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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1131.95038" ID-GBIF-Dataset="46e05d1b-511b-4d8e-8b1e-0c733d66f878" ID-PMC="PMC9836511" ID-Pensoft-Pub="1313-2970-1131-99" ID-Pensoft-UUID="1ACF6E07EE8A54DB9906143C0532304D" ID-PubMed="36761460" ID-ZooBank="EE89E8CF4B764FBC821A79BC51D28D67" ModsDocID="1313-2970-1131-99" checkinTime="1669182088444" checkinUser="pensoft" docAuthor="Xia, Mei-Hua, Tang, Liang &amp; Schillhammer, Harald" docDate="2022" docId="3238285259A7537596B73DE45E4021DD" docLanguage="en" docName="ZooKeys 1131: 99-134" docOrigin="ZooKeys 1131" docPubDate="2022-11-22" docSource="http://dx.doi.org/10.3897/zookeys.1131.95038" docTitle="Thoracostrongylus aduncatus Yang, Zhou &amp; Schillhammer 2011" docType="treatment" docVersion="4" id="1ACF6E07EE8A54DB9906143C0532304D" lastPageNumber="99" masterDocId="1ACF6E07EE8A54DB9906143C0532304D" masterDocTitle="Review of Chinese species of the genus Thoracostrongylus Bernhauer, 1915 (Coleoptera, Staphylinidae, Staphylininae)" masterLastPageNumber="134" masterPageNumber="99" pageNumber="99" updateTime="1732489169057" updateUser="ExternalLinkService">
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<mods:title id="189E8D739F7B4699C7AA808073D261DB">Review of Chinese species of the genus Thoracostrongylus Bernhauer, 1915 (Coleoptera, Staphylinidae, Staphylininae)</mods:title>
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<mods:namePart id="A76EB50CE8B9BC010E81D8B83D3B3BF4">Xia, Mei-Hua</mods:namePart>
<mods:affiliation id="AF7FC88226CE33609F42900B6B7AE075">College of Life Sciences, Shanghai Normal University, 100 Guilin Road, 1</mods:affiliation>
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<mods:namePart id="EABD46325A3CC44D34C2EFE1E2348826">Tang, Liang</mods:namePart>
<mods:affiliation id="24DA4841416F80A666CBC51004C128D4">College of Life Sciences, Shanghai Normal University, 100 Guilin Road, 1</mods:affiliation>
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<mods:namePart id="CCEA3BE4677442CC8D8DF6D2CCA4DA66">Schillhammer, Harald</mods:namePart>
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<paragraph id="46CF6925B234EE813149B385998F3075" pageId="0" pageNumber="99">
<taxonomicName id="2F8FD62116CBD6C2BF0DB66EAB7FC8EE" ID-CoL="56KH4" LSID="32382852-59A7-5375-96B7-3DE45E4021DD" authority="Yang, Zhou &amp; Schillhammer, 2011" authorityName="Yang, Zhou &amp; Schillhammer" authorityYear="2011" class="Insecta" family="Staphylinidae" genus="Thoracostrongylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thoracostrongylus aduncatus" order="Coleoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="aduncatus">Thoracostrongylus aduncatus Yang, Zhou &amp; Schillhammer, 2011</taxonomicName>
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<paragraph id="A0692548549C4FE6ACC0872B83A5A8AE" pageId="0" pageNumber="99">
<figureCitation id="315094755D4195DDCC3FCC4474582BA3" captionStart="Figures 1520" captionStartId="F3" captionText="Figures 15 - 20. Thoracostrongylus aduncatus 15, 16 habitus 17 - 20 aedeagus, ventral (17, 18) and lateral (19, 20) views. Scale bars: 1 mm (15, 16); 0.2 mm (17 - 20)." figureDoi="10.3897/zookeys.1131.95038.figures15-20" httpUri="https://binary.pensoft.net/fig/772739" pageId="0" pageNumber="99">Figs 15-20</figureCitation>
<figureCitation id="2913F45B6457D1F86857883E41645422" captionStart="Figures 2128" captionStartId="F4" captionText="Figures 21 - 28. Thoracostrongylus aduncatus 21 - 24 aedeagus from Baihualing, ventral (21, 22) and lateral (23, 24) views 25 - 28 aedeagus from Baihualing, ventral (25, 26) and lateral (27, 28) views. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1131.95038.figures21-28" httpUri="https://binary.pensoft.net/fig/772740" pageId="0" pageNumber="99">, 21-28</figureCitation>
<figureCitation id="35069C747B7DFFDDE3C202FE277DBB6C" captionStart="Figures 2932" captionStartId="F5" captionText="Figures 29 - 32. Thoracostrongylus aduncatus 29 - 32 aedeagus from Nabanhe, ventral (29, 30) and lateral (31, 32) views. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1131.95038.figures29-32" httpUri="https://binary.pensoft.net/fig/772741" pageId="0" pageNumber="99">, 29-32</figureCitation>
<figureCitation id="B7B19D35FB2451FABEDC47C9B1483000" captionStart="Figures 108122" captionStartId="F18" captionText="Figures 108 - 122. Distribution map of Thoracostrongylus species of China 108 T. acerosus 109 T. aduncatus 110 T. birmanus 111 T. diaoluoensis 112 T. formosanus formosanus 113 T. formosanus flavipes 114 T. fujianensis 115 T. malaisei 116 T. miyakei 117 T. sarawakensis 118 T. velutinus 119 T. baishanzuensis 120 T. bicolor 121 T. brachypterus 122 T. chrysites. Triangle, localities of specimens examined in this paper; square, localities of specimens listed in previous papers. Red, doubtful localities; blue, trustede localities." figureDoi="10.3897/zookeys.1131.95038.figures108-122" httpUri="https://binary.pensoft.net/fig/772754" pageId="0" pageNumber="99">
, 109
<normalizedToken id="0B636D5DE47221C53DA4A583DE6F0532" originalValue="钩茎钝胸隐翅虫">钩茎钝胸隐翅虫</normalizedToken>
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<subSubSection id="E11EB0B8BAFE086D17542A0F7BB69646" pageId="0" pageNumber="99" type="reference_group">
<paragraph id="7ECE13FFEE0B6C48350DBE86154A401C" pageId="0" pageNumber="99">
<taxonomicName id="D440F41970738833A7869C6A27B47D8B" authorityName="Yang, Zhou &amp; Schillhammer" authorityYear="2011" class="Insecta" family="Staphylinidae" genus="Thoracostrongylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thoracostrongylus aduncatus" order="Coleoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="aduncatus">Thoracostrongylus aduncatus</taxonomicName>
Yang, Zhou &amp; Schillhammer, 2011: 413.
</paragraph>
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<subSubSection id="09A4579894DE4FE7FE3C49D0AE22E57D" pageId="0" pageNumber="99" type="materials_examined">
<paragraph id="11B82FA693AAA1F0BF31EC522CA35FDB" pageId="0" pageNumber="99">Material examined.</paragraph>
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<materialsCitation id="0B894C2D403449C620F8BA48B5F46285" collectingDate="2018-04-03" collectorName="Peng, Shen, Cheng" country="China" county="Xishuangbanna" elevation="1700" latitude="20.511389" location="Mengsong" longLatPrecision="21" longitude="100.50528" municipality="Menglong Town" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan Prov.">
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,
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&amp;
<collectorName id="B107E6A78162120BF7228470E1CE311F">Cheng</collectorName>
leg.; SHNU
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<materialsCitation id="1C64FD72384A48037E95366CD7A4430A" collectingDate="2009-04-28" collectorName="Nabanhe N. R., Hu, Yin" country="China" elevation="1750" location="Bengganghani" municipality="Chuguohe" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan Prov.">
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<location id="80067DFA59778D745E96077D8809795D" LSID="urn:lsid:plazi:treatment:3238285259A7537596B73DE45E4021DD:80067DFA59778D745E96077D8809795D" country="China" municipality="Chuguohe" name="Bengganghani" stateProvince="Yunnan Prov.">Bengganghani</location>
; alt.
<quantity id="B8EADE3A7A13F158D2C28E7F441219B6" metricMagnitude="3" metricUnit="m" metricValue="1.75" unit="m" value="1750.0">
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&amp;
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leg.; SHNU
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<materialsCitation id="D001BD1BE1C87C121892B1C778AE6747" collectingDate="2009-04-30" collectorName="Nabanhe N. R., Hu, Yin" country="China" elevation="1650" location="Nanmugaha" municipality="Bengganghani" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan Prov.">
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<location id="3E0B9B45E045C6E0D8D4F17FA01D44F6" LSID="urn:lsid:plazi:treatment:3238285259A7537596B73DE45E4021DD:3E0B9B45E045C6E0D8D4F17FA01D44F6" country="China" municipality="Bengganghani" name="Nanmugaha" stateProvince="Yunnan Prov.">Nanmugaha</location>
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<collectorName id="8E5B0B1F611E3CFEF82A4730841356BE">Hu</collectorName>
&amp;
<collectorName id="E5C7B079AFE3E317A2AEF7CEE485DB90">Yin</collectorName>
leg.; SHNU
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<materialsCitation id="B1E9C57A1588C09060EA048A500DEA1E" collectingDate="2009-04-27" collectorName="Nabanhe N. R., Hu, Yin" country="China" elevation="1700" location="Bengganghani" municipality="Shanshenmiao" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Yunnan Prov.">
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;
<collectorName id="ABA21CE541E116E13D6D2290CEAC71DA">Hu</collectorName>
&amp;
<collectorName id="DA3B5E5FE87419262B90A5FD98BAB589">Yin</collectorName>
leg.; SHNU
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<materialsCitation id="0F4E3D885236F0D5874F4946E6DCC2BF" collectingDate="2009-05-03" collectorName="Nabanhe N. R., Hu, Yin" country="China" elevation="1750" location="Bengganghani" specimenCount="2" specimenCount-female="2" stateProvince="Yunnan Prov.">
<specimenCount id="C5CDB0641FA59A54AED81D14C317F81E" type="female">2♀♀</specimenCount>
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<collectorName id="8CFC5407A7D3972F59A69F0D921C6424">Nabanhe N.R.</collectorName>
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<collectorName id="F2A8B888FEC5C6ADF6DF2C5B4EF6EF06">Hu</collectorName>
&amp;
<collectorName id="9440DDE7AD0C245EC546EF014E84FD4D">Yin</collectorName>
leg.; SHNU
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<materialsCitation id="2CA8A44DE9E88B07ACCF2F6E6BF52907" collectingDate="2013-04-21" collectorName="Dai, Peng, Song" country="China" county="Baoshan City" elevation="1900" latitude="25.303055" location="Baihualing" longLatPrecision="21" longitude="98.79389" municipality="Mangkuan Town" specimenCount="5" specimenCount-female="1" specimenCount-male="4" stateProvince="Yunnan Prov.">
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</quantity>
;
<collectingDate id="DC14106C18B9988AA0BF318E6929DE11" value="2013-04-21">21 April 2013</collectingDate>
;
<collectorName id="5ADF8C5AD53F6034178D9E6D2CCEE4E8">Dai</collectorName>
,
<collectorName id="1CF8F7A251D65889804C129069B66E09">Peng</collectorName>
&amp;
<collectorName id="74C3C59B628E3A8E8FD4D6FCE57E83EA">Song</collectorName>
leg.; SHNU
</materialsCitation>
.
</paragraph>
<caption id="DACC169CE6F59229A5294C2348EE8F03" doi="10.3897/zookeys.1131.95038.figures15-20" httpUri="https://binary.pensoft.net/fig/772739" pageId="0" pageNumber="99" start="Figures 1520" startId="F3">
<paragraph id="018809525078BCA72469C33B4A089349" pageId="0" pageNumber="99">
<emphasis id="5FC9685A039A2DE4A0E4309ECBDD2FC9" bold="true" pageId="0" pageNumber="99">Figures 15-20.</emphasis>
<taxonomicName id="D2557FC308489E4F3A11908536209DA2" authorityName="Yang, Zhou &amp; Schillhammer" authorityYear="2011" class="Insecta" family="Staphylinidae" genus="Thoracostrongylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thoracostrongylus aduncatus" order="Coleoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="aduncatus">
<emphasis id="650D8FD58E58D65F409E99EBD0054FE9" italics="true" pageId="0" pageNumber="99">Thoracostrongylus aduncatus</emphasis>
</taxonomicName>
<emphasis id="DED05D99E165BBF923A7EF87D1B82DAB" bold="true" pageId="0" pageNumber="99">15, 16</emphasis>
habitus
<emphasis id="6C99614171AB58120A4BAEBC1AF16278" bold="true" pageId="0" pageNumber="99">17-20</emphasis>
aedeagus, ventral (
<emphasis id="83BA8F7DC1D033822F269BF2F26D9703" bold="true" pageId="0" pageNumber="99">17, 18</emphasis>
) and lateral (
<emphasis id="5C7D604DA65A14DDFFB5B2BE2AA52CE9" bold="true" pageId="0" pageNumber="99">19, 20</emphasis>
) views. Scale bars:
<quantity id="6DA1BFC586B1E39A9E22F33E403B0531" metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
(
<emphasis id="50628A00CB40E85AB726A72A4249D07B" bold="true" pageId="0" pageNumber="99">15, 16</emphasis>
); 0.2 mm (
<emphasis id="83877F3B4C7253DF0B96D792104BA1B0" bold="true" pageId="0" pageNumber="99">17-20</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="ECC8B9AFD039A58878B78F123F206920" pageId="0" pageNumber="99" type="description">
<paragraph id="DB9503A1A226FA8512F074740EF18964" pageId="0" pageNumber="99">Measurements.</paragraph>
<paragraph id="54813FD5B16DAB4C6C20E2A03EDA408D" pageId="0" pageNumber="99">
<emphasis id="E006E48301FAE17439A0B0F62D87D89E" bold="true" pageId="0" pageNumber="99">Male</emphasis>
: BL: 7.0-8.3 mm, FL: 4.2-5.1 mm. HL: 1.17-1.39 mm, HW: 1.61-1.89 mm, CL: 0.83-0.95 mm, PO: 0.17-0.28 mm, PL: 1.50-1.78 mm, PW: 1.22-1.50 mm, EL: 1.78-2.11 mm, EW: 1.78-2.11 mm. HL/HW: 0.70-0.77, CL/PO: 3.00-5.00, PL/PW: 1.12-1.27, EL/EW: 0.97-1.00, HW/EW: 0.84-0.91, PW/EW: 0.66-0.71, HW/PW: 1.24-1.32.
<emphasis id="B10C7B5E139BBAB31931A33039BC992B" bold="true" pageId="0" pageNumber="99">Female</emphasis>
: BL: 8.0-9.6 mm, FL: 4.8-5.3 mm. HL: 1.33-1.50 mm, HW: 1.83-2.06 mm, CL: 0.95-1.00 mm, PO: 0.22-0.28 mm, PL: 1.72-1.83 mm, PW: 1.39-1.56 mm, EL: 2.00-2.11 mm, EW: 2.11-2.22 mm. HL/HW: 0.73, CL/PO: 3.60-4.25, PL/PW: 1.18-1.24, EL/EW: 0.95, HW/EW: 0.87-0.93, PW/EW: 0.66-0.70, HW/PW: 1.32.
</paragraph>
<caption id="AA86D24C8C7665D6E43DDBABF7F4866D" doi="10.3897/zookeys.1131.95038.figures21-28" httpUri="https://binary.pensoft.net/fig/772740" pageId="0" pageNumber="99" start="Figures 2128" startId="F4">
<paragraph id="5B22FDDEFF0083DB08E551962BA6A792" pageId="0" pageNumber="99">
<emphasis id="925D8349549F55EB7F0720AE7E20902A" bold="true" pageId="0" pageNumber="99">Figures 21-28.</emphasis>
<taxonomicName id="52849EAF92DEF78B9925B0BBCC52F788" authorityName="Yang, Zhou &amp; Schillhammer" authorityYear="2011" class="Insecta" family="Staphylinidae" genus="Thoracostrongylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thoracostrongylus aduncatus" order="Coleoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="aduncatus">
<emphasis id="8FC208C70F8939D7C4B28A7CFB689BFE" italics="true" pageId="0" pageNumber="99">Thoracostrongylus aduncatus</emphasis>
</taxonomicName>
<emphasis id="B6B28D01AD4B4FD77628EE0623055988" bold="true" pageId="0" pageNumber="99">21-24</emphasis>
aedeagus from Baihualing, ventral (
<emphasis id="F5CAA374E9C0214901B76D1D7361608E" bold="true" pageId="0" pageNumber="99">21, 22</emphasis>
) and lateral (
<emphasis id="5BF9CB454CAB58CFDBED268123AC03DA" bold="true" pageId="0" pageNumber="99">23, 24</emphasis>
) views
<emphasis id="973BF1B89B567AE5DEB6E1F8BE51BEB2" bold="true" pageId="0" pageNumber="99">25-28</emphasis>
aedeagus from Baihualing, ventral (
<emphasis id="7816A73FA9903071614F44952C06D034" bold="true" pageId="0" pageNumber="99">25, 26</emphasis>
) and lateral (
<emphasis id="05D161B9F7A081DE7758EEA8792527CD" bold="true" pageId="0" pageNumber="99">27, 28</emphasis>
) views. Scale bars: 0.2 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="6D92BEB524E1C22A67B57ED2A5231658" pageId="0" pageNumber="99" type="diagnosis">
<paragraph id="CBC88E5ACA19C23DAEC7D4ADD09756E8" pageId="0" pageNumber="99">Diagnosis.</paragraph>
<paragraph id="99790CFFC31FE4040DB11E85963FB553" pageId="0" pageNumber="99">
The species is similar to
<taxonomicName id="FB2D8D19556CC0892515D97A70779557" class="Insecta" kingdom="Animalia" lsidName="T. acerosus" order="Coleoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="acerosus">
<emphasis id="596BB6B4AB664C31DEBE571C9BF173F7" italics="true" pageId="0" pageNumber="99">T. acerosus</emphasis>
</taxonomicName>
Yang, Zhou &amp; Schillhammer, 2011,
<taxonomicName id="34E81D8850F7D68D80A2AA57B2E6F5BB" class="Insecta" kingdom="Animalia" lsidName="T. fujianensis" order="Coleoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="fujianensis">
<emphasis id="9F803BBCE2D8AF4F4ED944230126EDF1" italics="true" pageId="0" pageNumber="99">T. fujianensis</emphasis>
</taxonomicName>
Yang, Zhou &amp; Schillhammer, 2011, and
<taxonomicName id="A8C82CDE95624AD019AC245DFBFC4F35" class="Insecta" kingdom="Animalia" lsidName="T. diaoluoensis" order="Coleoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="diaoluoensis">
<emphasis id="DA2B3FC23F90ED7ADC19A66E2748FE7E" italics="true" pageId="0" pageNumber="99">T. diaoluoensis</emphasis>
</taxonomicName>
Yang, Zhou &amp; Schillhammer, 2011 in general appearance, but it can be distinguished from them by the apex of median lobe pointed dorsad forming an apical tooth in lateral view, and from
<taxonomicName id="991D2170EDF64337DCB64EEB30E5A230" class="Insecta" kingdom="Animalia" lsidName="T. diaoluoensis" order="Coleoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="diaoluoensis">
<emphasis id="12E205D05CE7C02F2CA82DC9490E28E6" italics="true" pageId="0" pageNumber="99">T. diaoluoensis</emphasis>
</taxonomicName>
also by the aedeagal median lobe with a subapical tooth on the dorsal side. Aedeagal variation (Figs
<figureCitation id="C796C87DE6EF892E55B6CA3DB17302A4" captionStart="Figures 1520" captionStartId="F3" captionText="Figures 15 - 20. Thoracostrongylus aduncatus 15, 16 habitus 17 - 20 aedeagus, ventral (17, 18) and lateral (19, 20) views. Scale bars: 1 mm (15, 16); 0.2 mm (17 - 20)." figureDoi="10.3897/zookeys.1131.95038.figures15-20" httpUri="https://binary.pensoft.net/fig/772739" pageId="0" pageNumber="99">17</figureCitation>
-
<figureCitation id="2BE243CCB6944D3686FED02A90C05ABE" captionStart="Figures 2932" captionStartId="F5" captionText="Figures 29 - 32. Thoracostrongylus aduncatus 29 - 32 aedeagus from Nabanhe, ventral (29, 30) and lateral (31, 32) views. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1131.95038.figures29-32" httpUri="https://binary.pensoft.net/fig/772741" pageId="0" pageNumber="99">32</figureCitation>
) occurs in the apical parts of median lobe and paramere.
</paragraph>
<caption id="C03993FCEFFB3FC41E2D1C931350F319" doi="10.3897/zookeys.1131.95038.figures29-32" httpUri="https://binary.pensoft.net/fig/772741" pageId="0" pageNumber="99" start="Figures 2932" startId="F5">
<paragraph id="750A146CD184F37A7660290563E1B5E1" pageId="0" pageNumber="99">
<emphasis id="90D2026C1261D08C81B3C22580CB480F" bold="true" pageId="0" pageNumber="99">Figures 29-32.</emphasis>
<taxonomicName id="4D657328D4AFC5D18B30E4189C0F2A10" authorityName="Yang, Zhou &amp; Schillhammer" authorityYear="2011" class="Insecta" family="Staphylinidae" genus="Thoracostrongylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thoracostrongylus aduncatus" order="Coleoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="aduncatus">
<emphasis id="F31CDC719FEAD31912FFEC6249F09079" italics="true" pageId="0" pageNumber="99">Thoracostrongylus aduncatus</emphasis>
</taxonomicName>
<emphasis id="4E43B68574A3FE548DF03A5A206C8379" bold="true" pageId="0" pageNumber="99">29-32</emphasis>
aedeagus from Nabanhe, ventral (
<emphasis id="8D33B0041C27622ABD4B2589F5606059" bold="true" pageId="0" pageNumber="99">29, 30</emphasis>
) and lateral (
<emphasis id="064B4595DD47DC80DB4DF9EFE1A04119" bold="true" pageId="0" pageNumber="99">31, 32</emphasis>
) views. Scale bars: 0.2 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="BBF69CA4424FC9765B1424F3B5C51084" pageId="0" pageNumber="99" type="distribution">
<paragraph id="249E87955F9B86C7D6458CABA515D632" pageId="0" pageNumber="99">Distribution.</paragraph>
<paragraph id="CF6C9EA0D69A801C0DB995044440EDED" pageId="0" pageNumber="99">China (Yunnan).</paragraph>
</subSubSection>
</treatment>
</document>