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<document id="4282B25999326E8251073D52014454C5" ID-CLB-Dataset="4613" ID-DOI="10.5281/zenodo.5332860" ID-GBIF-Dataset="cc461ce1-8e44-4862-9ff2-9c9ef7f73cd9" ID-ISSN="0253-116X" ID-Zenodo-Dep="5332860" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630172711352" checkinUser="carolina" docAuthor="Guéorguiev, B." docDate="2010" docId="03E887AB595F3155FF3D6EE9FCC9EF11" docLanguage="en" docName="LinzerbiolBeitr.2010.42.1.659-669.pdf" docOrigin="Linzer biologische Beiträge 42 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Pterostichus (Oreoplatysma) stomoides" docType="treatment" docVersion="7" lastPageNumber="663" masterDocId="FFD1FFD3595B3151FF8F6C75FF94E951" masterDocTitle="The &quot; colchicus &quot; group of Pterostichus (Coleoptera, Carabidae)" masterLastPageNumber="669" masterPageNumber="659" pageNumber="663" updateTime="1699187604789" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="37C4BF291174486F8382F78C99E9D913">The &quot; colchicus &quot; group of Pterostichus (Coleoptera, Carabidae)</mods:title>
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<treatment id="03E887AB595F3155FF3D6EE9FCC9EF11" ID-DOI="http://doi.org/10.5281/zenodo.5411006" ID-GBIF-Taxon="185227616" ID-Zenodo-Dep="5411006" LSID="urn:lsid:plazi:treatment:03E887AB595F3155FF3D6EE9FCC9EF11" httpUri="http://treatment.plazi.org/id/03E887AB595F3155FF3D6EE9FCC9EF11" lastPageNumber="663" pageId="4" pageNumber="663">
<subSubSection id="C35B6536595F3155FF3D6EE9FC05EBE3" box="[178,913,668,690]" pageId="4" pageNumber="663" type="nomenclature">
<paragraph id="8BFE36BD595F3155FF3D6EE9FC05EBE3" blockId="4.[178,913,668,690]" box="[178,913,668,690]" pageId="4" pageNumber="663">
<heading id="D0B681D1595F3155FF3D6EE9FC05EBE3" box="[178,913,668,690]" fontSize="8" level="2" pageId="4" pageNumber="663" reason="2">
<emphasis id="B935EAAF595F3155FF3D6EE9FC05EBE3" bold="true" box="[178,913,668,690]" pageId="4" pageNumber="663">
<taxonomicName id="4C414D3E595F3155FF3D6EE9FC9FEBE3" ID-CoL="4Q45Z" authority="(CHAUDOIR 1868)" baseAuthorityName="CHAUDOIR" baseAuthorityYear="1868" box="[178,779,668,690]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="663" phylum="Arthropoda" rank="species" species="stomoides" subGenus="Oreoplatysma">
<emphasis id="B935EAAF595F3155FF3D6EE9FEA7EBE3" bold="true" box="[178,307,668,690]" italics="true" pageId="4" pageNumber="663">Pterostichus</emphasis>
(
<emphasis id="B935EAAF595F3155FECD6EE9FE46EBE3" bold="true" box="[322,466,668,690]" italics="true" pageId="4" pageNumber="663">Oreoplatysma</emphasis>
)
<emphasis id="B935EAAF595F3155FE6E6EE9FDD3EBE3" bold="true" box="[481,583,668,690]" italics="true" pageId="4" pageNumber="663">stomoides</emphasis>
(
<bibRefCitation id="EFD04B4C595F3155FDD96EE9FD5EEBE3" author="CHAUDOIR M" box="[598,714,668,690]" pageId="4" pageNumber="663" pagination="219 - 260" refId="ref4258" refString="CHAUDOIR M. de (1868): Description d'especes nouvelles ou peu connues de Feronia d'Europe et du bassin mediterraneen. - L'Abeille, Memoires d'Entomologie 5: 219 - 260." type="journal article" year="1868">CHAUDOIR</bibRefCitation>
1868)
</taxonomicName>
(
<figureCitation id="137A2A38595F3155FC976EE9FC1DEBE3" box="[792,905,668,690]" captionStart="Figs 11-14" captionStartId="10.[178,218,1246,1266]" captionTargetBox="[200,1074,225,1217]" captionTargetId="figure-2@10.[178,1085,220,1236]" captionTargetPageId="10" captionText="Figs 11-14: Feronia stomoides CHAUDOIR, lectotype: Fig. (11) aedeagus, left lateral view; Fig. (12) apex of aedeagus, dorsal view; Fig. (13) right paramere, internal view; Fig. (14) right paramere, external view. Scale bar 0.5 mm (11-14)." figureDoi="http://doi.org/10.5281/zenodo.5332866" httpUri="https://zenodo.org/record/5332866/files/figure.png" pageId="4" pageNumber="663">Figs 11-14</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35B6536595F3155FF3D6EB9FE3DEA31" pageId="4" pageNumber="663" type="reference_group">
<paragraph id="8BFE36BD595F3155FF3D6EB9FD89EBB1" blockId="4.[178,1086,716,896]" box="[178,541,716,736]" pageId="4" pageNumber="663">
<treatmentCitationGroup id="AB511193595F3155FF3D6EB9FD89EBB1" box="[178,541,716,736]" pageId="4" pageNumber="663">
<emphasis id="B935EAAF595F3155FF3D6EB9FF6CEBB1" box="[178,248,716,736]" italics="true" pageId="4" pageNumber="663">Feronia</emphasis>
<treatmentCitation id="0AE010AC595F3155FF726EB9FD8DEBB1" author="CHAUDOIR M" box="[253,537,716,736]" page="251" pageId="4" pageNumber="663" year="1868">
<taxonomicName id="4C414D3E595F3155FF726EB9FD8DEBB1" ID-CoL="4Q45Z" authority="CHAUDOIR 1868: 251" authorityName="CHAUDOIR" authorityPageNumber="251" authorityYear="1868" box="[253,537,716,736]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="663" phylum="Arthropoda" rank="species" species="stomoides">
<emphasis id="B935EAAF595F3155FF726EB9FEC0EBB1" box="[253,340,716,736]" italics="true" pageId="4" pageNumber="663">stomoides</emphasis>
<bibRefCitation id="EFD04B4C595F3155FED76EB9FD8DEBB1" author="CHAUDOIR M" box="[344,537,716,736]" pageId="4" pageNumber="663" pagination="219 - 260" refId="ref4258" refString="CHAUDOIR M. de (1868): Description d'especes nouvelles ou peu connues de Feronia d'Europe et du bassin mediterraneen. - L'Abeille, Memoires d'Entomologie 5: 219 - 260." type="journal article" year="1868">CHAUDOIR 1868: 251</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFE36BD595F3155FF3D6E99FE3DEA31" blockId="4.[178,1086,716,896]" pageId="4" pageNumber="663">
T y p e m a t e r i a l 1,
<specimenCount id="9D47FD34595F3155FE546E99FDC4EA51" box="[475,592,748,768]" pageId="4" pageNumber="663" type="generic" typeStatus="syntypes">1 syntypes</specimenCount>
(male
<typeStatus id="54FA881F595F3155FD196E99FD71EA51" box="[662,741,748,768]" pageId="4" pageNumber="663" type="lectotype">lectotype</typeStatus>
and female
<typeStatus id="54FA881F595F3155FCEF6E99FC40EA51" box="[864,980,748,768]" pageId="4" pageNumber="663" type="paralectotype">paralectotype</typeStatus>
by present designation), well preserved; male bears label &quot;Touchètie&quot; [handwritten], and the female label &quot;Grusia 1870&quot; [handwritten]. Close to both specimens, the following handwritten label is pinned on the bottom of box: &quot;
<taxonomicName id="4C414D3E595F3155FE1A6F41FC9BEA19" ID-CoL="4Q45Z" authority="MOTSCH. Caucase Motschulsky" authorityName="MOTSCH. Caucase Motschulsky" box="[405,783,820,840]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="663" phylum="Arthropoda" rank="species" species="stomoides">
<emphasis id="B935EAAF595F3155FE1A6F41FE78EA19" box="[405,492,820,840]" italics="true" pageId="4" pageNumber="663">stomoides</emphasis>
MOTSCH. Caucase Motschulsky
</taxonomicName>
&quot; (MNHN, &quot;Collection Chaudoir&quot;, box no 216 &quot;
<emphasis id="B935EAAF595F3155FEC06F39FE07EA31" box="[335,403,844,864]" italics="true" pageId="4" pageNumber="663">Argutor</emphasis>
&quot;).
</paragraph>
</subSubSection>
<subSubSection id="C35B6536595F3155FF3D6F19FCC9EF11" pageId="4" pageNumber="663" type="description">
<paragraph id="8BFE36BD595F3155FF3D6F19FBAEEAD1" blockId="4.[178,1086,716,896]" box="[178,1082,876,896]" pageId="4" pageNumber="663">
T y p e l o c a l i t y.&quot;AlpesdelaTouchétie(Caucasecentraletoriental)&quot;(
<bibRefCitation id="EFD04B4C595F3155FCE26F19FBBBEAD1" author="CHAUDOIR M" box="[877,1071,876,896]" pageId="4" pageNumber="663" pagination="219 - 260" refId="ref4258" refString="CHAUDOIR M. de (1868): Description d'especes nouvelles ou peu connues de Feronia d'Europe et du bassin mediterraneen. - L'Abeille, Memoires d'Entomologie 5: 219 - 260." type="journal article" year="1868">CHAUDOIR 1868: 251</bibRefCitation>
).
</paragraph>
<paragraph id="8BFE36BD595F3155FF3D6FFAFD94ED1E" blockId="4.[178,1086,910,1103]" pageId="4" pageNumber="663">
M a l e g e n i t a l i a. Median lobe of aedeagus in lateral view (
<figureCitation id="137A2A38595F3155FCE46FFBFC20EAF5" box="[875,948,910,932]" captionStart="Figs 11-14" captionStartId="10.[178,218,1246,1266]" captionTargetBox="[200,1074,225,1217]" captionTargetId="figure-2@10.[178,1085,220,1236]" captionTargetPageId="10" captionText="Figs 11-14: Feronia stomoides CHAUDOIR, lectotype: Fig. (11) aedeagus, left lateral view; Fig. (12) apex of aedeagus, dorsal view; Fig. (13) right paramere, internal view; Fig. (14) right paramere, external view. Scale bar 0.5 mm (11-14)." figureDoi="http://doi.org/10.5281/zenodo.5332866" httpUri="https://zenodo.org/record/5332866/files/figure.png" pageId="4" pageNumber="663">Fig. 11</figureCitation>
) with ostium deflected to left, apical part longer than basal part and with straight ventral margin, tip of apex somewhat bent upwards; basal part with strongly developed and prominent ventral process; internal sac (inverted state) with single strongly chitinised piece. Apex of median lobe in dorsal view (
<figureCitation id="137A2A38595F3155FE316875FD9DED47" box="[446,521,1024,1046]" captionStart="Figs 11-14" captionStartId="10.[178,218,1246,1266]" captionTargetBox="[200,1074,225,1217]" captionTargetId="figure-2@10.[178,1085,220,1236]" captionTargetPageId="10" captionText="Figs 11-14: Feronia stomoides CHAUDOIR, lectotype: Fig. (11) aedeagus, left lateral view; Fig. (12) apex of aedeagus, dorsal view; Fig. (13) right paramere, internal view; Fig. (14) right paramere, external view. Scale bar 0.5 mm (11-14)." figureDoi="http://doi.org/10.5281/zenodo.5332866" httpUri="https://zenodo.org/record/5332866/files/figure.png" pageId="4" pageNumber="663">Fig. 12</figureCitation>
) with tip deflected to left. Left paramere (
<figureCitation id="137A2A38595F3155FC4B6875FBA1ED47" box="[964,1077,1024,1046]" captionStart="Figs 11-14" captionStartId="10.[178,218,1246,1266]" captionTargetBox="[200,1074,225,1217]" captionTargetId="figure-2@10.[178,1085,220,1236]" captionTargetPageId="10" captionText="Figs 11-14: Feronia stomoides CHAUDOIR, lectotype: Fig. (11) aedeagus, left lateral view; Fig. (12) apex of aedeagus, dorsal view; Fig. (13) right paramere, internal view; Fig. (14) right paramere, external view. Scale bar 0.5 mm (11-14)." figureDoi="http://doi.org/10.5281/zenodo.5332866" httpUri="https://zenodo.org/record/5332866/files/figure.png" pageId="4" pageNumber="663">Figs 13-14</figureCitation>
) sickle-shaped, regularly round in middle; basal part forming big widened bulb; apical part with round (not pointed) tip.
</paragraph>
<paragraph id="8BFE36BD595F3155FF3D6828FE6AEDC1" blockId="4.[178,1085,1117,1168]" pageId="4" pageNumber="663">
After drawing, the genitalia were included in Euparal and put on separate transparent label pinned under the
<typeStatus id="54FA881F595F3155FE15680FFE6EEDC1" box="[410,506,1146,1168]" pageId="4" pageNumber="663" type="lectotype">lectotype</typeStatus>
.
</paragraph>
<paragraph id="8BFE36BD595F3155FF3D68EAFCBCEC0E" blockId="4.[178,1085,1182,1375]" pageId="4" pageNumber="663">
T a x o n o m i c n o t e s a n d r e l a t i o n s h i p s The presence of pubescence on the distal part of third antennomeres, dense and continuous punctuation on the basal part of pronotum, absence of medial setiferous punctures on hind coxae, glabrous ventral side of onychium, and sickle-shaped left paramere demonstrate that this species belongs to subgenus
<taxonomicName id="4C414D3E595F3155FE046965FD8DEC77" authorityName="JACOBSON" authorityYear="1907" box="[395,537,1296,1318]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="663" phylum="Arthropoda" rank="subGenus" subGenus="Oreoplatysma">
<emphasis id="B935EAAF595F3155FE046965FD8DEC77" box="[395,537,1296,1318]" italics="true" pageId="4" pageNumber="663">Oreoplatysma</emphasis>
</taxonomicName>
, and not to
<taxonomicName id="4C414D3E595F3155FD186965FD66EC77" authorityName="CHAUDOIR" authorityYear="1838" box="[663,754,1296,1318]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="666" phylum="Arthropoda" rank="subGenus" subGenus="Cryobius">
<emphasis id="B935EAAF595F3155FD186965FD66EC77" box="[663,754,1296,1318]" italics="true" pageId="4" pageNumber="663">Cryobius</emphasis>
</taxonomicName>
. Therefore, the following transfer is proposed:
<taxonomicName id="4C414D3E595F3155FED26959FC21EC13" authority="(CHAUDOIR 1868)" baseAuthorityName="CHAUDOIR" baseAuthorityYear="1868" box="[349,949,1324,1346]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="663" phylum="Arthropoda" rank="species" species="stomoides" subGenus="Oreoplatysma">
<emphasis id="B935EAAF595F3155FED26959FE4FEC13" box="[349,475,1324,1346]" italics="true" pageId="4" pageNumber="663">Pterostichus</emphasis>
(
<emphasis id="B935EAAF595F3155FE636959FDEFEC13" box="[492,635,1324,1346]" italics="true" pageId="4" pageNumber="663">Oreoplatysma</emphasis>
)
<emphasis id="B935EAAF595F3155FD036959FD65EC13" box="[652,753,1324,1346]" italics="true" pageId="4" pageNumber="663">stomoides</emphasis>
(
<bibRefCitation id="EFD04B4C595F3155FC8D6959FC3AEC13" author="CHAUDOIR M" box="[770,942,1324,1346]" pageId="4" pageNumber="663" pagination="219 - 260" refId="ref4258" refString="CHAUDOIR M. de (1868): Description d'especes nouvelles ou peu connues de Feronia d'Europe et du bassin mediterraneen. - L'Abeille, Memoires d'Entomologie 5: 219 - 260." type="journal article" year="1868">CHAUDOIR 1868</bibRefCitation>
)
</taxonomicName>
, new assignment of
<taxonomicName id="4C414D3E595F3155FE89693CFCB0EC0E" authority="(CHAUDOIR 1868)" baseAuthorityName="CHAUDOIR" baseAuthorityYear="1868" box="[262,804,1353,1375]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="663" phylum="Arthropoda" rank="species" species="stomoides" subGenus="Cryobius">
<emphasis id="B935EAAF595F3155FE89693CFE10EC0E" box="[262,388,1353,1375]" italics="true" pageId="4" pageNumber="663">Pterostichus</emphasis>
(
<emphasis id="B935EAAF595F3155FE1C693CFE7BEC0E" box="[403,495,1353,1375]" italics="true" pageId="4" pageNumber="663">Cryobius</emphasis>
)
<emphasis id="B935EAAF595F3155FE70693CFDF0EC0E" box="[511,612,1353,1375]" italics="true" pageId="4" pageNumber="663">stomoides</emphasis>
(
<bibRefCitation id="EFD04B4C595F3155FDFE693CFC89EC0E" author="CHAUDOIR M" box="[625,797,1353,1375]" pageId="4" pageNumber="663" pagination="219 - 260" refId="ref4258" refString="CHAUDOIR M. de (1868): Description d'especes nouvelles ou peu connues de Feronia d'Europe et du bassin mediterraneen. - L'Abeille, Memoires d'Entomologie 5: 219 - 260." type="journal article" year="1868">CHAUDOIR 1868</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8BFE36BD595F3155FF3D6918FC0AECA4" blockId="4.[178,1086,1389,1525]" pageId="4" pageNumber="663">
According to the external morphology and structure of the male genitalia, this species seems closer to the taxa from the group &quot;
<emphasis id="B935EAAF595F3155FDD069FFFD49ECF1" box="[607,733,1418,1440]" italics="true" pageId="4" pageNumber="663">daghestanus</emphasis>
&quot;, and especially to
<taxonomicName id="4C414D3E595F3155FC2169FEFEDDECED" authority="REITTER 1896" authorityName="REITTER" authorityYear="1896" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="663" phylum="Arthropoda" rank="species" species="chefsuricus">
<emphasis id="B935EAAF595F3155FC2169FEFC28ECF1" box="[942,956,1419,1440]" italics="true" pageId="4" pageNumber="663">P</emphasis>
.
