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<document id="3CFBEBF8EA53699F890E58F6F7B678F1" ID-CLB-Dataset="30864" ID-DOI="10.11646/zootaxa.4377.2.6" ID-GBIF-Dataset="fb507bc4-0a9b-4caf-8f1e-d9d34aa6cd8e" ID-ISSN="1175-5326" ID-Zenodo-Dep="1163937" ID-ZooBank="961E0020-EE4A-4FE3-902C-0478F8AE1449" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1517471885320" checkinUser="plazi" docAuthor="Almeida, Flavio Roberto De Albuquerque, Nunes, Benedito Mendes &amp; Fernandes, Jose Antonio Marin" docDate="2018" docId="AF3087D9FF98FFFA5FFEFBE4FA7E8D80" docLanguage="en" docName="zootaxa.4377.2.6.pdf" docOrigin="Zootaxa 4377 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Edessinae Amyot &amp; Serville 1843" docType="key" docVersion="7" lastPageNumber="256" masterDocId="5309FFA1FF99FFF85F69FF85FFE28E27" masterDocTitle="A new genus and new species of Edessinae (Hemiptera: Heteroptera: Pentatomidae)" masterLastPageNumber="268" masterPageNumber="254" pageNumber="255" updateTime="1698518168574" updateUser="ExternalLinkService">
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<mods:title id="661ED628D185987119342CB6EC3C3C74">A new genus and new species of Edessinae (Hemiptera: Heteroptera: Pentatomidae)</mods:title>
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<mods:namePart id="7039A802624B264AC7C72E21CF12A720">Almeida, Flavio Roberto De Albuquerque</mods:namePart>
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<mods:namePart id="D9B25CFAD4F2FB16C4C2EFC4E9C4540B">Nunes, Benedito Mendes</mods:namePart>
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<mods:namePart id="6AB3D5D2EE32A0C7E76B78B62BAD09A9">Fernandes, Jose Antonio Marin</mods:namePart>
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<mods:title id="7FAAB15B431EA286A29ADE5B6C88E7AD">Zootaxa</mods:title>
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<mods:date id="5AF8DA2BBD63E914428405D0CE8E78C7">2018</mods:date>
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<mods:number id="516EC2567661BA5E657CF66503497DD7">2018-02-01</mods:number>
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<mods:number id="EFD220647FCACAC5B439FC6CCC89E6FE">4377</mods:number>
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<mods:identifier id="7F1D06C97BDF6D93964B20DE531A54AA" type="DOI">10.11646/zootaxa.4377.2.6</mods:identifier>
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<treatment id="AF3087D9FF98FFFA5FFEFBE4FA7E8D80" ID-DOI="http://doi.org/10.5281/zenodo.5979854" ID-GBIF-Taxon="150737139" ID-Zenodo-Dep="5979854" LSID="urn:lsid:plazi:treatment:AF3087D9FF98FFFA5FFEFBE4FA7E8D80" httpUri="http://treatment.plazi.org/id/AF3087D9FF98FFFA5FFEFBE4FA7E8D80" lastPageId="2" lastPageNumber="256" pageId="1" pageNumber="255">
<subSubSection id="6F836544FF98FFF95FFEFBE4FB2A8A5C" box="[151,1224,1121,1147]" pageId="1" pageNumber="255" type="nomenclature">
<paragraph id="272636CFFF98FFF95FFEFBE4FB2A8A5C" blockId="1.[151,1224,1121,1147]" box="[151,1224,1121,1147]" pageId="1" pageNumber="255">
<heading id="7C6E81A3FF98FFF95FFEFBE4FB2A8A5C" bold="true" box="[151,1224,1121,1147]" fontSize="11" level="1" pageId="1" pageNumber="255" reason="1">
<emphasis id="15EDEADDFF98FFF95FFEFBE4FB2A8A5C" bold="true" box="[151,1224,1121,1147]" pageId="1" pageNumber="255">
Key to genera of
<taxonomicName id="E0994D4CFF98FFF95E05FBE4FE0A8A5C" authorityName="Amyot &amp; Serville" authorityYear="1843" box="[364,488,1121,1147]" class="Insecta" family="Pentatomidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="255" phylum="Arthropoda" rank="subFamily" subFamily="Edessinae">Edessinae</taxonomicName>
and subgenera of
<taxonomicName id="E0994D4CFF98FFF95DA4FBE4FCC08A5C" authorityName="Fabricius" authorityYear="1803" box="[717,802,1121,1147]" class="Insecta" family="Pentatomidae" genus="Edessa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="255" phylum="Arthropoda" rank="genus">
<emphasis id="15EDEADDFF98FFF95DA4FBE4FCC08A5C" bold="true" box="[717,802,1121,1147]" italics="true" pageId="1" pageNumber="255">Edessa</emphasis>
</taxonomicName>
. Modified from
<bibRefCitation id="43084B3EFF98FFF95C87FBE4FB2A8A5C" author="Grazia" box="[1006,1224,1121,1147]" pageId="1" pageNumber="255" refString="Grazia, J., Panizzi, A. R., Greve, C., Schwertner, C. F., Campos, L. A., Garbelotto, T. A. &amp; Fernandes, J. A. M. (2015) Stink Bugs (Pentatomidae). In: Panizzi, A. R. &amp; Grazia, J. (Eds.), True bugs (Heteroptera) of the Neotropics. Vol. 2. Springer, Rotterdam, pp. 681 - 756. https: // doi. org / 10.1007 / 978 - 94 - 017 - 9861 - 7 _ 22" type="book" year="2015">
Grazia
<emphasis id="15EDEADDFF98FFF95B25FBE4FB9D8A5C" bold="true" box="[1100,1151,1121,1147]" italics="true" pageId="1" pageNumber="255">et al</emphasis>
. 2015
</bibRefCitation>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="6F836544FF98FFFA5FFEFB22FA7E8D80" lastPageId="2" lastPageNumber="256" pageId="1" pageNumber="255" type="key">
<keyStep id="9C6D2E6AFF98FFF95FFEFB22FA798B32" pageId="1" pageNumber="255">
<paragraph id="272636CFFF98FFF95FFEFB22FA748AFD" blockId="1.[151,1436,1191,2034]" pageId="1" pageNumber="255">
<keyLead id="9C6895FAFF98FFF95FFEFB22FA748AFD" pageId="1" pageNumber="255">
1. Body rounded; anterior apex of metasternal process not surpassing mesocoxae, sulcate but not clearly bifurcated (
<figureCitation id="BFA22A4AFF98FFF95A4AFB22FAB98A9A" box="[1315,1371,1191,1213]" captionStart="FIGURES 1–10. 1–4" captionStartId="4.[151,264,1881,1904]" captionTargetBox="[251,1335,195,1845]" captionTargetId="figure@4.[226,1365,195,1870]" captionTargetPageId="4" captionText="FIGURES 1–10. 1–4 Metasternal process; 1—Brachystethus geniculatus (Fabricius, 1787); 2—Peromatus bolivianus Fallou, 1887; 3—Pantochlora vivida Stål, 1870; 4—Doesburguedessa nigrolimbata Fernandes, 2010. 5–6 Dorsal view of head and pronotum; 5—Peromatus bolivianus; 6—Edessa (Pygoda) sp. 7–8 Dorsal view; 7—Olbia caprina (Stål, 1862); 8—Edessa (Hypoxys) boerneri Breddin, 1904. 9–10 Dorsal view of head and pronotum; 9—Grammedessa sp.; 10—Edessa cervus (Fabricius, 1787). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163939/files/figure.png" pageId="1" pageNumber="255">Fig. 1</figureCitation>
)......................................................................................
