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<document id="0E3045EA4E3B10B7A590873E4A68A882" ID-CLB-Dataset="25313" ID-DOI="10.11646/zootaxa.4679.3.7" ID-GBIF-Dataset="0a75e36a-4254-44be-9a1a-2c2bc6f115fa" ID-ISSN="1175-5326" ID-Zenodo-Dep="3471264" ID-ZooBank="1FF46982-622E-4C27-AB8E-8F8A993A419A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1570099354004" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2019" docId="03F287B9FFD71B16FF10FA5009E524D2" docLanguage="en" docName="zootaxa.4679.3.7.pdf" docOrigin="Zootaxa 4679 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Microscydmus (Microscydmus) yusuharanus Jałoszyński 2019, sp. n." docType="treatment" docVersion="7" lastPageNumber="549" masterDocId="FFCBFFC1FFDF1B1CFF87FFF80864257D" masterDocTitle="Unexpected diversity of Microscydmus Saulcy &amp; Croissandeau in Japan (Coleoptera: Staphylinidae: Scydmaeninae)" masterLastPageNumber="552" masterPageNumber="539" pageNumber="547" updateTime="1698757590553" updateUser="ExternalLinkService">
<mods:mods id="69534AF8AAA122514A8F7C18BEEDE77F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="BF5E0723E3E112AD97C67AB7D76885B1">Unexpected diversity of Microscydmus Saulcy &amp; Croissandeau in Japan (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
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<mods:namePart id="2FDA2ECD8F19D345AC268DDAE3EC24D7">Jałoszyński, Paweł</mods:namePart>
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<mods:title id="DE7629E14A9A31CF57D53604BD578891">Zootaxa</mods:title>
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<mods:part id="19F57D7103BCBF0047762A6C93B949E2">
<mods:date id="BEDB7BFBF61842A448835B9B63A6FEA2">2019</mods:date>
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<mods:number id="F39B86627E526A858BECCB275738CE1E">2019-10-03</mods:number>
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<mods:number id="5A255E778FF14B9D016ECF76A77CD7DE">4679</mods:number>
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<mods:number id="F4937953FA7AE26F8A6AE9F55813483E">3</mods:number>
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<mods:classification id="D7E56513D4867303A62CCD529AEAA2FC">journal article</mods:classification>
<mods:identifier id="CDEF706795CC3DDEC29ECB4133F8AA7B" type="CLB-Dataset">25313</mods:identifier>
<mods:identifier id="8058C1564B12EEB1DD390CE4B97FCF46" type="DOI">10.11646/zootaxa.4679.3.7</mods:identifier>
<mods:identifier id="B66AFD8AE1FB50F4C31C4455CEF10FB8" type="GBIF-Dataset">0a75e36a-4254-44be-9a1a-2c2bc6f115fa</mods:identifier>
<mods:identifier id="4EAEF0A301092ECA87ED48C63DC81066" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03F287B9FFD71B16FF10FA5009E524D2" ID-DOI="http://doi.org/10.5281/zenodo.5925434" ID-GBIF-Taxon="159628636" ID-Zenodo-Dep="5925434" LSID="urn:lsid:plazi:treatment:03F287B9FFD71B16FF10FA5009E524D2" httpUri="http://treatment.plazi.org/id/03F287B9FFD71B16FF10FA5009E524D2" lastPageId="10" lastPageNumber="549" pageId="8" pageNumber="547">
<subSubSection id="C3416524FFD71B14FF10FA500B6520BE" box="[151,769,1448,1475]" pageId="8" pageNumber="547" type="nomenclature">
<paragraph id="8BE436AFFFD71B14FF10FA500B6520BE" blockId="8.[151,769,1448,1511]" box="[151,769,1448,1475]" pageId="8" pageNumber="547">
<heading id="D0AC81C3FFD71B14FF10FA500B6520BE" bold="true" box="[151,769,1448,1475]" fontSize="11" level="1" pageId="8" pageNumber="547" reason="1">
<emphasis id="B92FEABDFFD71B14FF10FA500B6520BE" bold="true" box="[151,769,1448,1475]" pageId="8" pageNumber="547">
<taxonomicName id="4C5B4D2CFFD71B14FF10FA500AD320BE" ID-CoL="8Y6FL" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[151,695,1448,1475]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="547" phylum="Arthropoda" rank="species" species="yusuharanus" status="sp. nov." subGenus="Microscydmus">
<emphasis id="B92FEABDFFD71B14FF10FA50092320BE" bold="true" box="[151,327,1448,1475]" italics="true" pageId="8" pageNumber="547">Microscydmus</emphasis>
(
<emphasis id="B92FEABDFFD71B14FEDFFA500A6C20BE" bold="true" box="[344,520,1448,1475]" italics="true" pageId="8" pageNumber="547">Microscydmus</emphasis>
)
<emphasis id="B92FEABDFFD71B14FD9EFA500AD320BE" bold="true" box="[537,695,1448,1475]" italics="true" pageId="8" pageNumber="547">yusuharanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21C57C6FFD71B14FD38FA510B6520BE" box="[703,769,1449,1475]" pageId="8" pageNumber="547" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3416524FFD71B14FF10FA340909209A" box="[151,365,1484,1511]" pageId="8" pageNumber="547" type="description">
<paragraph id="8BE436AFFFD71B14FF10FA340909209A" blockId="8.[151,769,1448,1511]" box="[151,365,1484,1511]" pageId="8" pageNumber="547">
(
<figureCitation id="13602A2AFFD71B14FF18FA34088C209A" box="[159,232,1484,1511]" captionStart="FIGURES 1–8" captionStartId="4.[151,264,1753,1778]" captionTargetBox="[170,1420,187,1717]" captionTargetId="figure@4.[151,1436,174,1734]" captionTargetPageId="4" captionText="FIGURES 1–8. Dorsal habitus of males. Microscydmus okinawanus sp. n. (1); M. oharaensis sp. n. (2); M. nishimeanus sp. n. (3); M. kochiensis sp. n. (4); M. elytratus sp. n. (5); M. tibialis sp. n. (6); M. yusuharanus sp. n. (7); M. omogonus sp. n. (8)." figureDoi="http://doi.org/10.5281/zenodo.3471266" httpUri="https://zenodo.org/record/3471266/files/figure.png" pageId="8" pageNumber="547">Figs 7</figureCitation>
,
<figureCitation id="13602A2AFFD71B14FF73FA340976209A" box="[244,274,1484,1511]" captionStart="FIGURES 9–16" captionStartId="7.[151,264,1648,1673]" captionTargetBox="[151,1436,769,1629]" captionTargetId="figure@7.[151,1436,769,1629]" captionTargetPageId="7" captionText="FIGURES 9–16. Heads of males in anterodorsal view. Microscydmus okinawanus sp. n. (9); M. oharaensis sp. n. (10); M. nishimeanus sp. n. (11); M. kochiensis sp. n. (12); M. elytratus sp. n. (13); M. tibialis sp. n. (14); M. yusuharanus sp. n. (15); M. omogonus sp. n. (16). Arrowheads show posterolateral vertexal setae." figureDoi="http://doi.org/10.5281/zenodo.3471268" httpUri="https://zenodo.org/record/3471268/files/figure.png" pageId="8" pageNumber="547">15</figureCitation>
,
<figureCitation id="13602A2AFFD71B14FE99FA350958209A" box="[286,316,1485,1511]" captionStart="FIGURES 17–24" captionStartId="9.[151,264,1941,1966]" captionTargetBox="[155,1427,699,1921]" captionTargetId="figure@9.[151,1436,697,1921]" captionTargetPageId="9" captionText="FIGURES 17–24. Apices of male protibia in dorsal (17–21, 22–24) and ventral (22) views. Microscydmus okinawanus sp. n. (17); M. oharaensis sp. n. (18); M. nishimeanus sp. n. (19); M. kochiensis sp. n. (20); M. elytratus sp. n. (21); M. tibialis sp. n. (22); M. yusuharanus sp. n. (23); M. omogonus sp. n. (24)." figureDoi="http://doi.org/10.5281/zenodo.3471270" httpUri="https://zenodo.org/record/3471270/files/figure.png" pageId="8" pageNumber="547">23</figureCitation>
,
<figureCitation id="13602A2AFFD71B14FECFFA350901209A" box="[328,357,1485,1511]" captionStart="FIGURES 25–32" captionStartId="11.[151,264,1484,1509]" captionTargetBox="[156,1424,237,1452]" captionTargetId="figure@11.[151,1436,229,1465]" captionTargetPageId="11" captionText="FIGURES 25–32. Aedeagus in ventral view. Microscydmus okinawanus sp. n. (25); M. oharaensis sp. n. (26); M. nishimeanus sp. n. (27); M. kochiensis sp. n. (28); M. elytratus sp. n. (29); M. tibialis sp. n. (30); M. yusuharanus sp. n. (31); M. omogonus sp. n. (32)." figureDoi="http://doi.org/10.5281/zenodo.3471272" httpUri="https://zenodo.org/record/3471272/files/figure.png" pageId="8" pageNumber="547">31</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3416524FFD71B14FF10F9ED0C37230A" pageId="8" pageNumber="547" type="materials_examined">
<paragraph id="8BE436AFFFD71B14FF10F9ED0C37230A" blockId="8.[151,1437,1556,2015]" pageId="8" pageNumber="547">
<emphasis id="B92FEABDFFD71B14FF10F9ED09232352" bold="true" box="[151,327,1556,1583]" pageId="8" pageNumber="547">Type material.</emphasis>
<materialsCitation id="3B333CF2FFD71B14FECAF9ED0C37230A" ID-GBIF-Occurrence="2423812454" collectingDate="1997-06-14" collectionCode="JAPAN, EUMJ" collectorName="M. Sakai" country="Japan" location="Kochi" pageId="8" pageNumber="547" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B92FEABDFFD71B14FECAF9ED0AB52352" bold="true" box="[333,721,1556,1583]" pageId="8" pageNumber="547">
<typeStatus id="54E0880DFFD71B14FECAF9ED09DA2352" box="[333,446,1557,1583]" pageId="8" pageNumber="547" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F34C763FFFD71B14FE4DF9EC0A422352" box="[458,550,1556,1583]" name="Japan" pageId="8" pageNumber="547">JAPAN</collectingCountry>
(Kôchi Pref.):
</emphasis>
<specimenCount id="9D5DFD26FFD71B14FD5FF9ED0A8B2352" box="[728,751,1557,1583]" pageId="8" pageNumber="547" type="male">♂</specimenCount>
, two labels: &quot;[
<location id="8E846074FFD71B14FC10F9EC0B842353" LSID="urn:lsid:plazi:treatment:03F287B9FFD71B16FF10FA5009E524D2:8E846074FFD71B14FC10F9EC0B842353" box="[919,992,1556,1582]" country="Japan" name="Kochi" pageId="8" pageNumber="547">Kôchi</location>
:
<collectionCode id="ED4AAE6AFFD71B14FC6EF9ED0C242352" box="[1001,1088,1557,1583]" pageId="8" pageNumber="547">JAPAN</collectionCode>
] / Kami-oriwatari / Yusuharacho /
<date id="FFE5106FFFD71B14FF51F9C10939232E" box="[214,349,1592,1619]" pageId="8" pageNumber="547" value="1997-06-14">
<collectingDate id="EFA1E987FFD71B14FF51F9C10939232E" box="[214,349,1592,1619]" pageId="8" pageNumber="547" value="1997-06-14">14. vi. 1997</collectingDate>
</date>
/
<collectorName id="26AE5379FFD71B14FEF3F9C109BE232F" box="[372,474,1592,1619]" pageId="8" pageNumber="547">M. Sakai</collectorName>
(½ * {= Japanese
<emphasis id="B92FEABDFFD71B14FD4EF9C00B7E232F" box="[713,794,1592,1618]" italics="true" pageId="8" pageNumber="547">kuchiki</emphasis>
, rotten wood})&quot; [white, printed]; &quot;
<taxonomicName id="4C5B4D2CFFD71B14FB24F9C10D1E232E" authorityName="Saulcy &amp; Croissandeau" authorityYear="1893" box="[1187,1402,1593,1619]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="547" phylum="Arthropoda" rank="genus">
<emphasis id="B92FEABDFFD71B14FB24F9C10D1E232E" box="[1187,1402,1593,1619]" italics="true" pageId="8" pageNumber="547">MICROSCYDMUS</emphasis>
</taxonomicName>
(s. str.) /
<emphasis id="B92FEABDFFD71B14FF5EF9A4090C230B" box="[217,360,1628,1654]" italics="true" pageId="8" pageNumber="547">yusuharanus</emphasis>
m. / P.
<bibRefCitation id="EFCA4B5EFFD71B14FE47F9A40AE3230A" author="Jaloszynski, P." box="[448,647,1628,1655]" pageId="8" pageNumber="547" pagination="205 - 220" refId="ref9055" refString="Jaloszynski, P. (2019) Taxonomy of the &quot; Afroeudesis group &quot; of glandulariine ant-like stone beetles (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 4612 (2), 205 - 220. https: // doi. org / 10.11646 / zootaxa. 4612.2.4" type="journal article" year="2019">Jałoszyński, 2019</bibRefCitation>
/
<typeStatus id="54E0880DFFD71B14FD1BF9A50B22230A" box="[668,838,1629,1655]" pageId="8" pageNumber="547" type="holotype">HOLOTYPUS</typeStatus>
&quot; [red, printed] (
<collectionCode id="ED4AAE6AFFD71B14FC7FF9A50C23230A" box="[1016,1095,1629,1655]" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="8" pageNumber="547">EUMJ</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3416524FFD71B14FF40F9780C9D227B" pageId="8" pageNumber="547" type="diagnosis">
<paragraph id="8BE436AFFFD71B14FF40F9780C9D227B" blockId="8.[151,1437,1556,2015]" pageId="8" pageNumber="547">
<emphasis id="B92FEABDFFD71B14FF40F978092623E6" bold="true" box="[199,322,1664,1691]" pageId="8" pageNumber="547">Diagnosis.</emphasis>
Body length ~
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; frons and vertex with fine and shallow but distinct, dense punctures; bristles on tempora and vertex well-visible; protibia not thickened distad, with apical portion forming a slightly protruding, short subrectangular lobe with its apical margin oblique in relation to the long axis of tibia, apical comb absent; aedeagus with a subapical group of dense, needle-like sclerites; parameres slightly recurved.
</paragraph>
</subSubSection>
<subSubSection id="C3416524FFD71B16FF40F8E809FA2462" lastPageId="10" lastPageNumber="549" pageId="8" pageNumber="547" type="description">
<paragraph id="8BE436AFFFD71B14FF40F8E80B102232" blockId="8.[151,1437,1556,2015]" pageId="8" pageNumber="547">
<emphasis id="B92FEABDFFD71B14FF40F8E8093D2256" bold="true" box="[199,345,1808,1835]" pageId="8" pageNumber="547">Description.</emphasis>
Body of male (
<figureCitation id="13602A2AFFD71B14FD84F8E80A222256" box="[515,582,1808,1835]" captionStart="FIGURES 1–8" captionStartId="4.[151,264,1753,1778]" captionTargetBox="[170,1420,187,1717]" captionTargetId="figure@4.[151,1436,174,1734]" captionTargetPageId="4" captionText="FIGURES 1–8. Dorsal habitus of males. Microscydmus okinawanus sp. n. (1); M. oharaensis sp. n. (2); M. nishimeanus sp. n. (3); M. kochiensis sp. n. (4); M. elytratus sp. n. (5); M. tibialis sp. n. (6); M. yusuharanus sp. n. (7); M. omogonus sp. n. (8)." figureDoi="http://doi.org/10.5281/zenodo.3471266" httpUri="https://zenodo.org/record/3471266/files/figure.png" pageId="8" pageNumber="547">Fig. 7</figureCitation>
) moderately slender, slightly flattened, pigmentation moderately dark brown with indistinctly darker head, vestiture light brown; BL
<quantity id="4CA39B4AFFD71B14FC8EF8CD0B142232" box="[777,880,1845,1871]" metricMagnitude="-4" metricUnit="m" metricValue="7.4" pageId="8" pageNumber="547" unit="mm" value="0.74">0.74 mm</quantity>
.
</paragraph>
<paragraph id="8BE436AFFFD71B15FF40F8A00ABE2462" blockId="8.[151,1437,1556,2015]" lastBlockId="9.[151,1436,153,683]" lastPageId="9" lastPageNumber="548" pageId="8" pageNumber="547">
Head (
<figureCitation id="13602A2AFFD71B14FE95F8A00907220E" box="[274,355,1880,1907]" captionStart="FIGURES 9–16" captionStartId="7.[151,264,1648,1673]" captionTargetBox="[151,1436,769,1629]" captionTargetId="figure@7.[151,1436,769,1629]" captionTargetPageId="7" captionText="FIGURES 9–16. Heads of males in anterodorsal view. Microscydmus okinawanus sp. n. (9); M. oharaensis sp. n. (10); M. nishimeanus sp. n. (11); M. kochiensis sp. n. (12); M. elytratus sp. n. (13); M. tibialis sp. n. (14); M. yusuharanus sp. n. (15); M. omogonus sp. n. (16). Arrowheads show posterolateral vertexal setae." figureDoi="http://doi.org/10.5281/zenodo.3471268" httpUri="https://zenodo.org/record/3471268/files/figure.png" pageId="8" pageNumber="547">Fig. 15</figureCitation>
) broadest at eyes, HL
<quantity id="4CA39B4AFFD71B14FDD8F8A10AA1220E" box="[607,709,1881,1907]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="8" pageNumber="547" unit="mm" value="0.13">0.13 mm</quantity>
, HW
<quantity id="4CA39B4AFFD71B14FC81F8A10B09220E" box="[774,877,1881,1907]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="8" pageNumber="547" unit="mm" value="0.18">0.18 mm</quantity>
; tempora weakly arcuate, much shorter than eyes; posterior margin of vertex nearly straight, vertex and frons confluent and convex, except for distinctly flattened frontal area between and slightly behind supraantennal tubercles; supraantennal tubercles weakly elevated but welldefined; eyes large, strongly convex and coarsely faceted. Frons and vertex uniformly and evenly covered with small, shallow but relatively distinct and dense punctures; setae sparse, short and suberect, bristles on tempora and vertex well-visible; long posterolateral setae on vertex present (only the left seta visible in the studied specimen, the other one presumably broken off). Antennae slender, AnL
<quantity id="4CA39B4AFFD61B15FCA5FF190BEC2586" box="[802,904,225,251]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="548" unit="mm" value="0.3">0.30 mm</quantity>
, antennomeres I and II strongly elongate, III–X each strongly transverse, XI about as long as broad.
</paragraph>
<paragraph id="8BE436AFFFD61B15FF40FED10BD824AE" blockId="9.[151,1436,153,683]" pageId="9" pageNumber="548">
Pronotum in dorsal view nearly circular, broadest distinctly in front of middle, PL
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, PW
<quantity id="4CA39B4AFFD61B15FAA9FED10DFC243E" box="[1326,1432,297,323]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="9" pageNumber="548" unit="mm" value="0.23">0.23 mm</quantity>
; anterior margin weakly arcuate; anterior corners not marked; sides strongly rounded in anterior half, weakly so in posterior half; posterior pronotal corners distinct, obtuse-angled; posterior margin weakly arcuate; pronotal base with distinct, relatively long transverse groove, lateral pits indiscernible. Punctures on disc fine, inconspicuous; setae sparse, short, suberect; lateral bristles sparse and moderately long.
</paragraph>
<paragraph id="8BE436AFFFD61B15FF40FE250B77271E" blockId="9.[151,1436,153,683]" pageId="9" pageNumber="548">
Elytra together oval, broadest distinctly in front of middle, EL
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, EW
<quantity id="4CA39B4AFFD61B15FBB4FE250CFE248A" box="[1075,1178,477,503]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="9" pageNumber="548" unit="mm" value="0.33">0.33 mm</quantity>
, EI 1.31; humeral calli distinct; elytral base with large and deep basal impression; adsutural area indistinctly impressed in anterior half; apices of elytra separately rounded. Punctures on elytra dense but small, very shallow and with diffuse margins, inconspicuous; setae short, sparse and nearly recumbent.
</paragraph>
<paragraph id="8BE436AFFFD61B15FF40FD950CE827D6" blockId="9.[151,1436,153,683]" pageId="9" pageNumber="548">
Protibia (
<figureCitation id="13602A2AFFD61B15FEB7FD9509E627FA" box="[304,386,621,647]" captionStart="FIGURES 17–24" captionStartId="9.[151,264,1941,1966]" captionTargetBox="[155,1427,699,1921]" captionTargetId="figure@9.[151,1436,697,1921]" captionTargetPageId="9" captionText="FIGURES 17–24. Apices of male protibia in dorsal (17–21, 22–24) and ventral (22) views. Microscydmus okinawanus sp. n. (17); M. oharaensis sp. n. (18); M. nishimeanus sp. n. (19); M. kochiensis sp. n. (20); M. elytratus sp. n. (21); M. tibialis sp. n. (22); M. yusuharanus sp. n. (23); M. omogonus sp. n. (24)." figureDoi="http://doi.org/10.5281/zenodo.3471270" httpUri="https://zenodo.org/record/3471270/files/figure.png" pageId="9" pageNumber="548">Fig. 23</figureCitation>
) not thickened distad, with apical portion forming a slightly protruding, short subrectangular lobe with its apical margin oblique in relation to the long axis of tibia, apical comb absent.
</paragraph>
<caption id="DF246627FFD61B15FF10F86D0A942288" ID-DOI="http://doi.org/10.5281/zenodo.3471270" ID-Zenodo-Dep="3471270" httpUri="https://zenodo.org/record/3471270/files/figure.png" pageId="9" pageNumber="548" startId="9.[151,264,1941,1966]" targetBox="[155,1427,699,1921]" targetPageId="9">
<paragraph id="8BE436AFFFD61B15FF10F86D0A942288" blockId="9.[151,1436,1941,2037]" pageId="9" pageNumber="548">
<emphasis id="B92FEABDFFD61B15FF10F86D093122D3" bold="true" box="[151,341,1941,1966]" pageId="9" pageNumber="548">FIGURES 17–24.</emphasis>
Apices of male protibia in dorsal (17–21, 22–24) and ventral (22) views.
<taxonomicName id="4C5B4D2CFFD61B15FBCFF86D0D3822D0" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[1096,1372,1941,1965]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="548" phylum="Arthropoda" rank="species" species="okinawanus" status="sp. nov.">
<emphasis id="B92FEABDFFD61B15FBCFF86D0D3822D0" box="[1096,1372,1941,1965]" italics="true" pageId="9" pageNumber="548">Microscydmus okinawanus</emphasis>
</taxonomicName>
<emphasis id="B92FEABDFFD61B15FAE4F86D0DF822D0" bold="true" box="[1379,1436,1941,1965]" pageId="9" pageNumber="548">
<taxonomicNameLabel id="A21C57C6FFD61B15FAE4F86D0DF822D0" box="[1379,1436,1941,1965]" pageId="9" pageNumber="548" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(17);
<taxonomicName id="4C5B4D2CFFD61B15FF49F842093B22AC" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[206,351,1977,2001]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="548" phylum="Arthropoda" rank="species" species="oharaensis" status="sp. nov.">
<emphasis id="B92FEABDFFD61B15FF49F842093B22AC" box="[206,351,1977,2001]" italics="true" pageId="9" pageNumber="548">M. oharaensis</emphasis>
</taxonomicName>
<emphasis id="B92FEABDFFD61B15FEE2F84109FA22AC" bold="true" box="[357,414,1977,2001]" pageId="9" pageNumber="548">
<taxonomicNameLabel id="A21C57C6FFD61B15FEE2F84109FA22AC" box="[357,414,1977,2001]" pageId="9" pageNumber="548" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(18);
<taxonomicName id="4C5B4D2CFFD61B15FE5CF8420A1822AC" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[475,636,1977,2001]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="548" phylum="Arthropoda" rank="species" species="nishimeanus" status="sp. nov.">
<emphasis id="B92FEABDFFD61B15FE5CF8420A1822AC" box="[475,636,1977,2001]" italics="true" pageId="9" pageNumber="548">M. nishimeanus</emphasis>
</taxonomicName>
<emphasis id="B92FEABDFFD61B15FD05F8410ADE22AC" bold="true" box="[642,698,1977,2001]" pageId="9" pageNumber="548">
<taxonomicNameLabel id="A21C57C6FFD61B15FD05F8410ADE22AC" box="[642,698,1977,2001]" pageId="9" pageNumber="548" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(19);
<taxonomicName id="4C5B4D2CFFD61B15FD7FF8420BE722AC" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[760,899,1977,2001]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="548" phylum="Arthropoda" rank="species" species="kochiensis" status="sp. nov.">
<emphasis id="B92FEABDFFD61B15FD7FF8420BE722AC" box="[760,899,1977,2001]" italics="true" pageId="9" pageNumber="548">M. kochiensis</emphasis>
</taxonomicName>
<emphasis id="B92FEABDFFD61B15FC0EF8410BA622AC" bold="true" box="[905,962,1977,2001]" pageId="9" pageNumber="548">
<taxonomicNameLabel id="A21C57C6FFD61B15FC0EF8410BA622AC" box="[905,962,1977,2001]" pageId="9" pageNumber="548" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(20);
<taxonomicName id="4C5B4D2CFFD61B15FC78F8420C1E22AC" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[1023,1146,1977,2001]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="548" phylum="Arthropoda" rank="species" species="elytratus" status="sp. nov.">
<emphasis id="B92FEABDFFD61B15FC78F8420C1E22AC" box="[1023,1146,1977,2001]" italics="true" pageId="9" pageNumber="548">M. elytratus</emphasis>
</taxonomicName>
<emphasis id="B92FEABDFFD61B15FBF8F8410CDC22AC" bold="true" box="[1151,1208,1977,2001]" pageId="9" pageNumber="548">
<taxonomicNameLabel id="A21C57C6FFD61B15FBF8F8410CDC22AC" box="[1151,1208,1977,2001]" pageId="9" pageNumber="548" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(21);
<taxonomicName id="4C5B4D2CFFD61B15FB72F8420D3922AC" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[1269,1373,1977,2001]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="548" phylum="Arthropoda" rank="species" species="tibialis" status="sp. nov.">
<emphasis id="B92FEABDFFD61B15FB72F8420D3922AC" box="[1269,1373,1977,2001]" italics="true" pageId="9" pageNumber="548">M. tibialis</emphasis>
</taxonomicName>
<emphasis id="B92FEABDFFD61B15FAE4F8410DF822AC" bold="true" box="[1379,1436,1977,2001]" pageId="9" pageNumber="548">
<taxonomicNameLabel id="A21C57C6FFD61B15FAE4F8410DF822AC" box="[1379,1436,1977,2001]" pageId="9" pageNumber="548" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(22);
<taxonomicName id="4C5B4D2CFFD61B15FF48F82609162288" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[207,370,2013,2037]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="548" phylum="Arthropoda" rank="species" species="yusuharanus" status="sp. nov.">
<emphasis id="B92FEABDFFD61B15FF48F82609162288" box="[207,370,2013,2037]" italics="true" pageId="9" pageNumber="548">M. yusuharanus</emphasis>
</taxonomicName>
<emphasis id="B92FEABDFFD61B15FEFFF82509D52288" bold="true" box="[376,433,2013,2037]" pageId="9" pageNumber="548">
<taxonomicNameLabel id="A21C57C6FFD61B15FEFFF82509D52288" box="[376,433,2013,2037]" pageId="9" pageNumber="548" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(23);
<taxonomicName id="4C5B4D2CFFD61B15FE68F8260A1D2288" authority="Jałoszyński, 2019" authorityName="Jałoszyński" authorityYear="2019" box="[495,633,2014,2037]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="548" phylum="Arthropoda" rank="species" species="omogonus" status="sp. nov.">
<emphasis id="B92FEABDFFD61B15FE68F8260A1D2288" box="[495,633,2014,2037]" italics="true" pageId="9" pageNumber="548">M. omogonus</emphasis>
</taxonomicName>
<emphasis id="B92FEABDFFD61B15FD07F8250ADD2288" bold="true" box="[640,697,2013,2037]" pageId="9" pageNumber="548">
<taxonomicNameLabel id="A21C57C6FFD61B15FD07F8250ADD2288" box="[640,697,2013,2037]" pageId="9" pageNumber="548" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(24).
</paragraph>
</caption>
<paragraph id="8BE436AFFFD51B16FF40FF6108AE2586" blockId="10.[151,1436,153,431]" pageId="10" pageNumber="549">
Aedeagus (
<figureCitation id="13602A2AFFD51B16FEC3FF6109F125CE" box="[324,405,153,179]" captionStart="FIGURES 25–32" captionStartId="11.[151,264,1484,1509]" captionTargetBox="[156,1424,237,1452]" captionTargetId="figure@11.[151,1436,229,1465]" captionTargetPageId="11" captionText="FIGURES 25–32. Aedeagus in ventral view. Microscydmus okinawanus sp. n. (25); M. oharaensis sp. n. (26); M. nishimeanus sp. n. (27); M. kochiensis sp. n. (28); M. elytratus sp. n. (29); M. tibialis sp. n. (30); M. yusuharanus sp. n. (31); M. omogonus sp. n. (32)." figureDoi="http://doi.org/10.5281/zenodo.3471272" httpUri="https://zenodo.org/record/3471272/files/figure.png" pageId="10" pageNumber="549">Fig. 31</figureCitation>
) moderately slender, AeL
<quantity id="4CA39B4AFFD51B16FD3AFF610B4725CE" box="[701,803,153,179]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="549" unit="mm" value="0.1">0.10 mm</quantity>
, endophallus lightly sclerotized, in subapical region with a large group of dense, needle-like sclerites directed distomesad; parameres slightly sinuate, each with one apical seta.
</paragraph>
<paragraph id="8BE436AFFFD51B16FF40FEFD09FA2462" blockId="10.[151,1436,153,431]" box="[199,414,261,287]" pageId="10" pageNumber="549">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C3416524FFD51B16FF40FED00AE6243E" box="[199,642,296,323]" pageId="10" pageNumber="549" type="distribution">
<paragraph id="8BE436AFFFD51B16FF40FED00AE6243E" blockId="10.[151,1436,153,431]" box="[199,642,296,323]" pageId="10" pageNumber="549">
<emphasis id="B92FEABDFFD51B16FF40FED00905243E" bold="true" box="[199,353,296,323]" pageId="10" pageNumber="549">Distribution.</emphasis>
<collectingCountry id="F34C763FFFD51B16FEE0FED109CE243E" box="[359,426,297,323]" name="Japan" pageId="10" pageNumber="549">Japan</collectingCountry>
, southern Shikoku.
</paragraph>
</subSubSection>
<subSubSection id="C3416524FFD51B16FF40FEB50B56241A" box="[199,818,333,359]" pageId="10" pageNumber="549" type="etymology">
<paragraph id="8BE436AFFFD51B16FF40FEB50B56241A" blockId="10.[151,1436,153,431]" box="[199,818,333,359]" pageId="10" pageNumber="549">
<emphasis id="B92FEABDFFD51B16FF40FEB5092B241A" bold="true" box="[199,335,333,359]" pageId="10" pageNumber="549">Etymology.</emphasis>
Locotypical, after the village of Yusuhara.
</paragraph>
</subSubSection>
<subSubSection id="C3416524FFD51B16FF40FE8909E524D2" pageId="10" pageNumber="549" type="discussion">
<paragraph id="8BE436AFFFD51B16FF40FE8909E524D2" blockId="10.[151,1436,153,431]" pageId="10" pageNumber="549">
<emphasis id="B92FEABDFFD51B16FF40FE89095F24F6" bold="true" box="[199,315,369,395]" pageId="10" pageNumber="549">Remarks.</emphasis>
This species is most similar to
<taxonomicName id="4C5B4D2CFFD51B16FD11FE8A0B5224F6" authorityName="Jałoszyński" authorityYear="2019" box="[662,822,369,395]" class="Insecta" family="Staphylinidae" genus="Microscydmus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="549" phylum="Arthropoda" rank="species" species="oharaensis">
<emphasis id="B92FEABDFFD51B16FD11FE8A0B5224F6" box="[662,822,369,395]" italics="true" pageId="10" pageNumber="549">M. oharaensis</emphasis>
</taxonomicName>
; similarities and differences were discussed in remarks for the latter species.
</paragraph>
</subSubSection>
</treatment>
</document>
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