<emphasis id="B935EAAF595F3155FC4569FFFBA9ECF1" box="[970,1085,1418,1440]" italics="true" pageId="4" pageNumber="663">chefsuricus</emphasis>
<bibRefCitation id="EFD04B4C595F3155FF3D69D2FEDDECED" author="REITTER E" box="[178,329,1446,1468]" pageId="4" pageNumber="663" pagination="211 - 215" refId="ref4942" refString="REITTER E. (1896): Ubersicht der mit Pterostichus pulchellus FALDERM. verwandten Coleopteren-Arten, = Agnodemus CHAUD. - Entomologische Nachrichten (Berlin) 22: 211 - 215." type="journal article" year="1896">REITTER 1896</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C414D3E595F3155FE0469D2FCACECED" authority="KRYZHANOVSKIJ 1988" authorityName="KRYZHANOVSKIJ" authorityYear="1988" box="[395,824,1446,1468]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="663" phylum="Arthropoda" rank="species" species="kirschenblatti">
<emphasis id="B935EAAF595F3155FE0469D2FE0EECED" box="[395,410,1447,1468]" italics="true" pageId="4" pageNumber="663">P</emphasis>
.
<emphasis id="B935EAAF595F3155FE2069D3FDAEECED" box="[431,570,1446,1468]" italics="true" pageId="4" pageNumber="663">kirschenblatti</emphasis>
<bibRefCitation id="EFD04B4C595F3155FDC669D2FCACECED" author="KRYZHANOVSKIJ O" box="[585,824,1446,1468]" pageId="4" pageNumber="663" pagination="12 - 21" refId="ref4660" refString="KRYZHANOVSKIJ O. L. (1988): [A review of the Caucasian species of the subgenus Oreoplatysma JACOBSON, 1906 (Coleoptera, Carabidae, Pterostichus)]. - Trudy Vsesoiuznogo Entomologichesjogo Obshchestva 70: 12 - 21 [in Russian]." type="journal article" year="1988">KRYZHANOVSKIJ 1988</bibRefCitation>
</taxonomicName>
. However, the apex of aedeagus distinctly bent to left and somewhat turn upwards and different shape of the left paramere suppose that
<taxonomicName id="4C414D3E595F3155FE156995FD88ECA4" baseAuthorityName="CHAUDOIR" baseAuthorityYear="1868" box="[410,540,1503,1525]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="663" phylum="Arthropoda" rank="species" species="stomoides">
<emphasis id="B935EAAF595F3155FE156995FE3DECA4" box="[410,425,1504,1525]" italics="true" pageId="4" pageNumber="663">P</emphasis>
.
<emphasis id="B935EAAF595F3155FE3869AAFD88ECA4" box="[439,540,1503,1525]" italics="true" pageId="4" pageNumber="663">stomoides</emphasis>
</taxonomicName>
is distinct from the other two species.
</paragraph>
<paragraph id="8BFE36BD595F3155FF3D6A73FD06EF4B" blockId="4.[178,658,1540,1563]" box="[178,658,1540,1563]" pageId="4" pageNumber="663">
<taxonomicName id="4C414D3E595F3155FF3D6A73FEA7EF4B" baseAuthorityName="CHAUDOIR" baseAuthorityYear="1868" box="[178,307,1540,1563]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="663" phylum="Arthropoda" rank="species" species="stomoides">
<emphasis id="B935EAAF595F3155FF3D6A73FF55EF4A" box="[178,193,1542,1563]" italics="true" pageId="4" pageNumber="663">P</emphasis>
.
<emphasis id="B935EAAF595F3155FF416A71FEA7EF4B" box="[206,307,1540,1562]" italics="true" pageId="4" pageNumber="663">stomoides</emphasis>
</taxonomicName>
should be treated as valid species.
</paragraph>
<paragraph id="8BFE36BD595F3155FF3D6A5EFCC9EF11" blockId="4.[178,861,1578,1600]" box="[178,861,1578,1600]" pageId="4" pageNumber="663">D i s t r i b u t i o n.Tushetia(KRYZHANOVSKIJ et al. 1995: 102).</paragraph>
</subSubSection>
</treatment>
</document>