treatments-xml/data/7E/5B/6A/7E5B6A0BC24054D1860BEFB396008C61.xml

694 lines
74 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document ID-CLB-Dataset="121969" ID-DOI="http://dx.doi.org/10.3897/zookeys.1117.83539" ID-GBIF-Dataset="ed66c960-818b-48cb-a3e8-9c7c033800c2" ID-PMC="PMC9848781" ID-Pensoft-Pub="1313-2970-1117-203" ID-Pensoft-UUID="76BC5EBEBEA65F82903D460EBF22B366" ID-PubMed="36761382" ID-ZooBank="B2596F5CFE214B3EAAECDE33FF394CEC" ModsDocID="1313-2970-1117-203" checkinTime="1660625102094" checkinUser="pensoft" docAuthor="El Alami, Majida, Benlasri, Mokhtar, Sartori, Michel, Vuataz, Laurent &amp; Ghamizi, Mohamed" docDate="2022" docId="7E5B6A0BC24054D1860BEFB396008C61" docLanguage="en" docName="ZooKeys 1117: 203-218" docOrigin="ZooKeys 1117" docPubDate="2022-08-15" docSource="http://dx.doi.org/10.3897/zookeys.1117.83539" docTitle="Prosopistoma maroccanum El Alami, Benlasri &amp; Sartori 2022, sp. nov." docType="treatment" docUuid="9C03C91D-8E32-47C7-9FD1-DE01A7093072" docUuidSource="ZooBank" docVersion="6" id="76BC5EBEBEA65F82903D460EBF22B366" lastPageNumber="203" masterDocId="76BC5EBEBEA65F82903D460EBF22B366" masterDocTitle="A new species of the genus Prosopistoma Latreille, 1833 (Ephemeroptera, Prosopistomatidae) from Morocco" masterLastPageNumber="218" masterPageNumber="203" pageNumber="203" updateTime="1732940517473" updateUser="ExternalLinkService">
<mods:mods id="1C320C91B5131B2A67F1D3915558CDC0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="842AE0F7B1906308D8F04D55EAF98A5E">
<mods:title id="A5B23A189EF87769508EF2E79AB258B6">A new species of the genus Prosopistoma Latreille, 1833 (Ephemeroptera, Prosopistomatidae) from Morocco</mods:title>
</mods:titleInfo>
<mods:name id="07CBEA9743C50F28F5E428C5449341B1" type="personal">
<mods:role id="1D56BF351F0CF59419AF3BEFBA10DEB2">
<mods:roleTerm id="86C65CA944C7C4FDE88343D4B5FE88B5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4C9D35E11C9E6684E77F381B1F9A4B41">El Alami, Majida</mods:namePart>
<mods:nameIdentifier id="F4F9701CB782FD74B87BA5E6F88914AA" type="ORCID">https://orcid.org/0000-0003-2664-646X</mods:nameIdentifier>
<mods:affiliation id="95BDF8ABE1C19CB795C182749B0E9953">Laboratoire Ecologie, Systematique, Conservation de la Biodiversite (LESCB), Unite de Recherche Labellisee CNRST N ° 18, Universite Abdelmalek Essaadi, Faculte des Sciences, Departement de Biologie, B. P. 2121 93002, Tetouan, Morocco</mods:affiliation>
<mods:nameIdentifier id="EC26065F7063A54410D8B3469AF1A3E8" type="email">elalami.majida@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="11CFBED689E5F487E2F40618B0D018AD" type="personal">
<mods:role id="9E69BE39D3BF1A2809047D7294D8C679">
<mods:roleTerm id="E4C1CE277F3BA4ADF878088C93123BE3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="80D22EC6298234E47541F7C6F0D83BB8">Benlasri, Mokhtar</mods:namePart>
<mods:affiliation id="9BED735363F670067498C102F00C17D6">Museum d'Histoire Naturelle de Marrakech, Universite Cadi Ayyad, Faculte des Sciences, Semlalia, B. P. 2390, Marrakech, Morocco</mods:affiliation>
</mods:name>
<mods:name id="234419958336190A8393F18E5A272DE9" type="personal">
<mods:role id="AC69B211045B91985B957D7A024AFDB5">
<mods:roleTerm id="EA79A4ED715EC387AA20F67B9ACED4E3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="660F5DD02EA4FBE5949B96C79010D884">Sartori, Michel</mods:namePart>
<mods:nameIdentifier id="A141EF37052690EE2C03767C577348B5" type="ORCID">https://orcid.org/0000-0003-3397-3397</mods:nameIdentifier>
<mods:affiliation id="F0300C0F8DBA4474D7B1B480C6CD3724">Museum of Zoology, Palais de Rumine, Place Riponne 6, CH- 1005, Lausanne, Switzerland &amp; Department of Ecology and Evolution, Lausanne University, CH- 1015, Lausanne, Switzerland</mods:affiliation>
</mods:name>
<mods:name id="15F88C85DB64348FB994D5FA95BE8D52" type="personal">
<mods:role id="2506915EE3C1D4AD319921C51FA72BE0">
<mods:roleTerm id="3928BF4A2DEFD94AF120477BDA45203F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="60D0FA00417C38EC82A733FEBA176A82">Vuataz, Laurent</mods:namePart>
<mods:nameIdentifier id="72677D25284C652AC51E8FA6EB1D9149" type="ORCID">https://orcid.org/0000-0001-9193-8683</mods:nameIdentifier>
<mods:affiliation id="99A92F2E177DA6DE9D870468D99D4DE6">Museum of Zoology, Palais de Rumine, Place Riponne 6, CH- 1005, Lausanne, Switzerland &amp; Department of Ecology and Evolution, Lausanne University, CH- 1015, Lausanne, Switzerland</mods:affiliation>
</mods:name>
<mods:name id="206290DFFCAB18C50CCD14AB83EEEF3D" type="personal">
<mods:role id="550EBA053E311427EAFFFD967382AE11">
<mods:roleTerm id="79236A887639035EC451DA1288A611A0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="501C550F51BB763FB79BFDCDF9B7AFD5">Ghamizi, Mohamed</mods:namePart>
<mods:affiliation id="3F826EB105C804CBC13649EC83BF0872">Museum d'Histoire Naturelle de Marrakech, Universite Cadi Ayyad, Faculte des Sciences, Semlalia, B. P. 2390, Marrakech, Morocco</mods:affiliation>
</mods:name>
<mods:typeOfResource id="EF813D7CFEA904A190580F4ACFBEF6EC">text</mods:typeOfResource>
<mods:relatedItem id="02D5D5D585CA5620E077EC677D113AEE" type="host">
<mods:titleInfo id="CC1C74F11C7096315338F76616B0E3A3">
<mods:title id="DDCB6CE132A1D21EF2F06EC57645FE57">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="F15D6507B2D172F5D8D21DA3BD50C500">
<mods:date id="E980C7DE943D7664ED409AC019CFADBE">2022</mods:date>
<mods:detail id="A24A956846FA33015BD58B9A0133286F" type="pubDate">
<mods:number id="D9C1B7C0561AD669202C1F4369848BD5">2022-08-15</mods:number>
</mods:detail>
<mods:detail id="BA41658E0D8E83B6037806D0CE05AFF8" type="volume">
<mods:number id="7787145F4A8479317E674B9279C3F7C1">1117</mods:number>
</mods:detail>
<mods:extent id="FE6FFB2DDE78EC3B35E98569A7D217F1" unit="page">
<mods:start id="EEAA2B8A163A5AD234DB1DD7862DBDF3">203</mods:start>
<mods:end id="F4F8D63C0794EA3E97A87EC70D735536">218</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="23BB6645415F0AB11A5E460AECFB8E01">
<mods:url id="16D0BBA15B2AFFF9DCD7D9098002F4ED">http://dx.doi.org/10.3897/zookeys.1117.83539</mods:url>
</mods:location>
<mods:classification id="8721F52BAAC21F8BBBDDFE42DDDBDE88">journal article</mods:classification>
<mods:identifier id="714EE78617428C7CD180746C9C6E26C0" type="DOI">http://dx.doi.org/10.3897/zookeys.1117.83539</mods:identifier>
<mods:identifier id="1A6094EE9A79FA4109FAD96FC96EC810" type="Pensoft-Pub">1313-2970-1117-203</mods:identifier>
<mods:identifier id="0C6FB84E82C787393DD19591F4161AA7" type="ZooBank">B2596F5CFE214B3EAAECDE33FF394CEC</mods:identifier>
<mods:identifier id="9C9ABC03F0CC1064ED24E96DDE4E9FF0" type="Pensoft-UUID">76BC5EBEBEA65F82903D460EBF22B366</mods:identifier>
</mods:mods>
<treatment id="7E5B6A0BC24054D1860BEFB396008C61" ID-GBIF-Taxon="199352418" LSID="urn:lsid:zoobank.org:act:9C03C91D-8E32-47C7-9FD1-DE01A7093072" httpUri="http://treatment.plazi.org/id/7E5B6A0BC24054D1860BEFB396008C61" lastPageNumber="203" pageId="0" pageNumber="203" scope_family="Prosopistomatidae" scope_order="Ephemeroptera">
<subSubSection id="4C4C1495339C5BFED8A385D341005FA2" pageId="0" pageNumber="203" type="nomenclature">
<paragraph id="EF5DFFCDC4255A42C3A532930209EDD6" pageId="0" pageNumber="203">
<taxonomicName id="222B12E52FFAB852A0DC43390A811BC6" LSID="https://zoobank.org/9C03C91D-8E32-47C7-9FD1-DE01A7093072" authority="El Alami, Benlasri &amp; Sartori" authorityName="El Alami, Benlasri &amp; Sartori" authorityYear="2022" class="Insecta" family="Prosopistomatidae" genus="Prosopistoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosopistoma maroccanum" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="maroccanum" status="sp. nov.">Prosopistoma maroccanum El Alami, Benlasri &amp; Sartori</taxonomicName>
<taxonomicNameLabel id="9EE7A4E081ABE7B4F5410C7186A13173" pageId="0" pageNumber="203">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="ADD47020F403DCA5CC482F241565B5FD" pageId="0" pageNumber="203" type="materials_examined">
<paragraph id="C26C80D6C98E463949E7D27A057A4239" pageId="0" pageNumber="203">Material examined.</paragraph>
<paragraph id="85B0451BBF12083170885062CD43B571" pageId="0" pageNumber="203">
<materialsCitation id="87C672A95E4E0B68258A233967121DDB" collectingDate="2021-12-14" collectorName="M. Benlasri" country="Morocco" county="Bzou village" elevation="372" latitude="32.101265" location="Beni-Mellal Province" longLatPrecision="1" longitude="-7.0440664" municipality="Oued Laabid" specimenCount="1" typeStatus="Holotype">
<emphasis id="C81B5FFC7F75DB19A1E484A7D88B0D2D" bold="true" italics="true" pageId="0" pageNumber="203">
<typeStatus id="A8A3A727D123F7AD99C4248E1C28AC7C">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="AB1BD942B174A995D5F6CF2B7E3456D8" name="Morocco">Morocco</collectingCountry>
• 1 nymph in ethanol;
<location id="A51FB81E7F60A04596934B96E1AA358C" LSID="urn:lsid:plazi:treatment:7E5B6A0BC24054D1860BEFB396008C61:A51FB81E7F60A04596934B96E1AA358C" country="Morocco" county="Bzou village" latitude="32.101265" longLatPrecision="1" longitude="-7.0440664" municipality="Oued Laabid" name="Beni-Mellal Province">
<normalizedToken id="BD4E3F1132A9849D0867AE076010EDA9" originalValue="Béni-Mellal">Beni-Mellal</normalizedToken>
Province
</location>
,
<collectingMunicipality id="BA439B4E25CD9ED8FB1795CA6D874AE8">Oued Laabid</collectingMunicipality>
, in
<collectingCounty id="243D74EEF2FB2235B87E584BEA303767">Bzou village</collectingCounty>
,
<geoCoordinate id="ACAC8335C7D0788CFBFD5CF478255236" degrees="32" direction="north" minutes="6.076" orientation="latitude" precision="1" value="32.101265">32°6.076'N</geoCoordinate>
,
<geoCoordinate id="AE82FD74F9F6D6A13B70638F8D4E06FA" degrees="7" direction="west" minutes="2.644" orientation="longitude" precision="1" value="-7.0440664">7°2.644'W</geoCoordinate>
,
<quantity id="B96D21DA1984689B8869B432EB165E2D" metricMagnitude="2" metricUnit="m" metricValue="3.72" unit="m" value="372.0">
<elevation id="81F4D3E9B5DC94D833E2FFDF46303B61" metricMagnitude="2" metricUnit="m" metricValue="3.72" unit="m" value="372.0">372 m</elevation>
</quantity>
alt.,
<collectingDate id="D0EF842B7A03F2B182AB6F8818425A40" value="2021-12-14">14 December 2021</collectingDate>
, coll.
<collectorName id="48156DEC7931FBE07D12598D959B8416">M. Benlasri</collectorName>
(MZL GBIFCH01119080)
</materialsCitation>
.
<materialsCitation id="A590F6B1DBBB6EB817D50FE5C5917355" collectingDate="2016-03-06" collectingDateMax="2021-10-14" collectingDateMin="2016-03-06" collectorName="Coll. M. Benlasri, H. Hajjani" country="Morocco" county="Imdahen village" elevation="364" latitude="32.137535" location="Beni-Mellal Province" longLatPrecision="1" longitude="-7.0294" municipality="Oued Laabid" specimenCount="1" typeStatus="Paratypes">
<emphasis id="5E31F19AE49DEE09C04E8545E835204B" bold="true" italics="true" pageId="0" pageNumber="203">
<typeStatus id="E83BD2E15B5DFA228334B271FEFD060E">Paratypes</typeStatus>
</emphasis>
:
<collectingCountry id="DCEB107576EF3642ED6BB03A7C21C81B" name="Morocco">Morocco</collectingCountry>
• 1 nymph in ethanol, same data as holotype (MZL GBIFCH01119081); 1 nymph on slide, same data as
<typeStatus id="A31D8AB81E966DEE69A84730EF1159EE">holotype</typeStatus>
(MZL GBIFCH00608997; 7 nymphs in ethanol, same data as
<typeStatus id="2B38BB7F90AEA8DFDA912D9A1B1241E1">holotype</typeStatus>
(LESCB); 1 nymph on slide, same data (LESCB); 3 nymphs in ethanol, same sampling site,
<collectingDate id="CB94F20E30265176D52EC3DD649A37A6" value="2021-05-08">08 May 2021</collectingDate>
; 4 nymphs in ethanol, same sampling site,
<collectingDate id="CE2175A59D32AD2C51AD637C2C995107" value="2021-10-14">14 October 2021</collectingDate>
,
<collectorName id="210C848114798F1765119704406E133F">Coll. M. Benlasri</collectorName>
(MHNM) • 4 nymphs in ethanol,
<location id="127B8418CB3EDFBF8F775C409CBA23D6" LSID="urn:lsid:plazi:treatment:7E5B6A0BC24054D1860BEFB396008C61:127B8418CB3EDFBF8F775C409CBA23D6" country="Morocco" county="Imdahen village" latitude="32.137535" longLatPrecision="1" longitude="-7.0294" municipality="Oued Laabid" name="Beni-Mellal Province">
<normalizedToken id="B202F4E17E66F51D26163A96E6389F69" originalValue="Béni-Mellal">Beni-Mellal</normalizedToken>
Province
</location>
,
<collectingMunicipality id="C06AD6BEC36BB6F62B2514564E1AB251">Oued Laabid</collectingMunicipality>
, in
<collectingCounty id="31D20143DD984C4A57A8FA7DF4B13DCB">Imdahen village</collectingCounty>
,
<geoCoordinate id="E490BC754240A851D10867B68A72299B" degrees="32" direction="north" minutes="8.252" orientation="latitude" precision="1" value="32.137535">32°8.252'N</geoCoordinate>
,
<geoCoordinate id="A03A0B17417B4B052349252AE88CBAFC" degrees="7" direction="west" minutes="1.764" orientation="longitude" precision="1" value="-7.0294">7°1.764'W</geoCoordinate>
,
<quantity id="84C911D31D06ED99999C8F82A7014DCA" metricMagnitude="2" metricUnit="m" metricValue="3.6399999999999997" unit="m" value="364.0">
<elevation id="8D2BA67891339E0FC0B6B50952FFEBB9" metricMagnitude="2" metricUnit="m" metricValue="3.6399999999999997" unit="m" value="364.0">364 m</elevation>
</quantity>
alt.,
<collectingDate id="70F453664B57DEAF1B061AE37BEB354B" value="2016-03-06">06 March 2016</collectingDate>
, coll.
<collectorName id="76C672E2D0D1414F2A98F9D1FF51C1A5">H. Hajjani</collectorName>
(MZL GBIFCH00980869); 1 nymph on slide, same data (MZL GBIFCH00970951); 5 nymphs in ethanol, same data (LESCB); 5 nymphs in ethanol, same site,
<collectingDate id="0355EADF33F11160A0018BE9C6525096" value="2021-05-08">08 May 2021</collectingDate>
; 7 nymphs in ethanol, same site,
<collectingDate id="0BEB0E84A7777620F8A8A8395552BC6C" value="2021-10-14">14 October 2021</collectingDate>
,
<collectorName id="5D2C0FDA7A61055AF3509148CD38281B">Coll. M. Benlasri</collectorName>
(MHNM)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="91A283CAC1F1F1ED5979A053A5909B4E" pageId="0" pageNumber="203" type="description">
<paragraph id="1F852E890142F11C76779B92AD33D64A" pageId="0" pageNumber="203">Description.</paragraph>
<paragraph id="0C33153039523E3BBA45A918F0B7751A" pageId="0" pageNumber="203">
<emphasis id="848C9D4AFE271827066EB2986352508A" bold="true" pageId="0" pageNumber="203">Nymph</emphasis>
(in alcohol). Body length 3-4 mm excluding caudal filaments.
</paragraph>
<paragraph id="F95A39F44F66A9516DD82C7F0B29BEB3" pageId="0" pageNumber="203">
Notal shield (carapace) length along median suture 2 mm and total length of body 4.0 mm. Carapace (Fig.
<figureCitation id="74F185751C151BDE968CF0C45427DB95" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Nymphal dorsal view of Prosopistoma maroccanum sp. nov." figureDoi="10.3897/zookeys.1117.83539.figure3" httpUri="https://binary.pensoft.net/fig/729513" pageId="0" pageNumber="203">3</figureCitation>
), wider than long, width/length ratio: 1.15-1.25; head width/length ratio:2.5, head width/carapace width ratio: 0.58-0.60; distance between eyes/head width ratio: ca 0.53 for male nymphs and 0.58 for female nymphs. Carapace flange relatively wide (Fig.
<figureCitation id="E4A3F9155C6B211C6FC280EB7454C060" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Nymphal dorsal view of Prosopistoma maroccanum sp. nov." figureDoi="10.3897/zookeys.1117.83539.figure3" httpUri="https://binary.pensoft.net/fig/729513" pageId="0" pageNumber="203">3</figureCitation>
).
</paragraph>
<caption id="A2499482ACCF2DE5FB67C5AA030B1D8A" doi="10.3897/zookeys.1117.83539.figure3" httpUri="https://binary.pensoft.net/fig/729513" pageId="0" pageNumber="203" start="Figure 3" startId="F3">
<paragraph id="9824C449E25A90A845DDEF8922F6F1FC" pageId="0" pageNumber="203">
<emphasis id="17A437345CB414AB533094CDE8D98FEA" bold="true" pageId="0" pageNumber="203">Figure 3.</emphasis>
Nymphal dorsal view of
<taxonomicName id="F851846ABBFA625D4C5B463A7C224597" authorityName="El Alami, Benlasri &amp; Sartori" authorityYear="2022" class="Insecta" family="Prosopistomatidae" genus="Prosopistoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosopistoma maroccanum" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="maroccanum">
<emphasis id="60F1DFEA6638C84FC51E8A6B91B2B379" italics="true" pageId="0" pageNumber="203">Prosopistoma maroccanum</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</caption>
<paragraph id="83E9E4803F1EF10019981802022E7B0B" pageId="0" pageNumber="203">
<emphasis id="681EE2AFBB878FB818078565033101E0" bold="true" pageId="0" pageNumber="203">
<emphasis id="501BA9F5399551ADE3F5C6F7EA08A63D" italics="true" pageId="0" pageNumber="203">Head</emphasis>
.
</emphasis>
Yellowish-brown, with a brownish spot above the median ocellus and between antennae (Fig.
<figureCitation id="4F22D6EB93EFF9509E12EC326D35230C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Nymphal dorsal view of Prosopistoma maroccanum sp. nov." figureDoi="10.3897/zookeys.1117.83539.figure3" httpUri="https://binary.pensoft.net/fig/729513" pageId="0" pageNumber="203">3</figureCitation>
). Compound eyes roughly oval, blackish, slightly larger than oval lateral ocelli in male nymphs (Fig.
<figureCitation id="F1EB1AF57D3F0B7D07E1C01BED22A54B" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Prosopistoma maroccanum sp. nov. A head of larva B antennae with 7 segments C antennae with 6 segments." figureDoi="10.3897/zookeys.1117.83539.figure4" httpUri="https://binary.pensoft.net/fig/729514" pageId="0" pageNumber="203">4A</figureCitation>
). Epicranial suture hardly discernible crossing the middle part of lateral ocelli, and between compound eyes and antennal bases (Fig.
<figureCitation id="387E01D0BC73FBCCB5296E7A4856F3F3" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Prosopistoma maroccanum sp. nov. A head of larva B antennae with 7 segments C antennae with 6 segments." figureDoi="10.3897/zookeys.1117.83539.figure4" httpUri="https://binary.pensoft.net/fig/729514" pageId="0" pageNumber="203">4A</figureCitation>
) and progressing to the
<normalizedToken id="F73089334B58BB90D2952B7BB47D516A" originalValue="heads">head's</normalizedToken>
lateral margin. Antennae 7-segmented (including scape and pedicel) in mature nymphs (Fig.
<figureCitation id="6B0A4B1A8821EE543F84A5BA4EEC2E65" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Prosopistoma maroccanum sp. nov. A head of larva B antennae with 7 segments C antennae with 6 segments." figureDoi="10.3897/zookeys.1117.83539.figure4" httpUri="https://binary.pensoft.net/fig/729514" pageId="0" pageNumber="203">4B</figureCitation>
), 6-segmented in younger specimens (Fig.
<figureCitation id="4566D468B72B5AB5D7602948842A0981" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Prosopistoma maroccanum sp. nov. A head of larva B antennae with 7 segments C antennae with 6 segments." figureDoi="10.3897/zookeys.1117.83539.figure4" httpUri="https://binary.pensoft.net/fig/729514" pageId="0" pageNumber="203">4C</figureCitation>
); segment III shorter than the total length of segment IV-VI or VII (respectively 0.64
<normalizedToken id="BCB177E8951F1C1B6B116A272BCED78A" originalValue="×">x</normalizedToken>
and 0.95
<normalizedToken id="0C3AEC7B679E50C24A4D3B198B1292BF" originalValue="×">x</normalizedToken>
), length of segment III/segments IV-V ratio: 1.18-1.21; antennae not extending beyond the
<normalizedToken id="1456A579CDA05566D12FA8B2955C0964" originalValue="heads">head's</normalizedToken>
anterior edge.
</paragraph>
<caption id="2EA11291440F2FEFF031190F9B293D45" doi="10.3897/zookeys.1117.83539.figure4" httpUri="https://binary.pensoft.net/fig/729514" pageId="0" pageNumber="203" start="Figure 4" startId="F4">
<paragraph id="0E0C50C35826B695FAD9CC512C33B183" pageId="0" pageNumber="203">
<emphasis id="CD78FD446C5469D5DEDB7D1F5A3EED9D" bold="true" pageId="0" pageNumber="203">Figure 4.</emphasis>
<taxonomicName id="AC293DD6B97293FFA74729A47C96888B" authorityName="El Alami, Benlasri &amp; Sartori" authorityYear="2022" class="Insecta" family="Prosopistomatidae" genus="Prosopistoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosopistoma maroccanum" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="maroccanum">
<emphasis id="B8994C6AF74720870755AF91FE8649D6" italics="true" pageId="0" pageNumber="203">Prosopistoma maroccanum</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="97230F8C32C01BC0B23AAD826D085E81" bold="true" pageId="0" pageNumber="203">A</emphasis>
head of larva
<emphasis id="D805A698EA7FF5B6EA9659A71D5D5E55" bold="true" pageId="0" pageNumber="203">B</emphasis>
antennae with 7 segments
<emphasis id="3F502B1AF251C5DB813EFB7B556032CD" bold="true" pageId="0" pageNumber="203">C</emphasis>
antennae with 6 segments.
</paragraph>
</caption>
<paragraph id="C69D0BF91300A4306688FBB010397704" pageId="0" pageNumber="203">
Labrum (Fig.
<figureCitation id="236FA7A2421BBB169751B8B3A98BF01C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Prosopistoma maroccanum sp. nov. A labrum (dorsal view) B right mandible (ventral view) C bristles arising from base of inner canine D left maxilla (ventral view) E canine (c) and moveable strong dentisetae (d 1, d 2, d 3) F labium (ventral view) G labial-palp." figureDoi="10.3897/zookeys.1117.83539.figure5" httpUri="https://binary.pensoft.net/fig/729515" pageId="0" pageNumber="203">5A</figureCitation>
) narrow, 3.25
<normalizedToken id="36AD02A1F416A405964CC3E5BD703795" originalValue="×">x</normalizedToken>
broader than long, anterior margin convex in medial section and straight laterally, anterior margin fringed with fine setae.
</paragraph>
<caption id="28A73B7EC11A0A54B8A35E3FAEBB7183" doi="10.3897/zookeys.1117.83539.figure5" httpUri="https://binary.pensoft.net/fig/729515" pageId="0" pageNumber="203" start="Figure 5" startId="F5">
<paragraph id="3163EF0A5F11E4608E8CE4B95AC17C68" pageId="0" pageNumber="203">
<emphasis id="BF1E1F868CEBAA2D529A3E632766BF24" bold="true" pageId="0" pageNumber="203">Figure 5.</emphasis>
<taxonomicName id="5D1D9ED01CFBFEAF7208CD30866B7564" authorityName="El Alami, Benlasri &amp; Sartori" authorityYear="2022" class="Insecta" family="Prosopistomatidae" genus="Prosopistoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosopistoma maroccanum" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="maroccanum">
<emphasis id="03C1F386FC4A2816A44814D1780D629F" italics="true" pageId="0" pageNumber="203">Prosopistoma maroccanum</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="471213EFB8169D4D2DBB5B9274C02BA4" bold="true" pageId="0" pageNumber="203">A</emphasis>
labrum (dorsal view)
<emphasis id="0B43564740CDB1C096B5838EDCC04187" bold="true" pageId="0" pageNumber="203">B</emphasis>
right mandible (ventral view)
<emphasis id="CD2F4A6FFB49B28C87FD9084251B33CC" bold="true" pageId="0" pageNumber="203">C</emphasis>
bristles arising from base of inner canine
<emphasis id="B58083215E11D221E7A39C7EC6AC1EC5" bold="true" pageId="0" pageNumber="203">D</emphasis>
left maxilla (ventral view)
<emphasis id="DEEA2943CA8F7566ECA505A2CF0E71AB" bold="true" pageId="0" pageNumber="203">E</emphasis>
canine (c) and moveable strong dentisetae (d1, d2, d3)
<emphasis id="1230B4941F7C4A8193CCDBB3D726C429" bold="true" pageId="0" pageNumber="203">F</emphasis>
labium (ventral view)
<emphasis id="CD271B7945A59BA9C88CF9DC38F8FC70" bold="true" pageId="0" pageNumber="203">G</emphasis>
labial-palp.
</paragraph>
</caption>
<paragraph id="D21E753E64E2268BBAF8EB33E767B69B" pageId="0" pageNumber="203">
Left and right mandibles similar (Fig.
<figureCitation id="0E396262826E734940E4549C51A4745D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Prosopistoma maroccanum sp. nov. A labrum (dorsal view) B right mandible (ventral view) C bristles arising from base of inner canine D left maxilla (ventral view) E canine (c) and moveable strong dentisetae (d 1, d 2, d 3) F labium (ventral view) G labial-palp." figureDoi="10.3897/zookeys.1117.83539.figure5" httpUri="https://binary.pensoft.net/fig/729515" pageId="0" pageNumber="203">5B</figureCitation>
). Outer canine distinctly longer and wider than inner canine with three apical teeth, inner one slightly longer than the two others. Outer margin of outer canine with 4-6 micro teeth, inner margin of outer canine with 4 or 5 micro teeth; first distal micro tooth large, conspicuous, more than twice as long and wider than others. Mandible inner canine, one-third shorter than outer canine, rod-shaped, apically bifurcate and forming a pair of elongate and thin pointed teeth with three subapical sharply pointed teeth. 6 or 7 long serrated bristles arising from base of inner canine (Fig.
<figureCitation id="1EA5AEB9850C6010EDC8830F6B553A44" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Prosopistoma maroccanum sp. nov. A labrum (dorsal view) B right mandible (ventral view) C bristles arising from base of inner canine D left maxilla (ventral view) E canine (c) and moveable strong dentisetae (d 1, d 2, d 3) F labium (ventral view) G labial-palp." figureDoi="10.3897/zookeys.1117.83539.figure5" httpUri="https://binary.pensoft.net/fig/729515" pageId="0" pageNumber="203">5C</figureCitation>
); two first anterior bristles shorter than remaining, and one long simple bristle in the middle of outer margin of mandible.
</paragraph>
<paragraph id="1DB78978FEA558BFC215E96549BFE60A" pageId="0" pageNumber="203">
Maxillae (Fig.
<figureCitation id="AA535F82A63E7C39EFF3CD23C579B7C0" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Prosopistoma maroccanum sp. nov. A labrum (dorsal view) B right mandible (ventral view) C bristles arising from base of inner canine D left maxilla (ventral view) E canine (c) and moveable strong dentisetae (d 1, d 2, d 3) F labium (ventral view) G labial-palp." figureDoi="10.3897/zookeys.1117.83539.figure5" httpUri="https://binary.pensoft.net/fig/729515" pageId="0" pageNumber="203">5D, E</figureCitation>
) crowned by a rigid canine and three moveable, strong dentisetae of subequal length (Fig.
<figureCitation id="4D33DC832083C34638AA1DF6377F3DC9" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Prosopistoma maroccanum sp. nov. A labrum (dorsal view) B right mandible (ventral view) C bristles arising from base of inner canine D left maxilla (ventral view) E canine (c) and moveable strong dentisetae (d 1, d 2, d 3) F labium (ventral view) G labial-palp." figureDoi="10.3897/zookeys.1117.83539.figure5" httpUri="https://binary.pensoft.net/fig/729515" pageId="0" pageNumber="203">5E</figureCitation>
). A row of 2 or 3 strong serrated bristles appear below the dentisetae and a single finer bristle on proximal part of sclerotized galea (Fig.
<figureCitation id="78E7D7A5903A171ED32477D5B79C11F2" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Prosopistoma maroccanum sp. nov. A labrum (dorsal view) B right mandible (ventral view) C bristles arising from base of inner canine D left maxilla (ventral view) E canine (c) and moveable strong dentisetae (d 1, d 2, d 3) F labium (ventral view) G labial-palp." figureDoi="10.3897/zookeys.1117.83539.figure5" httpUri="https://binary.pensoft.net/fig/729515" pageId="0" pageNumber="203">5E</figureCitation>
). Maxillary palp 3-segmented; segment II long and clearly longer than segment I (1.3-1.5 times) (Fig.
<figureCitation id="62B7F9086B5B280AA1539BEE39869C29" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Prosopistoma maroccanum sp. nov. A labrum (dorsal view) B right mandible (ventral view) C bristles arising from base of inner canine D left maxilla (ventral view) E canine (c) and moveable strong dentisetae (d 1, d 2, d 3) F labium (ventral view) G labial-palp." figureDoi="10.3897/zookeys.1117.83539.figure5" httpUri="https://binary.pensoft.net/fig/729515" pageId="0" pageNumber="203">5D</figureCitation>
).
</paragraph>
<paragraph id="DB9EB52A3CB0BEBBB76340E285285D81" pageId="0" pageNumber="203">
Labial palps 3-segmented, reaching front margin of labium (Fig.
<figureCitation id="2226D3FA4D22352EDCFCF6F64DF21CA9" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Prosopistoma maroccanum sp. nov. A labrum (dorsal view) B right mandible (ventral view) C bristles arising from base of inner canine D left maxilla (ventral view) E canine (c) and moveable strong dentisetae (d 1, d 2, d 3) F labium (ventral view) G labial-palp." figureDoi="10.3897/zookeys.1117.83539.figure5" httpUri="https://binary.pensoft.net/fig/729515" pageId="0" pageNumber="203">5F</figureCitation>
). Labial palp segment II 0.9
<normalizedToken id="163B673F613AA241E898AB863E6B447A" originalValue="×">x</normalizedToken>
length of segment I, segment III 0.5
<normalizedToken id="1B3FFA8619DAFB2F5EC8EF24F89027DF" originalValue="×">x</normalizedToken>
length of segment II (Fig.
<figureCitation id="5ED51947AB27BE52BE1301289C2DAAF2" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Prosopistoma maroccanum sp. nov. A labrum (dorsal view) B right mandible (ventral view) C bristles arising from base of inner canine D left maxilla (ventral view) E canine (c) and moveable strong dentisetae (d 1, d 2, d 3) F labium (ventral view) G labial-palp." figureDoi="10.3897/zookeys.1117.83539.figure5" httpUri="https://binary.pensoft.net/fig/729515" pageId="0" pageNumber="203">5G</figureCitation>
).
</paragraph>
<paragraph id="1E64CCAFA9F325BDF5F3706E52A2DF84" pageId="0" pageNumber="203">
<emphasis id="A604CF59BD59BB27FC8B8274DA24D46E" bold="true" pageId="0" pageNumber="203">
<emphasis id="6F59A6158A852F7A432B41A2E3B467A5" italics="true" pageId="0" pageNumber="203">Thorax</emphasis>
.
</emphasis>
Carapace yellowish brown, with distinct ornamentation (Fig.
<figureCitation id="73025E9E210043A97B39A856C1648026" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Prosopistoma maroccanum sp. nov. A posteromedial part of carapace B scale-like pattern on femur (ventral view) C ventral margin of fore tibia D gill I E gill II F gill III G abdominal segments VII-IX." figureDoi="10.3897/zookeys.1117.83539.figure6" httpUri="https://binary.pensoft.net/fig/729516" pageId="0" pageNumber="203">6A</figureCitation>
) and four irregular brownish markings, two on each side of midline of central region of carapace at approximately 0.20
<normalizedToken id="04127811B29BA2E7220ACE99008A3183" originalValue="×">x</normalizedToken>
length of carapace from posterior margin of head (Fig.
<figureCitation id="29C37F3292954B9E168A15790D2E9DC3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Nymphal dorsal view of Prosopistoma maroccanum sp. nov." figureDoi="10.3897/zookeys.1117.83539.figure3" httpUri="https://binary.pensoft.net/fig/729513" pageId="0" pageNumber="203">3</figureCitation>
); also, there are markings from the anterior end of the carapace at approximately 60° angle to the midline; carapace flange and distal part of carapace transparent but not apparently translucent; relatively wide. Posteromedial part of carapace distinctly concave (Figs
<figureCitation id="55AC3FB08C1CB1FA8E60B4F9344BC85E" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Nymphal dorsal view of Prosopistoma maroccanum sp. nov." figureDoi="10.3897/zookeys.1117.83539.figure3" httpUri="https://binary.pensoft.net/fig/729513" pageId="0" pageNumber="203">3</figureCitation>
,
<figureCitation id="B5E223E5243FDB6CE34F9AB604E50995" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Prosopistoma maroccanum sp. nov. A posteromedial part of carapace B scale-like pattern on femur (ventral view) C ventral margin of fore tibia D gill I E gill II F gill III G abdominal segments VII-IX." figureDoi="10.3897/zookeys.1117.83539.figure6" httpUri="https://binary.pensoft.net/fig/729516" pageId="0" pageNumber="203">6A</figureCitation>
).
</paragraph>
<paragraph id="C0D1AECED0A902DB1568B2FF8B173A19" pageId="0" pageNumber="203">
<emphasis id="61F44EE85554E9B2098E57AE6194123C" bold="true" pageId="0" pageNumber="203">
<emphasis id="817CD3DB8CCB81FD6DC04D62197A1B6F" italics="true" pageId="0" pageNumber="203">Legs</emphasis>
.
</emphasis>
Fore femora with reticulate pattern on dorsal surface consisting of scale-like structures more accentuated at the anterior and posterior border (Fig.
<figureCitation id="CD83B6181B9E0A2FB3BD2890841F56BF" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Prosopistoma maroccanum sp. nov. A posteromedial part of carapace B scale-like pattern on femur (ventral view) C ventral margin of fore tibia D gill I E gill II F gill III G abdominal segments VII-IX." figureDoi="10.3897/zookeys.1117.83539.figure6" httpUri="https://binary.pensoft.net/fig/729516" pageId="0" pageNumber="203">6B</figureCitation>
); dorsal margin of femora with simple and fine setae; ventral margin of fore tibia with 6-8 pectinate setae (Fig.
<figureCitation id="D58E39307C6A6CB926589E5278DB2A7F" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Prosopistoma maroccanum sp. nov. A posteromedial part of carapace B scale-like pattern on femur (ventral view) C ventral margin of fore tibia D gill I E gill II F gill III G abdominal segments VII-IX." figureDoi="10.3897/zookeys.1117.83539.figure6" httpUri="https://binary.pensoft.net/fig/729516" pageId="0" pageNumber="203">6C</figureCitation>
). All tarsal claws sharp, slender, smooth, and without denticles.
</paragraph>
<paragraph id="0A8FFA06551817D673541128281B6FB8" pageId="0" pageNumber="203">
<emphasis id="9EAECA24B14BEC25C0ABBE242C51460C" bold="true" pageId="0" pageNumber="203">
<emphasis id="9181D1C4911C52FC89864B285079AB1E" italics="true" pageId="0" pageNumber="203">Abdomen</emphasis>
.
</emphasis>
Abdominal gills (Fig.
<figureCitation id="6E88D265369307930C601B37734EB0B8" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Prosopistoma maroccanum sp. nov. A posteromedial part of carapace B scale-like pattern on femur (ventral view) C ventral margin of fore tibia D gill I E gill II F gill III G abdominal segments VII-IX." figureDoi="10.3897/zookeys.1117.83539.figure6" httpUri="https://binary.pensoft.net/fig/729516" pageId="0" pageNumber="203">6D-F</figureCitation>
). Gill I upper portion lamellate with serrated margin, apically slightly asymmetric and rounded; lower section cleaved in numerous filaments with about 6-8 major branches divided into 17-22 filaments (Fig.
<figureCitation id="64FE0BF9BF85A0650907B9F6056F9335" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Prosopistoma maroccanum sp. nov. A posteromedial part of carapace B scale-like pattern on femur (ventral view) C ventral margin of fore tibia D gill I E gill II F gill III G abdominal segments VII-IX." figureDoi="10.3897/zookeys.1117.83539.figure6" httpUri="https://binary.pensoft.net/fig/729516" pageId="0" pageNumber="203">6D</figureCitation>
). Gill II with rectangular lamella (ratio width/length ca 1.10); posterior margin and outer lateral margin concave with short, pointed spines; inner lateral margin convex without spines (Fig.
<figureCitation id="696BB707E4C37073EA7EE3E86000DFEA" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Prosopistoma maroccanum sp. nov. A posteromedial part of carapace B scale-like pattern on femur (ventral view) C ventral margin of fore tibia D gill I E gill II F gill III G abdominal segments VII-IX." figureDoi="10.3897/zookeys.1117.83539.figure6" httpUri="https://binary.pensoft.net/fig/729516" pageId="0" pageNumber="203">6E</figureCitation>
), covering gills III-V appearing with multiple branching filaments, decreasing in number towards gill V (Fig.
<figureCitation id="02813E3D2AC3EA3A3DEE2CFE989451A1" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Prosopistoma maroccanum sp. nov. A posteromedial part of carapace B scale-like pattern on femur (ventral view) C ventral margin of fore tibia D gill I E gill II F gill III G abdominal segments VII-IX." figureDoi="10.3897/zookeys.1117.83539.figure6" httpUri="https://binary.pensoft.net/fig/729516" pageId="0" pageNumber="203">6F</figureCitation>
); gill III with 6 main stems; gill VI conical in shape, very small and unbranched. Abdominal segments VII-IX apparently angular with straight posterior margins; posterolateral projections nearly symmetric, with straight inner margins and relatively pointed apex (Fig.
<figureCitation id="3C2B62666B2A489685320B8486650750" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Prosopistoma maroccanum sp. nov. A posteromedial part of carapace B scale-like pattern on femur (ventral view) C ventral margin of fore tibia D gill I E gill II F gill III G abdominal segments VII-IX." figureDoi="10.3897/zookeys.1117.83539.figure6" httpUri="https://binary.pensoft.net/fig/729516" pageId="0" pageNumber="203">6G</figureCitation>
). Segment X rectangular, relatively longer than wider. Caudal filaments, retractile, short, plume-like.
</paragraph>
<caption id="A2689BC0D3C543DA492EB33ED912D20D" doi="10.3897/zookeys.1117.83539.figure6" httpUri="https://binary.pensoft.net/fig/729516" pageId="0" pageNumber="203" start="Figure 6" startId="F6">
<paragraph id="FCD73BB261E7E3DC08A31987642B3FA3" pageId="0" pageNumber="203">
<emphasis id="5D60ABE6F79CB1A218E8B50D3AA6346F" bold="true" pageId="0" pageNumber="203">Figure 6.</emphasis>
<taxonomicName id="29345AF93B1C357735EF991FFB6062AE" authorityName="El Alami, Benlasri &amp; Sartori" authorityYear="2022" class="Insecta" family="Prosopistomatidae" genus="Prosopistoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosopistoma maroccanum" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="maroccanum">
<emphasis id="3A1AC3990696C6332D96002E88B2FA84" italics="true" pageId="0" pageNumber="203">Prosopistoma maroccanum</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="1C4A92E23944437C6620B87F2FAF07C1" bold="true" pageId="0" pageNumber="203">A</emphasis>
posteromedial part of carapace
<emphasis id="431056C34A7F30D6DC76B7476227E27A" bold="true" pageId="0" pageNumber="203">B</emphasis>
scale-like pattern on femur (ventral view)
<emphasis id="3D424FE519B3A6BCF44F718D23FEC4CA" bold="true" pageId="0" pageNumber="203">C</emphasis>
ventral margin of fore tibia
<emphasis id="2D64998C336DAA2CED9954D92EA71420" bold="true" pageId="0" pageNumber="203">D</emphasis>
gill I
<emphasis id="511970B2585814881CED224111096628" bold="true" pageId="0" pageNumber="203">E</emphasis>
gill II
<emphasis id="109FC87E1BCD88A156841C4330227468" bold="true" pageId="0" pageNumber="203">F</emphasis>
gill III
<emphasis id="87A5BABCB04F2BBE8EA76A6A92C52E05" bold="true" pageId="0" pageNumber="203">G</emphasis>
abdominal segments VII-IX.
</paragraph>
</caption>
<paragraph id="7D08C2FCA21EB08424E4B877D13E31B5" pageId="0" pageNumber="203">
<emphasis id="186C331F5B4F5BD7FC5E5AF610EB7810" bold="true" pageId="0" pageNumber="203">
<emphasis id="E5AA98689017CE25991ACF7E2B0CFCD4" italics="true" pageId="0" pageNumber="203">Imago</emphasis>
.
</emphasis>
Not known.
</paragraph>
</subSubSection>
<subSubSection id="DF85D8601D2585AEF7244304FD5CC3CE" pageId="0" pageNumber="203" type="morphological remark">
<paragraph id="4C3DA526754740B3B5DBE4F46A33AC79" pageId="0" pageNumber="203">Morphological remark.</paragraph>
<paragraph id="2FC788EF76B1A0B07ECE3206BD432586" pageId="0" pageNumber="203">
As already mentioned by
<bibRefCitation id="F8E430DE63A5B1FA44EF0E02013FF7DD" DOI="https://doi.org/10.1111/j.1365-2311.1954.tb00768.x" author="Gillies, MT" journalOrPublisher="Transactions of the Royal Entomological Society of London" pageId="0" pageNumber="203" pagination="355 - 372" refId="B18" refString="Gillies, MT, 1954. The adult stages of Prosopistoma Latreille (Ephemeroptera) with descriptions of two new species from Africa. Transactions of the Royal Entomological Society of London 105 (15): 355 - 372, DOI: https://doi.org/10.1111/j.1365-2311.1954.tb00768.x" title="The adult stages of Prosopistoma Latreille (Ephemeroptera) with descriptions of two new species from Africa." url="https://doi.org/10.1111/j.1365-2311.1954.tb00768.x" volume="105" year="1954">Gillies (1954)</bibRefCitation>
and
<bibRefCitation id="D751CF6BC54F456C161477DB51E20ED0" author="Peters, WL" journalOrPublisher="Tijdschrift voor Entomologie" pageId="0" pageNumber="203" pagination="207 - 222" refId="B26" refString="Peters, WL, 1967. New species of Prosopistoma from the Oriental Region (Prosopistomatidae: Ephemeroptera). Tijdschrift voor Entomologie 110: 207 - 222" title="New species of Prosopistoma from the Oriental Region (Prosopistomatidae: Ephemeroptera)." volume="110" year="1967">Peters (1967)</bibRefCitation>
, and well documented by
<bibRefCitation id="DABF670633155C03BDFBDFE35D345984" DOI="https://doi.org/10.1080/01650420802642414" author="Dalkiran, N" journalOrPublisher="Aquatic Insects" pageId="0" pageNumber="203" pagination="119 - 131" refId="B12" refString="Dalkiran, N, 2009. A new species of Prosopistoma Latreille, 1833 (Ephemeroptera: Prosopistomatidae) from northwestern Turkey. Aquatic Insects 31 (2): 119 - 131, DOI: https://doi.org/10.1080/01650420802642414" title="A new species of Prosopistoma Latreille, 1833 (Ephemeroptera: Prosopistomatidae) from northwestern Turkey." url="https://doi.org/10.1080/01650420802642414" volume="31" year="2009">Dalkiran (2009)</bibRefCitation>
and
<bibRefCitation id="93668DB053B63E87674EABF616149FD1" DOI="https://doi.org/10.11646/zoosymposia.11.1.6" author="Schletterer, M" journalOrPublisher="Zoosymposia" pageId="0" pageNumber="203" pagination="15 - 27" refId="B33" refString="Schletterer, M, Bauernfeind, E, Lechthaler, W, 2016. Larval redescription of Prosopistoma pennigerum (Mueller, 1785) from the River Volga near Rzhev, Tver Region, Russia (Insecta: Ephemeroptera). Zoosymposia 11: 15 - 27, DOI: https://doi.org/10.11646/zoosymposia.11.1.6" title="Larval redescription of Prosopistoma pennigerum (Mueller, 1785) from the River Volga near Rzhev, Tver Region, Russia (Insecta: Ephemeroptera)." url="https://doi.org/10.11646/zoosymposia.11.1.6" volume="11" year="2016">Schletterer et al. (2016)</bibRefCitation>
, the number of segments forming the antenna is subject to some variation. In
<taxonomicName id="709ABFA3A14C8116C6BB0482258AF210" class="Insecta" kingdom="Animalia" lsidName="P. maroccanum" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="maroccanum">
<emphasis id="6E4CC2AFF0E43DA2BC6641E611436B23" italics="true" pageId="0" pageNumber="203">P. maroccanum</emphasis>
</taxonomicName>
, mature nymphs possess 7 segments, while younger ones have only 6 segments. Therefore, the ratio length of segment III vs length of the remaining segments is to be use cautiously and may have a taxonomic value only when applying to mature nymphs with 7 segments, since this ratio is around 0.95 (subequal) for 6-segmented antennae vs 0.65 (much shorter) for 7-segmented antennae.
</paragraph>
</subSubSection>
<subSubSection id="967F1A6FB04D8630BE899AC0BE51E08A" pageId="0" pageNumber="203" type="diagnosis">
<paragraph id="2CB52F8F102D78E7178AD84F506A7BED" pageId="0" pageNumber="203">Diagnosis.</paragraph>
<paragraph id="56266693CE1D7FDCE7D8C2108F3402D9" pageId="0" pageNumber="203">
The nymph of
<taxonomicName id="E31786C7F55357E20702C71CD1E4E9FB" class="Insecta" kingdom="Animalia" lsidName="P. maroccanum" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="maroccanum">
<emphasis id="839B4181D7F6A31D79B9BE29B42BA230" italics="true" pageId="0" pageNumber="203">P. maroccanum</emphasis>
</taxonomicName>
sp. nov. appears to be more closely similar to
<taxonomicName id="10054FC9684A5ACC73174EE66A3FAE27" class="Insecta" kingdom="Animalia" lsidName="P. pennigerum" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="pennigerum">
<emphasis id="3232AA28A4110E0C59E013B534609883" italics="true" pageId="0" pageNumber="203">P. pennigerum</emphasis>
</taxonomicName>
than to
<taxonomicName id="01E73F40BD5030732BA10EE2F5F8FE4B" class="Insecta" kingdom="Animalia" lsidName="P. alaini" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="alaini">
<emphasis id="8643F41435E0E81DC1099415627B9E2E" italics="true" pageId="0" pageNumber="203">P. alaini</emphasis>
</taxonomicName>
from Algeria (Table
<tableCitation id="E52CFABC3809E359BFBB203A1C242235" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphological discriminant characters between the seven Prosopistoma species from the Western Palearctic." httpUri="http://table.plazi.org/id/90B3871DB0C2BDEEECFAD0AE425B5F37" pageId="0" pageNumber="203" tableUuid="90B3871DB0C2BDEEECFAD0AE425B5F37">2</tableCitation>
).
</paragraph>
<caption id="90B3871DB0C2BDEEECFAD0AE425B5F37" ID-Table-UUID="90B3871DB0C2BDEEECFAD0AE425B5F37" httpUri="http://table.plazi.org/id/90B3871DB0C2BDEEECFAD0AE425B5F37" pageId="0" pageNumber="203" start="Table 2" startId="T2">
<paragraph id="D966BA04283A0A73540B551EBC8274ED" pageId="0" pageNumber="203">
<emphasis id="FFA7044B1AFDD0690667DD1319EEC0EC" bold="true" pageId="0" pageNumber="203">Table 2.</emphasis>
Morphological discriminant characters between the seven
<taxonomicName id="BB8EEA490D7E3A1FBE451E6073B71DA2" authorityName="Latreille" authorityYear="1833" class="Insecta" family="Prosopistomatidae" genus="Prosopistoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosopistoma" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="genus">
<emphasis id="32A343C7F8DAEF3ECC91F65D53D20EE1" italics="true" pageId="0" pageNumber="203">Prosopistoma</emphasis>
</taxonomicName>
species from the Western Palearctic.
</paragraph>
</caption>
<paragraph id="A51CDD748E0EA3C007A1AF0983A190CD" pageId="0" pageNumber="203">
<table id="24CC2A21ECFD9727FFF13C2BD40F6C45" pageId="0" pageNumber="203">
<tr id="58B892EDAFFFEB64779F2871385B9AC2" pageId="0" pageNumber="203">
<th id="1D3A706926F1A1B0F95782EB87635ABA" colspan="1" pageId="0" pageNumber="203" rowspan="1">Characters</th>
<th id="AD54FA0CCF1A39A7B4C989FF6FC76BB9" colspan="1" pageId="0" pageNumber="203" rowspan="1">
<taxonomicName id="1CF33E803EA54016FE42206C5AEFF184" class="Insecta" kingdom="Animalia" lsidName="P. maroccanum" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="maroccanum">
<emphasis id="EF1A63F09C80AB28637BEE608FC7166E" italics="true" pageId="0" pageNumber="203">P. maroccanum</emphasis>
</taxonomicName>
</th>
<th id="91B34776907FD87444D14F50803316C5" colspan="1" pageId="0" pageNumber="203" rowspan="1">
<taxonomicName id="4DED686F2BAE1CF778051A0C26D770AF" class="Insecta" kingdom="Animalia" lsidName="P. pennigerum" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="pennigerum">
<emphasis id="5FCA9782A2A7C2F998219F6405774DDA" italics="true" pageId="0" pageNumber="203">P. pennigerum</emphasis>
</taxonomicName>
</th>
<th id="D46198EB658A4813C3758EE44D61EEBF" colspan="1" pageId="0" pageNumber="203" rowspan="1">
<taxonomicName id="A615AB3A32F15A028A7BD62084CAFCF8" class="Insecta" kingdom="Animalia" lsidName="P. alaini" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="alaini">
<emphasis id="5D1691184E319480ADEE482EC3E0B2A8" italics="true" pageId="0" pageNumber="203">P. alaini</emphasis>
</taxonomicName>
</th>
<th id="2EC4196067AD244E602932E84AB7A416" colspan="1" pageId="0" pageNumber="203" rowspan="1">
<taxonomicName id="0AB4A63D9AE48742A61BEAC54524BBEB" class="Insecta" kingdom="Animalia" lsidName="P. oronti" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="oronti">
<emphasis id="962185183BEB23B0165C6C9BE55236B8" italics="true" pageId="0" pageNumber="203">P. oronti</emphasis>
</taxonomicName>
</th>
<th id="6E85A647347E09B967A10AA165B7BD5F" colspan="1" pageId="0" pageNumber="203" rowspan="1">
<taxonomicName id="223AE36302AA3A51AB7C01105211A631" class="Insecta" kingdom="Animalia" lsidName="P. orhanelicum" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="orhanelicum">
<emphasis id="B5589305FFB88CDBC178398FCFD4FB2D" italics="true" pageId="0" pageNumber="203">P. orhanelicum</emphasis>
</taxonomicName>
</th>
<th id="6D3A08A368B1367C08A3DB355B29EC0F" colspan="1" pageId="0" pageNumber="203" rowspan="1">
<taxonomicName id="C85D2F2352AF87A66E8B84CFFAA19CD7" class="Insecta" kingdom="Animalia" lsidName="P. turcica" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="turcica">
<emphasis id="4CA2A7A736C1F39C2E2F9609E110CA00" italics="true" pageId="0" pageNumber="203">P. turcica</emphasis>
</taxonomicName>
</th>
<th id="9F0CA759C9A44D0E93A816032BE28700" colspan="1" pageId="0" pageNumber="203" rowspan="1">
<taxonomicName id="0B48743F6E1215E833E3A893DEE7FAD5" class="Insecta" kingdom="Animalia" lsidName="P. helenae" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="helenae">
<emphasis id="6FF90C9A06B44992D963EE0D2603986F" italics="true" pageId="0" pageNumber="203">P. helenae</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="DDEB5DEBBA6610F861F52841FA400CA9" pageId="0" pageNumber="203">
<td id="0439CBA5A13BD2695CC89A53C5BB956B" colspan="1" pageId="0" pageNumber="203" rowspan="1">
Antennal segments in mature nymphs (
<emphasis id="DFDCE4F33E3925F9E56F38550ACAAF20" italics="true" pageId="0" pageNumber="203">N</emphasis>
)
</td>
<td id="A456E1E7A1CA4862AADBEB1723F3B6FB" colspan="1" pageId="0" pageNumber="203" rowspan="1">7</td>
<td id="872C1A3A25C9D277F95A5E62B2A314C1" colspan="1" pageId="0" pageNumber="203" rowspan="1">6</td>
<td id="C8181B00CC55483816189403C095C515" colspan="1" pageId="0" pageNumber="203" rowspan="1">7</td>
<td id="6131CC7FBE02A4E87BB34C1C9598055C" colspan="1" pageId="0" pageNumber="203" rowspan="1">5</td>
<td id="3C77C373352E4CC010C78130443131E4" colspan="1" pageId="0" pageNumber="203" rowspan="1">7</td>
<td id="5D60316937CC7F2ED6C6D292F6F67D5B" colspan="1" pageId="0" pageNumber="203" rowspan="1">6</td>
<td id="E62B07E2BBEB1EBA5C7B9C2BB79A46D9" colspan="1" pageId="0" pageNumber="203" rowspan="1">6</td>
</tr>
<tr id="422ACA6C36BC65F07509A6D4BFFF79FD" pageId="0" pageNumber="203">
<td id="07543BF620F4EB015153D0AA64D692DB" colspan="1" pageId="0" pageNumber="203" rowspan="1">Antenna reaching/not reaching anterior margin of head</td>
<td id="52AC8BEEAEB925E01F08C90F509495C2" colspan="1" pageId="0" pageNumber="203" rowspan="1">not reaching</td>
<td id="7C77E80AD4818723BDD5005614C6891E" colspan="1" pageId="0" pageNumber="203" rowspan="1">reaching</td>
<td id="02325FB7B5930FE1F06937360F15C5ED" colspan="1" pageId="0" pageNumber="203" rowspan="1">not reaching</td>
<td id="6DCBD69AC40C4557AC58AE913E79C19A" colspan="1" pageId="0" pageNumber="203" rowspan="1">not reaching</td>
<td id="D2925BA02D964D13D100D429EC71CA30" colspan="1" pageId="0" pageNumber="203" rowspan="1">not reaching</td>
<td id="31DD9A77AB3CF8DE9B036D42FD3C8C2E" colspan="1" pageId="0" pageNumber="203" rowspan="1">not reaching</td>
<td id="BA5A45886E068BC19EFAAC63516D1F80" colspan="1" pageId="0" pageNumber="203" rowspan="1">reaching</td>
</tr>
<tr id="5E8546B7AD3E0BBFF36553A1E257C75D" pageId="0" pageNumber="203">
<td id="AFCB3D8B4E9706347719012D2A208707" colspan="1" pageId="0" pageNumber="203" rowspan="1">
Setae on the right mandible (
<emphasis id="453451793B92C6EFCE960812F7862BB8" italics="true" pageId="0" pageNumber="203">N</emphasis>
)
</td>
<td id="35B6A7833AD5246CC809C355049F723A" colspan="1" pageId="0" pageNumber="203" rowspan="1">6-7</td>
<td id="578DDA5D8F542DDA52F825F3871D7CE7" colspan="1" pageId="0" pageNumber="203" rowspan="1">7-8</td>
<td id="0720ACC3328339BA1DC4666177E1E51B" colspan="1" pageId="0" pageNumber="203" rowspan="1">8-9</td>
<td id="3160CDC639EE0E50388708817EE75155" colspan="1" pageId="0" pageNumber="203" rowspan="1">7-9</td>
<td id="3EBCEF1FDBEA1F3BD20A355347D59F90" colspan="1" pageId="0" pageNumber="203" rowspan="1">7</td>
<td id="44EA9DE6924625D540BA43B182B66185" colspan="1" pageId="0" pageNumber="203" rowspan="1">5</td>
<td id="974346340D21AC430EF291EAFBAC9EEF" colspan="1" pageId="0" pageNumber="203" rowspan="1">5</td>
</tr>
<tr id="66728A86035C7D58AEFB541603A49004" pageId="0" pageNumber="203">
<td id="A1959FF471434D9FD08AC764DB5548A5" colspan="1" pageId="0" pageNumber="203" rowspan="1">
Subapical teeth on outer margin of outer canine (
<emphasis id="B438B090815F5C490EB54EC8F1111B0B" italics="true" pageId="0" pageNumber="203">N</emphasis>
)
</td>
<td id="0DAF5D762E9F3CBFD28E59C98936CEFC" colspan="1" pageId="0" pageNumber="203" rowspan="1">6-7</td>
<td id="97B95A69AA34741AD3F39A51D84AA8B2" colspan="1" pageId="0" pageNumber="203" rowspan="1">6-8</td>
<td id="637EC50DB2D917C99730FABCAD09A112" colspan="1" pageId="0" pageNumber="203" rowspan="1">5</td>
<td id="DD444A05566E251CA392F99A87ED1EDD" colspan="1" pageId="0" pageNumber="203" rowspan="1">6-9</td>
<td id="C879629D09FFDB89F3F88CC764FFF6EC" colspan="1" pageId="0" pageNumber="203" rowspan="1">7-8</td>
<td id="1EE5F5BC9FC2FDC9500B71E18BFB77B8" colspan="1" pageId="0" pageNumber="203" rowspan="1">4-5</td>
<td id="AF1E6F3B65918048D1B2DE357209CAA8" colspan="1" pageId="0" pageNumber="203" rowspan="1">4-6</td>
</tr>
<tr id="FFA5F0F3A1E9D567D7D0D4AE17A103EA" pageId="0" pageNumber="203">
<td id="8D32990DB880BA8C0C7B0467545CCFE2" colspan="1" pageId="0" pageNumber="203" rowspan="1">Pectinate setae on inner margin of fore tibia (N)</td>
<td id="F41BF9747AF4231AF236F6D431B7CB5E" colspan="1" pageId="0" pageNumber="203" rowspan="1">6-8</td>
<td id="F5E3C46A8D1FA0AFA0D280B9CD273A2C" colspan="1" pageId="0" pageNumber="203" rowspan="1">10-11</td>
<td id="9555629C501B7252F800DD6738EBFDB5" colspan="1" pageId="0" pageNumber="203" rowspan="1">10-14</td>
<td id="B4BA9F497A07366992CC97DB7A7E63B9" colspan="1" pageId="0" pageNumber="203" rowspan="1">6-7</td>
<td id="BB11D7445BFD15A3347269CA14A9A35F" colspan="1" pageId="0" pageNumber="203" rowspan="1">9-10</td>
<td id="C65B1FECDAF21F323C5484BC8278A520" colspan="1" pageId="0" pageNumber="203" rowspan="1">7</td>
<td id="DA4EAB6B4B4F78FAEBFF4DB0A7B20051" colspan="1" pageId="0" pageNumber="203" rowspan="1">6-7</td>
</tr>
<tr id="4C2147CBB289F892EF816FF38281EF49" pageId="0" pageNumber="203">
<td id="652B36B06D08B440A5885029596B5968" colspan="1" pageId="0" pageNumber="203" rowspan="1">Filaments on gill I (N)</td>
<td id="D78BC3D1DA211D4C19FF54A937DFF880" colspan="1" pageId="0" pageNumber="203" rowspan="1">17-22</td>
<td id="2921F0817AF73F54832F2CB1CAD80D66" colspan="1" pageId="0" pageNumber="203" rowspan="1">24-28</td>
<td id="AA7C42CEE476D273568D3C10966F0DA5" colspan="1" pageId="0" pageNumber="203" rowspan="1">21-23</td>
<td id="E88E471CFFD8E875B7C195480C56329C" colspan="1" pageId="0" pageNumber="203" rowspan="1">12-14</td>
<td id="C94E9B3758511F60199078554293D886" colspan="1" pageId="0" pageNumber="203" rowspan="1">&gt;40</td>
<td id="AC916A499FD66E0A65B02083E8FE2A32" colspan="1" pageId="0" pageNumber="203" rowspan="1">20-22</td>
<td id="FBC019262825D18EE5B7C818A351AC5E" colspan="1" pageId="0" pageNumber="203" rowspan="1">15-17</td>
</tr>
<tr id="8C4082E0C87D5CCE74A32CAB1CC8DA15" pageId="0" pageNumber="203">
<td id="19AC8D9D8BA26D5A35B78B7001AE296E" colspan="1" pageId="0" pageNumber="203" rowspan="1">Lateral outer margin of gill II</td>
<td id="E62E100D8D2C13632A9C00DD8A537E8B" colspan="1" pageId="0" pageNumber="203" rowspan="1">concave</td>
<td id="409EF80FA0390CA2881773656A3548C8" colspan="1" pageId="0" pageNumber="203" rowspan="1">concave</td>
<td id="4942635AB4F20C4495ACB52DC5704467" colspan="1" pageId="0" pageNumber="203" rowspan="1">concave</td>
<td id="CF35E33175966C513A1254FB45B52133" colspan="1" pageId="0" pageNumber="203" rowspan="1">concave</td>
<td id="89EC74833EE1CF201342AACB87581CE3" colspan="1" pageId="0" pageNumber="203" rowspan="1">concave</td>
<td id="435BCE026B0EA9C71E8E08B8DE38F639" colspan="1" pageId="0" pageNumber="203" rowspan="1">straight</td>
<td id="C0A38600E1D32A9A82099D34E1987C21" colspan="1" pageId="0" pageNumber="203" rowspan="1">straight</td>
</tr>
<tr id="58583A2802FF712AFE257FB545236256" pageId="0" pageNumber="203">
<td id="2EC153634CAA9762EB5D955C44B1F348" colspan="1" pageId="0" pageNumber="203" rowspan="1">Shape of distal medial margin of carapace</td>
<td id="01AB388291249D986D42772D4F93AD4C" colspan="1" pageId="0" pageNumber="203" rowspan="1">distinctly concave</td>
<td id="46A5C26F2BFDE5C4B15147E5AF6DD750" colspan="1" pageId="0" pageNumber="203" rowspan="1">shallowly concave</td>
<td id="1FC38D0B68E48C3CA55573919DEE9F9C" colspan="1" pageId="0" pageNumber="203" rowspan="1">straight</td>
<td id="543C4FF83A3A96898E4EA7579BE96565" colspan="1" pageId="0" pageNumber="203" rowspan="1">distinctly concave</td>
<td id="0B88C2F47087A34D4E17EC28E3A16422" colspan="1" pageId="0" pageNumber="203" rowspan="1">convex</td>
<td id="82CDB183DF3E8048F340B3567F4C15FE" colspan="1" pageId="0" pageNumber="203" rowspan="1">distinctly concave</td>
<td id="55B8CA0D3E6FCDEC63E3D025E41948D5" colspan="1" pageId="0" pageNumber="203" rowspan="1">distinctly concave</td>
</tr>
<tr id="3D2264B02464A217915ACDEAF13F0F87" pageId="0" pageNumber="203">
<td id="5713F66B253AEDE4A826CE4ED9EC6FA7" colspan="1" pageId="0" pageNumber="203" rowspan="1">Ornamentation of the carapace</td>
<td id="2502256201FD449E1C701FBCD00AE9BA" colspan="1" pageId="0" pageNumber="203" rowspan="1">distinct</td>
<td id="6E687B59D3D8845448E2081E3D38C096" colspan="1" pageId="0" pageNumber="203" rowspan="1">distinct</td>
<td id="1F0383B04B4CCA9392DFD461F671329C" colspan="1" pageId="0" pageNumber="203" rowspan="1">indistinct</td>
<td id="0966BA816E9F123CAC3E19711E36ADD0" colspan="1" pageId="0" pageNumber="203" rowspan="1">distinct</td>
<td id="2FA71295BDE4D193981237A2583E6615" colspan="1" pageId="0" pageNumber="203" rowspan="1">indistinct</td>
<td id="D54E00674648F1E68D369DB188FD9B05" colspan="1" pageId="0" pageNumber="203" rowspan="1">distinct</td>
<td id="26D9E867F98D8469AE92FCD19A824937" colspan="1" pageId="0" pageNumber="203" rowspan="1">indistinct</td>
</tr>
<tr id="B473872091E5AA3ED6569B85A782F8DF" pageId="0" pageNumber="203">
<td id="56B46A13658B3FAC5EAD39187C8BDA23" colspan="1" pageId="0" pageNumber="203" rowspan="1">Ratio carapace width / length</td>
<td id="558BD84DD0CCFD07E78C800C96C06594" colspan="1" pageId="0" pageNumber="203" rowspan="1">1.15-1.25</td>
<td id="D2A19A1B4866E9CE96B6D9D2A6894592" colspan="1" pageId="0" pageNumber="203" rowspan="1">0.8-0.9</td>
<td id="89DCE75EB97030F9340E9291242063A6" colspan="1" pageId="0" pageNumber="203" rowspan="1">1.1</td>
<td id="F3CF645997209F86D1D19FE5B2EC9303" colspan="1" pageId="0" pageNumber="203" rowspan="1">1.1-1.2</td>
<td id="B7397D12DA3758BA655309CBE80A77C3" colspan="1" pageId="0" pageNumber="203" rowspan="1">1.1-1.4</td>
<td id="1FF29F8340E095A2D258E9288776F88F" colspan="1" pageId="0" pageNumber="203" rowspan="1">1.13</td>
<td id="02CFCD46513F97C870242BA172B5BEC7" colspan="1" pageId="0" pageNumber="203" rowspan="1">1.2-1.3</td>
</tr>
<tr id="0F67BD79FC13513AFCD5D0BC52FC075B" pageId="0" pageNumber="203">
<td id="11CD4FFD4014A63B48B7206980204CE5" colspan="1" pageId="0" pageNumber="203" rowspan="1">Distribution</td>
<td id="0F9A6352D9FC974EA0896F7CC4E021B9" colspan="1" pageId="0" pageNumber="203" rowspan="1">Morocco</td>
<td id="7CDD36B1E05E147322EF62B96847E6F6" colspan="1" pageId="0" pageNumber="203" rowspan="1">Europe</td>
<td id="FED0622D25BEB4BEBA914568533EEBDE" colspan="1" pageId="0" pageNumber="203" rowspan="1">Algeria</td>
<td id="536D16EA5E870FA56B9BF4AF408BE6E4" colspan="1" pageId="0" pageNumber="203" rowspan="1">Levant</td>
<td id="FE37C9CA53D9FFAD0096150EA4CDA84A" colspan="1" pageId="0" pageNumber="203" rowspan="1">Turkey</td>
<td id="4BD5A313DF4AA850E7DCC3C563465C3F" colspan="1" pageId="0" pageNumber="203" rowspan="1">Turkey</td>
<td id="17590DE60713B3474017BA05736EB0CB" colspan="1" pageId="0" pageNumber="203" rowspan="1">Iraq</td>
</tr>
</table>
</paragraph>
<paragraph id="7D498A9B5540DB92A792E42952A2B9FB" pageId="0" pageNumber="203">
Indeed, it differs from the latter in several aspects, mainly the distinct ornamentation of the carapace, the lower number of setae on the right mandible (6 vs 8-9), the more numerous subapical teeth on outer margin of the outer canine (6-7 vs 5), the number of maxillary dentisetae (3 vs 4), the distinctly concave distal medial margin of carapace (almost straight in
<taxonomicName id="C0A2DA500B5CE066B7E58C8F2B95A242" class="Insecta" kingdom="Animalia" lsidName="P. alaini" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="alaini">
<emphasis id="51D277536FE286E85F44C3617C7C7D28" italics="true" pageId="0" pageNumber="203">P. alaini</emphasis>
</taxonomicName>
), and above all by the fewer pectinate setae on the inner margin of fore tibia (7-8 vs 10-14). This last character also separates
<taxonomicName id="A51D848DB7344E8D090BAFAB9F95E979" class="Insecta" kingdom="Animalia" lsidName="P. maroccanum" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="maroccanum">
<emphasis id="FC75C80805F5CB5A690A43F42E31A5EB" italics="true" pageId="0" pageNumber="203">P. maroccanum</emphasis>
</taxonomicName>
from
<taxonomicName id="ACEF130298233B9432D2386EAE2F8179" class="Insecta" kingdom="Animalia" lsidName="P. pennigerum" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="pennigerum">
<emphasis id="53CDF8497691A97381491FED53FBF0DD" italics="true" pageId="0" pageNumber="203">P. pennigerum</emphasis>
</taxonomicName>
(10-11), as well as the number of antennal segments in mature nymphs (7 vs 6), the antenna not reaching the anterior margin of the head (reaching in
<taxonomicName id="5B6779CB9096125F97B98FDE1DDF9433" class="Insecta" kingdom="Animalia" lsidName="P. pennigerum" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="pennigerum">
<emphasis id="BDBAFE5179F92C995E8740D29783D1FC" italics="true" pageId="0" pageNumber="203">P. pennigerum</emphasis>
</taxonomicName>
), and the ratio width/length of the carapace higher in
<taxonomicName id="DE533ABCF77D040D13B522A5A0BCAA37" class="Insecta" kingdom="Animalia" lsidName="P. maroccanum" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="maroccanum">
<emphasis id="B1F194C4CC6699094BACD237B6D2E8FC" italics="true" pageId="0" pageNumber="203">P. maroccanum</emphasis>
</taxonomicName>
(1.25 vs 0.9). Furthermore, gill I has the apical tip of the dorsal lamina shorter in
<taxonomicName id="5DDC4132DCB665132FACE800B58477F8" class="Insecta" kingdom="Animalia" lsidName="P. maroccanum" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="maroccanum">
<emphasis id="9094F642372A0EBB8E36E470B61DB04A" italics="true" pageId="0" pageNumber="203">P. maroccanum</emphasis>
</taxonomicName>
than in
<taxonomicName id="41620EE5CA7A436899EC49B5D5A37408" class="Insecta" kingdom="Animalia" lsidName="P. pennigerum" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="pennigerum">
<emphasis id="CDC8C9FC76E04545717A5045BE8ED50E" italics="true" pageId="0" pageNumber="203">P. pennigerum</emphasis>
</taxonomicName>
. In addition, the ventral filamentous part possesses a number of main stems which overlap (7-8 vs 8-10) but with a greater number of filaments in
<taxonomicName id="E950B9B983C3B59E63A8ED047040B663" class="Insecta" kingdom="Animalia" lsidName="P. pennigerum" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="pennigerum">
<emphasis id="F254C1417E91E0B876D65BA12A0EB82C" italics="true" pageId="0" pageNumber="203">P. pennigerum</emphasis>
</taxonomicName>
(24-28 vs 17-22).
<taxonomicName id="5DFC3A943FB8630A836FDC973246A0A7" authorityName="El Alami, Benlasri &amp; Sartori" authorityYear="2022" class="Insecta" family="Prosopistomatidae" genus="Prosopistoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosopistoma maroccanum" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="maroccanum">
<emphasis id="F139661497E1DF862AF39F483C119845" italics="true" pageId="0" pageNumber="203">Prosopistoma maroccanum</emphasis>
</taxonomicName>
differs from
<taxonomicName id="737E8D29E218C4E11CAC93741B70CF5D" class="Insecta" kingdom="Animalia" lsidName="P. oronti" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="oronti">
<emphasis id="524C9ABCE039A1651FF9AD84075DE4BD" italics="true" pageId="0" pageNumber="203">P. oronti</emphasis>
</taxonomicName>
mainly by the number of antennal segments (5 in
<taxonomicName id="E28B20D0B0A0A05F0C1768F4E04CD439" class="Insecta" kingdom="Animalia" lsidName="P. oronti" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="oronti">
<emphasis id="B1CF4A55A2F0072D8CCDF9DC00D639B8" italics="true" pageId="0" pageNumber="203">P. oronti</emphasis>
</taxonomicName>
) and the length of segment II of the antenna shorter compared to the following segments (longer in
<taxonomicName id="D64D1A57057B39E53937DDAF194F395D" class="Insecta" kingdom="Animalia" lsidName="P. oronti" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="oronti">
<emphasis id="150ECA6710732BBD114881C4BF48851F" italics="true" pageId="0" pageNumber="203">P. oronti</emphasis>
</taxonomicName>
). It differs from
<taxonomicName id="6F40E6D17EA7C11E2CD7496EBED74D97" class="Insecta" kingdom="Animalia" lsidName="P. turcica" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="turcica">
<emphasis id="93F56D71D27535DB1F73B64C6830D4A2" italics="true" pageId="0" pageNumber="203">P. turcica</emphasis>
</taxonomicName>
mainly by the number of antennal segments (6) and the setation of the right mandible, from
<taxonomicName id="BB0B9F6C2A5F761F6C836A24723939A9" class="Insecta" kingdom="Animalia" lsidName="P. orhanelicum" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="orhanelicum">
<emphasis id="D7AFC681FD12C31A89C026D849854CB9" italics="true" pageId="0" pageNumber="203">P. orhanelicum</emphasis>
</taxonomicName>
by the fewer subapical teeth on inner margin of outer canine (4-5 vs 6-7), the shape of the distal medial margin of the carapace (convex in
<taxonomicName id="A158BE3E308A078A1B0053D813C9DE84" class="Insecta" kingdom="Animalia" lsidName="P. orhanelicum" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="orhanelicum">
<emphasis id="EE391DC2DCFD6BB7B6456CCD2087C8A7" italics="true" pageId="0" pageNumber="203">P. orhanelicum</emphasis>
</taxonomicName>
), and the fewer pectinate setae on the inner margin of fore tibiae (9-10 in
<taxonomicName id="45B0A41E7920A9AE5DC8419B25E7FC93" class="Insecta" kingdom="Animalia" lsidName="P. orhanelicum" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="orhanelicum">
<emphasis id="F303961022ABDF7CF6F5D8458D81A37A" italics="true" pageId="0" pageNumber="203">P. orhanelicum</emphasis>
</taxonomicName>
). From
<taxonomicName id="5E556F70CF9666AE0782C7797D0523F4" class="Insecta" kingdom="Animalia" lsidName="P. helenae" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="helenae">
<emphasis id="0C87838AEA781BA686DA5DC2349C2A11" italics="true" pageId="0" pageNumber="203">P. helenae</emphasis>
</taxonomicName>
,
<taxonomicName id="001CA3F70347279C077CE56DBCB4EDBF" class="Insecta" kingdom="Animalia" lsidName="P. maroccanum" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="maroccanum">
<emphasis id="3EE1732321D739C2AC4F44DB0BA03617" italics="true" pageId="0" pageNumber="203">P. maroccanum</emphasis>
</taxonomicName>
differs in having abdominal segments VII-IX angular (rounded in
<taxonomicName id="5F4723AE84E0F6A01E287B4E21ACFE46" class="Insecta" kingdom="Animalia" lsidName="P. helenae" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="helenae">
<emphasis id="936F6334C9785793803C17F754C7720F" italics="true" pageId="0" pageNumber="203">P. helenae</emphasis>
</taxonomicName>
) and in the number of antennal segments and length (antenna 6-segmented and reaching the anterior margin of the head in
<taxonomicName id="055CF88F00C7FA1A56929DC818D8505B" class="Insecta" kingdom="Animalia" lsidName="P. helenae" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="helenae">
<emphasis id="1562AD10947417EFDA4C54F7C67EC64B" italics="true" pageId="0" pageNumber="203">P. helenae</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="23611F9364743CA8E22B5EE960366EF7" pageId="0" pageNumber="203" type="ecology">
<paragraph id="FC046F707688A8C1C4DD4C48235E498B" pageId="0" pageNumber="203">Ecology.</paragraph>
<paragraph id="8C9CA7C41C90C983E493CD521191CE8B" pageId="0" pageNumber="203">
<taxonomicName id="961ECAB622E0D968C5BE30C06C2C3B16" authorityName="El Alami, Benlasri &amp; Sartori" authorityYear="2022" class="Insecta" family="Prosopistomatidae" genus="Prosopistoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosopistoma maroccanum" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="maroccanum">
<emphasis id="5D3DFED9C27809D76E7EE1C6FB557F5E" italics="true" pageId="0" pageNumber="203">Prosopistoma maroccanum</emphasis>
</taxonomicName>
sp. nov. was collected at two sites in the Laabid River, 117 km from Marrakech. The greatest density, 10 specimens, was recorded in December 2021 at site 2 (Imdahen locality; Fig.
<figureCitation id="F8B121A3FDE6DB9512A21A7E2BCF667C" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Sampling sites of Prosopistoma maroccanum sp. nov. A site 1 (Bzou locality) B site 2 (Imdahen locality)." figureDoi="10.3897/zookeys.1117.83539.figure7" httpUri="https://binary.pensoft.net/fig/729517" pageId="0" pageNumber="203">7B</figureCitation>
). At this site, the bottom structure was composed of 70% pebbles, 20% gravel, and 10% silt. During the sampling campaign, water temperatures were 22.4-23.7 °C, dissolved oxygen 6.57-8.26 mg/l, pH 8.05-8.17, and conductivity 977-999
<normalizedToken id="233132CA95FECB7600270BCF8EA851F8" originalValue="µS">µS</normalizedToken>
/cm. The channel, about 6 m wide and about 20 cm deep, had a moderate current velocity and turbid water. There was almost no riparian vegetation along the stream banks and no submerged macrophyte cover.
<taxonomicName id="366C9CA0E4FB3BFC1AE6CB8C4F7BD5BF" authorityName="Latreille" authorityYear="1833" class="Insecta" family="Prosopistomatidae" genus="Prosopistoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosopistoma" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="genus">
<emphasis id="3BF0FC2277059F95145C50FE2E317485" italics="true" pageId="0" pageNumber="203">Prosopistoma</emphasis>
</taxonomicName>
species are very sensitive to organic pollution and habitat degradation (
<bibRefCitation id="76E1D9532CE2BAC32EDBB9AD31DAF5C5" DOI="https://doi.org/10.4001/003.018.0112" author="Barber-James, HM" journalOrPublisher="African Entomology" pageId="0" pageNumber="203" pagination="147 - 165" refId="B5" refString="Barber-James, HM, 2010a. Two new species of Prosopistomatidae (Ephemeroptera) from South Africa and Swaziland. African Entomology 18 (1): 147 - 165, DOI: https://doi.org/10.4001/003.018.0112" title="Two new species of Prosopistomatidae (Ephemeroptera) from South Africa and Swaziland." url="https://doi.org/10.4001/003.018.0112" volume="18" year="2010 a">Barber-James 2010a</bibRefCitation>
). However, there are small villages near the two sampled locations (Fig.
<figureCitation id="296FECBED93FC3343F0D866BBF75F100" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Sampling sites of Prosopistoma maroccanum sp. nov. A site 1 (Bzou locality) B site 2 (Imdahen locality)." figureDoi="10.3897/zookeys.1117.83539.figure7" httpUri="https://binary.pensoft.net/fig/729517" pageId="0" pageNumber="203">7A, B</figureCitation>
) which probably increased the turbidity and organic pollution in the water. These impacts probably explain the low density, or the absence of this species, at other sampled sites. The ecological aspects such as microhabitat, nutrition, life history, and phenology of
<taxonomicName id="F1E7148EEEAE6DEEA9E9C857449467A8" class="Insecta" kingdom="Animalia" lsidName="P. maroccanum" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="maroccanum">
<emphasis id="F5CA9A2C253692ECD0A0E47C12E4FDEE" italics="true" pageId="0" pageNumber="203">P. maroccanum</emphasis>
</taxonomicName>
sp. nov. should be further investigated.
</paragraph>
<caption id="FB0822ED96D6A2CB77E1BD5D49484D99" doi="10.3897/zookeys.1117.83539.figure7" httpUri="https://binary.pensoft.net/fig/729517" pageId="0" pageNumber="203" start="Figure 7" startId="F7">
<paragraph id="05AA34D688317105B8272CB110619F0F" pageId="0" pageNumber="203">
<emphasis id="8B41AEF9CF6137699C2466C2BFC560A9" bold="true" pageId="0" pageNumber="203">Figure 7.</emphasis>
Sampling sites of
<taxonomicName id="F21DACF32365B7B9AB2F259711DADF4C" authorityName="El Alami, Benlasri &amp; Sartori" authorityYear="2022" class="Insecta" family="Prosopistomatidae" genus="Prosopistoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosopistoma maroccanum" order="Ephemeroptera" pageId="0" pageNumber="203" phylum="Arthropoda" rank="species" species="maroccanum">
<emphasis id="43B16FD5BCB80C3B5976107B41194A56" italics="true" pageId="0" pageNumber="203">Prosopistoma maroccanum</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="744FD485067B90B61BA71C47BED36505" bold="true" pageId="0" pageNumber="203">A</emphasis>
site 1 (Bzou locality)
<emphasis id="E170CB9B2CEC7E9CAC6174079EAF4615" bold="true" pageId="0" pageNumber="203">B</emphasis>
site 2 (Imdahen locality).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>