127 lines
16 KiB
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127 lines
16 KiB
XML
<document id="564B12F6D34DC0685B5CC5512DE75A77" ID-DOI="10.5281/zenodo.186069" ID-GBIF-Dataset="99d44e7b-46c3-47a2-9695-3ea43c4186a0" ID-ISSN="1175-5326" ID-Zenodo-Dep="186069" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460068170820" checkinUser="plazi" docAuthor="Maddison, Wayne P." docDate="2009" docId="335D87D75E071F1DFF115E8F265EAE54" docLanguage="en" docName="zt02021p022.pdf" docOrigin="Zootaxa 2021" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Cucudeta Maddison, 2009, new genus" docType="treatment" docVersion="9" lastPageNumber="15" masterDocId="CF64FFAF5E091F13FF865E182756AA37" masterDocTitle="New cocalodine jumping spiders from Papua New Guinea (Araneae: Salticidae: Cocalodinae)" masterLastPageNumber="22" masterPageNumber="1" pageNumber="15" updateTime="1698234705597" updateUser="plazi">
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<mods:mods id="579566BB18FDE279680D67C56D6C0A99" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B103B7FA8F150B46ADCBC903AA87C32C">
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<mods:title id="1140E2EAD20950EB141039D88CD97B65">New cocalodine jumping spiders from Papua New Guinea (Araneae: Salticidae: Cocalodinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="9EA17560167892BD5AA3F4822AC2DFE4" type="personal">
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<mods:roleTerm id="813C7BD51EF0C8D4797CBE44254CD32D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="AD8AF7FBF13677ED246327D13447A617">Maddison, Wayne P.</mods:namePart>
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<mods:typeOfResource id="378F43925E3E1F6F6531F2064C31371A">text</mods:typeOfResource>
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<mods:titleInfo id="FF32A9708BDD99B896D9845BF5C14054">
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<mods:title id="4CA34A74D363FF3FDEB7568EDFD2F2A4">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="96A6B13BD862CE6E69CD9BB6E6BEFC43">
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<mods:date id="8D2214288915260CDB0DCB9B724E5D8F">2009</mods:date>
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<mods:detail id="3C39C64A3FCE5C1735C837FD32B0BB67" type="volume">
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<mods:number id="68ACDF2FB31FE59F1CA4B00E14798A82">2021</mods:number>
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<mods:start id="18D1E8CAEECD04E1198B0692CAEEB5A2">1</mods:start>
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<mods:end id="54F1CAC09B3F41B4A4F0092B79074FB3">22</mods:end>
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<mods:classification id="680DA13FFAEC6B5C38E82E403A6BFE27">journal article</mods:classification>
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<mods:identifier id="D12277A802CC21E0C9A10071226B5036" type="DOI">10.5281/zenodo.186069</mods:identifier>
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<mods:identifier id="C519B1E1DDA70114C92DB621B0E2444A" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="D2905A0610E185B83C1F0234394E4C8B" type="Zenodo-Dep">186069</mods:identifier>
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<treatment id="335D87D75E071F1DFF115E8F265EAE54" ID-DOI="http://doi.org/10.5281/zenodo.6226964" ID-GBIF-Taxon="124874388" ID-Zenodo-Dep="6226964" LSID="urn:lsid:plazi:treatment:335D87D75E071F1DFF115E8F265EAE54" httpUri="http://treatment.plazi.org/id/335D87D75E071F1DFF115E8F265EAE54" lastPageNumber="15" pageId="14" pageNumber="15">
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<subSubSection id="F3EE654A5E071F1DFF115E8F26BCAA86" box="[151,490,151,177]" pageId="14" pageNumber="15" type="nomenclature">
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<paragraph id="BB4B36C15E071F1DFF115E8F26BCAA86" blockId="14.[151,490,151,177]" box="[151,490,151,177]" pageId="14" pageNumber="15">
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<heading id="E00381AD5E071F1DFF115E8F26BCAA86" bold="true" box="[151,490,151,177]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
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<emphasis id="8980EAD35E071F1DFF115E8F26BCAA86" bold="true" box="[151,490,151,177]" pageId="14" pageNumber="15">
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Genus
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<taxonomicName id="7CF44D425E071F1DFF6B5E8F2637AA86" ID-CoL="3X83" ID-ENA="1560424" box="[237,353,151,177]" class="Arachnida" family="Salticidae" genus="Cucudeta" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="8980EAD35E071F1DFF6B5E8F2637AA86" bold="true" box="[237,353,151,177]" italics="true" pageId="14" pageNumber="15">Cucudeta</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="92B357A85E071F1DFEEE5E8F26BCAA86" box="[360,490,151,177]" pageId="14" pageNumber="15" rank="genus">new genus</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="F3EE654A5E071F1DFF115ECF252CAAD8" box="[151,634,215,239]" pageId="14" pageNumber="15" type="materials_examined">
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<paragraph id="BB4B36C15E071F1DFF115ECF252CAAD8" blockId="14.[151,634,215,239]" box="[151,634,215,239]" pageId="14" pageNumber="15">
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<emphasis id="8980EAD35E071F1DFF115ECF267BAAD8" bold="true" box="[151,301,215,239]" pageId="14" pageNumber="15">
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<typeStatus id="644F88635E071F1DFF115ECF2799AAD8" box="[151,207,215,239]" pageId="14" pageNumber="15">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="7CF44D425E071F1DFEB55EC026B0AAD8" box="[307,486,216,239]" class="Arachnida" family="Salticidae" genus="Cucudeta" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="zabkai" status="sp. nov.">
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<emphasis id="8980EAD35E071F1DFEB55EC026B0AAD8" box="[307,486,216,239]" italics="true" pageId="14" pageNumber="15">Cucudeta zabkai</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="92B357A85E071F1DFE755EC02520AAD8" box="[499,630,216,239]" pageId="14" pageNumber="15" rank="species">new species</taxonomicNameLabel>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="F3EE654A5E071F1DFF115F0226F6AB9C" pageId="14" pageNumber="15" type="etymology">
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<paragraph id="BB4B36C15E071F1DFF115F0226F6AB9C" blockId="14.[151,1437,282,1123]" pageId="14" pageNumber="15">
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<emphasis id="8980EAD35E071F1DFF115F02267EAB03" bold="true" box="[151,296,282,308]" pageId="14" pageNumber="15">Etymology.</emphasis>
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The name is based on "
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<emphasis id="8980EAD35E071F1DFDC95F0225FBAB04" box="[591,685,282,307]" italics="true" pageId="14" pageNumber="15">kukudet</emphasis>
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", the word for spider in the language of the Hewa people, on whose land I first collected these spiders. "
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<emphasis id="8980EAD35E071F1DFD255F592451AB6D" box="[675,775,321,346]" italics="true" pageId="14" pageNumber="15">Kukudet</emphasis>
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" represents the pronunciation as interpreted by me on hearing the word spoken;
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<bibRefCitation id="DF654B305E071F1DFE4A5F7225D7ABB3" author="Vollrath" box="[460,641,362,388]" pageId="14" pageNumber="22" refString="Vollrath, P. W. (1985) Hewa phonemes: a tentative statement. In: Clifton, J. M. (Ed.), Five phonological studies, Workpapers in Papua New Guinea Languages 31, Ukarumpa: Summer Institute of Linguistics, 51 - 83." type="journal article" year="1985">Vollrath (1985)</bibRefCitation>
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lists the word as having the phonemes /'ku.ku.tε.tε/. The name is to be treated as feminine.
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</paragraph>
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</subSubSection>
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<subSubSection id="F3EE654A5E071F1DFF405FA0265EAE54" pageId="14" pageNumber="15" type="diagnosis">
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<paragraph id="BB4B36C15E071F1DFF405FA025ADA963" blockId="14.[151,1437,282,1123]" pageId="14" pageNumber="15">
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<emphasis id="8980EAD35E071F1DFF405FA02610ABE5" bold="true" box="[198,326,440,466]" pageId="14" pageNumber="15">Diagnosis.</emphasis>
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Small dark brown to black leaf litter salticids, sparsely covered with relatively small setae. No other known salticid has the following combination of features: (1) First metatarsus with four prominent pairs of ventral macrosetae (
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<figureCitation id="23CF2A445E071F1DFE2B5C1D255EA828" box="[429,520,517,543]" captionStart="FIGURES 72 – 86" captionStartId="18.[151,271,1831,1855]" captionTargetBox="[199,1385,207,1802]" captionTargetId="figure@18.[190,1396,197,1807]" captionTargetPageId="18" captionText="FIGURES 72 – 86. Cucudeta species. 72 – 78 C. zabkai, new species. 72 – 73 Palp of holotype male (72 ventral view, 73 retrolateral view). 74 Metatarsus and tarsus of first leg, female. 75 – 76 female carapace. 77 Epigynum, ventral view. 78 Cleared epigynum, dorsal view. 79 – 82 C. uzet, new species. 79 – 80 Palp of holotype male (79 ventral view, 80 retrolateral view). 81 Epigynum, ventral view. 82 Cleared epigynum, dorsal view. 83 – 86 C. gahavisuka, new species. 83 – 84 Palp of holotype male (83 ventral view, 84 retrolateral view). Figure 85 Epigynum, ventral view. 86 Cleared epigynum, dorsal view. Scale bars 0.1 mm on genitalia and first leg, 1 mm on carapace." httpUri="https://zenodo.org/record/186080/files/figure.png" pageId="14" pageNumber="15">Fig. 74</figureCitation>
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). This contrasts with the remaining leg segments which have relatively few macrosetae compared to other salticids. (2) Posterior median eyes displaced medially so as to be on the top of the ocular quadrangle, pointing upwards rather than diagonally as in most salticids. The medial displacement brings the PME to touch the line between the medial edges of the PLE and ALE. Other salticids known to me with PME so far medial are lyssomanines and hisponines. (3) Male palpus with median apophysis. (4) Lacking distinctive constriction between PME and PLE present in hisponines. (5) Sternum with raised mound at each of the leg bases, especially the first. Among non-salticoid salticids with a median apophysis,
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<taxonomicName id="7CF44D425E071F1DFAAA5CF622CAA930" box="[1324,1436,750,775]" class="Arachnida" family="Salticidae" genus="Cucudeta" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
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<emphasis id="8980EAD35E071F1DFAAA5CF622CAA930" box="[1324,1436,750,775]" italics="true" pageId="14" pageNumber="15">Cucudeta</emphasis>
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</taxonomicName>
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is most similar in habits and body form to
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<taxonomicName id="7CF44D425E071F1DFD285D0C25A8A91A" box="[686,766,788,813]" class="Arachnida" family="Salticidae" genus="Eupoa" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
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<emphasis id="8980EAD35E071F1DFD285D0C25A8A91A" box="[686,766,788,813]" italics="true" pageId="14" pageNumber="15">Eupoa</emphasis>
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</taxonomicName>
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, from which it differs by features (1) and (2), a more elongate body, and the simpler form of the embolus.
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</paragraph>
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<paragraph id="BB4B36C15E071F1DFF435D79265EAE54" blockId="14.[151,1437,282,1123]" pageId="14" pageNumber="15">
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<emphasis id="8980EAD35E071F1DFF435D792511A94C" bold="true" box="[197,583,865,891]" pageId="14" pageNumber="15">Natural history and behaviour.</emphasis>
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All three species of this genus were collected in mid to high elevation in association with leaf litter in rainforest.
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<taxonomicName id="7CF44D425E071F1DFDEC5D902478A996" box="[618,814,904,929]" class="Arachnida" family="Salticidae" genus="Cucudeta" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="zabkai">
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<emphasis id="8980EAD35E071F1DFDEC5D902478A996" box="[618,814,904,929]" italics="true" pageId="14" pageNumber="15">Cucudeta zabkai</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="7CF44D425E071F1DFCE05D9024EEA996" box="[870,952,904,929]" class="Arachnida" family="Salticidae" genus="Cucudeta" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="uzet">
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<emphasis id="8980EAD35E071F1DFCE05D9024EEA996" box="[870,952,904,929]" italics="true" pageId="14" pageNumber="15">C. uzet</emphasis>
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</taxonomicName>
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were collected from leaf litter suspended a few centimetres above the ground in the base of small
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<taxonomicName id="7CF44D425E071F1DFCB25DB724FFA9FF" box="[820,937,943,968]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="genus">
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<emphasis id="8980EAD35E071F1DFCB25DB724FFA9FF" box="[820,937,943,968]" italics="true" pageId="14" pageNumber="15">Pandanus</emphasis>
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</taxonomicName>
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plants (
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<figureCitation id="23CF2A445E071F1DFB8E5DB723C4A9FE" box="[1032,1170,943,969]" captionStart="FIGURES 87 – 92" captionStartId="19.[151,270,1328,1352]" captionTargetBox="[151,1434,194,1303]" captionTargetId="figure@19.[151,1436,194,1304]" captionTargetPageId="19" captionText="FIGURES 87 – 92. Habitats of cocalodines. Microhabitat indicated by arrows. 87 Habitat of Allococalodes madidus, on mosses on small tree trunks along stream (Umgé, S 5.3054 E 142.5098); approximate width of field 1.5 m. 88 Habitat of Yamangalea frewana, on tree branches and trunks (Mt. Gahavisuka, S 6.015 E 145.412); approximate diameter of branch indicated 5 cm. Most were found on larger, vertical tree trunks. 89 Habitat of Tabuina varirata, on conifer in clearing (Varirata National Park, S 9.436 E 147.364); approximate width of field 0.7 m. 90 Habitat of Cucudeta zabkai, in leaf litter at base of Pandanus (near Tualapa, S 5.291 E 142.500); approximate width of field 1 m. 91 Habitat of Cucudeta uzet, in leaf litter at base of Pandanus (Umgé, S 5.304 E 142.512); approximate width of field 1 m. 92 Habitat of Cucudeta gahavisuka, in leaf litter on terraces on slope in forest (Mt. Gahavisuka, S 6.01451 E 145.41171); approximate width of field 1.5 m." httpUri="https://zenodo.org/record/186081/files/figure.png" pageId="14" pageNumber="15">Figs 90–91</figureCitation>
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). Later,
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<taxonomicName id="7CF44D425E071F1DFB755DB722CAA9FF" box="[1267,1436,943,968]" class="Arachnida" family="Salticidae" genus="Cucudeta" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="gahavisuka">
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<emphasis id="8980EAD35E071F1DFB755DB722CAA9FF" box="[1267,1436,943,968]" italics="true" pageId="14" pageNumber="15">C. gahavisuka</emphasis>
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</taxonomicName>
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was found by sifting leaf litter in well-drained terraces on a small hillside (
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<figureCitation id="23CF2A445E071F1DFBCB5DCD23FBA9D8" box="[1101,1197,981,1007]" captionStart="FIGURES 87 – 92" captionStartId="19.[151,270,1328,1352]" captionTargetBox="[151,1434,194,1303]" captionTargetId="figure@19.[151,1436,194,1304]" captionTargetPageId="19" captionText="FIGURES 87 – 92. Habitats of cocalodines. Microhabitat indicated by arrows. 87 Habitat of Allococalodes madidus, on mosses on small tree trunks along stream (Umgé, S 5.3054 E 142.5098); approximate width of field 1.5 m. 88 Habitat of Yamangalea frewana, on tree branches and trunks (Mt. Gahavisuka, S 6.015 E 145.412); approximate diameter of branch indicated 5 cm. Most were found on larger, vertical tree trunks. 89 Habitat of Tabuina varirata, on conifer in clearing (Varirata National Park, S 9.436 E 147.364); approximate width of field 0.7 m. 90 Habitat of Cucudeta zabkai, in leaf litter at base of Pandanus (near Tualapa, S 5.291 E 142.500); approximate width of field 1 m. 91 Habitat of Cucudeta uzet, in leaf litter at base of Pandanus (Umgé, S 5.304 E 142.512); approximate width of field 1 m. 92 Habitat of Cucudeta gahavisuka, in leaf litter on terraces on slope in forest (Mt. Gahavisuka, S 6.01451 E 145.41171); approximate width of field 1.5 m." httpUri="https://zenodo.org/record/186081/files/figure.png" pageId="14" pageNumber="15">Fig. 92</figureCitation>
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). It is possible that
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<taxonomicName id="7CF44D425E071F1DFF115DE42646AE22" box="[151,272,1020,1045]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="genus">
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<emphasis id="8980EAD35E071F1DFF115DE42646AE22" box="[151,272,1020,1045]" italics="true" pageId="14" pageNumber="15">Pandanus</emphasis>
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</taxonomicName>
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is not a requirement for
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<taxonomicName id="7CF44D425E071F1DFDC75DE425E3AE22" box="[577,693,1020,1045]" class="Arachnida" family="Salticidae" genus="Cucudeta" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="zabkai">
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<emphasis id="8980EAD35E071F1DFDC75DE425E3AE22" box="[577,693,1020,1045]" italics="true" pageId="14" pageNumber="15">C. zabkai</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="7CF44D425E071F1DFD775DE42410AE22" box="[753,838,1020,1045]" class="Arachnida" family="Salticidae" genus="Cucudeta" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="uzet">
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<emphasis id="8980EAD35E071F1DFD775DE42410AE22" box="[753,838,1020,1045]" italics="true" pageId="14" pageNumber="15">C. uzet</emphasis>
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</taxonomicName>
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, and that the common habitat requirement of the genus is moist but well-drained leaf litter. No individuals were found walking exposed on the surface of the leaf litter.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |