treatments-xml/data/03/E8/87/03E887BFFFBFFFC5FF01FACFFDADFE37.xml

<document id="04EE715E9FA878DB1FEBDD57D549296D" ID-CLB-Dataset="21736" ID-DOI="10.37520/aemnp.2020.020" ID-GBIF-Dataset="4eddf6bc-be69-4711-b8e6-88301f143906" ID-ISSN="1804-6487" ID-Zenodo-Dep="3879983" ID-ZooBank="D491AC43-3847-4C9C-9C03-B9EABB679A19" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1591390439932" checkinUser="tatiana" docAuthor="Kondorosy, Előd, Zámbó, András, Kovács, Szilvia, Baňař, Petr &amp; Km, Recently" docDate="2020" docId="03E887BFFFBFFFC5FF01FACFFDADFE37" docLanguage="en" docName="ActaEntMusNatPra.60.1.343-352.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 60 (1)" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Scobinigaster Kondorosy &amp; Banar 2020, gen. nov." docType="treatment" docUuid="BE7CA4BC-E76A-4ACA-B4E4-CF3B0DCCDECE" docUuidSource="ZooBank" docVersion="7" lastPageNumber="352" masterDocId="FFD1FFC7FFBEFFCCFFCAFFF2FF98FFC2" masterDocTitle="Scobinigaster, a new genus of Lethaeini from Madagascar (Hemiptera: Heteroptera: Rhyparochromidae)" masterLastPageNumber="352" masterPageNumber="343" pageNumber="344" updateTime="1698842130349" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="A97E44C7DF0D0A6AF3D241BDBA889F72">Scobinigaster, a new genus of Lethaeini from Madagascar (Hemiptera: Heteroptera: Rhyparochromidae)</mods:title>
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<mods:namePart id="2C0E149033A1C3E80F26C383330B8C55">Kondorosy, Előd</mods:namePart>
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<mods:namePart id="806A37F00D5A6FF8E450B53A61006B0E">Zámbó, András</mods:namePart>
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<mods:namePart id="31B84DD7760BAE15E16B07FEECAE8FE1">Kovács, Szilvia</mods:namePart>
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<mods:namePart id="613F394EE9698F3E5571FE42E78773FD">Baňař, Petr</mods:namePart>
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<paragraph id="8BFE36A9FFBFFFCDFF01FACFFD59FA97" blockId="1.[203,705,1293,1365]" box="[203,705,1341,1365]" pageId="1" pageNumber="344">
<heading id="D0B681C5FFBFFFCDFF01FACFFD59FA97" box="[203,705,1341,1365]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="344" reason="3">
<emphasis id="B935EABBFFBFFFCDFF01FACFFD59FA97" bold="true" box="[203,705,1341,1365]" pageId="1" pageNumber="344">
<taxonomicName id="4C414D2AFFBFFFCDFF01FACFFDC9FA97" ID-CoL="8Q9BG" authority="Kondorosy &amp; Banar" authorityName="Kondorosy &amp; Banar" authorityYear="2020" box="[203,593,1341,1365]" class="Insecta" family="Rhyparochromidae" genus="Scobinigaster" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="344" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B935EABBFFBFFFCDFF01FACFFEF9FA97" bold="true" box="[203,353,1341,1365]" italics="true" pageId="1" pageNumber="344">Scobinigaster</emphasis>
Kondorosy &amp; Baňař
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, 
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<emphasis id="B935EABBFFBFFFCDFF5CFA9FFEB2FA46" bold="true" box="[150,298,1389,1412]" pageId="1" pageNumber="344">
<typeStatus id="54FA880BFFBFFFCDFF5CFA9FFF56FA46" box="[150,206,1389,1412]" pageId="1" pageNumber="344">Type</typeStatus>
species.
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<taxonomicName id="4C414D2AFFBFFFCDFEFBFA9FFD6AFA46" ID-CoL="8QND8" authority="Kondorosy &amp; Banar" authorityName="Kondorosy &amp; Banar" authorityYear="2020" box="[305,754,1389,1412]" class="Insecta" family="Rhyparochromidae" genus="Scobinigaster" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="344" phylum="Arthropoda" rank="species" species="henryi" status="sp. nov.">
<emphasis id="B935EABBFFBFFFCDFEFBFA9FFD97FA46" box="[305,527,1389,1412]" italics="true" pageId="1" pageNumber="344">Scobinigaster henryi</emphasis>
Kondorosy &amp; Baňař
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, 
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, present designation.
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<paragraph id="8BFE36A9FFBFFFCDFF5CFA4EFDF1F8D1" blockId="1.[150,759,1468,2067]" pageId="1" pageNumber="344">
<emphasis id="B935EABBFFBFFFCDFF5CFA4EFE86FA11" bold="true" box="[150,286,1468,1491]" pageId="1" pageNumber="344">Description.</emphasis>
<emphasis id="B935EABBFFBFFFCDFEE9FA4EFEC4FA11" box="[291,348,1468,1491]" italics="true" pageId="1" pageNumber="344">Head</emphasis>
moderately short, wider than long, with laterally protruding, moderately big and well-developed eyes, ocelli situated close to them. Antennae rather short, all antennomeres covered with fine semidecumbent setae. Scape in apical half abruptly widened, pedicel linear, basiflagellum gradually slightly widened, distiflagellum almost linear. Ventral surface of head densely punctate reaching to eyes. Maxillary plates broadened with a vertical keel on apex, appearing as minute, tooth-shaped projections (Figs 9–10, arrows). Labium long, extending to base of abdomen, labiomere I reaching prosternum.
</paragraph>
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<emphasis id="B935EABBFFBFFFCDFF7EF8EEFE96F8F1" box="[180,270,1820,1843]" italics="true" pageId="1" pageNumber="344">Thorax.</emphasis>
Pronotum with wide and medially much broader collar, delimited with an impressed row of punctures (variable among specimens); lateral margin well developed, about as wide as basal part of scape; anterolateral trichobothria situated on lateral margin, its distance from anterior edge equal to the width of lateral margin; midline of pronotum variable among specimens – always raised on collar and most of posterior lobe, often vanished on anterior half of pronotum and near the posterior margin (Fig. 21). Anterior lobe of pronotum slightly longer than posterior lobe, transverse impression between anterior and posterior lobe absent. Surface with some small, variably developed impunctate spots: two pairs of dark, finely wrinkled spots on calli, and one usually visible shiny pair on posterior margin.
</paragraph>
<paragraph id="8BFE36A9FFBFFFCDFC92FE6CFAECFDF7" blockId="1.[826,1435,190,2069]" pageId="1" pageNumber="344">Scutellum elongate, triangular, coarsely punctate, with two very finely wrinkled, dark impunctate spots on anterolateral corners; with a low midline, being more prominent on posterior half, and more or less developed sublateral smooth shiny yellow keels in central part.</paragraph>
<paragraph id="8BFE36A9FFBFFFCDFC92FDCCFB98FC17" blockId="1.[826,1435,190,2069]" pageId="1" pageNumber="344">Clavus in brachypterous specimens not separated from corium, with four rows of punctures (the inner three ones being irregular). Corium with dense but irregularly organised punctation, only punctures along claval suture forming a regular row. Vein R strongly elevated along its entire length. Apical margin of corium convex in brachypterous specimens, with arched rudiment of membrane only slightly wider than tibiae, leaving tergite VI in greater part and VII fully exposed. Macropterous specimen similar, but apical margin of corium slightly S-shaped, membrane well developed, almost reaching apex of abdomen, with cross-veins forming two almost closed basal cells.</paragraph>
<paragraph id="8BFE36A9FFBFFFCDFC92FC2CFB26FA97" blockId="1.[826,1435,190,2069]" pageId="1" pageNumber="344">
Thoracic venter densely punctate, appearing wrinkled; mesosternum impunctate with fine transverse wrinkles, metasternum with longitudinal median keel, dorsal parts of all pleura impunctate as well, with similar surface as basal spots of scutellum. Peritreme of metathoracic scent glands moderately long, with apical part curving posteriad. Evaporatorium (
<figureCitation id="137A2A2CFFBFFFCDFBE6FB6CFB34FB77" box="[1068,1196,1182,1205]" captionStart="Figs 13–18" captionStartId="5.[151,188,1665,1684]" captionTargetBox="[151,1436,191,1646]" captionTargetId="figure@5.[151,1437,191,1646]" captionTargetPageId="5" captionText="Figs 13–18. Scobinigaster henryi Kondorosy &amp; Baňař sp. nov., scanning electron micrographs. 13–17 – paratype, female: 13 – metathorax with position of peritreme and evaporatium; 14 – detail of peritreme and evaporatorium; 15 – trichobothria on segments III and IV, lateroventral view; 16 – detail of trichobothria on segment IV, lateroventral view; 17 – trichobothria on segment VI, lateroventral view.18 – holotype, male, pygophore, posterodorsal view." figureDoi="http://doi.org/10.5281/zenodo.3879985" httpUri="https://zenodo.org/record/3879985/files/figure.png" pageId="1" pageNumber="344">Figs 13–14</figureCitation>
) rather small, lateral margin convex, not surpassing half of metapleuron, anterolaterally narrowly extended along anterior metapleural margin, reaching lateral end of metathoracic spiracle, and on posterior margin of mesopleuron, reaching about midlength of metathoracic spiracle.
</paragraph>
<paragraph id="8BFE36A9FFBFFFCDFC92FAACFAB9F8D7" blockId="1.[826,1435,190,2069]" pageId="1" pageNumber="344">Coxae unarmed, femora with two complete ventral rows of strong stiff setae, similar to those of tibiae; on metafemora also with some additional strong dorsal setae in posterior half. Profemora with three very short toothlike setae subapically in anteroventral row; both rows finer than on other legs (posteroventral row especially fine). Profemora moderately thickened; metafemora of males thicker than pro- and mesofemora, with several tiny teeth between stiff setae along all its length (on females thinner and without teeth). Tibiae straight, protibia with one ventral row, meso- and metatibiae with four rows of strong stiff setae being at least so long as width of tibiae. Metatarsus longer than remaining tarsi, tarsomere I about twice as long as tarsomeres II–III combined.</paragraph>
<paragraph id="8BFE36A9FFBFFFCEFC92F8ECFBF3F7D8" blockId="1.[826,1435,190,2069]" lastBlockId="2.[828,1437,1891,2074]" lastPageId="2" lastPageNumber="345" pageId="1" pageNumber="344">
<emphasis id="B935EABBFFBFFFCDFC92F8ECFC27F8F7" box="[856,959,1822,1845]" italics="true" pageId="1" pageNumber="344">Abdomen</emphasis>
robust, rather high in lateral view especially at segments IV–V, ventrally convex with smooth median keel (Figs 3, 8), abdomen in both sexes strongly dorsoventrally flattened posteriad of segment V. Connexivum of males ending abruptly usually with a tiny spine at 1/3 length of tergite VII (Figs 1, 4), in females running narrowly to tergite VIII, without prominent posterior angle. Tergite VII of females posteromedially arcuately excavated, in males basally with lateral keel vanishing at tip of connexivum, convexly bending posteriorly and laterally, with sternite VII fully enclosing genital segments; surface of tergite VII coarsely granulate at base of setae (finer than on ventral side). Ventral side of abdomen covered with long setae; their base provided with rasp-like coarse granules. Sternite VII of males with 3 hardly visible apical teeth. Ovipositor (Fig. 8) almost dividing sternite VI, sternite V not narrowed medially. Positions of abdominal spiracles, and number and pattern of abdominal trichobothria, as typical for 
<taxonomicName id="4C414D2AFFBCFFCEFAC9F811FAF2F838" authorityName="Stål" authorityYear="1872" box="[1283,1386,2019,2042]" class="Insecta" family="Rhyparochromidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="345" phylum="Arthropoda" rank="tribe" tribe="Lethaeini">Lethaeini</taxonomicName>
(see also Differential diagnosis).
</paragraph>
<paragraph id="8BFE36A9FFBDFFC8FF7FF8B8FEECFA81" blockId="3.[151,760,1866,2081]" lastBlockId="4.[151,760,1292,1347]" lastPageId="4" lastPageNumber="347" pageId="3" pageNumber="346">
<emphasis id="B935EABBFFBDFFCFFF7FF8B8FEBCF8A3" box="[181,292,1866,1889]" italics="true" pageId="3" pageNumber="346">Genitalia.</emphasis>
Male genital capsule (
<figureCitation id="137A2A2CFFBDFFCFFDD2F8B8FDF3F8A3" box="[536,619,1866,1889]" captionStart="Figs 13–18" captionStartId="5.[151,188,1665,1684]" captionTargetBox="[151,1436,191,1646]" captionTargetId="figure@5.[151,1437,191,1646]" captionTargetPageId="5" captionText="Figs 13–18. Scobinigaster henryi Kondorosy &amp; Baňař sp. nov., scanning electron micrographs. 13–17 – paratype, female: 13 – metathorax with position of peritreme and evaporatium; 14 – detail of peritreme and evaporatorium; 15 – trichobothria on segments III and IV, lateroventral view; 16 – detail of trichobothria on segment IV, lateroventral view; 17 – trichobothria on segment VI, lateroventral view.18 – holotype, male, pygophore, posterodorsal view." figureDoi="http://doi.org/10.5281/zenodo.3879985" httpUri="https://zenodo.org/record/3879985/files/figure.png" pageId="3" pageNumber="346">Figs 18</figureCitation>
, 22, 26) subglobose, posterior half with fine decumbent pubescence, posterior aperture nearly quadrangular, anterior half with two tiny, blunt teeth on each side, cup-like sclerite large, reaching more than half length of aperture, with one medial and two lateral apices (
<figureCitation id="137A2A2CFFBDFFCFFDECF818FDE3F7C3" box="[550,635,2026,2049]" captionStart="Figs 23–27" captionStartId="7.[150,187,1038,1057]" captionTargetBox="[251,1259,206,1012]" captionTargetId="figure@7.[221,1366,189,1019]" captionTargetPageId="7" captionText="Figs 23–27. Scobinigaster gen. nov. species. 23–24, 26–27 – S. henryi Kondorosy &amp; Baňař sp. nov., holotype, male; 23 – paramere, ventral view; 24 – paramere, lateral view; 26 – pygophore, posterior view; 27 – paratype, female, spermatheca. 25 – S. paveli Kondorosy &amp; Baňař sp. nov., holotype, male, paramere, ventral view, hairs omitted." figureDoi="http://doi.org/10.5281/zenodo.3879987" httpUri="https://zenodo.org/record/3879987/files/figure.png" pageId="3" pageNumber="346">Fig. 26</figureCitation>
). Paramere broad, shank short, blade subquadrate, ventral side (
<figureCitation id="137A2A2CFFBDFFCFFD0DF7F8FC18F8A3" box="[711,896,1866,2081]" captionStart="Figs 23–27" captionStartId="7.[150,187,1038,1057]" captionTargetBox="[251,1259,206,1012]" captionTargetId="figure@7.[221,1366,189,1019]" captionTargetPageId="7" captionText="Figs 23–27. Scobinigaster gen. nov. species. 23–24, 26–27 – S. henryi Kondorosy &amp; Baňař sp. nov., holotype, male; 23 – paramere, ventral view; 24 – paramere, lateral view; 26 – pygophore, posterior view; 27 – paratype, female, spermatheca. 25 – S. paveli Kondorosy &amp; Baňař sp. nov., holotype, male, paramere, ventral view, hairs omitted." figureDoi="http://doi.org/10.5281/zenodo.3879987" httpUri="https://zenodo.org/record/3879987/files/figure.png" pageId="3" pageNumber="346">Figs 23, 25</figureCitation>
) more or less flat, with a fine medial keel in apical one-third, apical tip curving ventrally (
<figureCitation id="137A2A2CFFBDFFCFFB3DF898FAD5F843" box="[1271,1357,1898,1921]" captionStart="Figs 23–27" captionStartId="7.[150,187,1038,1057]" captionTargetBox="[251,1259,206,1012]" captionTargetId="figure@7.[221,1366,189,1019]" captionTargetPageId="7" captionText="Figs 23–27. Scobinigaster gen. nov. species. 23–24, 26–27 – S. henryi Kondorosy &amp; Baňař sp. nov., holotype, male; 23 – paramere, ventral view; 24 – paramere, lateral view; 26 – pygophore, posterior view; 27 – paratype, female, spermatheca. 25 – S. paveli Kondorosy &amp; Baňař sp. nov., holotype, male, paramere, ventral view, hairs omitted." figureDoi="http://doi.org/10.5281/zenodo.3879987" httpUri="https://zenodo.org/record/3879987/files/figure.png" pageId="3" pageNumber="346">Fig. 24</figureCitation>
); inner projection (on left side of 
<figureCitation id="137A2A2CFFBDFFCFFB9EF878FB55F863" box="[1108,1229,1930,1953]" captionStart="Figs 23–27" captionStartId="7.[150,187,1038,1057]" captionTargetBox="[251,1259,206,1012]" captionTargetId="figure@7.[221,1366,189,1019]" captionTargetPageId="7" captionText="Figs 23–27. Scobinigaster gen. nov. species. 23–24, 26–27 – S. henryi Kondorosy &amp; Baňař sp. nov., holotype, male; 23 – paramere, ventral view; 24 – paramere, lateral view; 26 – pygophore, posterior view; 27 – paratype, female, spermatheca. 25 – S. paveli Kondorosy &amp; Baňař sp. nov., holotype, male, paramere, ventral view, hairs omitted." figureDoi="http://doi.org/10.5281/zenodo.3879987" httpUri="https://zenodo.org/record/3879987/files/figure.png" pageId="3" pageNumber="346">Figs 23, 25</figureCitation>
) rectangular; outer projection slightly rounded; in lateral view almost evenly thickened to base of blade to a strong hump (on left side of 
<figureCitation id="137A2A2CFFBDFFCFFC90F818FC34F7C3" box="[858,940,2026,2049]" captionStart="Figs 23–27" captionStartId="7.[150,187,1038,1057]" captionTargetBox="[251,1259,206,1012]" captionTargetId="figure@7.[221,1366,189,1019]" captionTargetPageId="7" captionText="Figs 23–27. Scobinigaster gen. nov. species. 23–24, 26–27 – S. henryi Kondorosy &amp; Baňař sp. nov., holotype, male; 23 – paramere, ventral view; 24 – paramere, lateral view; 26 – pygophore, posterior view; 27 – paratype, female, spermatheca. 25 – S. paveli Kondorosy &amp; Baňař sp. nov., holotype, male, paramere, ventral view, hairs omitted." figureDoi="http://doi.org/10.5281/zenodo.3879987" httpUri="https://zenodo.org/record/3879987/files/figure.png" pageId="3" pageNumber="346">Fig. 24</figureCitation>
); dorsal side with dense, relatively long semi -decumbent or erect pubescence. Female spermatheca with a ring-like structure broadened to a globular apical receptacle (
<figureCitation id="137A2A2CFFBAFFC8FEDFFADEFEFFFA81" box="[277,359,1324,1347]" captionStart="Figs 23–27" captionStartId="7.[150,187,1038,1057]" captionTargetBox="[251,1259,206,1012]" captionTargetId="figure@7.[221,1366,189,1019]" captionTargetPageId="7" captionText="Figs 23–27. Scobinigaster gen. nov. species. 23–24, 26–27 – S. henryi Kondorosy &amp; Baňař sp. nov., holotype, male; 23 – paramere, ventral view; 24 – paramere, lateral view; 26 – pygophore, posterior view; 27 – paratype, female, spermatheca. 25 – S. paveli Kondorosy &amp; Baňař sp. nov., holotype, male, paramere, ventral view, hairs omitted." figureDoi="http://doi.org/10.5281/zenodo.3879987" httpUri="https://zenodo.org/record/3879987/files/figure.png" pageId="4" pageNumber="347">Fig. 27</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35B6522FFBAFFCAFF5DFAA9FD76F8AD" lastPageId="6" lastPageNumber="349" pageId="4" pageNumber="347" type="diagnosis">
<paragraph id="8BFE36A9FFBAFFC8FF5DFAA9FD18FA10" blockId="4.[151,760,1371,2066]" pageId="4" pageNumber="347">
<emphasis id="B935EABBFFBAFFC8FF5DFAA9FD58FAB0" bold="true" box="[151,704,1371,1394]" pageId="4" pageNumber="347">Differential diagnosis and systematic placement.</emphasis>
<taxonomicName id="4C414D2AFFBAFFC8FD02FAAEFE78FA50" authority="Kondorosy &amp; Banar" authorityName="Kondorosy &amp; Banar" authorityYear="2020" class="Insecta" family="Rhyparochromidae" genus="Scobinigaster" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="347" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B935EABBFFBAFFC8FD02FAAEFE9BFA50" italics="true" pageId="4" pageNumber="347">Scobinigaster</emphasis>
Kondorosy &amp; Baňař
</taxonomicName>
, 
<taxonomicNameLabel id="A20657C0FFBAFFC8FE20FA89FDD2FA50" box="[490,586,1403,1426]" pageId="4" pageNumber="347" rank="genus">gen. nov.</taxonomicNameLabel>
is unique within 
<taxonomicName id="4C414D2AFFBAFFC8FF5DFA69FEFBFA70" authorityName="Amyot &amp; Audinet-Serville" authorityYear="1843" box="[151,355,1435,1458]" class="Insecta" family="Rhyparochromidae" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="347" phylum="Arthropoda" rank="family">Rhyparochromidae</taxonomicName>
by the ventral side of abdomen provided with rasp-like, coarse setiferous granules.
</paragraph>
<paragraph id="8BFE36A9FFBAFFC8FF7FFA29FD6AF950" blockId="4.[151,760,1371,2066]" pageId="4" pageNumber="347">
<taxonomicName id="4C414D2AFFBAFFC8FF7FFA29FEDEFA30" authorityName="Kondorosy &amp; Banar" authorityYear="2020" box="[181,326,1499,1522]" class="Insecta" family="Rhyparochromidae" genus="Scobinigaster" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="B935EABBFFBAFFC8FF7FFA29FEDEFA30" box="[181,326,1499,1522]" italics="true" pageId="4" pageNumber="347">Scobinigaster</emphasis>
</taxonomicName>
is a typical member of the tribe 
<taxonomicName id="4C414D2AFFBAFFC8FD44FA29FD6CFA30" authorityName="Stål" authorityYear="1872" box="[654,756,1499,1522]" class="Insecta" family="Rhyparochromidae" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="347" phylum="Arthropoda" rank="tribe" tribe="Lethaeini">Lethaeini</taxonomicName>
, having two iridescent spots on the base of the head (Western Hemisphere species can have one spot instead, see e.g. O’DƟNNൾĿĿ 1991); a pair of anterolateral trichobothria on the pronotum; and the linear arrangement of trichobothria on sternum V (3 
<superScript id="7C349BE1FFBAFFC8FE8EF98BFEC9F945" attach="left" box="[324,337,1657,1671]" fontSize="6" pageId="4" pageNumber="347">rd</superScript>
trichobothrium always postspiracular).
</paragraph>
<paragraph id="8BFE36A9FFBAFFC8FF7FF969FB3DF881" blockId="4.[151,760,1371,2066]" lastBlockId="4.[828,1437,1292,2083]" pageId="4" pageNumber="347">
The genus belongs to the largest group of genera in the tribe, which has cross-veins on the membrane of the hemelytra (although the majority of the 
<taxonomicName id="4C414D2AFFBAFFC8FDFCF929FD5DF930" authorityName="Kondorosy &amp; Banar" authorityYear="2020" box="[566,709,1755,1778]" class="Insecta" family="Rhyparochromidae" genus="Scobinigaster" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="B935EABBFFBAFFC8FDFCF929FD5DF930" box="[566,709,1755,1778]" italics="true" pageId="4" pageNumber="347">Scobinigaster</emphasis>
</taxonomicName>
specimens are brachypterous), anterior collar on the pronotum and anterolateral trichobothria on the somewhat widened and not very concave lateral margin of the pronotum. This group contains five previously described genera, which are not always easy to separate, and some of them might be paraphyletic (i.e., 
<emphasis id="B935EABBFFBAFFC8FE93F869FE42F870" box="[345,474,1947,1970]" italics="true" pageId="4" pageNumber="347">Neolethaeus</emphasis>
Distant, 1909 and 
<emphasis id="B935EABBFFBAFFC8FD66F869FF7AF810" italics="true" pageId="4" pageNumber="347">Lophoraglius</emphasis>
Wagner, 1961). In the genera 
<emphasis id="B935EABBFFBAFFC8FDFDF849FD02F810" box="[567,666,1979,2002]" italics="true" pageId="4" pageNumber="347">Adauctus</emphasis>
Distant, 1909, 
<emphasis id="B935EABBFFBAFFC8FF10F829FEFFF830" box="[218,359,2011,2034]" italics="true" pageId="4" pageNumber="347">Lophoraglius</emphasis>
and 
<emphasis id="B935EABBFFBAFFC8FE54F829FDC8F830" box="[414,592,2011,2034]" italics="true" pageId="4" pageNumber="347">Porrectolethaeus</emphasis>
Scudder, 1971, the pronotal collar is separated by a strong groove, not only with inserted punctures as in 
<emphasis id="B935EABBFFBAFFC8FBAEFAFEFB7DFAE1" box="[1124,1253,1292,1315]" italics="true" pageId="4" pageNumber="347">Neolethaeus</emphasis>
, 
<emphasis id="B935EABBFFBAFFC8FB3AFAFEFADDFAE1" box="[1264,1349,1292,1315]" italics="true" pageId="4" pageNumber="347">Orbellis</emphasis>
Distant, 1913 and 
<taxonomicName id="4C414D2AFFBAFFC8FC6BFADEFBA8FA81" authorityName="Kondorosy &amp; Banar" authorityYear="2020" box="[929,1072,1324,1347]" class="Insecta" family="Rhyparochromidae" genus="Scobinigaster" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="B935EABBFFBAFFC8FC6BFADEFBA8FA81" box="[929,1072,1324,1347]" italics="true" pageId="4" pageNumber="347">Scobinigaster</emphasis>
</taxonomicName>
. A further common feature of the 3 latter genera is that in many (but not in all) species the males have more or less widened metafemora with several small teeth and 3 stronger or smaller teeth on the posteroventral margin of abdominal sternite VII. Within these genera only 
<taxonomicName id="4C414D2AFFBAFFC8FCF6FA3EFC4AFA21" authorityName="Kondorosy &amp; Banar" authorityYear="2020" box="[828,978,1484,1507]" class="Insecta" family="Rhyparochromidae" genus="Scobinigaster" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="B935EABBFFBAFFC8FCF6FA3EFC4AFA21" box="[828,978,1484,1507]" italics="true" pageId="4" pageNumber="347">Scobinigaster</emphasis>
</taxonomicName>
possesses the characteristic densely and strongly punctate anterior lobe of pronotum. There are some other 
<taxonomicName id="4C414D2AFFBAFFC8FCB1F9FEFC46F9E1" authorityName="Stål" authorityYear="1872" box="[891,990,1548,1571]" class="Insecta" family="Rhyparochromidae" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="347" phylum="Arthropoda" rank="tribe" tribe="Lethaeini">Lethaeini</taxonomicName>
taxa with densely punctate anterior lobe (
<emphasis id="B935EABBFFBAFFC8FC8EF9DEFC58F981" box="[836,960,1580,1603]" italics="true" pageId="4" pageNumber="347">Afromydrus</emphasis>
<bibRefCitation id="EFD04B58FFBAFFC8FC00F9DEFBFFF981" author="SCUDDER G. G. E." box="[970,1127,1580,1603]" pageId="4" pageNumber="347" pagination="205 - 213" refId="ref5776" refString="SCUDDER G. G. E. 1968: The identity of Naphius bug pests in Africa (Hemiptera: Lygaeidae). Bulletin of Entomological Research 58: 205 - 213." type="journal article" year="1968">Scudder, 1968</bibRefCitation>
, 
<emphasis id="B935EABBFFBAFFC8FBBCF9DEFB62F981" box="[1142,1274,1580,1603]" italics="true" pageId="4" pageNumber="347">Aristaenetus</emphasis>
Distant, 1901, 
<emphasis id="B935EABBFFBAFFC8FCF6F9BEFBD9F9A1" box="[828,1089,1612,1635]" italics="true" pageId="4" pageNumber="347">Lophoraglius punctatulus</emphasis>
Linnavuori, 1978, 
<emphasis id="B935EABBFFBAFFC8FACBF9BEFA03F9A1" box="[1281,1435,1612,1635]" italics="true" pageId="4" pageNumber="347">Margolethaeus</emphasis>
<bibRefCitation id="EFD04B58FFBAFFC8FCF6F99EFB02F941" author="ZSALAKOVICS L. &amp; KONDOROSY E." box="[828,1178,1644,1667]" pageId="4" pageNumber="347" pagination="307 - 312" refId="ref6165" refString="ZSALAKOVICS L. &amp; KONDOROSY E. 2014: Margolethaeus, a new Lethaeini genus (Hemiptera: Rhyparochromidae) from the Oriental Region. Acta Zoologica Academiae Scientiarum Hungaricae 60: 307 - 312." type="journal article" year="2014">Zsalakovics &amp; Kondorosy, 2014</bibRefCitation>
, 
<emphasis id="B935EABBFFBAFFC8FB62F99EFAA5F941" box="[1192,1341,1644,1667]" italics="true" pageId="4" pageNumber="347">Noteolethaeus</emphasis>
and 
<emphasis id="B935EABBFFBAFFC8FABCF99FFC48F961" italics="true" pageId="4" pageNumber="347">Porrectolethaeus</emphasis>
), but in these, the punctures are always rather fine and distinct, while in 
<taxonomicName id="4C414D2AFFBAFFC8FB9AF95EFB47F901" authorityName="Kondorosy &amp; Banar" authorityYear="2020" box="[1104,1247,1708,1731]" class="Insecta" family="Rhyparochromidae" genus="Scobinigaster" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="B935EABBFFBAFFC8FB9AF95EFB47F901" box="[1104,1247,1708,1731]" italics="true" pageId="4" pageNumber="347">Scobinigaster</emphasis>
</taxonomicName>
the punctures are coarse and partially confluent. Only 
<emphasis id="B935EABBFFBAFFC8FB70F93EFAACF921" box="[1210,1332,1740,1763]" italics="true" pageId="4" pageNumber="347">Afromydrus</emphasis>
is similar in this feature, but it differs e.g. by wide lateral margins of pronotum without anterolateral trichobothria and lack of veins on the membrane except Cu.
</paragraph>
<paragraph id="8BFE36A9FFBAFFC8FC90F8BEFBC3F821" blockId="4.[828,1437,1292,2083]" pageId="4" pageNumber="347">
Another very specific character is represented by the tiny teeth on the maxillary plates (Figs 9, 10: arrows) and the rasp-like structure formed by conspicuous setiferous tubercles, of the abdominal venter. These features seem to be unique within 
<taxonomicName id="4C414D2AFFBAFFC8FC3BF83EFBCFF821" authorityName="Stål" authorityYear="1872" box="[1009,1111,1996,2019]" class="Insecta" family="Rhyparochromidae" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="347" phylum="Arthropoda" rank="tribe" tribe="Lethaeini">Lethaeini</taxonomicName>
.
</paragraph>
<paragraph id="8BFE36A9FFBAFFC9FC90F81EFD60F801" blockId="4.[828,1437,1292,2083]" lastBlockId="5.[151,760,1804,2084]" lastPageId="5" lastPageNumber="348" pageId="4" pageNumber="347">
In most 
<taxonomicName id="4C414D2AFFBAFFC8FC64F81EFB88F7C1" authorityName="Stål" authorityYear="1872" box="[942,1040,2028,2051]" class="Insecta" family="Rhyparochromidae" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="347" phylum="Arthropoda" rank="tribe" tribe="Lethaeini">Lethaeini</taxonomicName>
genera, some strong and/or long setae can be found on the profemora, but the additional presence of one series of strong stiff setae (similar to the regular setae of the tibiae) on the meso- and metafemora is quite rare. Two parallel rows of such setae on the meso- and metafemora are found only in Afrotropical species of 
<emphasis id="B935EABBFFBBFFC9FD83F89EFD52F841" box="[585,714,1900,1923]" italics="true" pageId="5" pageNumber="348">Neolethaeus</emphasis>
and 
<emphasis id="B935EABBFFBBFFC9FF5DF87EFF74F861" box="[151,236,1932,1955]" italics="true" pageId="5" pageNumber="348">Orbellis</emphasis>
. Based on the characters mentioned above, those two genera seem to be the nearest relatives of 
<taxonomicName id="4C414D2AFFBBFFC9FDA9F85EFD69F801" authorityName="Kondorosy &amp; Banar" authorityYear="2020" box="[611,753,1964,1987]" class="Insecta" family="Rhyparochromidae" genus="Scobinigaster" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="B935EABBFFBBFFC9FDA9F85EFD69F801" box="[611,753,1964,1987]" italics="true" pageId="5" pageNumber="348">Scobinigaster</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="DF3E6621FFBBFFC9FF5DF973FA06F90B" ID-DOI="http://doi.org/10.5281/zenodo.3879985" ID-Zenodo-Dep="3879985" httpUri="https://zenodo.org/record/3879985/files/figure.png" pageId="5" pageNumber="348" startId="5.[151,188,1665,1684]" targetBox="[151,1436,191,1646]" targetPageId="5">
<paragraph id="8BFE36A9FFBBFFC9FF5DF973FA06F90B" blockId="5.[151,1438,1665,1737]" pageId="5" pageNumber="348">
Figs 13–18. 
<taxonomicName id="4C414D2AFFBBFFC9FECAF973FDF8F956" authority="Kondorosy &amp; Banar" authorityName="Kondorosy &amp; Banar" authorityYear="2020" box="[256,608,1665,1684]" class="Insecta" family="Rhyparochromidae" genus="Scobinigaster" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="species" species="henryi" status="sp. nov.">
<emphasis id="B935EABBFFBBFFC9FECAF973FE29F956" box="[256,433,1665,1684]" italics="true" pageId="5" pageNumber="348">Scobinigaster henryi</emphasis>
Kondorosy &amp; Baňař
</taxonomicName>
<taxonomicNameLabel id="A20657C0FFBBFFC9FDAFF973FD31F956" box="[613,681,1665,1684]" pageId="5" pageNumber="348" rank="species">sp. nov.</taxonomicNameLabel>
, scanning electron micrographs. 13–17 – paratype, female: 13 – metathorax with position of peritreme and evaporatium; 14 – detail of peritreme and evaporatorium; 15 – trichobothria on segments III and IV, lateroventral view; 16 – detail of trichobothria on segment IV, lateroventral view; 17 – trichobothria on segment VI, lateroventral view.18 – holotype, male, pygophore, posterodorsal view.
</paragraph>
</caption>
<paragraph id="8BFE36A9FFBBFFC9FF7FF83EFBFEF801" blockId="5.[151,760,1804,2084]" lastBlockId="5.[828,1437,1804,2083]" pageId="5" pageNumber="348">
Only in the continental African 
<emphasis id="B935EABBFFBBFFC9FDD1F83EFD38F821" box="[539,672,1996,2019]" italics="true" pageId="5" pageNumber="348">Neolethaeus</emphasis>
species (
<emphasis id="B935EABBFFBBFFC9FF6AF81FFEA0F7C1" box="[160,312,2028,2052]" italics="true" pageId="5" pageNumber="348">N. aethiopicus</emphasis>
Hesse, 1925, 
<emphasis id="B935EABBFFBBFFC9FE1BF81FFDCFF7C6" box="[465,599,2029,2052]" italics="true" pageId="5" pageNumber="348">N. giganteus</emphasis>
Scudder, 1963 and 
<emphasis id="B935EABBFFBBFFC9FF0EF7FFFED9F7E1" box="[196,321,2060,2084]" italics="true" pageId="5" pageNumber="348">N. ulugurus</emphasis>
Scudder, 1962) is the last tergite (VII) of the males so decumbent that it is closed together with the last sternite, and the genital segments are not at all visible. When comparing the parameres of 
<taxonomicName id="4C414D2AFFBBFFC9FB7AF8BEFAA7F8A1" authorityName="Kondorosy &amp; Banar" authorityYear="2020" box="[1200,1343,1868,1891]" class="Insecta" family="Rhyparochromidae" genus="Scobinigaster" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="B935EABBFFBBFFC9FB7AF8BEFAA7F8A1" box="[1200,1343,1868,1891]" italics="true" pageId="5" pageNumber="348">Scobinigaster</emphasis>
</taxonomicName>
with the figures of O’DƟNNൾĿĿ (1991), the most similar paramere is that of 
<emphasis id="B935EABBFFBBFFC9FC43F87EFB13F861" box="[905,1163,1932,1955]" italics="true" pageId="5" pageNumber="348">Neolethaeus aethiopicus</emphasis>
, though there are enough differences to separate them.
</paragraph>
<paragraph id="8BFE36A9FFBBFFCAFC90F83EFD76F8AD" blockId="5.[828,1437,1804,2083]" lastBlockId="6.[151,760,1656,1903]" lastPageId="6" lastPageNumber="349" pageId="5" pageNumber="348">
Brachyptery is not a very common character in 
<taxonomicName id="4C414D2AFFBBFFC9FAFFF83EFA01F821" authorityName="Stål" authorityYear="1872" box="[1333,1433,1996,2019]" class="Insecta" family="Rhyparochromidae" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="tribe" tribe="Lethaeini">Lethaeini</taxonomicName>
, except the taxa from the 
<collectingRegion id="4985F84BFFBBFFC9FBF1F81EFB21F7C1" box="[1083,1209,2028,2051]" country="South Africa" name="Western Cape" pageId="5" pageNumber="348">Cape region</collectingRegion>
of Africa (see SĿAŦൾR 1977), some Australian and Neotropical genera, and almost always it is facultative, as far as macropterous specimens can be also found (e.g. in 
<emphasis id="B935EABBFFB8FFCAFE75F968FDDBF973" box="[447,579,1690,1713]" italics="true" pageId="6" pageNumber="349">Camptocera</emphasis>
Jakovlev, 1877, 
<emphasis id="B935EABBFFB8FFCAFF5DF94AFF74F90D" box="[151,236,1720,1743]" italics="true" pageId="6" pageNumber="349">Diniella</emphasis>
and 
<emphasis id="B935EABBFFB8FFCAFED4F94AFE2BF90D" box="[286,435,1720,1743]" italics="true" pageId="6" pageNumber="349">Noteolethaeus</emphasis>
). Interestingly, the only known specimen of the ‘oldest’ but long forgotten 
<taxonomicName id="4C414D2AFFB8FFCAFDA8F92AFD5CF92D" authorityName="Stål" authorityYear="1872" box="[610,708,1752,1775]" class="Insecta" family="Rhyparochromidae" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="349" phylum="Arthropoda" rank="tribe" tribe="Lethaeini">Lethaeini</taxonomicName>
species, 
<emphasis id="B935EABBFFB8FFCAFF07F90AFEC4F8D3" box="[205,348,1784,1809]" italics="true" pageId="6" pageNumber="349">Lethaeus ater</emphasis>
(Thunberg, 1822) from 
<collectingCountry id="F3567639FFB8FFCAFD91F90AFD78F8CD" box="[603,736,1784,1807]" name="South Africa" pageId="6" pageNumber="349">South Africa</collectingCountry>
is also brachypterous (see KƟNĐƟRƟඌY et al. 2014). Now, with 
<taxonomicName id="4C414D2AFFB8FFCAFF5DF8CAFEBAF88D" authorityName="Kondorosy &amp; Banar" authorityYear="2020" box="[151,290,1848,1871]" class="Insecta" family="Rhyparochromidae" genus="Scobinigaster" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="B935EABBFFB8FFCAFF5DF8CAFEBAF88D" box="[151,290,1848,1871]" italics="true" pageId="6" pageNumber="349">Scobinigaster</emphasis>
</taxonomicName>
and 
<emphasis id="B935EABBFFB8FFCAFE9AF8CAFE79F88D" box="[336,481,1848,1871]" italics="true" pageId="6" pageNumber="349">Noteolethaeus</emphasis>
(see ZǍMƂó et al. 2019), we have discovered brachyptery in Madagascan taxa as well.
</paragraph>
</subSubSection>
<subSubSection id="C35B6522FFB8FFCAFF5DF87AFB2AF96B" pageId="6" pageNumber="349" type="etymology">
<paragraph id="8BFE36A9FFB8FFCAFF5DF87AFD07F7DD" blockId="6.[151,760,1928,2079]" pageId="6" pageNumber="349">
<emphasis id="B935EABBFFB8FFCAFF5DF87AFE80F85D" bold="true" box="[151,280,1928,1951]" pageId="6" pageNumber="349">Etymology.</emphasis>
The first part of the name of the genus refers to ‘ 
<emphasis id="B935EABBFFB8FFCAFF76F85AFE97F87D" box="[188,271,1960,1983]" italics="true" pageId="6" pageNumber="349">scobina</emphasis>
’, which means rasp in Latin; the latter part is ‘ 
<emphasis id="B935EABBFFB8FFCAFF6AF83BFF7BF822" box="[160,227,1993,2016]" italics="true" pageId="6" pageNumber="349">gaster</emphasis>
’ meaning abdomen in Latinized Greek (originally γᾰστήρ, meaning stomach
<emphasis id="B935EABBFFB8FFCAFE60F81AFE2BF83D" bold="true" box="[426,435,2024,2047]" pageId="6" pageNumber="349">)</emphasis>
, indicating the coarsely granulate surface of the abdomen. Gender is feminine.