<taxonomicName id="E0994D4CFF98FFF95BD4FB41FA748AFD" authority="Laporte" authorityName="Laporte" authorityYear="1832" box="[1213,1430,1220,1242]" class="Insecta" family="Pentatomidae" genus="Brachystethus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="255" phylum="Arthropoda" rank="genus">
<emphasis id="15EDEADDFF98FFF95BD4FB41FAA78AFD" box="[1213,1349,1220,1242]" italics="true" pageId="1" pageNumber="255">Brachystethus</emphasis>
Laporte
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="272636CFFF98FFF95FFEFB67FA798B32" blockId="1.[151,1436,1191,2034]" pageId="1" pageNumber="255">
<keyLead id="9C6895FAFF98FFF95FFEFB67FA798B32" pageId="1" pageNumber="255">
- Body with different shapes but usually elongate, lateral margins subparallel; metasternal process clearly surpassing mesocoxae, anterior part with different shapes but never only sulcate (
<figureCitation id="BFA22A4AFF98FFF95C26FB7AFC4F8B32" box="[847,941,1279,1301]" captionStart="FIGURES 1–10. 1–4" captionStartId="4.[151,264,1881,1904]" captionTargetBox="[251,1335,195,1845]" captionTargetId="figure@4.[226,1365,195,1870]" captionTargetPageId="4" captionText="FIGURES 1–10. 1–4 Metasternal process; 1—Brachystethus geniculatus (Fabricius, 1787); 2—Peromatus bolivianus Fallou, 1887; 3—Pantochlora vivida Stål, 1870; 4—Doesburguedessa nigrolimbata Fernandes, 2010. 5–6 Dorsal view of head and pronotum; 5—Peromatus bolivianus; 6—Edessa (Pygoda) sp. 7–8 Dorsal view; 7—Olbia caprina (Stål, 1862); 8—Edessa (Hypoxys) boerneri Breddin, 1904. 9–10 Dorsal view of head and pronotum; 9—Grammedessa sp.; 10—Edessa cervus (Fabricius, 1787). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163939/files/figure.png" pageId="1" pageNumber="255">Figs. 2–4</figureCitation>
)........................................ 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9C6D2E6AFF98FFF95FFEFA99FA798B68" pageId="1" pageNumber="255">
<paragraph id="272636CFFF98FFF95FFEFA99FA798B15" blockId="1.[151,1436,1191,2034]" box="[151,1435,1308,1330]" pageId="1" pageNumber="255">
<keyLead id="9C6895FAFF98FFF95FFEFA99FA798B15" box="[151,1435,1308,1330]" pageId="1" pageNumber="255">
2. Metasternal process anteriorly clearly bifurcated (
<figureCitation id="BFA22A4AFF98FFF95DD7FA99FD1A8B15" box="[702,760,1308,1330]" captionStart="FIGURES 1–10. 1–4" captionStartId="4.[151,264,1881,1904]" captionTargetBox="[251,1335,195,1845]" captionTargetId="figure@4.[226,1365,195,1870]" captionTargetPageId="4" captionText="FIGURES 1–10. 1–4 Metasternal process; 1—Brachystethus geniculatus (Fabricius, 1787); 2—Peromatus bolivianus Fallou, 1887; 3—Pantochlora vivida Stål, 1870; 4—Doesburguedessa nigrolimbata Fernandes, 2010. 5–6 Dorsal view of head and pronotum; 5—Peromatus bolivianus; 6—Edessa (Pygoda) sp. 7–8 Dorsal view; 7—Olbia caprina (Stål, 1862); 8—Edessa (Hypoxys) boerneri Breddin, 1904. 9–10 Dorsal view of head and pronotum; 9—Grammedessa sp.; 10—Edessa cervus (Fabricius, 1787). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163939/files/figure.png" pageId="1" pageNumber="255">Fig. 2</figureCitation>
)...................................................... 4
</keyLead>
</paragraph>
<paragraph id="272636CFFF98FFF95FFEFABFFA798B68" blockId="1.[151,1436,1191,2034]" box="[151,1435,1337,1359]" pageId="1" pageNumber="255">
<keyLead id="9C6895FAFF98FFF95FFEFABFFA798B68" box="[151,1435,1337,1359]" pageId="1" pageNumber="255">
- Metasternal process projected anteriorly in a simple process (
<figureCitation id="BFA22A4AFF98FFF95C45FABCFC658B68" box="[812,903,1337,1359]" captionStart="FIGURES 1–10. 1–4" captionStartId="4.[151,264,1881,1904]" captionTargetBox="[251,1335,195,1845]" captionTargetId="figure@4.[226,1365,195,1870]" captionTargetPageId="4" captionText="FIGURES 1–10. 1–4 Metasternal process; 1—Brachystethus geniculatus (Fabricius, 1787); 2—Peromatus bolivianus Fallou, 1887; 3—Pantochlora vivida Stål, 1870; 4—Doesburguedessa nigrolimbata Fernandes, 2010. 5–6 Dorsal view of head and pronotum; 5—Peromatus bolivianus; 6—Edessa (Pygoda) sp. 7–8 Dorsal view; 7—Olbia caprina (Stål, 1862); 8—Edessa (Hypoxys) boerneri Breddin, 1904. 9–10 Dorsal view of head and pronotum; 9—Grammedessa sp.; 10—Edessa cervus (Fabricius, 1787). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163939/files/figure.png" pageId="1" pageNumber="255">Figs. 3, 4</figureCitation>
).......................................... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9C6D2E6AFF98FFF95FFEFAD2FA798BE2" pageId="1" pageNumber="255">
<paragraph id="272636CFFF98FFF95FFEFAD2FA7E8BAD" blockId="1.[151,1436,1191,2034]" pageId="1" pageNumber="255">
<keyLead id="9C6895FAFF98FFF95FFEFAD2FA7E8BAD" pageId="1" pageNumber="255">
3. Metasternal process long, surpassing procoxae (
<figureCitation id="BFA22A4AFF98FFF95DDAFAD2FD0C8B4A" box="[691,750,1367,1389]" captionStart="FIGURES 1–10. 1–4" captionStartId="4.[151,264,1881,1904]" captionTargetBox="[251,1335,195,1845]" captionTargetId="figure@4.[226,1365,195,1870]" captionTargetPageId="4" captionText="FIGURES 1–10. 1–4 Metasternal process; 1—Brachystethus geniculatus (Fabricius, 1787); 2—Peromatus bolivianus Fallou, 1887; 3—Pantochlora vivida Stål, 1870; 4—Doesburguedessa nigrolimbata Fernandes, 2010. 5–6 Dorsal view of head and pronotum; 5—Peromatus bolivianus; 6—Edessa (Pygoda) sp. 7–8 Dorsal view; 7—Olbia caprina (Stål, 1862); 8—Edessa (Hypoxys) boerneri Breddin, 1904. 9–10 Dorsal view of head and pronotum; 9—Grammedessa sp.; 10—Edessa cervus (Fabricius, 1787). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163939/files/figure.png" pageId="1" pageNumber="255">Fig. 3</figureCitation>
), laterally compressed, fused with sterna; humeral angles slightly projected and acute..........................................................................
<emphasis id="15EDEADDFF98FFF95B90FAF1FA938BAD" box="[1273,1393,1396,1418]" italics="true" pageId="1" pageNumber="255">Pantochlora</emphasis>
Stål
</keyLead>
</paragraph>
<paragraph id="272636CFFF98FFF95FFEFA17FA798BE2" blockId="1.[151,1436,1191,2034]" pageId="1" pageNumber="255">
<keyLead id="9C6895FAFF98FFF95FFEFA17FA798BE2" pageId="1" pageNumber="255">
- Metasternal process short, not reaching procoxae (
<figureCitation id="BFA22A4AFF98FFF95DA4FA14FCEB8B80" box="[717,777,1425,1447]" captionStart="FIGURES 1–10. 1–4" captionStartId="4.[151,264,1881,1904]" captionTargetBox="[251,1335,195,1845]" captionTargetId="figure@4.[226,1365,195,1870]" captionTargetPageId="4" captionText="FIGURES 1–10. 1–4 Metasternal process; 1—Brachystethus geniculatus (Fabricius, 1787); 2—Peromatus bolivianus Fallou, 1887; 3—Pantochlora vivida Stål, 1870; 4—Doesburguedessa nigrolimbata Fernandes, 2010. 5–6 Dorsal view of head and pronotum; 5—Peromatus bolivianus; 6—Edessa (Pygoda) sp. 7–8 Dorsal view; 7—Olbia caprina (Stål, 1862); 8—Edessa (Hypoxys) boerneri Breddin, 1904. 9–10 Dorsal view of head and pronotum; 9—Grammedessa sp.; 10—Edessa cervus (Fabricius, 1787). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163939/files/figure.png" pageId="1" pageNumber="255">Fig. 4</figureCitation>
), conical, not in contact with sterna; humeral angles projected laterally, apices sharp................................................................
<taxonomicName id="E0994D4CFF98FFF95BFCFA2AFA798BE2" authority="Fernandes" authorityName="Fernandes" authorityYear="2010" box="[1173,1435,1455,1477]" class="Insecta" family="Pentatomidae" genus="Doesburgedessa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="255" phylum="Arthropoda" rank="genus">
<emphasis id="15EDEADDFF98FFF95BFCFA2AFAD08BE2" box="[1173,1330,1455,1477]" italics="true" pageId="1" pageNumber="255">Doesburgedessa</emphasis>
Fernandes
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9C6D2E6AFF98FFF95FFEFA49FA79881D" pageId="1" pageNumber="255">
<paragraph id="272636CFFF98FFF95FFEFA49FA7E8BD8" blockId="1.[151,1436,1191,2034]" pageId="1" pageNumber="255">
<keyLead id="9C6895FAFF98FFF95FFEFA49FA7E8BD8" pageId="1" pageNumber="255">
4. Antennae four-segmented; second antennomer very long, at least three times longer than the first (
<figureCitation id="BFA22A4AFF98FFF95BE6FA49FB2B8BC5" box="[1167,1225,1484,1506]" captionStart="FIGURES 1–10. 1–4" captionStartId="4.[151,264,1881,1904]" captionTargetBox="[251,1335,195,1845]" captionTargetId="figure@4.[226,1365,195,1870]" captionTargetPageId="4" captionText="FIGURES 1–10. 1–4 Metasternal process; 1—Brachystethus geniculatus (Fabricius, 1787); 2—Peromatus bolivianus Fallou, 1887; 3—Pantochlora vivida Stål, 1870; 4—Doesburguedessa nigrolimbata Fernandes, 2010. 5–6 Dorsal view of head and pronotum; 5—Peromatus bolivianus; 6—Edessa (Pygoda) sp. 7–8 Dorsal view; 7—Olbia caprina (Stål, 1862); 8—Edessa (Hypoxys) boerneri Breddin, 1904. 9–10 Dorsal view of head and pronotum; 9—Grammedessa sp.; 10—Edessa cervus (Fabricius, 1787). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163939/files/figure.png" pageId="1" pageNumber="255">Fig. 5</figureCitation>
); humeral angles globose.........................................................................
<taxonomicName id="E0994D4CFF98FFF95BEAFA6FFA7E8BD8" authority="Amyot &amp; Serville" authorityName="Amyot &amp; Serville" authorityYear="1843" box="[1155,1436,1513,1535]" class="Insecta" family="Pentatomidae" genus="Peromatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="255" phylum="Arthropoda" rank="genus">
<emphasis id="15EDEADDFF98FFF95BEAFA6FFB088BD8" box="[1155,1258,1514,1535]" italics="true" pageId="1" pageNumber="255">Peromatus</emphasis>
Amyot &amp; Serville
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="272636CFFF98FFF95FFEF982FA79881D" blockId="1.[151,1436,1191,2034]" pageId="1" pageNumber="255">
<keyLead id="9C6895FAFF98FFF95FFEF982FA79881D" pageId="1" pageNumber="255">
- Antennae five-segmented (
<figureCitation id="BFA22A4AFF98FFF95E81F982FDC6883A" box="[488,548,1543,1565]" captionStart="FIGURES 1–10. 1–4" captionStartId="4.[151,264,1881,1904]" captionTargetBox="[251,1335,195,1845]" captionTargetId="figure@4.[226,1365,195,1870]" captionTargetPageId="4" captionText="FIGURES 1–10. 1–4 Metasternal process; 1—Brachystethus geniculatus (Fabricius, 1787); 2—Peromatus bolivianus Fallou, 1887; 3—Pantochlora vivida Stål, 1870; 4—Doesburguedessa nigrolimbata Fernandes, 2010. 5–6 Dorsal view of head and pronotum; 5—Peromatus bolivianus; 6—Edessa (Pygoda) sp. 7–8 Dorsal view; 7—Olbia caprina (Stål, 1862); 8—Edessa (Hypoxys) boerneri Breddin, 1904. 9–10 Dorsal view of head and pronotum; 9—Grammedessa sp.; 10—Edessa cervus (Fabricius, 1787). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163939/files/figure.png" pageId="1" pageNumber="255">Fig. 6</figureCitation>
); second antennomer short, at most twice longer than the first; humeral angles variable in shape............................................................................................... 5
</keyLead>
</paragraph>
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<keyStep id="9C6D2E6AFF98FFF95FFEF9C4FA7E88B5" pageId="1" pageNumber="255">
<paragraph id="272636CFFF98FFF95FFEF9C4FA7E8852" blockId="1.[151,1436,1191,2034]" pageId="1" pageNumber="255">
<keyLead id="9C6895FAFF98FFF95FFEF9C4FA7E8852" pageId="1" pageNumber="255">
5. Posterior margin of pronotum with a large pair of spines overlapping basal angles of scutellum (
<figureCitation id="BFA22A4AFF98FFF95BEAF9C4FB5D8870" box="[1155,1215,1601,1623]" captionStart="FIGURES 1–10. 1–4" captionStartId="4.[151,264,1881,1904]" captionTargetBox="[251,1335,195,1845]" captionTargetId="figure@4.[226,1365,195,1870]" captionTargetPageId="4" captionText="FIGURES 1–10. 1–4 Metasternal process; 1—Brachystethus geniculatus (Fabricius, 1787); 2—Peromatus bolivianus Fallou, 1887; 3—Pantochlora vivida Stål, 1870; 4—Doesburguedessa nigrolimbata Fernandes, 2010. 5–6 Dorsal view of head and pronotum; 5—Peromatus bolivianus; 6—Edessa (Pygoda) sp. 7–8 Dorsal view; 7—Olbia caprina (Stål, 1862); 8—Edessa (Hypoxys) boerneri Breddin, 1904. 9–10 Dorsal view of head and pronotum; 9—Grammedessa sp.; 10—Edessa cervus (Fabricius, 1787). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163939/files/figure.png" pageId="1" pageNumber="255">Fig. 7</figureCitation>
); humeral angles long, sinuous, and laterally directed.....................................................................
<emphasis id="15EDEADDFF98FFF95A53F9DAFA938852" box="[1338,1393,1631,1653]" italics="true" pageId="1" pageNumber="255">Olbia</emphasis>
Stål
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</paragraph>
<paragraph id="272636CFFF98FFF95FFEF9F8FA7E88B5" blockId="1.[151,1436,1191,2034]" box="[151,1436,1660,1682]" pageId="1" pageNumber="255">
<keyLead id="9C6895FAFF98FFF95FFEF9F8FA7E88B5" box="[151,1436,1660,1682]" pageId="1" pageNumber="255">
- Posterior margin of pronotum without large spines or other projections (
<figureCitation id="BFA22A4AFF98FFF95CFAF9F9FC2F88B5" box="[915,973,1660,1682]" captionStart="FIGURES 1–10. 1–4" captionStartId="4.[151,264,1881,1904]" captionTargetBox="[251,1335,195,1845]" captionTargetId="figure@4.[226,1365,195,1870]" captionTargetPageId="4" captionText="FIGURES 1–10. 1–4 Metasternal process; 1—Brachystethus geniculatus (Fabricius, 1787); 2—Peromatus bolivianus Fallou, 1887; 3—Pantochlora vivida Stål, 1870; 4—Doesburguedessa nigrolimbata Fernandes, 2010. 5–6 Dorsal view of head and pronotum; 5—Peromatus bolivianus; 6—Edessa (Pygoda) sp. 7–8 Dorsal view; 7—Olbia caprina (Stål, 1862); 8—Edessa (Hypoxys) boerneri Breddin, 1904. 9–10 Dorsal view of head and pronotum; 9—Grammedessa sp.; 10—Edessa cervus (Fabricius, 1787). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163939/files/figure.png" pageId="1" pageNumber="255">Fig. 8</figureCitation>
); humeral angles variable in shape......... 6
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</keyStep>
<keyStep id="9C6D2E6AFF98FFF95FFEF91CFA798920" pageId="1" pageNumber="255">
<paragraph id="272636CFFF98FFF95FFEF91CFA7988EA" blockId="1.[151,1436,1191,2034]" pageId="1" pageNumber="255">
<keyLead id="9C6895FAFF98FFF95FFEF91CFA7988EA" pageId="1" pageNumber="255">
6. Head with 6-8 solid black stripes, punctures of the head concentrated on these stripes (
<figureCitation id="BFA22A4AFF98FFF95B40F91CFB878888" box="[1065,1125,1689,1711]" captionStart="FIGURES 1–10. 1–4" captionStartId="4.[151,264,1881,1904]" captionTargetBox="[251,1335,195,1845]" captionTargetId="figure@4.[226,1365,195,1870]" captionTargetPageId="4" captionText="FIGURES 1–10. 1–4 Metasternal process; 1—Brachystethus geniculatus (Fabricius, 1787); 2—Peromatus bolivianus Fallou, 1887; 3—Pantochlora vivida Stål, 1870; 4—Doesburguedessa nigrolimbata Fernandes, 2010. 5–6 Dorsal view of head and pronotum; 5—Peromatus bolivianus; 6—Edessa (Pygoda) sp. 7–8 Dorsal view; 7—Olbia caprina (Stål, 1862); 8—Edessa (Hypoxys) boerneri Breddin, 1904. 9–10 Dorsal view of head and pronotum; 9—Grammedessa sp.; 10—Edessa cervus (Fabricius, 1787). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163939/files/figure.png" pageId="1" pageNumber="255">Fig. 9</figureCitation>
); proctiger with a medial carina on posterior face..........................................................
<emphasis id="15EDEADDFF98FFF95B2BF932FB2E88EA" box="[1090,1228,1719,1741]" italics="true" pageId="1" pageNumber="255">Grammedessa</emphasis>
Correia &amp; Fernandes
</keyLead>
</paragraph>
<paragraph id="272636CFFF98FFF95FFEF950FA798920" blockId="1.[151,1436,1191,2034]" pageId="1" pageNumber="255">
<keyLead id="9C6895FAFF98FFF95FFEF950FA798920" pageId="1" pageNumber="255">
- Head with punctures sparse or concentrated but never on black solid lines (
<figureCitation id="BFA22A4AFF98FFF95CD8F951FC1B88CD" box="[945,1017,1748,1770]" captionStart="FIGURES 1–10. 1–4" captionStartId="4.[151,264,1881,1904]" captionTargetBox="[251,1335,195,1845]" captionTargetId="figure@4.[226,1365,195,1870]" captionTargetPageId="4" captionText="FIGURES 1–10. 1–4 Metasternal process; 1—Brachystethus geniculatus (Fabricius, 1787); 2—Peromatus bolivianus Fallou, 1887; 3—Pantochlora vivida Stål, 1870; 4—Doesburguedessa nigrolimbata Fernandes, 2010. 5–6 Dorsal view of head and pronotum; 5—Peromatus bolivianus; 6—Edessa (Pygoda) sp. 7–8 Dorsal view; 7—Olbia caprina (Stål, 1862); 8—Edessa (Hypoxys) boerneri Breddin, 1904. 9–10 Dorsal view of head and pronotum; 9—Grammedessa sp.; 10—Edessa cervus (Fabricius, 1787). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163939/files/figure.png" pageId="1" pageNumber="255">Fig. 10</figureCitation>
); posterior face of proctiger without medial carina............................................................................................... 7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9C6D2E6AFF98FFF95FFEF88AFA798965" pageId="1" pageNumber="255">
<paragraph id="272636CFFF98FFF95FFEF88AFA798902" blockId="1.[151,1436,1191,2034]" box="[151,1435,1807,1829]" pageId="1" pageNumber="255">
<keyLead id="9C6895FAFF98FFF95FFEF88AFA798902" box="[151,1435,1807,1829]" pageId="1" pageNumber="255">
7. Humeral angles rounded or acute but shorter than head width (
<figureCitation id="BFA22A4AFF98FFF95C51F88AFC4E8902" box="[824,940,1807,1829]" captionStart="FIGURES 11–22. 11–14" captionStartId="5.[151,264,1836,1859]" captionTargetBox="[271,1344,187,1807]" captionTargetId="figure@5.[239,1361,183,1833]" captionTargetPageId="5" captionText="FIGURES 11–22. 11–14 Dorsal view; 11—Paraedessa verhoeffi (Breddin, 1904); 12—Ascra bifida Say, 1832; 13—Edessa (Dorypleura) bubalus LePeletier &amp; Serville, 1825; 14—Edessa (Edessa) elaphus Breddin, 1905. 15–17 Female, genital plates, ventral view; 15—Edessa (Pygoda) polita (LePeletier &amp; Serville, 1825); 16—Edessa (Hypoxys) boerneri; 17—Edessa (Aceratodes) brasiliensis Silva et al., 2006. 18–19 Ventral view of head and thorax; 18—Plagaedessa celsa; 19—Edessa (Hypoxys) boerneri. 20—Dorsal view of Edessa (Pygoda) polita. 21–22 Metasternal process; 21—Ascra bifida; 22—Edessa (Aceratodes) rufomarginata (De Geer, 1773). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163941/files/figure.png" pageId="1" pageNumber="255">Figs. 11, 12</figureCitation>
)........................................ 8
</keyLead>
</paragraph>
<paragraph id="272636CFFF98FFF95FFEF8A8FA798965" blockId="1.[151,1436,1191,2034]" box="[151,1435,1836,1858]" pageId="1" pageNumber="255">
<keyLead id="9C6895FAFF98FFF95FFEF8A8FA798965" box="[151,1435,1836,1858]" pageId="1" pageNumber="255">
- Humeral angles globose, depressed dorsoventrally or acute and normally longer than head width (
