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<document id="BB9341F5BC4906E1FF2A5DB240658E24" ID-DOI="10.11646/phytotaxa.363.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13703414" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725581356715" checkinUser="felipe" docAuthor="Negaresh, Kazem &amp; Rahiminejad, Mohammad Reza" docDate="2018" docId="03E887F4DF7FFFA319D43B8CF01CF9FF" docLanguage="en" docName="phytotaxa.363.1.1.pdf" docOrigin="Phytotaxa 363 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.363.1.1" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.4:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="4" docTitle="Centaurea tomentella Handel-Mazzetti 1913" docType="treatment" docVersion="2" lastPageNumber="450" masterDocId="FFD1FF8CDF12FFCC195C3E3CF14EFF83" masterDocTitle="A revision of Centaurea sect. Cynaroides (Asteraceae, Cardueae-Centaureinae)" masterLastPageNumber="450" masterPageNumber="448" pageNumber="448" updateTime="1725593506943" updateUser="ExternalLinkService">
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<mods:title id="39DB9603A01DADFE03AED60088847D08">A revision of Centaurea sect. Cynaroides (Asteraceae, Cardueae-Centaureinae)</mods:title>
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<mods:name id="BAC76FD641EE0C178B2FC1FF4EBA2011" type="personal">
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<mods:namePart id="2B466B4E65C421B5755A351505BFAE15">Negaresh, Kazem</mods:namePart>
<mods:affiliation id="E9DBC5F2BBABEBF699F1487FD6290B68">Department of Horticulture, Faculty of Agriculture, Khuzestan Agricultural Sciences and Natural Resources University, Mollasani, Ahvaz, Iran. (* Corresponding author e-mail: negaresh @ ramin. ac. ir)</mods:affiliation>
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<mods:namePart id="49AA3AFDF2E608E6E696A96DB1048878">Rahiminejad, Mohammad Reza</mods:namePart>
<mods:affiliation id="5E49B7DA6FF5126723EC6C897EFAAF44">Department of Horticulture, Faculty of Agriculture, Khuzestan Agricultural Sciences and Natural Resources University, Mollasani, Ahvaz, Iran. (* Corresponding author e-mail: negaresh @ ramin. ac. ir) &amp; Department of Biology, University of Isfahan, Isfahan 81746 - 73441, Iran. &amp; Department of Horticulture, Faculty of Agriculture, Khuzestan Agricultural Sciences and Natural Resources University, Mollasani, Ahvaz, Iran. (* Corresponding author e-mail: negaresh @ ramin. ac. ir)</mods:affiliation>
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<mods:title id="2079A350504D64EA3515BA267FE8372F">Phytotaxa</mods:title>
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<paragraph id="8BFE36E2DF7FFFA119D43B8CF5B7F992" blockId="109.[136,1451,1454,1553]" pageId="109">
<emphasis id="B935EAF0DF7FFFA119D43B8CF1E5FA4A" bold="true" box="[136,171,1456,1481]" pageId="109">35.</emphasis>
<taxonomicName id="4C414D61DF7FFFA119E43B92F255FA4A" ID-CoL="69GDL" authority="Handel-Mazzetti (1913: 450)" authorityName="Handel-Mazzetti" authorityPageNumber="450" authorityYear="1913" box="[184,795,1454,1481]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="109" phylum="Tracheophyta" rank="species" species="tomentella">
<emphasis id="B935EAF0DF7FFFA119E43B92F0F9FA4A" bold="true" box="[184,439,1454,1481]" italics="true" pageId="109">Centaurea tomentella</emphasis>
<materialsCitation id="3B293CBFDF7FFFA118993B8CF255FA4A" box="[453,795,1456,1481]" collectingDate="1910-07-09" collectionCode="W, WU" collectorName="Trockene Hange &amp; von Kjachta &amp; den Euphrat &amp; Kalk &amp; Handel-Mazzetti" country="Turkey" county="Type" elevation="650" location="Lilan Tschai im Plateau" municipality="Karamuhara am Lilan Tschai" pageId="109" specimenCode="C7" specimenCount="2" stateProvince="Adiyaman" typeStatus="lectotype">Handel-Mazzetti (1913: 450)</materialsCitation>
</taxonomicName>
.
<collectingCounty id="629F4E6EDF7FFFA11A773B8CF22AFA4A" box="[811,868,1456,1481]" pageId="109">Type</collectingCounty>
:—
<collectingCountry id="F3567672DF7FFFA11AD93B8CF2B6FA4A" box="[901,1016,1456,1481]" name="Turkey" pageId="109">TURKEY</collectingCountry>
.
<specimenCode id="DBE79E99DF7FFFA11D543B8CF567FA4A" box="[1032,1065,1456,1481]" collectionCode="C" country="Denmark" name="University of Copenhagen" pageId="109" type="Herbarium">C7</specimenCode>
<collectingRegion id="4985F800DF7FFFA11D693B8CF5E0FA4A" box="[1077,1198,1456,1481]" country="Turkey" name="Adiyaman" pageId="109">Adiyaman</collectingRegion>
:
<collectorName id="26B45334DF7FFFA11DE13B8CF432FA4A" box="[1213,1404,1456,1481]" pageId="109">Trockene Hänge</collectorName>
bei
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Karamuhara am
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im Plateau südlich
<collectorName id="26B45334DF7FFFA11BED3BE8F274FA6E" box="[689,826,1492,1517]" pageId="109">von Kjachta</collectorName>
(Kahta) gegen
<collectorName id="26B45334DF7FFFA11AB73BE8F536FA6E" box="[1003,1144,1492,1517]" pageId="109">den Euphrat</collectorName>
,
<collectorName id="26B45334DF7FFFA11DDF3BE8F5F2FA6E" box="[1155,1212,1492,1517]" pageId="109">Kalk</collectorName>
,
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<elevation id="006CD1D1DF7FFFA11D953BE8F45DFA6E" box="[1225,1299,1492,1517]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="109" unit="m" value="650.0">650 m</elevation>
</quantity>
,
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<collectingDate id="EFBBE9CADF7FFFA11C7C3BE8F4E6FA6E" box="[1312,1448,1492,1517]" pageId="109" value="1910-07-09">9 July 1910</collectingDate>
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,
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<collectorName id="26B45334DF7FFFA119D43BC4F00DF992" box="[136,323,1528,1553]" pageId="109">Handel-Mazzetti</collectorName>
1953
</emphasis>
(
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W! designated by
<bibRefCitation id="EFD04B13DF7FFFA11B973BC4F2E1F992" author="Wagenitz, G." box="[715,943,1528,1553]" pageId="109" pagination="469 - 494" refId="ref69162" refString="Wagenitz, G. (1960) Centaurea L. sect. Cynaropsis, eine neue Sektion der Gattung aus Vorderasien. Willdenowia 2: 469 - 494." type="journal article" year="1960">Wagenitz 1960: 489</bibRefCitation>
,
<typeStatus id="54FA8840DF7FFFA11AE63BC4F50FF992" box="[954,1089,1528,1553]" pageId="109" type="isolectotype">isolectotype</typeStatus>
WU!). (
<figureCitation id="137A2A67DF7FFFA11DFC3BC4F5BFF992" box="[1184,1265,1528,1553]" captionStart="FIGURE 75" captionStartId="110.[136,229,1953,1974]" captionTargetBox="[228,1310,218,1902]" captionTargetId="figure-17@110.[228,1359,190,1929]" captionTargetPageId="110" captionText="FIGURE 75. Centaurea tomentella Handel-Mazzetti (Handel-Mazzetti 1953, W). Scale 2 cm." figureDoi="http://doi.org/10.5281/zenodo.13703618" httpUri="https://zenodo.org/record/13703618/files/figure.png" pageId="109">Fig. 75</figureCitation>
)
</paragraph>
</subSubSection>
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<paragraph id="8BFE36E2DF7FFFA319D4387CF536FE12" blockId="109.[136,1452,1600,2093]" lastBlockId="111.[136,1452,160,509]" lastPageId="111" pageId="109">
Biennial plants, with thick fleshy taproot, whole plant usually green opaque or green-brownish,
<quantity id="4CB99B07DF7FFFA11DEA387CF47DF9DA" box="[1206,1331,1600,1625]" metricMagnitude="-1" metricUnit="m" metricValue="6.25" metricValueMax="10.0" metricValueMin="2.5" pageId="109" unit="cm" value="62.5" valueMax="100.0" valueMin="25.0">25100 cm</quantity>
tall; collar of fibrous petiolar remains present at stem base. Stem erect, simple, sometimes branched from near base,
<quantity id="4CB99B07DF7FFFA11C633858F4E2F9FE" box="[1343,1452,1636,1661]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="15.0" metricValueMin="4.0" pageId="109" unit="mm" value="9.5" valueMax="15.0" valueMin="4.0">415 mm</quantity>
in diam. at base, cylindrical, whitish, loosely leafy throughout, densely covered with white-tomentose hairs. Leaves firm, undivided, basal and lower ones sparsely covered with hirsute-articulate hairs, more densely along midrib and veins, sometimes mixed with tomentose hairs, other leaves densely covered with white-tomentose hairs. Basal leaves usually with a long petiole (up to
<quantity id="4CB99B07DF7FFFA11B5938C8F304F88E" box="[517,586,1780,1805]" metricMagnitude="-1" metricUnit="m" metricValue="2.2" pageId="109" unit="cm" value="22.0">22 cm</quantity>
long), sometimes withered at anthesis, oblong-cordate, rarely elliptic, 1733 ×
<quantity id="4CB99B07DF7FFFA119FE3924F1B2F8B2" box="[162,252,1816,1841]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="109" unit="cm" value="7.0" valueMax="8.0" valueMin="6.0">68 cm</quantity>
, entire, acute at apex. Lower cauline leaves simple, lanceolate or oblong, 1320 ×
<quantity id="4CB99B07DF7FFFA11D973924F46BF8B2" box="[1227,1317,1816,1841]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="109" unit="cm" value="5.0" valueMax="6.0" valueMin="4.0">46 cm</quantity>
, sometimes with basal auricles, subacute or acute at apex. Median cauline leaves sessile, oblong or oblong-lanceolate, 713 ×
<quantity id="4CB99B07DF7FFFA119D4395CF1ACF8FA" box="[136,226,1888,1913]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="109" unit="cm" value="3.5" valueMax="5.0" valueMin="2.0">25 cm</quantity>
, narrowly decurrent, up to
<quantity id="4CB99B07DF7FFFA11B7E395CF328F8FA" box="[546,614,1888,1913]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="109" unit="mm" value="6.0">6 mm</quantity>
along stem, entire, sometimes denticulate, acute at apex. Upper cauline leaves increasingly smaller, sessile, lanceolate, narrowly lanceolate or lanceolate-linear, 39 ×
<quantity id="4CB99B07DF7FFFA11D9D39B8F460F81E" box="[1217,1326,1924,1949]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="3.0" metricValueMin="0.5" pageId="109" unit="cm" value="1.75" valueMax="3.0" valueMin="0.5">0.53 cm</quantity>
, decurrent, up to
<quantity id="4CB99B07DF7FFFA119953994F057F842" box="[201,281,1960,1985]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="109" unit="mm" value="12.0">12 mm</quantity>
along stem, entire, mucronate at apex (
<quantity id="4CB99B07DF7FFFA11B8B3994F20DF842" box="[727,835,1960,1985]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="109" unit="mm" value="7.5" valueMax="10.0" valueMin="5.0">510 mm</quantity>
long). Capitula numerous, variable in number, usually 310 sometimes up to 70, arranged in a raceme or spike, sessile or with peduncles usually short, up to
<quantity id="4CB99B07DF7FFFA11C6C39F0F427F866" box="[1328,1385,1996,2021]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="109" unit="cm" value="8.0">8 cm</quantity>
long. Involucres broadly ovoid or ovoid-subglobose, truncate at base, 3040 ×
<quantity id="4CB99B07DF7FFFA11A9F39CCF50EF78A" box="[963,1088,2032,2057]" metricMagnitude="-2" metricUnit="m" metricValue="3.15" metricValueMax="3.5" metricValueMin="2.8" pageId="109" unit="mm" value="31.5" valueMax="35.0" valueMin="28.0">2835 mm</quantity>
. Phyllaries multiseriate, yellow-greenish, imbricate, coriaceous-scarious, glabrous. Appendages very firm, totally concealing phyllaries, not decurrent, triangular, yellow and shiny,
<quantity id="4CB99B07DF7DFFA318953E9CF36BFF3A" box="[457,549,160,185]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="111" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">45 mm</quantity>
wide at base (excluding cilia); cilia patent, numerous, 610 on each side, 56(8) mm long; spines narrowly triangular, (7)912(17) mm long, longer than nearest cilia. Flowers pink or purple; central florets hermaphroditic,
<quantity id="4CB99B07DF7DFFA318D13ED4F34BFE82" box="[397,517,232,257]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.2" metricValueMin="2.8" pageId="111" unit="mm" value="30.0" valueMax="32.0" valueMin="28.0">2832 mm</quantity>
long, corolla
<quantity id="4CB99B07DF7DFFA31BC03ED4F25AFE82" box="[668,788,232,257]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="111" unit="mm" value="12.5" valueMax="13.0" valueMin="12.0">1213 mm</quantity>
long, 5-lobed, lobes
<quantity id="4CB99B07DF7DFFA31AA73ED4F519FE82" box="[1019,1111,232,257]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="111" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">56 mm</quantity>
long; peripheral florets sterile, nearly equal to central ones, numerous (
<quantity id="4CB99B07DF7DFFA31B0E3F30F3F6FEA6" box="[594,696,268,293]" metricMagnitude="-1" metricUnit="m" metricValue="4.445" metricValueMax="5.08" metricValueMin="3.81" pageId="111" unit="in" value="17.5" valueMax="20.0" valueMin="15.0">1520 in</quantity>
each capitulum) and conspicuous, finely dissected, not radiant, 4- lobed, limb lobes linear. Achenes lanceolate, ca.
<quantity id="4CB99B07DF7DFFA31BF53F0CF255FECA" box="[681,795,304,329]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="111" unit="mm" value="5.25" valueMax="5.5" valueMin="5.0">55.5 mm</quantity>
long,
<quantity id="4CB99B07DF7DFFA31A3D3F0CF29DFECA" box="[865,979,304,329]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" metricValueMax="3.0" metricValueMin="2.8" pageId="111" unit="mm" value="2.9" valueMax="3.0" valueMin="2.8">2.83 mm</quantity>
wide, smooth and shiny, whitish, rounded at apex, glabrous; insertion areole lateral, ca.
<quantity id="4CB99B07DF7DFFA31B233F68F39BFEEE" box="[639,725,340,365]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="111" unit="mm" value="0.8">0.8 mm</quantity>
long. Pappus double, persistent, multiseriate, scabrous, brownish or whitish,
<quantity id="4CB99B07DF7DFFA3185B3F44F03CFE12" box="[263,370,376,401]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="111" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">810 mm</quantity>
long, bristles of inner rows much shorter than others, ca.
<quantity id="4CB99B07DF7DFFA31AAB3F44F576FE12" box="[1015,1080,376,401]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="111" unit="mm" value="2.0">2 mm</quantity>
long.
</paragraph>
<caption id="DF3E666ADF7CFFA219D4399DF55FF835" ID-DOI="http://doi.org/10.5281/zenodo.13703618" ID-Zenodo-Dep="13703618" box="[136,1041,1953,1974]" httpUri="https://zenodo.org/record/13703618/files/figure.png" pageId="110" startId="110.[136,229,1953,1974]" targetBox="[228,1310,218,1902]" targetPageId="110" targetType="figure">
<paragraph id="8BFE36E2DF7CFFA219D4399DF55FF835" blockId="110.[136,1041,1953,1974]" box="[136,1041,1953,1974]" pageId="110">
<emphasis id="B935EAF0DF7CFFA219D4399DF044F835" bold="true" box="[136,266,1953,1974]" pageId="110">FIGURE 75.</emphasis>
<taxonomicName id="4C414D61DF7CFFA21853399DF3C8F835" authority="Handel-Mazzetti" authorityName="Handel-Mazzetti" authorityYear="1913" box="[271,646,1953,1974]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="110" phylum="Tracheophyta" rank="species" species="tomentella">
<emphasis id="B935EAF0DF7CFFA21853399DF091F835" box="[271,479,1953,1974]" italics="true" pageId="110">Centaurea tomentella</emphasis>
Handel-Mazzetti
</taxonomicName>
(
<emphasis id="B935EAF0DF7CFFA21BC8399DF224F835" box="[660,874,1953,1974]" italics="true" pageId="110">Handel-Mazzetti 1953</emphasis>
, W). Scale 2 cm.
</paragraph>
</caption>
<paragraph id="8BFE36E2DF7DFFA319E13FA0F5DFFE7E" blockId="111.[136,1452,160,509]" pageId="111">
<emphasis id="B935EAF0DF7DFFA319E13FA0F3C6FE36" bold="true" box="[189,648,412,437]" pageId="111">Taxonomic and distribution remarks:</emphasis>
<taxonomicName id="4C414D61DF7DFFA31BF83FA0F2D7FE36" authorityName="Handel-Mazzetti" authorityYear="1913" box="[676,921,412,437]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="111" phylum="Tracheophyta" rank="species" species="tomentella">
<emphasis id="B935EAF0DF7DFFA31BF83FA0F2D7FE36" box="[676,921,412,437]" italics="true" pageId="111">Centaurea tomentella</emphasis>
</taxonomicName>
<materialsCitation id="3B293CBFDF7DFFA31AFF3FA0F5C0FE7E" collectionCode="S" country="Turkey" county="Irano-Turanian" elevation="1025" location="Maras Provinces" municipality="It" pageId="111" specimenCount="1" stateProvince="Adiyaman">
is endemic to
<collectingRegion id="4985F800DF7DFFA31D1B3FA0F5F1FE36" box="[1095,1215,412,437]" country="Turkey" name="Adiyaman" pageId="111">Adiyaman</collectingRegion>
,
<collectingRegion id="4985F800DF7DFFA31D913FA0F467FE36" box="[1229,1321,412,437]" country="Turkey" name="Malatya" pageId="111">Malatya</collectingRegion>
and
<location id="8E9E6039DF7DFFA31C393FA0F1B7FE5A" LSID="urn:lsid:plazi:treatment:03E887F4DF7FFFA319D43B8CF01CF9FF:8E9E6039DF7DFFA31C393FA0F1B7FE5A" country="Turkey" county="Irano-Turanian" municipality="It" name="Maras Provinces" pageId="111" stateProvince="Adiyaman">Maraş Provinces</location>
,
<collectionCode id="ED50AE27DF7DFFA318593FFCF05BFE5A" box="[261,277,448,473]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="111" type="Herbarium">S</collectionCode>
<collectingCountry id="F3567672DF7DFFA318403FFCF022FE5A" box="[284,364,448,473]" name="Turkey" pageId="111">Turkey</collectingCountry>
(
<figureCitation id="137A2A67DF7DFFA318203FFCF09FFE5A" box="[380,465,448,473]" captionStart="FIGURE 76" captionStartId="111.[136,229,1314,1335]" captionTargetBox="[159,1391,546,1268]" captionTargetId="figure-184@111.[151,1436,533,1289]" captionTargetPageId="111" captionText="FIGURE 76. Distribution of Centaurea tomentella (squares), C. zaferii (circles), and C. zarrei (triangles)." figureDoi="http://doi.org/10.5281/zenodo.13703620" httpUri="https://zenodo.org/record/13703620/files/figure.png" pageId="111">Fig. 76</figureCitation>
).
<collectingMunicipality id="6B9AAC98DF7DFFA318B93FFCF0B8FE5A" box="[485,502,448,473]" pageId="111">It</collectingMunicipality>
is an
<collectingCounty id="629F4E6EDF7DFFA31B663FFCF3ADFE5A" box="[570,739,448,473]" pageId="111">Irano-Turanian</collectingCounty>
element, and grows on dry slopes, fallow fields and vineyards, at elevations of
<quantity id="4CB99B07DF7DFFA318663FD8F084FE7E" box="[314,458,484,509]" metricMagnitude="3" metricUnit="m" metricValue="1.025" metricValueMax="1.45" metricValueMin="0.6" pageId="111" unit="m" value="1025.0" valueMax="1450.0" valueMin="600.0">
<elevation id="006CD1D1DF7DFFA318663FD8F084FE7E" box="[314,458,484,509]" metricMagnitude="3" metricUnit="m" metricValue="1.025" metricValueMax="1.45" metricValueMin="0.6" pageId="111" unit="m" value="1025.0" valueMax="1450.0" valueMin="600.0">6001450 m</elevation>
</quantity>
. It closely resembles
<taxonomicName id="4C414D61DF7DFFA31BE63FD8F542FE7E" box="[698,1036,484,509]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="111" phylum="Tracheophyta" rank="subSpecies" species="cataonica" subSpecies="cataonica">
<emphasis id="B935EAF0DF7DFFA31BE63FD8F207FE7E" box="[698,841,484,509]" italics="true" pageId="111">C. cataonica</emphasis>
subsp.
<emphasis id="B935EAF0DF7DFFA31AC23FD8F542FE7E" box="[926,1036,484,509]" italics="true" pageId="111">cataonica</emphasis>
</taxonomicName>
in its habit
</materialsCitation>
.
</paragraph>
<caption id="DF3E666ADF7DFFA319D43B1EF5CAFAB4" ID-DOI="http://doi.org/10.5281/zenodo.13703620" ID-Zenodo-Dep="13703620" box="[136,1156,1314,1335]" httpUri="https://zenodo.org/record/13703620/files/figure.png" pageId="111" startId="111.[136,229,1314,1335]" targetBox="[159,1391,546,1268]" targetPageId="111" targetType="figure">
<paragraph id="8BFE36E2DF7DFFA319D43B1EF5CAFAB4" blockId="111.[136,1156,1314,1335]" box="[136,1156,1314,1335]" pageId="111">
<emphasis id="B935EAF0DF7DFFA319D43B1EF04DFAB4" bold="true" box="[136,259,1314,1335]" pageId="111">FIGURE 76</emphasis>
. Distribution of
<taxonomicName id="4C414D61DF7DFFA318FF3B1EF33DFAB4" authorityName="Handel-Mazzetti" authorityYear="1913" box="[419,627,1314,1335]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="111" phylum="Tracheophyta" rank="species" species="tomentella">
<emphasis id="B935EAF0DF7DFFA318FF3B1EF33DFAB4" box="[419,627,1314,1335]" italics="true" pageId="111">Centaurea tomentella</emphasis>
</taxonomicName>
(squares),
<taxonomicName id="4C414D61DF7DFFA31B813B1EF278FAB4" authorityName="Negaresh" authorityYear="2017" box="[733,822,1314,1335]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="111" phylum="Tracheophyta" rank="species" species="zaferii">
<emphasis id="B935EAF0DF7DFFA31B813B1EF278FAB4" box="[733,822,1314,1335]" italics="true" pageId="111">C. zaferii</emphasis>
</taxonomicName>
(circles), and
<taxonomicName id="4C414D61DF7DFFA31A9C3B1EF55AFAB4" box="[960,1044,1314,1335]" class="Magnoliopsida" family="Asteraceae" genus="Cynaroides" higherTaxonomySource="GBIF" kingdom="Plantae" order="Asterales" pageId="111" phylum="Tracheophyta" rank="species" species="zarrei">
<emphasis id="B935EAF0DF7DFFA31A9C3B1EF55AFAB4" box="[960,1044,1314,1335]" italics="true" pageId="111">C. zarrei</emphasis>
</taxonomicName>
(triangles).
</paragraph>
</caption>
<paragraph id="8BFE36E2DF7DFFA319E13B5BF35AFA27" blockId="111.[136,1452,1383,1660]" pageId="111">
<emphasis id="B935EAF0DF7DFFA319E13B5BF098FA03" bold="true" box="[189,470,1383,1408]" pageId="111">Chromosome numbers:</emphasis>
<bibRefCitation id="EFD04B13DF7DFFA318AF3B5BF38CFA03" author="Uysal, T. &amp; Ertugrul, K. &amp; Susanna, A. &amp; Garcia-Jacas, N." box="[499,706,1383,1408]" pageId="111" pagination="280 - 286" refId="ref69101" refString="Uysal, T., Ertugrul, K., Susanna, A. &amp; Garcia-Jacas, N. (2009) New chromosome counts in the genus Centaurea (Asteraceae) from Turkey. Botanical Journal of the Linnean Society 159: 280 - 286." type="journal article" year="2009">
Uysal
<emphasis id="B935EAF0DF7DFFA31B663B5BF33FFA03" box="[570,625,1383,1408]" italics="true" pageId="111">et al.</emphasis>
(2009)
</bibRefCitation>
and
<bibRefCitation id="EFD04B13DF7DFFA31BA93B5BF282FA03" author="Askoy, H. &amp; Berber, A. A. &amp; Celik, S. &amp; Baser, B. &amp; Karaveliogullari, F. A." box="[757,972,1383,1408]" pageId="111" pagination="285 - 294" refId="ref65175" refString="Askoy, H., Berber, A. A., Celik, S., Baser, B. &amp; Karaveliogullari, F. A. (2016) Karyological study on 12 Centaurea (Asteraceae) species from Turkey. Caryologia 69 (4): 285 - 294. http: // doi. org / 10.1080 / 00087114.2016.1188357" type="journal article" year="2016">
Askoy
<emphasis id="B935EAF0DF7DFFA31A183B5BF235FA03" box="[836,891,1383,1408]" italics="true" pageId="111">et al.</emphasis>
(2016)
</bibRefCitation>
reported the same chromosome number 2
<emphasis id="B935EAF0DF7DFFA31CC13B5BF4E5FA03" box="[1437,1451,1383,1408]" italics="true" pageId="111">n</emphasis>
= 2
<emphasis id="B935EAF0DF7DFFA319F13BB7F1F7FA27" box="[173,185,1419,1444]" italics="true" pageId="111">x</emphasis>
=18 for
<taxonomicName id="4C414D61DF7DFFA318473BB7F343FA27" authorityName="Handel-Mazzetti" authorityYear="1913" box="[283,525,1419,1444]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="111" phylum="Tracheophyta" rank="species" species="tomentella">
<emphasis id="B935EAF0DF7DFFA318473BB7F343FA27" box="[283,525,1419,1444]" italics="true" pageId="111">Centaurea tomentella</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C35B6569DF7DFFA319E13B93F01CF9FF" pageId="111" type="materials_examined">
<paragraph id="8BFE36E2DF7DFFA319E13B93F01CF9FF" blockId="111.[136,1452,1383,1660]" pageId="111">
<emphasis id="B935EAF0DF7DFFA319E13B93F309FA4B" bold="true" box="[189,583,1455,1480]" pageId="111">Selected specimens examined:—</emphasis>
<materialsCitation id="3B293CBFDF7DFFA31B1B3B93F3BFFA4B" box="[583,753,1455,1480]" collectionCode="B" country="Turkey" location="TURKEY" pageId="111" specimenCode="B6" specimenCount="1">
<emphasis id="B935EAF0DF7DFFA31B1B3B93F3F1FA4B" bold="true" box="[583,703,1455,1480]" pageId="111">
<collectingCountry id="F3567672DF7DFFA31B1B3B93F3F1FA4B" box="[583,703,1455,1480]" name="Turkey" pageId="111">TURKEY</collectingCountry>
</emphasis>
.
<specimenCode id="DBE79E99DF7DFFA31B8C3B93F3BFFA4B" box="[720,753,1455,1480]" collectionCode="B" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="111" type="Herbarium">B6</specimenCode>
</materialsCitation>
<materialsCitation id="3B293CBFDF7DFFA31BA03B93F02CF9B7" collectingDate="1952-07-23" collectingDateMax="2001-08-19" collectingDateMin="1952-07-23" collectionCode="E, S" collectorName="Reeves &amp; Adiguzel" country="Turkey" county="Maras" elevation="1270" location="Davis" municipality="Findik" pageId="111" specimenCount="1" stateProvince="Kahramanmaras">
<collectingCounty id="629F4E6EDF7DFFA31BA03B93F20CFA4B" box="[764,834,1455,1480]" pageId="111">Maraş</collectingCounty>
(
<collectingRegion id="4985F800DF7DFFA31A083B93F543FA4B" box="[852,1037,1455,1480]" country="Turkey" name="Kahramanmaras" pageId="111">Kahramanmaraş</collectingRegion>
): near
<collectingMunicipality id="6B9AAC98DF7DFFA31D383B93F5FEFA4B" box="[1124,1200,1455,1480]" pageId="111">Findik</collectingMunicipality>
,
<locationDeviation id="9A4C2FC0DF7DFFA31DE33B93F1ACFA6F" firstLocation="Cardak" pageId="111" secondLocation="Goksun">between Çardak and Göksun</locationDeviation>
,
<date id="FFFF1022DF7DFFA319B33BEFF0C6FA6F" box="[239,392,1491,1516]" pageId="111" value="1952-07-23">
<collectingDate id="EFBBE9CADF7DFFA319B33BEFF0C6FA6F" box="[239,392,1491,1516]" pageId="111" value="1952-07-23">23 July 1952</collectingDate>
</date>
,
<emphasis id="B935EAF0DF7DFFA318C93BEFF36BFA6F" box="[405,549,1491,1516]" italics="true" pageId="111">
<location id="8E9E6039DF7DFFA318C93BEFF098FA6F" LSID="urn:lsid:plazi:treatment:03E887F4DF7FFFA319D43B8CF01CF9FF:8E9E6039DF7DFFA318C93BEFF098FA6F" box="[405,470,1491,1516]" country="Turkey" county="Maras" municipality="Findik" name="Davis" pageId="111" stateProvince="Kahramanmaras">Davis</location>
20240
</emphasis>
(
<collectionCode id="ED50AE27DF7DFFA31B653BEFF31EFA6F" box="[569,592,1491,1516]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="111" type="Herbarium">E</collectionCode>
!);
<collectionCode id="ED50AE27DF7DFFA31B313BEFF333FA6F" box="[621,637,1491,1516]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="111" type="Herbarium">S</collectionCode>
of
<location id="8E9E6039DF7DFFA31BFA3BEFF22CFA6F" LSID="urn:lsid:plazi:treatment:03E887F4DF7FFFA319D43B8CF01CF9FF:8E9E6039DF7DFFA31BFA3BEFF22CFA6F" box="[678,866,1491,1516]" country="Turkey" county="Maras" municipality="Findik" name="Goksun-Elbistan" pageId="111" stateProvince="Kahramanmaras">Göksun-Elbistan</location>
road,
<collectionCode id="ED50AE27DF7DFFA31AF23BEFF2F1FA6F" box="[942,959,1491,1516]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="111" type="Herbarium">E</collectionCode>
of
<location id="8E9E6039DF7DFFA31AB53BEFF573FA6F" LSID="urn:lsid:plazi:treatment:03E887F4DF7FFFA319D43B8CF01CF9FF:8E9E6039DF7DFFA31AB53BEFF573FA6F" box="[1001,1085,1491,1516]" country="Turkey" county="Maras" municipality="Findik" name="Cardak" pageId="111" stateProvince="Kahramanmaras">Çardak</location>
,
<locationDeviation id="9A4C2FC0DF7DFFA31D163BEFF4E9FA6F" box="[1098,1447,1491,1516]" firstLocation="Dagdere" pageId="111" secondLocation="Soysalli">between Dagdere and Soysalli</locationDeviation>
,
<quantity id="4CB99B07DF7DFFA319D43BCBF1AFF993" box="[136,225,1527,1552]" metricMagnitude="3" metricUnit="m" metricValue="1.27" pageId="111" unit="m" value="1270.0">
<elevation id="006CD1D1DF7DFFA319D43BCBF1AFF993" box="[136,225,1527,1552]" metricMagnitude="3" metricUnit="m" metricValue="1.27" pageId="111" unit="m" value="1270.0">1270 m</elevation>
</quantity>
,
<date id="FFFF1022DF7DFFA319B33BCBF0E6F993" box="[239,424,1527,1552]" pageId="111" value="2001-08-19">
<collectingDate id="EFBBE9CADF7DFFA319B33BCBF0E6F993" box="[239,424,1527,1552]" pageId="111" value="2001-08-19">19 August 2001</collectingDate>
</date>
,
<emphasis id="B935EAF0DF7DFFA318EA3BCBF381F993" box="[438,719,1527,1552]" italics="true" pageId="111">
<collectorName id="26B45334DF7DFFA318EA3BCBF34AF993" box="[438,516,1527,1552]" pageId="111">Reeves</collectorName>
&amp;
<collectorName id="26B45334DF7DFFA31B773BCBF3C3F993" box="[555,653,1527,1552]" pageId="111">Adigüzel</collectorName>
2320
</emphasis>
(
<collectionCode id="ED50AE27DF7DFFA31BBE3BCBF3B7F993" box="[738,761,1527,1552]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="111" type="Herbarium">E</collectionCode>
!); Çardak-Esence köyü,
<quantity id="4CB99B07DF7DFFA31D4D3BCBF524F993" box="[1041,1130,1527,1552]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="111" unit="m" value="1350.0">
<elevation id="006CD1D1DF7DFFA31D4D3BCBF524F993" box="[1041,1130,1527,1552]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="111" unit="m" value="1350.0">1350 m</elevation>
</quantity>
,
<date id="FFFF1022DF7DFFA31D243BCBF47FF993" box="[1144,1329,1527,1552]" pageId="111" value="2001-08-18">
<collectingDate id="EFBBE9CADF7DFFA31D243BCBF47FF993" box="[1144,1329,1527,1552]" pageId="111" value="2001-08-18">18 August 2001</collectingDate>
</date>
,
<emphasis id="B935EAF0DF7DFFA31C633BCBF064F9B7" italics="true" pageId="111">
<collectorName id="26B45334DF7DFFA31C633BCBF4C3F993" box="[1343,1421,1527,1552]" pageId="111">Reeves</collectorName>
&amp;
<collectorName id="26B45334DF7DFFA319D43827F1A4F9B7" box="[136,234,1563,1588]" pageId="111">Adigüzel</collectorName>
2317
</emphasis>
(
<collectionCode id="ED50AE27DF7DFFA318613827F01AF9B7" box="[317,340,1563,1588]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="111" type="Herbarium">E</collectionCode>
!)
</materialsCitation>
.
<materialsCitation id="3B293CBFDF7DFFA318333827F0DEF9B7" box="[367,400,1563,1588]" collectionCode="C" country="Turkey" pageId="111" specimenCode="C6" specimenCount="1">
<specimenCode id="DBE79E99DF7DFFA318333827F0DEF9B7" box="[367,400,1563,1588]" collectionCode="C" country="Denmark" name="University of Copenhagen" pageId="111" type="Herbarium">C6</specimenCode>
</materialsCitation>
<materialsCitation id="3B293CBFDF7DFFA318C43827F31FF9DB" collectingDate="1865-07-12" collectionCode="JE, ISTE" collectorName="Kahraman &amp; Marasch &amp; Haussknecht" country="Turkey" county="Maras" location="Haussknecht" municipality="Maras" pageId="111" specimenCount="1">
<collectingCounty id="629F4E6EDF7DFFA318C43827F090F9B7" box="[408,478,1563,1588]" pageId="111">Maraş</collectingCounty>
: near
<collectingMunicipality id="6B9AAC98DF7DFFA31B7A3827F323F9B7" box="[550,621,1563,1588]" pageId="111">Maraş</collectingMunicipality>
,
<date id="FFFF1022DF7DFFA31B203827F25CF9B7" box="[636,786,1563,1588]" pageId="111" value="1865-07-12">
<collectingDate id="EFBBE9CADF7DFFA31B203827F25CF9B7" box="[636,786,1563,1588]" pageId="111" value="1865-07-12">12 July 1865</collectingDate>
</date>
,
<emphasis id="B935EAF0DF7DFFA31A423827F292F9B7" box="[798,988,1563,1588]" italics="true" pageId="111">
<location id="8E9E6039DF7DFFA31A423827F2E3F9B7" LSID="urn:lsid:plazi:treatment:03E887F4DF7FFFA319D43B8CF01CF9FF:8E9E6039DF7DFFA31A423827F2E3F9B7" box="[798,941,1563,1588]" country="Turkey" county="Maras" municipality="Maras" name="Haussknecht" pageId="111">Haussknecht</location>
s.n.
</emphasis>
(
<collectionCode id="ED50AE27DF7DFFA31AB23827F55CF9B7" box="[1006,1042,1563,1588]" country="Germany" lsid="urn:lsid:biocol.org:col:15444" name="Friedrich-Schiller-Universität Jena" pageId="111" type="Herbarium">JE</collectionCode>
!);
<collectorName id="26B45334DF7DFFA31D713827F5ECF9B7" box="[1069,1186,1563,1588]" pageId="111">Kahraman</collectorName>
maraş, prope
<collectorName id="26B45334DF7DFFA31C183827F4E9F9B7" box="[1348,1447,1563,1588]" pageId="111">Marasch</collectorName>
,
<emphasis id="B935EAF0DF7DFFA319D43803F009F9DB" box="[136,327,1599,1624]" italics="true" pageId="111">
<collectorName id="26B45334DF7DFFA319D43803F059F9DB" box="[136,279,1599,1624]" pageId="111">Haussknecht</collectorName>
s.n.
</emphasis>
(
<collectionCode id="ED50AE27DF7DFFA318053803F033F9DB" box="[345,381,1599,1624]" country="Germany" lsid="urn:lsid:biocol.org:col:15444" name="Friedrich-Schiller-Universität Jena" pageId="111" type="Herbarium">JE</collectionCode>
!);
<emphasis id="B935EAF0DF7DFFA318C43803F0A3F9DB" box="[408,493,1599,1624]" italics="true" pageId="111">s.c. s.n.</emphasis>
(
<collectionCode id="ED50AE27DF7DFFA318A33803F30AF9DB" box="[511,580,1599,1624]" country="Turkey" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14530" lsid="urn:lsid:biocol.org:col:14530" name="University of Istanbul" pageId="111" type="Herbarium">ISTE</collectionCode>
!)
</materialsCitation>
.
<materialsCitation id="3B293CBFDF7DFFA31B013803F330F9DB" box="[605,638,1599,1624]" collectionCode="B" country="Turkey" pageId="111" specimenCode="B7" specimenCount="1">
<specimenCode id="DBE79E99DF7DFFA31B013803F330F9DB" box="[605,638,1599,1624]" collectionCode="B" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="111" type="Herbarium">B7</specimenCode>
</materialsCitation>
<materialsCitation id="3B293CBFDF7DFFA31BDB3803F000F9FF" collectorName="Huber-Morath &amp; Hb. Hub. &amp; Mor." country="Turkey" elevation="950" location="Puturge" pageId="111" specimenCount="1" stateProvince="Malatya">
<collectingRegion id="4985F800DF7DFFA31BDB3803F3A9F9DB" box="[647,743,1599,1624]" country="Turkey" name="Malatya" pageId="111">Malatya</collectingRegion>
:
<quantity id="4CB99B07DF7DFFA31BAF3803F272F9DB" box="[755,828,1599,1624]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="111" unit="km" value="18.0">18 km</quantity>
from
<collectingRegion id="4985F800DF7DFFA31AD83803F2AEF9DB" box="[900,992,1599,1624]" country="Turkey" name="Malatya" pageId="111">Malatya</collectingRegion>
to
<location id="8E9E6039DF7DFFA31D5B3803F52FF9DB" LSID="urn:lsid:plazi:treatment:03E887F4DF7FFFA319D43B8CF01CF9FF:8E9E6039DF7DFFA31D5B3803F52FF9DB" box="[1031,1121,1599,1624]" country="Turkey" name="Puturge" pageId="111" stateProvince="Malatya">Pütürge</location>
,
<quantity id="4CB99B07DF7DFFA31D323803F5F7F9DB" box="[1134,1209,1599,1624]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="111" unit="m" value="950.0">
<elevation id="006CD1D1DF7DFFA31D323803F5F7F9DB" box="[1134,1209,1599,1624]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="111" unit="m" value="950.0">950 m</elevation>
</quantity>
,
<emphasis id="B935EAF0DF7DFFA31D9A3803F4E5F9DB" box="[1222,1451,1599,1624]" italics="true" pageId="111">
<collectorName id="26B45334DF7DFFA31D9A3803F424F9DB" box="[1222,1386,1599,1624]" pageId="111">Huber-Morath</collectorName>
9297
</emphasis>
(
<collectorName id="26B45334DF7DFFA319CC385FF1B3F9FF" box="[144,253,1635,1660]" pageId="111">Hb. Hub.</collectorName>
-
<collectorName id="26B45334DF7DFFA31859385FF00CF9FF" box="[261,322,1635,1660]" pageId="111">Mor.</collectorName>
!)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>