</paragraph>
<paragraph id="8BFE36A9FFB8FFCAFC9FF985FB2AF96B" blockId="6.[853,1418,1655,1705]" pageId="6" pageNumber="349">
<emphasis id="B935EABBFFB8FFCAFC9FF985FA12F94D" bold="true" box="[853,1418,1655,1679]" pageId="6" pageNumber="349">
<taxonomicName id="4C414D2AFFB8FFCAFC9FF985FAB1F94D" authority="Kondorosy &amp; Banar" authorityName="Kondorosy &amp; Banar" authorityYear="2020" box="[853,1321,1655,1679]" class="Insecta" family="Rhyparochromidae" genus="Scobinigaster" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="349" phylum="Arthropoda" rank="species" species="henryi" status="sp. nov.">
<emphasis id="B935EABBFFB8FFCAFC9FF985FBA0F94D" bold="true" box="[853,1080,1655,1679]" italics="true" pageId="6" pageNumber="349">Scobinigaster henryi</emphasis>
Kondorosy &amp; Baňař
</taxonomicName>
, 
<taxonomicNameLabel id="A20657C0FFB8FFCAFAFEF98AFA12F94D" box="[1332,1418,1656,1679]" pageId="6" pageNumber="349" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(Figs 1–3, 5–18)
</paragraph>
</subSubSection>
<subSubSection id="C35B6522FFB8FFCBFCF6F932FF77FA82" lastPageId="7" lastPageNumber="350" pageId="6" pageNumber="349" type="materials_examined">
<paragraph id="8BFE36A9FFB8FFCAFCF6F932FC35F8CA" blockId="6.[828,1438,1728,2067]" pageId="6" pageNumber="349">
<emphasis id="B935EABBFFB8FFCAFCF6F932FC21F911" bold="true" box="[828,953,1728,1747]" pageId="6" pageNumber="349">
<typeStatus id="54FA880BFFB8FFCAFCF6F932FCF1F911" box="[828,873,1728,1747]" pageId="6" pageNumber="349">Type</typeStatus>
locality.
</emphasis>
Southwestern 
<collectingCountry id="F3567639FFB8FFCAFB8AF932FB32F911" box="[1088,1194,1728,1747]" name="Madagascar" pageId="6" pageNumber="349">Madagascar</collectingCountry>
, Atsimo-Andrefana region, Zombitse-Vohibasia National Park, Zombitse forest, 
<geoCoordinate id="EE75506EFFB8FFCAFAECF929FA04F92C" box="[1318,1436,1755,1774]" degrees="22" direction="south" minutes="53.171" orientation="latitude" pageId="6" pageNumber="349" precision="1" value="-22.886183">22°53.171′ S</geoCoordinate>
<geoCoordinate id="EE75506EFFB8FFCAFCF6F907FC30F8CA" box="[828,936,1781,1800]" degrees="44" direction="east" minutes="41.489" orientation="longitude" pageId="6" pageNumber="349" precision="1" value="44.691483">44°41.489′E</geoCoordinate>
.
</paragraph>
<paragraph id="8BFE36A9FFB8FFCBFCF6F8E2FF77FA82" blockId="6.[828,1438,1728,2067]" lastBlockId="7.[151,760,1188,1344]" lastPageId="7" lastPageNumber="350" pageId="6" pageNumber="349">
<emphasis id="B935EABBFFB8FFCAFCF6F8E2FC58F8E1" bold="true" box="[828,960,1808,1827]" pageId="6" pageNumber="349">Type material.</emphasis>
HƟĿƟŦYPൾ: (
<collectionCode id="ED50AE6CFFB8FFCAFB8CF8E2FB15F8E1" box="[1094,1165,1808,1827]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="6" pageNumber="349">MMBC</collectionCode>
), ‘SW 
<collectingCountry id="F3567639FFB8FFCAFB00F8E2FAF3F8E1" box="[1226,1387,1808,1827]" name="Madagascar" pageId="6" pageNumber="349">MADAGASCAR</collectingCountry>
, 
<collectingRegion id="4985F84BFFB8FFCAFAB9F8E2FCCEF8FC" country="Madagascar" name="Toliara" pageId="6" pageNumber="349">Toliara</collectingRegion>
pr. / Zombitse-Vohibasia N.P. / Zombitse forest, 
<quantity id="4CB99B4CFFB8FFCAFAC8F8D9FAAAF8FC" box="[1282,1330,1835,1854]" metricMagnitude="2" metricUnit="m" metricValue="8.24" pageId="6" pageNumber="349" unit="m" value="824.0">824m</quantity>
main / entr. 22°53’171”S 44°41’489”E / 
<date id="FFFF1069FFB8FFCAFBE4F8B7FB05F89A" box="[1070,1181,1861,1880]" pageId="6" pageNumber="349" value="2014-01-22" valueMax="2014-01-25" valueMin="2014-01-22">22-25.i.2014</date>
, M.Trýzna leg. [printed, white label] // 
<typeStatus id="54FA880BFFB8FFCAFC48F892FB99F8B1" box="[898,1025,1888,1907]" pageId="6" pageNumber="349" type="holotype">HOLOTYPUS</typeStatus>
[printed] / 
<taxonomicName id="4C414D2AFFB8FFCAFBAAF892FB4CF8B1" authorityName="Kondorosy &amp; Banar" authorityYear="2020" box="[1120,1236,1888,1907]" class="Insecta" family="Rhyparochromidae" genus="Scobinigaster" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="351" phylum="Arthropoda" rank="genus">Scobinigaster</taxonomicName>
/ henryi 
<taxonomicNameLabel id="A20657C0FFB8FFCAFAD5F892FAF9F8B1" box="[1311,1377,1888,1907]" pageId="6" pageNumber="349" rank="species">sp. nov.</taxonomicNameLabel>
[handwritten] / det. Kondorosy &amp; Baňař [printed, red label]’. PARAŦYPൾඌ: 1 (
<collectionCode id="ED50AE6CFFB8FFCAFC88F867FC1BF86A" box="[834,899,1941,1960]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="6" pageNumber="349">NMPC</collectionCode>
), the same data as 
<typeStatus id="54FA880BFFB8FFCAFBEBF867FBF6F86A" box="[1057,1134,1941,1960]" pageId="6" pageNumber="349" type="holotype">holotype</typeStatus>
; 
<specimenCount id="9D47FD20FFB8FFCAFBBDF867FB0FF86B" box="[1143,1175,1938,1961]" pageId="6" pageNumber="349" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED50AE6CFFB8FFCAFB6BF867FB70F86A" box="[1185,1256,1941,1960]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="6" pageNumber="349">MMBC</collectionCode>
), ‘SW MADAGAS- 
<collectingCountry id="F3567639FFB8FFCAFCF6F842FCF2F801" box="[828,874,1968,1987]" name="Central African Republic" pageId="6" pageNumber="349">CAR</collectingCountry>
/ Isalo NP, Zahavola for. / cca 
<quantity id="4CB99B4CFFB8FFCAFBB9F842FB3EF801" box="[1139,1190,1968,1987]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="6" pageNumber="349" unit="m" value="850.0">850m</quantity>
; 
<date id="FFFF1069FFB8FFCAFB65F842FB67F801" box="[1199,1279,1968,1987]" pageId="6" pageNumber="349" value="2013-01-21">21.i.2013</date>
/ M. Trýzna leg.’; 
<specimenCount id="9D47FD20FFB8FFCAFCF6F839FCC5F81C" box="[828,861,1991,2014]" pageId="6" pageNumber="349" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED50AE6CFFB8FFCAFCA3F839FC28F81C" box="[873,944,1995,2014]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="6" pageNumber="349">MMBC</collectionCode>
), ‘ISL/ 
<date id="FFFF1069FFB8FFCAFC21F839FBA3F81C" box="[1003,1083,1995,2014]" pageId="6" pageNumber="349" value="2013-01">Jan 2013</date>
/19 
<collectingCountry id="F3567639FFB8FFCAFB96F839FB65F81C" box="[1116,1277,1995,2014]" name="Madagascar" pageId="6" pageNumber="349">MADAGASCAR</collectingCountry>
, Isalo NP; Piscine Naturelle; 
<quantity id="4CB99B4CFFB8FFCAFC50F817FC4BF83A" box="[922,979,2021,2040]" metricMagnitude="2" metricUnit="m" metricValue="8.52" pageId="6" pageNumber="349" unit="m" value="852.0">852 m</quantity>
/ 
<geoCoordinate id="EE75506EFFB8FFCAFC23F817FBF0F83A" box="[1001,1128,2021,2040]" degrees="22" direction="south" minutes="33" orientation="latitude" pageId="6" pageNumber="349" precision="1" seconds="43.5" value="-22.562082">S 22°33’43.5”</geoCoordinate>
<geoCoordinate id="EE75506EFFB8FFCAFBBAF817FB6AF83A" box="[1136,1266,2021,2040]" degrees="45" direction="east" minutes="23" orientation="longitude" pageId="6" pageNumber="349" precision="1" seconds="00.4" value="45.383446">E 45°23’00.4”</geoCoordinate>
; 
<date id="FFFF1069FFB8FFCAFB37F817FAD7F83A" box="[1277,1359,2021,2040]" pageId="6" pageNumber="349" value="2013-01-21">21.i.2013</date>
/ sifting litter, Winkler app. extr.; L.S. / Rahanitriniaina &amp; E. M. Rabotoson lgt.’; 
<specimenCount id="9D47FD20FFB9FFCBFF5DFB5AFF2EFB79" box="[151,182,1188,1211]" pageId="7" pageNumber="350" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED50AE6CFFB9FFCBFF75FB5AFE9CFB79" box="[191,260,1192,1211]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="7" pageNumber="350">MMBC</collectionCode>
) 1 (
<collectionCode id="ED50AE6CFFB9FFCBFEFDFB5AFEE5FB79" box="[311,381,1192,1211]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="7" pageNumber="350">HNHM</collectionCode>
), ‘ISL/ 
<date id="FFFF1069FFB9FFCBFE7FFB5AFD99FB79" box="[437,513,1192,1211]" pageId="7" pageNumber="350" value="2013-01">Jan 2013</date>
/09 
<collectingCountry id="F3567639FFB9FFCBFDEBFB5AFD58FB79" box="[545,704,1192,1211]" name="Madagascar" pageId="7" pageNumber="350">MADAGASCAR</collectingCountry>
/ Isalo N.P., Analalava forest; 
<quantity id="4CB99B4CFFB9FFCBFE97FB30FE16FB17" box="[349,398,1218,1237]" metricMagnitude="2" metricUnit="m" metricValue="7.35" pageId="7" pageNumber="350" unit="m" value="735.0">735m</quantity>
/ 
<geoCoordinate id="EE75506EFFB9FFCBFE55FB30FD81FB17" box="[415,537,1218,1237]" degrees="22" direction="south" minutes="34" orientation="latitude" pageId="7" pageNumber="350" precision="1" seconds="45.5" value="-22.579306">S 22°34’45.5”</geoCoordinate>
<geoCoordinate id="EE75506EFFB9FFCBFDEAFB30FD0DFB17" box="[544,661,1218,1237]" degrees="45" direction="east" minutes="08" orientation="longitude" pageId="7" pageNumber="350" precision="1" seconds="22.5" value="45.139584">E45°08’22.5”</geoCoordinate>
<date id="FFFF1069FFB9FFCBFD51FB30FD74FB17" box="[667,748,1218,1237]" pageId="7" pageNumber="350" value="2013-01-19">19.i.2013</date>
/ sifting litter, Winkler app.extr.; L.S. / Rahanitriniaina &amp; E.M. Rabotoson lgt.’ Each 
<typeStatus id="54FA880BFFB9FFCBFF24FB0AFEA0FAC9" box="[238,312,1272,1291]" pageId="7" pageNumber="350" type="paratype">paratype</typeStatus>
bears following red label: ‘ 
<typeStatus id="54FA880BFFB9FFCBFDEAFB0AFD04FAC9" box="[544,668,1272,1291]" pageId="7" pageNumber="350" type="paratype">PARATYPUS</typeStatus>
[printed] / 
<taxonomicName id="4C414D2AFFB9FFCBFF5DFAE0FE93FAE7" authorityName="Kondorosy &amp; Banar" authorityYear="2020" box="[151,267,1298,1317]" class="Insecta" family="Rhyparochromidae" genus="Scobinigaster" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="351" phylum="Arthropoda" rank="genus">Scobinigaster</taxonomicName>
/ henryi 
<taxonomicNameLabel id="A20657C0FFB9FFCBFE92FAE0FE04FAE7" box="[344,412,1298,1317]" pageId="7" pageNumber="350" rank="species">sp. nov.</taxonomicNameLabel>
[handwritten] / det. Kondorosy &amp; Baňař [printed]’.
</paragraph>
</subSubSection>
<caption id="DF3E6621FFB9FFCBFF5CFBFCFE43FB95" ID-DOI="http://doi.org/10.5281/zenodo.3879987" ID-Zenodo-Dep="3879987" httpUri="https://zenodo.org/record/3879987/files/figure.png" pageId="7" pageNumber="350" startId="7.[150,187,1038,1057]" targetBox="[251,1259,206,1012]" targetPageId="7">
<paragraph id="8BFE36A9FFB9FFCBFF5CFBFCFE43FB95" blockId="7.[150,1436,1038,1111]" pageId="7" pageNumber="350">
Figs 23–27. 
<taxonomicName id="4C414D2AFFB9FFCBFECBFBFCFEEEFBE3" authority="Kondorosy &amp; Zámbó &amp; Kovács &amp; Baňař &amp; Km, 2020" authorityName="Kondorosy &amp; Zámbó &amp; Kovács &amp; Baňař &amp; Km" authorityYear="2020" box="[257,374,1038,1057]" class="Insecta" family="Rhyparochromidae" genus="Scobinigaster" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="350" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B935EABBFFB9FFCBFECBFBFCFEEEFBE3" box="[257,374,1038,1057]" italics="true" pageId="7" pageNumber="350">Scobinigaster</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20657C0FFB9FFCBFEB1FBFCFE52FBE3" box="[379,458,1038,1057]" pageId="7" pageNumber="350" rank="genus">gen. nov.</taxonomicNameLabel>
species. 23–24, 26–27 – 
<taxonomicName id="4C414D2AFFB9FFCBFD6BFBFDFC38FBE3" authority="Kondorosy &amp; Banar" authorityName="Kondorosy &amp; Banar" authorityYear="2020" box="[673,928,1038,1058]" class="Insecta" family="Rhyparochromidae" genus="Scobinigaster" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="350" phylum="Arthropoda" rank="species" species="henryi" status="sp. nov.">
<emphasis id="B935EABBFFB9FFCBFD6BFBFDFD2AFBE0" box="[673,690,1039,1058]" italics="true" pageId="7" pageNumber="350">S.</emphasis>
<emphasis id="B935EABBFFB9FFCBFD7DFBFCFD76FBE3" box="[695,750,1038,1057]" italics="true" pageId="7" pageNumber="350">henryi</emphasis>
Kondorosy &amp; Baňař
</taxonomicName>
<taxonomicNameLabel id="A20657C0FFB9FFCBFC6FFBFCFC72FBE3" box="[933,1002,1038,1057]" pageId="7" pageNumber="350" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, male; 23 – paramere, ventral view; 24 – paramere, lateral view; 26 – pygophore, posterior view; 27 – paratype, female, spermatheca. 25 – 
<taxonomicName id="4C414D2AFFB9FFCBFC05FBD8FB50FBFE" authority="Kondorosy &amp; Banar" authorityName="Kondorosy &amp; Banar" authorityYear="2020" box="[975,1224,1065,1085]" class="Insecta" family="Rhyparochromidae" genus="Scobinigaster" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="350" phylum="Arthropoda" rank="species" species="paveli" status="sp. nov.">
<emphasis id="B935EABBFFB9FFCBFC05FBD8FB80FBFE" box="[975,1048,1065,1085]" italics="true" pageId="7" pageNumber="350">S. paveli</emphasis>
Kondorosy &amp; Baňař
</taxonomicName>
<taxonomicNameLabel id="A20657C0FFB9FFCBFB07FBDBFA89FBFE" box="[1229,1297,1065,1084]" pageId="7" pageNumber="350" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, male, paramere, ventral view, hairs omitted.
</paragraph>
</caption>
<subSubSection id="C35B6522FFB9FFCBFF5DFAAAFBF5F959" pageId="7" pageNumber="350" type="description">
<paragraph id="8BFE36A9FFB9FFCBFF5DFAAAFE93F96D" blockId="7.[151,760,1368,2063]" pageId="7" pageNumber="350">
<emphasis id="B935EABBFFB9FFCBFF5DFAAAFEBAFAAD" bold="true" box="[151,290,1368,1391]" pageId="7" pageNumber="350">Description.</emphasis>
<emphasis id="B935EABBFFB9FFCBFEE2FAAAFEECFAAD" box="[296,372,1368,1391]" italics="true" pageId="7" pageNumber="350">Colour</emphasis>
brown (Figs 1–4); head and anterior lobe of pronotum fuscous; antennae yellowish, apex of pedicel, basal half of basiflagellum and distiflagellum brownish, apical half of basiflagellum white; labium yellow; anterior margin of pronotal collar except middle and lateral margin yellow; posterior lobe of pronotum and hemelytra yellow with irregular dark spots of variable extension; scutellum mostly dark with narrow pale midline and submedial stripes; legs pale, coxae, trochanters, very base of femora and strong setae of middle and hind legs often dark.
</paragraph>
<paragraph id="8BFE36A9FFB9FFCBFF7FF94BFE6EF86D" blockId="7.[151,760,1368,2063]" pageId="7" pageNumber="350">
<emphasis id="B935EABBFFB9FFCBFF7FF94BFE85F912" box="[181,285,1721,1744]" italics="true" pageId="7" pageNumber="350">Structure.</emphasis>
Body oval, its entire surface densely punctate, fully covered with fine, very short, decumbent pilosity (Figs 6, 9, 11), hardly visible on hemelytra, while abdomen with longer but also decumbent fine setae (Fig. 8). Always brachypterous, clavus and corium not separated (Fig. 5), apical margin of corium convex, with arched membranal margin being slightly wider than tibiae, leaving tergite VI in greater part and VII fully free.
</paragraph>
<paragraph id="8BFE36A9FFB9FFCBFF7FF84AFE48F7CD" blockId="7.[151,760,1368,2063]" pageId="7" pageNumber="350">
Male genitalia: cup-like sclerite with an acute tooth medially, much longer than blunt lateral apices. Paramere (
<figureCitation id="137A2A2CFFB9FFCBFF55F80AFE81F7CD" box="[159,281,2040,2063]" captionStart="Figs 23–27" captionStartId="7.[150,187,1038,1057]" captionTargetBox="[251,1259,206,1012]" captionTargetId="figure@7.[221,1366,189,1019]" captionTargetPageId="7" captionText="Figs 23–27. Scobinigaster gen. nov. species. 23–24, 26–27 – S. henryi Kondorosy &amp; Baňař sp. nov., holotype, male; 23 – paramere, ventral view; 24 – paramere, lateral view; 26 – pygophore, posterior view; 27 – paratype, female, spermatheca. 25 – S. paveli Kondorosy &amp; Baňař sp. nov., holotype, male, paramere, ventral view, hairs omitted." figureDoi="http://doi.org/10.5281/zenodo.3879987" httpUri="https://zenodo.org/record/3879987/files/figure.png" pageId="7" pageNumber="350">Figs 23, 24</figureCitation>
) with long setae.
</paragraph>
<paragraph id="8BFE36A9FFB9FFCBFC90FB57FBF5F959" blockId="7.[828,1437,1188,2075]" pageId="7" pageNumber="350">
<emphasis id="B935EABBFFB9FFCBFC90FB57FC6EFB7E" box="[858,1014,1189,1212]" italics="true" pageId="7" pageNumber="350">Measurements</emphasis>
(all in mm, 
<specimenCount id="9D47FD20FFB9FFCBFB42FB56FB47FB79" box="[1160,1247,1188,1211]" pageId="7" pageNumber="350" type="male">3 males</specimenCount>
[
<typeStatus id="54FA880BFFB9FFCBFB25FB56FAC8FB79" box="[1263,1360,1188,1211]" pageId="7" pageNumber="350" type="holotype">holotype</typeStatus>
in parentheses], 
<specimenCount id="9D47FD20FFB9FFCBFC7CFB36FB86FB19" box="[950,1054,1220,1243]" pageId="7" pageNumber="350" type="female">3 females</specimenCount>
[basi- and distiflagellum on largest female absent]): Total body length: 4.56–4.86 (4.83), 4.27–5.39; head: length 0.75–0.78 (0.85), 0.72–0.85, width 0.94–0.98 (0.95), 0.85–1.05, interocular space 0.56–0.59 (0.58), 0.50–0.64; length of eye 0.27–0.29 (0.29), 0.28– 0.32, length of antenniferous tubercle 0.15–0.17 (0.17), 0.16–0.19; length of antennomeres: I 0.61–0.74 (0.67), 0.54–0.69, II 0.92–1.09 (0.96), 0.75–1.04, III 0.69–0.74 (0.67), 0.56–0.75, IV 0.72–0.76 (0.69), 0.62–0.78; length of labiomeres: I 0.72–0.78 (0.77), 0.82–0.90, II 0.71–0.76 (0.73), 0.81–0.88, III 0.57–0.64 (0.61), 0.68–0.81, IV 0.44–0.48 (0.42), 0.45–0.46; pronotum: length 0.92–1.02 (0.97), 0.86–1.02, width 1.36–1.45 (1.42), 1.19–1.49; scutellum: length 0.88–0.89 (0.83), 0.75–0.97, width 0.79–0.81 (0.78), 0.69–0.86.
</paragraph>
</subSubSection>
<subSubSection id="C35B6522FFB9FFCBFCF6F956FC7AF919" pageId="7" pageNumber="350" type="diagnosis">
<paragraph id="8BFE36A9FFB9FFCBFCF6F956FC7AF919" blockId="7.[828,1437,1188,2075]" pageId="7" pageNumber="350">
<emphasis id="B935EABBFFB9FFCBFCF6F956FBDCF979" bold="true" box="[828,1092,1700,1723]" pageId="7" pageNumber="350">Differential diagnosis.</emphasis>
See the differential diagnosis under 
<taxonomicName id="4C414D2AFFB9FFCBFCB5F937FC43F919" authorityName="Kondorosy &amp; Banar" authorityYear="2020" box="[895,987,1732,1756]" class="Insecta" family="Rhyparochromidae" genus="Scobinigaster" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="350" phylum="Arthropoda" rank="species" species="paveli">
<emphasis id="B935EABBFFB9FFCBFCB5F937FC43F919" box="[895,987,1732,1756]" italics="true" pageId="7" pageNumber="350">S. paveli</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C35B6522FFB9FFCBFCF6F916FB66F839" pageId="7" pageNumber="350" type="etymology">
<paragraph id="8BFE36A9FFB9FFCBFCF6F916FA82F8B9" blockId="7.[828,1437,1188,2075]" pageId="7" pageNumber="350">
<emphasis id="B935EABBFFB9FFCBFCF6F916FC24F939" bold="true" box="[828,956,1764,1787]" pageId="7" pageNumber="350">Etymology.</emphasis>
We dedicate this species to Thomas Henry, the excellent specialist in many groups of 
<taxonomicName id="4C414D2AFFB9FFCBFADDF8F6FA00F8D9" box="[1303,1432,1796,1819]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heteroptera" pageId="7" pageNumber="350" phylum="Arthropoda" rank="order">Heteroptera</taxonomicName>
, on the occasion of his 70 
<superScript id="7C349BE1FFB9FFCBFB9FF8D0FBF9F8F2" attach="left" box="[1109,1121,1826,1840]" fontSize="6" pageId="7" pageNumber="350">th</superScript>
birthday (for biography and bibliography see WIJൾൾĿൾR 2018). He helped us always very kindly when we asked him for a favour.
</paragraph>
<paragraph id="8BFE36A9FFB9FFCBFCF6F876FB66F839" blockId="7.[828,1437,1188,2075]" pageId="7" pageNumber="350">
<emphasis id="B935EABBFFB9FFCBFCF6F876FBF8F859" bold="true" box="[828,1120,1924,1947]" pageId="7" pageNumber="350">Collecting circumstances.</emphasis>
Three specimens were sifted from forest leaf litter of dry seasonal forest of low canopy height (Isalo NP, 
<figureCitation id="137A2A2CFFB9FFCBFC38F836FBA7F819" box="[1010,1087,1988,2011]" captionStart="Figs 28–29. 28" captionStartId="8.[151,188,2038,2057]" captionTargetBox="[311,1276,942,2021]" captionTargetId="figure@8.[311,1276,779,2021]" captionTargetPageId="8" captionText="Figs 28–29. 28 – typical habitat in Isalo NP, where part of the type series of Scobinigaster henryi Kondorosy &amp; Baňař sp. nov. was collected; 29 – habitat in Mahafaly plateau, type locality of S. paveli Kondorosy &amp; Baňař sp. nov. Photo: Miloš Trýzna." figureDoi="http://doi.org/10.5281/zenodo.3879991" httpUri="https://zenodo.org/record/3879991/files/figure.png" pageId="7" pageNumber="350">Fig. 28</figureCitation>
), three specimens were collected by other, non-specified collecting method.
</paragraph>
</subSubSection>
<subSubSection id="C35B6522FFB9FFC4FCF6F7F6FD81FF2D" lastPageId="8" lastPageNumber="351" pageId="7" pageNumber="350" type="distribution">
<paragraph id="8BFE36A9FFB9FFCBFCF6F7F6FB42F7D9" blockId="7.[828,1437,1188,2075]" box="[828,1242,2052,2075]" pageId="7" pageNumber="350">
<emphasis id="B935EABBFFB9FFCBFCF6F7F6FC56F7D9" bold="true" box="[828,974,2052,2075]" pageId="7" pageNumber="350">Distribution.</emphasis>
South-west 
<collectingCountry id="F3567639FFB9FFCBFB99F7F6FB4EF7D9" box="[1107,1238,2052,2075]" name="Madagascar" pageId="7" pageNumber="350">Madagascar</collectingCountry>
.
</paragraph>
<paragraph id="8BFE36A9FFB6FFC4FF7AFF4FFD81FF2D" blockId="8.[151,759,189,307]" pageId="8" pageNumber="351">
<emphasis id="B935EABBFFB6FFC4FF7AFF4FFD47FF17" bold="true" box="[176,735,189,213]" pageId="8" pageNumber="351">
<taxonomicName id="4C414D2AFFB6FFC4FF7AFF4FFDE6FF17" authority="Kondorosy &amp; Banar" authorityName="Kondorosy &amp; Banar" authorityYear="2020" box="[176,638,189,213]" class="Insecta" family="Rhyparochromidae" genus="Scobinigaster" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="351" phylum="Arthropoda" rank="species" species="paveli" status="sp. nov.">
<emphasis id="B935EABBFFB6FFC4FF7AFF4FFE15FF17" bold="true" box="[176,397,189,213]" italics="true" pageId="8" pageNumber="351">Scobinigaster paveli</emphasis>
Kondorosy &amp; Baňař
</taxonomicName>
, 
<taxonomicNameLabel id="A20657C0FFB6FFC4FD40FF4CFD47FF17" box="[650,735,190,213]" pageId="8" pageNumber="351" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(Figs 4, 19–22, 25)
</paragraph>
</subSubSection>
<subSubSection id="C35B6522FFB6FFC4FF5DFEF4FDBEFE6F" pageId="8" pageNumber="351" type="materials_examined">
<paragraph id="8BFE36A9FFB6FFC4FF5DFEF4FDC3FEF1" blockId="8.[151,759,189,307]" pageId="8" pageNumber="351">
<emphasis id="B935EABBFFB6FFC4FF5DFEF4FE94FEDB" bold="true" box="[151,268,262,281]" pageId="8" pageNumber="351">
<typeStatus id="54FA880BFFB6FFC4FF5DFEF4FF5BFEDB" box="[151,195,262,281]" pageId="8" pageNumber="351">Type</typeStatus>
locality.
</emphasis>
Southwestern 
<collectingCountry id="F3567639FFB6FFC4FE4FFEF4FE74FEDB" box="[389,492,262,281]" name="Madagascar" pageId="8" pageNumber="351">Madagascar</collectingCountry>
, Mahafaly Plateau,Antanambao near Bezaha, approximately 
<geoCoordinate id="EE75506EFFB6FFC4FE41FED2FE77FEF1" box="[395,495,288,307]" degrees="23" direction="south" minutes="29" orientation="latitude" pageId="8" pageNumber="351" precision="15" seconds="00" value="-23.483334">23°29′00″S</geoCoordinate>
, 
<geoCoordinate id="EE75506EFFB6FFC4FE32FED2FDC3FEF1" box="[504,603,288,307]" degrees="44" direction="east" minutes="28" orientation="longitude" pageId="8" pageNumber="351" precision="15" seconds="40" value="44.47778">44°28′40″E</geoCoordinate>
</paragraph>
<paragraph id="8BFE36A9FFB6FFC4FF5DFEB8FDBEFE6F" blockId="8.[151,760,330,429]" pageId="8" pageNumber="351">
<emphasis id="B935EABBFFB6FFC4FF5DFEB8FE8DFE9F" bold="true" box="[151,277,330,349]" pageId="8" pageNumber="351">Type material.</emphasis>
HƟĿƟŦYPൾ: (
<collectionCode id="ED50AE6CFFB6FFC4FE5BFEB8FE4EFE9F" box="[401,470,330,349]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="8" pageNumber="351">MMBC</collectionCode>
),‘S 
<collectingCountry id="F3567639FFB6FFC4FE33FEB8FD0CFE9F" box="[505,660,330,349]" name="Madagascar" pageId="8" pageNumber="351">MADAGASCAR</collectingCountry>
<date id="FFFF1069FFB6FFC4FD52FEB8FD6CFE9F" box="[664,756,330,349]" pageId="8" pageNumber="351" value="2013-01-14">2013 /14.i.</date>
; Mahafaly Plateau, / Antanambao near Bezaha / vill., M.Trýzna leg.’ // ‘ 
<typeStatus id="54FA880BFFB6FFC4FF50FE8DFE85FE50" box="[154,285,383,402]" pageId="8" pageNumber="351" type="holotype">HOLOTYPUS</typeStatus>
[printed] / 
<taxonomicName id="4C414D2AFFB6FFC4FEB4FE8DFE6AFE50" authorityName="Kondorosy &amp; Zámbó &amp; Kovács &amp; Baňař &amp; Km" authorityYear="2020" box="[382,498,383,402]" class="Insecta" family="Rhyparochromidae" genus="Scobinigaster" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="351" phylum="Arthropoda" rank="genus">Scobinigaster</taxonomicName>
/ paveli 
<taxonomicNameLabel id="A20657C0FFB6FFC4FDF6FE8DFDE7FE50" box="[572,639,383,402]" pageId="8" pageNumber="351" rank="species">sp. nov.</taxonomicNameLabel>
[handwritten] / det. Kondorosy &amp; Baňař [printed, red label]’.
</paragraph>
</subSubSection>
<subSubSection id="C35B6522FFB6FFC5FF5DFE34FD85FF37" lastPageId="9" lastPageNumber="352" pageId="8" pageNumber="351" type="description">
<paragraph id="8BFE36A9FFB6FFC4FF5DFE34FE3AFD9F" blockId="8.[151,761,454,733]" pageId="8" pageNumber="351">
<emphasis id="B935EABBFFB6FFC4FF5DFE34FEBAFE1F" bold="true" box="[151,290,454,477]" pageId="8" pageNumber="351">Description.</emphasis>
<emphasis id="B935EABBFFB6FFC4FEE3FE34FEEDFE1F" box="[297,373,454,477]" italics="true" pageId="8" pageNumber="351">Colour</emphasis>
similar to previous species, slightly paler; scape and pedicel almost uniformly pale; pale part of pronotum and hemelytra more extensive; scutellum pale except dark basal triangular spots along pale midline and lateral corners; legs pale.
</paragraph>
<paragraph id="8BFE36A9FFB6FFC4FF7FFD95FE5FFD1F" blockId="8.[151,761,454,733]" pageId="8" pageNumber="351">
<emphasis id="B935EABBFFB6FFC4FF7FFD95FE87FDBC" box="[181,287,615,638]" italics="true" pageId="8" pageNumber="351">Structure.</emphasis>
Pronotum trapezoidal, anterior lobe shorter. Hemelytra macropterous, having a fully developed membrane, almost reaching the end of the abdomen. Profemora without stiff subapical setae.
</paragraph>
<paragraph id="8BFE36A9FFB6FFC4FC90FF4CFAF0FED7" blockId="8.[828,1437,190,725]" pageId="8" pageNumber="351">Male genitalia: cup-like sclerite of pygophore with medial projection broad, not longer and similarly blunt as lateral apices. Paramere with moderately long setae.</paragraph>
<paragraph id="8BFE36A9FFB6FFC5FC90FEEDFD85FF37" blockId="8.[828,1437,190,725]" lastBlockId="9.[151,760,190,501]" lastPageId="9" lastPageNumber="352" pageId="8" pageNumber="351">
<emphasis id="B935EABBFFB6FFC4FC90FEEDFC6DFEF4" box="[858,1013,287,310]" italics="true" pageId="8" pageNumber="351">Measurements</emphasis>
(in mm): Total body length: 4.19; head: length 0.72, width 0.89, interocular space 0.49; length of eye 0.28, length of antenniferous tubercle 0.14; length of antennomeres: I 0.58, II 0.83, III 0.55, IV 0.61; length of labiomeres: I 0.61, II 0.49, III 0.50, IV 0.36; pronotum: length 0.91, width 1.41; scutellum: length 0.91, width 0.58. 
<emphasis id="B935EABBFFB6FFC4FCF6FE2CFBDAFE37" bold="true" box="[828,1090,478,501]" pageId="8" pageNumber="351">Differential diagnosis.</emphasis>
Both species are very similar but 
<taxonomicName id="4C414D2AFFB6FFC4FCADFE0DFC5EFDD7" authorityName="Kondorosy &amp; Banar" authorityYear="2020" box="[871,966,510,534]" class="Insecta" family="Rhyparochromidae" genus="Scobinigaster" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="351" phylum="Arthropoda" rank="species" species="paveli">
<emphasis id="B935EABBFFB6FFC4FCADFE0DFC5EFDD7" box="[871,966,510,534]" italics="true" pageId="8" pageNumber="351">S. paveli</emphasis>
</taxonomicName>
is macropterous, having a fully developed membrane, almost reaching the end of the abdomen. It is slightly smaller than 
<taxonomicName id="4C414D2AFFB6FFC4FBD2FDCDFBEDFD97" authorityName="Kondorosy &amp; Banar" authorityYear="2020" box="[1048,1141,574,598]" class="Insecta" family="Rhyparochromidae" genus="Scobinigaster" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="351" phylum="Arthropoda" rank="species" species="henryi">
<emphasis id="B935EABBFFB6FFC4FBD2FDCDFBEDFD97" box="[1048,1141,574,598]" italics="true" pageId="8" pageNumber="351">S. henryi</emphasis>
</taxonomicName>
, and with paler colouration. The form of the pronotum is clearly different, trapezoidal; the anterior lobe of the pronotum is also shorter than of 
<taxonomicName id="4C414D2AFFB6FFC4FA42FD8DFC18FD77" authorityName="Kondorosy &amp; Banar" authorityYear="2020" class="Insecta" family="Rhyparochromidae" genus="Scobinigaster" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="351" phylum="Arthropoda" rank="species" species="henryi">
<emphasis id="B935EABBFFB6FFC4FA42FD8DFC18FD77" italics="true" pageId="8" pageNumber="351">S. henryi</emphasis>
</taxonomicName>
. The most important differences are that in 
<taxonomicName id="4C414D2AFFB6FFC4FA88FD6DFA04FD77" authorityName="Kondorosy &amp; Banar" authorityYear="2020" box="[1346,1436,670,694]" class="Insecta" family="Rhyparochromidae" genus="Scobinigaster" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="351" phylum="Arthropoda" rank="species" species="paveli">
<emphasis id="B935EABBFFB6FFC4FA88FD6DFA04FD77" box="[1346,1436,670,694]" italics="true" pageId="8" pageNumber="351">S. paveli</emphasis>
</taxonomicName>
the stiff subapical setae of the profemora are absent, the paramere has shorter pubescence, and the cup-like sclerite of the pygophore is clearly different.
</paragraph>
</subSubSection>
<caption id="DF3E6621FFB6FFC4FF5DF804FC49F7E6" ID-DOI="http://doi.org/10.5281/zenodo.3879991" ID-Zenodo-Dep="3879991" httpUri="https://zenodo.org/record/3879991/files/figure.png" pageId="8" pageNumber="351" startId="8.[151,188,2038,2057]" targetBox="[311,1276,942,2021]" targetPageId="8">
<paragraph id="8BFE36A9FFB6FFC4FF5DF804FC49F7E6" blockId="8.[151,1436,2038,2085]" pageId="8" pageNumber="351">
Figs 28–29. 28 – typical habitat in Isalo NP, where part of the type series of 
<taxonomicName id="4C414D2AFFB6FFC4FCD9F804FBEBF7CB" authority="Kondorosy &amp; Banar" authorityName="Kondorosy &amp; Banar" authorityYear="2020" box="[787,1139,2038,2057]" class="Insecta" family="Rhyparochromidae" genus="Scobinigaster" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="351" phylum="Arthropoda" rank="species" species="henryi" status="sp. nov.">
<emphasis id="B935EABBFFB6FFC4FCD9F804FC5BF7CB" box="[787,963,2038,2057]" italics="true" pageId="8" pageNumber="351">Scobinigaster henryi</emphasis>
Kondorosy &amp; Baňař
</taxonomicName>
<taxonomicNameLabel id="A20657C0FFB6FFC4FBBDF804FB21F7CB" box="[1143,1209,2038,2057]" pageId="8" pageNumber="351" rank="species">sp. nov.</taxonomicNameLabel>
was collected; 29 – habitat in Mahafaly plateau, type locality of 
<taxonomicName id="4C414D2AFFB6FFC4FE19F7E0FD57F7E6" authority="Kondorosy &amp; Banar" authorityName="Kondorosy &amp; Banar" authorityYear="2020" box="[467,719,2065,2085]" class="Insecta" family="Rhyparochromidae" genus="Scobinigaster" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="351" phylum="Arthropoda" rank="species" species="paveli" status="sp. nov.">
<emphasis id="B935EABBFFB6FFC4FE19F7E0FD84F7E6" box="[467,540,2065,2085]" italics="true" pageId="8" pageNumber="351">S. paveli</emphasis>
Kondorosy &amp; Baňař
</taxonomicName>
<taxonomicNameLabel id="A20657C0FFB6FFC4FD1EF7E3FC8FF7E6" box="[724,791,2065,2084]" pageId="8" pageNumber="351" rank="species">sp. nov.</taxonomicNameLabel>
Photo: Miloš Trýzna.
</paragraph>
</caption>
<subSubSection id="C35B6522FFB7FFC5FF5DFF0CFD6EFE17" pageId="9" pageNumber="352" type="etymology">
<paragraph id="8BFE36A9FFB7FFC5FF5DFF0CFDD7FEB7" blockId="9.[151,760,190,501]" pageId="9" pageNumber="352">
<emphasis id="B935EABBFFB7FFC5FF5DFF0CFE80FED7" bold="true" box="[151,280,254,277]" pageId="9" pageNumber="352">Etymology.</emphasis>
We dedicate this species to the late Professor Pavel Štys, the eminent heteropterist, our teacher, mentor and friend, who died unexpectedly in 2018 (for biography and bibliography see KMൾNŦ et al. 2019).
</paragraph>
<paragraph id="8BFE36A9FFB7FFC5FF5DFE8CFD6EFE17" blockId="9.[151,760,190,501]" pageId="9" pageNumber="352">
<emphasis id="B935EABBFFB7FFC5FF5DFE8CFE2DFE57" bold="true" box="[151,437,382,405]" pageId="9" pageNumber="352">Collecting circumstances.</emphasis>
The single known specimen of this species was collected by a light trap in the Mahafaly plateau, an arid area in south-west 
<collectingCountry id="F3567639FFB7FFC5FDC0FE4CFD13FE17" box="[522,651,446,469]" name="Madagascar" pageId="9" pageNumber="352">Madagascar</collectingCountry>
(
<figureCitation id="137A2A2CFFB7FFC5FD50FE4CFD71FE17" box="[666,745,446,469]" captionStart="Figs 28–29. 28" captionStartId="8.[151,188,2038,2057]" captionTargetBox="[311,1276,942,2021]" captionTargetId="figure@8.[311,1276,779,2021]" captionTargetPageId="8" captionText="Figs 28–29. 28 – typical habitat in Isalo NP, where part of the type series of Scobinigaster henryi Kondorosy &amp; Baňař sp. nov. was collected; 29 – habitat in Mahafaly plateau, type locality of S. paveli Kondorosy &amp; Baňař sp. nov. Photo: Miloš Trýzna." figureDoi="http://doi.org/10.5281/zenodo.3879991" httpUri="https://zenodo.org/record/3879991/files/figure.png" pageId="9" pageNumber="352">Fig. 29</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35B6522FFB7FFC5FF5DFE2CFDADFE37" box="[151,565,478,501]" pageId="9" pageNumber="352" type="distribution">
<paragraph id="8BFE36A9FFB7FFC5FF5DFE2CFDADFE37" blockId="9.[151,760,190,501]" box="[151,565,478,501]" pageId="9" pageNumber="352">
<emphasis id="B935EABBFFB7FFC5FF5DFE2CFEB2FE37" bold="true" box="[151,298,478,501]" pageId="9" pageNumber="352">Distribution.</emphasis>
South-west 
<collectingCountry id="F3567639FFB7FFC5FE64FE2CFDA9FE37" box="[430,561,478,501]" name="Madagascar" pageId="9" pageNumber="352">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>