<figureCitation id="BFA22A4AFF98FFF95BE3F8A9FB1F8965" box="[1162,1277,1836,1858]" captionStart="FIGURES 11–22. 11–14" captionStartId="5.[151,264,1836,1859]" captionTargetBox="[271,1344,187,1807]" captionTargetId="figure@5.[239,1361,183,1833]" captionTargetPageId="5" captionText="FIGURES 11–22. 11–14 Dorsal view; 11—Paraedessa verhoeffi (Breddin, 1904); 12—Ascra bifida Say, 1832; 13—Edessa (Dorypleura) bubalus LePeletier &amp; Serville, 1825; 14—Edessa (Edessa) elaphus Breddin, 1905. 15–17 Female, genital plates, ventral view; 15—Edessa (Pygoda) polita (LePeletier &amp; Serville, 1825); 16—Edessa (Hypoxys) boerneri; 17—Edessa (Aceratodes) brasiliensis Silva et al., 2006. 18–19 Ventral view of head and thorax; 18—Plagaedessa celsa; 19—Edessa (Hypoxys) boerneri. 20—Dorsal view of Edessa (Pygoda) polita. 21–22 Metasternal process; 21—Ascra bifida; 22—Edessa (Aceratodes) rufomarginata (De Geer, 1773). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163941/files/figure.png" pageId="1" pageNumber="255">Figs. 13, 14</figureCitation>
).......... 12
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9C6D2E6AFF98FFF95FFEF8CCFA79899F" pageId="1" pageNumber="255">
<paragraph id="272636CFFF98FFF95FFEF8CCFA79895A" blockId="1.[151,1436,1191,2034]" pageId="1" pageNumber="255">
<keyLead id="9C6895FAFF98FFF95FFEF8CCFA79895A" pageId="1" pageNumber="255">
8. Species elongate; humeral angles acute (
<figureCitation id="BFA22A4AFF98FFF95D05F8CCFD518978" box="[620,691,1865,1887]" captionStart="FIGURES 11–22. 11–14" captionStartId="5.[151,264,1836,1859]" captionTargetBox="[271,1344,187,1807]" captionTargetId="figure@5.[239,1361,183,1833]" captionTargetPageId="5" captionText="FIGURES 11–22. 11–14 Dorsal view; 11—Paraedessa verhoeffi (Breddin, 1904); 12—Ascra bifida Say, 1832; 13—Edessa (Dorypleura) bubalus LePeletier &amp; Serville, 1825; 14—Edessa (Edessa) elaphus Breddin, 1905. 15–17 Female, genital plates, ventral view; 15—Edessa (Pygoda) polita (LePeletier &amp; Serville, 1825); 16—Edessa (Hypoxys) boerneri; 17—Edessa (Aceratodes) brasiliensis Silva et al., 2006. 18–19 Ventral view of head and thorax; 18—Plagaedessa celsa; 19—Edessa (Hypoxys) boerneri. 20—Dorsal view of Edessa (Pygoda) polita. 21–22 Metasternal process; 21—Ascra bifida; 22—Edessa (Aceratodes) rufomarginata (De Geer, 1773). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163941/files/figure.png" pageId="1" pageNumber="255">Fig. 11</figureCitation>
, red arrow); abdominal segment VII projected posteriorly (
<figureCitation id="BFA22A4AFF98FFF95B9CF8CCFA888978" box="[1269,1386,1865,1887]" captionStart="FIGURES 11–22. 11–14" captionStartId="5.[151,264,1836,1859]" captionTargetBox="[271,1344,187,1807]" captionTargetId="figure@5.[239,1361,183,1833]" captionTargetPageId="5" captionText="FIGURES 11–22. 11–14 Dorsal view; 11—Paraedessa verhoeffi (Breddin, 1904); 12—Ascra bifida Say, 1832; 13—Edessa (Dorypleura) bubalus LePeletier &amp; Serville, 1825; 14—Edessa (Edessa) elaphus Breddin, 1905. 15–17 Female, genital plates, ventral view; 15—Edessa (Pygoda) polita (LePeletier &amp; Serville, 1825); 16—Edessa (Hypoxys) boerneri; 17—Edessa (Aceratodes) brasiliensis Silva et al., 2006. 18–19 Ventral view of head and thorax; 18—Plagaedessa celsa; 19—Edessa (Hypoxys) boerneri. 20—Dorsal view of Edessa (Pygoda) polita. 21–22 Metasternal process; 21—Ascra bifida; 22—Edessa (Aceratodes) rufomarginata (De Geer, 1773). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163941/files/figure.png" pageId="1" pageNumber="255">Figs. 15, 16</figureCitation>
) and contrastingly dark at least in distal half (
<figureCitation id="BFA22A4AFF98FFF95D0AF8E2FD37895A" box="[611,725,1895,1917]" captionStart="FIGURES 11–22. 11–14" captionStartId="5.[151,264,1836,1859]" captionTargetBox="[271,1344,187,1807]" captionTargetId="figure@5.[239,1361,183,1833]" captionTargetPageId="5" captionText="FIGURES 11–22. 11–14 Dorsal view; 11—Paraedessa verhoeffi (Breddin, 1904); 12—Ascra bifida Say, 1832; 13—Edessa (Dorypleura) bubalus LePeletier &amp; Serville, 1825; 14—Edessa (Edessa) elaphus Breddin, 1905. 15–17 Female, genital plates, ventral view; 15—Edessa (Pygoda) polita (LePeletier &amp; Serville, 1825); 16—Edessa (Hypoxys) boerneri; 17—Edessa (Aceratodes) brasiliensis Silva et al., 2006. 18–19 Ventral view of head and thorax; 18—Plagaedessa celsa; 19—Edessa (Hypoxys) boerneri. 20—Dorsal view of Edessa (Pygoda) polita. 21–22 Metasternal process; 21—Ascra bifida; 22—Edessa (Aceratodes) rufomarginata (De Geer, 1773). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163941/files/figure.png" pageId="1" pageNumber="255">Figs. 11, 20</figureCitation>
)......................................................... 9
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</paragraph>
<paragraph id="272636CFFF98FFF95FFEF800FA79899F" blockId="1.[151,1436,1191,2034]" pageId="1" pageNumber="255">
<keyLead id="9C6895FAFF98FFF95FFEF800FA79899F" pageId="1" pageNumber="255">
- Species rounded; humeral angles rounded (
<figureCitation id="BFA22A4AFF98FFF95DE9F801FD2589BD" box="[640,711,1924,1946]" captionStart="FIGURES 11–22. 11–14" captionStartId="5.[151,264,1836,1859]" captionTargetBox="[271,1344,187,1807]" captionTargetId="figure@5.[239,1361,183,1833]" captionTargetPageId="5" captionText="FIGURES 11–22. 11–14 Dorsal view; 11—Paraedessa verhoeffi (Breddin, 1904); 12—Ascra bifida Say, 1832; 13—Edessa (Dorypleura) bubalus LePeletier &amp; Serville, 1825; 14—Edessa (Edessa) elaphus Breddin, 1905. 15–17 Female, genital plates, ventral view; 15—Edessa (Pygoda) polita (LePeletier &amp; Serville, 1825); 16—Edessa (Hypoxys) boerneri; 17—Edessa (Aceratodes) brasiliensis Silva et al., 2006. 18–19 Ventral view of head and thorax; 18—Plagaedessa celsa; 19—Edessa (Hypoxys) boerneri. 20—Dorsal view of Edessa (Pygoda) polita. 21–22 Metasternal process; 21—Ascra bifida; 22—Edessa (Aceratodes) rufomarginata (De Geer, 1773). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163941/files/figure.png" pageId="1" pageNumber="255">Fig. 12</figureCitation>
, red arrow); abdominal segment VII not posteriorly projected (
<figureCitation id="BFA22A4AFF98FFF95A4CF801FA8989BD" box="[1317,1387,1924,1946]" captionStart="FIGURES 11–22. 11–14" captionStartId="5.[151,264,1836,1859]" captionTargetBox="[271,1344,187,1807]" captionTargetId="figure@5.[239,1361,183,1833]" captionTargetPageId="5" captionText="FIGURES 11–22. 11–14 Dorsal view; 11—Paraedessa verhoeffi (Breddin, 1904); 12—Ascra bifida Say, 1832; 13—Edessa (Dorypleura) bubalus LePeletier &amp; Serville, 1825; 14—Edessa (Edessa) elaphus Breddin, 1905. 15–17 Female, genital plates, ventral view; 15—Edessa (Pygoda) polita (LePeletier &amp; Serville, 1825); 16—Edessa (Hypoxys) boerneri; 17—Edessa (Aceratodes) brasiliensis Silva et al., 2006. 18–19 Ventral view of head and thorax; 18—Plagaedessa celsa; 19—Edessa (Hypoxys) boerneri. 20—Dorsal view of Edessa (Pygoda) polita. 21–22 Metasternal process; 21—Ascra bifida; 22—Edessa (Aceratodes) rufomarginata (De Geer, 1773). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163941/files/figure.png" pageId="1" pageNumber="255">Fig. 17</figureCitation>
) and concolorous with connexivum (
<figureCitation id="BFA22A4AFF98FFF95D79F827FDB7899F" box="[528,597,1954,1976]" captionStart="FIGURES 11–22. 11–14" captionStartId="5.[151,264,1836,1859]" captionTargetBox="[271,1344,187,1807]" captionTargetId="figure@5.[239,1361,183,1833]" captionTargetPageId="5" captionText="FIGURES 11–22. 11–14 Dorsal view; 11—Paraedessa verhoeffi (Breddin, 1904); 12—Ascra bifida Say, 1832; 13—Edessa (Dorypleura) bubalus LePeletier &amp; Serville, 1825; 14—Edessa (Edessa) elaphus Breddin, 1905. 15–17 Female, genital plates, ventral view; 15—Edessa (Pygoda) polita (LePeletier &amp; Serville, 1825); 16—Edessa (Hypoxys) boerneri; 17—Edessa (Aceratodes) brasiliensis Silva et al., 2006. 18–19 Ventral view of head and thorax; 18—Plagaedessa celsa; 19—Edessa (Hypoxys) boerneri. 20—Dorsal view of Edessa (Pygoda) polita. 21–22 Metasternal process; 21—Ascra bifida; 22—Edessa (Aceratodes) rufomarginata (De Geer, 1773). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163941/files/figure.png" pageId="1" pageNumber="255">Fig. 12</figureCitation>
)................................................................... 13
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</paragraph>
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<keyStep id="9C6D2E6AFF98FFFA5FFEF83AFA798EC0" lastPageId="2" lastPageNumber="256" pageId="1" pageNumber="255">
<paragraph id="272636CFFF98FFF95FFEF83AFA7E89D5" blockId="1.[151,1436,1191,2034]" pageId="1" pageNumber="255">
<keyLead id="9C6895FAFF98FFF95FFEF83AFA7E89D5" pageId="1" pageNumber="255">
9. Legs with black stripes (
<figureCitation id="BFA22A4AFF98FFF95EBBF83AFDF889F2" box="[466,538,1983,2005]" captionStart="FIGURES 11–22. 11–14" captionStartId="5.[151,264,1836,1859]" captionTargetBox="[271,1344,187,1807]" captionTargetId="figure@5.[239,1361,183,1833]" captionTargetPageId="5" captionText="FIGURES 11–22. 11–14 Dorsal view; 11—Paraedessa verhoeffi (Breddin, 1904); 12—Ascra bifida Say, 1832; 13—Edessa (Dorypleura) bubalus LePeletier &amp; Serville, 1825; 14—Edessa (Edessa) elaphus Breddin, 1905. 15–17 Female, genital plates, ventral view; 15—Edessa (Pygoda) polita (LePeletier &amp; Serville, 1825); 16—Edessa (Hypoxys) boerneri; 17—Edessa (Aceratodes) brasiliensis Silva et al., 2006. 18–19 Ventral view of head and thorax; 18—Plagaedessa celsa; 19—Edessa (Hypoxys) boerneri. 20—Dorsal view of Edessa (Pygoda) polita. 21–22 Metasternal process; 21—Ascra bifida; 22—Edessa (Aceratodes) rufomarginata (De Geer, 1773). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163941/files/figure.png" pageId="1" pageNumber="255">Fig. 18</figureCitation>
); antennae black, except first segment with a ventral yellow spot (
<figureCitation id="BFA22A4AFF98FFF95BCFF83AFB0D89F2" box="[1190,1263,1983,2005]" captionStart="FIGURES 11–22. 11–14" captionStartId="5.[151,264,1836,1859]" captionTargetBox="[271,1344,187,1807]" captionTargetId="figure@5.[239,1361,183,1833]" captionTargetPageId="5" captionText="FIGURES 11–22. 11–14 Dorsal view; 11—Paraedessa verhoeffi (Breddin, 1904); 12—Ascra bifida Say, 1832; 13—Edessa (Dorypleura) bubalus LePeletier &amp; Serville, 1825; 14—Edessa (Edessa) elaphus Breddin, 1905. 15–17 Female, genital plates, ventral view; 15—Edessa (Pygoda) polita (LePeletier &amp; Serville, 1825); 16—Edessa (Hypoxys) boerneri; 17—Edessa (Aceratodes) brasiliensis Silva et al., 2006. 18–19 Ventral view of head and thorax; 18—Plagaedessa celsa; 19—Edessa (Hypoxys) boerneri. 20—Dorsal view of Edessa (Pygoda) polita. 21–22 Metasternal process; 21—Ascra bifida; 22—Edessa (Aceratodes) rufomarginata (De Geer, 1773). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163941/files/figure.png" pageId="1" pageNumber="255">Fig. 18</figureCitation>
); pygophore with posterolateral angles developed and posterolaterally directed (
<figureCitation id="BFA22A4AFF98FFF95C21F859FC2089D5" box="[840,962,2012,2034]" captionStart="FIGURES 23–27" captionStartId="7.[151,264,1906,1929]" captionTargetBox="[221,1359,197,1852]" captionTargetId="figure@7.[219,1368,195,1885]" captionTargetPageId="7" captionText="FIGURES 23–27. Plagaedessa celsa. 23–25 Male, pygophore; 23—dorsal view; 24—posterior view; 25—ventral view. 26— Female, genital plates. 27—Metasternal process, ventral view.(pyg–pygophore; dr–dorsal rim; vr–ventral rim; proc–proctiger; gp–genital superior process; pa–parameres; gc8–gonocoxite 8; gc9–gonocoxite 9; la8–laterotergite 8; la9–laterotergite 9; x– abdominal segment X. Scale = 1mm)." httpUri="https://zenodo.org/record/1163943/files/figure.png" pageId="1" pageNumber="255">Figs. 23–25</figureCitation>
); females with gonapophysis 9 bearing a small
</keyLead>
</paragraph>
<paragraph id="272636CFFF9BFFFA5F88FF12FA798EC0" blockId="2.[151,1436,151,935]" pageId="2" pageNumber="256">
tumescence at the base of gonocoxites 9 (
<figureCitation id="BFA22A4AFF9BFFFA5D07FF12FD038E8A" box="[622,737,151,173]" captionStart="FIGURES 40–43. 40–41" captionStartId="11.[151,264,1885,1908]" captionTargetBox="[283,1311,205,1858]" captionTargetId="figure@11.[262,1335,193,1868]" captionTargetPageId="11" captionText="FIGURES 40–43. 40–41 Laterotergites, gonocoxites and gonapophyses of ninth segment, and ectodermical genital ducts, ventral: 40—Plagaedessa celsa; 41—P. distanti. 42—Female, genital plates of P. celsa. 43—Phallus of P. celsa, lateral view. (aaf–anterior annular flange; ch–chitinellipsen; cs–capsula seminalis; dr–ductus receptaculi; paf–posterior annular flange; pc– pars comunis; ph–phallus; pi–pars intermedialis; tdr–thickening of ductus receptaculi; tg9–thickening of gonapophyses 9; tvi– thickening of vaginal intima; ves–vesical.Scale = 1mm)." httpUri="https://zenodo.org/record/1163951/files/figure.png" pageId="2" pageNumber="256">Figs. 40, 41</figureCitation>
)........................................
<taxonomicName id="E0994D4CFF9BFFFA5BBBFF12FAA98E8A" authority="Almeida &amp; Nunes &amp; Fernandes, 2018" authorityName="Almeida &amp; Nunes &amp; Fernandes" authorityYear="2018" box="[1234,1355,151,173]" class="Insecta" family="Pentatomidae" genus="Plagaedessa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="256" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="15EDEADDFF9BFFFA5BBBFF12FAA98E8A" box="[1234,1355,151,173]" italics="true" pageId="2" pageNumber="256">Plagaedessa</emphasis>
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<emphasis id="15EDEADDFF9BFFFA5A3BFF12FA748E8A" bold="true" box="[1362,1430,151,173]" pageId="2" pageNumber="256">gen. n.</emphasis>
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<keyLead id="9C6895FAFF9BFFFA5FFEFF30FA798EC0" pageId="2" pageNumber="256">
- Legs unicolorous (
<figureCitation id="BFA22A4AFF9BFFFA5EFFFF31FE3F8EED" box="[406,477,180,202]" captionStart="FIGURES 11–22. 11–14" captionStartId="5.[151,264,1836,1859]" captionTargetBox="[271,1344,187,1807]" captionTargetId="figure@5.[239,1361,183,1833]" captionTargetPageId="5" captionText="FIGURES 11–22. 11–14 Dorsal view; 11—Paraedessa verhoeffi (Breddin, 1904); 12—Ascra bifida Say, 1832; 13—Edessa (Dorypleura) bubalus LePeletier &amp; Serville, 1825; 14—Edessa (Edessa) elaphus Breddin, 1905. 15–17 Female, genital plates, ventral view; 15—Edessa (Pygoda) polita (LePeletier &amp; Serville, 1825); 16—Edessa (Hypoxys) boerneri; 17—Edessa (Aceratodes) brasiliensis Silva et al., 2006. 18–19 Ventral view of head and thorax; 18—Plagaedessa celsa; 19—Edessa (Hypoxys) boerneri. 20—Dorsal view of Edessa (Pygoda) polita. 21–22 Metasternal process; 21—Ascra bifida; 22—Edessa (Aceratodes) rufomarginata (De Geer, 1773). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163941/files/figure.png" pageId="2" pageNumber="256">Fig. 19</figureCitation>
); antennae yellow to light brown (
<figureCitation id="BFA22A4AFF9BFFFA5C47FF31FC948EED" box="[814,886,180,202]" captionStart="FIGURES 11–22. 11–14" captionStartId="5.[151,264,1836,1859]" captionTargetBox="[271,1344,187,1807]" captionTargetId="figure@5.[239,1361,183,1833]" captionTargetPageId="5" captionText="FIGURES 11–22. 11–14 Dorsal view; 11—Paraedessa verhoeffi (Breddin, 1904); 12—Ascra bifida Say, 1832; 13—Edessa (Dorypleura) bubalus LePeletier &amp; Serville, 1825; 14—Edessa (Edessa) elaphus Breddin, 1905. 15–17 Female, genital plates, ventral view; 15—Edessa (Pygoda) polita (LePeletier &amp; Serville, 1825); 16—Edessa (Hypoxys) boerneri; 17—Edessa (Aceratodes) brasiliensis Silva et al., 2006. 18–19 Ventral view of head and thorax; 18—Plagaedessa celsa; 19—Edessa (Hypoxys) boerneri. 20—Dorsal view of Edessa (Pygoda) polita. 21–22 Metasternal process; 21—Ascra bifida; 22—Edessa (Aceratodes) rufomarginata (De Geer, 1773). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163941/files/figure.png" pageId="2" pageNumber="256">Fig. 19</figureCitation>
); pygophore with posterolateral angles barely developed and posteriorly directed; females without tumescence on gonapophysis 9........................................ 10
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9C6D2E6AFF9BFFFA5FFEFF6AFA798FB0" pageId="2" pageNumber="256">
<paragraph id="272636CFFF9BFFFA5FFEFF6AFA748F18" blockId="2.[151,1436,151,935]" pageId="2" pageNumber="256">
<keyLead id="9C6895FAFF9BFFFA5FFEFF6AFA748F18" pageId="2" pageNumber="256">
10. Corium uniformly brown with contrastingly yellow veins sometimes strongly branched (
<figureCitation id="BFA22A4AFF9BFFFA5B28FF6AFB688F22" box="[1089,1162,239,261]" captionStart="FIGURES 11–22. 11–14" captionStartId="5.[151,264,1836,1859]" captionTargetBox="[271,1344,187,1807]" captionTargetId="figure@5.[239,1361,183,1833]" captionTargetPageId="5" captionText="FIGURES 11–22. 11–14 Dorsal view; 11—Paraedessa verhoeffi (Breddin, 1904); 12—Ascra bifida Say, 1832; 13—Edessa (Dorypleura) bubalus LePeletier &amp; Serville, 1825; 14—Edessa (Edessa) elaphus Breddin, 1905. 15–17 Female, genital plates, ventral view; 15—Edessa (Pygoda) polita (LePeletier &amp; Serville, 1825); 16—Edessa (Hypoxys) boerneri; 17—Edessa (Aceratodes) brasiliensis Silva et al., 2006. 18–19 Ventral view of head and thorax; 18—Plagaedessa celsa; 19—Edessa (Hypoxys) boerneri. 20—Dorsal view of Edessa (Pygoda) polita. 21–22 Metasternal process; 21—Ascra bifida; 22—Edessa (Aceratodes) rufomarginata (De Geer, 1773). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163941/files/figure.png" pageId="2" pageNumber="256">Fig. 20</figureCitation>
); abdominal segment VII at least 1.5 times longer than wide at base (
<figureCitation id="BFA22A4AFF9BFFFA5D0CFE89FD498F05" box="[613,683,268,290]" captionStart="FIGURES 11–22. 11–14" captionStartId="5.[151,264,1836,1859]" captionTargetBox="[271,1344,187,1807]" captionTargetId="figure@5.[239,1361,183,1833]" captionTargetPageId="5" captionText="FIGURES 11–22. 11–14 Dorsal view; 11—Paraedessa verhoeffi (Breddin, 1904); 12—Ascra bifida Say, 1832; 13—Edessa (Dorypleura) bubalus LePeletier &amp; Serville, 1825; 14—Edessa (Edessa) elaphus Breddin, 1905. 15–17 Female, genital plates, ventral view; 15—Edessa (Pygoda) polita (LePeletier &amp; Serville, 1825); 16—Edessa (Hypoxys) boerneri; 17—Edessa (Aceratodes) brasiliensis Silva et al., 2006. 18–19 Ventral view of head and thorax; 18—Plagaedessa celsa; 19—Edessa (Hypoxys) boerneri. 20—Dorsal view of Edessa (Pygoda) polita. 21–22 Metasternal process; 21—Ascra bifida; 22—Edessa (Aceratodes) rufomarginata (De Geer, 1773). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163941/files/figure.png" pageId="2" pageNumber="256">Fig. 15</figureCitation>
); proctiger with posterior face rounded; phallus with a distal constriction;
<emphasis id="15EDEADDFF9BFFFA5A18FE88FE888F18" italics="true" pageId="2" pageNumber="256">capsula seminalis</emphasis>
as big as or bigger than laterotergites 8 and 9...........................subg.
<emphasis id="15EDEADDFF9BFFFA5BF2FEACFB078F18" box="[1179,1253,297,319]" italics="true" pageId="2" pageNumber="256">Pygoda</emphasis>
Amyot &amp; Serville
</keyLead>
</paragraph>
<paragraph id="272636CFFF9BFFFA5FFEFEC2FA798FB0" blockId="2.[151,1436,151,935]" pageId="2" pageNumber="256">
<keyLead id="9C6895FAFF9BFFFA5FFEFEC2FA798FB0" pageId="2" pageNumber="256">
- Corium brown variegate with whitish and blackish spots (
<figureCitation id="BFA22A4AFF9BFFFA5C7CFEC2FCB88F7A" box="[789,858,327,349]" captionStart="FIGURES 1–10. 1–4" captionStartId="4.[151,264,1881,1904]" captionTargetBox="[251,1335,195,1845]" captionTargetId="figure@4.[226,1365,195,1870]" captionTargetPageId="4" captionText="FIGURES 1–10. 1–4 Metasternal process; 1—Brachystethus geniculatus (Fabricius, 1787); 2—Peromatus bolivianus Fallou, 1887; 3—Pantochlora vivida Stål, 1870; 4—Doesburguedessa nigrolimbata Fernandes, 2010. 5–6 Dorsal view of head and pronotum; 5—Peromatus bolivianus; 6—Edessa (Pygoda) sp. 7–8 Dorsal view; 7—Olbia caprina (Stål, 1862); 8—Edessa (Hypoxys) boerneri Breddin, 1904. 9–10 Dorsal view of head and pronotum; 9—Grammedessa sp.; 10—Edessa cervus (Fabricius, 1787). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163939/files/figure.png" pageId="2" pageNumber="256">Figs. 8</figureCitation>
,
<figureCitation id="BFA22A4AFF9BFFFA5C0FFEC2FC9D8F7A" box="[870,895,327,349]" captionStart="FIGURES 11–22. 11–14" captionStartId="5.[151,264,1836,1859]" captionTargetBox="[271,1344,187,1807]" captionTargetId="figure@5.[239,1361,183,1833]" captionTargetPageId="5" captionText="FIGURES 11–22. 11–14 Dorsal view; 11—Paraedessa verhoeffi (Breddin, 1904); 12—Ascra bifida Say, 1832; 13—Edessa (Dorypleura) bubalus LePeletier &amp; Serville, 1825; 14—Edessa (Edessa) elaphus Breddin, 1905. 15–17 Female, genital plates, ventral view; 15—Edessa (Pygoda) polita (LePeletier &amp; Serville, 1825); 16—Edessa (Hypoxys) boerneri; 17—Edessa (Aceratodes) brasiliensis Silva et al., 2006. 18–19 Ventral view of head and thorax; 18—Plagaedessa celsa; 19—Edessa (Hypoxys) boerneri. 20—Dorsal view of Edessa (Pygoda) polita. 21–22 Metasternal process; 21—Ascra bifida; 22—Edessa (Aceratodes) rufomarginata (De Geer, 1773). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163941/files/figure.png" pageId="2" pageNumber="256">11</figureCitation>
); abdominal segment VII at most 1.2 times longer than wide at base (
<figureCitation id="BFA22A4AFF9BFFFA5E00FEE1FE508F5D" box="[361,434,356,378]" captionStart="FIGURES 11–22. 11–14" captionStartId="5.[151,264,1836,1859]" captionTargetBox="[271,1344,187,1807]" captionTargetId="figure@5.[239,1361,183,1833]" captionTargetPageId="5" captionText="FIGURES 11–22. 11–14 Dorsal view; 11—Paraedessa verhoeffi (Breddin, 1904); 12—Ascra bifida Say, 1832; 13—Edessa (Dorypleura) bubalus LePeletier &amp; Serville, 1825; 14—Edessa (Edessa) elaphus Breddin, 1905. 15–17 Female, genital plates, ventral view; 15—Edessa (Pygoda) polita (LePeletier &amp; Serville, 1825); 16—Edessa (Hypoxys) boerneri; 17—Edessa (Aceratodes) brasiliensis Silva et al., 2006. 18–19 Ventral view of head and thorax; 18—Plagaedessa celsa; 19—Edessa (Hypoxys) boerneri. 20—Dorsal view of Edessa (Pygoda) polita. 21–22 Metasternal process; 21—Ascra bifida; 22—Edessa (Aceratodes) rufomarginata (De Geer, 1773). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163941/files/figure.png" pageId="2" pageNumber="256">Fig. 16</figureCitation>
); proctiger with posterior face flat; phallus not constricted distally;
<emphasis id="15EDEADDFF9BFFFA5B20FEE1FB148F5D" box="[1097,1270,356,378]" italics="true" pageId="2" pageNumber="256">capsula seminalis</emphasis>
smaller than laterotergitos 8 and 9.................................................................................... 11
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9C6D2E6AFF9BFFFA5FFEFE1AFA7E8C60" pageId="2" pageNumber="256">
<paragraph id="272636CFFF9BFFFA5FFEFE1AFA798FC8" blockId="2.[151,1436,151,935]" pageId="2" pageNumber="256">
<keyLead id="9C6895FAFF9BFFFA5FFEFE1AFA798FC8" pageId="2" pageNumber="256">
11. Pronotum unicolorous (
<figureCitation id="BFA22A4AFF9BFFFA5EAAFE1AFDEA8F92" box="[451,520,415,437]" captionStart="FIGURES 11–22. 11–14" captionStartId="5.[151,264,1836,1859]" captionTargetBox="[271,1344,187,1807]" captionTargetId="figure@5.[239,1361,183,1833]" captionTargetPageId="5" captionText="FIGURES 11–22. 11–14 Dorsal view; 11—Paraedessa verhoeffi (Breddin, 1904); 12—Ascra bifida Say, 1832; 13—Edessa (Dorypleura) bubalus LePeletier &amp; Serville, 1825; 14—Edessa (Edessa) elaphus Breddin, 1905. 15–17 Female, genital plates, ventral view; 15—Edessa (Pygoda) polita (LePeletier &amp; Serville, 1825); 16—Edessa (Hypoxys) boerneri; 17—Edessa (Aceratodes) brasiliensis Silva et al., 2006. 18–19 Ventral view of head and thorax; 18—Plagaedessa celsa; 19—Edessa (Hypoxys) boerneri. 20—Dorsal view of Edessa (Pygoda) polita. 21–22 Metasternal process; 21—Ascra bifida; 22—Edessa (Aceratodes) rufomarginata (De Geer, 1773). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163941/files/figure.png" pageId="2" pageNumber="256">Fig. 11</figureCitation>
); dorsal surface of abdomen mostly metallic blue (
<figureCitation id="BFA22A4AFF9BFFFA5C85FE1AFBD08F92" box="[1004,1074,415,437]" captionStart="FIGURES 11–22. 11–14" captionStartId="5.[151,264,1836,1859]" captionTargetBox="[271,1344,187,1807]" captionTargetId="figure@5.[239,1361,183,1833]" captionTargetPageId="5" captionText="FIGURES 11–22. 11–14 Dorsal view; 11—Paraedessa verhoeffi (Breddin, 1904); 12—Ascra bifida Say, 1832; 13—Edessa (Dorypleura) bubalus LePeletier &amp; Serville, 1825; 14—Edessa (Edessa) elaphus Breddin, 1905. 15–17 Female, genital plates, ventral view; 15—Edessa (Pygoda) polita (LePeletier &amp; Serville, 1825); 16—Edessa (Hypoxys) boerneri; 17—Edessa (Aceratodes) brasiliensis Silva et al., 2006. 18–19 Ventral view of head and thorax; 18—Plagaedessa celsa; 19—Edessa (Hypoxys) boerneri. 20—Dorsal view of Edessa (Pygoda) polita. 21–22 Metasternal process; 21—Ascra bifida; 22—Edessa (Aceratodes) rufomarginata (De Geer, 1773). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163941/files/figure.png" pageId="2" pageNumber="256">Fig. 11</figureCitation>
); posterior part of proctiger expanded laterally in a flat plate (see
<bibRefCitation id="43084B3EFF9BFFFA5E86FE39FD718FF5" author="Silva" box="[495,659,444,466]" pageId="2" pageNumber="256" refString="Silva, V. J., Nunes, B. M. &amp; Fernandes, J. A. M. (2013) Paraedessa, a new genus of Edessinae (Hemiptera: Heteroptera: Pentatomidae). Zootaxa, 3716 (3), 395 - 416. https: // doi. org / 10.11646 / zootaxa. 3716.3.4" type="journal article" year="2013">
Silva
<emphasis id="15EDEADDFF9BFFFA5D41FE38FDB68FF5" box="[552,596,444,466]" italics="true" pageId="2" pageNumber="256">et al</emphasis>
. 2013
</bibRefCitation>
—
<figureCitation id="BFA22A4AFF9BFFFA5DC5FE39FD0C8FF5" box="[684,750,444,466]" captionStart="FIGURES 1–10. 1–4" captionStartId="4.[151,264,1881,1904]" captionTargetBox="[251,1335,195,1845]" captionTargetId="figure@4.[226,1365,195,1870]" captionTargetPageId="4" captionText="FIGURES 1–10. 1–4 Metasternal process; 1—Brachystethus geniculatus (Fabricius, 1787); 2—Peromatus bolivianus Fallou, 1887; 3—Pantochlora vivida Stål, 1870; 4—Doesburguedessa nigrolimbata Fernandes, 2010. 5–6 Dorsal view of head and pronotum; 5—Peromatus bolivianus; 6—Edessa (Pygoda) sp. 7–8 Dorsal view; 7—Olbia caprina (Stål, 1862); 8—Edessa (Hypoxys) boerneri Breddin, 1904. 9–10 Dorsal view of head and pronotum; 9—Grammedessa sp.; 10—Edessa cervus (Fabricius, 1787). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163939/files/figure.png" pageId="2" pageNumber="256">Figs. 2</figureCitation>
,
<figureCitation id="BFA22A4AFF9BFFFA5D93FE39FCF78FF5" box="[762,789,444,466]" captionStart="FIGURES 23–27" captionStartId="7.[151,264,1906,1929]" captionTargetBox="[221,1359,197,1852]" captionTargetId="figure@7.[219,1368,195,1885]" captionTargetPageId="7" captionText="FIGURES 23–27. Plagaedessa celsa. 23–25 Male, pygophore; 23—dorsal view; 24—posterior view; 25—ventral view. 26— Female, genital plates. 27—Metasternal process, ventral view.(pyg–pygophore; dr–dorsal rim; vr–ventral rim; proc–proctiger; gp–genital superior process; pa–parameres; gc8–gonocoxite 8; gc9–gonocoxite 9; la8–laterotergite 8; la9–laterotergite 9; x– abdominal segment X. Scale = 1mm)." httpUri="https://zenodo.org/record/1163943/files/figure.png" pageId="2" pageNumber="256">27</figureCitation>
); gonocoxites 8 somewhat reduced, leaving visible the sclerotized gonapophyses 8...............................................................
<taxonomicName id="E0994D4CFF9BFFFA5B1BFE5CFA798FC8" authority="Silva &amp; Fernandes" authorityName="Silva &amp; Fernandes" authorityYear="2013" box="[1138,1435,473,495]" class="Insecta" family="Pentatomidae" genus="Paraedessa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="256" phylum="Arthropoda" rank="genus">
<emphasis id="15EDEADDFF9BFFFA5B1BFE5CFB018FC8" box="[1138,1251,473,495]" italics="true" pageId="2" pageNumber="256">Paraedessa</emphasis>
Silva &amp; Fernandes
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="272636CFFF9BFFFA5FFEFE72FA7E8C60" blockId="2.[151,1436,151,935]" pageId="2" pageNumber="256">
<keyLead id="9C6895FAFF9BFFFA5FFEFE72FA7E8C60" pageId="2" pageNumber="256">
- Pronotum with dark transversal stripe (
<figureCitation id="BFA22A4AFF9BFFFA5D35FE72FD758C2A" box="[604,663,503,525]" captionStart="FIGURES 1–10. 1–4" captionStartId="4.[151,264,1881,1904]" captionTargetBox="[251,1335,195,1845]" captionTargetId="figure@4.[226,1365,195,1870]" captionTargetPageId="4" captionText="FIGURES 1–10. 1–4 Metasternal process; 1—Brachystethus geniculatus (Fabricius, 1787); 2—Peromatus bolivianus Fallou, 1887; 3—Pantochlora vivida Stål, 1870; 4—Doesburguedessa nigrolimbata Fernandes, 2010. 5–6 Dorsal view of head and pronotum; 5—Peromatus bolivianus; 6—Edessa (Pygoda) sp. 7–8 Dorsal view; 7—Olbia caprina (Stål, 1862); 8—Edessa (Hypoxys) boerneri Breddin, 1904. 9–10 Dorsal view of head and pronotum; 9—Grammedessa sp.; 10—Edessa cervus (Fabricius, 1787). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163939/files/figure.png" pageId="2" pageNumber="256">Fig. 8</figureCitation>
); dorsal surface of abdomen never metallic blue; posterior part of proctiger not expanded laterally, posterior face thick; gonocoxites 8 never reduced, leaving visible at most a tiny membranous part of gonapophyses 8............................................................ subg.
<emphasis id="15EDEADDFF9BFFFA5BFEFDB7FB088C60" box="[1175,1258,562,583]" italics="true" pageId="2" pageNumber="256">Hypoxys</emphasis>
Amyot &amp; Serville
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9C6D2E6AFF9BFFFA5FFEFDCAFA778CD0" pageId="2" pageNumber="256">
<paragraph id="272636CFFF9BFFFA5FFEFDCAFA7E8CB8" blockId="2.[151,1436,151,935]" pageId="2" pageNumber="256">
<keyLead id="9C6895FAFF9BFFFA5FFEFDCAFA7E8CB8" pageId="2" pageNumber="256">
12. Humeral angles long, apex acute (
<figureCitation id="BFA22A4AFF9BFFFA5D47FDCAFD948C42" box="[558,630,591,613]" captionStart="FIGURES 11–22. 11–14" captionStartId="5.[151,264,1836,1859]" captionTargetBox="[271,1344,187,1807]" captionTargetId="figure@5.[239,1361,183,1833]" captionTargetPageId="5" captionText="FIGURES 11–22. 11–14 Dorsal view; 11—Paraedessa verhoeffi (Breddin, 1904); 12—Ascra bifida Say, 1832; 13—Edessa (Dorypleura) bubalus LePeletier &amp; Serville, 1825; 14—Edessa (Edessa) elaphus Breddin, 1905. 15–17 Female, genital plates, ventral view; 15—Edessa (Pygoda) polita (LePeletier &amp; Serville, 1825); 16—Edessa (Hypoxys) boerneri; 17—Edessa (Aceratodes) brasiliensis Silva et al., 2006. 18–19 Ventral view of head and thorax; 18—Plagaedessa celsa; 19—Edessa (Hypoxys) boerneri. 20—Dorsal view of Edessa (Pygoda) polita. 21–22 Metasternal process; 21—Ascra bifida; 22—Edessa (Aceratodes) rufomarginata (De Geer, 1773). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163941/files/figure.png" pageId="2" pageNumber="256">Fig. 13</figureCitation>
, red arrow); abdominal segment VII twisted and strongly projected posteriorly, far surpassing genital plates or pygophore; pronotum with an anterior yellow stripe (
<figureCitation id="BFA22A4AFF9BFFFA5C81FDE9FBD28CA5" box="[1000,1072,620,642]" captionStart="FIGURES 11–22. 11–14" captionStartId="5.[151,264,1836,1859]" captionTargetBox="[271,1344,187,1807]" captionTargetId="figure@5.[239,1361,183,1833]" captionTargetPageId="5" captionText="FIGURES 11–22. 11–14 Dorsal view; 11—Paraedessa verhoeffi (Breddin, 1904); 12—Ascra bifida Say, 1832; 13—Edessa (Dorypleura) bubalus LePeletier &amp; Serville, 1825; 14—Edessa (Edessa) elaphus Breddin, 1905. 15–17 Female, genital plates, ventral view; 15—Edessa (Pygoda) polita (LePeletier &amp; Serville, 1825); 16—Edessa (Hypoxys) boerneri; 17—Edessa (Aceratodes) brasiliensis Silva et al., 2006. 18–19 Ventral view of head and thorax; 18—Plagaedessa celsa; 19—Edessa (Hypoxys) boerneri. 20—Dorsal view of Edessa (Pygoda) polita. 21–22 Metasternal process; 21—Ascra bifida; 22—Edessa (Aceratodes) rufomarginata (De Geer, 1773). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163941/files/figure.png" pageId="2" pageNumber="256">Fig. 13</figureCitation>
)...................................................................................................... subg.
<emphasis id="15EDEADDFF9BFFFA5B13FD0CFB088CB8" box="[1146,1258,649,671]" italics="true" pageId="2" pageNumber="256">Dorypleura</emphasis>
Amyot &amp; Serville
</keyLead>
</paragraph>
<paragraph id="272636CFFF9BFFFA5FFEFD22FA778CD0" blockId="2.[151,1436,151,935]" pageId="2" pageNumber="256">
<keyLead id="9C6895FAFF9BFFFA5FFEFD22FA778CD0" pageId="2" pageNumber="256">
- Humeral angles variable in length, apex globose (
<figureCitation id="BFA22A4AFF9BFFFA5DD7FD22FCE78C9A" box="[702,773,679,701]" captionStart="FIGURES 11–22. 11–14" captionStartId="5.[151,264,1836,1859]" captionTargetBox="[271,1344,187,1807]" captionTargetId="figure@5.[239,1361,183,1833]" captionTargetPageId="5" captionText="FIGURES 11–22. 11–14 Dorsal view; 11—Paraedessa verhoeffi (Breddin, 1904); 12—Ascra bifida Say, 1832; 13—Edessa (Dorypleura) bubalus LePeletier &amp; Serville, 1825; 14—Edessa (Edessa) elaphus Breddin, 1905. 15–17 Female, genital plates, ventral view; 15—Edessa (Pygoda) polita (LePeletier &amp; Serville, 1825); 16—Edessa (Hypoxys) boerneri; 17—Edessa (Aceratodes) brasiliensis Silva et al., 2006. 18–19 Ventral view of head and thorax; 18—Plagaedessa celsa; 19—Edessa (Hypoxys) boerneri. 20—Dorsal view of Edessa (Pygoda) polita. 21–22 Metasternal process; 21—Ascra bifida; 22—Edessa (Aceratodes) rufomarginata (De Geer, 1773). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163941/files/figure.png" pageId="2" pageNumber="256">Fig. 14</figureCitation>
, red arrow); abdominal segment VII not twisted and not projected as far posteriorly as above, not reaching to slightly surpassing genital plates or pygophore; pronotum without anterior yellow stripe (
<figureCitation id="BFA22A4AFF9BFFFA5E4AFD64FE8B8CD0" box="[291,361,737,759]" captionStart="FIGURES 11–22. 11–14" captionStartId="5.[151,264,1836,1859]" captionTargetBox="[271,1344,187,1807]" captionTargetId="figure@5.[239,1361,183,1833]" captionTargetPageId="5" captionText="FIGURES 11–22. 11–14 Dorsal view; 11—Paraedessa verhoeffi (Breddin, 1904); 12—Ascra bifida Say, 1832; 13—Edessa (Dorypleura) bubalus LePeletier &amp; Serville, 1825; 14—Edessa (Edessa) elaphus Breddin, 1905. 15–17 Female, genital plates, ventral view; 15—Edessa (Pygoda) polita (LePeletier &amp; Serville, 1825); 16—Edessa (Hypoxys) boerneri; 17—Edessa (Aceratodes) brasiliensis Silva et al., 2006. 18–19 Ventral view of head and thorax; 18—Plagaedessa celsa; 19—Edessa (Hypoxys) boerneri. 20—Dorsal view of Edessa (Pygoda) polita. 21–22 Metasternal process; 21—Ascra bifida; 22—Edessa (Aceratodes) rufomarginata (De Geer, 1773). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163941/files/figure.png" pageId="2" pageNumber="256">Fig. 14</figureCitation>
) (except
<taxonomicName id="E0994D4CFF9BFFFA5EABFD64FD028CD0" authority="Stal, 1872" authorityName="Stal" authorityYear="1872" box="[450,736,737,759]" class="Insecta" family="Pentatomidae" genus="Edessa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="256" phylum="Arthropoda" rank="species" species="pictiventris">
<emphasis id="15EDEADDFF9BFFFA5EABFD64FD958CD0" box="[450,631,737,759]" italics="true" pageId="2" pageNumber="256">Edessa pictiventris</emphasis>
<bibRefCitation id="43084B3EFF9BFFFA5D17FD64FD028CD0" author="Stal" box="[638,736,737,759]" pageId="2" pageNumber="256" refString="Stal, C. (1872) Enumeratio Hemipterorum. Bidrag till em forteckning ofver alla hittils kanda Hemiptera, Jemte systematiska meddelanden. 2. Kongliga Svenska Vetenskaps-Akademien Handlingar, 10 (4), 1 - 159." type="journal article" year="1872">Stål, 1872</bibRefCitation>
</taxonomicName>
)...................................... subg.
<taxonomicName id="E0994D4CFF9BFFFA5B9AFD64FA778CD0" ID-CoL="632LJ" authority="Fabricius" authorityName="Fabricius" authorityYear="1803" box="[1267,1429,737,759]" class="Insecta" family="Pentatomidae" genus="Edessa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="256" phylum="Arthropoda" rank="genus">
<emphasis id="15EDEADDFF9BFFFA5B9AFD64FAD58CD0" box="[1267,1335,737,759]" italics="true" pageId="2" pageNumber="256">Edessa</emphasis>
Fabricius
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="9C6D2E6AFF9BFFFA5FFEFD7AFA7E8D80" pageId="2" pageNumber="256">
<paragraph id="272636CFFF9BFFFA5FFEFD7AFA7E8D68" blockId="2.[151,1436,151,935]" pageId="2" pageNumber="256">
<keyLead id="9C6895FAFF9BFFFA5FFEFD7AFA7E8D68" pageId="2" pageNumber="256">
13. Pronotal disk tumid; body covered with dark punctures; arms of the metasternal process developed and sometimes slightly laterally compressed (
<figureCitation id="BFA22A4AFF9BFFFA5EF5FC99FE018D15" box="[412,483,796,818]" captionStart="FIGURES 11–22. 11–14" captionStartId="5.[151,264,1836,1859]" captionTargetBox="[271,1344,187,1807]" captionTargetId="figure@5.[239,1361,183,1833]" captionTargetPageId="5" captionText="FIGURES 11–22. 11–14 Dorsal view; 11—Paraedessa verhoeffi (Breddin, 1904); 12—Ascra bifida Say, 1832; 13—Edessa (Dorypleura) bubalus LePeletier &amp; Serville, 1825; 14—Edessa (Edessa) elaphus Breddin, 1905. 15–17 Female, genital plates, ventral view; 15—Edessa (Pygoda) polita (LePeletier &amp; Serville, 1825); 16—Edessa (Hypoxys) boerneri; 17—Edessa (Aceratodes) brasiliensis Silva et al., 2006. 18–19 Ventral view of head and thorax; 18—Plagaedessa celsa; 19—Edessa (Hypoxys) boerneri. 20—Dorsal view of Edessa (Pygoda) polita. 21–22 Metasternal process; 21—Ascra bifida; 22—Edessa (Aceratodes) rufomarginata (De Geer, 1773). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163941/files/figure.png" pageId="2" pageNumber="256">Fig. 21</figureCitation>
); Central America (except
<emphasis id="15EDEADDFF9BFFFA5D8EFC99FCBE8D15" box="[743,860,796,818]" italics="true" pageId="2" pageNumber="256">Ascra bifida</emphasis>
Say, 1832 also found in north part of South America)..................................................................................................
<emphasis id="15EDEADDFF9BFFFA5A5DFCBCFA938D68" box="[1332,1393,825,847]" italics="true" pageId="2" pageNumber="256">Ascra</emphasis>
Say
</keyLead>
</paragraph>
<paragraph id="272636CFFF9BFFFA5FFEFCD2FA7E8D80" blockId="2.[151,1436,151,935]" pageId="2" pageNumber="256">
<keyLead id="9C6895FAFF9BFFFA5FFEFCD2FA7E8D80" pageId="2" pageNumber="256">
- Pronotal disk not tumid; body covered with concolorous or light brown punctures; arms of the metasternal process strongly laterally compressed (
<figureCitation id="BFA22A4AFF9BFFFA5EF6FCF1FE0B8DAD" box="[415,489,884,906]" captionStart="FIGURES 11–22. 11–14" captionStartId="5.[151,264,1836,1859]" captionTargetBox="[271,1344,187,1807]" captionTargetId="figure@5.[239,1361,183,1833]" captionTargetPageId="5" captionText="FIGURES 11–22. 11–14 Dorsal view; 11—Paraedessa verhoeffi (Breddin, 1904); 12—Ascra bifida Say, 1832; 13—Edessa (Dorypleura) bubalus LePeletier &amp; Serville, 1825; 14—Edessa (Edessa) elaphus Breddin, 1905. 15–17 Female, genital plates, ventral view; 15—Edessa (Pygoda) polita (LePeletier &amp; Serville, 1825); 16—Edessa (Hypoxys) boerneri; 17—Edessa (Aceratodes) brasiliensis Silva et al., 2006. 18–19 Ventral view of head and thorax; 18—Plagaedessa celsa; 19—Edessa (Hypoxys) boerneri. 20—Dorsal view of Edessa (Pygoda) polita. 21–22 Metasternal process; 21—Ascra bifida; 22—Edessa (Aceratodes) rufomarginata (De Geer, 1773). (Scale = 2mm)." httpUri="https://zenodo.org/record/1163941/files/figure.png" pageId="2" pageNumber="256">Fig. 22</figureCitation>
), barely developed in some species; South America (except the widespread species
<taxonomicName id="E0994D4CFF9BFFFA5A49FCF1FE0F8D80" authority="(De Geer, 1773))" authorityName="" authorityYear="1773" baseAuthorityName="De Geer" baseAuthorityYear="1773" class="Insecta" family="Pentatomidae" genus="Edessa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="256" phylum="Arthropoda" rank="species" species="rufomarginata">
<emphasis id="15EDEADDFF9BFFFA5A49FCF1FEA78D80" italics="true" pageId="2" pageNumber="256">Edessa rufomarginata</emphasis>
(De Geer, 1773))
</taxonomicName>
................................................. subg.
<taxonomicName id="E0994D4CFF9BFFFA5B17FC14FA7E8D80" authority="Amyot &amp; Serville" authorityName="Amyot &amp; Serville" authorityYear="1843" box="[1150,1436,913,935]" class="Insecta" family="Pentatomidae" genus="Aceratodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="256" phylum="Arthropoda" rank="genus">
<emphasis id="15EDEADDFF9BFFFA5B17FC14FB088D80" box="[1150,1258,913,935]" italics="true" pageId="2" pageNumber="256">Aceratodes</emphasis>
Amyot &amp; Serville
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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