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<document id="5F2F2037276C07996C3C8F1BAB73033D" ID-DOI="10.11646/zootaxa.5414.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10681263" ID-ZooBank="3B1DFADA-C20E-473B-A5E9-843548B54AD3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1708377054888" checkinUser="plazi" docAuthor="Paredes-Munguía, Williams, Brescovit, Antonio D. &amp; Teixeira, Renato A." docDate="2024" docId="03E887D2BA0BFE2AFDDBFCF860C5FAFA" docLanguage="en" docName="zootaxa.5414.1.1.pdf" docOrigin="Zootaxa 5414 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5414.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.18:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="18" docTitle="Arctosa cinerea" docType="key" docVersion="1" lastPageNumber="7" masterDocId="FFD1FFAABA0DFE2CFD4CFF8E6559FFD2" masterDocTitle="Revision of Neotropical wolf spider genus Arctosa C. L. Koch, 1847 (Araneae: Lycosidae), with description of seven new species" masterLastPageNumber="83" masterPageNumber="1" pageNumber="7" updateTime="1708377875290" updateUser="ExternalLinkService">
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<mods:title id="8EC2B3893E9EDEF38CB486B8BBB733BE">Revision of Neotropical wolf spider genus Arctosa C. L. Koch, 1847 (Araneae: Lycosidae), with description of seven new species</mods:title>
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<mods:namePart id="8220ADB8EA77BE4A201B71D689D18740">Paredes-Munguía, Williams</mods:namePart>
<mods:nameIdentifier id="379F86F1A5D54A1E4616171C8E53DAB1" type="ORCID">0000-0002-3320-4611</mods:nameIdentifier>
<mods:affiliation id="6CA923A63B3F64838D7B884287C2C248">Pontifícia Universidade Católica do Rio Grande do Sul, Escola de Ciências da Saúde e da Vida, Museu de Ciências e Tecnologia, Laboratório de Aracnologia, Porto Alegre, Brazil. &amp; Museo de Historia Natural de la Universidad Nacional Mayor de San Marcos, Laboratorio de Entomología, Av. Arenales 1256, Lima 14, Peru. &amp; williams. paredes @ acad. pucrs. br; https: // orcid. org / 0000 - 0002 - 3320 - 4611</mods:affiliation>
<mods:nameIdentifier id="5B2EDDEAEF32B6D11702B5AD07EFDA30" type="email">williams.paredes@acad.pucrs.br</mods:nameIdentifier>
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<mods:namePart id="742FD54EF30B4DDF6BC8699D74F2B6FD">Brescovit, Antonio D.</mods:namePart>
<mods:nameIdentifier id="55A2B5090B28816A98D20AF3DD5D6CC6" type="ORCID">0000-0002-1511-5324</mods:nameIdentifier>
<mods:affiliation id="FF7294DA1766A6BDDBF71B8901B9F32F">Laboratório de Coleções Zoológicas, Av. Vital Brasil, 1500, Butantã, 05503 - 900, São Paulo, SP, Brazil. &amp; antonio. brescovit @ butantan. gov. br; https: // orcid. org / 0000 - 0002 - 1511 - 5324</mods:affiliation>
<mods:nameIdentifier id="56D7F37BC958A15DE520A17A7C5F5D95" type="email">antonio.brescovit@butantan.gov.br</mods:nameIdentifier>
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<mods:namePart id="D4D950C701CE719E00FDC405FE367598">Teixeira, Renato A.</mods:namePart>
<mods:nameIdentifier id="A436F1DAF42AB2A0BB08B2DC3D87D753" type="ORCID">0000-0002-1756-9821</mods:nameIdentifier>
<mods:affiliation id="AE6B40A7FA1AF6D1B0160885B20E0F12">Pontifícia Universidade Católica do Rio Grande do Sul, Escola de Ciências da Saúde e da Vida, Museu de Ciências e Tecnologia, Laboratório de Aracnologia, Porto Alegre, Brazil. &amp; Pontifícia Universidade Católica do Rio Grande do Sul, Escola de Ciências da Saúde e da Vida, Museu de Ciências e Tecnologia, Laboratório de Aracnologia, Porto Alegre, Brazil. &amp; renatoaug. tx @ gmail. com; https: // orcid. org / 0000 - 0002 - 1756 - 9821</mods:affiliation>
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<mods:date id="3F011E0C9EC42DED2ADDC7D44996653A">2024</mods:date>
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<treatment id="03E887D2BA0BFE2AFDDBFCF860C5FAFA" LSID="urn:lsid:plazi:treatment:03E887D2BA0BFE2AFDDBFCF860C5FAFA" httpUri="http://treatment.plazi.org/id/03E887D2BA0BFE2AFDDBFCF860C5FAFA" lastPageNumber="7" pageId="6" pageNumber="7">
<subSubSection id="C35B654FBA0BFE2AFDDBFCF864CEFC5D" box="[151,407,885,911]" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="8BFE36C4BA0BFE2AFDDBFCF864CEFC5D" blockId="6.[151,1437,885,1321]" box="[151,407,885,911]" pageId="6" pageNumber="7">
<heading id="D0B681A8BA0BFE2AFDDBFCF864CEFC5D" box="[151,407,885,911]" fontSize="11" level="2" pageId="6" pageNumber="7" reason="5">
<taxonomicName id="4C414D47BA0BFE2AFDDBFCF865B3FC5D" ID-CoL="67RS3" baseAuthorityName="Fabricius" baseAuthorityYear="1777" box="[151,234,886,911]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="cinerea">
<emphasis id="B935EAD6BA0BFE2AFDDBFCF865B3FC5D" box="[151,234,886,911]" italics="true" pageId="6" pageNumber="7">cinerea</emphasis>
</taxonomicName>
group, females
</heading>
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<key id="FFD061F5BA0BFE2AFDDBFC1860C5FAFA" pageId="6" pageNumber="7">
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<paragraph id="8BFE36C4BA0BFE2AFDDBFC1860C5FC79" blockId="6.[151,1437,885,1321]" box="[151,1436,918,940]" pageId="6" pageNumber="7">
<keyLead id="30B095F1BA0BFE2AFDDBFC1860C5FC79" box="[151,1436,918,940]" pageId="6" pageNumber="7">
1 Median septum (MS) completely flat (
<figureCitation id="137A2A41BA0BFE2AFF1FFC1867CEFC7E" box="[595,663,918,940]" captionStart="FIGURE 5" captionStartId="12.[151,250,1317,1341]" captionTargetBox="[151,1436,181,1293]" captionTargetId="figure-25@12.[151,1436,181,1293]" captionTargetPageId="12" captionText="FIGURE 5. Arctosa ayaymama sp. nov. Epigynum (MUSM-ENT 506993) a–c (a, ventral; b, dorsal; c, dictynoid pore) d, Male leg I, prolateral. Scale bars: a, 150 µm, b, 400 µm, c, 15 µm, d, 200 µm. CO copulatory opening, FD fertilization duct, H hood, HS head of spermatheca, DP dictynoid pore, MS median septum." figureDoi="http://doi.org/10.5281/zenodo.10681273" httpUri="https://zenodo.org/record/10681273/files/figure.png" pageId="6" pageNumber="7">Fig. 5a</figureCitation>
)........................................
<emphasis id="B935EAD6BA0BFE2AF9C1FC1860C5FC79" bold="true" box="[1165,1436,918,939]" pageId="6" pageNumber="7">
<taxonomicName id="4C414D47BA0BFE2AF9C1FC186013FC79" authority="Paredes-Munguía &amp; Brescovit &amp; Teixeira, 2024" authorityName="Paredes-Munguía &amp; Brescovit &amp; Teixeira" authorityYear="2024" box="[1165,1354,918,939]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="ayaymama" status="sp. nov.">
<emphasis id="B935EAD6BA0BFE2AF9C1FC186013FC79" bold="true" box="[1165,1354,918,939]" italics="true" pageId="6" pageNumber="7">Arctosa ayaymama</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20657ADBA0BFE2AF81CFC1860C5FC79" box="[1360,1436,918,939]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
<paragraph id="8BFE36C4BA0BFE2AFDDBFC3A60C5FC1A" blockId="6.[151,1437,885,1321]" box="[151,1436,947,969]" pageId="6" pageNumber="7">
<keyLead id="30B095F1BA0BFE2AFDDBFC3A60C5FC1A" box="[151,1436,947,969]" pageId="6" pageNumber="7">
- Median septum (MS) with a longitudinal raised ridge (
<figureCitation id="137A2A41BA0BFE2AFFA2FC3D6662FC1B" box="[750,827,947,969]" captionStart="FIGURE 28" captionStartId="35.[151,250,1908,1932]" captionTargetBox="[191,1397,191,1875]" captionTargetId="figure-21@35.[186,1402,181,1885]" captionTargetPageId="35" captionText="FIGURE 28. Arctosa sapiranga Silva &amp; Lise, 2009. Male pedipalp. a–c. Pedipalp, holotype, Rio Grande do Sul (MCTP 19999) (a, ventral; b, retrolateral; c, prolateral). d–e. Palpal bulb (d, anterior; e posterior). f–h. Paratype, São Paulo (MNRJ 6501), (f, retrolateral, g, ventral, h, anterior). Scale bars: a–h, 0.25mm. DMA depression of MA, E embolus, MA median apophysis, Pa Posterior arm, Ra Retrolateral arm, SD seminal duct, ST subtegulum, T tegulum, TA Terminal Apophysis." figureDoi="http://doi.org/10.5281/zenodo.10681321" httpUri="https://zenodo.org/record/10681321/files/figure.png" pageId="6" pageNumber="7">Fig. 28i</figureCitation>
).................................................
<emphasis id="B935EAD6BA0BFE2AF8DCFC3D60C5FC1A" bold="true" box="[1424,1436,947,968]" pageId="6" pageNumber="7">2</emphasis>
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<keyStep id="30B52E61BA0BFE2AFDDBFC5F60C5FBD1" pageId="6" pageNumber="7">
<paragraph id="8BFE36C4BA0BFE2AFDDBFC5F60C4FC34" blockId="6.[151,1437,885,1321]" box="[151,1437,976,998]" pageId="6" pageNumber="7">
<keyLead id="30B095F1BA0BFE2AFDDBFC5F60C4FC34" box="[151,1437,976,998]" pageId="6" pageNumber="7">
2 Transversal septum projected posteriorly (
<bibRefCitation id="EFD04B35BA0BFE2AFF3AFC5E6628FC34" author="Dondale, C. D. &amp; Redner, J. H." box="[630,881,976,998]" pageId="6" pageNumber="7" pagination="1 - 30" refId="ref48997" refString="Dondale, C. D. &amp; Redner, J. H. (1983 a) Revision of the wolf spiders of the genus Arctosa C. L. Koch in North and Central America (Araneae: Lycosidae). The Journal of Arachnology, 11 (1), 1 - 30." type="journal article" year="1983">Dondale &amp; Redner 1983a</bibRefCitation>
: fig. 70).............
<taxonomicName id="4C414D47BA0BFE2AF924FC5E60C4FC34" ID-CoL="GBGJ" authority="(Hentz, 1844)" baseAuthorityName="Hentz" baseAuthorityYear="1844" box="[1128,1437,976,998]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="littoralis">
<emphasis id="B935EAD6BA0BFE2AF924FC5E60C4FC34" bold="true" box="[1128,1437,976,998]" pageId="6" pageNumber="7">
<emphasis id="B935EAD6BA0BFE2AF924FC5E6054FC34" bold="true" box="[1128,1293,976,998]" italics="true" pageId="6" pageNumber="7">Arctosa littoralis</emphasis>
(
<bibRefCitation id="EFD04B35BA0BFE2AF856FC5E60CCFC34" author="Hentz, N. M." box="[1306,1429,976,998]" pageId="6" pageNumber="7" pagination="386 - 396" refId="ref49735" refString="Hentz, N. M. (1844) Descriptions and figures of the araneides of the United States. Boston Journal of Natural History, 4, 386 - 396. https: // doi. org / 10.5962 / bhl. part. 16998" type="journal article" year="1844">Hentz, 1844</bibRefCitation>
)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFE36C4BA0BFE2AFDDBFC6060C5FBD1" blockId="6.[151,1437,885,1321]" box="[151,1436,1006,1028]" pageId="6" pageNumber="7">
<keyLead id="30B095F1BA0BFE2AFDDBFC6060C5FBD1" box="[151,1436,1006,1028]" pageId="6" pageNumber="7">
- Transversal septum aligned with epigastric furrow (
<figureCitation id="137A2A41BA0BFE2AFF9DFC60667EFBD1" box="[721,807,1006,1028]" captionStart="FIGURE 44" captionStartId="51.[151,250,1909,1933]" captionTargetBox="[202,1384,190,1876]" captionTargetId="figure-21@51.[194,1394,181,1885]" captionTargetPageId="51" captionText="FIGURE 44. Arctosa villa sp. nov. Male pedipalp (MUSM-ENT 500104) a–c, f (a, ventral; b, retrolateral; c, sub anterior; f, anterior). Male prosoma d–e (d, lateral; e, frontal). Scale bars: a–c, f 0.25 mm, d–e 1 mm. CHI chillum, CND condylum, DMA depression of median apophysis, E embolus, Pa posterior arm, Pp palea protuberance, Ra retrolateral arm, TA terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.10681355" httpUri="https://zenodo.org/record/10681355/files/figure.png" pageId="6" pageNumber="7">Figs 44a</figureCitation>
,
<figureCitation id="137A2A41BA0BFE2AFE7CFC60660CFBD6" box="[816,853,1006,1028]" captionStart="FIGURE 45" captionStartId="52.[151,250,1851,1875]" captionTargetBox="[151,1436,181,1827]" captionTargetId="figure-25@52.[151,1436,181,1827]" captionTargetPageId="52" captionText="FIGURE 45. Arctosa villa sp. nov. Male pedipalp (MUSM-ENT 500104) a–c (a, retrolateral; b, ventral; c, prolateral). Pedipalp bulb (MUSM-ENT 505002) d–f, h (d, retrolateral; e, ventral; f, prolateral; h, anterior). g, spinnerets; i, chelicerae. Scale bars: a–c, 0.2 mm, d–f, h, 250 µm, g, 200 µm, i, 500 µm. DMA depression of median apophysis, E embolus, Pa posterior arm, Pp palea protuberance, Ra retrolateral arm, SD seminal duct, SF striae file, TA terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.10681357" httpUri="https://zenodo.org/record/10681357/files/figure.png" pageId="6" pageNumber="7">45b</figureCitation>
)..............................................
<emphasis id="B935EAD6BA0BFE2AF8DCFC6060C5FBD1" bold="true" box="[1424,1436,1006,1027]" pageId="6" pageNumber="7">3</emphasis>
</keyLead>
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</keyStep>
<keyStep id="30B52E61BA0BFE2AFDDBFB8560C5FB89" pageId="6" pageNumber="7">
<paragraph id="8BFE36C4BA0BFE2AFDDBFB8560C5FBEF" blockId="6.[151,1437,885,1321]" pageId="6" pageNumber="7">
<keyLead id="30B095F1BA0BFE2AFDDBFB8560C5FBEF" pageId="6" pageNumber="7">
3 Vulval chamber positioned below to the head of spermatheca, i.e., visible in dorsal view (
<bibRefCitation id="EFD04B35BA0BFE2AF970FB85605FFBF3" author="Roth, V. D. &amp; Brown, W. L." box="[1084,1286,1035,1057]" pageId="6" pageNumber="7" pagination="61 - 63" refId="ref52104" refString="Roth, V. D. &amp; Brown, W. L. (1976) A new wolf spider from the Gulf of California beaches. Journal of the Arizona Academy of Science, 11 (2), 61 - 63." type="journal article" year="1976">Roth &amp; Brown 1976</bibRefCitation>
: fig. 7)..............................................................................
<taxonomicName id="4C414D47BA0BFE2AF90AFBA660C5FBEF" ID-CoL="67S3X" authority="Roth &amp; Brown, 1976" authorityName="Roth &amp; Brown" authorityYear="1976" box="[1094,1436,1064,1086]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="serii">
<emphasis id="B935EAD6BA0BFE2AF90AFBA660C5FBEF" bold="true" box="[1094,1436,1064,1086]" pageId="6" pageNumber="7">
<emphasis id="B935EAD6BA0BFE2AF90AFBA6619AFBEF" bold="true" box="[1094,1219,1064,1085]" italics="true" pageId="6" pageNumber="7">Arctosa serii</emphasis>
<bibRefCitation id="EFD04B35BA0BFE2AF985FBA760C5FBEF" author="Roth, V. D. &amp; Brown, W. L." box="[1225,1436,1064,1086]" pageId="6" pageNumber="7" pagination="61 - 63" refId="ref52104" refString="Roth, V. D. &amp; Brown, W. L. (1976) A new wolf spider from the Gulf of California beaches. Journal of the Arizona Academy of Science, 11 (2), 61 - 63." type="journal article" year="1976">Roth &amp; Brown, 1976</bibRefCitation>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFE36C4BA0BFE2AFDDBFBC860C5FB89" blockId="6.[151,1437,885,1321]" box="[151,1436,1094,1116]" pageId="6" pageNumber="7">
<keyLead id="30B095F1BA0BFE2AFDDBFBC860C5FB89" box="[151,1436,1094,1116]" pageId="6" pageNumber="7">
- Vulval chamber positioned behind to the head of spermatheca, i.e., not visible in dorsal vew (
<figureCitation id="137A2A41BA0BFE2AF92DFBC861EDFB8E" box="[1121,1204,1094,1116]" captionStart="FIGURE 28" captionStartId="35.[151,250,1908,1932]" captionTargetBox="[191,1397,191,1875]" captionTargetId="figure-21@35.[186,1402,181,1885]" captionTargetPageId="35" captionText="FIGURE 28. Arctosa sapiranga Silva &amp; Lise, 2009. Male pedipalp. a–c. Pedipalp, holotype, Rio Grande do Sul (MCTP 19999) (a, ventral; b, retrolateral; c, prolateral). d–e. Palpal bulb (d, anterior; e posterior). f–h. Paratype, São Paulo (MNRJ 6501), (f, retrolateral, g, ventral, h, anterior). Scale bars: a–h, 0.25mm. DMA depression of MA, E embolus, MA median apophysis, Pa Posterior arm, Ra Retrolateral arm, SD seminal duct, ST subtegulum, T tegulum, TA Terminal Apophysis." figureDoi="http://doi.org/10.5281/zenodo.10681321" httpUri="https://zenodo.org/record/10681321/files/figure.png" pageId="6" pageNumber="7">Fig. 28h</figureCitation>
)................ ..
<emphasis id="B935EAD6BA0BFE2AF8DCFBC860C5FB89" bold="true" box="[1424,1436,1094,1115]" pageId="6" pageNumber="7">4</emphasis>
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</keyStep>
<keyStep id="30B52E61BA0BFE2AFDDBFBED60C5FB47" pageId="6" pageNumber="7">
<paragraph id="8BFE36C4BA0BFE2AFDDBFBED60C5FBAA" blockId="6.[151,1437,885,1321]" box="[151,1436,1123,1145]" pageId="6" pageNumber="7">
<keyLead id="30B095F1BA0BFE2AFDDBFBED60C5FBAA" box="[151,1436,1123,1145]" pageId="6" pageNumber="7">
4 Head of spermatheca rectangular (
<figureCitation id="137A2A41BA0BFE2AFF67FBED67D8FBAB" box="[555,641,1123,1145]" captionStart="FIGURE 12" captionStartId="17.[151,250,1236,1260]" captionTargetBox="[162,1425,190,1201]" captionTargetId="figure-21@17.[151,1436,181,1211]" captionTargetPageId="17" captionText="FIGURE 12. Arctosa costenola sp. nov. Male pedipalp holotype (ICN-Ar 1378) a–c (a, anterior; b, ventral; c, retrolateral). Scale bars: a–c, 0.25 mm. CMS cymbial macrosetae, DMA depression of median apophysis, E embolus, Pa posterior arm, Pp palea protuberance, Ra retrolateral arm, ST subtegulum, T tegulum, TA terminal apophysis" figureDoi="http://doi.org/10.5281/zenodo.10681289" httpUri="https://zenodo.org/record/10681289/files/figure.png" pageId="6" pageNumber="7">Figs 12b</figureCitation>
,
<figureCitation id="137A2A41BA0BFE2AFFC0FBED67F6FBAA" box="[652,687,1123,1144]" captionStart="FIGURE 13" captionStartId="18.[151,250,1909,1933]" captionTargetBox="[177,1411,181,1885]" captionTargetId="figure-25@18.[177,1411,181,1885]" captionTargetPageId="18" captionText="FIGURE 13. Arctosa costenola sp. nov. Male pedipalp (ICN-Ar 1378) a–c (a, prolateral; b, ventral; c, retrolateral). Pedipalp bulb d–h (d, prolateral; e, ventral; f, retrolateral; g posterior; h, anterior). i dorsal cymbium macrosetae. Scale bars: a–c, 0.20 mm, d–h, 100 µm, i, 200 µm. C conductor, DMA depression of median apophysis, E embolus, Ra retrolateral arm, Pa posterior arm, T tegulum, TA terminal apophysis, TK tegular keel, ST subtegulum," figureDoi="http://doi.org/10.5281/zenodo.10681291" httpUri="https://zenodo.org/record/10681291/files/figure.png" pageId="6" pageNumber="7">13e</figureCitation>
)............................................................
<emphasis id="B935EAD6BA0BFE2AF8DCFBED60C5FBAA" bold="true" box="[1424,1436,1123,1144]" pageId="6" pageNumber="7">5</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BFE36C4BA0BFE2AFDDBFB0F60C5FB47" blockId="6.[151,1437,885,1321]" box="[151,1436,1152,1174]" pageId="6" pageNumber="7">
<keyLead id="30B095F1BA0BFE2AFDDBFB0F60C5FB47" box="[151,1436,1152,1174]" pageId="6" pageNumber="7">
- Head of spermatheca triangular or pear-shaped (
<figureCitation id="137A2A41BA0BFE2AFFFCFB0E665CFB44" box="[688,773,1152,1174]" captionStart="FIGURE 28" captionStartId="35.[151,250,1908,1932]" captionTargetBox="[191,1397,191,1875]" captionTargetId="figure-21@35.[186,1402,181,1885]" captionTargetPageId="35" captionText="FIGURE 28. Arctosa sapiranga Silva &amp; Lise, 2009. Male pedipalp. a–c. Pedipalp, holotype, Rio Grande do Sul (MCTP 19999) (a, ventral; b, retrolateral; c, prolateral). d–e. Palpal bulb (d, anterior; e posterior). f–h. Paratype, São Paulo (MNRJ 6501), (f, retrolateral, g, ventral, h, anterior). Scale bars: a–h, 0.25mm. DMA depression of MA, E embolus, MA median apophysis, Pa Posterior arm, Ra Retrolateral arm, SD seminal duct, ST subtegulum, T tegulum, TA Terminal Apophysis." figureDoi="http://doi.org/10.5281/zenodo.10681321" httpUri="https://zenodo.org/record/10681321/files/figure.png" pageId="6" pageNumber="7">Figs 28e</figureCitation>
,
<figureCitation id="137A2A41BA0BFE2AFE43FB0E666DFB44" box="[783,820,1152,1174]" captionStart="FIGURE 44" captionStartId="51.[151,250,1909,1933]" captionTargetBox="[202,1384,190,1876]" captionTargetId="figure-21@51.[194,1394,181,1885]" captionTargetPageId="51" captionText="FIGURE 44. Arctosa villa sp. nov. Male pedipalp (MUSM-ENT 500104) a–c, f (a, ventral; b, retrolateral; c, sub anterior; f, anterior). Male prosoma d–e (d, lateral; e, frontal). Scale bars: a–c, f 0.25 mm, d–e 1 mm. CHI chillum, CND condylum, DMA depression of median apophysis, E embolus, Pa posterior arm, Pp palea protuberance, Ra retrolateral arm, TA terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.10681355" httpUri="https://zenodo.org/record/10681355/files/figure.png" pageId="6" pageNumber="7">44d</figureCitation>
)................................................ ..
<emphasis id="B935EAD6BA0BFE2AF8DCFB0E60C5FB47" bold="true" box="[1424,1436,1152,1173]" pageId="6" pageNumber="7">6</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B52E61BA0BFE2AFDDBFB1060C5FB3F" pageId="6" pageNumber="7">
<paragraph id="8BFE36C4BA0BFE2AFDDBFB1060C5FB61" blockId="6.[151,1437,885,1321]" box="[151,1436,1181,1204]" pageId="6" pageNumber="7">
<keyLead id="30B095F1BA0BFE2AFDDBFB1060C5FB61" box="[151,1436,1181,1204]" pageId="6" pageNumber="7">
5 Epigynal plate projected into the atrium (
<figureCitation id="137A2A41BA0BFE2AFF21FB10679BFB61" box="[621,706,1182,1204]" captionStart="FIGURE 12" captionStartId="17.[151,250,1236,1260]" captionTargetBox="[162,1425,190,1201]" captionTargetId="figure-21@17.[151,1436,181,1211]" captionTargetPageId="17" captionText="FIGURE 12. Arctosa costenola sp. nov. Male pedipalp holotype (ICN-Ar 1378) a–c (a, anterior; b, ventral; c, retrolateral). Scale bars: a–c, 0.25 mm. CMS cymbial macrosetae, DMA depression of median apophysis, E embolus, Pa posterior arm, Pp palea protuberance, Ra retrolateral arm, ST subtegulum, T tegulum, TA terminal apophysis" figureDoi="http://doi.org/10.5281/zenodo.10681289" httpUri="https://zenodo.org/record/10681289/files/figure.png" pageId="6" pageNumber="7">Figs 12a</figureCitation>
,
<figureCitation id="137A2A41BA0BFE2AFF80FB1067A9FB66" box="[716,752,1182,1204]" captionStart="FIGURE 13" captionStartId="18.[151,250,1909,1933]" captionTargetBox="[177,1411,181,1885]" captionTargetId="figure-25@18.[177,1411,181,1885]" captionTargetPageId="18" captionText="FIGURE 13. Arctosa costenola sp. nov. Male pedipalp (ICN-Ar 1378) a–c (a, prolateral; b, ventral; c, retrolateral). Pedipalp bulb d–h (d, prolateral; e, ventral; f, retrolateral; g posterior; h, anterior). i dorsal cymbium macrosetae. Scale bars: a–c, 0.20 mm, d–h, 100 µm, i, 200 µm. C conductor, DMA depression of median apophysis, E embolus, Ra retrolateral arm, Pa posterior arm, T tegulum, TA terminal apophysis, TK tegular keel, ST subtegulum," figureDoi="http://doi.org/10.5281/zenodo.10681291" httpUri="https://zenodo.org/record/10681291/files/figure.png" pageId="6" pageNumber="7">13b</figureCitation>
)...................................
<emphasis id="B935EAD6BA0BFE2AF9D6FB1060C5FB61" bold="true" box="[1178,1436,1181,1203]" pageId="6" pageNumber="7">
<taxonomicName id="4C414D47BA0BFE2AF9D6FB106010FB61" authority="Paredes-Munguía &amp; Brescovit &amp; Teixeira, 2024" authorityName="Paredes-Munguía &amp; Brescovit &amp; Teixeira" authorityYear="2024" box="[1178,1353,1181,1203]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="costenola" status="sp. nov.">
<emphasis id="B935EAD6BA0BFE2AF9D6FB106010FB61" bold="true" box="[1178,1353,1181,1203]" italics="true" pageId="6" pageNumber="7">Arctosa costenola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20657ADBA0BFE2AF81CFB1060C5FB61" box="[1360,1436,1182,1203]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BFE36C4BA0BFE2AFDDBFB3260C5FB3F" blockId="6.[151,1437,885,1321]" pageId="6" pageNumber="7">
<keyLead id="30B095F1BA0BFE2AFDDBFB3260C5FB3F" pageId="6" pageNumber="7">
- Epyginal plate not projected (
<bibRefCitation id="EFD04B35BA0BFE2AFCB2FB3567A1FB02" author="Jimenez, M. L. &amp; Dondale, C. D." box="[510,760,1211,1233]" pageId="6" pageNumber="7" pagination="115 - 116" refId="ref49943" refString="Jimenez, M. L. &amp; Dondale, C. D. (1984) A new species of Arctosa from Guerrero, Mexico (Araneae, Lycosidae). The Journal of Arachnology, 12 (1), 115 - 116." type="journal article" year="1984">Jiménez &amp; Dondale 1984</bibRefCitation>
: fig. 1)................................................................................................................
<taxonomicName id="4C414D47BA0BFE2AFE9CFB5660C5FB3F" ID-CoL="68443" authority="Jimenez &amp; Dondale, 1984" authorityName="Jimenez &amp; Dondale" authorityYear="1984" box="[976,1436,1240,1262]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="denticulata">
<emphasis id="B935EAD6BA0BFE2AFE9CFB5660C5FB3F" bold="true" box="[976,1436,1240,1262]" pageId="6" pageNumber="7">
<emphasis id="B935EAD6BA0BFE2AFE9CFB5661C8FB3C" bold="true" box="[976,1169,1240,1262]" italics="true" pageId="6" pageNumber="7">Arctosa denticulata</emphasis>
<bibRefCitation id="EFD04B35BA0BFE2AF9DBFB5660C5FB3F" author="Jimenez, M. L. &amp; Dondale, C. D." box="[1175,1436,1240,1262]" pageId="6" pageNumber="7" pagination="115 - 116" refId="ref49943" refString="Jimenez, M. L. &amp; Dondale, C. D. (1984) A new species of Arctosa from Guerrero, Mexico (Araneae, Lycosidae). The Journal of Arachnology, 12 (1), 115 - 116." type="journal article" year="1984">Jiménez &amp; Dondale, 1984</bibRefCitation>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B52E61BA0BFE2AFDDBFB7860C5FAFA" pageId="6" pageNumber="7">
<paragraph id="8BFE36C4BA0BFE2AFDDBFB7860C5FAD9" blockId="6.[151,1437,885,1321]" box="[151,1436,1270,1292]" pageId="6" pageNumber="7">
<keyLead id="30B095F1BA0BFE2AFDDBFB7860C5FAD9" box="[151,1436,1270,1292]" pageId="6" pageNumber="7">
6 Head of spermatheca pear-shaped (
<figureCitation id="137A2A41BA0BFE2AFF7FFB7867D1FAD9" box="[563,648,1270,1292]" captionStart="FIGURE 28" captionStartId="35.[151,250,1908,1932]" captionTargetBox="[191,1397,191,1875]" captionTargetId="figure-21@35.[186,1402,181,1885]" captionTargetPageId="35" captionText="FIGURE 28. Arctosa sapiranga Silva &amp; Lise, 2009. Male pedipalp. a–c. Pedipalp, holotype, Rio Grande do Sul (MCTP 19999) (a, ventral; b, retrolateral; c, prolateral). d–e. Palpal bulb (d, anterior; e posterior). f–h. Paratype, São Paulo (MNRJ 6501), (f, retrolateral, g, ventral, h, anterior). Scale bars: a–h, 0.25mm. DMA depression of MA, E embolus, MA median apophysis, Pa Posterior arm, Ra Retrolateral arm, SD seminal duct, ST subtegulum, T tegulum, TA Terminal Apophysis." figureDoi="http://doi.org/10.5281/zenodo.10681321" httpUri="https://zenodo.org/record/10681321/files/figure.png" pageId="6" pageNumber="7">Figs 28e</figureCitation>
,
<figureCitation id="137A2A41BA0BFE2AFFDEFB7867EEFADE" box="[658,695,1270,1292]" captionStart="FIGURE 32" captionStartId="39.[151,250,1401,1425]" captionTargetBox="[151,1436,181,1378]" captionTargetId="figure-21@39.[151,1436,181,1378]" captionTargetPageId="39" captionText="FIGURE 32. Arctosa sapiranga Silva &amp; Lise, 2009. Male. Pedipalp (MCTP 28548) (a, ventral; b, detail TA; c, close up of MA; d, detail Pa; e, protuberance on tegulum). Scale bars: a, 250 µm; b, e 50 µm; c, 100 µm; d, 25 µm. CMS cymbium seate, E embolus, MA median apophysis, Pa posterior arm, Pp palea protuberance, Ra retrolateral arm, ST subtegulum, T tegulum, TA terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.10681329" httpUri="https://zenodo.org/record/10681329/files/figure.png" pageId="6" pageNumber="7">32b</figureCitation>
)...............................
<taxonomicName id="4C414D47BA0BFE2AF962FB7860C5FAD9" ID-CoL="GBJ6" authority="Silva &amp; Lise 2009" authorityName="Silva &amp; Lise" authorityYear="2009" box="[1070,1436,1270,1292]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="sapiranga">
<emphasis id="B935EAD6BA0BFE2AF962FB7860C5FAD9" bold="true" box="[1070,1436,1270,1292]" pageId="6" pageNumber="7">
<emphasis id="B935EAD6BA0BFE2AF962FB7861BAFAD9" bold="true" box="[1070,1251,1270,1291]" italics="true" pageId="6" pageNumber="7">Arctosa sapiranga</emphasis>
<bibRefCitation id="EFD04B35BA0BFE2AF9A5FB7860C5FAD9" author="Silva, E. L. C. da &amp; Lise, A. A." box="[1257,1436,1270,1292]" pageId="6" pageNumber="7" pagination="27 - 32" refId="ref52382" refString="Silva, E. L. C. da &amp; Lise, A. A. (2009) A new species of the wolf spider genus Arctosa (Araneae, Lycosidae) from Southern Brazil. Revista Peruana de Biologia, 15 (2), 27 - 32. https: // doi. org / 10.15381 / rpb. v 15 i 2.1698" type="journal article" year="2009">Silva &amp; Lise 2009</bibRefCitation>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BFE36C4BA0BFE2AFDDBFA9A60C5FAFA" blockId="6.[151,1437,885,1321]" box="[151,1436,1299,1321]" pageId="6" pageNumber="7">
<keyLead id="30B095F1BA0BFE2AFDDBFA9A60C5FAFA" box="[151,1436,1299,1321]" pageId="6" pageNumber="7">
- Head of spermatheca triangular (
<figureCitation id="137A2A41BA0BFE2AFF50FA9D672AFAFB" box="[540,627,1299,1321]" captionStart="FIGURE 44" captionStartId="51.[151,250,1909,1933]" captionTargetBox="[202,1384,190,1876]" captionTargetId="figure-21@51.[194,1394,181,1885]" captionTargetPageId="51" captionText="FIGURE 44. Arctosa villa sp. nov. Male pedipalp (MUSM-ENT 500104) a–c, f (a, ventral; b, retrolateral; c, sub anterior; f, anterior). Male prosoma d–e (d, lateral; e, frontal). Scale bars: a–c, f 0.25 mm, d–e 1 mm. CHI chillum, CND condylum, DMA depression of median apophysis, E embolus, Pa posterior arm, Pp palea protuberance, Ra retrolateral arm, TA terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.10681355" httpUri="https://zenodo.org/record/10681355/files/figure.png" pageId="6" pageNumber="7">Figs 44d</figureCitation>
,
<figureCitation id="137A2A41BA0BFE2AFF31FA9D67FBFAFB" box="[637,674,1299,1321]" captionStart="FIGURE 45" captionStartId="52.[151,250,1851,1875]" captionTargetBox="[151,1436,181,1827]" captionTargetId="figure-25@52.[151,1436,181,1827]" captionTargetPageId="52" captionText="FIGURE 45. Arctosa villa sp. nov. Male pedipalp (MUSM-ENT 500104) a–c (a, retrolateral; b, ventral; c, prolateral). Pedipalp bulb (MUSM-ENT 505002) d–f, h (d, retrolateral; e, ventral; f, prolateral; h, anterior). g, spinnerets; i, chelicerae. Scale bars: a–c, 0.2 mm, d–f, h, 250 µm, g, 200 µm, i, 500 µm. DMA depression of median apophysis, E embolus, Pa posterior arm, Pp palea protuberance, Ra retrolateral arm, SD seminal duct, SF striae file, TA terminal apophysis." figureDoi="http://doi.org/10.5281/zenodo.10681357" httpUri="https://zenodo.org/record/10681357/files/figure.png" pageId="6" pageNumber="7">45b</figureCitation>
).............................................
<emphasis id="B935EAD6BA0BFE2AF981FA9D60C5FAFA" bold="true" box="[1229,1436,1299,1321]" pageId="6" pageNumber="7">
<taxonomicName id="4C414D47BA0BFE2AF981FA9D6013FAFB" authority="Paredes-Munguía &amp; Brescovit &amp; Teixeira, 2024" authorityName="Paredes-Munguía &amp; Brescovit &amp; Teixeira" authorityYear="2024" box="[1229,1354,1299,1321]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="villa" status="sp. nov.">
<emphasis id="B935EAD6BA0BFE2AF981FA9D6013FAFB" bold="true" box="[1229,1354,1299,1321]" italics="true" pageId="6" pageNumber="7">Arctosa villa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20657ADBA0BFE2AF81CFA9D60C5FAFA" box="[1360,1436,1299,1320]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>
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