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<mods:title id="5027EB418E848D8D6024FE4305B48DC9">Haplohyphes (Ephemeroptera: Leptohyphidae), new species and stage descriptions with a key to separate the species of the genus</mods:title>
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<taxonomicName id="79FC4D11E16CAC2FFE80FA07FD201525" ID-CoL="3JMT4" authority="Dominguez" authorityName="Dominguez" box="[264,654,1524,1550]" class="Insecta" family="Leptohyphidae" genus="Haplohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="baritu">
<emphasis id="8C88EA80E16CAC2FFE80FA07FE561525" bold="true" box="[264,504,1524,1550]" italics="true" pageId="3" pageNumber="4">Haplohyphes baritu</emphasis>
Domínguez
</taxonomicName>
(
<figureCitation id="26C72A17E16CAC2FFD15FA07FCB61525" box="[669,792,1524,1550]" captionStart="FIGURES 1 ­ 19" captionStartId="2.[264,382,1681,1705]" captionTargetBox="[273,1303,333,1649]" captionTargetId="figure@2.[273,1314,321,1654]" captionTargetPageId="2" captionText="FIGURES 1 ­ 19. Haplohyphes aquilonius Lugo­Ortiz and McCafferty: 1 a, male fore wing; 1 b, male hind wing; 1 c, idem, detail; 2 a, female fore wing; 2 b, female hind wing; 2 c, idem, detail; 3, nymphal head, d. v.; 4, male genitalia, v. v. H. dominguezi sp. nov.: 5 a, male fore wing; 5 b, male hind wing; 5 c, idem, detail; 6, male genitalia, v. v. H. huallaga Allen: 7, male hind wing, detail. H. mithras (Traver): 8, male hind wing, detail. H. baritu Domínguez: 9 a, female fore wing; 9 b, female hind wing; 9 c, idem, detail; 10 a, male fore wing; 10 b, male hind wing; 10 c, idem, detail; 11, male genitalia, v. v.; 12, detail of forceps, l. v.; 13, nymphal head, d. v.; 14, gill II, d. v.; 15 ­ 19, gills II­VI, v. v." httpUri="https://zenodo.org/record/156637/files/figure.png" pageId="3" pageNumber="4">Figs. 9­19</figureCitation>
, 
<figureCitation id="26C72A17E16CAC2FFCAEFA07FCEB1525" box="[806,837,1524,1550]" captionStart="FIGURES 42 ­ 43" captionStartId="8.[264,382,1806,1830]" captionTargetBox="[284,1303,1116,1795]" captionTargetId="figure@8.[276,1310,1108,1799]" captionTargetPageId="8" captionText="FIGURES 42 ­ 43. Haplohyphes baritu Domínguez, SEM photographs: 42, group of eggs; 43, general view of egg; 44, a pair of eggs attached by adhesive filaments; 45, micropylar area. Scale bars = 10 µ." httpUri="https://zenodo.org/record/156638/files/figure.png" pageId="3" pageNumber="4">42</figureCitation>
­45)
</emphasis>
</heading>
</paragraph>
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<paragraph id="BE433692E16CAC2FFE80F9CCFDAF155E" blockId="3.[264,1322,1598,1750]" pageId="3" pageNumber="4">
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<taxonomicName id="79FC4D11E16CAC2FFE80F9CCFDC6157E" ID-CoL="3JMT4" authority="Dominguez, 1984: 106" authorityName="Dominguez" authorityPageNumber="106" authorityYear="1984" box="[264,616,1598,1622]" class="Insecta" family="Leptohyphidae" genus="Haplohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="baritu">
<emphasis id="8C88EA80E16CAC2FFE80F9CCFEC7157D" box="[264,361,1599,1622]" italics="true" pageId="3" pageNumber="4">H. baritu</emphasis>
<treatmentCitation id="3F5D1083E16CAC2FFEF8F9CDFDC6157E" author="Dominguez" box="[368,616,1598,1621]" page="106" pageId="3" pageNumber="4" year="1984">
<bibRefCitation id="DA6D4B63E16CAC2FFEF8F9CDFD9D157E" author="Dominguez" box="[368,563,1598,1621]" pageId="3" pageNumber="11" refString="Dominguez, E. (1984) Dos especies nuevas del genero Haplohyphes Allen (Ephemeroptera: Tricorythidae) de la Argentina. Revista de la Sociedad Entomologica Argentina, 43 (1 - 4), 103 - 112." type="journal article" year="1984">Domínguez, 1984</bibRefCitation>
: 106
</treatmentCitation>
</taxonomicName>
; 
<treatmentCitation id="3F5D1083E16CAC2FFDFBF9CDFC3F157E" author="Dominguez" box="[627,913,1598,1621]" page="97" pageId="3" pageNumber="4" year="1994">
<bibRefCitation id="DA6D4B63E16CAC2FFDFBF9CDFCC7157E" author="Dominguez" box="[627,873,1598,1621]" pageId="3" pageNumber="11" refString="Dominguez, E.; Hubbard M.. &amp; Pescador, M. L. (1994) Los Ephemeroptera en Argentina. Fauna de Agua Dulce de la Republica Argentina, 33 (1), 1 - 142." type="journal article" year="1994">Domínguez et al. 1994</bibRefCitation>
: 97
</treatmentCitation>
; 
<treatmentCitation id="3F5D1083E16CAC2FFC14F9CDFA88157E" author="Lugo-Ortiz" box="[924,1318,1598,1621]" page="169" pageId="3" pageNumber="4" year="1995">
<bibRefCitation id="DA6D4B63E16CAC2FFC14F9CDFB5F157E" author="Lugo-Ortiz" box="[924,1265,1598,1621]" pageId="3" pageNumber="11" refString="Lugo-Ortiz, C. R. &amp; McCafferty, W. P. (1995) Contribution to the Taxonomy of the Leptohyphidae (Insecta: Ephemeroptera) of Central America. Studies on Neotropical Fauna and Environment 30 (3), 165 - 176." type="journal article" year="1995">Lugo­Ortiz &amp; McCafferty 1995</bibRefCitation>
: 169
</treatmentCitation>
; 
<treatmentCitation id="3F5D1083E16CAC2FFEB0F9ADFE53155E" author="Molineri" box="[312,509,1630,1653]" page="29" pageId="3" pageNumber="4" year="1999">
<bibRefCitation id="DA6D4B63E16CAC2FFEB0F9ADFE7B155E" author="Molineri" box="[312,469,1630,1653]" pageId="3" pageNumber="11" refString="Molineri, C. (1999) Descripcion de una nueva especie de Haplohyphes Allen (Ephemeroptera: Leptohyphidae) y clave para separar los imagos machos del genero. Acta Zoologica Lilloana, 45 (1), 29 - 32." type="journal article" year="1999">Molineri 1999</bibRefCitation>
: 29
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.
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</paragraph>
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<taxonomicName id="79FC4D11E16CAC2FFE80F98CFD4F15BE" authority="Dominguez, 1984: 108" authorityName="Dominguez" authorityPageNumber="108" authorityYear="1984" box="[264,737,1662,1686]" class="Insecta" family="Leptohyphidae" genus="Haplohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="furtiva">
<emphasis id="8C88EA80E16CAC2FFE80F98CFE4C15BD" box="[264,482,1663,1686]" italics="true" pageId="3" pageNumber="4">Haplohyphes furtiva</emphasis>
<treatmentCitation id="3F5D1083E16CAC2FFE61F98DFD4F15BE" author="Dominguez" box="[489,737,1662,1685]" page="108" pageId="3" pageNumber="4" year="1984">
<bibRefCitation id="DA6D4B63E16CAC2FFE61F98DFD0215BE" author="Dominguez" box="[489,684,1662,1685]" pageId="3" pageNumber="11" refString="Dominguez, E. (1984) Dos especies nuevas del genero Haplohyphes Allen (Ephemeroptera: Tricorythidae) de la Argentina. Revista de la Sociedad Entomologica Argentina, 43 (1 - 4), 103 - 112." type="journal article" year="1984">Domínguez, 1984</bibRefCitation>
: 108
</treatmentCitation>
</taxonomicName>
. SYN. NOV. 
<taxonomicName id="79FC4D11E16CAC2FFE80F96CFD3B159E" authority="Dominguez et al. 1994: 98" authorityName="Dominguez et al." authorityPageNumber="98" authorityYear="1994" box="[264,661,1694,1718]" class="Insecta" family="Leptohyphidae" genus="Haplohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="furtiva">
<emphasis id="8C88EA80E16CAC2FFE80F96CFEC1159D" box="[264,367,1695,1718]" italics="true" pageId="3" pageNumber="4">H. furtiva</emphasis>
<treatmentCitation id="3F5D1083E16CAC2FFEFEF96DFD3B159E" author="Dominguez" box="[374,661,1694,1717]" page="98" pageId="3" pageNumber="4" year="1994">
<bibRefCitation id="DA6D4B63E16CAC2FFEFEF96DFDC3159E" author="Dominguez" box="[374,621,1694,1717]" pageId="3" pageNumber="11" refString="Dominguez, E.; Hubbard M.. &amp; Pescador, M. L. (1994) Los Ephemeroptera en Argentina. Fauna de Agua Dulce de la Republica Argentina, 33 (1), 1 - 142." type="journal article" year="1994">Domínguez et al. 1994</bibRefCitation>
: 98
</treatmentCitation>
</taxonomicName>
; 
<treatmentCitation id="3F5D1083E16CAC2FFD28F96DFCCB159E" author="Molineri" box="[672,869,1694,1717]" page="29" pageId="3" pageNumber="4" year="1999">
<bibRefCitation id="DA6D4B63E16CAC2FFD28F96DFC93159E" author="Molineri" box="[672,829,1694,1717]" pageId="3" pageNumber="11" refString="Molineri, C. (1999) Descripcion de una nueva especie de Haplohyphes Allen (Ephemeroptera: Leptohyphidae) y clave para separar los imagos machos del genero. Acta Zoologica Lilloana, 45 (1), 29 - 32." type="journal article" year="1999">Molineri 1999</bibRefCitation>
: 29
</treatmentCitation>
. 
<taxonomicName id="79FC4D11E16CAC2FFE80F94CFCA215FE" authority="Lugo-Ortiz &amp; McCafferty 1995: 169" authorityName="Lugo-Ortiz &amp; McCafferty" authorityPageNumber="169" authorityYear="1995" box="[264,780,1726,1750]" class="Insecta" family="Leptohyphidae" genus="Haplohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="furtivus">
<emphasis id="8C88EA80E16CAC2FFE80F94CFED715FD" box="[264,377,1727,1750]" italics="true" pageId="3" pageNumber="4">H. furtivus</emphasis>
<treatmentCitation id="3F5D1083E16CAC2FFE09F94DFCA215FE" author="Lugo-Ortiz" box="[385,780,1726,1749]" page="169" pageId="3" pageNumber="4" year="1995">
<bibRefCitation id="DA6D4B63E16CAC2FFE09F94DFD7915FE" author="Lugo-Ortiz" box="[385,727,1726,1749]" pageId="3" pageNumber="11" refString="Lugo-Ortiz, C. R. &amp; McCafferty, W. P. (1995) Contribution to the Taxonomy of the Leptohyphidae (Insecta: Ephemeroptera) of Central America. Studies on Neotropical Fauna and Environment 30 (3), 165 - 176." type="journal article" year="1995">Lugo­Ortiz &amp; McCafferty 1995</bibRefCitation>
: 169
</treatmentCitation>
</taxonomicName>
.
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</paragraph>
<paragraph id="BE433692E16CAC28FE80F8F4FE3012AD" blockId="3.[264,1323,1799,1905]" lastBlockId="4.[264,1325,284,1870]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
Egg (
<figureCitation id="26C72A17E16CAC2FFEC3F8F4FE1D140A" box="[331,435,1799,1825]" captionStart="FIGURES 42 ­ 43" captionStartId="8.[264,382,1806,1830]" captionTargetBox="[284,1303,1116,1795]" captionTargetId="figure@8.[276,1310,1108,1799]" captionTargetPageId="8" captionText="FIGURES 42 ­ 43. Haplohyphes baritu Domínguez, SEM photographs: 42, group of eggs; 43, general view of egg; 44, a pair of eggs attached by adhesive filaments; 45, micropylar area. Scale bars = 10 µ." httpUri="https://zenodo.org/record/156638/files/figure.png" pageId="3" pageNumber="4">Figs. 42</figureCitation>
­45). Length, 110­130 µ; maximum width, 65­75 µ. One blunt polar cap present (
<figureCitation id="26C72A17E16CAC2FFEFBF8DCFDA81462" box="[371,518,1839,1865]" captionStart="FIGURES 42 ­ 43" captionStartId="8.[264,382,1806,1830]" captionTargetBox="[284,1303,1116,1795]" captionTargetId="figure@8.[276,1310,1108,1799]" captionTargetPageId="8" captionText="FIGURES 42 ­ 43. Haplohyphes baritu Domínguez, SEM photographs: 42, group of eggs; 43, general view of egg; 44, a pair of eggs attached by adhesive filaments; 45, micropylar area. Scale bars = 10 µ." httpUri="https://zenodo.org/record/156638/files/figure.png" pageId="3" pageNumber="4">Figs. 42­43</figureCitation>
). One subcircular micropylar area, surrounded by five chorionic plates (Fig. 45). Polygonal chorionic plates (Figs. 44­45) contiguous; with the margin opposed to capped pole, elevated. Adhesive filaments (probably derived KTC) relatively long and thin (Fig. 44), sometimes widening at the apex very slightly, scattered on chorionic surface.
</paragraph>
</subSubSection>
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<paragraph id="BE433692E16BAC28FEB0FE67FC161705" blockId="4.[264,1325,284,1870]" pageId="4" pageNumber="5">
Material. 
<taxonomicName id="79FC4D11E16BAC28FE27FE67FD301286" box="[431,670,404,429]" class="Insecta" family="Leptohyphidae" genus="Haplohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="baritu">
<emphasis id="8C88EA80E16BAC28FE27FE67FD301286" box="[431,670,404,429]" italics="true" pageId="4" pageNumber="5">Haplohyphes baritu</emphasis>
</taxonomicName>
: 
<typeStatus id="61478830E16BAC28FD3BFE67FCB41285" box="[691,794,404,430]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
male subimago, allotype female subimago and 28 
<typeStatus id="61478830E16BAC28FED7FE4FFE6A12FD" box="[351,452,444,470]" pageId="4" pageNumber="5" type="paratype">paratype</typeStatus>
nymphs from 
<collectingCountry id="C6EB7602E16BAC28FDF9FE4FFC8F12FD" box="[625,801,444,470]" name="Argentina" pageId="4" pageNumber="5">ARGENTINA</collectingCountry>
: Salta, Santa Victoria, Parque Nacional 
<taxonomicName id="79FC4D11E16BAC28FB7EFE4FFE8412D5" class="Insecta" family="Leptohyphidae" genus="Haplohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="9" phylum="Arthropoda" rank="species" species="baritu">Baritu</taxonomicName>
, arroyo 
<taxonomicName id="79FC4D11E16BAC28FE02FE17FD9012D5" box="[394,574,484,510]" class="Insecta" family="Leptohyphidae" genus="Haplohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="9" phylum="Arthropoda" rank="species" species="baritu">
Baritu, 
<quantity id="79049B77E16BAC28FE6BFE17FD9012D5" box="[483,574,484,510]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="5" unit="m" value="1500.0">1500 m</quantity>
</taxonomicName>
, 
<date id="CA421052E16BAC28FDC3FE17FD5C12D5" box="[587,754,484,510]" pageId="4" pageNumber="5" value="1981-10-10" valueMax="1981-10-17" valueMin="1981-10-10">10­17/X/1981</date>
, E. Domínguez Col. 
<taxonomicName id="79FC4D11E16BAC28FC64FE17FB7212D6" box="[1004,1244,484,509]" class="Insecta" family="Leptohyphidae" genus="Haplohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="furtiva">
<emphasis id="8C88EA80E16BAC28FC64FE17FB7212D6" box="[1004,1244,484,509]" italics="true" pageId="4" pageNumber="5">Haplohyphes furtiva</emphasis>
</taxonomicName>
: 
<typeStatus id="61478830E16BAC28FB64FE17FE94110D" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
male imago and 15 
<typeStatus id="61478830E16BAC28FDBDFDFFFD34110D" box="[565,666,524,550]" pageId="4" pageNumber="5" type="paratype">paratype</typeStatus>
male imagos from 
<collectingCountry id="C6EB7602E16BAC28FC0FFDFFFB9B110D" box="[903,1077,524,550]" name="Argentina" pageId="4" pageNumber="5">ARGENTINA</collectingCountry>
, Salta, Anta, Parque Nacional El Rey, arroyo Las Salas, 
<date id="CA421052E16BAC28FD46FDC7FCF61165" box="[718,856,564,590]" pageId="4" pageNumber="5" value="1982-04-25">25/IV/1982</date>
, E. Domínguez Col.; allotype female imago and 1 nymph, idem except date, 
<date id="CA421052E16BAC28FD5DFDAFFCC4115D" box="[725,874,604,630]" pageId="4" pageNumber="5" value="1981-12-20">20/XII/1981</date>
. Other material. 
<collectingCountry id="C6EB7602E16BAC28FBA5FDAFFB73115D" box="[1069,1245,604,630]" name="Argentina" pageId="4" pageNumber="5">ARGENTINA</collectingCountry>
: Salta, P. N. El Rey, arroyo Los Puestos, 
<date id="CA421052E16BAC28FD24FD77FCF211B5" box="[684,860,644,670]" pageId="4" pageNumber="5" value="1999-03-24" valueMax="1999-03-25" valueMin="1999-03-24">24­25/III/1999</date>
, C. Molineri Col. (
<specimenCount id="A8FAFD1BE16BAC28FBC0FD77FB0411B5" box="[1096,1194,644,670]" pageId="4" pageNumber="5" type="male">19 male</specimenCount>
imagos, 
<specimenCount id="A8FAFD1BE16BAC28FA93FD77FEF611ED" pageId="4" pageNumber="5" type="female">5 female</specimenCount>
imagos, 2 reared female subimagos, 5 nymphs); Salta, Santa Victoria, Los Toldos, río Huaico Grande, 
<date id="CA421052E16BAC28FD8AFD27FD2C11C5" box="[514,642,724,750]" pageId="4" pageNumber="5" value="1999-10-26">26/X/1999</date>
, C. Molineri Col. (
<specimenCount id="A8FAFD1BE16BAC28FCF1FD27FC6011C5" box="[889,974,724,750]" pageId="4" pageNumber="5" type="male">5 male</specimenCount>
and 
<specimenCount id="A8FAFD1BE16BAC28FB9AFD27FBD311C5" box="[1042,1149,724,750]" pageId="4" pageNumber="5" type="female">4 female</specimenCount>
imagos, eggs extracted from one of these females); Jujuy, río Yala, 
<date id="CA421052E16BAC28FC10FD0FFB8C103D" box="[920,1058,764,790]" pageId="4" pageNumber="5" value="1997-06-26">26/VI/1997</date>
, C. Molineri Col. (25 nymphs); Jujuy, El Carmen, arroyo Las Lanzas, 
<geoCoordinate id="DBC85055E16BAC28FCC2FCD7FC4A1015" box="[842,996,804,830]" direction="south" orientation="latitude" pageId="4" pageNumber="5" precision="15" value="-24.454723">S 24º 27' 17&quot;</geoCoordinate>
­ 
<geoCoordinate id="DBC85055E16BAC28FC65FCD7FB331015" box="[1005,1181,804,830]" direction="west" orientation="longitude" pageId="4" pageNumber="5" precision="15" value="-65.29667">W 65º 17' 48&quot;</geoCoordinate>
, 
<quantity id="79049B77E16BAC28FB22FCD7FAA61015" box="[1194,1288,804,830]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="4" pageNumber="5" unit="m" value="1250.0">1250 m</quantity>
, 
<date id="CA421052E16BAC28FA9DFCD7FEC2104D" pageId="4" pageNumber="5" value="2000-03-03">3/ III/2000</date>
, E. Domínguez &amp; C. Molineri Cols. (
<specimenCount id="A8FAFD1BE16BAC28FCB0FCBFFC37104D" box="[824,921,844,870]" pageId="4" pageNumber="5" type="male">13 male</specimenCount>
and 
<specimenCount id="A8FAFD1BE16BAC28FC5FFCBFFBEE104D" box="[983,1088,844,870]" pageId="4" pageNumber="5" type="female">7 female</specimenCount>
imagos, 1 nymph). 
<collectingCountry id="C6EB7602E16BAC28FE80FC87FE2B10A5" box="[264,389,884,910]" name="Bolivia" pageId="4" pageNumber="5">BOLIVIA</collectingCountry>
: dept. La Paz, stream near Caranavi, 
<geoCoordinate id="DBC85055E16BAC28FCC8FC87FC7710A5" box="[832,985,884,910]" direction="south" orientation="latitude" pageId="4" pageNumber="5" precision="15" value="-15.6075">S 15º 36' 27&quot;</geoCoordinate>
­Wº 67 45' 58&quot;, 
<date id="CA421052E16BAC28FB14FC87FA8810A5" box="[1180,1318,884,910]" pageId="4" pageNumber="5" value="2000-11-27">27/XI/2000</date>
, E. Domínguez, C. Molineri y C. Nieto Cols. (24 nymphs); dept. La Paz, Reserva Carrasco, 
<geoCoordinate id="DBC85055E16BAC28FE80FC37FE0B10F5" box="[264,421,964,990]" direction="south" orientation="latitude" pageId="4" pageNumber="5" precision="15" value="-15.719166">S 15º 43' 09&quot;</geoCoordinate>
­ 
<geoCoordinate id="DBC85055E16BAC28FE25FC37FDF210F5" box="[429,604,964,990]" direction="west" orientation="longitude" pageId="4" pageNumber="5" precision="15" value="-67.51833">W 67º 31' 06&quot;</geoCoordinate>
, 
<quantity id="79049B77E16BAC28FDE2FC37FD1710F5" box="[618,697,964,990]" metricMagnitude="2" metricUnit="m" metricValue="9.4" pageId="4" pageNumber="5" unit="m" value="940.0">940 m</quantity>
, 
<date id="CA421052E16BAC28FD4FFC37FCFF10F5" box="[711,849,964,990]" pageId="4" pageNumber="5" value="2000-11-29">29/XI/2000</date>
, E. Domínguez, C. Molineri y C. Nieto Cols. (
<specimenCount id="A8FAFD1BE16BAC28FED2FC1FFE00172D" box="[346,430,1004,1030]" pageId="4" pageNumber="5" type="male">6 male</specimenCount>
and 
<specimenCount id="A8FAFD1BE16BAC28FE78FC1FFDF4172D" box="[496,602,1004,1030]" pageId="4" pageNumber="5" type="female">3 female</specimenCount>
imagos, 14 nymphs). All the material is housed in 
<collectionCode id="D8EDAE57E16BAC28FB55FC1FFA88172D" box="[1245,1318,1004,1030]" pageId="4" pageNumber="5">IFML</collectionCode>
, except 15 nymphs, 
<specimenCount id="A8FAFD1BE16BAC28FE64FBE7FD951705" box="[492,571,1044,1070]" pageId="4" pageNumber="5" type="male">3 male</specimenCount>
and 
<specimenCount id="A8FAFD1BE16BAC28FDFCFBE7FD741705" box="[628,730,1044,1070]" pageId="4" pageNumber="5" type="female">1 female</specimenCount>
imagos in 
<collectionCode id="D8EDAE57E16BAC28FCD3FBE7FC1A1705" box="[859,948,1044,1070]" httpUri="http://grbio.org/cool/jxh3-1jar" name="Instituto de Ecologia" pageId="4" pageNumber="5">UMSA</collectionCode>
.
</paragraph>
<paragraph id="BE433692E16BAC28FEB0FBCFFAAE166D" blockId="4.[264,1325,284,1870]" pageId="4" pageNumber="5">
Discussion. 
<bibRefCitation id="DA6D4B63E16BAC28FE45FBCFFD1E177D" author="Dominguez" box="[461,688,1084,1110]" pageId="4" pageNumber="11" refString="Dominguez, E. (1984) Dos especies nuevas del genero Haplohyphes Allen (Ephemeroptera: Tricorythidae) de la Argentina. Revista de la Sociedad Entomologica Argentina, 43 (1 - 4), 103 - 112." type="journal article" year="1984">Domínguez (1984)</bibRefCitation>
described 
<taxonomicName id="79FC4D11E16BAC28FCB3FBCFFB85177E" box="[827,1067,1084,1109]" class="Insecta" family="Leptohyphidae" genus="Haplohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="baritu">
<emphasis id="8C88EA80E16BAC28FCB3FBCFFB85177E" box="[827,1067,1084,1109]" italics="true" pageId="4" pageNumber="5">Haplohyphes baritu</emphasis>
</taxonomicName>
from adults of both sexes and nymphs. Adults were described as imagos but all the 
<typeStatus id="61478830E16BAC28FC62FB97FBB31755" box="[1002,1053,1124,1150]" pageId="4" pageNumber="5">type</typeStatus>
material is represented by subimagos. The characters used to distinguish this species from 
<taxonomicName id="79FC4D11E16BAC28FBB0FB7FFA85178E" box="[1080,1323,1164,1189]" class="Insecta" family="Leptohyphidae" genus="Haplohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="furtiva">
<emphasis id="8C88EA80E16BAC28FBB0FB7FFA85178E" box="[1080,1323,1164,1189]" italics="true" pageId="4" pageNumber="5">Haplohyphes furtiva</emphasis>
</taxonomicName>
are attributable to stage differences (dissimilar fore tarsal claws and division of penes). New material from all the stages collected at or near the 
<typeStatus id="61478830E16BAC28FC4BFB2FFC5B17DD" box="[963,1013,1244,1270]" pageId="4" pageNumber="5">type</typeStatus>
localities of both species show them to be synonyms. 
<taxonomicName id="79FC4D11E16BAC28FDD0FAF7FCE61636" box="[600,840,1284,1309]" class="Insecta" family="Leptohyphidae" genus="Haplohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="furtiva">
<emphasis id="8C88EA80E16BAC28FDD0FAF7FCE61636" box="[600,840,1284,1309]" italics="true" pageId="4" pageNumber="5">Haplohyphes furtiva</emphasis>
</taxonomicName>
is considered junior synonym of 
<taxonomicName id="79FC4D11E16BAC28FB5CFAF7FE88166E" class="Insecta" family="Leptohyphidae" genus="Haplohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="baritu">
<emphasis id="8C88EA80E16BAC28FB5CFAF7FE88166E" italics="true" pageId="4" pageNumber="5">H. baritu</emphasis>
</taxonomicName>
due to page priority. 
<taxonomicName id="79FC4D11E16BAC28FDA9FADFFCA4166E" box="[545,778,1324,1349]" class="Insecta" family="Leptohyphidae" genus="Haplohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="baritu">
<emphasis id="8C88EA80E16BAC28FDA9FADFFCA4166E" box="[545,778,1324,1349]" italics="true" pageId="4" pageNumber="5">Haplohyphes baritu</emphasis>
</taxonomicName>
is recorded from 
<collectingCountry id="C6EB7602E16BAC28FC53FADFFB9F166D" box="[987,1073,1324,1350]" name="Bolivia" pageId="4" pageNumber="5">Bolivia</collectingCountry>
for the first time.
</paragraph>
<paragraph id="BE433692E16BAC28FEB0FAA7FBAE15AD" blockId="4.[264,1325,284,1870]" pageId="4" pageNumber="5">
The eggs of 
<taxonomicName id="79FC4D11E16BAC28FE43FAA7FD181646" box="[459,694,1364,1389]" class="Insecta" family="Leptohyphidae" genus="Haplohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="baritu">
<emphasis id="8C88EA80E16BAC28FE43FAA7FD181646" box="[459,694,1364,1389]" italics="true" pageId="4" pageNumber="5">Haplohyphes baritu</emphasis>
</taxonomicName>
show similarities with those known for some species of 
<taxonomicName id="79FC4D11E16BAC28FEA4FA8FFD8116BD" authority="Ulmer" authorityName="Ulmer" box="[300,559,1404,1430]" class="Insecta" family="Leptohyphidae" genus="Tricorythodes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="8C88EA80E16BAC28FEA4FA8FFE6116BE" box="[300,463,1404,1429]" italics="true" pageId="4" pageNumber="5">Tricorythodes</emphasis>
Ulmer
</taxonomicName>
(
<taxonomicName id="79FC4D11E16BAC28FDCCFA8FFC1C16BD" authority="Hofmann &amp; Sartori" authorityName="Hofmann &amp; Sartori" box="[580,946,1404,1430]" class="Insecta" family="Leptohyphidae" genus="Tricorythodes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="griseus">
<emphasis id="8C88EA80E16BAC28FDCCFA8FFD1916BE" box="[580,695,1404,1429]" italics="true" pageId="4" pageNumber="5">T. griseus</emphasis>
Hofmann &amp; Sartori
</taxonomicName>
, 
<taxonomicName id="79FC4D11E16BAC28FC49FA8FFB1716BD" authority="Molineri" authorityName="Molineri" box="[961,1209,1404,1430]" class="Insecta" family="Leptohyphidae" genus="Tricorythodes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="hiemalis">
<emphasis id="8C88EA80E16BAC28FC49FA8FFBEC16BE" box="[961,1090,1404,1429]" italics="true" pageId="4" pageNumber="5">T. hiemalis</emphasis>
Molineri
</taxonomicName>
, unpubl), showing similar chorionic sculpture and adhesive filaments. Other genera of 
<taxonomicName id="79FC4D11E16BAC28FB35FA57FEF816CD" class="Insecta" family="Leptohyphidae" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Leptohyphidae</taxonomicName>
(
<taxonomicName id="79FC4D11E16BAC28FEE1FA3FFD4316CD" authority="Hofmann &amp; Sartori" authorityName="Hofmann &amp; Sartori" box="[361,749,1484,1510]" class="Insecta" family="Leptohyphidae" genus="Allenhyphes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="8C88EA80E16BAC28FEE1FA3FFE5616CE" box="[361,504,1484,1509]" italics="true" pageId="4" pageNumber="5">Allenhyphes</emphasis>
Hofmann &amp; Sartori
</taxonomicName>
, in 
<bibRefCitation id="DA6D4B63E16BAC28FC93FA3FFBA016CD" author="Hofmann" box="[795,1038,1484,1510]" pageId="4" pageNumber="11" refString="Hofmann, C.; Sartori, M. &amp; Thomas, A. (1999) Les Ephemeropteres (Ephemeroptera) de la Guadeloupe (petites Antilles francaises). Memoires de la Societe Vaudoise des Sciences Naturelles, 20 (1), 1 - 96." type="journal article" year="1999">Hofmann et al 1999</bibRefCitation>
; 
<taxonomicName id="79FC4D11E16BAC28FB94FA3FFEE81525" authority="Molineri 2001" authorityName="Molineri" authorityYear="2001" class="Insecta" family="Leptohyphidae" genus="Traverhyphes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="8C88EA80E16BAC28FB94FA3FFB1516CE" box="[1052,1211,1484,1509]" italics="true" pageId="4" pageNumber="5">Traverhyphes</emphasis>
<bibRefCitation id="DA6D4B63E16BAC28FB4CFA3FFEE81525" author="Molineri" pageId="4" pageNumber="11" refString="Molineri, C. (2001) Traverhyphes: a new genus of Leptohyphidae for Leptohyphes indicator and related species (Insecta: Ephemeroptera). Spixiana, 24 (2), 129 - 140." type="journal article" year="2001">Molineri 2001</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="79FC4D11E16BAC28FEDCFA07FD991525" authority="Eaton" authorityName="Eaton" box="[340,567,1524,1550]" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="8C88EA80E16BAC28FEDCFA07FE461526" box="[340,488,1524,1549]" italics="true" pageId="4" pageNumber="5">Leptohyphes</emphasis>
Eaton
</taxonomicName>
, in 
<bibRefCitation id="DA6D4B63E16BAC28FDECFA07FCA11525" author="Molineri" box="[612,783,1524,1550]" pageId="4" pageNumber="11" refString="Molineri, C. (2003) Revision of South American species of Leptohyphes (Ephemeroptera: Leptohyphidae) with a key to the nymphs. Studies on Neotropical Fauna and Environment, 38 (1), 47 - 70. &quot;" type="journal article" year="2003">Molineri 2003</bibRefCitation>
) show adhesive filaments much shorter, widened at the apex. This suggest a closer relationship between 
<taxonomicName id="79FC4D11E16BAC28FC55F9EFFBD6151E" box="[989,1144,1564,1589]" class="Insecta" family="Leptohyphidae" genus="Haplohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="8C88EA80E16BAC28FC55F9EFFBD6151E" box="[989,1144,1564,1589]" italics="true" pageId="4" pageNumber="5">Haplohyphes</emphasis>
</taxonomicName>
and 
<taxonomicName id="79FC4D11E16BAC28FB3CF9EFFE931576" class="Insecta" family="Leptohyphidae" genus="Tricorythodes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="8C88EA80E16BAC28FB3CF9EFFE931576" italics="true" pageId="4" pageNumber="5">Tricorythodes</emphasis>
</taxonomicName>
than previously accepted (
<bibRefCitation id="DA6D4B63E16BAC28FDFDF9B7FC701575" author="Wiersema" box="[629,990,1604,1630]" pageId="4" pageNumber="11" refString="Wiersema, N. A. &amp; McCafferty, W. P. (2000) Generic Revision of the North and Central American Leptohyphidae (Ephemeroptera: Pannota). Transactions of the American Entomological Society, 126 (3 - 4), 337 - 371." type="journal article" year="2000">Wiersema &amp; McCafferty 2000</bibRefCitation>
), situation supported also by other morphological characters in gills and legs (
<bibRefCitation id="DA6D4B63E16BAC28FCCCF99FFC5A15AD" author="Molineri" box="[836,1012,1644,1670]" pageId="4" pageNumber="11" refString="Molineri, C. (2002) Cladistic analysis of the South American species of Tricorythodes (Ephemeroptera: Leptohyphidae) with the description of new species and stages. Aquatic Insects, 24 (4), 273 - 308." type="journal article" year="2002">Molineri 2002</bibRefCitation>
).
</paragraph>
<paragraph id="BE433692E16BAC29FEB0F967FB4B12AD" blockId="4.[264,1325,284,1870]" lastBlockId="5.[264,1323,284,390]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
<taxonomicName id="79FC4D11E16BAC28FEB0F967FD8A1586" box="[312,548,1684,1709]" class="Insecta" family="Leptohyphidae" genus="Haplohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="baritu">
<emphasis id="8C88EA80E16BAC28FEB0F967FD8A1586" box="[312,548,1684,1709]" italics="true" pageId="4" pageNumber="5">Haplohyphes baritu</emphasis>
</taxonomicName>
can be distinguished from the other species of the genus by the following combination of characters: 1) three longitudinal veins present on hind wings (
<figureCitation id="26C72A17E16BAC28FE99F917FEDC15D5" box="[273,370,1764,1790]" captionStart="FIGURES 1 ­ 19" captionStartId="2.[264,382,1681,1705]" captionTargetBox="[273,1303,333,1649]" captionTargetId="figure@2.[273,1314,321,1654]" captionTargetPageId="2" captionText="FIGURES 1 ­ 19. Haplohyphes aquilonius Lugo­Ortiz and McCafferty: 1 a, male fore wing; 1 b, male hind wing; 1 c, idem, detail; 2 a, female fore wing; 2 b, female hind wing; 2 c, idem, detail; 3, nymphal head, d. v.; 4, male genitalia, v. v. H. dominguezi sp. nov.: 5 a, male fore wing; 5 b, male hind wing; 5 c, idem, detail; 6, male genitalia, v. v. H. huallaga Allen: 7, male hind wing, detail. H. mithras (Traver): 8, male hind wing, detail. H. baritu Domínguez: 9 a, female fore wing; 9 b, female hind wing; 9 c, idem, detail; 10 a, male fore wing; 10 b, male hind wing; 10 c, idem, detail; 11, male genitalia, v. v.; 12, detail of forceps, l. v.; 13, nymphal head, d. v.; 14, gill II, d. v.; 15 ­ 19, gills II­VI, v. v." httpUri="https://zenodo.org/record/156637/files/figure.png" pageId="4" pageNumber="5">Figs. 10</figureCitation>
b­c); 2) segment 1 of forceps as long as or longer than segment 2 (
<figureCitation id="26C72A17E16BAC28FB28F917FB5715D5" box="[1184,1273,1764,1790]" captionStart="FIGURES 1 ­ 19" captionStartId="2.[264,382,1681,1705]" captionTargetBox="[273,1303,333,1649]" captionTargetId="figure@2.[273,1314,321,1654]" captionTargetPageId="2" captionText="FIGURES 1 ­ 19. Haplohyphes aquilonius Lugo­Ortiz and McCafferty: 1 a, male fore wing; 1 b, male hind wing; 1 c, idem, detail; 2 a, female fore wing; 2 b, female hind wing; 2 c, idem, detail; 3, nymphal head, d. v.; 4, male genitalia, v. v. H. dominguezi sp. nov.: 5 a, male fore wing; 5 b, male hind wing; 5 c, idem, detail; 6, male genitalia, v. v. H. huallaga Allen: 7, male hind wing, detail. H. mithras (Traver): 8, male hind wing, detail. H. baritu Domínguez: 9 a, female fore wing; 9 b, female hind wing; 9 c, idem, detail; 10 a, male fore wing; 10 b, male hind wing; 10 c, idem, detail; 11, male genitalia, v. v.; 12, detail of forceps, l. v.; 13, nymphal head, d. v.; 14, gill II, d. v.; 15 ­ 19, gills II­VI, v. v." httpUri="https://zenodo.org/record/156637/files/figure.png" pageId="4" pageNumber="5">Fig. 12</figureCitation>
); 3) penes divided on apical 0.23­0.26 (
<figureCitation id="26C72A17E16BAC28FD2BF8FFFD56140D" box="[675,760,1804,1830]" captionStart="FIGURES 1 ­ 19" captionStartId="2.[264,382,1681,1705]" captionTargetBox="[273,1303,333,1649]" captionTargetId="figure@2.[273,1314,321,1654]" captionTargetPageId="2" captionText="FIGURES 1 ­ 19. Haplohyphes aquilonius Lugo­Ortiz and McCafferty: 1 a, male fore wing; 1 b, male hind wing; 1 c, idem, detail; 2 a, female fore wing; 2 b, female hind wing; 2 c, idem, detail; 3, nymphal head, d. v.; 4, male genitalia, v. v. H. dominguezi sp. nov.: 5 a, male fore wing; 5 b, male hind wing; 5 c, idem, detail; 6, male genitalia, v. v. H. huallaga Allen: 7, male hind wing, detail. H. mithras (Traver): 8, male hind wing, detail. H. baritu Domínguez: 9 a, female fore wing; 9 b, female hind wing; 9 c, idem, detail; 10 a, male fore wing; 10 b, male hind wing; 10 c, idem, detail; 11, male genitalia, v. v.; 12, detail of forceps, l. v.; 13, nymphal head, d. v.; 14, gill II, d. v.; 15 ­ 19, gills II­VI, v. v." httpUri="https://zenodo.org/record/156637/files/figure.png" pageId="4" pageNumber="5">Fig. 11</figureCitation>
); 4) membranous lobes of penes obliquely truncated at apex (
<figureCitation id="26C72A17E16BAC28FE3CF8C7FDA51465" box="[436,523,1844,1870]" captionStart="FIGURES 1 ­ 19" captionStartId="2.[264,382,1681,1705]" captionTargetBox="[273,1303,333,1649]" captionTargetId="figure@2.[273,1314,321,1654]" captionTargetPageId="2" captionText="FIGURES 1 ­ 19. Haplohyphes aquilonius Lugo­Ortiz and McCafferty: 1 a, male fore wing; 1 b, male hind wing; 1 c, idem, detail; 2 a, female fore wing; 2 b, female hind wing; 2 c, idem, detail; 3, nymphal head, d. v.; 4, male genitalia, v. v. H. dominguezi sp. nov.: 5 a, male fore wing; 5 b, male hind wing; 5 c, idem, detail; 6, male genitalia, v. v. H. huallaga Allen: 7, male hind wing, detail. H. mithras (Traver): 8, male hind wing, detail. H. baritu Domínguez: 9 a, female fore wing; 9 b, female hind wing; 9 c, idem, detail; 10 a, male fore wing; 10 b, male hind wing; 10 c, idem, detail; 11, male genitalia, v. v.; 12, detail of forceps, l. v.; 13, nymphal head, d. v.; 14, gill II, d. v.; 15 ­ 19, gills II­VI, v. v." httpUri="https://zenodo.org/record/156637/files/figure.png" pageId="4" pageNumber="5">Fig. 11</figureCitation>
); 5) apical spines of penes shorter than membranous lobes; 6) male fore wing length, 5.0­ 
<quantity id="79049B77E16AAC29FD8FFEEFFDC9121D" box="[519,615,284,310]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="5" pageNumber="6" unit="mm" value="7.2">7.2 mm</quantity>
; 7) abdomen uniformly shaded with gray. In the nymphs: 1) fore wingpads shaded with black on basal half of costal margin; 2) head shaded with gray as in 
<figureCitation id="26C72A17E16AAC29FECEFE9FFE3212AD" box="[326,412,364,390]" captionStart="FIGURES 1 ­ 19" captionStartId="2.[264,382,1681,1705]" captionTargetBox="[273,1303,333,1649]" captionTargetId="figure@2.[273,1314,321,1654]" captionTargetPageId="2" captionText="FIGURES 1 ­ 19. Haplohyphes aquilonius Lugo­Ortiz and McCafferty: 1 a, male fore wing; 1 b, male hind wing; 1 c, idem, detail; 2 a, female fore wing; 2 b, female hind wing; 2 c, idem, detail; 3, nymphal head, d. v.; 4, male genitalia, v. v. H. dominguezi sp. nov.: 5 a, male fore wing; 5 b, male hind wing; 5 c, idem, detail; 6, male genitalia, v. v. H. huallaga Allen: 7, male hind wing, detail. H. mithras (Traver): 8, male hind wing, detail. H. baritu Domínguez: 9 a, female fore wing; 9 b, female hind wing; 9 c, idem, detail; 10 a, male fore wing; 10 b, male hind wing; 10 c, idem, detail; 11, male genitalia, v. v.; 12, detail of forceps, l. v.; 13, nymphal head, d. v.; 14, gill II, d. v.; 15 ­ 19, gills II­VI, v. v." httpUri="https://zenodo.org/record/156637/files/figure.png" pageId="5" pageNumber="6">Fig. 13</figureCitation>
. Abdominal gills are illustrated for comparative purposes (
<figureCitation id="26C72A17E16AAC29FBC6FE9FFB7612AD" box="[1102,1240,364,390]" captionStart="FIGURES 1 ­ 19" captionStartId="2.[264,382,1681,1705]" captionTargetBox="[273,1303,333,1649]" captionTargetId="figure@2.[273,1314,321,1654]" captionTargetPageId="2" captionText="FIGURES 1 ­ 19. Haplohyphes aquilonius Lugo­Ortiz and McCafferty: 1 a, male fore wing; 1 b, male hind wing; 1 c, idem, detail; 2 a, female fore wing; 2 b, female hind wing; 2 c, idem, detail; 3, nymphal head, d. v.; 4, male genitalia, v. v. H. dominguezi sp. nov.: 5 a, male fore wing; 5 b, male hind wing; 5 c, idem, detail; 6, male genitalia, v. v. H. huallaga Allen: 7, male hind wing, detail. H. mithras (Traver): 8, male hind wing, detail. H. baritu Domínguez: 9 a, female fore wing; 9 b, female hind wing; 9 c, idem, detail; 10 a, male fore wing; 10 b, male hind wing; 10 c, idem, detail; 11, male genitalia, v. v.; 12, detail of forceps, l. v.; 13, nymphal head, d. v.; 14, gill II, d. v.; 15 ­ 19, gills II­VI, v. v." httpUri="https://zenodo.org/record/156637/files/figure.png" pageId="5" pageNumber="6">Figs. 14­19</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>