treatments-xml/data/03/F2/87/03F287C94B50FF8FFF34FAE1FE974F93.xml

<document id="D856F3053BCFF59EE2A26AB4579B20C6" ID-CLB-Dataset="46654" ID-DOI="10.5281/zenodo.207593" ID-GBIF-Dataset="edca0207-f631-494d-97f6-aa74c6a199d8" ID-ISSN="1175-5326" ID-Zenodo-Dep="207593" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460369910365" checkinUser="plazi" docAuthor="Fujita, Yoshihisa &amp; Naruse, Tohru" docDate="2011" docId="03F287C94B50FF8FFF34FAE1FE974F93" docLanguage="en" docName="zt02918p038.pdf" docOrigin="Zootaxa 2918" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Catoptrus iejima Fujita &amp; Naruse, 2011, n. sp." docType="treatment" docVersion="11" lastPageNumber="37" masterDocId="FFCBFFB14B51FF86FFA3FFB0FFA74E14" masterDocTitle="Catoptrus iejima, a new species of cavernicolous swimming crab (Crustacea: Brachyura: Portunidae) from a submarine cave at Ie Island, Ryukyu Islands, Japan" masterLastPageNumber="38" masterPageNumber="29" pageNumber="30" updateTime="1698295639478" updateUser="ExternalLinkService">
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<mods:title id="D3C341DF8AEE0FDC1779E0F23B8620D0">Catoptrus iejima, a new species of cavernicolous swimming crab (Crustacea: Brachyura: Portunidae) from a submarine cave at Ie Island, Ryukyu Islands, Japan</mods:title>
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<mods:namePart id="1ECBFFDBFC619C42624539158BC5BDF6">Fujita, Yoshihisa</mods:namePart>
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<mods:namePart id="1E472EB9E59BFA6877D186B4054AF942">Naruse, Tohru</mods:namePart>
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<mods:date id="9759CC53227A771817C57562C6799251">2011</mods:date>
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<treatment id="03F287C94B50FF8FFF34FAE1FE974F93" ID-DOI="http://doi.org/10.5281/zenodo.6191874" ID-GBIF-Taxon="119411597" ID-Zenodo-Dep="6191874" LSID="urn:lsid:plazi:treatment:03F287C94B50FF8FFF34FAE1FE974F93" httpUri="http://treatment.plazi.org/id/03F287C94B50FF8FFF34FAE1FE974F93" lastPageId="9" lastPageNumber="37" pageId="1" pageNumber="30">
<subSubSection id="C34165544B50FF87FF34FAE1FEB54B98" pageId="1" pageNumber="30" type="nomenclature">
<paragraph id="8BE436DF4B50FF87FF34FAE1FE174B7F" blockId="1.[151,432,1361,1420]" box="[151,432,1361,1387]" pageId="1" pageNumber="30">
<heading id="D0AC81B34B50FF87FF34FAE1FE174B7F" bold="true" box="[151,432,1361,1387]" fontSize="11" level="1" pageId="1" pageNumber="30" reason="1">
<emphasis id="B92FEACD4B50FF87FF34FAE1FE174B7F" bold="true" box="[151,432,1361,1387]" pageId="1" pageNumber="30">
<taxonomicName id="4C5B4D5C4B50FF87FF34FAE1FEC74B7F" ID-CoL="RVX3" box="[151,352,1361,1387]" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="30" phylum="Arthropoda" rank="species" species="iejima" status="sp. nov.">
<emphasis id="B92FEACD4B50FF87FF34FAE1FEC74B7F" bold="true" box="[151,352,1361,1387]" italics="true" pageId="1" pageNumber="30">Catoptrus iejima</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A21C57B64B50FF87FECDFAE1FE174B7F" box="[366,432,1361,1387]" pageId="1" pageNumber="30" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE436DF4B50FF87FF34FAC4FEB54B98" blockId="1.[151,432,1361,1420]" box="[151,274,1396,1420]" pageId="1" pageNumber="30">
(
<figureCitation id="13602A5A4B50FF87FF3CFAC4FEAE4B98" box="[159,265,1396,1420]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStartId-0="2.[151,250,1970,1993]" captionStartId-1="3.[151,250,1913,1936]" captionStartId-2="4.[151,250,1949,1972]" captionStartId-3="5.[151,250,1417,1440]" captionStartId-4="6.[151,250,1763,1786]" captionStartId-5="7.[151,250,1817,1840]" captionTargetBox-0="[151,1436,985,1948]" captionTargetBox-1="[245,1341,193,1855]" captionTargetBox-2="[380,1206,193,1928]" captionTargetBox-3="[197,1416,208,1378]" captionTargetBox-4="[151,1436,289,1741]" captionTargetBox-5="[182,1386,324,1770]" captionTargetId-0="figure@2.[151,1436,985,1949]" captionTargetId-1="figure@3.[245,1341,193,1856]" captionTargetId-2="figure@4.[380,1208,193,1929]" captionTargetId-3="figure@5.[151,1436,193,1396]" captionTargetId-4="figure@6.[151,1436,289,1742]" captionTargetId-5="figure@7.[151,1436,301,1797]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionText-0="FIGURE 2. Live colouration of Catoptrus iejima, n. sp., in situ, photo taken by Yusuke Yamada. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317." captionText-1="FIGURE 3. Live colouration of Catoptrus iejima, n. sp. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A, habitus, dorsal view; B, habitus, ventral view." captionText-2="FIGURE 4. Live colouration of Catoptrus iejima, n. sp. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A, carapace, dorsal view; B, cephalothorax, anterior view; C, cephalothorax, ventral view." captionText-3="FIGURE 5. Catoptrus iejima, n. sp. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A, cephalothorax, anterior view; B, right eye, dorsal view; C, cephalothorax, dorsal view; D, left third maxilliped; E, abdomen and telson. Scale bars: A, C, E = 5.0 mm; B, D = 1.0 mm." captionText-4="FIGURE 6. Chelipeds of Catoptrus iejima, n. sp. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A – C, left cheliped; D – F, right cheliped. A, D, upper view; B, E, chela, outer view; C, F, chela, inner view." captionText-5="FIGURE 7. Upper views of right ambulatory legs of Catoptrus iejima, n. sp. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A, E, P 2 (first ambulatory leg = second pereopod) and its dactylus; B, F, P 3 and its dactylus; C, G, P 4 and its dactylus; D, H, P 5 and its dactylus. Scale bars: A – D = 5.0 mm; E – H = 1.0 mm." httpUri-0="https://zenodo.org/record/207595/files/figure.png" httpUri-1="https://zenodo.org/record/207596/files/figure.png" httpUri-2="https://zenodo.org/record/207597/files/figure.png" httpUri-3="https://zenodo.org/record/207598/files/figure.png" httpUri-4="https://zenodo.org/record/207599/files/figure.png" httpUri-5="https://zenodo.org/record/207600/files/figure.png" pageId="1" pageNumber="30">Figs. 2–7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34165544B50FF87FF34FA0CFD10490C" pageId="1" pageNumber="30" type="materials_examined">
<paragraph id="8BE436DF4B50FF87FF34FA0CFE0B4808" blockId="1.[151,1436,1468,2032]" pageId="1" pageNumber="30">
<emphasis id="B92FEACD4B50FF87FF34FA0CFE274BC1" bold="true" box="[151,384,1468,1493]" pageId="1" pageNumber="30">Material examined.</emphasis>
<typeStatus id="54E0887D4B50FF87FE25FA0CFE564BC0" box="[390,497,1468,1492]" pageId="1" pageNumber="30" type="holotype">Holotype</typeStatus>
, female, 21.8 × 
<quantity id="4CA39B3A4B50FF87FD07FA0CFCAD4BC0" box="[676,778,1468,1492]" metricMagnitude="-2" metricUnit="m" metricValue="3.56" pageId="1" pageNumber="30" unit="mm" value="35.6">35.6 mm</quantity>
, RUMF-ZC-1317, point “Ohoba No. 1”, Ie 
<collectingCountry id="F34C764F4B50FF87FB50FA0CFA9D4BC0" box="[1267,1338,1468,1492]" name="Iceland" pageId="1" pageNumber="30">Island</collectingCountry>
, Ryukyu Islands, 
<collectingCountry id="F34C764F4B50FF87FF5FFA51FE984BEC" box="[252,319,1505,1528]" name="Japan" pageId="1" pageNumber="30">Japan</collectingCountry>
, - 
<quantity id="4CA39B3A4B50FF87FEF9FA50FE284BEC" box="[346,399,1504,1528]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="30" unit="m" value="10.0">10m</quantity>
, submarine cave, baited with Pacific saury (
<taxonomicName id="4C5B4D5C4B50FF87FC16FA50FBCD4BEC" box="[949,1130,1504,1528]" class="Actinopterygii" family="Scomberesocidae" genus="Cololabis" kingdom="Animalia" order="Beloniformes" pageId="1" pageNumber="30" phylum="Chordata" rank="species" species="saira">
<emphasis id="B92FEACD4B50FF87FC16FA50FBCD4BEC" box="[949,1130,1504,1528]" italics="true" pageId="1" pageNumber="30">Cololabis saira</emphasis>
</taxonomicName>
), coll. Y. Yamada &amp; K. Yunokawa, 
<date id="FFE5101F4B50FF87FEB8F9B4FE0F4808" box="[283,424,1540,1564]" pageId="1" pageNumber="30" value="2010-07-15">15 Jul. 2010</date>
.
</paragraph>
<paragraph id="8BE436DF4B50FF87FF66F998FD10490C" blockId="1.[151,1436,1468,2032]" pageId="1" pageNumber="30">
<emphasis id="B92FEACD4B50FF87FF66F998FE724855" bold="true" box="[197,469,1576,1601]" pageId="1" pageNumber="30">Comparative material.</emphasis>
<taxonomicName id="4C5B4D5C4B50FF87FE7FF998FC054854" authority="Takeda, 2003: 1" authorityName="Takeda" authorityPageNumber="1" authorityYear="2003" box="[476,930,1576,1600]" class="Malacostraca" family="Portunidae" genus="Atoportunus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="30" phylum="Arthropoda" rank="species" species="dolichopus">
<emphasis id="B92FEACD4B50FF87FE7FF998FD4F4854" box="[476,744,1576,1600]" italics="true" pageId="1" pageNumber="30">Atoportunus dolichopus</emphasis>
Takeda, 2003: 
<specimenCount id="9D5DFD564B50FF87FC37F998FC054854" box="[916,930,1576,1600]" pageId="1" pageNumber="30" type="female">1</specimenCount>
</taxonomicName>
female, 23.9 × 36.0 mm, RUMF-ZC-1318, 
<specimenCount id="9D5DFD564B50FF87FA2DF998FF414870" pageId="1" pageNumber="30" type="female">1 female</specimenCount>
, 20.6 × 
<quantity id="4CA39B3A4B50FF87FE9CF9FCFE034870" box="[319,420,1612,1636]" metricMagnitude="-2" metricUnit="m" metricValue="3.09" pageId="1" pageNumber="30" unit="mm" value="30.9">30.9 mm</quantity>
, 
<collectionCode id="ED4AAE1A4B50FF87FE13F9FCFE414870" box="[432,486,1612,1636]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="30">ZRC</collectionCode>
2011.0014, KUMEJIMA 2009 stn. Diving 38, Hidenchigama, Kume Island, Ryukyu Islands, 
<collectingCountry id="F34C764F4B50FF87FF54F9C1FE9D489C" box="[247,314,1649,1672]" name="Japan" pageId="1" pageNumber="30">Japan</collectingCountry>
, submarine limestone cave, - 
<quantity id="4CA39B3A4B50FF87FD2BF9C0FD55489C" box="[648,754,1648,1672]" metricMagnitude="1" metricUnit="m" metricValue="3.75" metricValueMax="4.0" metricValueMin="3.5" pageId="1" pageNumber="30" unit="m" value="37.5" valueMax="40.0" valueMin="35.0">35–40 m</quantity>
, coll. Y. Fujita, 
<date id="FFE5101F4B50FF87FC0DF9C0FBE9489C" box="[942,1102,1648,1672]" pageId="1" pageNumber="30" value="2009-11-19">19 Nov. 2009</date>
. 
<taxonomicName id="4C5B4D5C4B50FF87FBF8F9C1FA86489C" box="[1115,1313,1648,1672]" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="30" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="B92FEACD4B50FF87FBF8F9C1FA86489C" box="[1115,1313,1648,1672]" italics="true" pageId="1" pageNumber="30">Catoptrus nitidus</emphasis>
</taxonomicName>
A. Milne- 
<bibRefCitation id="EFCA4B2E4B50FF87FF34F924FEE248B8" author="Edwards" box="[151,325,1684,1708]" pageId="1" pageNumber="30" refString="Milne-Edwards, A. (1870) Note sur le Catoptrus nouveau genre appartenant a la division des crustaces brachyoures catometopes. Annales des Sciences naturelles, Zoologie (Paris), ser. 5, 13, 82." type="journal article" year="1870">Edwards, 1870</bibRefCitation>
: 
<specimenCount id="9D5DFD564B50FF87FEF1F924FE0548B8" box="[338,418,1684,1708]" pageId="1" pageNumber="30" type="male">1 male</specimenCount>
, 15.6 × 
<quantity id="4CA39B3A4B50FF87FDA1F924FDCD48B8" box="[514,618,1684,1708]" metricMagnitude="-2" metricUnit="m" metricValue="2.3899999999999997" pageId="1" pageNumber="30" unit="mm" value="23.9">23.9 mm</quantity>
, 
<collectionCode id="ED4AAE1A4B50FF87FDDBF925FD6A48B8" LSID="urn:lsid:biocol.org:col:34988" box="[632,717,1685,1708]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="30">MNHN</collectionCode>
<collectionCode id="ED4AAE1A4B50FF87FD74F924FD5548B8" box="[727,754,1684,1708]" pageId="1" pageNumber="30">B-</collectionCode>
4639, Iles 
<collectingCountry id="F34C764F4B50FF87FCCFF924FC3E48B8" box="[876,921,1684,1708]" name="Fiji" pageId="1" pageNumber="30">Viti</collectingCountry>
; 
<specimenCount id="9D5DFD564B50FF87FC04F924FBA548B8" box="[935,1026,1684,1708]" pageId="1" pageNumber="30" type="male">2 males</specimenCount>
, 7.8 × 
<quantity id="4CA39B3A4B50FF87FBF6F924FB1A48B8" box="[1109,1213,1684,1708]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="1" pageNumber="30" unit="mm" value="12.1">12.1 mm</quantity>
, 6.5 x ca. 
<quantity id="4CA39B3A4B50FF87FA95F924FA3B48B8" box="[1334,1436,1684,1708]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="1" pageNumber="30" unit="mm" value="10.1">10.1 mm</quantity>
(damaged), 
<specimenCount id="9D5DFD564B50FF87FE83F908FE3048C4" box="[288,407,1720,1744]" pageId="1" pageNumber="30" type="juvenile">1 juvenile</specimenCount>
, 4.2 x 
<quantity id="4CA39B3A4B50FF87FE4FF908FDEF48C4" box="[492,584,1720,1744]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="1" pageNumber="30" unit="mm" value="6.4">6.4 mm</quantity>
, 
<collectionCode id="ED4AAE1A4B50FF87FDFBF909FD0948C4" LSID="urn:lsid:biocol.org:col:34988" box="[600,686,1721,1744]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="30">MNHN</collectionCode>
<collectionCode id="ED4AAE1A4B50FF87FD1AF908FD7348C4" box="[697,724,1720,1744]" pageId="1" pageNumber="30">B-</collectionCode>
22342, 
<collectingCountry id="F34C764F4B50FF87FC8EF908FCDA48C4" box="[813,893,1720,1744]" name="Samoa" pageId="1" pageNumber="30">Samoa</collectingCountry>
. 
<taxonomicName id="4C5B4D5C4B50FF87FC28F909FAB948C4" authority="Serene, 1966" authorityName="Serene" authorityYear="1966" box="[907,1310,1720,1744]" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="30" phylum="Arthropoda" rank="species" species="rathbunae">
<emphasis id="B92FEACD4B50FF87FC28F909FBDE48C4" box="[907,1145,1720,1744]" italics="true" pageId="1" pageNumber="30">Catoptrus rathbunae</emphasis>
<bibRefCitation id="EFCA4B2E4B50FF87FB27F908FAB948C4" author="Serene" box="[1156,1310,1720,1744]" pageId="1" pageNumber="30" refString="Serene, R. (1966) Notes sur les genres Catoptrus et Lybistes et les Catoptrinae. Bulletin du Museum national d'Histoire naturelle, (2) 37 (6), 929 - 1000." type="journal article" year="1966">Serène, 1966</bibRefCitation>
</taxonomicName>
: 
<typeStatus id="54E0887D4B50FF87FA8EF908FA3F48C4" box="[1325,1432,1720,1744]" pageId="1" pageNumber="30" type="holotype">Holotype</typeStatus>
, male, 5.0 × 7.0 mm, 
<collectionCode id="ED4AAE1A4B50FF87FE21F96DFE7F48E0" LSID="urn:lsid:biocol.org:col:34988" box="[386,472,1757,1780]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="30">MNHN</collectionCode>
<collectionCode id="ED4AAE1A4B50FF87FE7CF96CFE5D48E0" box="[479,506,1756,1780]" pageId="1" pageNumber="30">B-</collectionCode>
5564, Nhatrang, 
<collectingCountry id="F34C764F4B50FF87FD1BF96CFCBD48E0" box="[696,794,1756,1780]" name="Vietnam" pageId="1" pageNumber="30">Vietnam</collectingCountry>
, coll. R. Serène, 
<date id="FFE5101F4B50FF87FC7AF96CFBD148E0" box="[985,1142,1756,1781]" pageId="1" pageNumber="30" value="1964-08-14">14 Aug. 1964</date>
; 
<typeStatus id="54E0887D4B50FF87FB21F96DFB4248E1" box="[1154,1253,1757,1781]" pageId="1" pageNumber="30" type="paratype">paratype</typeStatus>
, 
<specimenCount id="9D5DFD564B50FF87FB51F96CFAF148E0" box="[1266,1366,1756,1780]" pageId="1" pageNumber="30" type="female">1 female</specimenCount>
, 6.0 × 
<quantity id="4CA39B3A4B50FF87FF34F8B0FF48490C" box="[151,239,1792,1816]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="1" pageNumber="30" unit="mm" value="8.5">8.5 mm</quantity>
, 
<collectionCode id="ED4AAE1A4B50FF87FF59F8B1FEF7490C" LSID="urn:lsid:biocol.org:col:34988" box="[250,336,1793,1816]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="30">MNHN</collectionCode>
<collectionCode id="ED4AAE1A4B50FF87FEF4F8B0FED4490C" box="[343,371,1792,1816]" pageId="1" pageNumber="30">B-</collectionCode>
5563, same data as 
<typeStatus id="54E0887D4B50FF87FDEDF8B0FD14490C" box="[590,691,1792,1816]" pageId="1" pageNumber="30" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C34165544B50FF84FF66F894FD554EE3" lastPageId="2" lastPageNumber="31" pageId="1" pageNumber="30" type="diagnosis">
<paragraph id="8BE436DF4B50FF84FF66F894FD554EE3" blockId="1.[151,1436,1468,2032]" lastBlockId="2.[151,1437,151,932]" lastPageId="2" lastPageNumber="31" pageId="1" pageNumber="30">
<emphasis id="B92FEACD4B50FF87FF66F894FE984929" bold="true" box="[197,319,1828,1853]" pageId="1" pageNumber="30">Diagnosis.</emphasis>
Carapace ovoid, longitudinally, transversely convex. Supraorbital margin beaded on convex part, without fissure; infraorbital margin discontinuous. External orbital angle dorso-ventrally flattened, distally rounded, continuous to anterolateral margin. Anterolateral margins with 5 teeth excluding external orbital angle, first and second teeth, third and fourth teeth placed closer to each other, fifth tooth largest; margins between bases of teeth lined with granules, forming ridge-like structure. Eyes small, not completely occupying orbit. Chelipeds slender; anterior margin of merus with 1 subproximal small tooth, 1 subdistal small tooth; palm swollen with no striae or granules; fingers slender, as long as palm, distally hooked, 3–4 long sharp teeth and few small sharp teeth between them. Ambulatory legs slender, long; propodus of P5 with subparallel inner, outer margins, slender; P5 dactylus lanceolate, dactylus narrower than propodus.
</paragraph>
</subSubSection>
<subSubSection id="C34165544B53FF80FF66FEB3FC494EE3" lastPageId="6" lastPageNumber="35" pageId="2" pageNumber="31" type="description">
<paragraph id="8BE436DF4B53FF84FF66FEB3FD504C4B" blockId="2.[151,1437,151,932]" pageId="2" pageNumber="31">
<emphasis id="B92FEACD4B53FF84FF66FEB3FEF14F08" bold="true" box="[197,342,259,284]" pageId="2" pageNumber="31">Description.</emphasis>
Carapace ovoid, longitudinally, transversely convex, dorsal surface smooth, regions poorly defined (
<figureCitation id="13602A5A4B53FF84FF59FE97FEEE4F2B" box="[250,329,295,319]" captionStart="FIGURE 2" captionStartId="2.[151,250,1970,1993]" captionTargetBox="[151,1436,985,1948]" captionTargetId="figure@2.[151,1436,985,1949]" captionTargetPageId="2" captionText="FIGURE 2. Live colouration of Catoptrus iejima, n. sp., in situ, photo taken by Yusuke Yamada. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317." httpUri="https://zenodo.org/record/207595/files/figure.png" pageId="2" pageNumber="31">Figs. 2</figureCitation>
, 
<figureCitation id="13602A5A4B53FF84FEF6FE97FEC54F2B" box="[341,354,295,319]" captionStart="FIGURE 3" captionStartId="3.[151,250,1913,1936]" captionTargetBox="[245,1341,193,1855]" captionTargetId="figure@3.[245,1341,193,1856]" captionTargetPageId="3" captionText="FIGURE 3. Live colouration of Catoptrus iejima, n. sp. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A, habitus, dorsal view; B, habitus, ventral view." httpUri="https://zenodo.org/record/207596/files/figure.png" pageId="2" pageNumber="31">3</figureCitation>
A, 4A, B, 5A). Front bilobed, with small median notch, frontal margin straight, lined with 2 parallel rows of granules; supraorbital margin beaded on convex part, without fissure; infraorbital margin discontinuous, inner orbital lobe dorso-ventrally flattened, produced anteriorly, lobe followed laterally by few, small, granules, one relatively large granule, not continuous to external orbital angle (
<figureCitation id="13602A5A4B53FF84FBA6FE24FBF44FB8" box="[1029,1107,404,428]" captionStart="FIGURE 4" captionStartId="4.[151,250,1949,1972]" captionTargetBox="[380,1206,193,1928]" captionTargetId="figure@4.[380,1208,193,1929]" captionTargetPageId="4" captionText="FIGURE 4. Live colouration of Catoptrus iejima, n. sp. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A, carapace, dorsal view; B, cephalothorax, anterior view; C, cephalothorax, ventral view." httpUri="https://zenodo.org/record/207597/files/figure.png" pageId="2" pageNumber="31">Figs. 4</figureCitation>
B, 5A). External orbital angle dorso-ventrally flattened, distally rounded, reaching slightly beyond level of frontal margin (
<figureCitation id="13602A5A4B53FF84FB0EFE07FB574FDB" box="[1197,1264,439,463]" captionStart="FIGURE 5" captionStartId="5.[151,250,1417,1440]" captionTargetBox="[197,1416,208,1378]" captionTargetId="figure@5.[151,1436,193,1396]" captionTargetPageId="5" captionText="FIGURE 5. Catoptrus iejima, n. sp. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A, cephalothorax, anterior view; B, right eye, dorsal view; C, cephalothorax, dorsal view; D, left third maxilliped; E, abdomen and telson. Scale bars: A, C, E = 5.0 mm; B, D = 1.0 mm." httpUri="https://zenodo.org/record/207598/files/figure.png" pageId="2" pageNumber="31">Fig. 5</figureCitation>
A, C), continuous to anterolateral margin. Antero- and posterolateral margins (
<figureCitation id="13602A5A4B53FF84FCCCFE6CFC1B4FE0" box="[879,956,476,500]" captionStart="FIGURE 3" captionStartId="3.[151,250,1913,1936]" captionTargetBox="[245,1341,193,1855]" captionTargetId="figure@3.[245,1341,193,1856]" captionTargetPageId="3" captionText="FIGURE 3. Live colouration of Catoptrus iejima, n. sp. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A, habitus, dorsal view; B, habitus, ventral view." httpUri="https://zenodo.org/record/207596/files/figure.png" pageId="2" pageNumber="31">Figs. 3</figureCitation>
A, 4A, 5C) almost of same length; anterolateral margin with 5 teeth excluding external orbital angle, first and second teeth, third and fourth teeth placed closer to each other, fifth tooth largest, first to fourth teeth small, almost of same size; margins between bases of teeth lined with granules, forming ridge-like structure.
</paragraph>
<paragraph id="8BE436DF4B53FF84FF66FDDCFD3C4CB3" blockId="2.[151,1437,151,932]" pageId="2" pageNumber="31">
Epistome (
<figureCitation id="13602A5A4B53FF84FE9CFDDCFE254C90" box="[319,386,620,644]" captionStart="FIGURE 5" captionStartId="5.[151,250,1417,1440]" captionTargetBox="[197,1416,208,1378]" captionTargetId="figure@5.[151,1436,193,1396]" captionTargetPageId="5" captionText="FIGURE 5. Catoptrus iejima, n. sp. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A, cephalothorax, anterior view; B, right eye, dorsal view; C, cephalothorax, dorsal view; D, left third maxilliped; E, abdomen and telson. Scale bars: A, C, E = 5.0 mm; B, D = 1.0 mm." httpUri="https://zenodo.org/record/207598/files/figure.png" pageId="2" pageNumber="31">Fig. 5</figureCitation>
A) with posterior margin medially produced, produced part medially notched, posterior margin rimmed, beaded, except for median notch.
</paragraph>
<paragraph id="8BE436DF4B53FF84FF66FD05FEE44DB0" blockId="2.[151,1437,151,932]" pageId="2" pageNumber="31">
Eyes small, not completely occupying orbit, slightly mobile; eyestalk with L-shaped prominence along proximal margin of cornea of anterior surface to concave part of cornea on upper surface; cornea small, width as long as length (
<figureCitation id="13602A5A4B53FF84FF4EFD4CFE9A4D00" box="[237,317,764,788]" captionStart="FIGURE 4" captionStartId="4.[151,250,1949,1972]" captionTargetBox="[380,1206,193,1928]" captionTargetId="figure@4.[380,1208,193,1929]" captionTargetPageId="4" captionText="FIGURE 4. Live colouration of Catoptrus iejima, n. sp. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A, carapace, dorsal view; B, cephalothorax, anterior view; C, cephalothorax, ventral view." httpUri="https://zenodo.org/record/207597/files/figure.png" pageId="2" pageNumber="31">Figs. 4</figureCitation>
B, 5A, B). Distomesial angle of basal antennal segment continuous with ventral extension of distolateral angle of front (
<figureCitation id="13602A5A4B53FF84FE36FCAFFE7D4D23" box="[405,474,799,823]" captionStart="FIGURE 5" captionStartId="5.[151,250,1417,1440]" captionTargetBox="[197,1416,208,1378]" captionTargetId="figure@5.[151,1436,193,1396]" captionTargetPageId="5" captionText="FIGURE 5. Catoptrus iejima, n. sp. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A, cephalothorax, anterior view; B, right eye, dorsal view; C, cephalothorax, dorsal view; D, left third maxilliped; E, abdomen and telson. Scale bars: A, C, E = 5.0 mm; B, D = 1.0 mm." httpUri="https://zenodo.org/record/207598/files/figure.png" pageId="2" pageNumber="31">Fig. 5</figureCitation>
A). First maxilliped with distinct “portunoid lobe”. Third maxilliped (
<figureCitation id="13602A5A4B53FF84FB5DFCAFFAE54D23" box="[1278,1346,799,823]" captionStart="FIGURE 5" captionStartId="5.[151,250,1417,1440]" captionTargetBox="[197,1416,208,1378]" captionTargetId="figure@5.[151,1436,193,1396]" captionTargetPageId="5" captionText="FIGURE 5. Catoptrus iejima, n. sp. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A, cephalothorax, anterior view; B, right eye, dorsal view; C, cephalothorax, dorsal view; D, left third maxilliped; E, abdomen and telson. Scale bars: A, C, E = 5.0 mm; B, D = 1.0 mm." httpUri="https://zenodo.org/record/207598/files/figure.png" pageId="2" pageNumber="31">Fig. 5</figureCitation>
D) with ischium to merus semi-rectangular; ischium with median sinuous groove, distal margin sinuous, with produced distomesial angle, mid-length about 2 times that of merus; merus with distolateral angle almost right angle, dorso-ventrally flattened.
</paragraph>
<caption id="DF2466574B53FF84FF34F802FED949F3" httpUri="https://zenodo.org/record/207595/files/figure.png" pageId="2" pageNumber="31" targetBox="[151,1436,985,1948]" targetPageId="2">
<paragraph id="8BE436DF4B53FF84FF34F802FED949F3" blockId="2.[151,1436,1970,2023]" pageId="2" pageNumber="31">
<emphasis id="B92FEACD4B53FF84FF34F802FEB549DC" bold="true" box="[151,274,1970,1993]" pageId="2" pageNumber="31">FIGURE 2.</emphasis>
Live colouration of 
<taxonomicName id="4C5B4D5C4B53FF84FE43F803FD2D49DC" box="[480,650,1971,1992]" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="31" phylum="Arthropoda" rank="species" species="iejima" status="sp. nov.">
<emphasis id="B92FEACD4B53FF84FE43F803FD2D49DC" box="[480,650,1971,1992]" italics="true" pageId="2" pageNumber="31">Catoptrus iejima</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A21C57B64B53FF84FD35F803FD6849DC" box="[662,719,1971,1992]" pageId="2" pageNumber="31" rank="species">
<emphasis id="B92FEACD4B53FF84FD35F803FD6849DC" bold="true" box="[662,719,1971,1992]" pageId="2" pageNumber="31">n. sp.</emphasis>
</taxonomicNameLabel>
, in situ, photo taken by Yusuke Yamada. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC-1317.
</paragraph>
</caption>
<caption id="DF2466574B52FF85FF34F8C9FE4849BA" httpUri="https://zenodo.org/record/207596/files/figure.png" pageId="3" pageNumber="32" targetBox="[245,1341,193,1855]" targetPageId="3">
<paragraph id="8BE436DF4B52FF85FF34F8C9FE4849BA" blockId="3.[151,1436,1913,1966]" pageId="3" pageNumber="32">
<emphasis id="B92FEACD4B52FF85FF34F8C9FEB5499B" bold="true" box="[151,274,1913,1936]" pageId="3" pageNumber="32">FIGURE 3.</emphasis>
Live colouration of 
<taxonomicName id="4C5B4D5C4B52FF85FE40F8CBFD2A4984" box="[483,653,1915,1936]" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="iejima" status="sp. nov.">
<emphasis id="B92FEACD4B52FF85FE40F8CBFD2A4984" box="[483,653,1915,1936]" italics="true" pageId="3" pageNumber="32">Catoptrus iejima</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A21C57B64B52FF85FD38F8CAFD73499B" box="[667,724,1914,1935]" pageId="3" pageNumber="32" rank="species">
<emphasis id="B92FEACD4B52FF85FD38F8CAFD73499B" bold="true" box="[667,724,1914,1935]" pageId="3" pageNumber="32">n. sp.</emphasis>
</taxonomicNameLabel>
Holotype, female, 21.8 × 35.6 mm, RUMF-ZC-1317. A, habitus, dorsal view; B, habitus, ventral view.
</paragraph>
</caption>
<caption id="DF2466574B55FF82FF34F82DFC0349C6" httpUri="https://zenodo.org/record/207597/files/figure.png" pageId="4" pageNumber="33" targetBox="[380,1206,193,1928]" targetPageId="4">
<paragraph id="8BE436DF4B55FF82FF34F82DFC0349C6" blockId="4.[151,1436,1949,2002]" pageId="4" pageNumber="33">
<emphasis id="B92FEACD4B55FF82FF34F82DFEB349A7" bold="true" box="[151,276,1949,1972]" pageId="4" pageNumber="33">FIGURE 4.</emphasis>
Live colouration of 
<taxonomicName id="4C5B4D5C4B55FF82FE4FF82FFD3E49A0" box="[492,665,1951,1972]" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="iejima" status="sp. nov.">
<emphasis id="B92FEACD4B55FF82FE4FF82FFD3E49A0" box="[492,665,1951,1972]" italics="true" pageId="4" pageNumber="33">Catoptrus iejima</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A21C57B64B55FF82FD0BF82EFD4449A7" box="[680,739,1950,1971]" pageId="4" pageNumber="33" rank="species">
<emphasis id="B92FEACD4B55FF82FD0BF82EFD4449A7" bold="true" box="[680,739,1950,1971]" pageId="4" pageNumber="33">n. sp.</emphasis>
</taxonomicNameLabel>
Holotype, female, 21.8 × 35.6 mm, RUMF-ZC-1317. A, carapace, dorsal view; B, cephalothorax, anterior view; C, cephalothorax, ventral view.
</paragraph>
</caption>
<caption id="DF2466574B54FF83FF34FA39FECC4BC9" httpUri="https://zenodo.org/record/207598/files/figure.png" pageId="5" pageNumber="34" targetBox="[197,1416,208,1378]" targetPageId="5">
<paragraph id="8BE436DF4B54FF83FF34FA39FECC4BC9" blockId="5.[151,1435,1417,1501]" pageId="5" pageNumber="34">
<emphasis id="B92FEACD4B54FF83FF34FA39FEB44B8B" bold="true" box="[151,275,1417,1440]" pageId="5" pageNumber="34">FIGURE 5.</emphasis>
<taxonomicName id="4C5B4D5C4B54FF83FEBAFA3BFE634BB4" box="[281,452,1419,1440]" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="iejima" status="sp. nov.">
<emphasis id="B92FEACD4B54FF83FEBAFA3BFE634BB4" box="[281,452,1419,1440]" italics="true" pageId="5" pageNumber="34">Catoptrus iejima</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A21C57B64B54FF83FE72FA3AFDAD4B8B" box="[465,522,1418,1439]" pageId="5" pageNumber="34" rank="species">
<emphasis id="B92FEACD4B54FF83FE72FA3AFDAD4B8B" bold="true" box="[465,522,1418,1439]" pageId="5" pageNumber="34">n. sp.</emphasis>
</taxonomicNameLabel>
Holotype, female, 21.8 × 35.6 mm, RUMF-ZC-1317. A, cephalothorax, anterior view; B, right eye, dorsal view; C, cephalothorax, dorsal view; D, left third maxilliped; E, abdomen and telson. Scale bars: A, C, E = 5.0 mm; B, D = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BE436DF4B54FF83FF66F9B9FA31489A" blockId="5.[151,1436,1545,2001]" pageId="5" pageNumber="34">Thoracic sternites 3, 4 weakly divided by lateral short grooves; suture of thoracic sternites 4, 5 mesially end in elongated slit; vulva transversely elongated, longitudinally placed about middle of sternite 6; suture between sternites 6, 7 reaching to below inner end of vulva, but not connected to posterior margin of transverse median “ridge” connects sternites; median longitudinal groove between sternites 7, 8 reaching as far as median “ridge” near vulva.</paragraph>
<paragraph id="8BE436DF4B54FF83FF66F929FB40499D" blockId="5.[151,1436,1545,2001]" pageId="5" pageNumber="34">
Chelipeds slender, longer than ambulatory legs (
<figureCitation id="13602A5A4B54FF83FD42F929FC9748A5" box="[737,816,1689,1713]" captionStart="FIGURE 2" captionStartId="2.[151,250,1970,1993]" captionTargetBox="[151,1436,985,1948]" captionTargetId="figure@2.[151,1436,985,1949]" captionTargetPageId="2" captionText="FIGURE 2. Live colouration of Catoptrus iejima, n. sp., in situ, photo taken by Yusuke Yamada. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317." httpUri="https://zenodo.org/record/207595/files/figure.png" pageId="5" pageNumber="34">Figs. 2</figureCitation>
, 
<figureCitation id="13602A5A4B54FF83FC9FF929FCEC48A5" box="[828,843,1689,1713]" captionStart="FIGURE 3" captionStartId="3.[151,250,1913,1936]" captionTargetBox="[245,1341,193,1855]" captionTargetId="figure@3.[245,1341,193,1856]" captionTargetPageId="3" captionText="FIGURE 3. Live colouration of Catoptrus iejima, n. sp. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A, habitus, dorsal view; B, habitus, ventral view." httpUri="https://zenodo.org/record/207596/files/figure.png" pageId="5" pageNumber="34">3</figureCitation>
); merus (
<figureCitation id="13602A5A4B54FF83FC16F929FBA348A5" box="[949,1028,1689,1713]" captionStart="FIGURE 3" captionStartId="3.[151,250,1913,1936]" captionTargetBox="[245,1341,193,1855]" captionTargetId="figure@3.[245,1341,193,1856]" captionTargetPageId="3" captionText="FIGURE 3. Live colouration of Catoptrus iejima, n. sp. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A, habitus, dorsal view; B, habitus, ventral view." httpUri="https://zenodo.org/record/207596/files/figure.png" pageId="5" pageNumber="34">Figs. 3</figureCitation>
, 
<figureCitation id="13602A5A4B54FF83FBB3F929FBB848A5" box="[1040,1055,1689,1713]" captionStart="FIGURE 6" captionStartId="6.[151,250,1763,1786]" captionTargetBox="[151,1436,289,1741]" captionTargetId="figure@6.[151,1436,289,1742]" captionTargetPageId="6" captionText="FIGURE 6. Chelipeds of Catoptrus iejima, n. sp. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A – C, left cheliped; D – F, right cheliped. A, D, upper view; B, E, chela, outer view; C, F, chela, inner view." httpUri="https://zenodo.org/record/207599/files/figure.png" pageId="5" pageNumber="34">6</figureCitation>
) slightly shorter than chela, subovate in cross section with slightly concave lower surface, anterior margin with 1 subproximal small tooth, 1 subdistal small tooth; left chela (
<figureCitation id="13602A5A4B54FF83FE61F951FDA348ED" box="[450,516,1761,1785]" captionStart="FIGURE 6" captionStartId="6.[151,250,1763,1786]" captionTargetBox="[151,1436,289,1741]" captionTargetId="figure@6.[151,1436,289,1742]" captionTargetPageId="6" captionText="FIGURE 6. Chelipeds of Catoptrus iejima, n. sp. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A – C, left cheliped; D – F, right cheliped. A, D, upper view; B, E, chela, outer view; C, F, chela, inner view." httpUri="https://zenodo.org/record/207599/files/figure.png" pageId="5" pageNumber="34">Fig. 6</figureCitation>
A–C) relatively stouter; chela with swollen palm, surfaces smooth, with no striae or granules; fingers slender, as long as palm, distally hooked, 3–4 long sharp teeth and few small sharp teeth between them along cutting edges which interdigitate. Ambulatory legs (
<figureCitation id="13602A5A4B54FF83FC61F899FBB64955" box="[962,1041,1833,1857]" captionStart="FIGURE 3" captionStartId="3.[151,250,1913,1936]" captionTargetBox="[245,1341,193,1855]" captionTargetId="figure@3.[245,1341,193,1856]" captionTargetPageId="3" captionText="FIGURE 3. Live colouration of Catoptrus iejima, n. sp. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A, habitus, dorsal view; B, habitus, ventral view." httpUri="https://zenodo.org/record/207596/files/figure.png" pageId="5" pageNumber="34">Figs. 3</figureCitation>
, 
<figureCitation id="13602A5A4B54FF83FBBFF899FB8C4955" box="[1052,1067,1833,1857]" captionStart="FIGURE 7" captionStartId="7.[151,250,1817,1840]" captionTargetBox="[182,1386,324,1770]" captionTargetId="figure@7.[151,1436,301,1797]" captionTargetPageId="7" captionText="FIGURE 7. Upper views of right ambulatory legs of Catoptrus iejima, n. sp. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A, E, P 2 (first ambulatory leg = second pereopod) and its dactylus; B, F, P 3 and its dactylus; C, G, P 4 and its dactylus; D, H, P 5 and its dactylus. Scale bars: A – D = 5.0 mm; E – H = 1.0 mm." httpUri="https://zenodo.org/record/207600/files/figure.png" pageId="5" pageNumber="34">7</figureCitation>
) slender, P3 longest, P5 shortest, with long, fine setae, ratios of lengths of P2–P5 meri to CL 0.98, 1.03, 0.93, 0.67 respectively; propodus of P5 with subparallel inner, outer margins, slender; P5 dactylus lanceolate, dactylus narrower than propodus.
</paragraph>
<paragraph id="8BE436DF4B54FF83FF66F826FD5149C5" blockId="5.[151,1436,1545,2001]" pageId="5" pageNumber="34">
All abdominal somites (
<figureCitation id="13602A5A4B54FF83FE74F826FDBD49BA" box="[471,538,1942,1966]" captionStart="FIGURE 5" captionStartId="5.[151,250,1417,1440]" captionTargetBox="[197,1416,208,1378]" captionTargetId="figure@5.[151,1436,193,1396]" captionTargetPageId="5" captionText="FIGURE 5. Catoptrus iejima, n. sp. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A, cephalothorax, anterior view; B, right eye, dorsal view; C, cephalothorax, dorsal view; D, left third maxilliped; E, abdomen and telson. Scale bars: A, C, E = 5.0 mm; B, D = 1.0 mm." httpUri="https://zenodo.org/record/207598/files/figure.png" pageId="5" pageNumber="34">Fig. 5</figureCitation>
E) free, somite 4 to telson triangular in shape; telson longer than broad, lateral margins slightly concave; pleopods setose, developed.
</paragraph>
<paragraph id="8BE436DF4B57FF80FF66FF27FC494EE3" blockId="6.[151,1436,151,247]" pageId="6" pageNumber="35">
<emphasis id="B92FEACD4B57FF80FF66FF27FEEA4EA4" bold="true" box="[197,333,151,176]" pageId="6" pageNumber="35">Coloration.</emphasis>
Dorsal surfaces of carapace and chelipeds orange, except for the tip of the fifth anterolateral tooth, tips of two teeth on the anterior margin of cheliped merus, and occlusal margins and about distal third of cheliped fingers; ventral surfaces and ambulatory legs yellowish-white (
<figureCitation id="13602A5A4B57FF80FCFAFF6FFC624EE3" box="[857,965,223,247]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="2.[151,250,1970,1993]" captionStartId-1="3.[151,250,1913,1936]" captionStartId-2="4.[151,250,1949,1972]" captionTargetBox-0="[151,1436,985,1948]" captionTargetBox-1="[245,1341,193,1855]" captionTargetBox-2="[380,1206,193,1928]" captionTargetId-0="figure@2.[151,1436,985,1949]" captionTargetId-1="figure@3.[245,1341,193,1856]" captionTargetId-2="figure@4.[380,1208,193,1929]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 2. Live colouration of Catoptrus iejima, n. sp., in situ, photo taken by Yusuke Yamada. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317." captionText-1="FIGURE 3. Live colouration of Catoptrus iejima, n. sp. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A, habitus, dorsal view; B, habitus, ventral view." captionText-2="FIGURE 4. Live colouration of Catoptrus iejima, n. sp. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A, carapace, dorsal view; B, cephalothorax, anterior view; C, cephalothorax, ventral view." httpUri-0="https://zenodo.org/record/207595/files/figure.png" httpUri-1="https://zenodo.org/record/207596/files/figure.png" httpUri-2="https://zenodo.org/record/207597/files/figure.png" pageId="6" pageNumber="35">Figs. 2–4</figureCitation>
, 
<figureCitation id="13602A5A4B57FF80FC73FF6FFC464EE3" box="[976,993,223,247]" captionStart="FIGURE 6" captionStartId="6.[151,250,1763,1786]" captionTargetBox="[151,1436,289,1741]" captionTargetId="figure@6.[151,1436,289,1742]" captionTargetPageId="6" captionText="FIGURE 6. Chelipeds of Catoptrus iejima, n. sp. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A – C, left cheliped; D – F, right cheliped. A, D, upper view; B, E, chela, outer view; C, F, chela, inner view." httpUri="https://zenodo.org/record/207599/files/figure.png" pageId="6" pageNumber="35">6</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF2466574B57FF80FF34F953FC44490C" httpUri="https://zenodo.org/record/207599/files/figure.png" pageId="6" pageNumber="35" targetBox="[151,1436,289,1741]" targetPageId="6">
<paragraph id="8BE436DF4B57FF80FF34F953FC44490C" blockId="6.[151,1436,1763,1817]" pageId="6" pageNumber="35">
<emphasis id="B92FEACD4B57FF80FF34F953FEB548ED" bold="true" box="[151,274,1763,1786]" pageId="6" pageNumber="35">FIGURE 6.</emphasis>
Chelipeds of 
<taxonomicName id="4C5B4D5C4B57FF80FE3CF955FDEE48EE" box="[415,585,1765,1786]" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="iejima" status="sp. nov.">
<emphasis id="B92FEACD4B57FF80FE3CF955FDEE48EE" box="[415,585,1765,1786]" italics="true" pageId="6" pageNumber="35">Catoptrus iejima</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A21C57B64B57FF80FDF5F954FD2848ED" box="[598,655,1764,1785]" pageId="6" pageNumber="35" rank="species">
<emphasis id="B92FEACD4B57FF80FDF5F954FD2848ED" bold="true" box="[598,655,1764,1785]" pageId="6" pageNumber="35">n. sp.</emphasis>
</taxonomicNameLabel>
Holotype, female, 21.8 × 35.6 mm, RUMF-ZC-1317. A–C, left cheliped; D– F, right cheliped. A, D, upper view; B, E, chela, outer view; C, F, chela, inner view.
</paragraph>
</caption>
<subSubSection id="C34165544B57FF81FF66F8F5FBDC4EE3" lastPageId="7" lastPageNumber="36" pageId="6" pageNumber="35" type="etymology">
<paragraph id="8BE436DF4B57FF80FF66F8F5FD0F4995" blockId="6.[151,1437,1861,2030]" pageId="6" pageNumber="35">
<emphasis id="B92FEACD4B57FF80FF66F8F5FEEC4949" bold="true" box="[197,331,1861,1885]" pageId="6" pageNumber="35">Etymology.</emphasis>
The new species is named after the 
<taxonomicName id="4C5B4D5C4B57FF80FD7DF8F5FC824949" box="[734,805,1861,1885]" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="38" phylum="Arthropoda" rank="species" species="iejima" status="sp. nov.">Iejima</taxonomicName>
(= Ie 
<collectingCountry id="F34C764F4B57FF80FCCBF8F5FC164949" box="[872,945,1861,1885]" name="Iceland" pageId="6" pageNumber="35">Island</collectingCountry>
), where the 
<typeStatus id="54E0887D4B57FF80FB9BF8F5FB3D4949" box="[1080,1178,1861,1885]" pageId="6" pageNumber="35" type="holotype">holotype</typeStatus>
of the species was collected. The name is used as noun in apposition.
</paragraph>
<paragraph id="8BE436DF4B57FF81FF66F83DFBDC4EE3" blockId="6.[151,1437,1861,2030]" lastBlockId="7.[151,1436,151,247]" lastPageId="7" lastPageNumber="36" pageId="6" pageNumber="35">
<emphasis id="B92FEACD4B57FF80FF66F83DFE2449B1" bold="true" box="[197,387,1933,1958]" pageId="6" pageNumber="35">Common name.</emphasis>
The standard Japanese name for this species is given here as Iejima-gama-gazami. 
<emphasis id="B92FEACD4B57FF80FF66F801FE2349DE" bold="true" box="[197,388,1969,1994]" pageId="6" pageNumber="35">Ecological note.</emphasis>
<taxonomicName id="4C5B4D5C4B57FF80FE2DF802FDE949DD" box="[398,590,1970,1993]" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="iejima" status="sp. nov.">
<emphasis id="B92FEACD4B57FF80FE2DF802FDE949DD" box="[398,590,1970,1993]" italics="true" pageId="6" pageNumber="35">Catoptrus iejima</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A21C57B64B57FF80FDFCF802FD0649DE" box="[607,673,1970,1994]" pageId="6" pageNumber="35" rank="species">
<emphasis id="B92FEACD4B57FF80FDFCF802FD0649DE" bold="true" box="[607,673,1970,1994]" pageId="6" pageNumber="35">n. sp.</emphasis>
</taxonomicNameLabel>
, was collected from a submarine cave with depth of about 
<specimenCount id="9D5DFD564B57FF80FAFAF801FA3B49DD" box="[1369,1436,1969,1993]" pageId="6" pageNumber="35" type="generic">
<quantity id="4CA39B3A4B57FF80FAFAF801FA3249DD" box="[1369,1429,1969,1993]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="35" unit="m" value="10.0">10 m</quantity>
.
</specimenCount>
When the collectors checked the set trap baited with Pacific saury (
<taxonomicName id="4C5B4D5C4B57FF80FC34F866FBE249FA" box="[919,1093,2006,2030]" class="Actinopterygii" family="Scomberesocidae" genus="Cololabis" kingdom="Animalia" order="Beloniformes" pageId="6" pageNumber="35" phylum="Chordata" rank="species" species="saira">
<emphasis id="B92FEACD4B57FF80FC34F866FBE249FA" box="[919,1093,2006,2030]" italics="true" pageId="6" pageNumber="35">Cololabis saira</emphasis>
</taxonomicName>
; “Sanma” in Japanese), about five individuals of this species, including a very small one, were observed to be wandering outside the trap. The crabs observed all appear to move very slowly. Although no individual was actually trapped, one female was caught by hand from outside the trap (Y. Yamada &amp; K. Yunokawa, pers. comm.) (
<figureCitation id="13602A5A4B56FF81FB8AFF6FFBC94EE3" box="[1065,1134,223,247]" captionStart="FIGURE 2" captionStartId="2.[151,250,1970,1993]" captionTargetBox="[151,1436,985,1948]" captionTargetId="figure@2.[151,1436,985,1949]" captionTargetPageId="2" captionText="FIGURE 2. Live colouration of Catoptrus iejima, n. sp., in situ, photo taken by Yusuke Yamada. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317." httpUri="https://zenodo.org/record/207595/files/figure.png" pageId="7" pageNumber="36">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF2466574B56FF81FF34F8A9FC9C4979" httpUri="https://zenodo.org/record/207600/files/figure.png" pageId="7" pageNumber="36" targetBox="[182,1386,324,1770]" targetPageId="7">
<paragraph id="8BE436DF4B56FF81FF34F8A9FC9C4979" blockId="7.[151,1436,1817,1901]" pageId="7" pageNumber="36">
<emphasis id="B92FEACD4B56FF81FF34F8A9FEB4493B" bold="true" box="[151,275,1817,1840]" pageId="7" pageNumber="36">FIGURE 7.</emphasis>
Upper views of right ambulatory legs of 
<taxonomicName id="4C5B4D5C4B56FF81FD18F8ABFCC04924" box="[699,871,1819,1840]" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="iejima" status="sp. nov.">
<emphasis id="B92FEACD4B56FF81FD18F8ABFCC04924" box="[699,871,1819,1840]" italics="true" pageId="7" pageNumber="36">Catoptrus iejima</emphasis>
</taxonomicName>
, 
<emphasis id="B92FEACD4B56FF81FCD7F8AAFC09493B" bold="true" box="[884,942,1818,1839]" pageId="7" pageNumber="36">
<taxonomicNameLabel id="A21C57B64B56FF81FCD7F8AAFC09493B" box="[884,942,1818,1839]" pageId="7" pageNumber="36" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A, E, P2 (first ambulatory leg = second pereopod) and its dactylus; B, F, P3 and its dactylus; C, G, P4 and its dactylus; D, H, P5 and its dactylus. Scale bars: A–D = 5.0 mm; E–H = 1.0 mm.
</paragraph>
</caption>
<subSubSection id="C34165544B56FF8EFF66F825FCE64F2B" lastPageId="8" lastPageNumber="37" pageId="7" pageNumber="36" type="discussion">
<paragraph id="8BE436DF4B56FF8EFF66F825FCE64F2B" blockId="7.[151,1437,1941,2038]" lastBlockId="8.[151,1436,151,825]" lastPageId="8" lastPageNumber="37" pageId="7" pageNumber="36">
<emphasis id="B92FEACD4B56FF81FF66F825FE9E49B9" bold="true" box="[197,313,1941,1965]" pageId="7" pageNumber="36">Remarks.</emphasis>
<taxonomicName id="4C5B4D5C4B56FF81FEE3F826FE5A49B9" box="[320,509,1942,1965]" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="iejima" status="sp. nov.">
<emphasis id="B92FEACD4B56FF81FEE3F826FE5A49B9" box="[320,509,1942,1965]" italics="true" pageId="7" pageNumber="36">Catoptrus iejima</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A21C57B64B56FF81FDA8F825FDED49B9" box="[523,586,1941,1965]" pageId="7" pageNumber="36" rank="species">
<emphasis id="B92FEACD4B56FF81FDA8F825FDED49B9" bold="true" box="[523,586,1941,1965]" pageId="7" pageNumber="36">n. sp.</emphasis>
</taxonomicNameLabel>
, superficially resembles 
<taxonomicName id="4C5B4D5C4B56FF81FCC0F825FB7349B9" authority="Ng &amp; Takeda, 2003" authorityName="Ng &amp; Takeda" authorityYear="2003" box="[867,1236,1941,1966]" class="Malacostraca" family="Portunidae" genus="Atoportunus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B92FEACD4B56FF81FCC0F825FC4949B9" box="[867,1006,1941,1965]" italics="true" pageId="7" pageNumber="36">Atoportunus</emphasis>
Ng &amp; Takeda, 2003
</taxonomicName>
, in its remarkable characters, such as the long and slender pereopods (
<figureCitation id="13602A5A4B56FF81FD42F809FC9649C5" box="[737,817,1977,2001]" captionStart="FIGURE 3" captionStartId="3.[151,250,1913,1936]" captionTargetBox="[245,1341,193,1855]" captionTargetId="figure@3.[245,1341,193,1856]" captionTargetPageId="3" captionText="FIGURE 3. Live colouration of Catoptrus iejima, n. sp. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A, habitus, dorsal view; B, habitus, ventral view." httpUri="https://zenodo.org/record/207596/files/figure.png" pageId="7" pageNumber="36">Figs. 3</figureCitation>
, 
<figureCitation id="13602A5A4B56FF81FC9DF809FCEA49C5" box="[830,845,1977,2001]" captionStart="FIGURE 7" captionStartId="7.[151,250,1817,1840]" captionTargetBox="[182,1386,324,1770]" captionTargetId="figure@7.[151,1436,301,1797]" captionTargetPageId="7" captionText="FIGURE 7. Upper views of right ambulatory legs of Catoptrus iejima, n. sp. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A, E, P 2 (first ambulatory leg = second pereopod) and its dactylus; B, F, P 3 and its dactylus; C, G, P 4 and its dactylus; D, H, P 5 and its dactylus. Scale bars: A – D = 5.0 mm; E – H = 1.0 mm." httpUri="https://zenodo.org/record/207600/files/figure.png" pageId="7" pageNumber="36">7</figureCitation>
) and relatively small eyes (
<figureCitation id="13602A5A4B56FF81FB26F809FB7249C5" box="[1157,1237,1977,2001]" captionStart="FIGURE 4" captionStartId="4.[151,250,1949,1972]" captionTargetBox="[380,1206,193,1928]" captionTargetId="figure@4.[380,1208,193,1929]" captionTargetPageId="4" captionText="FIGURE 4. Live colouration of Catoptrus iejima, n. sp. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A, carapace, dorsal view; B, cephalothorax, anterior view; C, cephalothorax, ventral view." httpUri="https://zenodo.org/record/207597/files/figure.png" pageId="7" pageNumber="36">Figs. 4</figureCitation>
B, 5A). It is noteworthy that the new species possesses all the diagnostic characters of 
<taxonomicName id="4C5B4D5C4B56FF81FC0AF86EFA9C49E1" authority="A. Milne-Edwards, 1870" authorityName="A. Milne-Edwards" authorityYear="1870" box="[937,1339,2013,2037]" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B92FEACD4B56FF81FC0AF86EFBBE49E1" box="[937,1049,2014,2037]" italics="true" pageId="7" pageNumber="36">Catoptrus</emphasis>
<bibRefCitation id="EFCA4B2E4B56FF81FB82F86DFA9C49E1" author="Milne-Edwards" box="[1057,1339,2013,2037]" pageId="7" pageNumber="36" refString="Milne-Edwards, A. (1870) Note sur le Catoptrus nouveau genre appartenant a la division des crustaces brachyoures catometopes. Annales des Sciences naturelles, Zoologie (Paris), ser. 5, 13, 82." type="journal article" year="1870">A. Milne-Edwards, 1870</bibRefCitation>
</taxonomicName>
, e.g. laterally ovate or rounded carapace (
<figureCitation id="13602A5A4B59FF8EFD84FF27FDC84EBB" box="[551,623,151,175]" captionStart="FIGURE 4" captionStartId="4.[151,250,1949,1972]" captionTargetBox="[380,1206,193,1928]" captionTargetId="figure@4.[380,1208,193,1929]" captionTargetPageId="4" captionText="FIGURE 4. Live colouration of Catoptrus iejima, n. sp. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A, carapace, dorsal view; B, cephalothorax, anterior view; C, cephalothorax, ventral view." httpUri="https://zenodo.org/record/207597/files/figure.png" pageId="8" pageNumber="37">Fig. 4</figureCitation>
A), subproximal and subdistal teeth present on the anterior margin of cheliped merus (
<figureCitation id="13602A5A4B59FF8EFEF3FF0CFE354EC0" box="[336,402,188,212]" captionStart="FIGURE 6" captionStartId="6.[151,250,1763,1786]" captionTargetBox="[151,1436,289,1741]" captionTargetId="figure@6.[151,1436,289,1742]" captionTargetPageId="6" captionText="FIGURE 6. Chelipeds of Catoptrus iejima, n. sp. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A – C, left cheliped; D – F, right cheliped. A, D, upper view; B, E, chela, outer view; C, F, chela, inner view." httpUri="https://zenodo.org/record/207599/files/figure.png" pageId="8" pageNumber="37">Fig. 6</figureCitation>
A, D) (except in 
<taxonomicName id="4C5B4D5C4B59FF8EFDEDFF0DFC064EC0" authority="Rathbun, 1906" authorityName="Rathbun" authorityYear="1906" box="[590,929,188,212]" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="B92FEACD4B59FF8EFDEDFF0DFD444EC0" box="[590,739,188,212]" italics="true" pageId="8" pageNumber="37">C. inaequalis</emphasis>
(
<bibRefCitation id="EFCA4B2E4B59FF8EFD52FF0CFC3F4EC0" author="Rathbun" box="[753,920,188,212]" pageId="8" pageNumber="37" refString="Rathbun, M. J. (1906) The Brachyura and Macrura of the Hawaiian Islands. Bulletin of the United States Fish Commission, 23 (3) [1903], 827 - 930, pls. 1 - 24." type="journal article" year="1906">Rathbun, 1906</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="4C5B4D5C4B59FF8EFC74FF0DFF744EE3" authority="Yang, Chen &amp; Tang, 2006" authorityName="Yang, Chen &amp; Tang" authorityYear="2006" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="quinquedentatus">
<emphasis id="B92FEACD4B59FF8EFC74FF0DFB174EC0" box="[983,1200,188,212]" italics="true" pageId="8" pageNumber="37">C. quinquedentatus</emphasis>
<bibRefCitation id="EFCA4B2E4B59FF8EFB14FF0DFF744EE3" author="Yang" pageId="8" pageNumber="37" refString="Yang, S. - L., Chen, H. - L. &amp; Tang, B. - P. (2006) On two new species of Catoptrus A. Milne-Edwards, 1870 (Decapoda, Brachyura, Portunidae) from China. Crustaceana, 79 (7), 835 - 842." type="journal article" year="2006">Yang, Chen &amp; Tang, 2006</bibRefCitation>
</taxonomicName>
), almost right angled distolateral angle of the merus of the third maxilliped (
<figureCitation id="13602A5A4B59FF8EFB91FF6FFBD34EE3" box="[1074,1140,223,247]" captionStart="FIGURE 5" captionStartId="5.[151,250,1417,1440]" captionTargetBox="[197,1416,208,1378]" captionTargetId="figure@5.[151,1436,193,1396]" captionTargetPageId="5" captionText="FIGURE 5. Catoptrus iejima, n. sp. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A, cephalothorax, anterior view; B, right eye, dorsal view; C, cephalothorax, dorsal view; D, left third maxilliped; E, abdomen and telson. Scale bars: A, C, E = 5.0 mm; B, D = 1.0 mm." httpUri="https://zenodo.org/record/207598/files/figure.png" pageId="8" pageNumber="37">Fig. 5</figureCitation>
D), and straight to lanceolate dactylus of P5 (
<figureCitation id="13602A5A4B59FF8EFED0FEB4FE124F08" box="[371,437,260,284]" captionStart="FIGURE 7" captionStartId="7.[151,250,1817,1840]" captionTargetBox="[182,1386,324,1770]" captionTargetId="figure@7.[151,1436,301,1797]" captionTargetPageId="7" captionText="FIGURE 7. Upper views of right ambulatory legs of Catoptrus iejima, n. sp. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A, E, P 2 (first ambulatory leg = second pereopod) and its dactylus; B, F, P 3 and its dactylus; C, G, P 4 and its dactylus; D, H, P 5 and its dactylus. Scale bars: A – D = 5.0 mm; E – H = 1.0 mm." httpUri="https://zenodo.org/record/207600/files/figure.png" pageId="8" pageNumber="37">Fig. 7</figureCitation>
H) (
<bibRefCitation id="EFCA4B2E4B59FF8EFE42FEB4FDD04F08" author="Serene" box="[481,631,260,284]" pageId="8" pageNumber="37" refString="Serene, R. (1966) Notes sur les genres Catoptrus et Lybistes et les Catoptrinae. Bulletin du Museum national d'Histoire naturelle, (2) 37 (6), 929 - 1000." type="journal article" year="1966">Serène, 1966</bibRefCitation>
; Ng, 2011). Although no male specimen was collected, the characters of 
<taxonomicName id="4C5B4D5C4B59FF8EFF34FE98FF5A4F54" box="[151,253,296,320]" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="iejima" status="sp. nov.">
<emphasis id="B92FEACD4B59FF8EFF34FE98FF5A4F54" box="[151,253,296,320]" italics="true" pageId="8" pageNumber="37">C. iejima</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A21C57B64B59FF8EFEAFFE98FEED4F54" box="[268,330,296,320]" pageId="8" pageNumber="37" rank="species">
<emphasis id="B92FEACD4B59FF8EFEAFFE98FEED4F54" bold="true" box="[268,330,296,320]" pageId="8" pageNumber="37">n. sp.</emphasis>
</taxonomicNameLabel>
, best fit 
<emphasis id="B92FEACD4B59FF8EFE0FFE98FDBB4F2B" box="[428,540,296,319]" italics="true" pageId="8" pageNumber="37">Catoprtus</emphasis>
<bibRefCitation id="EFCA4B2E4B59FF8EFD87FE98FC9A4F2B" author="Milne-Edwards" box="[548,829,295,320]" pageId="8" pageNumber="37" refString="Milne-Edwards, A. (1870) Note sur le Catoptrus nouveau genre appartenant a la division des crustaces brachyoures catometopes. Annales des Sciences naturelles, Zoologie (Paris), ser. 5, 13, 82." type="journal article" year="1870">A. Milne-Edwards, 1870</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34165544B59FF8FFF66FEFDFE974F93" lastPageId="9" lastPageNumber="38" pageId="8" pageNumber="37" type="reference_group">
<paragraph id="8BE436DF4B59FF8EFF66FEFDFAB04D23" blockId="8.[151,1436,151,825]" pageId="8" pageNumber="37">
<taxonomicName id="4C5B4D5C4B59FF8EFF66FEFDFE924F70" ID-CoL="7NS3T" box="[197,309,333,356]" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B92FEACD4B59FF8EFF66FEFDFE924F70" box="[197,309,333,356]" italics="true" pageId="8" pageNumber="37">Catoptrus</emphasis>
</taxonomicName>
currently contains five species: 
<taxonomicName id="4C5B4D5C4B59FF8EFD0AFEFDFBA24F70" ID-CoL="RVX4" authority="Rathbun, 1906" authorityName="Rathbun" authorityYear="1906" box="[681,1029,332,356]" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="B92FEACD4B59FF8EFD0AFEFDFCE74F70" box="[681,832,332,356]" italics="true" pageId="8" pageNumber="37">C. inaequalis</emphasis>
(
<bibRefCitation id="EFCA4B2E4B59FF8EFCF2FEFCFC594F70" author="Rathbun" box="[849,1022,332,356]" pageId="8" pageNumber="37" refString="Rathbun, M. J. (1906) The Brachyura and Macrura of the Hawaiian Islands. Bulletin of the United States Fish Commission, 23 (3) [1903], 827 - 930, pls. 1 - 24." type="journal article" year="1906">Rathbun, 1906</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C5B4D5C4B59FF8EFBB0FEFDFEBC4F93" ID-CoL="RVX7" authority="Yang, Chen &amp; Tang, 2006" authorityName="Yang, Chen &amp; Tang" authorityYear="2006" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="quinquedentatus">
<emphasis id="B92FEACD4B59FF8EFBB0FEFDFB484F70" box="[1043,1263,332,356]" italics="true" pageId="8" pageNumber="37">C. quinquedentatus</emphasis>
<bibRefCitation id="EFCA4B2E4B59FF8EFB5BFEFDFEBC4F93" author="Yang" pageId="8" pageNumber="37" refString="Yang, S. - L., Chen, H. - L. &amp; Tang, B. - P. (2006) On two new species of Catoptrus A. Milne-Edwards, 1870 (Decapoda, Brachyura, Portunidae) from China. Crustaceana, 79 (7), 835 - 842." type="journal article" year="2006">Yang, Chen &amp; Tang, 2006</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C5B4D5C4B59FF8EFE8AFEC0FD664F93" ID-CoL="RVX6" authority="A. Milne-Edwards, 1870" authorityName="A. Milne-Edwards" authorityYear="1870" box="[297,705,367,392]" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="B92FEACD4B59FF8EFE8AFEC0FE3E4F9C" box="[297,409,368,392]" italics="true" pageId="8" pageNumber="37">C. nitidus</emphasis>
<bibRefCitation id="EFCA4B2E4B59FF8EFE07FEC0FD664F93" author="Milne-Edwards" box="[420,705,367,392]" pageId="8" pageNumber="37" refString="Milne-Edwards, A. (1870) Note sur le Catoptrus nouveau genre appartenant a la division des crustaces brachyoures catometopes. Annales des Sciences naturelles, Zoologie (Paris), ser. 5, 13, 82." type="journal article" year="1870">A. Milne-Edwards, 1870</bibRefCitation>
</taxonomicName>
[
<typeStatus id="54E0887D4B59FF8EFD72FEC0FCA34F9C" box="[721,772,368,392]" pageId="8" pageNumber="37">type</typeStatus>
species], 
<taxonomicName id="4C5B4D5C4B59FF8EFCD9FEC0FB134F93" ID-CoL="RVX8" authority="Serene, 1966" authorityName="Serene" authorityYear="1966" box="[890,1204,367,392]" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="rathbunae">
<emphasis id="B92FEACD4B59FF8EFCD9FEC0FBB64F9C" box="[890,1041,368,392]" italics="true" pageId="8" pageNumber="37">C. rathbunae</emphasis>
<bibRefCitation id="EFCA4B2E4B59FF8EFBBFFEDFFB134F93" author="Serene" box="[1052,1204,367,391]" pageId="8" pageNumber="37" refString="Serene, R. (1966) Notes sur les genres Catoptrus et Lybistes et les Catoptrinae. Bulletin du Museum national d'Histoire naturelle, (2) 37 (6), 929 - 1000." type="journal article" year="1966">Serène, 1966</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="4C5B4D5C4B59FF8EFB56FEC0FD344FB8" ID-CoL="RVX9" authority="Yang, Chen &amp; Tang, 2006 (Ng et al., 2008)" authorityName="Yang, Chen &amp; Tang, 2006 (Ng et al." authorityYear="2008" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="undulatipes">
<emphasis id="B92FEACD4B59FF8EFB56FEC0FA3B4F9C" box="[1269,1436,368,392]" italics="true" pageId="8" pageNumber="37">C. undulatipes</emphasis>
<bibRefCitation id="EFCA4B2E4B59FF8EFF34FE25FE604FB8" author="Yang" box="[151,455,404,429]" pageId="8" pageNumber="37" refString="Yang, S. - L., Chen, H. - L. &amp; Tang, B. - P. (2006) On two new species of Catoptrus A. Milne-Edwards, 1870 (Decapoda, Brachyura, Portunidae) from China. Crustaceana, 79 (7), 835 - 842." type="journal article" year="2006">Yang, Chen &amp; Tang, 2006</bibRefCitation>
(Ng et al., 2008)
</taxonomicName>
. 
<taxonomicName id="4C5B4D5C4B59FF8EFD06FE25FCC24FB8" ID-CoL="RVX3" box="[677,869,405,428]" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="iejima" status="sp. nov.">
<emphasis id="B92FEACD4B59FF8EFD06FE25FCC24FB8" box="[677,869,405,428]" italics="true" pageId="8" pageNumber="37">Catoptrus iejima</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A21C57B64B59FF8EFCD5FE24FC1F4FB8" box="[886,952,404,428]" pageId="8" pageNumber="37" rank="species">
<emphasis id="B92FEACD4B59FF8EFCD5FE24FC1F4FB8" bold="true" box="[886,952,404,428]" pageId="8" pageNumber="37">n. sp.</emphasis>
</taxonomicNameLabel>
most closely resembles 
<taxonomicName id="4C5B4D5C4B59FF8EFB75FE25FAE14FB8" ID-CoL="RVX6" box="[1238,1350,404,428]" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="B92FEACD4B59FF8EFB75FE25FAE14FB8" box="[1238,1350,404,428]" italics="true" pageId="8" pageNumber="37">C. nitidus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C5B4D5C4B59FF8EFA21FE25FEBD4FC4" ID-CoL="RVX9" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="undulatipes">
<emphasis id="B92FEACD4B59FF8EFA21FE25FEBD4FC4" italics="true" pageId="8" pageNumber="37">C. undulatipes</emphasis>
</taxonomicName>
in their laterally ovate carapace outlines and the presence of two teeth on the anterior margin of each cheliped merus. However, 
<taxonomicName id="4C5B4D5C4B59FF8EFE65FE6DFD894FE0" ID-CoL="RVX3" box="[454,558,477,500]" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="iejima" status="sp. nov.">
<emphasis id="B92FEACD4B59FF8EFE65FE6DFD894FE0" box="[454,558,477,500]" italics="true" pageId="8" pageNumber="37">C. iejima</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A21C57B64B59FF8EFD9EFE6CFDDA4FE0" box="[573,637,476,500]" pageId="8" pageNumber="37" rank="species">
<emphasis id="B92FEACD4B59FF8EFD9EFE6CFDDA4FE0" bold="true" box="[573,637,476,500]" pageId="8" pageNumber="37">n. sp.</emphasis>
</taxonomicNameLabel>
can be readily distinguished from 
<taxonomicName id="4C5B4D5C4B59FF8EFBA9FE6DFBDF4FE0" ID-CoL="RVX6" box="[1034,1144,476,500]" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="B92FEACD4B59FF8EFBA9FE6DFBDF4FE0" box="[1034,1144,476,500]" italics="true" pageId="8" pageNumber="37">C. nitidus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C5B4D5C4B59FF8EFB13FE6DFAF24FE0" ID-CoL="RVX9" box="[1200,1365,476,500]" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="undulatipes">
<emphasis id="B92FEACD4B59FF8EFB13FE6DFAF24FE0" box="[1200,1365,476,500]" italics="true" pageId="8" pageNumber="37">C. undulatipes</emphasis>
</taxonomicName>
by its proportionately more elongated pereopods and the shape of the teeth on the anterolateral margin of the carapace. In 
<taxonomicName id="4C5B4D5C4B59FF8EFF34FD95FF594C28" ID-CoL="RVX3" box="[151,254,549,572]" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="iejima">
<emphasis id="B92FEACD4B59FF8EFF34FD95FF594C28" box="[151,254,549,572]" italics="true" pageId="8" pageNumber="37">C. iejima</emphasis>
</taxonomicName>
, the merus of P3, for example, is 1.03 times as long as the CL (
<figureCitation id="13602A5A4B59FF8EFC71FD94FBB84C28" box="[978,1055,548,572]" captionStart="FIGURE 3" captionStartId="3.[151,250,1913,1936]" captionTargetBox="[245,1341,193,1855]" captionTargetId="figure@3.[245,1341,193,1856]" captionTargetPageId="3" captionText="FIGURE 3. Live colouration of Catoptrus iejima, n. sp. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A, habitus, dorsal view; B, habitus, ventral view." httpUri="https://zenodo.org/record/207596/files/figure.png" pageId="8" pageNumber="37">Figs. 3</figureCitation>
A, 7B), while that of 
<taxonomicName id="4C5B4D5C4B59FF8EFAB7FD95FA264C28" ID-CoL="RVX6" box="[1300,1409,548,572]" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="B92FEACD4B59FF8EFAB7FD95FA264C28" box="[1300,1409,548,572]" italics="true" pageId="8" pageNumber="37">C. nitidus</emphasis>
</taxonomicName>
is only 0.74 (
<figureCitation id="13602A5A4B59FF8EFEB1FDF7FEF04C4B" box="[274,343,583,607]" captionStart="FIGURE 8" captionStartId="8.[151,250,1780,1803]" captionTargetBox="[151,1436,877,1759]" captionTargetId="figure@8.[151,1436,877,1760]" captionTargetPageId="8" captionText="FIGURE 8. Catoptrus nitidus A. Milne-Edwards, 1870. Male, 15.6 x 23.9 mm, MNHN B- 4639, Iles Viti." httpUri="https://zenodo.org/record/207601/files/figure.png" pageId="8" pageNumber="37">Fig. 8</figureCitation>
). 
<bibRefCitation id="EFCA4B2E4B59FF8EFECEFDF8FD9C4C4B" box="[365,571,583,608]" pageId="8" pageNumber="37" refString="Yang, S. - L., Chen, H. - L. &amp; Tang, B. - P. (2006) On two new species of Catoptrus A. Milne-Edwards, 1870 (Decapoda, Brachyura, Portunidae) from China. Crustaceana, 79 (7), 835 - 842." type="journal article">
Yang 
<emphasis id="B92FEACD4B59FF8EFE0CFDF9FE4E4C74" box="[431,489,584,608]" italics="true" pageId="8" pageNumber="37">et al.</emphasis>
(2006)
</bibRefCitation>
did not describe or indicate how long the ambulatory legs of 
<taxonomicName id="4C5B4D5C4B59FF8EFB54FDF8FA3B4C74" ID-CoL="RVX9" box="[1271,1436,584,608]" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="undulatipes">
<emphasis id="B92FEACD4B59FF8EFB54FDF8FA3B4C74" box="[1271,1436,584,608]" italics="true" pageId="8" pageNumber="37">C. undulatipes</emphasis>
</taxonomicName>
were, but their figure of its cheliped (
<bibRefCitation id="EFCA4B2E4B59FF8EFD95FDDCFD5C4C90" author="Yang" box="[566,763,620,644]" pageId="8" pageNumber="37" refString="Yang, S. - L., Chen, H. - L. &amp; Tang, B. - P. (2006) On two new species of Catoptrus A. Milne-Edwards, 1870 (Decapoda, Brachyura, Portunidae) from China. Crustaceana, 79 (7), 835 - 842." type="journal article" year="2006">
Yang 
<emphasis id="B92FEACD4B59FF8EFDDAFDDDFD164C90" box="[633,689,620,644]" italics="true" pageId="8" pageNumber="37">et al.</emphasis>
, 2006
</bibRefCitation>
: fig. 3D) shows a merus which is clearly shorter than that of 
<taxonomicName id="4C5B4D5C4B59FF8EFF34FD20FF594CBC" ID-CoL="RVX3" box="[151,254,656,680]" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="iejima">
<emphasis id="B92FEACD4B59FF8EFF34FD20FF594CBC" box="[151,254,656,680]" italics="true" pageId="8" pageNumber="37">C. iejima</emphasis>
</taxonomicName>
(
<figureCitation id="13602A5A4B59FF8EFEAEFD3FFEFB4CB3" box="[269,348,655,679]" captionStart="FIGURE 2" captionStartId="2.[151,250,1970,1993]" captionTargetBox="[151,1436,985,1948]" captionTargetId="figure@2.[151,1436,985,1949]" captionTargetPageId="2" captionText="FIGURE 2. Live colouration of Catoptrus iejima, n. sp., in situ, photo taken by Yusuke Yamada. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317." httpUri="https://zenodo.org/record/207595/files/figure.png" pageId="8" pageNumber="37">Figs. 2</figureCitation>
, 
<figureCitation id="13602A5A4B59FF8EFECBFD3FFEDF4CB3" box="[360,376,655,679]" captionStart="FIGURE 3" captionStartId="3.[151,250,1913,1936]" captionTargetBox="[245,1341,193,1855]" captionTargetId="figure@3.[245,1341,193,1856]" captionTargetPageId="3" captionText="FIGURE 3. Live colouration of Catoptrus iejima, n. sp. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A, habitus, dorsal view; B, habitus, ventral view." httpUri="https://zenodo.org/record/207596/files/figure.png" pageId="8" pageNumber="37">3</figureCitation>
). Eyes of 
<taxonomicName id="4C5B4D5C4B59FF8EFE49FD20FDF64CBC" ID-CoL="RVX3" box="[490,593,656,680]" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="iejima">
<emphasis id="B92FEACD4B59FF8EFE49FD20FDF64CBC" box="[490,593,656,680]" italics="true" pageId="8" pageNumber="37">C. iejima</emphasis>
</taxonomicName>
is reduced and not occupying the orbit (
<figureCitation id="13602A5A4B59FF8EFBB5FD3FFBC14CB3" box="[1046,1126,655,679]" captionStart="FIGURE 2" captionStartId="2.[151,250,1970,1993]" captionTargetBox="[151,1436,985,1948]" captionTargetId="figure@2.[151,1436,985,1949]" captionTargetPageId="2" captionText="FIGURE 2. Live colouration of Catoptrus iejima, n. sp., in situ, photo taken by Yusuke Yamada. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317." httpUri="https://zenodo.org/record/207595/files/figure.png" pageId="8" pageNumber="37">Figs. 2</figureCitation>
, 
<figureCitation id="13602A5A4B59FF8EFBD1FD3FFB264CB3" box="[1138,1153,655,679]" captionStart="FIGURE 4" captionStartId="4.[151,250,1949,1972]" captionTargetBox="[380,1206,193,1928]" captionTargetId="figure@4.[380,1208,193,1929]" captionTargetPageId="4" captionText="FIGURE 4. Live colouration of Catoptrus iejima, n. sp. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A, carapace, dorsal view; B, cephalothorax, anterior view; C, cephalothorax, ventral view." httpUri="https://zenodo.org/record/207597/files/figure.png" pageId="8" pageNumber="37">4</figureCitation>
A, B, 5A–C). In contrast, the eyes of 
<taxonomicName id="4C5B4D5C4B59FF8EFEB8FD05FE2E4CD8" ID-CoL="RVX6" box="[283,393,692,716]" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="B92FEACD4B59FF8EFEB8FD05FE2E4CD8" box="[283,393,692,716]" italics="true" pageId="8" pageNumber="37">C. nitidus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C5B4D5C4B59FF8EFE60FD05FDCF4CD8" ID-CoL="RVX9" box="[451,616,692,716]" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="undulatipes">
<emphasis id="B92FEACD4B59FF8EFE60FD05FDCF4CD8" box="[451,616,692,716]" italics="true" pageId="8" pageNumber="37">C. undulatipes</emphasis>
</taxonomicName>
are developed and fully occupying the orbits (
<figureCitation id="13602A5A4B59FF8EFBDEFD04FB654CD8" box="[1149,1218,692,716]" captionStart="FIGURE 8" captionStartId="8.[151,250,1780,1803]" captionTargetBox="[151,1436,877,1759]" captionTargetId="figure@8.[151,1436,877,1760]" captionTargetPageId="8" captionText="FIGURE 8. Catoptrus nitidus A. Milne-Edwards, 1870. Male, 15.6 x 23.9 mm, MNHN B- 4639, Iles Viti." httpUri="https://zenodo.org/record/207601/files/figure.png" pageId="8" pageNumber="37">Fig. 8</figureCitation>
; 
<bibRefCitation id="EFCA4B2E4B59FF8EFB72FD05FA3F4CD8" author="Yang" box="[1233,1432,692,717]" pageId="8" pageNumber="37" refString="Yang, S. - L., Chen, H. - L. &amp; Tang, B. - P. (2006) On two new species of Catoptrus A. Milne-Edwards, 1870 (Decapoda, Brachyura, Portunidae) from China. Crustaceana, 79 (7), 835 - 842." type="journal article" year="2006">
Yang 
<emphasis id="B92FEACD4B59FF8EFAB0FD05FAEA4CD8" box="[1299,1357,692,716]" italics="true" pageId="8" pageNumber="37">et al.</emphasis>
, 2006
</bibRefCitation>
: fig. 3A, B). Furthermore, in 
<taxonomicName id="4C5B4D5C4B59FF8EFE75FD68FD9A4CE4" ID-CoL="RVX3" box="[470,573,728,752]" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="iejima" status="sp. nov.">
<emphasis id="B92FEACD4B59FF8EFE75FD68FD9A4CE4" box="[470,573,728,752]" italics="true" pageId="8" pageNumber="37">C. iejima</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A21C57B64B59FF8EFDEFFD68FD2C4CE4" box="[588,651,728,752]" pageId="8" pageNumber="37" rank="species">
<emphasis id="B92FEACD4B59FF8EFDEFFD68FD2C4CE4" bold="true" box="[588,651,728,752]" pageId="8" pageNumber="37">n. sp.</emphasis>
</taxonomicNameLabel>
, the teeth of the anterolateral margin are relatively small, so that their bases are well apart from each other. In contrast, the teeth of 
<taxonomicName id="4C5B4D5C4B59FF8EFCC5FD4DFC714D00" ID-CoL="RVX6" box="[870,982,764,788]" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="B92FEACD4B59FF8EFCC5FD4DFC714D00" box="[870,982,764,788]" italics="true" pageId="8" pageNumber="37">C. nitidus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C5B4D5C4B59FF8EFBB0FD4DFB1E4D00" ID-CoL="RVX9" box="[1043,1209,764,788]" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="undulatipes">
<emphasis id="B92FEACD4B59FF8EFBB0FD4DFB1E4D00" box="[1043,1209,764,788]" italics="true" pageId="8" pageNumber="37">C. undulatipes</emphasis>
</taxonomicName>
are proportionately larger and wider, so the bases of the teeth are adjacent to each other (
<figureCitation id="13602A5A4B59FF8EFC39FCAFFC794D23" box="[922,990,799,823]" captionStart="FIGURE 8" captionStartId="8.[151,250,1780,1803]" captionTargetBox="[151,1436,877,1759]" captionTargetId="figure@8.[151,1436,877,1760]" captionTargetPageId="8" captionText="FIGURE 8. Catoptrus nitidus A. Milne-Edwards, 1870. Male, 15.6 x 23.9 mm, MNHN B- 4639, Iles Viti." httpUri="https://zenodo.org/record/207601/files/figure.png" pageId="8" pageNumber="37">Fig. 8</figureCitation>
; 
<bibRefCitation id="EFCA4B2E4B59FF8EFC48FC91FB0B4D23" author="Yang" box="[1003,1196,799,825]" pageId="8" pageNumber="37" refString="Yang, S. - L., Chen, H. - L. &amp; Tang, B. - P. (2006) On two new species of Catoptrus A. Milne-Edwards, 1870 (Decapoda, Brachyura, Portunidae) from China. Crustaceana, 79 (7), 835 - 842." type="journal article" year="2006">Yang et al., 2006</bibRefCitation>
: fig. 3A).
</paragraph>
<caption id="DF2466574B59FF8EFF34F944FB6D491F" box="[151,1226,1780,1803]" httpUri="https://zenodo.org/record/207601/files/figure.png" pageId="8" pageNumber="37" targetBox="[151,1436,877,1759]" targetPageId="8">
<paragraph id="8BE436DF4B59FF8EFF34F944FB6D491F" blockId="8.[151,1226,1780,1803]" box="[151,1226,1780,1803]" pageId="8" pageNumber="37">
<emphasis id="B92FEACD4B59FF8EFF34F944FEB5491E" bold="true" box="[151,274,1780,1803]" pageId="8" pageNumber="37">FIGURE 8.</emphasis>
<taxonomicName id="4C5B4D5C4B59FF8EFEBAF946FD68491E" ID-CoL="RVX6" authority="A. Milne-Edwards, 1870" authorityName="A. Milne-Edwards" authorityYear="1870" box="[281,719,1781,1803]" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="B92FEACD4B59FF8EFEBAF946FE6E491F" box="[281,457,1781,1803]" italics="true" pageId="8" pageNumber="37">Catoptrus nitidus</emphasis>
A. Milne-Edwards, 1870
</taxonomicName>
. Male, 15.6 x 23.9 mm, MNHN B-4639, Iles Viti.
</paragraph>
</caption>
<paragraph id="8BE436DF4B59FF8FFF66F882FE974F93" blockId="8.[151,1436,1842,2011]" lastBlockId="9.[151,1437,151,391]" lastPageId="9" lastPageNumber="38" pageId="8" pageNumber="37">
Although 
<taxonomicName id="4C5B4D5C4B59FF8EFE95F883FE3A495E" ID-CoL="RVX3" box="[310,413,1843,1866]" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="iejima" status="sp. nov.">
<emphasis id="B92FEACD4B59FF8EFE95F883FE3A495E" box="[310,413,1843,1866]" italics="true" pageId="8" pageNumber="37">C. iejima</emphasis>
</taxonomicName>
, 
<emphasis id="B92FEACD4B59FF8EFE0FF883FE4C495F" bold="true" box="[428,491,1843,1867]" pageId="8" pageNumber="37">
<taxonomicNameLabel id="A21C57B64B59FF8EFE0FF883FE4C495F" box="[428,491,1843,1867]" pageId="8" pageNumber="37" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="4C5B4D5C4B59FF8EFD80F882FD09495E" ID-CoL="7NRR4" box="[547,686,1842,1866]" class="Malacostraca" family="Portunidae" genus="Atoportunus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B92FEACD4B59FF8EFD80F882FD09495E" box="[547,686,1842,1866]" italics="true" pageId="8" pageNumber="37">Atoportunus</emphasis>
</taxonomicName>
species share remarkable characters, e.g. elongated pereopods and reduced eyes; they can be differentiated by the following characters. In 
<taxonomicName id="4C5B4D5C4B59FF8EFC62F8E7FB8F497A" ID-CoL="RVX3" box="[961,1064,1879,1902]" class="Malacostraca" family="Portunidae" genus="Catoptrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="iejima" status="sp. nov.">
<emphasis id="B92FEACD4B59FF8EFC62F8E7FB8F497A" box="[961,1064,1879,1902]" italics="true" pageId="8" pageNumber="37">C. iejima</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A21C57B64B59FF8EFB94F8E7FBD0497B" box="[1079,1143,1879,1903]" pageId="8" pageNumber="37" rank="species">
<emphasis id="B92FEACD4B59FF8EFB94F8E7FBD0497B" bold="true" box="[1079,1143,1879,1903]" pageId="8" pageNumber="37">n. sp.</emphasis>
</taxonomicNameLabel>
, the chelipeds are moderately stronger (
<figureCitation id="13602A5A4B59FF8EFEE1F8CAFE344986" box="[322,403,1914,1938]" captionStart="FIGURE 2" captionStartId="2.[151,250,1970,1993]" captionTargetBox="[151,1436,985,1948]" captionTargetId="figure@2.[151,1436,985,1949]" captionTargetPageId="2" captionText="FIGURE 2. Live colouration of Catoptrus iejima, n. sp., in situ, photo taken by Yusuke Yamada. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317." httpUri="https://zenodo.org/record/207595/files/figure.png" pageId="8" pageNumber="37">Figs. 2</figureCitation>
, 
<figureCitation id="13602A5A4B59FF8EFE02F8CAFE174986" box="[417,432,1914,1938]" captionStart="FIGURE 3" captionStartId="3.[151,250,1913,1936]" captionTargetBox="[245,1341,193,1855]" captionTargetId="figure@3.[245,1341,193,1856]" captionTargetPageId="3" captionText="FIGURE 3. Live colouration of Catoptrus iejima, n. sp. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A, habitus, dorsal view; B, habitus, ventral view." httpUri="https://zenodo.org/record/207596/files/figure.png" pageId="8" pageNumber="37">3</figureCitation>
), the palm of the chela is more swollen and the fingers are alternatively lined with long sharp teeth and a few small sharp teeth (
<figureCitation id="13602A5A4B59FF8EFDF0F82EFD3049A2" box="[595,663,1950,1974]" captionStart="FIGURE 6" captionStartId="6.[151,250,1763,1786]" captionTargetBox="[151,1436,289,1741]" captionTargetId="figure@6.[151,1436,289,1742]" captionTargetPageId="6" captionText="FIGURE 6. Chelipeds of Catoptrus iejima, n. sp. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A – C, left cheliped; D – F, right cheliped. A, D, upper view; B, E, chela, outer view; C, F, chela, inner view." httpUri="https://zenodo.org/record/207599/files/figure.png" pageId="8" pageNumber="37">Fig. 6</figureCitation>
); the outline of the carapace is elliptical in dorsal view (
<figureCitation id="13602A5A4B59FF8EFAA7F82EFAF749A2" box="[1284,1360,1950,1974]" captionStart="FIGURE 3" captionStartId="3.[151,250,1913,1936]" captionTargetBox="[245,1341,193,1855]" captionTargetId="figure@3.[245,1341,193,1856]" captionTargetPageId="3" captionText="FIGURE 3. Live colouration of Catoptrus iejima, n. sp. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A, habitus, dorsal view; B, habitus, ventral view." httpUri="https://zenodo.org/record/207596/files/figure.png" pageId="8" pageNumber="37">Figs. 3</figureCitation>
A, 4A, B, 5A); and the shape of the P5 propodus and dactylus are similar with those of the P2–P4 (
<figureCitation id="13602A5A4B59FF8EFB30F872FB7049CE" box="[1171,1239,1986,2010]" captionStart="FIGURE 7" captionStartId="7.[151,250,1817,1840]" captionTargetBox="[182,1386,324,1770]" captionTargetId="figure@7.[151,1436,301,1797]" captionTargetPageId="7" captionText="FIGURE 7. Upper views of right ambulatory legs of Catoptrus iejima, n. sp. Holotype, female, 21.8 × 35.6 mm, RUMF-ZC- 1317. A, E, P 2 (first ambulatory leg = second pereopod) and its dactylus; B, F, P 3 and its dactylus; C, G, P 4 and its dactylus; D, H, P 5 and its dactylus. Scale bars: A – D = 5.0 mm; E – H = 1.0 mm." httpUri="https://zenodo.org/record/207600/files/figure.png" pageId="8" pageNumber="37">Fig. 7</figureCitation>
). In 
<taxonomicName id="4C5B4D5C4B59FF8EFAA9F873FA3349CF" ID-CoL="7NRR4" box="[1290,1428,1987,2011]" class="Malacostraca" family="Portunidae" genus="Atoportunus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B92FEACD4B59FF8EFAA9F873FA3349CF" box="[1290,1428,1987,2011]" italics="true" pageId="8" pageNumber="37">Atoportunus</emphasis>
</taxonomicName>
, the cheliped is very slender (Ng &amp; Takeda, 2003: figs. 2A, 3A, 5A, 6A, 7; Takeda, 2003: figs. 1, 2E, 3); the palm is as narrow as the merus (Ng &amp; Takeda, 2003: figs. 2A, 3A, 5A, 6A, 7; Takeda, 2003: figs. 1, 2E, 3), the very slender fingers are sparsely provided with long and thin teeth (Ng &amp; Takeda, 2003: figs. 5B, 6C, 7; Takeda, 2003: fig. 2F); the outline of the carapace is more typical, been fan-shaped outline with stronger lobes or teeth along anterolateral margin (Ng &amp; Takeda, 2003: figs. 2B, 3A, 4A, 6B 8A; Takeda, 2003: figs. 1A, 2A); and the propodus and dactylus of P5 are always foliaceous and widely lanceolate, respectively (Ng &amp; Takeda, 2003: figs. 5C, 8C; Takeda, 2003: figs. 1B, 2G).
</paragraph>
</subSubSection>
</treatment>
</document>