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<document id="855A5E7B108E0D8138A0E9EFE344ABBB" ID-DOI="10.11646/phytotaxa.363.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13703414" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725581356715" checkinUser="felipe" docAuthor="Negaresh, Kazem &amp; Rahiminejad, Mohammad Reza" docDate="2018" docId="03E887F4DF23FFFF19D439F5F592FC3C" docLanguage="en" docName="phytotaxa.363.1.1.pdf" docOrigin="Phytotaxa 363 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.363.1.1" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.4:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="4" docTitle="Centaurea haussknechtii Boissier 1875" docType="treatment" docVersion="2" lastPageNumber="450" masterDocId="FFD1FF8CDF12FFCC195C3E3CF14EFF83" masterDocTitle="A revision of Centaurea sect. Cynaroides (Asteraceae, Cardueae-Centaureinae)" masterLastPageNumber="450" masterPageNumber="448" pageNumber="448" updateTime="1725593506943" updateUser="ExternalLinkService">
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<mods:title id="0D125CDEFA0BD6DF88283D79F8E2280C">A revision of Centaurea sect. Cynaroides (Asteraceae, Cardueae-Centaureinae)</mods:title>
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<mods:namePart id="F82A932FFC47446EE147A32D952B730D">Negaresh, Kazem</mods:namePart>
<mods:affiliation id="344E2DF28EFC8374D3D6FC6EFF8BAA9F">Department of Horticulture, Faculty of Agriculture, Khuzestan Agricultural Sciences and Natural Resources University, Mollasani, Ahvaz, Iran. (* Corresponding author e-mail: negaresh @ ramin. ac. ir)</mods:affiliation>
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<mods:namePart id="8A9CECBD2652D6D37B38051A41420874">Rahiminejad, Mohammad Reza</mods:namePart>
<mods:affiliation id="E15A3D88BF081B602A04FBA1F2442D16">Department of Horticulture, Faculty of Agriculture, Khuzestan Agricultural Sciences and Natural Resources University, Mollasani, Ahvaz, Iran. (* Corresponding author e-mail: negaresh @ ramin. ac. ir) &amp; Department of Biology, University of Isfahan, Isfahan 81746 - 73441, Iran. &amp; Department of Horticulture, Faculty of Agriculture, Khuzestan Agricultural Sciences and Natural Resources University, Mollasani, Ahvaz, Iran. (* Corresponding author e-mail: negaresh @ ramin. ac. ir)</mods:affiliation>
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<mods:title id="4E0F6C5B78DFA47A36D0DDBDEFC7DFF8">Phytotaxa</mods:title>
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<mods:date id="FDDC721EBB281114671BD9069F0E7CEB">2018</mods:date>
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<treatment id="03E887F4DF23FFFF19D439F5F592FC3C" ID-DOI="http://doi.org/10.5281/zenodo.13703754" ID-Zenodo-Dep="13703754" LSID="urn:lsid:plazi:treatment:03E887F4DF23FFFF19D439F5F592FC3C" httpUri="http://treatment.plazi.org/id/03E887F4DF23FFFF19D439F5F592FC3C" lastPageId="51" pageId="49">
<subSubSection id="C35B6569DF23FFFD19D439F5F26EF7A9" pageId="49" type="nomenclature">
<paragraph id="8BFE36E2DF23FFFD19D439F5F26EF7A9" blockId="49.[136,1451,1991,2090]" pageId="49">
<emphasis id="B935EAF0DF23FFFD19D439F5F1E2F861" bold="true" box="[136,172,1993,2018]" pageId="49">15.</emphasis>
<taxonomicName id="4C414D61DF23FFFD19EF39FBF386F861" ID-CoL="S6NX" authority="Boissier (1875: 670)" authorityName="Boissier" authorityPageNumber="670" authorityYear="1875" box="[179,712,1991,2018]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="49" phylum="Tracheophyta" rank="species" species="haussknechtii">
<emphasis id="B935EAF0DF23FFFD19EF39FBF09AF861" bold="true" box="[179,468,1991,2018]" italics="true" pageId="49">Centaurea haussknechtii</emphasis>
<bibRefCitation id="EFD04B13DF23FFFD188039F5F386F861" author="Boissier, E." box="[476,712,1993,2018]" pageId="49" refId="ref65414" refString="Boissier, E. (1875) Flora Orientalis, vol. 3. H. Georg, Geneva &amp; Basilea [Basel], 1033 pp." type="book" year="1875">Boissier (1875: 670)</bibRefCitation>
</taxonomicName>
. Type:—
<collectingCountry id="F3567672DF23FFFD1A7039F5F2D1F861" box="[812,927,1993,2018]" name="Turkey" pageId="49">TURKEY</collectingCountry>
. C6
<collectingRegion id="4985F800DF23FFFD1A8839F5F504F861" box="[980,1098,1993,2018]" country="Turkey" name="Gaziantep" pageId="49">Gaziantep</collectingRegion>
: in graminosis calcareis montis Ssoffdagh (So f Da.) Syriae borealis,
<quantity id="4CB99B07DF23FFFD1B6F39D1F3C5F785" box="[563,651,2029,2054]" metricMagnitude="3" metricUnit="m" metricValue="1.0699999999999998" pageId="49" unit="m" value="1070.0">1070 m</quantity>
,
<date id="FFFF1022DF23FFFD1BC439D1F260F785" box="[664,814,2029,2054]" pageId="49" value="1865-07-27">27 July 1865</date>
,
<emphasis id="B935EAF0DF23FFFD1A6639D1F2B6F785" box="[826,1016,2029,2054]" italics="true" pageId="49">Haussknecht s.n.</emphasis>
(
<typeStatus id="54FA8840DF23FFFD1D5439D1F53FF785" box="[1032,1137,2029,2054]" pageId="49" type="lectotype">lectotype</typeStatus>
G! designated by
<bibRefCitation id="EFD04B13DF23FFFD1C1E39D1F1B2F7A9" author="Wagenitz, G." pageId="49" pagination="469 - 494" refId="ref69162" refString="Wagenitz, G. (1960) Centaurea L. sect. Cynaropsis, eine neue Sektion der Gattung aus Vorderasien. Willdenowia 2: 469 - 494." type="journal article" year="1960">Wagenitz 1960: 490</bibRefCitation>
,
<typeStatus id="54FA8840DF23FFFD185B362DF0D7F7A9" box="[263,409,2065,2090]" pageId="49" type="isolectotype">isolectotypes</typeStatus>
G!, JE!: two sheets, W!). (
<figureCitation id="137A2A67DF23FFFD1B9A362DF259F7A9" box="[710,791,2065,2090]" captionStart="FIGURE 32" captionStartId="50.[136,229,1953,1974]" captionTargetBox="[196,1393,203,1924]" captionTargetId="figure-17@50.[194,1393,190,1929]" captionTargetPageId="50" captionText="FIGURE 32. Centaurea haussknechtii Boissier (Haussknecht s.n., G). A. habit. B. close up of capitule. Scale B = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.13703492" httpUri="https://zenodo.org/record/13703492/files/figure.png" pageId="49">Fig. 32</figureCitation>
)
</paragraph>
</subSubSection>
<caption id="DF3E666ADF20FFFE19D4399DF447F835" ID-DOI="http://doi.org/10.5281/zenodo.13703492" ID-Zenodo-Dep="13703492" box="[136,1289,1953,1974]" httpUri="https://zenodo.org/record/13703492/files/figure.png" pageId="50" startId="50.[136,229,1953,1974]" targetBox="[196,1393,203,1924]" targetPageId="50" targetType="figure">
<paragraph id="8BFE36E2DF20FFFE19D4399DF447F835" blockId="50.[136,1289,1953,1974]" box="[136,1289,1953,1974]" pageId="50">
<emphasis id="B935EAF0DF20FFFE19D4399DF047F835" bold="true" box="[136,265,1953,1974]" pageId="50">FIGURE 32.</emphasis>
<taxonomicName id="4C414D61DF20FFFE1853399DF31FF835" authority="Boissier" authorityName="Boissier" authorityYear="1875" box="[271,593,1953,1974]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="50" phylum="Tracheophyta" rank="species" species="haussknechtii">
<emphasis id="B935EAF0DF20FFFE1853399DF0B3F835" box="[271,509,1953,1974]" italics="true" pageId="50">Centaurea haussknechtii</emphasis>
Boissier
</taxonomicName>
(
<emphasis id="B935EAF0DF20FFFE1B03399DF24CF835" box="[607,770,1953,1974]" italics="true" pageId="50">Haussknecht s.n.</emphasis>
, G).
<emphasis id="B935EAF0DF20FFFE1A6E399DF204F835" bold="true" box="[818,842,1953,1974]" pageId="50">A.</emphasis>
habit.
<emphasis id="B935EAF0DF20FFFE1AD0399DF2ECF835" bold="true" box="[908,930,1953,1974]" pageId="50">B.</emphasis>
close up of capitule. Scale
<emphasis id="B935EAF0DF20FFFE1DF5399DF5F7F835" bold="true" box="[1193,1209,1953,1974]" pageId="50">B</emphasis>
= 1 cm.
</paragraph>
</caption>
<subSubSection id="C35B6569DF21FFFF19D43E9CF44FFC1F" pageId="51" type="description">
<paragraph id="8BFE36E2DF21FFFF19D43E9CF2D8FD6F" blockId="51.[136,1452,160,959]" pageId="51">
Biennial herbs, with long and stout taproot, whole plant usually grayish or green-yellowish,
<quantity id="4CB99B07DF21FFFF1DE63E9CF462FF3A" box="[1210,1324,160,185]" metricMagnitude="-1" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="51" unit="cm" value="55.0" valueMax="60.0" valueMin="50.0">5060 cm</quantity>
tall; collar of fibrous petiolar remains present at stem base. Stem erect, simple,
<quantity id="4CB99B07DF21FFFF1AC73EFFF2B4FF5F" box="[923,1018,195,220]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="51" unit="mm" value="3.5" valueMax="5.0" valueMin="2.0">25 mm</quantity>
in diam. at base, cylindrical, sparsely leafy throughout, lower part purplish and loosely covered with hirsute-articulate hairs, others ± densely covered with white-tomentose hairs. Leaves firm or rigid (on drying), usually divided, densely covered with white-tomentose hairs, sometimes basal and lower ones sparsely covered with hirsute-articulate hairs, denser along midrib and veins, and mixed with tomentose hairs. Basal and lower cauline leaves petiolate, (7)
<quantity id="4CB99B07DF21FFFF1A8F3F6CF50BFEEA" box="[979,1093,336,361]" metricMagnitude="-1" metricUnit="m" metricValue="2.1" metricValueMax="3.0" metricValueMin="1.2" pageId="51" unit="cm" value="21.0" valueMax="30.0" valueMin="12.0">1230 cm</quantity>
long, lyrate; terminal segments larger, oblong, triangular-ovate or lanceolate, 410 × (1.2)
<quantity id="4CB99B07DF21FFFF1A703F4FF2CBFE0F" box="[812,901,371,396]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="6.0" metricValueMin="2.0" pageId="51" unit="cm" value="4.0" valueMax="6.0" valueMin="2.0">26 cm</quantity>
, entire or shallowly serrulate, obtuse or truncate at base, acute or rounded at apex; lateral segments smaller, 12 pairs, lanceolate or ± retrorse. Median and upper cauline leaves sessile, simple, oblong or narrowly oblong, 210 ×
<quantity id="4CB99B07DF21FFFF1A283F85F2BBFE51" box="[884,1013,441,466]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="51" unit="cm" value="1.0" valueMax="1.5" valueMin="0.5">0.51.5 cm</quantity>
, very shortly decurrent, sometimes not decurrent, entire, acuminate or mucronate at apex. Capitula few, 4 to 7, arranged in a spike or raceme, often sessile or subsessile, sometimes with
<quantity id="4CB99B07DF21FFFF18E03C3CF340FD9A" box="[444,526,512,537]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="6.0" metricValueMin="2.0" pageId="51" unit="cm" value="4.0" valueMax="6.0" valueMin="2.0">26 cm</quantity>
peduncles. Involucres ovoid, truncate at base, (33)3540 ×
<quantity id="4CB99B07DF21FFFF1DEA3C3CF47CFD9A" box="[1206,1330,512,537]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="51" unit="mm" value="32.5" valueMax="35.0" valueMin="30.0">3035 mm</quantity>
. Phyllaries multiseriate, yellow-greenish, imbricate, coriaceous-scarious, glabrous.Appendages rigid, totally concealing phyllaries, not decurrent, lanceolate, straw-coloured or pale brown, 34(4.5) mm wide at base (excluding cilia), sometimes squarrose or reflexed; cilia erect, numerous, 79 on each side,
<quantity id="4CB99B07DF21FFFF1A0A3C55F2FAFD01" box="[854,948,617,642]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="51" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">57 mm</quantity>
long; spines narrowly triangular, (7)
<quantity id="4CB99B07DF21FFFF1C3A3C55F1FAFD26" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="51" unit="mm" value="16.0" valueMax="17.0" valueMin="15.0">1517 mm</quantity>
long, strongly longer than nearest cilia. Flowers pink-purple; central florets hermaphroditic, limb ca.
<quantity id="4CB99B07DF21FFFF1C423CB0F423FD26" box="[1310,1389,652,677]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="51" unit="mm" value="14.0">14 mm</quantity>
long, 5-lobed, lobes ca.
<quantity id="4CB99B07DF21FFFF180D3C8CF0DEFD4A" box="[337,400,688,713]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="51" unit="mm" value="6.0">6 mm</quantity>
long; peripheral florets sterile, slightly shorter than central ones, conspicuous, finely dissected, not radiant, 5-lobed, limb lobes linear. Achenes and pappus unknown.
</paragraph>
<paragraph id="8BFE36E2DF21FFFF19E13CCAF5F1FCD6" blockId="51.[136,1452,160,959]" pageId="51">
<emphasis id="B935EAF0DF21FFFF19E13CCAF3C5FC8C" bold="true" box="[189,651,758,783]" pageId="51">Taxonomic and distribution remarks:</emphasis>
<taxonomicName id="4C414D61DF21FFFF1BFB3CCAF2F1FC8C" authorityName="Boissier" authorityYear="1875" box="[679,959,758,783]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="51" phylum="Tracheophyta" rank="species" species="haussknechtii">
<emphasis id="B935EAF0DF21FFFF1BFB3CCAF2F1FC8C" box="[679,959,758,783]" italics="true" pageId="51">Centaurea haussknechtii</emphasis>
</taxonomicName>
is a rare endemic of
<collectingRegion id="4985F800DF21FFFF1D983CCAF4E9FC8C" box="[1220,1447,758,783]" country="Turkey" name="Gaziantep" pageId="51">Gaziantep Province</collectingRegion>
, S
<collectingCountry id="F3567672DF21FFFF19FD3D25F1BFFCB1" box="[161,241,793,818]" name="Turkey" pageId="51">Turkey</collectingCountry>
(
<figureCitation id="137A2A67DF21FFFF185F3D25F014FCB1" box="[259,346,793,818]" captionStart="FIGURE 30" captionStartId="47.[136,229,1389,1410]" captionTargetBox="[166,1381,628,1339]" captionTargetId="figure-193@47.[151,1436,605,1363]" captionTargetPageId="47" captionText="FIGURE 30. Distribution of Centaurea gudrunensis (circles), C. haradjianii (squares), and C. haussknechtii (triangles)." figureDoi="http://doi.org/10.5281/zenodo.13703484" httpUri="https://zenodo.org/record/13703484/files/figure.png" pageId="51">Fig. 30</figureCitation>
).
<taxonomicName id="4C414D61DF21FFFF182C3D25F3C8FCB1" authorityName="Boissier" authorityYear="1875" box="[368,646,793,818]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="51" phylum="Tracheophyta" rank="species" species="haussknechtii">
<emphasis id="B935EAF0DF21FFFF182C3D25F3C8FCB1" box="[368,646,793,818]" italics="true" pageId="51">Centaurea haussknechtii</emphasis>
</taxonomicName>
is an Irano-Turanian element, and grows on limestone, grassland and calcareous mountains, at elevations of
<quantity id="4CB99B07DF21FFFF1B663D00F399FCD6" box="[570,727,828,853]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="51" unit="m" value="1050.0" valueMax="1100.0" valueMin="1000.0">10001100 m</quantity>
. It closely resembles
<taxonomicName id="4C414D61DF21FFFF1A9B3D00F574FCD6" authorityName="Post" authorityYear="1888" box="[967,1082,828,853]" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="51" phylum="Tracheophyta" rank="species" species="doddsii">
<emphasis id="B935EAF0DF21FFFF1A9B3D00F574FCD6" box="[967,1082,828,853]" italics="true" pageId="51">C. doddsii</emphasis>
</taxonomicName>
in its habit.
</paragraph>
<paragraph id="8BFE36E2DF21FFFF19E13D5CF44FFC1F" blockId="51.[136,1452,160,959]" pageId="51">
<emphasis id="B935EAF0DF21FFFF19E13D5CF094FCFA" bold="true" box="[189,474,864,889]" pageId="51">Chromosome numbers:</emphasis>
<bibRefCitation id="EFD04B13DF21FFFF18AA3D5CF399FCFA" author="Askoy, H. &amp; Berber, A. A. &amp; Celik, S. &amp; Baser, B. &amp; Karaveliogullari, F. A." box="[502,727,864,889]" pageId="51" pagination="285 - 294" refId="ref65175" refString="Askoy, H., Berber, A. A., Celik, S., Baser, B. &amp; Karaveliogullari, F. A. (2016) Karyological study on 12 Centaurea (Asteraceae) species from Turkey. Caryologia 69 (4): 285 - 294. http: // doi. org / 10.1080 / 00087114.2016.1188357" type="journal article" year="2016">
Askoy
<emphasis id="B935EAF0DF21FFFF1B143D5CF3CDFCFA" box="[584,643,864,889]" italics="true" pageId="51">et al.</emphasis>
(2016)
</bibRefCitation>
reported the chromosome number 2
<emphasis id="B935EAF0DF21FFFF1D253D5CF5C9FCFA" box="[1145,1159,864,889]" italics="true" pageId="51">n</emphasis>
= 2
<emphasis id="B935EAF0DF21FFFF1DE43D5CF58AFCFA" box="[1208,1220,864,889]" italics="true" pageId="51">x</emphasis>
= 18 for
<taxonomicName id="4C414D61DF21FFFF1C6A3D5CF06EFC1F" authorityName="Boissier" authorityYear="1875" class="Magnoliopsida" family="Asteraceae" genus="Centaurea" kingdom="Plantae" order="Asterales" pageId="51" phylum="Tracheophyta" rank="species" species="haussknechtii">
<emphasis id="B935EAF0DF21FFFF1C6A3D5CF06EFC1F" italics="true" pageId="51">Centaurea haussknechtii</emphasis>
</taxonomicName>
collected from
<collectingCountry id="F3567672DF21FFFF188C3DBFF36FFC1F" box="[464,545,899,924]" name="Turkey" pageId="51">Turkey</collectingCountry>
, Siverek-Kahta (
<collectingRegion id="4985F800DF21FFFF1B813DBFF217FC1F" box="[733,857,899,924]" country="Turkey" name="Adiyaman" pageId="51">Adıyaman</collectingRegion>
). This is the only count for this taxon.
</paragraph>
</subSubSection>
<subSubSection id="C35B6569DF21FFFF19E13D9AF592FC3C" box="[189,1244,934,959]" pageId="51" type="materials_examined">
<paragraph id="8BFE36E2DF21FFFF19E13D9AF592FC3C" blockId="51.[136,1452,160,959]" box="[189,1244,934,959]" pageId="51">
<emphasis id="B935EAF0DF21FFFF19E13D9AF3F9FC3C" bold="true" box="[189,695,934,959]" pageId="51">
Selected specimens examined:—
<collectingCountry id="F3567672DF21FFFF1B633D9AF3F9FC3C" box="[575,695,934,959]" name="Turkey" pageId="51">TURKEY</collectingCountry>
</emphasis>
. Taurus cataonicus, 1865,
<emphasis id="B935EAF0DF21FFFF1ABD3D9AF5D0FC3C" box="[993,1182,934,959]" italics="true" pageId="51">Haussknecht s.n.</emphasis>
(
<collectionCode id="ED50AE27DF21FFFF1DF33D9AF584FC3C" box="[1199,1226,934,959]" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" pageId="51" type="Herbarium">G</collectionCode>
!).
</paragraph>
</subSubSection>
</treatment>
</document>