumber="64">H. papilio papilio</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="15EEEA8BFFCD6816FEB5DF6BEF34A2D9" bold="true" box="[317,513,1912,1934]" pageId="63" pageNumber="64">
|
||
<taxonomicNameLabel id="0EDD57F0FFCD6816FEB5DF6BEF34A2D9" box="[317,513,1912,1934]" pageId="63" pageNumber="64" rank="subSpecies">sp. et subsp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype; (L)
|
||
<taxonomicName id="E09A4D1AFFCD6816FD23DF6AEEE1A2D9" authorityName="YAMAZAKI & YAMAZAKI & UESHIMA" authorityYear="2013" box="[683,980,1912,1934]" class="Gastropoda" family="Diplommatinidae" genus="Hungerfordia" kingdom="Animalia" order="Architaenioglossa" pageId="63" pageNumber="64" phylum="Mollusca" rank="subSpecies" species="goniobasis" subSpecies="dmasechensis">
|
||
<emphasis id="15EEEA8BFFCD6816FD23DF6AEEE1A2D9" box="[683,980,1912,1934]" italics="true" pageId="63" pageNumber="64">H. goniobasis dmasechensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="15EEEA8BFFCD6816FC56DF6BE96FA2D9" bold="true" box="[990,1114,1912,1934]" pageId="63" pageNumber="64">subsp. nov.</emphasis>
|
||
, holotype; (M)
|
||
<taxonomicName id="E09A4D1AFFCD6816FA82DF6AEC30A2FA" authority="YAMAZAKI & YAMAZAKI & UESHIMA, 2013" authorityName="goniobasis K. Yamazaki and Ueshima" authorityYear="2013" class="Gastropoda" family="Diplommatinidae" genus="Hungerfordia" kingdom="Animalia" order="Architaenioglossa" pageId="63" pageNumber="64" phylum="Mollusca" rank="subSpecies" species="goniobasis" status="sp. et subsp. nov." subSpecies="goniobasis">
|
||
<emphasis id="15EEEA8BFFCD6816FA82DF6AEC30A2FA" italics="true" pageId="63" pageNumber="64">H. goniobasis goniobasis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="15EEEA8BFFCD6816FE84DF84ECF1A2FA" bold="true" box="[268,452,1943,1965]" pageId="63" pageNumber="64">
|
||
<taxonomicNameLabel id="0EDD57F0FFCD6816FE84DF84ECF1A2FA" box="[268,452,1943,1965]" pageId="63" pageNumber="64" rank="subSpecies">sp. et subsp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype; (N)
|
||
<taxonomicName id="E09A4D1AFFCD6816FDEBDF8BEE3BA2FA" authority="YAMAZAKI & YAMAZAKI & UESHIMA, 2013" authorityName="YAMAZAKI & YAMAZAKI & UESHIMA" authorityYear="2013" box="[611,782,1943,1965]" class="Gastropoda" family="Diplommatinidae" genus="Hungerfordia" kingdom="Animalia" order="Architaenioglossa" pageId="63" pageNumber="64" phylum="Mollusca" rank="species" species="elegantissima" status="sp. nov.">
|
||
<emphasis id="15EEEA8BFFCD6816FDEBDF8BEE3BA2FA" box="[611,782,1943,1965]" italics="true" pageId="63" pageNumber="64">H. elegantissima</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="15EEEA8BFFCD6816FC9EDF84EE52A2FA" bold="true" box="[790,871,1943,1965]" pageId="63" pageNumber="64">
|
||
<taxonomicNameLabel id="0EDD57F0FFCD6816FC9EDF84EE52A2FA" box="[790,871,1943,1965]" pageId="63" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype; (O
|
||
<subScript id="BB1E34DCFFCD6816FC7FDFB7EECBA2E7" attach="right" box="[1015,1022,1956,1968]" fontSize="5" pageId="63" pageNumber="64">1</subScript>
|
||
)
|
||
<taxonomicName id="E09A4D1AFFCD6816FB85DF8BE873A2FA" authority="Beddome, 1889" authorityName=": Beddome" authorityYear="1889" box="[1037,1350,1943,1965]" class="Gastropoda" family="Diplommatinidae" genus="Hungerfordia" kingdom="Animalia" order="Architaenioglossa" pageId="63" pageNumber="64" phylum="Mollusca" rank="species" species="pelewensis">
|
||
<emphasis id="15EEEA8BFFCD6816FB85DF8BE9AEA2FA" box="[1037,1179,1943,1965]" italics="true" pageId="63" pageNumber="64">H. pelewensis</emphasis>
|
||
<bibRefCitation id="430B4B68FFCD6816FB2ADF84E873A2FA" author="Beddome, R. H." box="[1186,1350,1943,1965]" pageId="63" pageNumber="64" pagination="112 - 116" refId="ref48107" refString="Beddome, R. H. (1889) Descriptions of land-shells from the island of Koror, Pelew Group. Proceedings of the Zoological Society of London, 1889, 112 - 116. http: // dx. doi. org / 10.1111 / j. 1469 - 7998.1889. tb 06760. x" type="journal article" year="1889">Beddome, 1889</bibRefCitation>
|
||
</taxonomicName>
|
||
, B0542; (O
|
||
<subScript id="BB1E34DCFFCD6816FF39DFD5ED8DA285" attach="left" box="[177,184,1990,2002]" fontSize="5" pageId="63" pageNumber="64">2</subScript>
|
||
)
|
||
<taxonomicName id="E09A4D1AFFCD6816FF4FDFAAECC9A299" authority="Beddome, 1889" authorityName=": Beddome" authorityYear="1889" box="[199,508,1976,1998]" class="Gastropoda" family="Diplommatinidae" genus="Hungerfordia" kingdom="Animalia" order="Architaenioglossa" pageId="63" pageNumber="64" phylum="Mollusca" rank="species" species="pelewensis">
|
||
<emphasis id="15EEEA8BFFCD6816FF4FDFAAEC61A299" box="[199,340,1976,1998]" italics="true" pageId="63" pageNumber="64">H. pelewensis</emphasis>
|
||
<bibRefCitation id="430B4B68FFCD6816FED2DFABECC9A299" author="Beddome, R. H." box="[346,508,1976,1998]" pageId="63" pageNumber="64" pagination="112 - 116" refId="ref48107" refString="Beddome, R. H. (1889) Descriptions of land-shells from the island of Koror, Pelew Group. Proceedings of the Zoological Society of London, 1889, 112 - 116. http: // dx. doi. org / 10.1111 / j. 1469 - 7998.1889. tb 06760. x" type="journal article" year="1889">Beddome, 1889</bibRefCitation>
|
||
</taxonomicName>
|
||
, B0383. Scale bar, 1mm. All specimens, UMUTZ-MG.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="27253699FFB26869FF4DD884EDC1A584" blockId="64.[151,1436,151,1279]" pageId="64" pageNumber="65">
|
||
<emphasis id="15EEEA8BFFB26869FF4DD884EC60A5E7" bold="true" box="[197,341,151,176]" pageId="64" pageNumber="65">
|
||
<emphasis id="15EEEA8BFFB26869FF4DD884EC7BA5E7" bold="true" box="[197,334,151,176]" italics="true" pageId="64" pageNumber="65">Dimensions</emphasis>
|
||
.
|
||
</emphasis>
|
||
Shell height 4.8–6.2 mm, diameter 3.6–5.0 mm, suture width 1.9–2.6 mm, peristome height 1.5– 2.1 mm.
|
||
</paragraph>
|
||
<paragraph id="27253699FFB26869FF4DD8CCE8BDA448" blockId="64.[151,1436,151,1279]" pageId="64" pageNumber="65">
|
||
<emphasis id="15EEEA8BFFB26869FF4DD8CCEC7DA5AF" bold="true" box="[197,328,223,248]" italics="true" pageId="64" pageNumber="65">Operculum</emphasis>
|
||
(
|
||
<figureCitation id="BFA12A1CFFB26869FED2D8CCECF7A5AF" box="[346,450,223,248]" captionStart="FIGURE 28" captionStartId="62.[151,249,1859,1881]" captionTargetBox="[151,1436,193,1837]" captionTargetId="figure-128@62.[151,1436,193,1837]" captionTargetPageId="62" captionText="FIGURE 28. Hungerfordia papilio stenoptera subsp. nov. (A1–A4) A topotypic paratype (B0223a); (B) columella (dorsal view), (C) axial ribs above the aperture, and (D) sculpture on the last 2–4 whorls of topotypic paratypes (B0223a, b); (E) a topotypic paratype after removal of the winged ribs, B0223c; (F) a paratype from Loc. Ik3-1, B0212; (G1–G4) holotype, B0223T; (H) a paratype from Loc. Mc4-1, B0334; (I) a paratype from Loc. Mc2-1, B0236; (J1, J2) outer surface, and (J3, J4) inner surface of operculum of a topotypic paratype, B0223. Scale bars 1mm (A–I), 0.5 mm (J). All specimens, UMUTZ-MG." figureDoi="http://doi.org/10.5281/zenodo.5271074" httpUri="https://zenodo.org/record/5271074/files/figure.png" pageId="64" pageNumber="65">Figs. 28J</figureCitation>
|
||
<figureCitation id="BFA12A1CFFB26869FE4AD8FDECD7A5A0" box="[450,482,224,252]" captionStart="FIGURE 1" captionStartId="4.[151,250,1978,2000]" captionTargetBox="[187,1407,193,1949]" captionTargetId="figure-53@4.[187,1407,193,1949]" captionTargetPageId="4" captionText="FIGURE 1. Map of Palau islands showing collecting sites. Small red circles show collecting sites. Areas encircled by blue rectangles are shown as enlarged maps in Figs. 2–5. Abbreviations of the collecting sites are listed in Table 2." figureDoi="http://doi.org/10.5281/zenodo.5271038" httpUri="https://zenodo.org/record/5271038/files/figure.png" pageId="64" pageNumber="65">
|
||
<subScript id="BB1E34DCFFB26869FE4AD8FDECFFA5AB" attach="left" box="[450,458,238,252]" fontSize="6" pageId="64" pageNumber="65">1</subScript>
|
||
–J
|
||
</figureCitation>
|
||
<figureCitation id="BFA12A1CFFB26869FE6BD8FDECDEA5AB" box="[483,491,238,252]" captionStart="FIGURES 4–5" captionStartId="6.[151,264,1911,1933]" captionTargetBox="[172,1416,193,1890]" captionTargetId="figure-64@6.[172,1416,193,1890]" captionTargetPageId="6" captionText="FIGURES 4–5. Enlarged maps of Palau islands showing collecting sites. Top (Fig. 4), map of Ngerkutabel island and its neighboring islands or islets; bottom (Fig. 5), map of Mecherchar island and its neighboring islands or islets. Abbreviations of the collecting sites are listed in Table 2." figureDoi="http://doi.org/10.5281/zenodo.5271042" httpUri="https://zenodo.org/record/5271042/files/figure.png" pageId="64" pageNumber="65">
|
||
<subScript id="BB1E34DCFFB26869FE6BD8FDECDEA5AB" attach="right" box="[483,491,238,252]" fontSize="6" pageId="64" pageNumber="65">4</subScript>
|
||
</figureCitation>
|
||
). Operculum corneous, multispiral, circular, transparent, amber colored, thin, flat, slightly concave; outer surface smooth; inner surface with a low, weak, arcuate ridge near the columellar margin.
|
||
</paragraph>
|
||
<paragraph id="27253699FFB26869FF4DD939EC90A414" blockId="64.[151,1436,151,1279]" box="[197,421,298,323]" pageId="64" pageNumber="65">
|
||
<emphasis id="15EEEA8BFFB26869FF4DD939EC3EA414" bold="true" box="[197,267,298,323]" pageId="64" pageNumber="65">
|
||
<emphasis id="15EEEA8BFFB26869FF4DD939EC30A414" bold="true" box="[197,261,298,323]" italics="true" pageId="64" pageNumber="65">Penis</emphasis>
|
||
.
|
||
</emphasis>
|
||
Penis absent.
|
||
</paragraph>
|
||
<paragraph id="27253699FFB26869FF4DD95DEEBBA74D" blockId="64.[151,1436,151,1279]" pageId="64" pageNumber="65">
|
||
<taxonomicName id="E09A4D1AFFB26869FF4DD95DEC2CA430" box="[197,281,334,359]" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Plantae" order="Porellales" pageId="64" pageNumber="65" phylum="Marchantiophyta" rank="genus">
|
||
<emphasis id="15EEEA8BFFB26869FF4DD95DEC2CA430" bold="true" box="[197,281,334,359]" italics="true" pageId="64" pageNumber="65">Radula</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="BFA12A1CFFB26869FEA0D95DECB4A430" box="[296,385,334,359]" captionStart="FIGURE 9" captionStartId="11.[151,250,1790,1812]" captionTargetBox="[274,1314,195,1747]" captionTargetId="figure-167@11.[273,1315,193,1748]" captionTargetPageId="11" captionText="FIGURE 9. Radulae of Hungerfordia species. (A) H. pelewensis Beddome, 1889, B0283; (B) H. triplochilus sp. nov., topotypic paratype, B0158; (C) H. expansilabris sp. nov., topotypic paratype, B1299; (D) H. nudicollum sp. nov., topotypic paratype, B0364; (E) H. echinata echinata sp. et subsp. nov., paratype, B1084; (F) H. echinata tubulispina subsp. nov., topotypic paratype, B0169; (G) H. elegantissima sp. nov., topotypic paratype, B0375; (H) H. goniobasis goniobasis sp. et subsp. nov., topotypic paratype, B0235; (I) H. goniobasis dmasechensis subsp. nov., topotypic paratype, B0579; (J) H. lamellata (Crosse, 1866), B0157; (K) H. subalata sp. nov., paratype, B0021; (L) H. pteropurpuroides sp. nov., topotypic paratype, B1116; (M) H. alata (Crosse, 1866), B1041; (N) H. papilio papilio sp. et subsp. nov., paratype, B0374; (O) H. papilio stenoptera subsp. nov., paratype, B0761. Scale bars, 10 µm. All specimens, UMUTZ-MG." figureDoi="http://doi.org/10.5281/zenodo.5271050" httpUri="https://zenodo.org/record/5271050/files/figure.png" pageId="64" pageNumber="65">Fig. 9O</figureCitation>
|
||
).
|
||
<taxonomicName id="E09A4D1AFFB26869FE1CD95DECD6A430" box="[404,483,334,359]" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Plantae" order="Porellales" pageId="64" pageNumber="65" phylum="Marchantiophyta" rank="genus">Radula</taxonomicName>
|
||
of specialized
|
||
<typeStatus id="F821883BFFB26869FD03D95CEF8EA430" box="[651,699,335,359]" pageId="64" pageNumber="65">type</typeStatus>
|
||
of taenioglossate; ribbon very long. Central tooth very large, with a very wide, large, and blunt central cusp, without lateral cup, basal plate longitudinally elongated and strongly constricted at the base. Lateral teeth also large, with a large, very wide and blunt major cusp, without inner nor outer cusp. Inner marginal teeth narrower and shorter than the lateral teeth, with a large major cusp, with two slender inner cusps, with a reduced outer cup. Outer marginal teeth shorter than the inner marginals, with a large and blunt major cusp, with a slender inner cusp, without outer cusp.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6F806512FFB26869FF4DDA35E9DCA734" pageId="64" pageNumber="65" type="distribution">
|
||
<paragraph id="27253699FFB26869FF4DDA35E9DCA734" blockId="64.[151,1436,151,1279]" pageId="64" pageNumber="65">
|
||
<emphasis id="15EEEA8BFFB26869FF4DDA35ECC5A768" bold="true" box="[197,496,550,575]" pageId="64" pageNumber="65">Distribution and ecology.</emphasis>
|
||
Endemic to
|
||
<collectingCountry id="5F8D7609FFB26869FDF5DA35EFF5A768" box="[637,704,550,575]" name="Palau" pageId="64" pageNumber="65">Palau</collectingCountry>
|
||
: Mecherchar and Euidelchol islands, and their satellite islets. The species inhabits vertical walls of large limestone rocks or cliffs, occasionally on limestone rubbles.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6F806512FFB26869FF4DDA7DEE4DA1E0" pageId="64" pageNumber="65" type="discussion">
|
||
<paragraph id="27253699FFB26869FF4DDA7DEEA3A138" blockId="64.[151,1436,151,1279]" pageId="64" pageNumber="65">
|
||
<emphasis id="15EEEA8BFFB26869FF4DDA7DEC0CA7D0" bold="true" box="[197,313,622,647]" pageId="64" pageNumber="65">Remarks.</emphasis>
|
||
<taxonomicName id="E09A4D1AFFB26869FEC9DA63EF04A7D0" authorityName="YAMAZAKI & YAMAZAKI & UESHIMA" authorityYear="2013" box="[321,561,623,647]" class="Gastropoda" family="Diplommatinidae" genus="Hungerfordia" kingdom="Animalia" order="Architaenioglossa" pageId="64" pageNumber="65" phylum="Mollusca" rank="subSpecies" species="papilio" subSpecies="stenoptera">
|
||
<emphasis id="15EEEA8BFFB26869FEC9DA63EF04A7D0" box="[321,561,623,647]" italics="true" pageId="64" pageNumber="65">H. papilio stenoptera</emphasis>
|
||
</taxonomicName>
|
||
differs from the nominal subspecies in the following points. First,
|
||
<taxonomicName id="E09A4D1AFFB26869FAA1DA63EC38A7FD" authorityName="YAMAZAKI & YAMAZAKI & UESHIMA" authorityYear="2013" class="Gastropoda" family="Diplommatinidae" genus="Hungerfordia" kingdom="Animalia" order="Architaenioglossa" pageId="64" pageNumber="65" phylum="Mollusca" rank="subSpecies" species="papilio" subSpecies="stenoptera">
|
||
<emphasis id="15EEEA8BFFB26869FAA1DA63EC38A7FD" italics="true" pageId="64" pageNumber="65">H. papilio stenoptera</emphasis>
|
||
</taxonomicName>
|
||
has narrower wing-like projections of the ribs on the penultimate and antepenultimate whorls, which are clearly separated from those on the previous whorls (
|
||
<figureCitation id="BFA12A1CFFB26869FD61DAA5EE6CA798" box="[745,857,694,719]" captionStart="FIGURE 28" captionStartId="62.[151,249,1859,1881]" captionTargetBox="[151,1436,193,1837]" captionTargetId="figure-128@62.[151,1436,193,1837]" captionTargetPageId="62" captionText="FIGURE 28. Hungerfordia papilio stenoptera subsp. nov. (A1–A4) A topotypic paratype (B0223a); (B) columella (dorsal view), (C) axial ribs above the aperture, and (D) sculpture on the last 2–4 whorls of topotypic paratypes (B0223a, b); (E) a topotypic paratype after removal of the winged ribs, B0223c; (F) a paratype from Loc. Ik3-1, B0212; (G1–G4) holotype, B0223T; (H) a paratype from Loc. Mc4-1, B0334; (I) a paratype from Loc. Mc2-1, B0236; (J1, J2) outer surface, and (J3, J4) inner surface of operculum of a topotypic paratype, B0223. Scale bars 1mm (A–I), 0.5 mm (J). All specimens, UMUTZ-MG." figureDoi="http://doi.org/10.5281/zenodo.5271074" httpUri="https://zenodo.org/record/5271074/files/figure.png" pageId="64" pageNumber="65">Figs. 28A</figureCitation>
|
||
<figureCitation id="BFA12A1CFFB26869FCD1DAD6EE54A784" box="[857,865,709,723]" captionStart="FIGURE 1" captionStartId="4.[151,250,1978,2000]" captionTargetBox="[187,1407,193,1949]" captionTargetId="figure-53@4.[187,1407,193,1949]" captionTargetPageId="4" captionText="FIGURE 1. Map of Palau islands showing collecting sites. Small red circles show collecting sites. Areas encircled by blue rectangles are shown as enlarged maps in Figs. 2–5. Abbreviations of the collecting sites are listed in Table 2." figureDoi="http://doi.org/10.5281/zenodo.5271038" httpUri="https://zenodo.org/record/5271038/files/figure.png" pageId="64" pageNumber="65">
|
||
<subScript id="BB1E34DCFFB26869FCD1DAD6EE54A784" attach="left" box="[857,865,709,723]" fontSize="6" pageId="64" pageNumber="65">1</subScript>
|
||
</figureCitation>
|
||
,
|
||
<figureCitation id="BFA12A1CFFB26869FCE6DAA5EEE9A798" box="[878,988,694,719]" captionStart="FIGURE 28" captionStartId="62.[151,249,1859,1881]" captionTargetBox="[151,1436,193,1837]" captionTargetId="figure-128@62.[151,1436,193,1837]" captionTargetPageId="62" captionText="FIGURE 28. Hungerfordia papilio stenoptera subsp. nov. (A1–A4) A topotypic paratype (B0223a); (B) columella (dorsal view), (C) axial ribs above the aperture, and (D) sculpture on the last 2–4 whorls of topotypic paratypes (B0223a, b); (E) a topotypic paratype after removal of the winged ribs, B0223c; (F) a paratype from Loc. Ik3-1, B0212; (G1–G4) holotype, B0223T; (H) a paratype from Loc. Mc4-1, B0334; (I) a paratype from Loc. Mc2-1, B0236; (J1, J2) outer surface, and (J3, J4) inner surface of operculum of a topotypic paratype, B0223. Scale bars 1mm (A–I), 0.5 mm (J). All specimens, UMUTZ-MG." figureDoi="http://doi.org/10.5281/zenodo.5271074" httpUri="https://zenodo.org/record/5271074/files/figure.png" pageId="64" pageNumber="65">28D, 28G</figureCitation>
|
||
<figureCitation id="BFA12A1CFFB26869FC55DAD6EED0A784" box="[989,997,709,723]" captionStart="FIGURES 2–3" captionStartId="5.[151,264,1935,1957]" captionTargetBox="[166,1410,193,1889]" captionTargetId="figure-62@5.[166,1410,193,1889]" captionTargetPageId="5" captionText="FIGURES 2–3. Enlarged maps of Palau islands showing collecting sites. Top (Fig. 2), map of southern part of Babeldaob island; bottom (Fig. 3), map of Koror island and its neighboring islands or islets. Abbreviations of the collecting sites are listed in Table 2." figureDoi="http://doi.org/10.5281/zenodo.5271040" httpUri="https://zenodo.org/record/5271040/files/figure.png" pageId="64" pageNumber="65">
|
||
<subScript id="BB1E34DCFFB26869FC55DAD6EED0A784" attach="right" box="[989,997,709,723]" fontSize="6" pageId="64" pageNumber="65">3</subScript>
|
||
</figureCitation>
|
||
). Whereas, the nominal subspecies has much wider wing-like projections on these whorls, which are overlapped with and attaching to those on the previous whorls (
|
||
<figureCitation id="BFA12A1CFFB26869FED4DB12ECFBA64D" box="[348,462,769,794]" captionStart="FIGURE 27" captionStartId="58.[151,249,1829,1851]" captionTargetBox="[164,1423,193,1807]" captionTargetId="figure-150@58.[163,1423,193,1807]" captionTargetPageId="58" captionText="FIGURE 27. Hungerfordia papilio papilio sp. et subsp. nov. (A1–A4) A paratype from Loc. Gk4-3, B0374a; (B) columella (dorsal view), (C) axial ribs above the aperture, and (D) sculpture on the last 2–4 whorls of paratypes from Loc. Gk4-3 (B0374a, b); (E) a paratype after removal of the winged ribs, Loc. Gk4-3, B0374c; (F) a paratype from Gk4-3, B0374d; (G1– G4) holotype, B0434T; (H) a paratype from Loc. Gk6-3, B0858; (I) a paratypes from Loc. Gk7-3, B0963; (J) umbilical view of a paratype after removal of the winged ribs, Loc. Gk4-3, B0374c; (K1, K2) outer surface, and (K3, K4) inner surface of operculum of a paratype, Loc. Gk4-3, B0374. Scale bars, 1mm (A–J), 0.5 mm (K). All specimens, UMUTZ-MG." figureDoi="http://doi.org/10.5281/zenodo.5271072" httpUri="https://zenodo.org/record/5271072/files/figure.png" pageId="64" pageNumber="65">Figs. 27A</figureCitation>
|
||
<figureCitation id="BFA12A1CFFB26869FE45DB03ECE0A649" box="[461,469,784,798]" captionStart="FIGURE 1" captionStartId="4.[151,250,1978,2000]" captionTargetBox="[187,1407,193,1949]" captionTargetId="figure-53@4.[187,1407,193,1949]" captionTargetPageId="4" captionText="FIGURE 1. Map of Palau islands showing collecting sites. Small red circles show collecting sites. Areas encircled by blue rectangles are shown as enlarged maps in Figs. 2–5. Abbreviations of the collecting sites are listed in Table 2." figureDoi="http://doi.org/10.5281/zenodo.5271038" httpUri="https://zenodo.org/record/5271038/files/figure.png" pageId="64" pageNumber="65">
|
||
<subScript id="BB1E34DCFFB26869FE45DB03ECE0A649" attach="left" box="[461,469,784,798]" fontSize="6" pageId="64" pageNumber="65">1</subScript>
|
||
</figureCitation>
|
||
,
|
||
<figureCitation id="BFA12A1CFFB26869FE6CDB12EF60A64D" box="[484,597,769,794]" captionStart="FIGURE 27" captionStartId="58.[151,249,1829,1851]" captionTargetBox="[164,1423,193,1807]" captionTargetId="figure-150@58.[163,1423,193,1807]" captionTargetPageId="58" captionText="FIGURE 27. Hungerfordia papilio papilio sp. et subsp. nov. (A1–A4) A paratype from Loc. Gk4-3, B0374a; (B) columella (dorsal view), (C) axial ribs above the aperture, and (D) sculpture on the last 2–4 whorls of paratypes from Loc. Gk4-3 (B0374a, b); (E) a paratype after removal of the winged ribs, Loc. Gk4-3, B0374c; (F) a paratype from Gk4-3, B0374d; (G1– G4) holotype, B0434T; (H) a paratype from Loc. Gk6-3, B0858; (I) a paratypes from Loc. Gk7-3, B0963; (J) umbilical view of a paratype after removal of the winged ribs, Loc. Gk4-3, B0374c; (K1, K2) outer surface, and (K3, K4) inner surface of operculum of a paratype, Loc. Gk4-3, B0374. Scale bars, 1mm (A–J), 0.5 mm (K). All specimens, UMUTZ-MG." figureDoi="http://doi.org/10.5281/zenodo.5271072" httpUri="https://zenodo.org/record/5271072/files/figure.png" pageId="64" pageNumber="65">27D, 27G</figureCitation>
|
||
<figureCitation id="BFA12A1CFFB26869FDDCDB03EF69A649" box="[596,604,784,798]" captionStart="FIGURES 2–3" captionStartId="5.[151,264,1935,1957]" captionTargetBox="[166,1410,193,1889]" captionTargetId="figure-62@5.[166,1410,193,1889]" captionTargetPageId="5" captionText="FIGURES 2–3. Enlarged maps of Palau islands showing collecting sites. Top (Fig. 2), map of southern part of Babeldaob island; bottom (Fig. 3), map of Koror island and its neighboring islands or islets. Abbreviations of the collecting sites are listed in Table 2." figureDoi="http://doi.org/10.5281/zenodo.5271040" httpUri="https://zenodo.org/record/5271040/files/figure.png" pageId="64" pageNumber="65">
|
||
<subScript id="BB1E34DCFFB26869FDDCDB03EF69A649" attach="left" box="[596,604,784,798]" fontSize="6" pageId="64" pageNumber="65">3</subScript>
|
||
</figureCitation>
|
||
). Second, the less preserved synchronization of the wing-like ribs is also diagnostic of this subspecies. In the nominal subspecies, wing-like ribs on all whorls except the last one are perfectly synchronized with those on the previous whorls (
|
||
<figureCitation id="BFA12A1CFFB26869FCC7DB5FEEF6A632" box="[847,963,844,869]" captionStart="FIGURE 27" captionStartId="58.[151,249,1829,1851]" captionTargetBox="[164,1423,193,1807]" captionTargetId="figure-150@58.[163,1423,193,1807]" captionTargetPageId="58" captionText="FIGURE 27. Hungerfordia papilio papilio sp. et subsp. nov. (A1–A4) A paratype from Loc. Gk4-3, B0374a; (B) columella (dorsal view), (C) axial ribs above the aperture, and (D) sculpture on the last 2–4 whorls of paratypes from Loc. Gk4-3 (B0374a, b); (E) a paratype after removal of the winged ribs, Loc. Gk4-3, B0374c; (F) a paratype from Gk4-3, B0374d; (G1– G4) holotype, B0434T; (H) a paratype from Loc. Gk6-3, B0858; (I) a paratypes from Loc. Gk7-3, B0963; (J) umbilical view of a paratype after removal of the winged ribs, Loc. Gk4-3, B0374c; (K1, K2) outer surface, and (K3, K4) inner surface of operculum of a paratype, Loc. Gk4-3, B0374. Scale bars, 1mm (A–J), 0.5 mm (K). All specimens, UMUTZ-MG." figureDoi="http://doi.org/10.5281/zenodo.5271072" httpUri="https://zenodo.org/record/5271072/files/figure.png" pageId="64" pageNumber="65">Figs. 27A</figureCitation>
|
||
<figureCitation id="BFA12A1CFFB26869FC4BDB48EEFEA63E" box="[963,971,859,873]" captionStart="FIGURES 2–3" captionStartId="5.[151,264,1935,1957]" captionTargetBox="[166,1410,193,1889]" captionTargetId="figure-62@5.[166,1410,193,1889]" captionTargetPageId="5" captionText="FIGURES 2–3. Enlarged maps of Palau islands showing collecting sites. Top (Fig. 2), map of southern part of Babeldaob island; bottom (Fig. 3), map of Koror island and its neighboring islands or islets. Abbreviations of the collecting sites are listed in Table 2." figureDoi="http://doi.org/10.5281/zenodo.5271040" httpUri="https://zenodo.org/record/5271040/files/figure.png" pageId="64" pageNumber="65">
|
||
<subScript id="BB1E34DCFFB26869FC4BDB48EEFEA63E" attach="left" box="[963,971,859,873]" fontSize="6" pageId="64" pageNumber="65">3</subScript>
|
||
</figureCitation>
|
||
,
|
||
<figureCitation id="BFA12A1CFFB26869FC56DB5FE964A632" box="[990,1105,844,869]" captionStart="FIGURE 27" captionStartId="58.[151,249,1829,1851]" captionTargetBox="[164,1423,193,1807]" captionTargetId="figure-150@58.[163,1423,193,1807]" captionTargetPageId="58" captionText="FIGURE 27. Hungerfordia papilio papilio sp. et subsp. nov. (A1–A4) A paratype from Loc. Gk4-3, B0374a; (B) columella (dorsal view), (C) axial ribs above the aperture, and (D) sculpture on the last 2–4 whorls of paratypes from Loc. Gk4-3 (B0374a, b); (E) a paratype after removal of the winged ribs, Loc. Gk4-3, B0374c; (F) a paratype from Gk4-3, B0374d; (G1– G4) holotype, B0434T; (H) a paratype from Loc. Gk6-3, B0858; (I) a paratypes from Loc. Gk7-3, B0963; (J) umbilical view of a paratype after removal of the winged ribs, Loc. Gk4-3, B0374c; (K1, K2) outer surface, and (K3, K4) inner surface of operculum of a paratype, Loc. Gk4-3, B0374. Scale bars, 1mm (A–J), 0.5 mm (K). All specimens, UMUTZ-MG." figureDoi="http://doi.org/10.5281/zenodo.5271072" httpUri="https://zenodo.org/record/5271072/files/figure.png" pageId="64" pageNumber="65">27D, 27G</figureCitation>
|
||
<figureCitation id="BFA12A1CFFB26869FBD9DB48E96CA63E" box="[1105,1113,859,873]" captionStart="FIGURE 1" captionStartId="4.[151,250,1978,2000]" captionTargetBox="[187,1407,193,1949]" captionTargetId="figure-53@4.[187,1407,193,1949]" captionTargetPageId="4" captionText="FIGURE 1. Map of Palau islands showing collecting sites. Small red circles show collecting sites. Areas encircled by blue rectangles are shown as enlarged maps in Figs. 2–5. Abbreviations of the collecting sites are listed in Table 2." figureDoi="http://doi.org/10.5281/zenodo.5271038" httpUri="https://zenodo.org/record/5271038/files/figure.png" pageId="64" pageNumber="65">
|
||
<subScript id="BB1E34DCFFB26869FBD9DB48E96CA63E" attach="left" box="[1105,1113,859,873]" fontSize="6" pageId="64" pageNumber="65">1</subScript>
|
||
</figureCitation>
|
||
). In contrast, in
|
||
<emphasis id="15EEEA8BFFB26869FAADDB5EEC21A6DC" italics="true" pageId="64" pageNumber="65">
|
||
<taxonomicName id="E09A4D1AFFB26869FAADDB5EEC25A6DC" authorityName="YAMAZAKI & YAMAZAKI & UESHIMA" authorityYear="2013" class="Gastropoda" family="Diplommatinidae" genus="Hungerfordia" kingdom="Animalia" order="Architaenioglossa" pageId="64" pageNumber="65" phylum="Mollusca" rank="subSpecies" species="papilio" subSpecies="stenoptera">H. papilio stenoptera</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
non-synchronized ribs occasionally occur on the apical whorls, and are frequently found on the penultimate and antepenultimate whorls (
|
||
<figureCitation id="BFA12A1CFFB26869FDFADB84EFD0A6E7" box="[626,741,919,944]" captionStart="FIGURE 28" captionStartId="62.[151,249,1859,1881]" captionTargetBox="[151,1436,193,1837]" captionTargetId="figure-128@62.[151,1436,193,1837]" captionTargetPageId="62" captionText="FIGURE 28. Hungerfordia papilio stenoptera subsp. nov. (A1–A4) A topotypic paratype (B0223a); (B) columella (dorsal view), (C) axial ribs above the aperture, and (D) sculpture on the last 2–4 whorls of topotypic paratypes (B0223a, b); (E) a topotypic paratype after removal of the winged ribs, B0223c; (F) a paratype from Loc. Ik3-1, B0212; (G1–G4) holotype, B0223T; (H) a paratype from Loc. Mc4-1, B0334; (I) a paratype from Loc. Mc2-1, B0236; (J1, J2) outer surface, and (J3, J4) inner surface of operculum of a topotypic paratype, B0223. Scale bars 1mm (A–I), 0.5 mm (J). All specimens, UMUTZ-MG." figureDoi="http://doi.org/10.5281/zenodo.5271074" httpUri="https://zenodo.org/record/5271074/files/figure.png" pageId="64" pageNumber="65">Figs. 28A</figureCitation>
|
||
<figureCitation id="BFA12A1CFFB26869FD6CDBB5EFD9A6E3" box="[740,748,934,948]" captionStart="FIGURES 2–3" captionStartId="5.[151,264,1935,1957]" captionTargetBox="[166,1410,193,1889]" captionTargetId="figure-62@5.[166,1410,193,1889]" captionTargetPageId="5" captionText="FIGURES 2–3. Enlarged maps of Palau islands showing collecting sites. Top (Fig. 2), map of southern part of Babeldaob island; bottom (Fig. 3), map of Koror island and its neighboring islands or islets. Abbreviations of the collecting sites are listed in Table 2." figureDoi="http://doi.org/10.5281/zenodo.5271040" httpUri="https://zenodo.org/record/5271040/files/figure.png" pageId="64" pageNumber="65">
|
||
<subScript id="BB1E34DCFFB26869FD6CDBB5EFD9A6E3" attach="left" box="[740,748,934,948]" fontSize="6" pageId="64" pageNumber="65">3</subScript>
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||
</figureCitation>
|
||
,
|
||
<figureCitation id="BFA12A1CFFB26869FD75DB84EE5BA6E7" box="[765,878,919,944]" captionStart="FIGURE 28" captionStartId="62.[151,249,1859,1881]" captionTargetBox="[151,1436,193,1837]" captionTargetId="figure-128@62.[151,1436,193,1837]" captionTargetPageId="62" captionText="FIGURE 28. Hungerfordia papilio stenoptera subsp. nov. (A1–A4) A topotypic paratype (B0223a); (B) columella (dorsal view), (C) axial ribs above the aperture, and (D) sculpture on the last 2–4 whorls of topotypic paratypes (B0223a, b); (E) a topotypic paratype after removal of the winged ribs, B0223c; (F) a paratype from Loc. Ik3-1, B0212; (G1–G4) holotype, B0223T; (H) a paratype from Loc. Mc4-1, B0334; (I) a paratype from Loc. Mc2-1, B0236; (J1, J2) outer surface, and (J3, J4) inner surface of operculum of a topotypic paratype, B0223. Scale bars 1mm (A–I), 0.5 mm (J). All specimens, UMUTZ-MG." figureDoi="http://doi.org/10.5281/zenodo.5271074" httpUri="https://zenodo.org/record/5271074/files/figure.png" pageId="64" pageNumber="65">28D, 28G</figureCitation>
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<figureCitation id="BFA12A1CFFB26869FCE6DBB5EE43A6E3" box="[878,886,934,948]" captionStart="FIGURE 1" captionStartId="4.[151,250,1978,2000]" captionTargetBox="[187,1407,193,1949]" captionTargetId="figure-53@4.[187,1407,193,1949]" captionTargetPageId="4" captionText="FIGURE 1. Map of Palau islands showing collecting sites. Small red circles show collecting sites. Areas encircled by blue rectangles are shown as enlarged maps in Figs. 2–5. Abbreviations of the collecting sites are listed in Table 2." figureDoi="http://doi.org/10.5281/zenodo.5271038" httpUri="https://zenodo.org/record/5271038/files/figure.png" pageId="64" pageNumber="65">
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<subScript id="BB1E34DCFFB26869FCE6DBB5EE43A6E3" attach="left" box="[878,886,934,948]" fontSize="6" pageId="64" pageNumber="65">1</subScript>
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||
</figureCitation>
|
||
). Third, in
|
||
<emphasis id="15EEEA8BFFB26869FC77DB8BE9CFA6F8" box="[1023,1274,919,943]" italics="true" pageId="64" pageNumber="65">
|
||
<taxonomicName id="E09A4D1AFFB26869FC77DB8BE9C3A6F8" authorityName="YAMAZAKI & YAMAZAKI & UESHIMA" authorityYear="2013" box="[1023,1270,919,943]" class="Gastropoda" family="Diplommatinidae" genus="Hungerfordia" kingdom="Animalia" order="Architaenioglossa" pageId="64" pageNumber="65" phylum="Mollusca" rank="subSpecies" species="papilio" subSpecies="stenoptera">H. papilio stenoptera</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
the wing-like projections of ribs are more strongly folded at the periphery than the nominal subspecies, particularly on the penultimate and antepenultimate whorls (
|
||
<figureCitation id="BFA12A1CFFB26869FDF6DBF1EFC6A6AC" box="[638,755,994,1019]" captionStart="FIGURE 27" captionStartId="58.[151,249,1829,1851]" captionTargetBox="[164,1423,193,1807]" captionTargetId="figure-150@58.[163,1423,193,1807]" captionTargetPageId="58" captionText="FIGURE 27. Hungerfordia papilio papilio sp. et subsp. nov. (A1–A4) A paratype from Loc. Gk4-3, B0374a; (B) columella (dorsal view), (C) axial ribs above the aperture, and (D) sculpture on the last 2–4 whorls of paratypes from Loc. Gk4-3 (B0374a, b); (E) a paratype after removal of the winged ribs, Loc. Gk4-3, B0374c; (F) a paratype from Gk4-3, B0374d; (G1– G4) holotype, B0434T; (H) a paratype from Loc. Gk6-3, B0858; (I) a paratypes from Loc. Gk7-3, B0963; (J) umbilical view of a paratype after removal of the winged ribs, Loc. Gk4-3, B0374c; (K1, K2) outer surface, and (K3, K4) inner surface of operculum of a paratype, Loc. Gk4-3, B0374. Scale bars, 1mm (A–J), 0.5 mm (K). All specimens, UMUTZ-MG." figureDoi="http://doi.org/10.5281/zenodo.5271072" httpUri="https://zenodo.org/record/5271072/files/figure.png" pageId="64" pageNumber="65">Figs. 27A</figureCitation>
|
||
<figureCitation id="BFA12A1CFFB26869FD7BDBE2EFCEA6A8" box="[755,763,1009,1023]" captionStart="FIGURES 2–3" captionStartId="5.[151,264,1935,1957]" captionTargetBox="[166,1410,193,1889]" captionTargetId="figure-62@5.[166,1410,193,1889]" captionTargetPageId="5" captionText="FIGURES 2–3. Enlarged maps of Palau islands showing collecting sites. Top (Fig. 2), map of southern part of Babeldaob island; bottom (Fig. 3), map of Koror island and its neighboring islands or islets. Abbreviations of the collecting sites are listed in Table 2." figureDoi="http://doi.org/10.5281/zenodo.5271040" httpUri="https://zenodo.org/record/5271040/files/figure.png" pageId="64" pageNumber="65">
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<subScript id="BB1E34DCFFB26869FD7BDBE2EFCEA6A8" attach="left" box="[755,763,1009,1023]" fontSize="6" pageId="64" pageNumber="65">3</subScript>
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||
</figureCitation>
|
||
,
|
||
<figureCitation id="BFA12A1CFFB26869FC86DBF1EE74A6AC" box="[782,833,994,1019]" captionStart="FIGURE 27" captionStartId="58.[151,249,1829,1851]" captionTargetBox="[164,1423,193,1807]" captionTargetId="figure-150@58.[163,1423,193,1807]" captionTargetPageId="58" captionText="FIGURE 27. Hungerfordia papilio papilio sp. et subsp. nov. (A1–A4) A paratype from Loc. Gk4-3, B0374a; (B) columella (dorsal view), (C) axial ribs above the aperture, and (D) sculpture on the last 2–4 whorls of paratypes from Loc. Gk4-3 (B0374a, b); (E) a paratype after removal of the winged ribs, Loc. Gk4-3, B0374c; (F) a paratype from Gk4-3, B0374d; (G1– G4) holotype, B0434T; (H) a paratype from Loc. Gk6-3, B0858; (I) a paratypes from Loc. Gk7-3, B0963; (J) umbilical view of a paratype after removal of the winged ribs, Loc. Gk4-3, B0374c; (K1, K2) outer surface, and (K3, K4) inner surface of operculum of a paratype, Loc. Gk4-3, B0374. Scale bars, 1mm (A–J), 0.5 mm (K). All specimens, UMUTZ-MG." figureDoi="http://doi.org/10.5281/zenodo.5271072" httpUri="https://zenodo.org/record/5271072/files/figure.png" pageId="64" pageNumber="65">27D</figureCitation>
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,
|
||
<figureCitation id="BFA12A1CFFB26869FCDADBF1EEB7A6AC" box="[850,898,994,1019]" captionStart="FIGURE 28" captionStartId="62.[151,249,1859,1881]" captionTargetBox="[151,1436,193,1837]" captionTargetId="figure-128@62.[151,1436,193,1837]" captionTargetPageId="62" captionText="FIGURE 28. Hungerfordia papilio stenoptera subsp. nov. (A1–A4) A topotypic paratype (B0223a); (B) columella (dorsal view), (C) axial ribs above the aperture, and (D) sculpture on the last 2–4 whorls of topotypic paratypes (B0223a, b); (E) a topotypic paratype after removal of the winged ribs, B0223c; (F) a paratype from Loc. Ik3-1, B0212; (G1–G4) holotype, B0223T; (H) a paratype from Loc. Mc4-1, B0334; (I) a paratype from Loc. Mc2-1, B0236; (J1, J2) outer surface, and (J3, J4) inner surface of operculum of a topotypic paratype, B0223. Scale bars 1mm (A–I), 0.5 mm (J). All specimens, UMUTZ-MG." figureDoi="http://doi.org/10.5281/zenodo.5271074" httpUri="https://zenodo.org/record/5271074/files/figure.png" pageId="64" pageNumber="65">28A</figureCitation>
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<figureCitation id="BFA12A1CFFB26869FC0ADBE2EEBFA6A8" box="[898,906,1009,1023]" captionStart="FIGURES 2–3" captionStartId="5.[151,264,1935,1957]" captionTargetBox="[166,1410,193,1889]" captionTargetId="figure-62@5.[166,1410,193,1889]" captionTargetPageId="5" captionText="FIGURES 2–3. Enlarged maps of Palau islands showing collecting sites. Top (Fig. 2), map of southern part of Babeldaob island; bottom (Fig. 3), map of Koror island and its neighboring islands or islets. Abbreviations of the collecting sites are listed in Table 2." figureDoi="http://doi.org/10.5281/zenodo.5271040" httpUri="https://zenodo.org/record/5271040/files/figure.png" pageId="64" pageNumber="65">
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<subScript id="BB1E34DCFFB26869FC0ADBE2EEBFA6A8" attach="left" box="[898,906,1009,1023]" fontSize="6" pageId="64" pageNumber="65">3</subScript>
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</figureCitation>
|
||
,
|
||
<figureCitation id="BFA12A1CFFB26869FC16DBF1EEE7A6AC" box="[926,978,994,1019]" captionStart="FIGURE 28" captionStartId="62.[151,249,1859,1881]" captionTargetBox="[151,1436,193,1837]" captionTargetId="figure-128@62.[151,1436,193,1837]" captionTargetPageId="62" captionText="FIGURE 28. Hungerfordia papilio stenoptera subsp. nov. (A1–A4) A topotypic paratype (B0223a); (B) columella (dorsal view), (C) axial ribs above the aperture, and (D) sculpture on the last 2–4 whorls of topotypic paratypes (B0223a, b); (E) a topotypic paratype after removal of the winged ribs, B0223c; (F) a paratype from Loc. Ik3-1, B0212; (G1–G4) holotype, B0223T; (H) a paratype from Loc. Mc4-1, B0334; (I) a paratype from Loc. Mc2-1, B0236; (J1, J2) outer surface, and (J3, J4) inner surface of operculum of a topotypic paratype, B0223. Scale bars 1mm (A–I), 0.5 mm (J). All specimens, UMUTZ-MG." figureDoi="http://doi.org/10.5281/zenodo.5271074" httpUri="https://zenodo.org/record/5271074/files/figure.png" pageId="64" pageNumber="65">28D</figureCitation>
|
||
). Fourth, in
|
||
<taxonomicName id="E09A4D1AFFB26869FBE4DBF0E853A6AD" authorityName="YAMAZAKI & YAMAZAKI & UESHIMA" authorityYear="2013" box="[1132,1382,994,1018]" class="Gastropoda" family="Diplommatinidae" genus="Hungerfordia" kingdom="Animalia" order="Architaenioglossa" pageId="64" pageNumber="65" phylum="Mollusca" rank="subSpecies" species="papilio" subSpecies="stenoptera">
|
||
<emphasis id="15EEEA8BFFB26869FBE4DBF0E853A6AD" box="[1132,1382,994,1018]" italics="true" pageId="64" pageNumber="65">H. papilio stenoptera</emphasis>
|
||
</taxonomicName>
|
||
, the angular margin of the inner peristome is more strongly protruded (
|
||
<figureCitation id="BFA12A1CFFB26869FCF3DC1BEEDFA176" box="[891,1002,1032,1057]" captionStart="FIGURE 27" captionStartId="58.[151,249,1829,1851]" captionTargetBox="[164,1423,193,1807]" captionTargetId="figure-150@58.[163,1423,193,1807]" captionTargetPageId="58" captionText="FIGURE 27. Hungerfordia papilio papilio sp. et subsp. nov. (A1–A4) A paratype from Loc. Gk4-3, B0374a; (B) columella (dorsal view), (C) axial ribs above the aperture, and (D) sculpture on the last 2–4 whorls of paratypes from Loc. Gk4-3 (B0374a, b); (E) a paratype after removal of the winged ribs, Loc. Gk4-3, B0374c; (F) a paratype from Gk4-3, B0374d; (G1– G4) holotype, B0434T; (H) a paratype from Loc. Gk6-3, B0858; (I) a paratypes from Loc. Gk7-3, B0963; (J) umbilical view of a paratype after removal of the winged ribs, Loc. Gk4-3, B0374c; (K1, K2) outer surface, and (K3, K4) inner surface of operculum of a paratype, Loc. Gk4-3, B0374. Scale bars, 1mm (A–J), 0.5 mm (K). All specimens, UMUTZ-MG." figureDoi="http://doi.org/10.5281/zenodo.5271072" httpUri="https://zenodo.org/record/5271072/files/figure.png" pageId="64" pageNumber="65">Figs. 27A</figureCitation>
|
||
<figureCitation id="BFA12A1CFFB26869FC62DC04EEC7A172" box="[1002,1010,1047,1061]" captionStart="FIGURES 4–5" captionStartId="6.[151,264,1911,1933]" captionTargetBox="[172,1416,193,1890]" captionTargetId="figure-64@6.[172,1416,193,1890]" captionTargetPageId="6" captionText="FIGURES 4–5. Enlarged maps of Palau islands showing collecting sites. Top (Fig. 4), map of Ngerkutabel island and its neighboring islands or islets; bottom (Fig. 5), map of Mecherchar island and its neighboring islands or islets. Abbreviations of the collecting sites are listed in Table 2." figureDoi="http://doi.org/10.5281/zenodo.5271042" httpUri="https://zenodo.org/record/5271042/files/figure.png" pageId="64" pageNumber="65">
|
||
<subScript id="BB1E34DCFFB26869FC62DC04EEC7A172" attach="left" box="[1002,1010,1047,1061]" fontSize="6" pageId="64" pageNumber="65">4</subScript>
|
||
</figureCitation>
|
||
,
|
||
<figureCitation id="BFA12A1CFFB26869FC77DC1BE905A176" box="[1023,1072,1032,1057]" captionStart="FIGURE 28" captionStartId="62.[151,249,1859,1881]" captionTargetBox="[151,1436,193,1837]" captionTargetId="figure-128@62.[151,1436,193,1837]" captionTargetPageId="62" captionText="FIGURE 28. Hungerfordia papilio stenoptera subsp. nov. (A1–A4) A topotypic paratype (B0223a); (B) columella (dorsal view), (C) axial ribs above the aperture, and (D) sculpture on the last 2–4 whorls of topotypic paratypes (B0223a, b); (E) a topotypic paratype after removal of the winged ribs, B0223c; (F) a paratype from Loc. Ik3-1, B0212; (G1–G4) holotype, B0223T; (H) a paratype from Loc. Mc4-1, B0334; (I) a paratype from Loc. Mc2-1, B0236; (J1, J2) outer surface, and (J3, J4) inner surface of operculum of a topotypic paratype, B0223. Scale bars 1mm (A–I), 0.5 mm (J). All specimens, UMUTZ-MG." figureDoi="http://doi.org/10.5281/zenodo.5271074" httpUri="https://zenodo.org/record/5271074/files/figure.png" pageId="64" pageNumber="65">28A</figureCitation>
|
||
<figureCitation id="BFA12A1CFFB26869FBA7DC04E902A172" box="[1071,1079,1047,1061]" captionStart="FIGURES 4–5" captionStartId="6.[151,264,1911,1933]" captionTargetBox="[172,1416,193,1890]" captionTargetId="figure-64@6.[172,1416,193,1890]" captionTargetPageId="6" captionText="FIGURES 4–5. Enlarged maps of Palau islands showing collecting sites. Top (Fig. 4), map of Ngerkutabel island and its neighboring islands or islets; bottom (Fig. 5), map of Mecherchar island and its neighboring islands or islets. Abbreviations of the collecting sites are listed in Table 2." figureDoi="http://doi.org/10.5281/zenodo.5271042" httpUri="https://zenodo.org/record/5271042/files/figure.png" pageId="64" pageNumber="65">
|
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<subScript id="BB1E34DCFFB26869FBA7DC04E902A172" attach="left" box="[1071,1079,1047,1061]" fontSize="6" pageId="64" pageNumber="65">4</subScript>
|
||
</figureCitation>
|
||
), and thus, the palatal margin is more strongly concave in left lateral view (
|
||
<figureCitation id="BFA12A1CFFB26869FD19DC3CEE31A11F" box="[657,772,1071,1096]" captionStart="FIGURE 27" captionStartId="58.[151,249,1829,1851]" captionTargetBox="[164,1423,193,1807]" captionTargetId="figure-150@58.[163,1423,193,1807]" captionTargetPageId="58" captionText="FIGURE 27. Hungerfordia papilio papilio sp. et subsp. nov. (A1–A4) A paratype from Loc. Gk4-3, B0374a; (B) columella (dorsal view), (C) axial ribs above the aperture, and (D) sculpture on the last 2–4 whorls of paratypes from Loc. Gk4-3 (B0374a, b); (E) a paratype after removal of the winged ribs, Loc. Gk4-3, B0374c; (F) a paratype from Gk4-3, B0374d; (G1– G4) holotype, B0434T; (H) a paratype from Loc. Gk6-3, B0858; (I) a paratypes from Loc. Gk7-3, B0963; (J) umbilical view of a paratype after removal of the winged ribs, Loc. Gk4-3, B0374c; (K1, K2) outer surface, and (K3, K4) inner surface of operculum of a paratype, Loc. Gk4-3, B0374. Scale bars, 1mm (A–J), 0.5 mm (K). All specimens, UMUTZ-MG." figureDoi="http://doi.org/10.5281/zenodo.5271072" httpUri="https://zenodo.org/record/5271072/files/figure.png" pageId="64" pageNumber="65">Figs. 27A</figureCitation>
|
||
<figureCitation id="BFA12A1CFFB26869FC8CDC2DEE39A11B" box="[772,780,1086,1100]" captionStart="FIGURES 2–3" captionStartId="5.[151,264,1935,1957]" captionTargetBox="[166,1410,193,1889]" captionTargetId="figure-62@5.[166,1410,193,1889]" captionTargetPageId="5" captionText="FIGURES 2–3. Enlarged maps of Palau islands showing collecting sites. Top (Fig. 2), map of southern part of Babeldaob island; bottom (Fig. 3), map of Koror island and its neighboring islands or islets. Abbreviations of the collecting sites are listed in Table 2." figureDoi="http://doi.org/10.5281/zenodo.5271040" httpUri="https://zenodo.org/record/5271040/files/figure.png" pageId="64" pageNumber="65">
|
||
<subScript id="BB1E34DCFFB26869FC8CDC2DEE39A11B" attach="left" box="[772,780,1086,1100]" fontSize="6" pageId="64" pageNumber="65">2</subScript>
|
||
</figureCitation>
|
||
,
|
||
<figureCitation id="BFA12A1CFFB26869FC95DC3CEE78A11F" box="[797,845,1071,1096]" captionStart="FIGURE 28" captionStartId="62.[151,249,1859,1881]" captionTargetBox="[151,1436,193,1837]" captionTargetId="figure-128@62.[151,1436,193,1837]" captionTargetPageId="62" captionText="FIGURE 28. Hungerfordia papilio stenoptera subsp. nov. (A1–A4) A topotypic paratype (B0223a); (B) columella (dorsal view), (C) axial ribs above the aperture, and (D) sculpture on the last 2–4 whorls of topotypic paratypes (B0223a, b); (E) a topotypic paratype after removal of the winged ribs, B0223c; (F) a paratype from Loc. Ik3-1, B0212; (G1–G4) holotype, B0223T; (H) a paratype from Loc. Mc4-1, B0334; (I) a paratype from Loc. Mc2-1, B0236; (J1, J2) outer surface, and (J3, J4) inner surface of operculum of a topotypic paratype, B0223. Scale bars 1mm (A–I), 0.5 mm (J). All specimens, UMUTZ-MG." figureDoi="http://doi.org/10.5281/zenodo.5271074" httpUri="https://zenodo.org/record/5271074/files/figure.png" pageId="64" pageNumber="65">28A</figureCitation>
|
||
<figureCitation id="BFA12A1CFFB26869FCC5DC2DEE60A11B" box="[845,853,1086,1100]" captionStart="FIGURES 2–3" captionStartId="5.[151,264,1935,1957]" captionTargetBox="[166,1410,193,1889]" captionTargetId="figure-62@5.[166,1410,193,1889]" captionTargetPageId="5" captionText="FIGURES 2–3. Enlarged maps of Palau islands showing collecting sites. Top (Fig. 2), map of southern part of Babeldaob island; bottom (Fig. 3), map of Koror island and its neighboring islands or islets. Abbreviations of the collecting sites are listed in Table 2." figureDoi="http://doi.org/10.5281/zenodo.5271040" httpUri="https://zenodo.org/record/5271040/files/figure.png" pageId="64" pageNumber="65">
|
||
<subScript id="BB1E34DCFFB26869FCC5DC2DEE60A11B" attach="left" box="[845,853,1086,1100]" fontSize="6" pageId="64" pageNumber="65">2</subScript>
|
||
</figureCitation>
|
||
). Fifth, the shell size is generally smaller and the umbilicus tends to be narrower compared to the nominal subspecies.
|
||
</paragraph>
|
||
<paragraph id="27253699FFB26869FF4DDC69EE4DA1E0" blockId="64.[151,1436,151,1279]" pageId="64" pageNumber="65">Mecherchar and Euidelchol populations are almost fixed for the characteristic phenotype, although there are few populations showing intermediate or exceptional phenotypes.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6F806512FFB26869FF4DDCD1E925A1A8" pageId="64" pageNumber="65" type="etymology">
|
||
<paragraph id="27253699FFB26869FF4DDCD1E925A1A8" blockId="64.[151,1436,151,1279]" pageId="64" pageNumber="65">
|
||
<emphasis id="15EEEA8BFFB26869FF4DDCD1EC79A18C" bold="true" box="[197,332,1218,1243]" pageId="64" pageNumber="65">Etymology.</emphasis>
|
||
The subspecific name, derived from Greek
|
||
<emphasis id="15EEEA8BFFB26869FCCADCD0EEBDA18D" box="[834,904,1219,1242]" italics="true" pageId="64" pageNumber="65">stenos</emphasis>
|
||
(= narrow) and
|
||
<emphasis id="15EEEA8BFFB26869FBCBDCD0E9BCA18D" box="[1091,1161,1219,1242]" italics="true" pageId="64" pageNumber="65">pterus</emphasis>
|
||
(= with wing), refers to the narrower wing-like ribs on the last two whorls than the nominal subspecies.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |