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<document id="13C9944809B88B6FD8A89EA789DDEFBD" ID-DOI="10.11646/zootaxa.3634.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5262081" ID-ZooBank="81906FA2-FB3C-4F02-9AF9-449BC73F3BEE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629850604360" checkinUser="felipe" docAuthor="Santos, Bernardo F. &amp; Aguiar, Alexandre P." docDate="2013" docId="3629866AFFF92E2EB8BFFD93FB18B205" docLanguage="en" docName="zt03634p284.pdf" docOrigin="Zootaxa 3634 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.3634.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Messatoporus depressifrons Santos &amp; Aguiar 2013, sp. nov." docType="treatment" docVersion="3" lastPageNumber="106" masterDocId="CA10FE12FF9E2E47B828FFFFFF9FB319" masterDocTitle="&lt;strong&gt; Phylogeny and revision of &lt;em&gt; Messatoporus &lt;/ em&gt; Cushman (Hymenoptera, Ichneumonidae, Cryptinae), with descriptions of sixty five new species &lt;/ strong&gt;" masterLastPageNumber="284" masterPageNumber="1" pageNumber="104" updateTime="1699121228478" updateUser="plazi">
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<mods:title id="943892E5E49F4EAACFA49E5EB23292CB">&lt;strong&gt; Phylogeny and revision of &lt;em&gt; Messatoporus &lt;/ em&gt; Cushman (Hymenoptera, Ichneumonidae, Cryptinae), with descriptions of sixty five new species &lt;/ strong&gt;</mods:title>
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<mods:namePart id="94BA8C38C3A1C4293EDD263E4D764670">Santos, Bernardo F.</mods:namePart>
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<mods:namePart id="F0352D7761368934E8521A8AD1FE1F4A">Aguiar, Alexandre P.</mods:namePart>
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<subSubSection id="F69A64F7FFF92E20B8BFFD93FD32B19C" box="[151,685,619,646]" pageId="103" pageNumber="104" type="nomenclature">
<paragraph id="BE3F377CFFF92E20B8BFFD93FD32B19C" blockId="103.[151,685,619,678]" box="[151,685,619,646]" pageId="103" pageNumber="104">
<heading id="E5778010FFF92E20B8BFFD93FD32B19C" bold="true" box="[151,685,619,646]" fontSize="11" level="2" pageId="103" pageNumber="104" reason="4">
<emphasis id="8CF4EB6EFFF92E20B8BFFD93FD32B19C" bold="true" box="[151,685,619,646]" pageId="103" pageNumber="104">
<taxonomicName id="79804CFFFFF92E20B8BFFD93FDDDB19C" authority="Santos" authorityName="Santos &amp; Aguiar" authorityYear="2013" box="[151,578,619,646]" class="Insecta" family="Ichneumonidae" genus="Messatoporus" kingdom="Animalia" order="Hymenoptera" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="depressifrons" status="sp. nov.">
<emphasis id="8CF4EB6EFFF92E20B8BFFD93FE78B19F" bold="true" box="[151,487,620,646]" italics="true" pageId="103" pageNumber="104">Messatoporus depressifrons</emphasis>
Santos
</taxonomicName>
,
<taxonomicNameLabel id="97C75615FFF92E20BA78FD94FD32B19C" box="[592,685,619,645]" pageId="103" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F69A64F7FFF92E2FB8BFFD71FC73B7B9" lastPageId="104" lastPageNumber="105" pageId="103" pageNumber="104" type="description">
<paragraph id="BE3F377CFFF92E20B8BFFD71FE3AB1BF" blockId="103.[151,685,619,678]" box="[151,421,654,678]" pageId="103" pageNumber="104">
(
<figureCitation id="26BB2BF9FFF92E20B8B7FD71FF6AB1BF" box="[159,245,654,678]" captionStart="FIGURES 7175" captionStartId="241.[151,258,1980,2001]" captionTargetBox="[172,1416,185,1967]" captionTargetId="figure-49@241.[172,1416,185,1967]" captionTargetPageId="241" captionText="FIGURES 7175. Lateral view. 71, M. compressicornis*. 72, M. nigriscapus*. 73, M. depressifrons*. 74, M. keraiopetalus. 75, M. arcuatus." figureDoi="http://doi.org/10.5281/zenodo.5262157" httpUri="https://zenodo.org/record/5262157/files/figure.png" pageId="103" pageNumber="104">Figs 73</figureCitation>
,
<figureCitation id="26BB2BF9FFF92E20B929FD71FEB2B1BF" box="[257,301,654,678]" captionStart="FIGURES 236241" captionStartId="268.[151,258,1967,1988]" captionTargetBox="[151,1436,193,1946]" captionTargetId="figure-56@268.[151,1436,193,1946]" captionTargetPageId="268" captionText="FIGURES 236241. Hind leg. 236, M. mesonotator. 237, M. arcuatus. 238, M. keraiopetalus. 239, M. mesosternalis. 240, M. albiaterartus*. 241, M. versicolor*." figureDoi="http://doi.org/10.5281/zenodo.5262211" httpUri="https://zenodo.org/record/5262211/files/figure.png" pageId="103" pageNumber="104">237</figureCitation>
,
<figureCitation id="26BB2BF9FFF92E20B911FD71FEF9B1BF" box="[313,358,654,678]" captionStart="FIGURES 272278. 272273" captionStartId="271.[151,258,1912,1933]" captionTargetBox="[151,1415,179,1897]" captionTargetId="figure-94@271.[151,1415,179,1897]" captionTargetPageId="271" captionText="FIGURES 272278. 272273, Gena and occiput, back view. 272, M. euryoikos. Occipital carina ending far from hypostomal carina. 273, M. lissonotus. Occiptal carina ending near hypostomal carina 274, M. depressifrons*, supraantennal area. 275276, Dorsal view. 275, M. complexifemur. 276, M. amarelus*. 277, M. keraiopetalus. Fore and mid femora with dorsal brown stripes. 278, M. complexifemur, mesopleuron." figureDoi="http://doi.org/10.5281/zenodo.5262217" httpUri="https://zenodo.org/record/5262217/files/figure.png" pageId="103" pageNumber="104">274</figureCitation>
,
<figureCitation id="26BB2BF9FFF92E20B959FD71FE03B1BF" box="[369,412,654,678]" captionStart="FIGURES 313320" captionStartId="276.[151,258,1962,1983]" captionTargetBox="[151,1398,195,1954]" captionTargetId="figure-69@276.[151,1398,193,1954]" captionTargetPageId="276" captionText="FIGURES 313320. Distribution maps for species of Messatoporus. 313, M. compressicornis. 314, M. concavivenus. 315, M. convexus. 316, M. copiosus. 317, M. covarrubiasi. 318, M. depressifrons. 319, M. dialeipsis. 320, M. discoidalis." figureDoi="http://doi.org/10.5281/zenodo.5262227" httpUri="https://zenodo.org/record/5262227/files/figure.png" pageId="103" pageNumber="104">318</figureCitation>
)
</paragraph>
<paragraph id="BE3F377CFFF92E20B8BFFD28FB16B717" blockId="103.[151,1437,726,1974]" pageId="103" pageNumber="104">
<emphasis id="8CF4EB6EFFF92E20B8BFFD28FF6AB1F6" bold="true" box="[151,245,727,751]" pageId="103" pageNumber="104">Female.</emphasis>
Fore wing
<quantity id="79789A99FFF92E20B950FD29FE7FB1F7" box="[376,480,726,750]" metricMagnitude="-3" metricUnit="m" metricValue="8.91" pageId="103" pageNumber="104" unit="mm" value="8.91">8.91 mm</quantity>
. HEAD: Mandible densely covered with long hairs; MLW 2.29; MWW 0.42; dorsal tooth in front view projected upwards, much longer than ventral tooth; ventral tooth triangular. MSM 0.25. Clypeus slightly and more or less uniformly convex, centrally concave; CHW 2.56; CWW 3.95; apical area medially concave, laterally slightly projected, its margin very sharp, raised. Antenna with 27 flagellomeres; flagellum strongly flattened and compressed on apical 0.3; apex of apical flagellomere about as wide as base, not distinctly tapered. Supra-antennal area shiny, with moderately dense punctation, not striate near antennal sockets, ventrally and around ocelli distinctly concave, medially with a distinct depression but without a distinct line. Occipital carina uniformly curved, fading out at distance about as long as basal width of mandible from the hypostomal carina. Temple and gena moderately wide, gena somewhat swollen and projected (as in
<figureCitation id="26BB2BF9FFF92E20BC33FC09FBE2B717" box="[1051,1149,1014,1038]" captionStart="FIGURES 260271. 260261" captionStartId="270.[151,258,1798,1819]" captionTargetBox="[151,1436,193,1777]" captionTargetId="figure-105@270.[151,1436,193,1777]" captionTargetPageId="270" captionText="FIGURES 260271. 260261, Gena, lateral. 260, M. longicaudis*. 261, M. compressicornis. 262264, Antenna. 262, M. tibiator. 263, M. compresscornis. 264, M. antennator, male. 265266, Hind t4. 265, M. paeneater*. 266, M. nigriscutus*. 267271, Metasoma and ovipositor. 267, M. opacus*. 268, M. discoidalis. 269, M. longicaudis*. 270, M. convexus. 271, M. townesi." figureDoi="http://doi.org/10.5281/zenodo.5262215" httpUri="https://zenodo.org/record/5262215/files/figure.png" pageId="103" pageNumber="104">Fig. 261</figureCitation>
).
</paragraph>
<paragraph id="BE3F377CFFF92E20B8EFFBE5FC39B6FB" blockId="103.[151,1437,726,1974]" pageId="103" pageNumber="104">THORAX: Pronotum glabrous, shiny, impunctate, ventrally with strong wrinkles; epomia moderately strong, distinct only after diverging from pronotal collar, short, ending far from dorsal margin of pronotum, after diverging from pronotal collar distinctly curved. Mesoscutum moderately convex, flattened posteriorly, ovoid, 1.37 × as long as wide, sparsely pilose, shiny, minutely punctulate; notaulus reaching about 0.76 of mesoscutum length, deeply impressed, surface over notaulus not wrinkled; scuto-scutellar groove deep, with distinct longitudinal wrinkles. Subalar ridge weakly projected; epicnemial carina reaching about 0.7 of distance to subalar ridge, irregular; sternaulus moderately strong, wider on posterior apex, strongly sinuous, smooth; scrobe moderately deep; mesopleural suture ventrally with strong longitudinal wrinkles, wrinkles dorsally weak. Mesosternum medially with short transverse wrinkles; median portion of posterior transverse carina of the mesothoracic venter short, arched backwards. Transverse sulcus at base of propodeum moderately wide, about 0.59 × as long as anterior area of propodeum, impunctate; metapleuron moderately punctate, without transverse wrinkles, moderately pilose; juxtacoxal carina absent. Fore tibia distinctly swollen. Mesal lobe of t4 with a distinct cluster of stout bristles. Hind coxa with weak punctures separated by more than their own diameter.</paragraph>
<paragraph id="BE3F377CFFF92E20B8EFFA11FCD1B56B" blockId="103.[151,1437,726,1974]" pageId="103" pageNumber="104">PROPODEUM: 1.04 × as long as wide, shiny, moderately pilose; anterior area scarcely punctate; spiracle elongate, SWL 6.25; anterior transverse carina medially straight. Propodeal wrinkles strong, closely spaced, anteriorly faint at median portion, posteriorly partly complete and partly incomplete, mostly slightly curved or somewhat sinuous; posterior transverse carina entirely absent.</paragraph>
<paragraph id="BE3F377CFFF92E20B8EFF981FD4BB4AF" blockId="103.[151,1437,726,1974]" pageId="103" pageNumber="104">WINGS: Fore wing vein 1-Rs+M distinctly sinuous, with bulla placed on basal 0.4; crossvein 1m-cu irregular, limit between 1m-cu and 1-Rs+M more or less traceable; vein 1M+Rs anteriorly straight, posteriorly weakly curved; fore wing crossvein 1 cu-a basad of 1M+Rs by 0.49 of its own length; vein 2Cua 0.32 × as long as crossvein 2cu-a; bulla of crossvein 2m-cu placed mostly on posterior half; cell 1+2Rs small, APH 0.71, pentagonal, about as high as wide, AWH 0.94; crossvein 3r-m fully spectral, 2r-m and 3r-m subparallel, about same length; vein 3-M about as long as 2-M; 4-Rs somewhat irregular, apically slightly upcurved; 4-M tubular, irregular. Hind wing vein 1-M forming distinctly obtuse angle with vein Cua; vein 2-Rs tubular, apically spectral, reaching wing margin; HW1C 1.03; vein Cub distinctly convex on posterior half, forming straight angle with Cua; vein 2-1A reaching 0.81 of distance to posterior wing margin.</paragraph>
<paragraph id="BE3F377CFFF62E2FB8EFFF68FC49B2B5" blockId="104.[151,1436,151,2012]" pageId="104" pageNumber="105">METASOMA: First tergite moderately long, about 0.45 × as long as T28, strongly depressed, ventrolaterally rounded, dorsally glabrate, laterally moderately pilose; T1LW 3.3; T1WW 1.74; spiracle on 0.55 of its length, distinctly prominent; median depression elongate, very shallow; lateral and median posterior depressions absent; ventrolateral carina absent, but traceable as limit between lateral and ventral portions. T2LW 1.29; T2WW 1.8; thyridium longer than wide, with small circular depression just behind it; T28 covered with small, shallow foveae, densely and uniformly pilose. OST 1.24; ovipositor moderately stout, straight, basally cylindric, apically slightly depressed; dorsal valve with four moderately strong ridges; ventral valve apex with 8 teeth, apical teeth progressively more closely spaced; surface anterior to first tooth rugulose.</paragraph>
<paragraph id="BE3F377CFFF62E2FB8EFFE47FBF2B716" blockId="104.[151,1436,151,2012]" pageId="104" pageNumber="105">
<emphasis id="8CF4EB6EFFF62E2FB8EFFE47FE98B2C9" box="[199,263,440,464]" italics="true" pageId="104" pageNumber="105">Color</emphasis>
. Head, mesosoma and metasoma black and whitish. Head: black; clypeus, basal 0.4 of mandible, mouthparts, supra-clypeal area, ventral spot at scape and complete orbital band, light yellow (226,208,098); f58 entirely white, f4 partially white, f9 almost entirely white and f1014 with dorsal white longitudinal band. Mesosoma: black; very narrow mark at collar, posterior 0.5 of dorsal margin of pronotum, propleuron, subcircular spot on mesoscutum, scutellar carina, scutellum, axillar carinae, postscutellum, tegula, subalar ridge, central mark on mesopleuron, mesosternum, dorsal division of metapleuron, metapleuron except posterior apex and arc-shaped, quadratic mark on propodeum, light yellow; fore and mid coxae and first trochanters whitish, coxae with apical dark brown marks, mid one also with posterior face dark brown; first trochanters with basal dark brown mark; fore and mid second trochanter, femur, tibia and tarsus bright yellow (222,153,049), with strong dorsal brown stripes on femora, fore one with weak ventral stripe; tibiae with ventral brown marks, fore one on basal 0.8, mid one on basal 0.3; mid tarsus fuscous, fore and mid t5 blackish; hind coxa whitish with three longitudinal stripes, anterior one reaching 0.5 of its length, lateral and posterior ones reaching coxa apex; hind first trochanter whitish with basal blackish mark; second trochanter blackish, with apical 0.2 dark yellow; hind femur dorsal and anterior faces bright yellow, ventral face blackish, posterior face blackish on basal 0.1 and apical 0.2, medially bright yellow; hind tibia bright yellow, hind tarsus basally bright yellow, whitish towards apex. Metasoma: black; T1 laterally with anterior and posterior 0.2 whitish, dorsally anterior 0.5 whitish; T27 with posterior and lateral whitish stripes, T8 only with lateral stripes; S16 whitish, S25 with progressively smaller brown lateral marks.
</paragraph>
<paragraph id="BE3F377CFFF62E2FB8EFFBE3FE17B72D" blockId="104.[151,1436,151,2012]" box="[199,392,1052,1076]" pageId="104" pageNumber="105">
<emphasis id="8CF4EB6EFFF62E2FB8EFFBE3FE9BB72D" bold="true" box="[199,260,1052,1076]" pageId="104" pageNumber="105">Male</emphasis>
. Unknown.
</paragraph>
<paragraph id="BE3F377CFFF62E2FB8EFFBC0FC73B7B9" blockId="104.[151,1436,151,2012]" pageId="104" pageNumber="105">
<emphasis id="8CF4EB6EFFF62E2FB8EFFBC0FEA8B741" bold="true" box="[199,311,1087,1112]" pageId="104" pageNumber="105">Variation</emphasis>
. Fore wing
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long. Arc-shaped mark on propodeum anteriorly either rounded or somewhat quadratic; hind tibia sometimes with very small blackish apical mark. Specimen from
<collectingRegion id="7C44F99EFFF62E2FBCD4FB9BFAC0B765" box="[1276,1375,1124,1148]" country="Argentina" name="La Rioja" pageId="104" pageNumber="105">La Rioja</collectingRegion>
with anterior propodeal wrinkles weak and median depression on T1 very strong.
</paragraph>
</subSubSection>
<subSubSection id="F69A64F7FFF62E2FB8EFFB54FAAAB41D" pageId="104" pageNumber="105" type="discussion">
<paragraph id="BE3F377CFFF62E2FB8EFFB54FD38B6D9" blockId="104.[151,1436,151,2012]" pageId="104" pageNumber="105">
<emphasis id="8CF4EB6EFFF62E2FB8EFFB54FED7B7DD" bold="true" box="[199,328,1195,1220]" pageId="104" pageNumber="105">Comments</emphasis>
.
<taxonomicName id="79804CFFFFF62E2FB97EFB52FD10B7DD" authorityName="Santos &amp; Aguiar" authorityYear="2013" box="[342,655,1196,1220]" class="Insecta" family="Ichneumonidae" genus="Messatoporus" kingdom="Animalia" order="Hymenoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="depressifrons">
<emphasis id="8CF4EB6EFFF62E2FB97EFB52FD10B7DD" box="[342,655,1196,1220]" italics="true" pageId="104" pageNumber="105">Messatoporus depressifrons</emphasis>
</taxonomicName>
is part of a group of four very close species, which apparently form a monophyletic group (
<figureCitation id="26BB2BF9FFF62E2FB9B7FB2FFDB3B7F1" box="[415,556,1232,1256]" captionStart="FIGURES 7175" captionStartId="241.[151,258,1980,2001]" captionTargetBox="[172,1416,185,1967]" captionTargetId="figure-49@241.[172,1416,185,1967]" captionTargetPageId="241" captionText="FIGURES 7175. Lateral view. 71, M. compressicornis*. 72, M. nigriscapus*. 73, M. depressifrons*. 74, M. keraiopetalus. 75, M. arcuatus." figureDoi="http://doi.org/10.5281/zenodo.5262157" httpUri="https://zenodo.org/record/5262157/files/figure.png" pageId="104" pageNumber="105">Figs 7175</figureCitation>
; see Comments under
<taxonomicName id="79804CFFFFF62E2FBB7FFB2EFBA1B7FE" authorityName=": Kasparyan &amp; Ruiz" authorityYear="2008" box="[855,1086,1232,1256]" class="Insecta" family="Ichneumonidae" genus="Messatoporus" kingdom="Animalia" order="Hymenoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="compressicornis">
<emphasis id="8CF4EB6EFFF62E2FBB7FFB2EFBA1B7FE" box="[855,1086,1232,1256]" italics="true" pageId="104" pageNumber="105">M. compressicornis</emphasis>
</taxonomicName>
). Among these species,
<taxonomicName id="79804CFFFFF62E2FBD55FB2EFEB3B615" authorityName="Santos &amp; Aguiar" authorityYear="2013" class="Insecta" family="Ichneumonidae" genus="Messatoporus" kingdom="Animalia" order="Hymenoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="depressifrons">
<emphasis id="8CF4EB6EFFF62E2FBD55FB2EFA0BB7F1" box="[1405,1428,1233,1256]" italics="true" pageId="104" pageNumber="105">M</emphasis>
.
<emphasis id="8CF4EB6EFFF62E2FB8BFFB0BFEB3B615" box="[151,300,1268,1292]" italics="true" pageId="104" pageNumber="105">depressifrons</emphasis>
</taxonomicName>
can be differentiated from
<taxonomicName id="79804CFFFFF62E2FBA73FB0AFC87B615" authority="Santos &amp; Aguiar, 2013" authorityName="Santos &amp; Aguiar" authorityYear="2013" box="[603,792,1268,1292]" class="Insecta" family="Ichneumonidae" genus="Messatoporus" kingdom="Animalia" order="Hymenoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="keraiopetalus" status="sp. nov.">
<emphasis id="8CF4EB6EFFF62E2FBA73FB0AFDEDB615" box="[603,626,1269,1292]" italics="true" pageId="104" pageNumber="105">M</emphasis>
.
<emphasis id="8CF4EB6EFFF62E2FBAA8FB0BFC87B615" box="[640,792,1268,1292]" italics="true" pageId="104" pageNumber="105">keraiopetalus</emphasis>
</taxonomicName>
<emphasis id="8CF4EB6EFFF62E2FBB37FB0BFCE6B615" bold="true" box="[799,889,1268,1292]" pageId="104" pageNumber="105">
<taxonomicNameLabel id="97C75615FFF62E2FBB37FB0BFCE6B615" box="[799,889,1268,1292]" pageId="104" pageNumber="105" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by having clypeus very slightly and more or less uniformly convex (vs. basally strongly convex, emarginated in its mid-length, abruptly flattened); supra-antennal area with a median depression (vs. with median smoother area); epomia moderately strong, distinct only after diverging from collar (vs. weak but complete); transverse sulcus of propodeum moderately long, 0.59 × as long as anterior area (vs. very long, 1.07 × as long as anterior area of propodeum); and fore wing vein 3-M and 2-M with about the same size (vs. 3-M distinctly longer).
</paragraph>
<paragraph id="BE3F377CFFF62E2FB8EFFA33FD72B56D" blockId="104.[151,1436,151,2012]" pageId="104" pageNumber="105">
It can be isolated from
<taxonomicName id="79804CFFFFF62E2FB9EEFA32FD3EB6FD" authorityName=": Kasparyan &amp; Ruiz" authorityYear="2008" box="[454,673,1484,1508]" class="Insecta" family="Ichneumonidae" genus="Messatoporus" kingdom="Animalia" order="Hymenoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="compressicornis">
<emphasis id="8CF4EB6EFFF62E2FB9EEFA32FE42B6FD" box="[454,477,1485,1508]" italics="true" pageId="104" pageNumber="105">M</emphasis>
.
<emphasis id="8CF4EB6EFFF62E2FB9C3FA32FD3EB6FD" box="[491,673,1485,1508]" italics="true" pageId="104" pageNumber="105">compressicornis</emphasis>
</taxonomicName>
by the hind femur ventrally black, dorsally deep yellow (vs. entirely orange); fore and mid femora dorsally with strong longitudinal brown stripes (vs. entirely orange); ovipositor dorsal valve with four moderately strong ridges (vs. five); supra-antennal area with median carina indistinct, represented only by a smoother area (vs. with slight median carina); and mesosternum medially with short transverse wrinkles (vs. without transverse wrinkles).
</paragraph>
<paragraph id="BE3F377CFFF62E2FB8EFF97FFAAAB41D" blockId="104.[151,1436,151,2012]" pageId="104" pageNumber="105">
It can be separated from
<taxonomicName id="79804CFFFFF62E2FB9D0F97EFD39B58E" authority="Santos &amp; Aguiar, 2013" authorityName="Santos &amp; Aguiar" authorityYear="2013" box="[504,678,1664,1688]" class="Insecta" family="Ichneumonidae" genus="Messatoporus" kingdom="Animalia" order="Hymenoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="nigriscapus" status="sp. nov.">
<emphasis id="8CF4EB6EFFF62E2FB9D0F97EFD90B581" box="[504,527,1665,1688]" italics="true" pageId="104" pageNumber="105">M</emphasis>
.
<emphasis id="8CF4EB6EFFF62E2FBA0BF97FFD39B58E" box="[547,678,1664,1687]" italics="true" pageId="104" pageNumber="105">nigriscapus</emphasis>
</taxonomicName>
<emphasis id="8CF4EB6EFFF62E2FBA9AF97FFC8EB581" bold="true" box="[690,785,1664,1688]" pageId="104" pageNumber="105">
<taxonomicNameLabel id="97C75615FFF62E2FBA9AF97FFC8EB581" box="[690,785,1664,1688]" pageId="104" pageNumber="105" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the orbital band complete (vs. shortly interrupted); mandible basally yellow (vs. entirely black); collar yellow (vs. black); supra-antennal area with median depression (vs. with slight median carina); median portion of posterior transverse carina of the mesothoracic venter arched backwards (vs. arched forwards); and spiracle of propodeum elongate, SWL 4.86 (vs. elliptic, SWL 3.37)
</paragraph>
</subSubSection>
<subSubSection id="F69A64F7FFF62E2FB8EFF8F0FE3DB455" pageId="104" pageNumber="105" type="biology_ecology">
<paragraph id="BE3F377CFFF62E2FB8EFF8F0FE3DB455" blockId="104.[151,1436,151,2012]" pageId="104" pageNumber="105">
<emphasis id="8CF4EB6EFFF62E2FB8EFF8F0FE80B431" bold="true" box="[199,287,1807,1832]" pageId="104" pageNumber="105">Biology</emphasis>
. One specimen from Moca,
<collectingCountry id="C69777ECFFF62E2FBA53F8EFFD68B431" box="[635,759,1808,1832]" name="Guatemala" pageId="104" pageNumber="105">Guatemala</collectingCountry>
, was reared from a nest of
<taxonomicName id="79804CFFFFF62E2FBC66F8EFFA03B431" authority="Smith" authorityName="Smith" authorityYear="1857" box="[1102,1436,1808,1832]" class="Insecta" family="Eumenidae" genus="Pachymenes" kingdom="Animalia" order="Hymenoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="8CF4EB6EFFF62E2FBC66F8EFFAD3B431" box="[1102,1356,1808,1832]" italics="true" pageId="104" pageNumber="105">Pachymenes obscurus</emphasis>
Smith
</taxonomicName>
(
<taxonomicName id="79804CFFFFF62E2FB8B6F8CBFE97B455" box="[158,264,1844,1868]" class="Insecta" family="Vespidae" kingdom="Animalia" order="Hymenoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="family">Vespidae</taxonomicName>
, Eumeninae).
</paragraph>
</subSubSection>
<subSubSection id="F69A64F7FFF62E2FB8EFF8A7FD8CB48A" pageId="104" pageNumber="105" type="etymology">
<paragraph id="BE3F377CFFF62E2FB8EFF8A7FD8CB48A" blockId="104.[151,1436,151,2012]" pageId="104" pageNumber="105">
<emphasis id="8CF4EB6EFFF62E2FB8EFF8A7FED9B469" bold="true" box="[199,326,1880,1904]" pageId="104" pageNumber="105">Etymology</emphasis>
. From the Latin
<emphasis id="8CF4EB6EFFF62E2FBA34F8A7FD16B469" box="[540,649,1880,1904]" italics="true" pageId="104" pageNumber="105">depressus</emphasis>
, meaning “depressed”, and
<emphasis id="8CF4EB6EFFF62E2FBBF4F8A7FB8AB469" box="[988,1045,1880,1904]" italics="true" pageId="104" pageNumber="105">frons</emphasis>
; in reference to the longitudinal depression at supra-antennal area.
</paragraph>
</subSubSection>
<subSubSection id="F69A64F7FFF62E2EB8EFF860FB18B205" lastPageId="105" lastPageNumber="106" pageId="104" pageNumber="105" type="materials_examined">
<paragraph id="BE3F377CFFF62E2FB8EFF860FB26B4C5" blockId="104.[151,1436,151,2012]" pageId="104" pageNumber="105">
<emphasis id="8CF4EB6EFFF62E2FB8EFF860FE32B4A1" bold="true" box="[199,429,1951,1976]" pageId="104" pageNumber="105">Material examined</emphasis>
.
<materialsCitation id="0EE83D21FFF62E2FB997F85FFB26B4C5" collectingDate="1976-08" collectionCode="AMNH" country="Guatemala" elevation="140" location="Santa Elena" pageId="104" pageNumber="105" specimenCount="5" specimenCount-female="5" stateProvince="Peten" typeStatus="holotype">
<specimenCount id="A886FCF5FFF62E2FB997F85FFE73B4A1" box="[447,492,1952,1976]" pageId="104" pageNumber="105" type="female">4 ♀</specimenCount>
.
<typeStatus id="613B89DEFFF62E2FBA3BF85FFDE4B4A1" box="[531,635,1952,1976]" pageId="104" pageNumber="105" type="holotype">Holotype</typeStatus>
<specimenCount id="A886FCF5FFF62E2FBAAEF85EFD04B4A1" box="[646,667,1953,1976]" pageId="104" pageNumber="105" type="female"></specimenCount>
from
<collectingCountry id="C69777ECFFF62E2FBACFF85FFC07B4A1" box="[743,920,1952,1976]" name="Guatemala" pageId="104" pageNumber="105">GUATEMALA</collectingCountry>
,
<collectingRegion id="7C44F99EFFF62E2FBB8FF85EFC77B4A1" box="[935,1000,1953,1976]" country="Guatemala" name="Peten" pageId="104" pageNumber="105">Peten</collectingRegion>
,
<location id="BB5F61A7FFF62E2FBBDFF85FFB1DB4A1" LSID="urn:lsid:plazi:treatment:3629866AFFF92E2EB8BFFD93FB18B205:BB5F61A7FFF62E2FBBDFF85FFB1DB4A1" box="[1015,1154,1952,1976]" country="Guatemala" name="Santa Elena" pageId="104" pageNumber="105" stateProvince="Peten">Santa Elena</location>
,
<quantity id="79789A99FFF62E2FBCB9F85FFA89B4A1" box="[1169,1302,1952,1976]" metricMagnitude="2" metricUnit="m" metricValue="1.4" metricValueMax="1.6" metricValueMin="1.2" pageId="104" pageNumber="105" unit="m" value="140.0" valueMax="160.0" valueMin="120.0">
<elevation id="35ADD04FFFF62E2FBCB9F85FFA89B4A1" box="[1169,1302,1952,1976]" metricMagnitude="2" metricUnit="m" metricValue="1.4" metricValueMax="1.6" metricValueMin="1.2" pageId="104" pageNumber="105" unit="m" value="140.0" valueMax="160.0" valueMin="120.0">120160 m</elevation>
</quantity>
,
<date id="CA3E11BCFFF62E2FBD0EF85FFA08B4A1" box="[1318,1431,1952,1976]" pageId="104" pageNumber="105" value="1976-08">
<collectingDate id="DA7AE854FFF62E2FBD0EF85FFA08B4A1" box="[1318,1431,1952,1976]" pageId="104" pageNumber="105" value="1976-08">VIII.1976</collectingDate>
</date>
, NLHKrauss (
<collectionCode id="D891AFB9FFF62E2FB905F83BFE16B4C5" box="[301,393,1988,2012]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="104" pageNumber="105">AMNH</collectionCode>
). Pinned. Apical third of right antenna missing; otherwise in good shape.
</materialsCitation>
</paragraph>
<paragraph id="BE3F377CFFF72E2EB8EFFF67FA03B3B6" blockId="105.[151,1436,151,284]" box="[199,1436,151,176]" pageId="105" pageNumber="106">
<materialsCitation id="0EE83D21FFF72E2EB8EFFF67FA03B3B6" box="[199,1436,151,176]" collectingDate="1931-03" collectingDateMax="1931-04-31" collectingDateMin="1931-03" country="Guatemala" location="Guatalon" municipality="Moca" pageId="105" pageNumber="106" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="613B89DEFFF72E2EB8EFFF67FEA7B3A9" box="[199,312,152,176]" pageId="105" pageNumber="106" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="A886FCF5FFF72E2EB96BFF68FEF2B3B6" box="[323,365,151,175]" pageId="105" pageNumber="106" type="female">1 ♀</specimenCount>
from
<collectingCountry id="C69777ECFFF72E2EB99AFF68FDFBB3B6" box="[434,612,151,175]" name="Guatemala" pageId="105" pageNumber="106">GUATEMALA</collectingCountry>
,
<location id="BB5F61A7FFF72E2EBA46FF68FD47B3B6" LSID="urn:lsid:plazi:treatment:3629866AFFF92E2EB8BFFD93FB18B205:BB5F61A7FFF72E2EBA46FF68FD47B3B6" box="[622,728,151,175]" country="Guatemala" municipality="Moca" name="Guatalon" pageId="105" pageNumber="106">Guatalon</location>
,
<collectingMunicipality id="5E5BAD06FFF72E2EBACBFF67FCBAB3B6" box="[739,805,152,175]" pageId="105" pageNumber="106">Moca</collectingMunicipality>
,
<date id="CA3E11BCFFF72E2EBB19FF68FC28B3B6" box="[817,951,151,175]" pageId="105" pageNumber="106" value="1931-03" valueMax="1931-04-31" valueMin="1931-03-00">
<collectingDate id="DA7AE854FFF72E2EBB19FF68FC28B3B6" box="[817,951,151,175]" pageId="105" pageNumber="106" value="1931-03" valueMax="1931-04-31" valueMin="1931-03-00">IIIIV.1931</collectingDate>
</date>
, “SC/ from nest of
<taxonomicName id="79804CFFFFF72E2EBCB8FF67FA17B3A9" authorityName="Smith" authorityYear="1857" box="[1168,1416,152,176]" class="Insecta" family="Eumenidae" genus="Pachymenes" kingdom="Animalia" order="Hymenoptera" pageId="105" pageNumber="106" phylum="Arthropoda" rank="species" species="obscurus">
<emphasis id="8CF4EB6EFFF72E2EBCB8FF67FA17B3A9" box="[1168,1416,152,176]" italics="true" pageId="105" pageNumber="106">Pachymenes obscurus</emphasis>
</taxonomicName>
.”
</materialsCitation>
</paragraph>
<paragraph id="BE3F377CFFF72E2EB8BFFF43FA03B3CD" blockId="105.[151,1436,151,284]" box="[151,1436,188,212]" pageId="105" pageNumber="106">
<materialsCitation id="0EE83D21FFF72E2EB8BFFF43FBC6B3CD" box="[151,1113,188,212]" collectingDate="1957-04-30" collectionCode="USNM" collectorName="Pab, N." country="El Salvador" location="San Salvador" pageId="105" pageNumber="106" specimenCount="1" specimenCount-female="1" stateProvince="San Salvador">
<specimenCount id="A886FCF5FFF72E2EB8BFFF43FF58B3CA" box="[151,199,188,212]" pageId="105" pageNumber="106" type="female">1 ♀</specimenCount>
from
<collectingCountry id="C69777ECFFF72E2EB930FF43FE41B3CD" box="[280,478,188,212]" name="El Salvador" pageId="105" pageNumber="106">EL SALVADOR</collectingCountry>
,
<collectingRegion id="7C44F99EFFF72E2EB9C7FF43FD14B3CD" box="[495,651,188,212]" country="El Salvador" name="San Salvador" pageId="105" pageNumber="106">San Salvador</collectingRegion>
,
<date id="CA3E11BCFFF72E2EBAB5FF43FC84B3CD" box="[669,795,188,212]" pageId="105" pageNumber="106" value="1957-04-30">
<collectingDate id="DA7AE854FFF72E2EBAB5FF43FC84B3CD" box="[669,795,188,212]" pageId="105" pageNumber="106" value="1957-04-30">30.IV.1957</collectingDate>
</date>
,
<collectorName id="137552AAFFF72E2EBB05FF43FC1AB3CD" box="[813,901,188,212]" pageId="105" pageNumber="106">Pab, N.</collectorName>
714-250 (
<collectionCode id="D891AFB9FFF72E2EBBD2FF43FBCEB3CD" box="[1018,1105,188,212]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="105" pageNumber="106" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="0EE83D21FFF72E2EBC41FF43FA07B3CD" box="[1129,1432,188,212]" country="Costa Rica" location="COSTA RICA" pageId="105" pageNumber="106" specimenCount="1" specimenCount-female="1">
<specimenCount id="A886FCF5FFF72E2EBC41FF43FB05B3CA" box="[1129,1178,188,212]" pageId="105" pageNumber="106" type="female">1 ♀</specimenCount>
from
<collectingCountry id="C69777ECFFF72E2EBCC3FF43FA07B3CD" box="[1259,1432,188,212]" name="Costa Rica" pageId="105" pageNumber="106">COSTA RICA</collectingCountry>
</materialsCitation>
,
</paragraph>
<paragraph id="BE3F377CFFF72E2EB8BFFF1FFB18B205" blockId="105.[151,1436,151,284]" pageId="105" pageNumber="106">
<materialsCitation id="0EE83D21FFF72E2EB8BFFF1FFB1DB205" collectingDate="1956-05-26" collectionCode="FSCA" collectorName="Puntarenas &amp; La Rioja" country="Guatemala" location="Guatemala" pageId="105" pageNumber="106" specimenCount="1">
<collectorName id="137552AAFFF72E2EB8BFFF1FFE8AB3EE" box="[151,277,224,247]" pageId="105" pageNumber="106">Puntarenas</collectorName>
,
<collectorName id="137552AAFFF72E2EB908FF1FFE1CB3EE" box="[288,387,223,247]" pageId="105" pageNumber="106">La Rioja</collectorName>
,
<date id="CA3E11BCFFF72E2EB9A7FF20FD9BB3EE" box="[399,516,223,247]" pageId="105" pageNumber="106" value="1956-05-26">
<collectingDate id="DA7AE854FFF72E2EB9A7FF20FD9BB3EE" box="[399,516,223,247]" pageId="105" pageNumber="106" value="1956-05-26">26.V.1956</collectingDate>
</date>
, LCKuitert, traps (
<collectionCode id="D891AFB9FFF72E2EBAFEFF20FCBDB3EE" box="[726,802,223,247]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="105" pageNumber="106" type="Museum">FSCA</collectionCode>
).
<emphasis id="8CF4EB6EFFF72E2EB8EFFEFCFEC6B205" bold="true" box="[199,345,259,284]" pageId="105" pageNumber="106">Distribution</emphasis>
. Recorded from three localities in
<collectingCountry id="C69777ECFFF72E2EBAF7FEFBFCC7B205" box="[735,856,260,284]" name="Guatemala" pageId="105" pageNumber="106">Guatemala</collectingCountry>
and
<collectingCountry id="C69777ECFFF72E2EBBA6FEFBFB97B205" box="[910,1032,260,284]" name="Costa Rica" pageId="105" pageNumber="106">Costa Rica</collectingCountry>
(
<figureCitation id="26BB2BF9FFF72E2EBC30FEFBFBE5B205" box="[1048,1146,260,284]" captionStart="FIGURES 313320" captionStartId="276.[151,258,1962,1983]" captionTargetBox="[151,1398,195,1954]" captionTargetId="figure-69@276.[151,1398,193,1954]" captionTargetPageId="276" captionText="FIGURES 313320. Distribution maps for species of Messatoporus. 313, M. compressicornis. 314, M. concavivenus. 315, M. convexus. 316, M. copiosus. 317, M. covarrubiasi. 318, M. depressifrons. 319, M. dialeipsis. 320, M. discoidalis." figureDoi="http://doi.org/10.5281/zenodo.5262227" httpUri="https://zenodo.org/record/5262227/files/figure.png" pageId="105" pageNumber="106">Fig. 318</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>