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<document id="8F71F7B50D5F3716E089B70E4BE1E554" ID-DOI="10.11646/phytotaxa.363.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13703414" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725581356715" checkinUser="felipe" docAuthor="Negaresh, Kazem &amp; Rahiminejad, Mohammad Reza" docDate="2018" docId="03E887F4DF52FF8F19D43D4CF327FDA2" docLanguage="en" docName="phytotaxa.363.1.1.pdf" docOrigin="Phytotaxa 363 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.363.1.1" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.4:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="4" docTitle="Centaurea kamalnejadii Negaresh 2018, sp. nov." docType="treatment" docVersion="2" lastPageNumber="450" masterDocId="FFD1FF8CDF12FFCC195C3E3CF14EFF83" masterDocTitle="A revision of Centaurea sect. Cynaroides (Asteraceae, Cardueae-Centaureinae)" masterLastPageNumber="450" masterPageNumber="448" pageNumber="448" updateTime="1725593506943" updateUser="ExternalLinkService">
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<mods:title id="1CAD57138B43587D6ED7EFA56F0C0317">A revision of Centaurea sect. Cynaroides (Asteraceae, Cardueae-Centaureinae)</mods:title>
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<mods:name id="879773AB46B8FC07A3D99B0DB0604B80" type="personal">
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<mods:namePart id="99B3BC9BD2E5A2A63C18153052D22F98">Negaresh, Kazem</mods:namePart>
<mods:affiliation id="69F4B172BEFE7E130EDC1F91A89C1F21">Department of Horticulture, Faculty of Agriculture, Khuzestan Agricultural Sciences and Natural Resources University, Mollasani, Ahvaz, Iran. (* Corresponding author e-mail: negaresh @ ramin. ac. ir)</mods:affiliation>
<mods:nameIdentifier id="4931D605F1B2BC35F76FD60436EEFBE0" type="email">negaresh@ramin.ac.ir</mods:nameIdentifier>
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<mods:namePart id="D362B0566C27DD2117241B88D61B95BF">Rahiminejad, Mohammad Reza</mods:namePart>
<mods:affiliation id="F7BEBA58F80D016BABE7F636B2F60BEA">Department of Horticulture, Faculty of Agriculture, Khuzestan Agricultural Sciences and Natural Resources University, Mollasani, Ahvaz, Iran. (* Corresponding author e-mail: negaresh @ ramin. ac. ir) &amp; Department of Biology, University of Isfahan, Isfahan 81746 - 73441, Iran. &amp; Department of Horticulture, Faculty of Agriculture, Khuzestan Agricultural Sciences and Natural Resources University, Mollasani, Ahvaz, Iran. (* Corresponding author e-mail: negaresh @ ramin. ac. ir)</mods:affiliation>
<mods:nameIdentifier id="4BBDA8DEAF0CE3CCA3B8CC61FB45159B" type="email">negaresh@ramin.ac.ir</mods:nameIdentifier>
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<mods:title id="A296C1985AEA9DD9A5BA999EDE52F4A7">Phytotaxa</mods:title>
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<mods:date id="C265BD5578EB83B82A570B268E49621E">2018</mods:date>
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<subSubSection id="C35B6569DF52FF8C19D43D4CF245FC0A" box="[136,779,878,905]" pageId="64" type="nomenclature">
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<emphasis id="B935EAF0DF52FF8C19D43D4CF1E5FC0A" bold="true" box="[136,171,880,905]" pageId="64">19.</emphasis>
<taxonomicName id="4C414D61DF52FF8C19EE3D52F373FC0A" ID-CoL="5XGB9" authority="Negaresh" authorityName="Negaresh" authorityYear="2018" box="[178,573,878,905]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="64" phylum="Tracheophyta" rank="species" species="kamalnejadii" status="sp. nov.">
<emphasis id="B935EAF0DF52FF8C19EE3D52F087FC0A" bold="true" box="[178,457,878,905]" italics="true" pageId="64">Centaurea kamalnejadii</emphasis>
Negaresh
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,
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<taxonomicNameLabel id="A206578BDF52FF8C1B143D4CF3ECFC0A" box="[584,674,880,905]" pageId="64" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="137A2A67DF52FF8C1BED3D4CF24CFC0A" box="[689,770,880,905]" captionStart="FIGURE 44" captionStartId="65.[136,229,1953,1974]" captionTargetBox="[185,1402,190,1929]" captionTargetId="figure-14@65.[185,1402,190,1929]" captionTargetPageId="65" captionText="FIGURE 44. Centaurea kamalnejadii Negaresh (Negaresh &amp; Kamalnejad 130, herb. Negaresh). A. close up of capitule. B. capitule (with above view). C. habit and habitat. D. inflorescence. Scale A = 2 cm, B = 5 cm, D = 4 cm." figureDoi="http://doi.org/10.5281/zenodo.13703524" httpUri="https://zenodo.org/record/13703524/files/figure.png" pageId="64">Fig. 44</figureCitation>
)
</paragraph>
</subSubSection>
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<emphasis id="B935EAF0DF52FF8C19D43DAAF27BFB24" italics="true" pageId="64">
<taxonomicName id="4C414D61DF52FF8C19D43DAAF088FC28" box="[136,454,918,939]" class="Magnoliopsida" family="Asteraceae" genus="Centaureae" kingdom="Plantae" order="Asterales" pageId="64" phylum="Tracheophyta" rank="subSpecies" species="kabirkuhense" subSpecies="affinis">Centaureae kabirkuhense affinis</taxonomicName>
, a qua foliis basalibus et inferioribus caulinis ovatis vel lanceolatis, integris, ca. 24 ×
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(nec late cordatis ad late ovatis, basi lobis 12 minoribus vel integris, 2032 ×
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), involucro
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(nec
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) lato, appendicibus exterioribus elongatis triangularibus,
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longis, albidis, spinulis utrinque 23,
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longis (nec elongatis lanceolatis vel oblongo-lanceolatis,
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longis, brunneis ad atrobrunneis, integris), appendicibus phyllorum mediorum elongatis triangularibus ad spiniformibus, 2530 ×
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, purpureis ad stramineis, spinulis utrinque 12 et
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longis (nec elongatis lanceolatis vel oblongo-lanceolatis, 1530 ×
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, brunneis ad atrobrunneis, integris), appendicibus interioribus spiniformibus, 1325 ×
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, purpureis ad brunneis (nec elongatis lanceolatis vel oblongo-lanceolatis, 1530 ×
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), pappo non doublo,
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longo (nec doublo,
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longo) differt.
</emphasis>
</paragraph>
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:—
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. Prov.
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:
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toward
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, fallow field,
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,
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,
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<collectorName id="26B45334DF52FF8C1DCD3A8AF5A2FB48" box="[1169,1260,1206,1227]" pageId="64">Negaresh</collectorName>
&amp;
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130
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(
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herb. Negaresh!,
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!,
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!)
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.
</paragraph>
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Biennial herbs, with a thick freshly taproot, whole plant usually green, up to
<quantity id="4CB99B07DF52FF8C1D593B1CF502FABA" box="[1029,1100,1312,1337]" metricMagnitude="-1" metricUnit="m" metricValue="4.5" pageId="64" unit="cm" value="45.0">45 cm</quantity>
tall; collum neck with fibrous remnants of petioles from previous year. Stem stout, erect, almost simple, yellowish, ca.
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in diam. at base, cylindrical, finely white-striate, densely leafy in median part, lower part reddish and subglabrous, or sparsely covered with deciduous hirsute-articulate hairs (up to
<quantity id="4CB99B07DF52FF8C1BCD3BB0F39AFA26" box="[657,724,1420,1445]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="64" unit="mm" value="3.0">3 mm</quantity>
long), median part loosely covered with arachnoid hairs mixed with hirsute-articulate hairs, upper part densely covered with arachnoid hairs. Leaves rigid, papyraceous (on drying), undivided, covered with hirsute-articulate hairs, denser along midrib and veins, veins prominent on both sides. Basal and lower cauline leaves petiolate, withered at anthesis, ovate or lanceolate, ca. 24 ×
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, acute at apex. Median cauline leaves sessile, lanceolate or oblanceolate, narrowed toward base, 1420 ×
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, broadly decurrent, up to
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along stem, scabrous or remotely denticulate, acute or subacuminate at apex. Upper cauline leaves increasingly smaller, sessile, lanceolate or narrowly lanceolate, (5)815 × (0.5)
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, loosely covered with arachnoid hairs, decurrent, up to
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along stem, entire, acuminate or cuspidate at apex. Capitula numerous, 7 to 10, arranged in raceme, upper capitula on peduncles (3)
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long, sometimes with bracts similar to phyllaries with small stramineous and chaffy appendages. Involucres ovoid to subglobose, convex at base, ca. 50 ×
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. Phyllaries multiseriate, green in vivo, green-yellowish in sicco, imbricate, coriaceous-scarious, densely floccose-tomentose; appendages partially concealing phyllaries, rigid or chartaceous, not decurrent. Outer phyllaries oblong, 78 × ca.
<quantity id="4CB99B07DF52FF8C19D43900F180F8D6" box="[136,206,1852,1877]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="64" unit="mm" value="6.0">6 mm</quantity>
; appendages elongate triangular, 1023 × ca.
<quantity id="4CB99B07DF52FF8C1B843900F276F8D6" box="[728,824,1852,1877]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="64" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">45 mm</quantity>
, with 23 (
<quantity id="4CB99B07DF52FF8C1AE63900F57EF8D6" box="[954,1072,1852,1877]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="64" unit="mm" value="2.25" valueMax="3.0" valueMin="1.5">1.53 mm</quantity>
) spinules on each side, gradually narrowed into a
<quantity id="4CB99B07DF52FF8C1818395CF0FFF8FA" box="[324,433,1888,1913]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="64" unit="mm" value="7.5" valueMax="10.0" valueMin="5.0">510 mm</quantity>
spine, white. Median phyllaries lanceolate, 1225 × ca.
<quantity id="4CB99B07DF52FF8C1D64395CF5FAF8FA" box="[1080,1204,1888,1913]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="64" unit="mm" value="11.5" valueMax="12.0" valueMin="11.0">1112 mm</quantity>
; appendages elongate triangular to spiniform, 2530 ×
<quantity id="4CB99B07DF52FF8C18A439B8F319F81E" box="[504,599,1924,1949]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="64" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
, with 12 (
<quantity id="4CB99B07DF52FF8C1B8939B8F27AF81E" box="[725,820,1924,1949]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="64" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">35 mm</quantity>
) spinules on each side, gradually narrowed into a
<quantity id="4CB99B07DF52FF8C1C3939B8F1FAF842" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="64" unit="mm" value="12.5" valueMax="15.0" valueMin="10.0">1015 mm</quantity>
spine, purple to straw-coloured. Inner phyllaries lanceolate or narrowly lanceolate, 59 ×
<quantity id="4CB99B07DF52FF8C1DFE3994F450F842" box="[1186,1310,1960,1985]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="64" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">1012 mm</quantity>
; appendages spiniform, 1325 ×
<quantity id="4CB99B07DF52FF8C183B39F0F088F866" box="[359,454,1996,2021]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="64" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">12 mm</quantity>
, entire, purple to brown. Flowers purple; central florets hermaphroditic,
<quantity id="4CB99B07DF52FF8C1DAF39F0F422F866" box="[1267,1388,1996,2021]" metricMagnitude="-2" metricUnit="m" metricValue="5.85" metricValueMax="6.0" metricValueMin="5.7" pageId="64" unit="mm" value="58.5" valueMax="60.0" valueMin="57.0">5760 mm</quantity>
long, corolla ±
<quantity id="4CB99B07DF52FF8C19AD39CCF071F78A" box="[241,319,2032,2057]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="64" unit="mm" value="22.0">22 mm</quantity>
long, 5-lobed, lobes
<quantity id="4CB99B07DF52FF8C1B7A39CCF3C1F78A" box="[550,655,2032,2057]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="64" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">810 mm</quantity>
long, with deep purple nerves, anthers pale purple, shorter than or equal to corolla, with apical appendages acute, stigma exserted from corolla up to
<quantity id="4CB99B07DF52FF8C1AB63628F573F7AE" box="[1002,1085,2068,2093]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="64" unit="mm" value="10.0">10 mm</quantity>
; peripheral florets sterile, nearly equal to central ones, numerous (
<quantity id="4CB99B07DF50FF8E18AA3E9CF314FF3A" box="[502,602,160,185]" metricMagnitude="-1" metricUnit="m" metricValue="4.445" metricValueMax="5.08" metricValueMin="3.81" pageId="66" unit="in" value="17.5" valueMax="20.0" valueMin="15.0">1520 in</quantity>
each capitulum) and conspicuous, finely dissected, slightly radiant, 5-lobed, limb lobe filiform, ca.
<quantity id="4CB99B07DF50FF8E18D03EF8F092FF5E" box="[396,476,196,221]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="66" unit="mm" value="10.0">10 mm</quantity>
long. Achenes oblong,
<quantity id="4CB99B07DF50FF8E1BB63EF8F210FF5E" box="[746,862,196,221]" metricMagnitude="-3" metricUnit="m" metricValue="6.75" metricValueMax="7.0" metricValueMin="6.5" pageId="66" unit="mm" value="6.75" valueMax="7.0" valueMin="6.5">6.57 mm</quantity>
long, ca.
<quantity id="4CB99B07DF50FF8E1A8C3EF8F2B2FF5E" box="[976,1020,196,221]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="66" unit="m" value="3.0">3 m</quantity>
wide, pale brown, glabrous; insertion areole lateral, yellow, ca.
<quantity id="4CB99B07DF50FF8E18F63ED4F0A2FE82" box="[426,492,232,257]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="66" unit="mm" value="1.0">1 mm</quantity>
long. Pappus persistent, multiseriate, scabrous, pale brown,
<quantity id="4CB99B07DF50FF8E1DC53ED4F45DFE82" box="[1177,1299,232,257]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="66" unit="mm" value="15.5" valueMax="16.0" valueMin="15.0">1516 mm</quantity>
long, bristles of inner rows slightly longer than others.
</paragraph>
</subSubSection>
<caption id="DF3E666ADF53FF8D19D4399DF2ADF859" ID-DOI="http://doi.org/10.5281/zenodo.13703524" ID-Zenodo-Dep="13703524" httpUri="https://zenodo.org/record/13703524/files/figure.png" pageId="65" startId="65.[136,229,1953,1974]" targetBox="[185,1402,190,1929]" targetPageId="65" targetType="figure">
<paragraph id="8BFE36E2DF53FF8D19D4399DF2ADF859" blockId="65.[136,1451,1953,2010]" pageId="65">
<emphasis id="B935EAF0DF53FF8D19D4399DF04DF835" bold="true" box="[136,259,1953,1974]" pageId="65">FIGURE 44</emphasis>
.
<taxonomicName id="4C414D61DF53FF8D1852399DF318F835" authority="Negaresh" authorityName="Negaresh" authorityYear="2018" box="[270,598,1953,1974]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="65" phylum="Tracheophyta" rank="species" species="kamalnejadii">
<emphasis id="B935EAF0DF53FF8D1852399DF0BBF835" box="[270,501,1953,1974]" italics="true" pageId="65">Centaurea kamalnejadii</emphasis>
Negaresh
</taxonomicName>
(
<emphasis id="B935EAF0DF53FF8D1B3F399DF234F835" box="[611,890,1953,1974]" italics="true" pageId="65">Negaresh &amp; Kamalnejad 130</emphasis>
, herb. Negaresh).
<emphasis id="B935EAF0DF53FF8D1D76399DF575F835" bold="true" box="[1066,1083,1953,1974]" pageId="65">A</emphasis>
. close up of capitule.
<emphasis id="B935EAF0DF53FF8D1C50399DF452F835" bold="true" box="[1292,1308,1953,1974]" pageId="65">B</emphasis>
. capitule (with above view).
<emphasis id="B935EAF0DF53FF8D185639F9F055F859" bold="true" box="[266,283,1989,2010]" pageId="65">C</emphasis>
. habit and habitat.
<emphasis id="B935EAF0DF53FF8D188F39F9F0AAF859" bold="true" box="[467,484,1989,2010]" pageId="65">D</emphasis>
. inflorescence. Scale
<emphasis id="B935EAF0DF53FF8D1BEB39F9F386F859" bold="true" box="[695,712,1989,2010]" pageId="65">A</emphasis>
= 2 cm,
<emphasis id="B935EAF0DF53FF8D1A4139F9F263F859" bold="true" box="[797,813,1989,2010]" pageId="65">B</emphasis>
= 5 cm,
<emphasis id="B935EAF0DF53FF8D1ADE39F9F2DDF859" bold="true" box="[898,915,1989,2010]" pageId="65">D</emphasis>
= 4 cm.
</paragraph>
</caption>
<subSubSection id="C35B6569DF50FF8F19E13F0CF327FDA2" lastPageId="67" pageId="66" type="etymology">
<paragraph id="8BFE36E2DF50FF8E19E13F0CF4C7FECA" blockId="66.[136,1452,160,581]" box="[189,1417,304,329]" pageId="66">
<emphasis id="B935EAF0DF50FF8E19E13F0CF070FECA" bold="true" box="[189,318,304,329]" pageId="66">Etymology</emphasis>
:—The specific epithet is given in honour of Mr. Jamal Al. Kamalnejad who collected the species.
</paragraph>
<paragraph id="8BFE36E2DF50FF8E19E13F68F030FDC6" blockId="66.[136,1452,160,581]" pageId="66">
<emphasis id="B935EAF0DF50FF8E19E13F68F332FEEE" bold="true" box="[189,636,340,365]" pageId="66">Taxonomic and distribution remarks:</emphasis>
<taxonomicName id="4C414D61DF50FF8E1BC43F68F2E8FEEE" authorityName="Negaresh" authorityYear="2018" box="[664,934,340,365]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="66" phylum="Tracheophyta" rank="species" species="kamalnejadii">
<emphasis id="B935EAF0DF50FF8E1BC43F68F2E8FEEE" box="[664,934,340,365]" italics="true" pageId="66">Centaurea kamalnejadii</emphasis>
</taxonomicName>
is a rare endemic of
<collectingRegion id="4985F800DF50FF8E1DD13F68F4C9FEEE" box="[1165,1415,340,365]" country="Iran" name="Kermanshah" pageId="66">Kermanshah Province</collectingRegion>
, W
<collectingCountry id="F3567672DF50FF8E19D43F44F1FBFE12" box="[136,181,376,401]" name="Iran" pageId="66">Iran</collectingCountry>
(
<figureCitation id="137A2A67DF50FF8E19993F44F054FE12" box="[197,282,376,401]" captionStart="FIGURE 43" captionStartId="63.[136,229,1711,1732]" captionTargetBox="[163,1373,958,1666]" captionTargetId="figure-352@63.[151,1436,929,1685]" captionTargetPageId="63" captionText="FIGURE 43. Distribution of Centaurea kabirkuhensis (circles), C. kamalnejadii (squares), and C. kamyaranensis (triangles)." figureDoi="http://doi.org/10.5281/zenodo.13703522" httpUri="https://zenodo.org/record/13703522/files/figure.png" pageId="66">Fig. 43</figureCitation>
). It is an Irano-Turanian element, and grows on fallow fields and roadsides, at elevations of
<quantity id="4CB99B07DF50FF8E1C713F44F1D0FE36" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="66" unit="m" value="1350.0" valueMax="1400.0" valueMin="1300.0">13001400 m</quantity>
(
<figureCitation id="137A2A67DF50FF8E19F23FA0F0C8FE36" box="[174,390,412,437]" captionStart="FIGURE 44" captionStartId="65.[136,229,1953,1974]" captionTargetBox="[185,1402,190,1929]" captionTargetId="figure-14@65.[185,1402,190,1929]" captionTargetPageId="65" captionText="FIGURE 44. Centaurea kamalnejadii Negaresh (Negaresh &amp; Kamalnejad 130, herb. Negaresh). A. close up of capitule. B. capitule (with above view). C. habit and habitat. D. inflorescence. Scale A = 2 cm, B = 5 cm, D = 4 cm." figureDoi="http://doi.org/10.5281/zenodo.13703524" httpUri="https://zenodo.org/record/13703524/files/figure.png" pageId="66">Figs. 44A and 44B</figureCitation>
).
<taxonomicName id="4C414D61DF50FF8E18C63FA0F3E7FE36" authorityName="Negaresh" authorityYear="2018" box="[410,681,412,437]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="66" phylum="Tracheophyta" rank="species" species="kamalnejadii">
<emphasis id="B935EAF0DF50FF8E18C63FA0F3E7FE36" box="[410,681,412,437]" italics="true" pageId="66">Centaurea kamalnejadii</emphasis>
</taxonomicName>
only found in Shahu Mountains; these mountains can be considered as a local diversity centre because many new species were described from it (Ranjbar
<emphasis id="B935EAF0DF50FF8E1D1F3FFCF535FE5A" box="[1091,1147,448,473]" italics="true" pageId="66">et al.</emphasis>
2013,
<bibRefCitation id="EFD04B13DF50FF8E1D9A3FFCF181FE7E" author="Ranjbar, M. &amp; Negaresh, K." pageId="66" pagination="51 - 56" refId="ref68791" refString="Ranjbar, M. &amp; Negaresh, K. (2014 a) The genus Centaurea (Asteraceae, Cardueae) in Iran: two new species and a lectotypification. Edinburgh Journal of Botany 71 (1): 51 - 56." type="journal article" year="2014">Ranjbar &amp; Negaresh 2014a</bibRefCitation>
,
<bibRefCitation id="EFD04B13DF50FF8E19863FD8F06DFE7E" author="Ranjbar, M. &amp; Negaresh, K." box="[218,291,484,509]" pageId="66" pagination="321 - 327" refId="ref68844" refString="Ranjbar, M. &amp; Negaresh, K. (2014 b) Taxonomic notes and two new species in Centaurea sect. Cynaroides (Compositae, Cardueae) from Iran. Novon 23 (3): 321 - 327." type="journal article" year="2014">2014b</bibRefCitation>
, Negaresh
<emphasis id="B935EAF0DF50FF8E18C13FD8F09BFE7E" box="[413,469,484,509]" italics="true" pageId="66">et al.</emphasis>
2014,
<bibRefCitation id="EFD04B13DF50FF8E1B7D3FD8F230FE7E" author="Negaresh, K. &amp; Rahiminejad, M. R." box="[545,894,484,509]" pageId="66" pagination="229 - 244" refId="ref67972" refString="Negaresh, K. &amp; Rahiminejad, M. R. (2014) A contribution to the taxonomy of Centaurea sect. Cynaroides (Asteraceae, Cardueae - Centaureinae) in Iran. Phytotaxa 158 (3): 229 - 244. http: // dx. doi. org / 10.11646 / phytotaxa. 158.3.3" type="journal article" year="2014">Negaresh &amp; Rahiminejad 2014</bibRefCitation>
,
<bibRefCitation id="EFD04B13DF50FF8E1AD43FD8F28BFE7E" author="Negaresh, K. &amp; Rahiminejad, M. R." box="[904,965,484,509]" pageId="66" pagination="15 - 22" refId="ref68264" refString="Negaresh, K. &amp; Rahiminejad, M. R. (2016) Centaurea rahiminejadii sp. nov. and a new record in the genus Centaurea (Asteraceae, Cardueae - Centaureinae) from Iran. Nordic Journal of Botany 34 (1): 15 - 22." type="journal article" year="2016">2016</bibRefCitation>
). It resembles
<taxonomicName id="4C414D61DF50FF8E1D393FD8F46FFE7E" authorityName="Mozaffarian, F. Ghahremaninejad &amp; Fereidounfar" authorityYear="2012" box="[1125,1313,484,509]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="66" phylum="Tracheophyta" rank="species" species="kabirkuhensis">
<emphasis id="B935EAF0DF50FF8E1D393FD8F46FFE7E" box="[1125,1313,484,509]" italics="true" pageId="66">C. kabirkuhensis</emphasis>
</taxonomicName>
in the shape of inflorescence, indumentum of leaves and color of flowers, although it differs from the latter in some pronounced features (see
<tableCitation id="C6C30359DF50FF8E18473C10F03FFDC6" box="[283,369,556,581]" captionStart="TABLE 2" captionStartId="66.[136,224,629,652]" captionText="TABLE 2. Diagnostic morphological characters of Centaurea kamalnejadii and C. kabirkuhensis." pageId="66">Table 2</tableCitation>
).
</paragraph>
<caption id="DF3E666ADF50FF8E19D43C49F5D3FD0F" box="[136,1181,629,652]" pageId="66" startId="66.[136,224,629,652]" targetType="table">
<paragraph id="8BFE36E2DF50FF8E19D43C49F5D3FD0F" blockId="66.[136,1181,629,652]" box="[136,1181,629,652]" pageId="66">
<emphasis id="B935EAF0DF50FF8E19D43C49F1BAFD0F" bold="true" box="[136,244,629,652]" pageId="66">TABLE 2</emphasis>
. Diagnostic morphological characters of
<taxonomicName id="4C414D61DF50FF8E1BF23C49F2E1FD0F" authorityName="Negaresh" authorityYear="2018" box="[686,943,629,652]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="66" phylum="Tracheophyta" rank="species" species="kamalnejadii">
<emphasis id="B935EAF0DF50FF8E1BF23C49F2E1FD0F" box="[686,943,629,652]" italics="true" pageId="66">Centaurea kamalnejadii</emphasis>
</taxonomicName>
and
<taxonomicName id="4C414D61DF50FF8E1ABF3C49F5D8FD0F" authorityName="Mozaffarian, F. Ghahremaninejad &amp; Fereidounfar" authorityYear="2012" box="[995,1174,629,652]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="66" phylum="Tracheophyta" rank="species" species="kabirkuhensis">
<emphasis id="B935EAF0DF50FF8E1ABF3C49F5D8FD0F" box="[995,1174,629,652]" italics="true" pageId="66">C. kabirkuhensis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BFE36E2DF50FF8E1B1A3CA5F421F90B" pageId="66">
<table id="F941C442DF50003319CC3CA5F4EAF90B" box="[144,1444,665,1672]" gridcols="4" gridrows="33" pageId="66">
<tr id="357134A0DF50003319CC3CA5F4EAFD2C" box="[144,1444,665,687]" gridrow="0" pageId="66" rowspan-0="1" rowspan-3="1">
<th id="76A05DDCDF5000331B1A3CA5F2A0FD2C" box="[582,1006,665,687]" gridcol="1" gridrow="0" pageId="66">
<taxonomicName id="4C414D61DF50FF8E1B1A3CA5F27BFD2C" authorityName="Negaresh" authorityYear="2018" box="[582,821,665,687]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="66" phylum="Tracheophyta" rank="species" species="kamalnejadii">
<emphasis id="B935EAF0DF50FF8E1B1A3CA5F27BFD2C" bold="true" box="[582,821,665,687]" italics="true" pageId="66">Centaurea kamalnejadii</emphasis>
</taxonomicName>
</th>
<th id="76A05DDCDF5000331AA03CA5F432FD2C" box="[1020,1404,665,687]" gridcol="2" gridrow="0" pageId="66">
<taxonomicName id="4C414D61DF50FF8E1AA03CA5F5B8FD2C" authority="Stem" authorityName="Stem" box="[1020,1270,665,687]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="66" phylum="Tracheophyta" rank="species" species="kabirkuhensis">
<emphasis id="B935EAF0DF50FF8E1AA03CA5F5B8FD2C" bold="true" box="[1020,1270,665,687]" italics="true" pageId="66">Centaurea kabirkuhensis</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="357134A0DF50003319CC3C85F4EAFD4C" box="[144,1444,697,719]" gridrow="1" pageId="66" rowspan-3="1">
<th id="76A05DDCDF50003319CC3C85F362FD4C" box="[144,556,697,719]" gridcol="0" gridrow="1" pageId="66">
<emphasis id="B935EAF0DF50FF8E19CC3C85F00AFD4D" bold="true" box="[144,324,697,718]" pageId="66">Stem, height (cm)</emphasis>
</th>
<td id="76A05DDCDF5000331B1A3C85F2A0FD4C" box="[582,1006,697,719]" gridcol="1" gridrow="1" pageId="66">up to 45</td>
<td id="76A05DDCDF5000331AA03C85F432FD4C" box="[1020,1404,697,719]" gridcol="2" gridrow="1" pageId="66">50150</td>
</tr>
<tr id="357134A0DF50003319CC3CE5F4EAFD6D" box="[144,1444,729,750]" gridrow="2" pageId="66">
<th id="76A05DDCDF50003319CC3CE5F362FD6D" box="[144,556,729,750]" gridcol="0" gridrow="2" pageId="66">
<emphasis id="B935EAF0DF50FF8E19CC3CE5F01AFD6D" bold="true" box="[144,340,729,750]" pageId="66">Stem, indumentum</emphasis>
</th>
<td id="76A05DDCDF5000331B1A3CE5F2A0FD6D" box="[582,1006,729,750]" gridcol="1" gridrow="2" pageId="66">lower part subglabrous or with sparse</td>
<td id="76A05DDCDF5000331AA03CE5F4EAFD6D" box="[1020,1444,729,750]" colspan="2" colspanRight="1" gridcol="2" gridrow="2" pageId="66">
lower to median parts with dense
<tr id="357134A0DF50003319CC3CCBF4EAFC8F" box="[144,1444,759,780]" gridrow="3" pageId="66" rowspan-0="1">
<td id="76A05DDCDF5000331B1A3CCBF2A0FC8F" box="[582,1006,759,780]" gridcol="1" gridrow="3" pageId="66">hirsutedeciduous hirsute-articulate hairs (up to 3</td>
<td id="76A05DDCDF5000331AA03CCBF432FC8F" box="[1020,1404,759,780]" gridcol="2" gridrow="3" pageId="66">articulate hairs, others subglabrous</td>
<td id="76A05DDCDF5000331CCC3CCBF4EAFC8F" box="[1424,1444,759,780]" gridcol="3" gridrow="3" pageId="66">or</td>
</tr>
</td>
</tr>
<tr id="357134A0DF50003319CC3D28F4EAFCAA" box="[144,1444,788,809]" gridrow="4" pageId="66" rowspan-0="1" rowspan-3="1">
<td id="76A05DDCDF5000331B1A3D28F2A0FCAA" box="[582,1006,788,809]" gridcol="1" gridrow="4" pageId="66">mm long), median part loosely arachnoid</td>
<td id="76A05DDCDF5000331AA03D28F432FCAA" box="[1020,1404,788,809]" gridcol="2" gridrow="4" pageId="66">sparsely arachnoid</td>
</tr>
<tr id="357134A0DF50003319CC3D0DF4EAFCC5" box="[144,1444,817,838]" gridrow="5" pageId="66" rowspan-0="1" rowspan-2="1" rowspan-3="1">
<td id="76A05DDCDF5000331B1A3D0DF2A0FCC5" box="[582,1006,817,838]" gridcol="1" gridrow="5" pageId="66">with hirsute-articulate hairs, upper part with</td>
</tr>
<tr id="357134A0DF50003319CC3D73F4EAFCE7" box="[144,1444,847,868]" gridrow="6" pageId="66" rowspan-0="1" rowspan-2="1" rowspan-3="1">
<td id="76A05DDCDF5000331B1A3D73F2A0FCE7" box="[582,1006,847,868]" gridcol="1" gridrow="6" pageId="66">dense arachnoid hairs</td>
</tr>
<tr id="357134A0DF50003319CC3D52F4EAFC00" box="[144,1444,878,899]" gridrow="7" pageId="66">
<th id="76A05DDCDF50003319CC3D52F362FC00" box="[144,556,878,899]" gridcol="0" gridrow="7" pageId="66">
<emphasis id="B935EAF0DF50FF8E19CC3D52F088FC00" bold="true" box="[144,454,878,899]" pageId="66">Basal and lower cauline leaves</emphasis>
</th>
<td id="76A05DDCDF5000331B1A3D52F2A0FC00" box="[582,1006,878,899]" gridcol="1" gridrow="7" pageId="66">undivided</td>
<td id="76A05DDCDF5000331AA03D52F4EAFC00" box="[1020,1444,878,899]" colspan="2" colspanRight="1" gridcol="2" gridrow="7" pageId="66">with 12 small lobes in base of lamina, or</td>
</tr>
<tr id="357134A0DF50003319CC3DB0F4EAFC22" box="[144,1444,908,929]" gridrow="8" pageId="66" rowspan-0="1" rowspan-1="1">
<td id="76A05DDCDF5000331AA03DB0F4EAFC22" box="[1020,1444,908,929]" colspan="2" colspanRight="1" gridcol="2" gridrow="8" pageId="66">undivided, lower leaves sometimes with</td>
</tr>
<tr id="357134A0DF50003319CC3D95F4EAFC3D" box="[144,1444,937,958]" gridrow="9" pageId="66" rowspan-0="1" rowspan-1="1" rowspan-3="1">
<td id="76A05DDCDF5000331AA03D95F432FC3D" box="[1020,1404,937,958]" gridcol="2" gridrow="9" pageId="66">auricles at base</td>
</tr>
<tr id="357134A0DF50003319CC3DF4F4EAFC5D" box="[144,1444,968,990]" gridrow="10" pageId="66" rowspan-3="1">
<th id="76A05DDCDF50003319CC3DF4F362FC5D" box="[144,556,968,990]" gridcol="0" gridrow="10" pageId="66">
<emphasis id="B935EAF0DF50FF8E19CC3DF4F343FC5E" bold="true" box="[144,525,968,989]" pageId="66">Basal and lower cauline leaves, shape</emphasis>
</th>
<td id="76A05DDCDF5000331B1A3DF4F2A0FC5D" box="[582,1006,968,990]" gridcol="1" gridrow="10" pageId="66">ovate or lanceolate</td>
<td id="76A05DDCDF5000331AA03DF4F432FC5D" box="[1020,1404,968,990]" gridcol="2" gridrow="10" pageId="66">broadly cordate to broadly ovate</td>
</tr>
<tr id="357134A0DF50003319CC3DD4F4EAFC7D" box="[144,1444,1000,1022]" gridrow="11" pageId="66" rowspan-3="1">
<th id="76A05DDCDF50003319CC3DD4F362FC7D" box="[144,556,1000,1022]" gridcol="0" gridrow="11" pageId="66">
<emphasis id="B935EAF0DF50FF8E19CC3DD4F362FC7E" bold="true" box="[144,556,1000,1021]" pageId="66">Basal and lower cauline leaves, size (cm)</emphasis>
</th>
<td id="76A05DDCDF5000331B1A3DD4F2A0FC7D" box="[582,1006,1000,1022]" gridcol="1" gridrow="11" pageId="66">ca. 24 × 810</td>
<td id="76A05DDCDF5000331AA03DD4F432FC7D" box="[1020,1404,1000,1022]" gridcol="2" gridrow="11" pageId="66">2032 × 1216</td>
</tr>
<tr id="357134A0DF50003319CC3A34F4EAFB9E" box="[144,1444,1032,1053]" gridrow="12" pageId="66" rowspan-3="1">
<th id="76A05DDCDF50003319CC3A34F362FB9E" box="[144,556,1032,1053]" gridcol="0" gridrow="12" pageId="66">
<emphasis id="B935EAF0DF50FF8E19CC3A34F1B2FB9E" bold="true" box="[144,252,1032,1053]" pageId="66">Involucres</emphasis>
</th>
<td id="76A05DDCDF5000331B1A3A34F2A0FB9E" box="[582,1006,1032,1053]" gridcol="1" gridrow="12" pageId="66">convex at base, 4550 mm wide</td>
<td id="76A05DDCDF5000331AA03A34F432FB9E" box="[1020,1404,1032,1053]" gridcol="2" gridrow="12" pageId="66">truncate at base, 3540 mm wide</td>
</tr>
<tr id="357134A0DF50003319CC3A14F4EAFBBE" box="[144,1444,1064,1085]" gridrow="13" pageId="66">
<th id="76A05DDCDF50003319CC3A14F362FBBE" box="[144,556,1064,1085]" gridcol="0" gridrow="13" pageId="66">
<emphasis id="B935EAF0DF50FF8E19CC3A14F005FBBE" bold="true" box="[144,331,1064,1085]" pageId="66">Outer appendages</emphasis>
</th>
<td id="76A05DDCDF5000331B1A3A14F2A0FBBE" box="[582,1006,1064,1085]" gridcol="1" gridrow="13" pageId="66">elongate triangular, with 23 (1.53 mm)</td>
<td id="76A05DDCDF5000331AA03A14F4EAFBBE" box="[1020,1444,1064,1085]" colspan="2" colspanRight="1" gridcol="2" gridrow="13" pageId="66">elongate-lanceolate or oblong-lanceolate,</td>
</tr>
<tr id="357134A0DF50003319CC3A79F4EAFBD9" box="[144,1444,1093,1114]" gridrow="14" pageId="66" rowspan-0="1" rowspan-3="1">
<td id="76A05DDCDF5000331B1A3A79F2A0FBD9" box="[582,1006,1093,1114]" gridcol="1" gridrow="14" pageId="66">spinules on each side, gradually narrowed</td>
<td id="76A05DDCDF5000331AA03A79F432FBD9" box="[1020,1404,1093,1114]" gridcol="2" gridrow="14" pageId="66">entire</td>
</tr>
<tr id="357134A0DF50003319CC3A5FF4EAFBFB" box="[144,1444,1123,1144]" gridrow="15" pageId="66" rowspan-0="1" rowspan-2="1" rowspan-3="1">
<td id="76A05DDCDF5000331B1A3A5FF2A0FBFB" box="[582,1006,1123,1144]" gridcol="1" gridrow="15" pageId="66">into a 510 mm spine</td>
</tr>
<tr id="357134A0DF50003319CC3ABEF4EAFB1B" box="[144,1444,1154,1176]" gridrow="16" pageId="66" rowspan-3="1">
<th id="76A05DDCDF50003319CC3ABEF362FB1B" box="[144,556,1154,1176]" gridcol="0" gridrow="16" pageId="66">
<emphasis id="B935EAF0DF50FF8E19CC3ABEF09BFB14" bold="true" box="[144,469,1154,1175]" pageId="66">Outer appendages, length (mm)</emphasis>
</th>
<td id="76A05DDCDF5000331B1A3ABEF2A0FB1B" box="[582,1006,1154,1176]" gridcol="1" gridrow="16" pageId="66">1023</td>
<td id="76A05DDCDF5000331AA03ABEF432FB1B" box="[1020,1404,1154,1176]" gridcol="2" gridrow="16" pageId="66">712</td>
</tr>
<tr id="357134A0DF50003319CC3A9EF4EAFB34" box="[144,1444,1186,1207]" gridrow="17" pageId="66" rowspan-3="1">
<th id="76A05DDCDF50003319CC3A9EF362FB34" box="[144,556,1186,1207]" gridcol="0" gridrow="17" pageId="66">
<emphasis id="B935EAF0DF50FF8E19CC3A9EF0C5FB34" bold="true" box="[144,395,1186,1207]" pageId="66">Outer appendages, color</emphasis>
</th>
<td id="76A05DDCDF5000331B1A3A9EF2A0FB34" box="[582,1006,1186,1207]" gridcol="1" gridrow="17" pageId="66">white</td>
<td id="76A05DDCDF5000331AA03A9EF432FB34" box="[1020,1404,1186,1207]" gridcol="2" gridrow="17" pageId="66">brown to dark brown</td>
</tr>
<tr id="357134A0DF50003319CC3AFEF4EAFB54" box="[144,1444,1218,1239]" gridrow="18" pageId="66">
<th id="76A05DDCDF50003319CC3AFEF362FB54" box="[144,556,1218,1239]" gridcol="0" gridrow="18" pageId="66">
<emphasis id="B935EAF0DF50FF8E19CC3AFEF012FB54" bold="true" box="[144,348,1218,1239]" pageId="66">Median appendages</emphasis>
</th>
<td id="76A05DDCDF5000331B1A3AFEF2A0FB54" box="[582,1006,1218,1239]" gridcol="1" gridrow="18" pageId="66">elongate triangular to spiniform, with 12</td>
<td id="76A05DDCDF5000331AA03AFEF4EAFB54" box="[1020,1444,1218,1239]" colspan="2" colspanRight="1" gridcol="2" gridrow="18" pageId="66">elongate-lanceolate or oblong-lanceolate,</td>
</tr>
<tr id="357134A0DF50003319CC3AE3F4EAFB77" box="[144,1444,1247,1268]" gridrow="19" pageId="66" rowspan-0="1" rowspan-3="1">
<td id="76A05DDCDF5000331B1A3AE3F2A0FB77" box="[582,1006,1247,1268]" gridcol="1" gridrow="19" pageId="66">(35 mm) spinules on each side, gradually</td>
<td id="76A05DDCDF5000331AA03AE3F432FB77" box="[1020,1404,1247,1268]" gridcol="2" gridrow="19" pageId="66">entire</td>
</tr>
<tr id="357134A0DF50003319CC3AC1F4EAFA91" box="[144,1444,1277,1298]" gridrow="20" pageId="66" rowspan-0="1" rowspan-2="1" rowspan-3="1">
<td id="76A05DDCDF5000331B1A3AC1F2A0FA91" box="[582,1006,1277,1298]" gridcol="1" gridrow="20" pageId="66">narrowed into a 1015 mm spine</td>
</tr>
<tr id="357134A0DF50003319CC3B20F4EAFAB1" box="[144,1444,1308,1330]" gridrow="21" pageId="66" rowspan-3="1">
<th id="76A05DDCDF50003319CC3B20F362FAB1" box="[144,556,1308,1330]" gridcol="0" gridrow="21" pageId="66">
<emphasis id="B935EAF0DF50FF8E19CC3B20F082FAB2" bold="true" box="[144,460,1308,1329]" pageId="66">Median appendages, size (mm)</emphasis>
</th>
<td id="76A05DDCDF5000331B1A3B20F2A0FAB1" box="[582,1006,1308,1330]" gridcol="1" gridrow="21" pageId="66">2530 × 23</td>
<td id="76A05DDCDF5000331AA03B20F432FAB1" box="[1020,1404,1308,1330]" gridcol="2" gridrow="21" pageId="66">1530 × 35</td>
</tr>
<tr id="357134A0DF50003319CC3B00F4EAFAD2" box="[144,1444,1340,1361]" gridrow="22" pageId="66" rowspan-3="1">
<th id="76A05DDCDF50003319CC3B00F362FAD2" box="[144,556,1340,1361]" gridcol="0" gridrow="22" pageId="66">
<emphasis id="B935EAF0DF50FF8E19CC3B00F0D2FAD2" bold="true" box="[144,412,1340,1361]" pageId="66">Median appendages, color</emphasis>
</th>
<td id="76A05DDCDF5000331B1A3B00F2A0FAD2" box="[582,1006,1340,1361]" gridcol="1" gridrow="22" pageId="66">purple to straw-coloured</td>
<td id="76A05DDCDF5000331AA03B00F432FAD2" box="[1020,1404,1340,1361]" gridcol="2" gridrow="22" pageId="66">brown to dark brown</td>
</tr>
<tr id="357134A0DF50003319CC3B60F4EAFAF2" box="[144,1444,1372,1393]" gridrow="23" pageId="66">
<th id="76A05DDCDF50003319CC3B60F362FAF2" box="[144,556,1372,1393]" gridcol="0" gridrow="23" pageId="66">
<emphasis id="B935EAF0DF50FF8E19CC3B60F009FAF2" bold="true" box="[144,327,1372,1393]" pageId="66">Inner appendages</emphasis>
</th>
<td id="76A05DDCDF5000331B1A3B60F2A0FAF2" box="[582,1006,1372,1393]" gridcol="1" gridrow="23" pageId="66">spiniform, entire</td>
<td id="76A05DDCDF5000331AA03B60F4EAFAF2" box="[1020,1444,1372,1393]" colspan="2" colspanRight="1" gridcol="2" gridrow="23" pageId="66">elongate-lanceolate or oblong-lanceolate,</td>
</tr>
<tr id="357134A0DF50003319CC3B45F4EAFA0D" box="[144,1444,1401,1422]" gridrow="24" pageId="66" rowspan-0="1" rowspan-1="1">
<td id="76A05DDCDF5000331AA03B45F432FA0D" box="[1020,1404,1401,1422]" gridcol="2" gridrow="24" pageId="66">entire, sometimes shallowly fringed</td>
<td id="76A05DDCDF5000331CCC3B45F4EAFA0D" box="[1424,1444,1401,1422]" gridcol="3" gridrow="24" pageId="66">or</td>
</tr>
<tr id="357134A0DF50003319CC3BABF4EAFA2F" box="[144,1444,1431,1452]" gridrow="25" pageId="66" rowspan-0="1" rowspan-1="1" rowspan-3="1">
<td id="76A05DDCDF5000331AA03BABF432FA2F" box="[1020,1404,1431,1452]" gridcol="2" gridrow="25" pageId="66">denticulate</td>
</tr>
<tr id="357134A0DF50003319CC3B8AF4EAFA48" box="[144,1444,1462,1483]" gridrow="26" pageId="66" rowspan-3="1">
<th id="76A05DDCDF50003319CC3B8AF362FA48" box="[144,556,1462,1483]" gridcol="0" gridrow="26" pageId="66">
<emphasis id="B935EAF0DF50FF8E19CC3B8AF0F8FA48" bold="true" box="[144,438,1462,1483]" pageId="66">Inner appendages, size (mm)</emphasis>
</th>
<td id="76A05DDCDF5000331B1A3B8AF2A0FA48" box="[582,1006,1462,1483]" gridcol="1" gridrow="26" pageId="66">1325 × 12</td>
<td id="76A05DDCDF5000331AA03B8AF432FA48" box="[1020,1404,1462,1483]" gridcol="2" gridrow="26" pageId="66">1530 × 35</td>
</tr>
<tr id="357134A0DF50003319CC3BEAF4EAFA68" box="[144,1444,1494,1515]" gridrow="27" pageId="66" rowspan-3="1">
<th id="76A05DDCDF50003319CC3BEAF362FA68" box="[144,556,1494,1515]" gridcol="0" gridrow="27" pageId="66">
<emphasis id="B935EAF0DF50FF8E19CC3BEAF0C9FA68" bold="true" box="[144,391,1494,1515]" pageId="66">Inner appendages, color</emphasis>
</th>
<td id="76A05DDCDF5000331B1A3BEAF2A0FA68" box="[582,1006,1494,1515]" gridcol="1" gridrow="27" pageId="66">purple to brown</td>
<td id="76A05DDCDF5000331AA03BEAF432FA68" box="[1020,1404,1494,1515]" gridcol="2" gridrow="27" pageId="66">brown to dark brown</td>
</tr>
<tr id="357134A0DF50003319CC3BCAF4EAF988" box="[144,1444,1526,1547]" gridrow="28" pageId="66" rowspan-3="1">
<th id="76A05DDCDF50003319CC3BCAF362F988" box="[144,556,1526,1547]" gridcol="0" gridrow="28" pageId="66">
<emphasis id="B935EAF0DF50FF8E19CC3BCAF025F988" bold="true" box="[144,363,1526,1547]" pageId="66">Flowers, length (mm)</emphasis>
</th>
<td id="76A05DDCDF5000331B1A3BCAF2A0F988" box="[582,1006,1526,1547]" gridcol="1" gridrow="28" pageId="66">5760</td>
<td id="76A05DDCDF5000331AA03BCAF432F988" box="[1020,1404,1526,1547]" gridcol="2" gridrow="28" pageId="66">5055</td>
</tr>
<tr id="357134A0DF50003319CC3829F4EAF9A8" box="[144,1444,1557,1579]" gridrow="29" pageId="66" rowspan-3="1">
<th id="76A05DDCDF50003319CC3829F362F9A8" box="[144,556,1557,1579]" gridcol="0" gridrow="29" pageId="66">
<emphasis id="B935EAF0DF50FF8E19CC3829F021F9A9" bold="true" box="[144,367,1557,1578]" pageId="66">Achenes, length (mm)</emphasis>
</th>
<td id="76A05DDCDF5000331B1A3829F2A0F9A8" box="[582,1006,1557,1579]" gridcol="1" gridrow="29" pageId="66">6.57</td>
<td id="76A05DDCDF5000331AA03829F432F9A8" box="[1020,1404,1557,1579]" gridcol="2" gridrow="29" pageId="66">78</td>
</tr>
<tr id="357134A0DF50003319CC3809F4EAF9C9" box="[144,1444,1589,1610]" gridrow="30" pageId="66" rowspan-3="1">
<th id="76A05DDCDF50003319CC3809F362F9C9" box="[144,556,1589,1610]" gridcol="0" gridrow="30" pageId="66">
<emphasis id="B935EAF0DF50FF8E19CC3809F011F9C9" bold="true" box="[144,351,1589,1610]" pageId="66">Achenes, wide (mm)</emphasis>
</th>
<td id="76A05DDCDF5000331B1A3809F2A0F9C9" box="[582,1006,1589,1610]" gridcol="1" gridrow="30" pageId="66">ca. 3</td>
<td id="76A05DDCDF5000331AA03809F432F9C9" box="[1020,1404,1589,1610]" gridcol="2" gridrow="30" pageId="66">34</td>
</tr>
<tr id="357134A0DF50003319CC3869F4EAF9E9" box="[144,1444,1621,1642]" gridrow="31" pageId="66">
<th id="76A05DDCDF50003319CC3869F362F9E9" box="[144,556,1621,1642]" gridcol="0" gridrow="31" pageId="66">
<emphasis id="B935EAF0DF50FF8E19CC3869F192F9E9" bold="true" box="[144,220,1621,1642]" pageId="66">Pappus</emphasis>
</th>
<td id="76A05DDCDF5000331B1A3869F2A0F9E9" box="[582,1006,1621,1642]" gridcol="1" gridrow="31" pageId="66">1516 mm long, bristles of inner rows</td>
<td id="76A05DDCDF5000331AA03869F4EAF9E9" box="[1020,1444,1621,1642]" colspan="2" colspanRight="1" gridcol="2" gridrow="31" pageId="66">79 mm long, bristles of inner rows much</td>
</tr>
<tr id="357134A0DF50003319CC384FF4EAF90B" box="[144,1444,1651,1672]" gridrow="32" pageId="66" rowspan-0="1" rowspan-3="1">
<td id="76A05DDCDF5000331B1A384FF2A0F90B" box="[582,1006,1651,1672]" gridcol="1" gridrow="32" pageId="66">slightly longer than others</td>
<td id="76A05DDCDF5000331AA0384FF432F90B" box="[1020,1404,1651,1672]" gridcol="2" gridrow="32" pageId="66">shorter than others, ca. 1.52 mm long</td>
</tr>
</table>
</paragraph>
<paragraph id="8BFE36E2DF50FF8F19E1388AF322FE12" blockId="66.[136,1452,1718,2067]" lastBlockId="67.[136,1452,160,545]" lastPageId="67" pageId="66">
In addition,
<taxonomicName id="4C414D61DF50FF8E1817388AF34FF94C" authorityName="Negaresh" authorityYear="2018" box="[331,513,1718,1743]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="66" phylum="Tracheophyta" rank="species" species="kamalnejadii">
<emphasis id="B935EAF0DF50FF8E1817388AF34FF94C" box="[331,513,1718,1743]" italics="true" pageId="66">C. kamalnejadii</emphasis>
</taxonomicName>
shares some characters such as shape of inflorescence, indumentum of stem and leaves, size and color of flowers with
<taxonomicName id="4C414D61DF50FF8E1B6D38E6F383F970" box="[561,717,1754,1779]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="66" phylum="Tracheophyta" rank="species" species="shahuensis">
<emphasis id="B935EAF0DF50FF8E1B6D38E6F383F970" box="[561,717,1754,1779]" italics="true" pageId="66">C. shahuensis</emphasis>
</taxonomicName>
. However, it markedly differs from the latter in stem up to
<quantity id="4CB99B07DF50FF8E1C3A38E6F4E5F970" box="[1382,1451,1754,1779]" metricMagnitude="-1" metricUnit="m" metricValue="4.5" pageId="66" unit="cm" value="45.0">45 cm</quantity>
(
<emphasis id="B935EAF0DF50FF8E19CD38C2F1E6F894" box="[145,168,1790,1815]" italics="true" pageId="66">vs</emphasis>
.
<quantity id="4CB99B07DF50FF8E19EA38C2F07BF894" box="[182,309,1790,1815]" metricMagnitude="-1" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="66" unit="cm" value="95.0" valueMax="100.0" valueMin="90.0">90100 cm</quantity>
) tall, stem indumentum in lower part subglabrous or with sparse deciduous hirsute-articulate hairs (up to
<quantity id="4CB99B07DF50FF8E19FB391EF1A4F8B8" box="[167,234,1826,1851]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="66" unit="mm" value="3.0">3 mm</quantity>
long), median part loosely arachnoid with hirsute-articulate hairs, upper part densely arachnoid (
<emphasis id="B935EAF0DF50FF8E1C18391EF415F8B8" box="[1348,1371,1826,1851]" italics="true" pageId="66">vs</emphasis>
. lower part glabrescent, median part with sparse hirsute-articulate hairs and sessile glands to subglabrous, upper part with scattered sessile glands sometimes mixed with a few arachnoid hairs), basal leaves ca. 24 ×
<quantity id="4CB99B07DF50FF8E1DF33956F45CF800" box="[1199,1298,1898,1923]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="66" unit="cm" value="9.0" valueMax="10.0" valueMin="8.0">810 cm</quantity>
(
<emphasis id="B935EAF0DF50FF8E1C793956F472F800" box="[1317,1340,1898,1923]" italics="true" pageId="66">vs</emphasis>
. 3035 ×
<quantity id="4CB99B07DF50FF8E19D439B2F041F824" box="[136,271,1934,1959]" metricMagnitude="-1" metricUnit="m" metricValue="1.325" metricValueMax="1.45" metricValueMin="1.2" pageId="66" unit="cm" value="13.25" valueMax="14.5" valueMin="12.0">1214.5 cm</quantity>
), involucres ovoid to subglobose, ca. 50 ×
<quantity id="4CB99B07DF50FF8E1BBA39B2F210F824" box="[742,862,1934,1959]" metricMagnitude="-2" metricUnit="m" metricValue="4.75" metricValueMax="5.0" metricValueMin="4.5" pageId="66" unit="mm" value="47.5" valueMax="50.0" valueMin="45.0">4550 mm</quantity>
(
<emphasis id="B935EAF0DF50FF8E1A3139B2F2CAF824" box="[877,900,1934,1959]" italics="true" pageId="66">vs</emphasis>
. globose-conical, abruptly contracted at apex and angustate (in flowering state), 3540 × 3242(45) mm), outer appendages elongate triangular, 1023 × ca.
<quantity id="4CB99B07DF50FF8E1C14398EF4E9F848" box="[1352,1447,1970,1995]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="66" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">45 mm</quantity>
, with 23 (
<quantity id="4CB99B07DF50FF8E19A239EAF03BF86C" box="[254,373,2006,2031]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="66" unit="mm" value="2.25" valueMax="3.0" valueMin="1.5">1.53 mm</quantity>
) spinules on each side, gradually narrowed into a
<quantity id="4CB99B07DF50FF8E1AE639EAF569F86C" box="[954,1063,2006,2031]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="66" unit="mm" value="7.5" valueMax="10.0" valueMin="5.0">510 mm</quantity>
spine, white (
<emphasis id="B935EAF0DF50FF8E1D9739EAF5ACF86C" box="[1227,1250,2006,2031]" italics="true" pageId="66">vs</emphasis>
. ovate-triangular, 79 × ca.
<quantity id="4CB99B07DF50FF8E19A639C6F014F790" box="[250,346,2042,2067]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="66" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">67 mm</quantity>
, margins entire or fringed in upper part, gradually narrowed into a
<quantity id="4CB99B07DF50FF8E1D0839C6F5FCF790" box="[1108,1202,2042,2067]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="8.0" metricValueMin="2.0" pageId="66" unit="mm" value="5.0" valueMax="8.0" valueMin="2.0">28 mm</quantity>
spine, straw-coloured to pale purple), median appendages elongate triangular to spiniform, 2530 ×
<quantity id="4CB99B07DF51FF8F1AA23E9CF510FF3A" box="[1022,1118,160,185]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="67" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">23 mm</quantity>
, with 12 (
<quantity id="4CB99B07DF51FF8F1DBC3E9CF40EFF3A" box="[1248,1344,160,185]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="67" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">35 mm</quantity>
) spinules on each side, gradually narrowed into a
<quantity id="4CB99B07DF51FF8F1B133EF8F387FF5E" box="[591,713,196,221]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="67" unit="mm" value="12.5" valueMax="15.0" valueMin="10.0">1015 mm</quantity>
spine, purple to straw-coloured (
<emphasis id="B935EAF0DF51FF8F1D1E3EF8F517FF5E" box="[1090,1113,196,221]" italics="true" pageId="67">vs</emphasis>
. triangular, ca. 15 ×
<quantity id="4CB99B07DF51FF8F1C1B3EF8F4E9FF5E" box="[1351,1447,196,221]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="67" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">57 mm</quantity>
, margin entire or sometimes with 12 cilia (
<quantity id="4CB99B07DF51FF8F1B263ED4F3BEFE82" box="[634,752,232,257]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" metricValueMax="3.0" metricValueMin="1.2" pageId="67" unit="mm" value="2.1" valueMax="3.0" valueMin="1.2">1.23 mm</quantity>
) on each side, irregular, gradually narrowed into a ca.
<quantity id="4CB99B07DF51FF8F1C353ED4F4E2FE82" box="[1385,1452,232,257]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="67" unit="mm" value="8.0">8 mm</quantity>
spine), inner appendages spiniform, 1325 ×
<quantity id="4CB99B07DF51FF8F1B213F30F394FEA6" box="[637,730,268,293]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="67" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">12 mm</quantity>
, entire, purple to brown (
<emphasis id="B935EAF0DF51FF8F1AAD3F30F546FEA6" box="[1009,1032,268,293]" italics="true" pageId="67">vs</emphasis>
. narrowly triangular or subspiniform, 1417 ×
<quantity id="4CB99B07DF51FF8F19B03F0CF02FFECA" box="[236,353,304,329]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" metricValueMax="3.2" metricValueMin="2.0" pageId="67" unit="mm" value="2.6" valueMax="3.2" valueMin="2.0">23.2 mm</quantity>
, margin entire or sometimes with up to 1 cilium (
<quantity id="4CB99B07DF51FF8F1AD13F0CF54CFECA" box="[909,1026,304,329]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" metricValueMax="3.0" metricValueMin="1.2" pageId="67" unit="mm" value="2.1" valueMax="3.0" valueMin="1.2">1.23 mm</quantity>
) on each side, irregular, deep purple), and also pappus
<quantity id="4CB99B07DF51FF8F181D3F68F0F4FEEE" box="[321,442,340,365]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="67" unit="mm" value="15.5" valueMax="16.0" valueMin="15.0">1516 mm</quantity>
(
<emphasis id="B935EAF0DF51FF8F18953F68F0AEFEEE" box="[457,480,340,365]" italics="true" pageId="67">vs</emphasis>
. ca.
<quantity id="4CB99B07DF51FF8F1B493F68F32BFEEE" box="[533,613,340,365]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="67" unit="mm" value="14.0">14 mm</quantity>
) long. This new species and its close relative (
<taxonomicName id="4C414D61DF51FF8F1D303F68F449FEEE" box="[1132,1287,340,365]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="67" phylum="Tracheophyta" rank="species" species="shahuensis">
<emphasis id="B935EAF0DF51FF8F1D303F68F531FEEE" box="[1132,1151,340,365]" italics="true" pageId="67">C</emphasis>
.
<emphasis id="B935EAF0DF51FF8F1DD03F68F449FEEE" box="[1164,1287,340,365]" italics="true" pageId="67">shahuensis</emphasis>
</taxonomicName>
) are occurring sympatrically in Shahu Mountains, W
<collectingCountry id="F3567672DF51FF8F1B643F44F327FE12" box="[568,617,376,401]" name="Iran" pageId="67">Iran</collectingCountry>
.
</paragraph>
<paragraph id="8BFE36E2DF51FF8F19E13FA0F52CFE7E" blockId="67.[136,1452,160,545]" pageId="67">
<materialsCitation id="3B293CBFDF51FF8F19E13FA0F29EFE5A" collectingDate="2017-06-15" collectionCode="RUH" country="Iran" county="Ravansar" elevation="1345" latitude="34.73214" location="Kamyaran" longLatPrecision="12" longitude="46.675556" municipality="Ravansar" pageId="67" specimenCount="1" stateProvince="Kermanshah" typeStatus="paratype">
<emphasis id="B935EAF0DF51FF8F19E13FA0F07BFE36" bold="true" box="[189,309,412,437]" pageId="67">
<typeStatus id="54FA8840DF51FF8F19E13FA0F07BFE36" box="[189,309,412,437]" pageId="67" type="paratype">Paratypes</typeStatus>
</emphasis>
:—
<collectingCountry id="F3567672DF51FF8F180B3FA0F0EFFE36" box="[343,417,412,437]" name="Iran" pageId="67">IRAN</collectingCountry>
. Prov.
<collectingRegion id="4985F800DF51FF8F18B13FA0F331FE36" box="[493,639,412,437]" country="Iran" name="Kermanshah" pageId="67">Kermanshah</collectingRegion>
:
<collectingCounty id="629F4E6EDF51FF8F1BD63FA0F3BAFE36" box="[650,756,412,437]" pageId="67">Ravansar</collectingCounty>
,
<quantity id="4CB99B07DF51FF8F1BA33FA0F277FE36" box="[767,825,412,437]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="67" unit="km" value="2.0">2 km</quantity>
from
<collectingMunicipality id="6B9AAC98DF51FF8F1A223FA0F2A8FE36" box="[894,998,412,437]" pageId="67">Ravansar</collectingMunicipality>
toward
<location id="8E9E6039DF51FF8F1D1E3FA0F5F4FE36" LSID="urn:lsid:plazi:treatment:03E887F4DF52FF8F19D43D4CF327FDA2:8E9E6039DF51FF8F1D1E3FA0F5F4FE36" box="[1090,1210,412,437]" country="Iran" county="Ravansar" latitude="34.73214" longLatPrecision="12" longitude="46.675556" municipality="Ravansar" name="Kamyaran" pageId="67" stateProvince="Kermanshah">Kamyaran</location>
,
<geoCoordinate id="EE755025DF51FF8F1D993FA0F437FE36" box="[1221,1401,412,437]" degrees="34" direction="north" minutes="43" orientation="latitude" pageId="67" precision="1" seconds="55.7" value="34.73214">34° 43 55.7″ N</geoCoordinate>
,
<geoCoordinate id="EE755025DF51FF8F1CD83FA0F1BBFE5A" degrees="46" direction="east" minutes="40" orientation="longitude" pageId="67" precision="15" seconds="32" value="46.675556">46° 40 32″ E</geoCoordinate>
,
<quantity id="4CB99B07DF51FF8F185E3FFCF014FE5A" box="[258,346,448,473]" metricMagnitude="3" metricUnit="m" metricValue="1.345" pageId="67" unit="m" value="1345.0">
<elevation id="006CD1D1DF51FF8F185E3FFCF014FE5A" box="[258,346,448,473]" metricMagnitude="3" metricUnit="m" metricValue="1.345" pageId="67" unit="m" value="1345.0">1345 m</elevation>
</quantity>
,
<date id="FFFF1022DF51FF8F183B3FFCF34CFE5A" box="[359,514,448,473]" pageId="67" value="2017-06-15">
<collectingDate id="EFBBE9CADF51FF8F183B3FFCF34CFE5A" box="[359,514,448,473]" pageId="67" value="2017-06-15">15 June 2017</collectingDate>
</date>
,
<emphasis id="B935EAF0DF51FF8F1B523FFCF3E4FE5A" box="[526,682,448,473]" italics="true" pageId="67">Negaresh 178</emphasis>
(herb. Negaresh!,
<collectionCode id="ED50AE27DF51FF8F1A223FFCF28DFE5A" box="[894,963,448,473]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15206" lsid="urn:lsid:biocol.org:col:15206" name="Rhodes University" pageId="67" type="Herbarium">RUH</collectionCode>
!)
</materialsCitation>
.
<materialsCitation id="3B293CBFDF51FF8F1A873FFCF510FE7E" collectingDate="2017-06-16" collectionCode="RUH" country="Iran" elevation="1370" latitude="34.717888" location="Sardem village" longLatPrecision="1" longitude="46.67739" municipality="Ravansar" pageId="67" specimenCount="1" typeStatus="paratype">
<collectingMunicipality id="6B9AAC98DF51FF8F1A873FFCF50DFE5A" box="[987,1091,448,473]" pageId="67">Ravansar</collectingMunicipality>
toward
<location id="8E9E6039DF51FF8F1DFD3FFCF400FE5A" LSID="urn:lsid:plazi:treatment:03E887F4DF52FF8F19D43D4CF327FDA2:8E9E6039DF51FF8F1DFD3FFCF400FE5A" box="[1185,1358,448,473]" country="Iran" latitude="34.717888" longLatPrecision="1" longitude="46.67739" municipality="Ravansar" name="Sardem village" pageId="67">Sardem village</location>
,
<geoCoordinate id="EE755025DF51FF8F1C063FFCF19BFE7E" degrees="34" direction="north" minutes="43" orientation="latitude" pageId="67" precision="1" seconds="4.4" value="34.717888">34° 43 4.4″ N</geoCoordinate>
,
<geoCoordinate id="EE755025DF51FF8F19BC3FD8F0C0FE7E" box="[224,398,484,509]" degrees="46" direction="east" minutes="40" orientation="longitude" pageId="67" precision="1" seconds="38.6" value="46.67739">46° 40 38.6″ E</geoCoordinate>
,
<quantity id="4CB99B07DF51FF8F18C63FD8F0BFFE7E" box="[410,497,484,509]" metricMagnitude="3" metricUnit="m" metricValue="1.3699999999999999" pageId="67" unit="m" value="1370.0">
<elevation id="006CD1D1DF51FF8F18C63FD8F0BFFE7E" box="[410,497,484,509]" metricMagnitude="3" metricUnit="m" metricValue="1.3699999999999999" pageId="67" unit="m" value="1370.0">1370 m</elevation>
</quantity>
,
<date id="FFFF1022DF51FF8F18A13FD8F3DBFE7E" box="[509,661,484,509]" pageId="67" value="2017-06-16">
<collectingDate id="EFBBE9CADF51FF8F18A13FD8F3DBFE7E" box="[509,661,484,509]" pageId="67" value="2017-06-16">16 June 2017</collectingDate>
</date>
,
<emphasis id="B935EAF0DF51FF8F1BFC3FD8F275FE7E" box="[672,827,484,509]" italics="true" pageId="67">Negaresh 179</emphasis>
(herb. Negaresh!,
<collectionCode id="ED50AE27DF51FF8F1D513FD8F51CFE7E" box="[1037,1106,484,509]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15206" lsid="urn:lsid:biocol.org:col:15206" name="Rhodes University" pageId="67" type="Herbarium">RUH</collectionCode>
!)
</materialsCitation>
.
</paragraph>
<paragraph id="8BFE36E2DF51FF8F19E13C34F327FDA2" blockId="67.[136,1452,160,545]" box="[189,617,520,545]" pageId="67">
<emphasis id="B935EAF0DF51FF8F19E13C34F099FDA2" bold="true" box="[189,471,520,545]" pageId="67">Chromosome numbers:</emphasis>
—Unknown.
</paragraph>
</subSubSection>
</treatment>
</document>