167 lines
19 KiB
XML
167 lines
19 KiB
XML
<document id="01B744EC15C7EDE981FC904E0CBDB593" ID-DOI="10.5281/zenodo.191040" ID-GBIF-Dataset="3e731a6c-b611-4044-8112-370881753ca2" ID-ISSN="1175-5326" ID-Zenodo-Dep="191040" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461122948366" checkinUser="plazi" docAuthor="Capellari, Renato Soares & Amorim, Dalton De Souza" docDate="2009" docId="0907EA79D1315157E0B9F943FDB9FD71" docLanguage="en" docName="zt02274p061.pdf" docOrigin="Zootaxa 2274" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Paraclius parenti Capellari & Amorim, 2009, sp. nov." docType="treatment" docVersion="9" lastPageNumber="55" masterDocId="F53E9201D1325153E02EFF88FF9AFFD9" masterDocTitle="Four new species of Paraclius Loew (Diptera: Dolichopodidae) from the Brazilian Atlantic Forest, and notes on the systematic position of Cheiromyia Dyte" masterLastPageNumber="61" masterPageNumber="51" pageNumber="54" updateTime="1698596254519" updateUser="plazi">
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<mods:mods id="5CD0C860A915970A9EA128853428C6FE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="53D78A2830F0E679CE6E6924FBB860F1">
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<mods:title id="49BDCE131FBB7B904BFD2DB6A9CDA07F">Four new species of Paraclius Loew (Diptera: Dolichopodidae) from the Brazilian Atlantic Forest, and notes on the systematic position of Cheiromyia Dyte</mods:title>
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</mods:titleInfo>
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<mods:name id="1A9C8D6C3F59DE4AA0496B362C814690" type="personal">
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<mods:roleTerm id="4BE9B4D9691C5F5E8CD892C9D65DBC63">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C43AC65C9314105247FA0B0F1493B3B2">Capellari, Renato Soares</mods:namePart>
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</mods:name>
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<mods:name id="2378BF04132198C62A72D3E50618C8F1" type="personal">
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<mods:role id="9B04E6F78932A476CCFCD9F45E101D1E">
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<mods:roleTerm id="3E3ACC6DD256D2470089EC103AE74399">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="69295B4904A8CBD01B9F6FDCBC18CE9C">Amorim, Dalton De Souza</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="8C27F052DC7D5006C1584DA44623E12B">text</mods:typeOfResource>
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<mods:titleInfo id="9D87E9F5838AAF2DA259D5D0CAD5C5B5">
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<mods:title id="95153CC26B12D2D4B60D6E1C773765DA">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="10AFE7338899049FE85D3E38FA1474A1">
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<mods:date id="AB3B4B6544EA5B8B6A5CD78AEB9802DF">2009</mods:date>
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<mods:detail id="25307191537246A7FC657B3C693F380F" type="volume">
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<mods:number id="212062FF44318F3B8841A2F596C3FB7B">2274</mods:number>
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<mods:start id="D09E3A28AF1496FF5E885F181CBDB158">51</mods:start>
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<mods:end id="37E61CD1504AE01A0958E06A3435BD85">61</mods:end>
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<mods:classification id="2CEF432D8A1D9A63C3A5D376F025FE95">journal article</mods:classification>
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<mods:identifier id="3864159BCB20D0BF227FEC6DA58AD157" type="DOI">10.5281/zenodo.191040</mods:identifier>
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<mods:identifier id="2C4F6D54E219A280B25878BD3B307F77" type="GBIF-Dataset">3e731a6c-b611-4044-8112-370881753ca2</mods:identifier>
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<mods:identifier id="DB54DE1B9C8CD0F000B659CC3315BED3" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="2593FD08AD23C8A4A59312543CF4DBD2" type="Zenodo-Dep">191040</mods:identifier>
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<treatment id="0907EA79D1315157E0B9F943FDB9FD71" ID-DOI="http://doi.org/10.5281/zenodo.6212824" ID-GBIF-Taxon="119625891" ID-Zenodo-Dep="6212824" LSID="urn:lsid:plazi:treatment:0907EA79D1315157E0B9F943FDB9FD71" httpUri="http://treatment.plazi.org/id/0907EA79D1315157E0B9F943FDB9FD71" lastPageId="4" lastPageNumber="55" pageId="3" pageNumber="54">
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<subSubSection id="C9B408E4D1315150E0B9F943FEEAF8D1" pageId="3" pageNumber="54" type="nomenclature">
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<paragraph id="81115B6FD1315150E0B9F943FE4EF93D" blockId="3.[151,468,1738,1800]" box="[151,468,1738,1765]" pageId="3" pageNumber="54">
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<heading id="DA59EC03D1315150E0B9F943FE4EF93D" bold="true" box="[151,468,1738,1765]" fontSize="11" level="1" pageId="3" pageNumber="54" reason="1">
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||
<emphasis id="B3DA877DD1315150E0B9F943FE4EF93D" bold="true" box="[151,468,1738,1765]" pageId="3" pageNumber="54">
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<taxonomicName id="46AE20ECD1315150E0B9F943FEF2F93C" ID-CoL="9HWFX" box="[151,360,1739,1765]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="parenti" status="sp. nov.">
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<emphasis id="B3DA877DD1315150E0B9F943FEF2F93C" bold="true" box="[151,360,1739,1765]" italics="true" pageId="3" pageNumber="54">Paraclius parenti</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A8E93A06D1315150E159F942FE4EF93D" box="[375,468,1738,1764]" pageId="3" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="81115B6FD1315150E0B9F966FEEAF8D1" blockId="3.[151,468,1738,1800]" box="[151,368,1774,1800]" pageId="3" pageNumber="54">
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(
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<figureCitation id="199547EAD1315150E0B1F966FE8AF8D1" box="[159,272,1774,1800]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,273,1943,1967]" captionTargetBox="[204,1382,183,1918]" captionTargetId="figure@2.[204,1382,183,1918]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Male hypopygium of Paraclius. 1 – 2. P. amphiatheratus, sp. nov., paratype (1, left lateral view; 2, ventral view). 3 – 4. P. parenti, sp. nov., paratype (3, left lateral view; 4, ventral view). In lateral view, left cercus removed; in ventral view, cerci not depicted." httpUri="https://zenodo.org/record/191041/files/figure.png" pageId="3" pageNumber="54">Figs. 3–4</figureCitation>
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,
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<figureCitation id="199547EAD1315150E132F966FEFCF8D1" box="[284,358,1774,1800]" captionStart="FIGURE 15 – 16" captionStartId="8.[151,255,1946,1970]" captionTargetBox="[177,1419,1266,1911]" captionTargetId="figure@8.[151,1436,1257,1921]" captionTargetPageId="8" captionText="FIGURE 15 – 16. Paraclius parenti, sp. nov., paratype. 15. Male antenna. 16. Male wing." httpUri="https://zenodo.org/record/191045/files/figure.png" pageId="3" pageNumber="54">15–16</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C9B408E4D1315150E0B9F8B2FAEBF87B" pageId="3" pageNumber="54" type="diagnosis">
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<paragraph id="81115B6FD1315150E0B9F8B2FAEBF87B" blockId="3.[151,1436,1850,1954]" pageId="3" pageNumber="54">
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<emphasis id="B3DA877DD1315150E0B9F8B2FE80F88D" bold="true" box="[151,282,1850,1876]" pageId="3" pageNumber="54">Diagnosis.</emphasis>
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First flagellomere almost rectangular, not pointed at apex; basal article of stylus about half the length of distal article. Two short black setae dorsally on proepisternum and scattered white setulae. Phallus without ornamentation except serrate ventral surface of apex; hypandrium harpoon-shaped in lateral view.
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</paragraph>
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</subSubSection>
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<subSubSection id="C9B408E4D1315157E0EBF85DFD4BFF01" lastPageId="4" lastPageNumber="55" pageId="3" pageNumber="54" type="materials_examined">
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<paragraph id="81115B6FD1315157E0EBF85DFD4BFF01" blockId="3.[197,1436,2005,2032]" lastBlockId="4.[151,1437,151,680]" lastPageId="4" lastPageNumber="55" pageId="3" pageNumber="54">
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<emphasis id="B3DA877DD1315150E0EBF85DFE20F836" bold="true" box="[197,442,2005,2031]" pageId="3" pageNumber="54">Material examined.</emphasis>
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<typeStatus id="5E15E5CDD1315150E1EFF85EFDB4F829" box="[449,558,2006,2032]" pageId="3" pageNumber="54" type="holotype">Holotype</typeStatus>
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3,
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<collectingCountry id="F9B91BFFD1315150E275F85EFD52F829" box="[603,712,2006,2032]" name="Brazil" pageId="3" pageNumber="54">BRAZIL</collectingCountry>
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,
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<emphasis id="B3DA877DD1315150E2FAF85DFCE9F836" bold="true" box="[724,883,2005,2031]" pageId="3" pageNumber="54">Pernambuco</emphasis>
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, Recife, Parque dos Dois Irmãos, 1 3, track 3, Malaise trap,
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<date id="F5107DAFD1365157E118FF10FE70FF6B" box="[310,490,152,178]" pageId="4" pageNumber="55" value="2002-07-17" valueMax="2002-07-20" valueMin="2002-07-17">17–20.vii.2002</date>
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, S. T. P. Amarante e eq. cols.
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<typeStatus id="5E15E5CDD1365157E36AFF10FC20FF6B" box="[836,954,152,178]" pageId="4" pageNumber="55" type="paratype">Paratypes</typeStatus>
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: 1 3, same data as
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<typeStatus id="5E15E5CDD1365157E4B7FF10FA99FF6B" box="[1177,1283,152,178]" pageId="4" pageNumber="55" type="holotype">holotype</typeStatus>
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, except track 5; same, but 1 3, track 3; same, but 2 3, track 1.
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</paragraph>
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</subSubSection>
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<subSubSection id="C9B408E4D1365157E0EBFF6DFF48FE43" pageId="4" pageNumber="55" type="description">
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<paragraph id="81115B6FD1365157E0EBFF6DFF48FE43" blockId="4.[151,1437,151,680]" pageId="4" pageNumber="55">
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<emphasis id="B3DA877DD1365157E0EBFF6DFEFEFF26" bold="true" box="[197,356,229,255]" pageId="4" pageNumber="55">Description.</emphasis>
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Very similar to
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<taxonomicName id="46AE20ECD1365157E203FF6DFC96FF27" box="[557,780,229,254]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="amphiatheratus">
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<emphasis id="B3DA877DD1365157E203FF6DFC96FF27" box="[557,780,229,254]" italics="true" pageId="4" pageNumber="55">P. amphiatheratus</emphasis>
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</taxonomicName>
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except by the features listed in the diagnosis and the following details.
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<emphasis id="B3DA877DD1365157E143FE83FE2EFEFC" bold="true" box="[365,436,267,293]" pageId="4" pageNumber="55">Male.</emphasis>
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Body length, 2.9–3.1. Wing (
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<figureCitation id="199547EAD1365157E33BFE84FCF6FEFF" box="[789,876,268,294]" captionStart="FIGURE 15 – 16" captionStartId="8.[151,255,1946,1970]" captionTargetBox="[177,1419,1266,1911]" captionTargetId="figure@8.[151,1436,1257,1921]" captionTargetPageId="8" captionText="FIGURE 15 – 16. Paraclius parenti, sp. nov., paratype. 15. Male antenna. 16. Male wing." httpUri="https://zenodo.org/record/191045/files/figure.png" pageId="4" pageNumber="55">Fig. 16</figureCitation>
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), 2.5–2.7 long, 0.9–1.0 wide. Male antennae as in
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<figureCitation id="199547EAD1365157E09BFEBAFE96FE95" box="[181,268,306,332]" captionStart="FIGURE 15 – 16" captionStartId="8.[151,255,1946,1970]" captionTargetBox="[177,1419,1266,1911]" captionTargetId="figure@8.[151,1436,1257,1921]" captionTargetPageId="8" captionText="FIGURE 15 – 16. Paraclius parenti, sp. nov., paratype. 15. Male antenna. 16. Male wing." httpUri="https://zenodo.org/record/191045/files/figure.png" pageId="4" pageNumber="55">Fig. 15</figureCitation>
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. Leg measures: I: 0.9, 0.9, 0.4, 0.1, 0.1, 0.1, 0.1. II: 1.2, 1.2, 0.5, 0.3, 0.2, 0.2, 0.1. III: 1.2, 1.3, 0.4, 0.5, 0.3, 0.2, 0.1. Hypopygium as in
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<figureCitation id="199547EAD1365157E27FFED1FD59FEAA" box="[593,707,345,371]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,273,1943,1967]" captionTargetBox="[204,1382,183,1918]" captionTargetId="figure@2.[204,1382,183,1918]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Male hypopygium of Paraclius. 1 – 2. P. amphiatheratus, sp. nov., paratype (1, left lateral view; 2, ventral view). 3 – 4. P. parenti, sp. nov., paratype (3, left lateral view; 4, ventral view). In lateral view, left cercus removed; in ventral view, cerci not depicted." httpUri="https://zenodo.org/record/191041/files/figure.png" pageId="4" pageNumber="55">Figs. 3–4</figureCitation>
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. Sperm pump slightly enlarged and weakly folded back over itself
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</paragraph>
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</subSubSection>
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<subSubSection id="C9B408E4D1365157E0EBFE2EFBD9FE3E" pageId="4" pageNumber="55" type="etymology">
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<paragraph id="81115B6FD1365157E0EBFE2EFBD9FE3E" blockId="4.[151,1437,151,680]" pageId="4" pageNumber="55">
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<emphasis id="B3DA877DD1365157E0EBFE2EFEC9FE19" bold="true" box="[197,339,422,448]" pageId="4" pageNumber="55">Etymology.</emphasis>
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This species is named after the French abbot and dipterist Octave Parent (
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<date id="F5107DAFD1365157E4E7FE2EFACEFE19" box="[1225,1364,422,448]" pageId="4" pageNumber="55" value="1882" valueMax="1942">1882–1942</date>
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), who extensively contributed to the development of the taxonomy of
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<taxonomicName id="46AE20ECD1365157E351FE45FBA4FE3E" box="[895,1086,461,487]" class="Insecta" family="Dolichopodidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="family">Dolichopodidae</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C9B408E4D1365157E0EBFE7BFDB9FD71" pageId="4" pageNumber="55" type="discussion">
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<paragraph id="81115B6FD1365157E0EBFE7BFDB9FD71" blockId="4.[151,1437,151,680]" pageId="4" pageNumber="55">
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<emphasis id="B3DA877DD1365157E0EBFE7BFECEFDD4" bold="true" box="[197,340,499,525]" pageId="4" pageNumber="55">Comments</emphasis>
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. This species resembles
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<taxonomicName id="46AE20ECD1365157E2BDFE7CFC2DFDD4" box="[659,951,500,525]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="dicrophallus" status="sp. nov.">
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<emphasis id="B3DA877DD1365157E2BDFE7CFC2DFDD4" box="[659,951,500,525]" italics="true" pageId="4" pageNumber="55">Paraclius dicrophallus</emphasis>
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</taxonomicName>
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,
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<emphasis id="B3DA877DD1365157E3E6FE7BFBB5FDD4" bold="true" box="[968,1071,499,525]" pageId="4" pageNumber="55">
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<taxonomicNameLabel id="A8E93A06D1365157E3E6FE7BFBB5FDD4" box="[968,1071,499,525]" pageId="4" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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in the rectangular male first flagellomere, but can be easily separated from it by the shape of the phallus (simple, not furcated in
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<taxonomicName id="46AE20ECD1365157E50AFD92FA0FFDEA" box="[1316,1429,538,563]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="parenti" status="sp. nov.">
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<emphasis id="B3DA877DD1365157E50AFD92FA0FFDEA" box="[1316,1429,538,563]" italics="true" pageId="4" pageNumber="55">P. parenti</emphasis>
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</taxonomicName>
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,
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<emphasis id="B3DA877DD1365157E0B9FDC8FF6EFD83" bold="true" box="[151,244,576,602]" pageId="4" pageNumber="55">
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<taxonomicNameLabel id="A8E93A06D1365157E0B9FDC8FF6EFD83" box="[151,244,576,602]" pageId="4" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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,
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<figureCitation id="199547EAD1365157E12CFDC9FEE8FD82" box="[258,370,577,603]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,273,1943,1967]" captionTargetBox="[204,1382,183,1918]" captionTargetId="figure@2.[204,1382,183,1918]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Male hypopygium of Paraclius. 1 – 2. P. amphiatheratus, sp. nov., paratype (1, left lateral view; 2, ventral view). 3 – 4. P. parenti, sp. nov., paratype (3, left lateral view; 4, ventral view). In lateral view, left cercus removed; in ventral view, cerci not depicted." httpUri="https://zenodo.org/record/191041/files/figure.png" pageId="4" pageNumber="55">Figs. 3–4</figureCitation>
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, and divided mesally at apex into a pair of stems in
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<taxonomicName id="46AE20ECD1365157E3E2FDC9FBE1FD83" box="[972,1147,577,602]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="dicrophallus" status="sp. nov.">
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<emphasis id="B3DA877DD1365157E3E2FDC9FBE1FD83" box="[972,1147,577,602]" italics="true" pageId="4" pageNumber="55">P. dicrophallus</emphasis>
|
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</taxonomicName>
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,
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<emphasis id="B3DA877DD1365157E4A7FDC8FB7CFD83" bold="true" box="[1161,1254,576,602]" pageId="4" pageNumber="55">
|
||
<taxonomicNameLabel id="A8E93A06D1365157E4A7FDC8FB7CFD83" box="[1161,1254,576,602]" pageId="4" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
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<figureCitation id="199547EAD1365157E4DAFDC9FAF8FD82" box="[1268,1378,577,603]" captionStart="FIGURES 5 – 8" captionStartId="5.[151,272,1930,1954]" captionTargetBox="[248,1339,181,1904]" captionTargetId="figure@5.[248,1340,181,1905]" captionTargetPageId="5" captionText="FIGURES 5 – 8. Male hypopygium of Paraclius. 5 – 6. P. sagittatus, sp. nov., holotype (5, left lateral view; 6, ventral view). 7 – 8. P. dicrophallus, sp. nov., holotype (7, left lateral view; 8, ventral view). In lateral view, left cercus removed; in ventral view, cerci not depicted." httpUri="https://zenodo.org/record/191042/files/figure.png" pageId="4" pageNumber="55">Figs. 7–8</figureCitation>
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) and of the hypandrium (harpoon-shaped in
|
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<taxonomicName id="46AE20ECD1365157E240FDE0FD78FD58" box="[622,738,616,641]" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="parenti" status="sp. nov.">
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<emphasis id="B3DA877DD1365157E240FDE0FD78FD58" box="[622,738,616,641]" italics="true" pageId="4" pageNumber="55">P. parenti</emphasis>
|
||
</taxonomicName>
|
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,
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<emphasis id="B3DA877DD1365157E2DCFDEFFCC8FD58" bold="true" box="[754,850,615,641]" pageId="4" pageNumber="55">
|
||
<taxonomicNameLabel id="A8E93A06D1365157E2DCFDEFFCC8FD58" box="[754,850,615,641]" pageId="4" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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,
|
||
<figureCitation id="199547EAD1365157E34DFDE0FC36FD5B" box="[867,940,616,642]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,273,1943,1967]" captionTargetBox="[204,1382,183,1918]" captionTargetId="figure@2.[204,1382,183,1918]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Male hypopygium of Paraclius. 1 – 2. P. amphiatheratus, sp. nov., paratype (1, left lateral view; 2, ventral view). 3 – 4. P. parenti, sp. nov., paratype (3, left lateral view; 4, ventral view). In lateral view, left cercus removed; in ventral view, cerci not depicted." httpUri="https://zenodo.org/record/191041/files/figure.png" pageId="4" pageNumber="55">Fig. 3</figureCitation>
|
||
, and laterally forked into two stems in
|
||
<taxonomicName id="46AE20ECD1365157E5A8FDE0FEB0FD71" class="Insecta" family="Dolichopodidae" genus="Paraclius" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="dicrophallus" status="sp. nov.">
|
||
<emphasis id="B3DA877DD1365157E5A8FDE0FEB0FD71" italics="true" pageId="4" pageNumber="55">P. dicrophallus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B3DA877DD1365157E116FD06FE0FFD71" bold="true" box="[312,405,654,680]" pageId="4" pageNumber="55">
|
||
<taxonomicNameLabel id="A8E93A06D1365157E116FD06FE0FFD71" box="[312,405,654,680]" pageId="4" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<figureCitation id="199547EAD1365157E18BFD06FD8FFD71" box="[421,533,654,680]" captionStart="FIGURES 5 – 8" captionStartId="5.[151,272,1930,1954]" captionTargetBox="[248,1339,181,1904]" captionTargetId="figure@5.[248,1340,181,1905]" captionTargetPageId="5" captionText="FIGURES 5 – 8. Male hypopygium of Paraclius. 5 – 6. P. sagittatus, sp. nov., holotype (5, left lateral view; 6, ventral view). 7 – 8. P. dicrophallus, sp. nov., holotype (7, left lateral view; 8, ventral view). In lateral view, left cercus removed; in ventral view, cerci not depicted." httpUri="https://zenodo.org/record/191042/files/figure.png" pageId="4" pageNumber="55">Figs. 7–8</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |