217 lines
32 KiB
XML
217 lines
32 KiB
XML
<document id="25998E9FDBEA4C2D36626D12DB922832" ID-DOI="10.11646/zootaxa.3599.5.3" ID-GBIF-Dataset="8d957ec1-f5ac-478f-b9fc-286ba1389922" ID-ISSN="1175-5326" ID-Zenodo-Dep="218814" ID-ZooBank="C668DAB3-5A86-4FAB-B14C-7A05BADF149D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460568024226" checkinUser="plazi" docAuthor="Villalobos, José Luis & Álvarez, Fernando" docDate="2013" docId="410487DCB71D1D71FF2352B6FC6650C3" docLanguage="en" docName="zt03599p470.pdf" docOrigin="Zootaxa 3599 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sylvathelphusa cavernicola Villalobos & Álvarez, 2013, n. sp." docType="treatment" docVersion="8" lastPageNumber="466" masterDocId="BD3DFFA4B71B1D78FFB4536DFF9B5247" masterDocTitle="Two new genera and three new species of freshwater crabs (Crustacea: Pseudothelphusidae: Potamocarcinini) from Chiapas, Mexico" masterLastPageNumber="470" masterPageNumber="457" pageNumber="463" updateTime="1698321545706" updateUser="plazi">
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<mods:title id="4BB7358E9FD0FFC8ABE3CC19EBDB0B56">Two new genera and three new species of freshwater crabs (Crustacea: Pseudothelphusidae: Potamocarcinini) from Chiapas, Mexico</mods:title>
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<mods:name id="72C442EE1FF4C3C6DE6E55E42E0A68CF" type="personal">
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<mods:namePart id="94319198F95E6CF62BC41673245F7102">Villalobos, José Luis</mods:namePart>
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<mods:name id="9BA2F347D61DAE3D8C4AB6F813CD03D7" type="personal">
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<mods:namePart id="E0C439FEFA41B7AE32FED602B81CC245">Álvarez, Fernando</mods:namePart>
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<mods:typeOfResource id="F0FE8D4AE61647565913B0BF5937F34F">text</mods:typeOfResource>
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<mods:title id="F41ECF2DD1BF9342ADA5DD021F83E004">Zootaxa</mods:title>
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<mods:part id="07CA8F31325DADC2D4EE63FB46B2EFA0">
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<mods:date id="019F7F44233BC929B61E859B345656A6">2013</mods:date>
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<mods:detail id="B1CB7AF3A3213F50B4498A1749BD8679" type="volume">
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<mods:number id="1BF4E2A24CF4BD2427E2004AEB13624B">3599</mods:number>
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<mods:detail id="5E39AFE653EB868AA3BDC8EF2D01A710" type="issue">
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<mods:number id="57FF9D84835ACD5EDFE5A3A71003DC02">5</mods:number>
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<mods:classification id="04E422B69153E54F1758664E1AB8FF4F">journal article</mods:classification>
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<mods:identifier id="05BC04B987330C8BBBF68A6A3155D489" type="DOI">10.11646/zootaxa.3599.5.3</mods:identifier>
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<mods:identifier id="C2ADCC28BC1986359BD1D0BF959CF9E2" type="ISSN">1175-5326</mods:identifier>
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<treatment id="410487DCB71D1D71FF2352B6FC6650C3" ID-DOI="http://doi.org/10.5281/zenodo.6146614" ID-GBIF-Taxon="119558133" ID-Zenodo-Dep="6146614" LSID="urn:lsid:plazi:treatment:410487DCB71D1D71FF2352B6FC6650C3" httpUri="http://treatment.plazi.org/id/410487DCB71D1D71FF2352B6FC6650C3" lastPageId="9" lastPageNumber="466" pageId="6" pageNumber="463">
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<subSubSection id="81B76541B71D1D7EFF2352B6FE895050" pageId="6" pageNumber="463" type="nomenclature">
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<paragraph id="C91236CAB71D1D7EFF2352B6FD8253B3" blockId="6.[151,537,475,535]" box="[151,537,475,500]" pageId="6" pageNumber="463">
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<heading id="925A81A6B71D1D7EFF2352B6FD8253B3" bold="true" box="[151,537,475,500]" fontSize="10" level="2" pageId="6" pageNumber="463" reason="2">
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<emphasis id="FBD9EAD8B71D1D7EFF2352B6FD8253B3" bold="true" box="[151,537,475,500]" pageId="6" pageNumber="463">
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<taxonomicName id="0EAD4D49B71D1D7EFF2352B6FE4853B3" ID-CoL="53N3D" box="[151,467,475,500]" class="Malacostraca" family="Pseudothelphusidae" genus="Sylvathelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="463" phylum="Arthropoda" rank="species" species="cavernicola" status="sp. nov.">
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<emphasis id="FBD9EAD8B71D1D7EFF2352B6FE4853B3" bold="true" box="[151,467,475,500]" italics="true" pageId="6" pageNumber="463">Sylvathelphusa cavernicola</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="E0EA57A3B71D1D7EFE6E52B1FD8253B3" box="[474,537,476,500]" pageId="6" pageNumber="463" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="C91236CAB71D1D7EFF235292FE895050" blockId="6.[151,537,475,535]" box="[151,274,511,535]" pageId="6" pageNumber="463">
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(
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<figureCitation id="51962A4FB71D1D7EFF2B5292FF745050" box="[159,239,511,535]" captionStart="FIGURE 4" captionStartId="7.[151,250,1946,1969]" captionTargetBox="[175,1414,187,1908]" captionTargetId="figure@7.[151,1436,159,1938]" captionTargetPageId="7" captionText="FIGURE 4. Sylvathelphusa cavernicola n. sp., male holotype. a, dorsal view; b, major chela; c, third maxillipeds; d, frontal portion of carapace; e, apical view of gonopod. Abbreviations: clb, caudolateral border; mep, mesial process; mes, mesial surface." httpUri="https://zenodo.org/record/218818/files/figure.png" pageId="6" pageNumber="463">Figs. 4</figureCitation>
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,
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<figureCitation id="51962A4FB71D1D7EFF4E5292FE935050" box="[250,264,511,535]" captionStart="FIGURE 5" captionStartId="8.[151,250,1952,1975]" captionTargetBox="[167,1411,182,1909]" captionTargetId="figure@8.[151,1436,163,1931]" captionTargetPageId="8" captionText="FIGURE 5. Sylvathelphusa cavernicola n. sp., male holotype. Left first gonopod: a, mesial view; b, cephalic view; c, lateral view; d, caudal view. Abbreviations: ac, apex cavity; clb, caudolateral border; las, lateral surface; map, marginal process; mas, marginal suture; mep, mesial process; mes, mesial surface." httpUri="https://zenodo.org/record/218819/files/figure.png" pageId="6" pageNumber="463">5</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="81B76541B71D1D7EFF235125FD6450A8" pageId="6" pageNumber="463" type="materials_examined">
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<paragraph id="C91236CAB71D1D7EFF235125FD6450A8" blockId="6.[151,1437,583,2012]" pageId="6" pageNumber="463">
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<emphasis id="FBD9EAD8B71D1D7EFF235125FEDE5027" bold="true" box="[151,325,583,608]" pageId="6" pageNumber="463">
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<typeStatus id="16168868B71D1D7EFF235125FF495027" box="[151,210,584,608]" pageId="6" pageNumber="463">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="16168868B71D1D7EFEF8512AFE225018" box="[332,441,583,607]" pageId="6" pageNumber="463" type="holotype">Holotype</typeStatus>
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: male, cb
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<quantity id="0E559B2FB71D1D7EFD9E512AFD0A5018" box="[554,657,583,607]" metricMagnitude="-2" metricUnit="m" metricValue="3.28" pageId="6" pageNumber="463" unit="mm" value="32.8">32.8 mm</quantity>
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, cl
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<quantity id="0E559B2FB71D1D7EFD0D512AFCBB5018" box="[697,800,583,607]" metricMagnitude="-2" metricUnit="m" metricValue="2.03" pageId="6" pageNumber="463" unit="mm" value="20.3">20.3 mm</quantity>
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; female allotype, cb
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<quantity id="0E559B2FB71D1D7EFBBE512AFBEA5018" box="[1034,1137,583,607]" metricMagnitude="-2" metricUnit="m" metricValue="4.66" pageId="6" pageNumber="463" unit="mm" value="46.6">46.6 mm</quantity>
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, cl
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<quantity id="0E559B2FB71D1D7EFB2D512AFA9B5018" box="[1177,1280,583,607]" metricMagnitude="-2" metricUnit="m" metricValue="2.86" pageId="6" pageNumber="463" unit="mm" value="28.6">28.6 mm</quantity>
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; Cueva de las Arañas, near the town of San Fernando (
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<geoCoordinate id="AC99500DB71D1D7EFDC65101FC8250C3" box="[626,793,620,644]" direction="north" orientation="latitude" pageId="6" pageNumber="463" precision="1" value="16.857224">16°51'26.0" N</geoCoordinate>
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,
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<geoCoordinate id="AC99500DB71D1D7EFC9C5101FC4F50C3" box="[808,980,620,644]" direction="west" orientation="longitude" pageId="6" pageNumber="463" precision="1" value="-93.26709">93°16'01.5" W</geoCoordinate>
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;
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<quantity id="0E559B2FB71D1D7EFC565101FBA750C3" box="[994,1084,620,644]" metricMagnitude="3" metricUnit="m" metricValue="1.14" pageId="6" pageNumber="463" unit="m" value="1140.0">1140 m</quantity>
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), Municipio de San Fernando, Chiapas,
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<collectingCountry id="B1BA765AB71D1D7EFF4B51E2FEC350E0" box="[255,344,655,679]" name="Mexico" pageId="6" pageNumber="463">Mexico</collectingCountry>
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;
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<date id="BD13100AB71D1D7EFED751E2FE7850E0" box="[355,483,655,679]" pageId="6" pageNumber="463" value="2008-09-05">5 Sep 2008</date>
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; colls: F. Álvarez, E. Torres, K. Zárate, G. Armendáriz, A. García and J.L. Villalobos;
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<collectionCode id="AFBCAE0FB71D1D7EFF2351D9FF78508B" box="[151,227,692,716]" pageId="6" pageNumber="463">CNCR</collectionCode>
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25324.
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<typeStatus id="16168868B71D1D7EFEF551D8FE29508A" box="[321,434,693,717]" pageId="6" pageNumber="463" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="DFABFD43B71D1D7EFE0B51D9FD82508B" box="[447,537,692,716]" pageId="6" pageNumber="463" type="male">3 males</specimenCount>
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, cb 28.6-
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<quantity id="0E559B2FB71D1D7EFD3051D9FD70508B" box="[644,747,692,716]" metricMagnitude="-2" metricUnit="m" metricValue="2.41" pageId="6" pageNumber="463" unit="mm" value="24.1">24.1 mm</quantity>
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, cl 17.1-
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<quantity id="0E559B2FB71D1D7EFCFA51D9FC22508B" box="[846,953,692,716]" metricMagnitude="-2" metricUnit="m" metricValue="1.44" pageId="6" pageNumber="463" unit="mm" value="14.4">14.4 mm</quantity>
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;
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<specimenCount id="DFABFD43B71D1D7EFC7251D9FBB7508B" box="[966,1068,692,716]" pageId="6" pageNumber="463" type="female">1 female</specimenCount>
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, cb 17.0 mm, cl 11.0 mm; same locality, date and collectors as
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<typeStatus id="16168868B71D1D7EFE5A51BAFDC850A8" box="[494,595,727,751]" pageId="6" pageNumber="463" type="holotype">holotype</typeStatus>
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,
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<collectionCode id="AFBCAE0FB71D1D7EFDEA51BAFD3150A8" box="[606,682,727,751]" pageId="6" pageNumber="463">CNCR</collectionCode>
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27210.
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</paragraph>
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</subSubSection>
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<subSubSection id="81B76541B71D1D71FF735196FC005323" lastPageId="9" lastPageNumber="466" pageId="6" pageNumber="463" type="description">
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<paragraph id="C91236CAB71D1D7EFF735196FC735657" blockId="6.[151,1437,583,2012]" pageId="6" pageNumber="463">
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<emphasis id="FBD9EAD8B71D1D7EFF735196FEC25153" bold="true" box="[199,345,763,788]" pageId="6" pageNumber="463">Description.</emphasis>
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Crabs of moderate size (cb <
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<quantity id="0E559B2FB71D1D7EFD1B5191FC985153" box="[687,771,764,788]" metricMagnitude="-2" metricUnit="m" metricValue="4.7" pageId="6" pageNumber="463" unit="mm" value="47.0">47 mm</quantity>
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). In dorsal view, carapace oval, widest in the middle (cb/cl average = 1.62); dorsal surface flat, punctated, regions faintly indicated, pair of gastric pits barely distinct. Cervical grooves shallow, straight, not reaching anterolateral margin (
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<figureCitation id="51962A4FB71D1D7EFCED5029FC3A511B" box="[857,929,836,860]" captionStart="FIGURE 4" captionStartId="7.[151,250,1946,1969]" captionTargetBox="[175,1414,187,1908]" captionTargetId="figure@7.[151,1436,159,1938]" captionTargetPageId="7" captionText="FIGURE 4. Sylvathelphusa cavernicola n. sp., male holotype. a, dorsal view; b, major chela; c, third maxillipeds; d, frontal portion of carapace; e, apical view of gonopod. Abbreviations: clb, caudolateral border; mep, mesial process; mes, mesial surface." httpUri="https://zenodo.org/record/218818/files/figure.png" pageId="6" pageNumber="463">Fig. 4</figureCitation>
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a). Postfrontal lobes indistinct, separated by shallow median groove. Frontal region granulated, slightly curved towards superior frontal margin; inferior border faintly granulated, projected, divided into two convex lobes; superior border bilobed with shallow mesial notch. Superior orbital margin carinate, continuous with inferior frontal border; inferior orbital margin marked by row of granules; exorbital angle projected, narrow and acute (
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<figureCitation id="51962A4FB71D1D7EFD4F50B9FCDB51AB" box="[763,832,980,1004]" captionStart="FIGURE 4" captionStartId="7.[151,250,1946,1969]" captionTargetBox="[175,1414,187,1908]" captionTargetId="figure@7.[151,1436,159,1938]" captionTargetPageId="7" captionText="FIGURE 4. Sylvathelphusa cavernicola n. sp., male holotype. a, dorsal view; b, major chela; c, third maxillipeds; d, frontal portion of carapace; e, apical view of gonopod. Abbreviations: clb, caudolateral border; mep, mesial process; mes, mesial surface." httpUri="https://zenodo.org/record/218818/files/figure.png" pageId="6" pageNumber="463">Fig. 4</figureCitation>
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d). Anterolateral margins serrated with sharp granules along anterior half, posterior half almost smooth. Posterior margin concave.
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</paragraph>
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<paragraph id="C91236CAB71D1D7EFF735770FED95683" blockId="6.[151,1437,583,2012]" pageId="6" pageNumber="463">
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Eyes with cornea pigmented, faintly faceted, not filling orbits (
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<figureCitation id="51962A4FB71D1D7EFC355771FC5F5673" box="[897,964,1052,1076]" captionStart="FIGURE 4" captionStartId="7.[151,250,1946,1969]" captionTargetBox="[175,1414,187,1908]" captionTargetId="figure@7.[151,1436,159,1938]" captionTargetPageId="7" captionText="FIGURE 4. Sylvathelphusa cavernicola n. sp., male holotype. a, dorsal view; b, major chela; c, third maxillipeds; d, frontal portion of carapace; e, apical view of gonopod. Abbreviations: clb, caudolateral border; mep, mesial process; mes, mesial surface." httpUri="https://zenodo.org/record/218818/files/figure.png" pageId="6" pageNumber="463">Fig. 4</figureCitation>
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d). Epistome narrow, epistomial tooth low, triangular, slightly deflexed, borders smooth. Pterygostomial region covered by dense pubescence around mouthparts. Endopod of third maxilliped with outer margin of ischium sinuous, inner margin straight; external margin of merus widely curved; ratio exopod/ischium 0.52 (
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<figureCitation id="51962A4FB71D1D7EFCD357EAFC2A56D8" box="[871,945,1159,1183]" captionStart="FIGURE 4" captionStartId="7.[151,250,1946,1969]" captionTargetBox="[175,1414,187,1908]" captionTargetId="figure@7.[151,1436,159,1938]" captionTargetPageId="7" captionText="FIGURE 4. Sylvathelphusa cavernicola n. sp., male holotype. a, dorsal view; b, major chela; c, third maxillipeds; d, frontal portion of carapace; e, apical view of gonopod. Abbreviations: clb, caudolateral border; mep, mesial process; mes, mesial surface." httpUri="https://zenodo.org/record/218818/files/figure.png" pageId="6" pageNumber="463">Fig. 4</figureCitation>
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c). Aperture of efferent branchial channel subrectangular.
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</paragraph>
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<paragraph id="C91236CAB71D1D7EFF7357A2FB51546B" blockId="6.[151,1437,583,2012]" pageId="6" pageNumber="463">
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Chelipeds similar in shape, subequal in length, same color as walking legs. Major cheliped with merus subtriangular in cross section; 3.3 times as long as wide; dorsal suface smooth, internal dorsal margin with row of conical, acute spines, increasing in size distally alternating with small spinules; external margin granulated. Carpus smooth, prominent and acute subdistal spine on inner surface, preceded by two acute spinules. Palm high, compressed, surface smooth, with minute squamous granules on dorsal and ventral margins. Fingers slender, shorter than palm, not gaping, tips crossing; both fingers with triangular sharp teeth on cutting edges, smaller distally; dactylus with smaller teeth sometimes interspersed with larger ones (
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<figureCitation id="51962A4FB71D1D7EFBBF56CAFBCA57F8" box="[1035,1105,1447,1471]" captionStart="FIGURE 4" captionStartId="7.[151,250,1946,1969]" captionTargetBox="[175,1414,187,1908]" captionTargetId="figure@7.[151,1436,159,1938]" captionTargetPageId="7" captionText="FIGURE 4. Sylvathelphusa cavernicola n. sp., male holotype. a, dorsal view; b, major chela; c, third maxillipeds; d, frontal portion of carapace; e, apical view of gonopod. Abbreviations: clb, caudolateral border; mep, mesial process; mes, mesial surface." httpUri="https://zenodo.org/record/218818/files/figure.png" pageId="6" pageNumber="463">Fig. 4</figureCitation>
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b). Thoracic sternites of third maxillipeds and first pereiopods completely fused, except for small notches at lateral edges of sternum; sternal sulci s4/s5, s5/s6 and s6/s7 distinct, just failing to reach midline of thoracic sternum. All abdominal segments free. Telson subtriangular, narrower than sixth abdominal segment; lateral margins straigth, tip acute.
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</paragraph>
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<paragraph id="C91236CAB71D1D71FF73555AFC005323" blockId="6.[151,1437,583,2012]" lastBlockId="9.[151,1437,151,644]" lastPageId="9" lastPageNumber="466" pageId="6" pageNumber="463">
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Male gonopod latero-mesially compressed, slightly arched in caudal and cephalic views, distal half wider. In mesial view, marginal suture almost straight along most of gonopod, slightly concave in the middle (
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<figureCitation id="51962A4FB71D1D7EFA8D5531FA1B5433" box="[1337,1408,1628,1652]" captionStart="FIGURE 5" captionStartId="8.[151,250,1952,1975]" captionTargetBox="[167,1411,182,1909]" captionTargetId="figure@8.[151,1436,163,1931]" captionTargetPageId="8" captionText="FIGURE 5. Sylvathelphusa cavernicola n. sp., male holotype. Left first gonopod: a, mesial view; b, cephalic view; c, lateral view; d, caudal view. Abbreviations: ac, apex cavity; clb, caudolateral border; las, lateral surface; map, marginal process; mas, marginal suture; mep, mesial process; mes, mesial surface." httpUri="https://zenodo.org/record/218819/files/figure.png" pageId="6" pageNumber="463">Fig. 5</figureCitation>
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a). Marginal plate between caudal and mesial surfaces, proximal half narrow, with few long marginal setae at base; distal half wider, forming rectangular projection, distal margin rounded with prominent spine (
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<figureCitation id="51962A4FB71D1D7EFB5455C9FABC54FB" box="[1248,1319,1700,1724]" captionStart="FIGURE 5" captionStartId="8.[151,250,1952,1975]" captionTargetBox="[167,1411,182,1909]" captionTargetId="figure@8.[151,1436,163,1931]" captionTargetPageId="8" captionText="FIGURE 5. Sylvathelphusa cavernicola n. sp., male holotype. Left first gonopod: a, mesial view; b, cephalic view; c, lateral view; d, caudal view. Abbreviations: ac, apex cavity; clb, caudolateral border; las, lateral surface; map, marginal process; mas, marginal suture; mep, mesial process; mes, mesial surface." httpUri="https://zenodo.org/record/218819/files/figure.png" pageId="6" pageNumber="463">Fig. 5</figureCitation>
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a). Caudal portion of marginal plate limited by straight and incipient lateral suture. Mesial surface ending apically in tonguelike apical projection, disto-cephalically directed and longer than apical projection of marginal plate, distal margin with acute and robust spines. Mesial process as a triangular, stout, acute spine, laterad directed, forming an 85º angle with respect to principal axis of gonopod. Caudal surface with narrow, deep medial notch. Lateral suture visible through proximal two thirds; on distal third interrupted and reappearing separating marginal plate from caudal surface. In cephalic view, mesial process conical, stout, acute (
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<figureCitation id="51962A4FB71D1D7EFC135411FC7155D3" box="[935,1002,1916,1940]" captionStart="FIGURE 5" captionStartId="8.[151,250,1952,1975]" captionTargetBox="[167,1411,182,1909]" captionTargetId="figure@8.[151,1436,163,1931]" captionTargetPageId="8" captionText="FIGURE 5. Sylvathelphusa cavernicola n. sp., male holotype. Left first gonopod: a, mesial view; b, cephalic view; c, lateral view; d, caudal view. Abbreviations: ac, apex cavity; clb, caudolateral border; las, lateral surface; map, marginal process; mas, marginal suture; mep, mesial process; mes, mesial surface." httpUri="https://zenodo.org/record/218819/files/figure.png" pageId="6" pageNumber="463">Fig. 5</figureCitation>
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b). Lateral surface broad, semicircular, with one slender, acute distomedial tubercle. Caudal margin with acute spinules and distal triangular, acute spine. Apical projection of mesial surface tongue-like, with thin serrate margin. In lateral view, distal margin of mesial surface with 7 strong teeth, spine on lateral surface acute, constriction of gonopod at middle portion evident (
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<figureCitation id="51962A4FB7121D71FADB53FAFF3D5293" captionStart="FIGURE 5" captionStartId="8.[151,250,1952,1975]" captionTargetBox="[167,1411,182,1909]" captionTargetId="figure@8.[151,1436,163,1931]" captionTargetPageId="8" captionText="FIGURE 5. Sylvathelphusa cavernicola n. sp., male holotype. Left first gonopod: a, mesial view; b, cephalic view; c, lateral view; d, caudal view. Abbreviations: ac, apex cavity; clb, caudolateral border; las, lateral surface; map, marginal process; mas, marginal suture; mep, mesial process; mes, mesial surface." httpUri="https://zenodo.org/record/218819/files/figure.png" pageId="9" pageNumber="466">Fig. 5</figureCitation>
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c). In caudal view, gonopod straight. Distal half with lateral margin expanded as rounded plate with spinular edge and distal, acute spine (
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<figureCitation id="51962A4FB7121D71FE1653B2FE7D52B0" box="[418,486,223,247]" captionStart="FIGURE 5" captionStartId="8.[151,250,1952,1975]" captionTargetBox="[167,1411,182,1909]" captionTargetId="figure@8.[151,1436,163,1931]" captionTargetPageId="8" captionText="FIGURE 5. Sylvathelphusa cavernicola n. sp., male holotype. Left first gonopod: a, mesial view; b, cephalic view; c, lateral view; d, caudal view. Abbreviations: ac, apex cavity; clb, caudolateral border; las, lateral surface; map, marginal process; mas, marginal suture; mep, mesial process; mes, mesial surface." httpUri="https://zenodo.org/record/218819/files/figure.png" pageId="9" pageNumber="466">Fig. 5</figureCitation>
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d). In apical view, apical projection of mesial surface with thin distal margin serrate (
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<figureCitation id="51962A4FB7121D71FF2B5269FF7E535B" box="[159,229,260,284]" captionStart="FIGURE 4" captionStartId="7.[151,250,1946,1969]" captionTargetBox="[175,1414,187,1908]" captionTargetId="figure@7.[151,1436,159,1938]" captionTargetPageId="7" captionText="FIGURE 4. Sylvathelphusa cavernicola n. sp., male holotype. a, dorsal view; b, major chela; c, third maxillipeds; d, frontal portion of carapace; e, apical view of gonopod. Abbreviations: clb, caudolateral border; mep, mesial process; mes, mesial surface." httpUri="https://zenodo.org/record/218818/files/figure.png" pageId="9" pageNumber="466">Fig. 4</figureCitation>
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e), caudal surface concave, with few short setae in the apical cavity. Mesial process conical, stout directed laterad. Apex cavity elongated, partially covered by hood formed by distal crest of lateral surface, with few setae. Caudal border of lateral plate ending distally in acute spine (
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<figureCitation id="51962A4FB7121D71FC885221FC1B5323" box="[828,896,332,356]" captionStart="FIGURE 4" captionStartId="7.[151,250,1946,1969]" captionTargetBox="[175,1414,187,1908]" captionTargetId="figure@7.[151,1436,159,1938]" captionTargetPageId="7" captionText="FIGURE 4. Sylvathelphusa cavernicola n. sp., male holotype. a, dorsal view; b, major chela; c, third maxillipeds; d, frontal portion of carapace; e, apical view of gonopod. Abbreviations: clb, caudolateral border; mep, mesial process; mes, mesial surface." httpUri="https://zenodo.org/record/218818/files/figure.png" pageId="9" pageNumber="466">Fig. 4</figureCitation>
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e).
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</paragraph>
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</subSubSection>
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<caption id="9DD26642B71C1D7FFF2354F7FF7C55A9" httpUri="https://zenodo.org/record/218818/files/figure.png" pageId="7" pageNumber="464" targetBox="[175,1414,187,1908]" targetPageId="7">
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<paragraph id="C91236CAB71C1D7FFF2354F7FF7C55A9" blockId="7.[151,1436,1946,2030]" pageId="7" pageNumber="464">
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<emphasis id="FBD9EAD8B71C1D7FFF2354F7FE8F55F7" bold="true" box="[151,276,1946,1969]" pageId="7" pageNumber="464">FIGURE 4.</emphasis>
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<taxonomicName id="0EAD4D49B71C1D7FFEA854F7FDAE55F7" box="[284,565,1946,1968]" class="Malacostraca" family="Pseudothelphusidae" genus="Sylvathelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="464" phylum="Arthropoda" rank="species" species="cavernicola" status="sp. nov.">
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<emphasis id="FBD9EAD8B71C1D7FFEA854F7FDAE55F7" box="[284,565,1946,1968]" italics="true" pageId="7" pageNumber="464">Sylvathelphusa cavernicola</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="E0EA57A3B71C1D7FFD8854F6FDEC55F7" box="[572,631,1947,1968]" pageId="7" pageNumber="464" rank="species">
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<emphasis id="FBD9EAD8B71C1D7FFD8854F6FDEC55F7" bold="true" box="[572,631,1947,1968]" pageId="7" pageNumber="464">n. sp.</emphasis>
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</taxonomicNameLabel>
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, male holotype. a, dorsal view; b, major chela; c, third maxillipeds; d, frontal portion of carapace; e, apical view of gonopod. Abbreviations: clb, caudolateral border; mep, mesial process; mes, mesial surface.
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</paragraph>
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</caption>
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<caption id="9DD26642B7131D70FF2354CDFD7055B4" httpUri="https://zenodo.org/record/218819/files/figure.png" pageId="8" pageNumber="465" targetBox="[167,1411,182,1909]" targetPageId="8">
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<paragraph id="C91236CAB7131D70FF2354CDFD7055B4" blockId="8.[151,1437,1952,2035]" pageId="8" pageNumber="465">
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<emphasis id="FBD9EAD8B7131D70FF2354CDFE8F55F1" bold="true" box="[151,276,1952,1975]" pageId="8" pageNumber="465">FIGURE 5.</emphasis>
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<taxonomicName id="0EAD4D49B7131D70FEAF54CDFDAF55F1" box="[283,564,1952,1974]" class="Malacostraca" family="Pseudothelphusidae" genus="Sylvathelphusa" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="465" phylum="Arthropoda" rank="species" species="cavernicola" status="sp. nov.">
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<emphasis id="FBD9EAD8B7131D70FEAF54CDFDAF55F1" box="[283,564,1952,1974]" italics="true" pageId="8" pageNumber="465">Sylvathelphusa cavernicola</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="E0EA57A3B7131D70FD8F54CCFDEE55F1" box="[571,629,1953,1974]" pageId="8" pageNumber="465" rank="species">
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<emphasis id="FBD9EAD8B7131D70FD8F54CCFDEE55F1" bold="true" box="[571,629,1953,1974]" pageId="8" pageNumber="465">n. sp.</emphasis>
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</taxonomicNameLabel>
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, male holotype. Left first gonopod: a, mesial view; b, cephalic view; c, lateral view; d, caudal view. Abbreviations: ac, apex cavity; clb, caudolateral border; las, lateral surface; map, marginal process; mas, marginal suture; mep, mesial process; mes, mesial surface.
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</paragraph>
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</caption>
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<subSubSection id="81B76541B7121D71FF73521DFC6650C3" pageId="9" pageNumber="466" type="discussion">
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<paragraph id="C91236CAB7121D71FF73521DFC6650C3" blockId="9.[151,1437,151,644]" pageId="9" pageNumber="466">
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<emphasis id="FBD9EAD8B7121D71FF73521DFEA053CF" bold="true" box="[199,315,368,392]" pageId="9" pageNumber="466">Remarks.</emphasis>
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<taxonomicName id="0EAD4D49B7121D71FEF2521DFD1B53CF" box="[326,640,368,392]" class="Malacostraca" family="Pseudothelphusidae" genus="Sylvathelphusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="466" phylum="Arthropoda" rank="species" species="cavernicola" status="sp. nov.">
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<emphasis id="FBD9EAD8B7121D71FEF2521DFD1B53CF" box="[326,640,368,392]" italics="true" pageId="9" pageNumber="466">Sylvathelphusa cavernicola</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="E0EA57A3B7121D71FD38521DFD4B53CF" box="[652,720,368,392]" pageId="9" pageNumber="466" rank="species">
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<emphasis id="FBD9EAD8B7121D71FD38521DFD4B53CF" bold="true" box="[652,720,368,392]" pageId="9" pageNumber="466">n. sp.</emphasis>
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</taxonomicNameLabel>
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represents the third stygobitic freshwater crab of the family
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<taxonomicName id="0EAD4D49B7121D71FF2352F9FEEE53EB" box="[151,373,404,428]" class="Malacostraca" family="Pseudothelphusidae" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="466" phylum="Arthropoda" rank="family">Pseudothelphusidae</taxonomicName>
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recorded in Chiapas after
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<taxonomicName id="0EAD4D49B7121D71FD1552F9FB0D53EB" authority="Rioja, 1952" authorityName="Rioja" authorityYear="1952" box="[673,1174,404,428]" class="Malacostraca" family="Pseudothelphusidae" genus="Typhlopseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="466" phylum="Arthropoda" rank="species" species="mocinoi">
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<emphasis id="FBD9EAD8B7121D71FD1552F9FB9153EB" box="[673,1034,404,428]" italics="true" pageId="9" pageNumber="466">Typhlopseudothelphusa mocinoi</emphasis>
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<bibRefCitation id="AD3C4B3BB7121D71FBA652F9FB0D53EB" author="Rioja" box="[1042,1174,404,428]" pageId="9" pageNumber="466" refString="Rioja, E. (1952). Estudios carcinologicos. XXVIII. Descripcion de un nuevo genero de potamonidos cavernicolas y ciegos de la Cueva del Tio Ticho, Comitan, Chis. Anales del Instituto de Biologia, Universidad Nacional Autonoma de Mexico, 23, 217 - 225." type="journal article" year="1952">Rioja, 1952</bibRefCitation>
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</taxonomicName>
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and
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<taxonomicName id="0EAD4D49B7121D71FB7952F8FED05388" authority="Rodriguez & Hobbs, 1989" authorityName="Rodriguez & Hobbs" authorityYear="1989" class="Malacostraca" family="Pseudothelphusidae" genus="Typhlopseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="466" phylum="Arthropoda" rank="species" species="hyba">
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<emphasis id="FBD9EAD8B7121D71FB7952F8FA8653EB" box="[1229,1309,404,428]" italics="true" pageId="9" pageNumber="466">T. hyba</emphasis>
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<bibRefCitation id="AD3C4B3BB7121D71FA9352F9FED05388" author="Rodriguez" pageId="9" pageNumber="466" refString="Rodriguez, G. & Hobbs, H. H. Jr. (1989) Freshwater crabs associated with caves in southern Mexico and Belize, with descriptions of three new species. Proceedings of the Biological Society of Washington, 102, 394 - 400." type="journal article" year="1989">Rodríguez & Hobbs, 1989</bibRefCitation>
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</taxonomicName>
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. Interestingly, two different lineages represented by these two genera have colonized the karstic environment in a relatively reduced area in central Chiapas. Morphologically the gonopod of
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<taxonomicName id="0EAD4D49B7121D71FB0052B0FACE53B3" box="[1204,1365,476,500]" class="Malacostraca" family="Pseudothelphusidae" genus="Sylvathelphusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="466" phylum="Arthropoda" rank="species" species="cavernicola" status="sp. nov.">
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<emphasis id="FBD9EAD8B7121D71FB0052B0FACE53B3" box="[1204,1365,476,500]" italics="true" pageId="9" pageNumber="466">S. cavernicola</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="E0EA57A3B7121D71FAE852B1FA0053B3" box="[1372,1435,476,500]" pageId="9" pageNumber="466" rank="species">
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<emphasis id="FBD9EAD8B7121D71FAE852B1FA0053B3" bold="true" box="[1372,1435,476,500]" pageId="9" pageNumber="466">n. sp.</emphasis>
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</taxonomicNameLabel>
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is different from those of the species of
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<taxonomicName id="0EAD4D49B7121D71FDE1516DFCC6505F" box="[597,861,512,536]" class="Malacostraca" family="Pseudothelphusidae" genus="Typhlopseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="466" phylum="Arthropoda" rank="genus">
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<emphasis id="FBD9EAD8B7121D71FDE1516DFCC6505F" box="[597,861,512,536]" italics="true" pageId="9" pageNumber="466">Typhlopseudothelphusa</emphasis>
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</taxonomicName>
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in that in the latter it is bent mesially, and there are no supra-apical processes, being more similar to
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<taxonomicName id="0EAD4D49B7121D71FD015148FCF0507B" box="[693,875,549,572]" class="Malacostraca" family="Pseudothelphusidae" genus="Potamocarcinus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="466" phylum="Arthropoda" rank="genus">
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<emphasis id="FBD9EAD8B7121D71FD015148FCF0507B" box="[693,875,549,572]" italics="true" pageId="9" pageNumber="466">Potamocarcinus</emphasis>
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</taxonomicName>
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where the gonopod is straight and stout. The most evident differences with
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<taxonomicName id="0EAD4D49B7121D71FE1B5125FD2C5027" box="[431,695,584,608]" class="Malacostraca" family="Pseudothelphusidae" genus="Typhlopseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="466" phylum="Arthropoda" rank="genus">
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<emphasis id="FBD9EAD8B7121D71FE1B5125FD2C5027" box="[431,695,584,608]" italics="true" pageId="9" pageNumber="466">Typhlopseudothelphusa</emphasis>
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</taxonomicName>
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are the stoutness of the gonopod, with the marginal plate and the distal crest of the mesial surface distally projected beyond the mesial process.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |