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<document id="9A993307B15937B50CAC280CB56689BD" ID-DOI="10.14446/AMNP.2014.55" ID-ISSN="2533-4069" ID-Zenodo-Dep="13183435" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722606817950" checkinUser="felipe" docAuthor="Sabol, Martin &amp; Zervanová, Júlia" docDate="2014" docId="03E8878BFF90FFDEFF6D50AAECEBAF32" docLanguage="en" docName="ActaMusNatPraSerB.70.1-2.55-58.pdf" docOrigin="Acta Musei Nationalis Pragae Series B 70 (1 - 2)" docSource="https://www.mendeley.com/catalogue/945d27fa-849f-393e-9432-de9565e98445/" docStyle="DocumentStyle:63483C499D9A2BE76998B4D5AD5DCB53.3:ActaMusNatPraSerB.2014-2015.journal_article" docStyleId="63483C499D9A2BE76998B4D5AD5DCB53" docStyleName="ActaMusNatPraSerB.2014-2015.journal_article" docStyleVersion="3" docTitle="Cricetus runtonensis NEWTON 1909" docType="treatment" docVersion="1" lastPageNumber="57" masterDocId="FFD1FFF3FF91FFDCFFBC566EEE6CAC73" masterDocTitle="Cricetus Cf. Runtonensis (Newton, 1909) (Cricetidae, Rodentia) From Za Hájovnou Cave (The Czech Republic)" masterLastPageNumber="58" masterPageNumber="55" pageNumber="56" updateTime="1722656563024" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="864CD1CE7774F449CC01F5BFC4B76476">Cricetus Cf. Runtonensis (Newton, 1909) (Cricetidae, Rodentia) From Za Hájovnou Cave (The Czech Republic)</mods:title>
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<mods:affiliation id="CC1D86BC062BA08CB660094B3B2D1802">Department of Geology and Palaeontology, Comenius University, Mlynská dolina, SK - 842 15 Bratislava, Slovak Republic; e-mails: sabol @ fns. uniba. sk; zervanova @ fns. uniba. sk</mods:affiliation>
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<taxonomicName id="4C414D1EFF90FFDDFF6D50AAECEDAAA8" authority="NEWTON, 1909" authorityName="NEWTON" authorityYear="1909" box="[209,641,1732,1756]" class="Mammalia" family="Cricetidae" genus="Cricetus" isUncertain="true" kingdom="Animalia" order="Rodentia" pageId="1" pageNumber="56" phylum="Chordata" rank="species" species="runtonensis">
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<emphasis id="B935EA8FFF90FFDDFF6D50AAEF41AAAF" bold="true" box="[209,301,1732,1756]" italics="true" pageId="1" pageNumber="56">Cricetus</emphasis>
cf.
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<bibRefCitation id="EFD04B6CFF90FFDDFE6350AAEC52AAA8" author="Newton, E. T." box="[479,574,1732,1755]" pageId="1" pageNumber="56" pagination="110 - 117" refId="ref4123" refString="Newton, E. T. (1909): Hamster remains from the Norfolk Forest Bed. - Geological Magazine, 6: 110 - 117. http: // dx. doi. org / 10.1017 / S 0016756800120552" type="journal article" year="1909">
<smallCapsWord id="8D18A041FF90FFDDFE6350AAEC52AAA8" baselines="1749,1750" box="[479,574,1732,1755]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Newton" pageId="1" pageNumber="56">NEWTON</smallCapsWord>
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, 1909
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<figureCitation id="137A2A18FF90FFDDFECD509DEF8DAB79" box="[369,481,1779,1802]" captionStart="Text-fig" captionStartId="1.[820,901,920,941]" captionTargetBox="[106,1459,151,1094]" captionTargetPageId="1" captionText="Text-fig. 2. Fossil remains of Cricetus cf. runtonensis NEWTON, 1909 from Za Hájovnou Cave (Moravia, Czech Republic), Middle Pleistocene. ab) lower molar row (a occlusal view, b lingual view); c) m1 dext., occlusal view; de) left mandible fragment with incisor (d lingual view, e buccal view); f) calcaneus sin., anterior view." figureDoi="http://doi.org/10.5281/zenodo.13183437" httpUri="https://zenodo.org/record/13183437/files/figure.png" pageId="1" pageNumber="56">Text-fig. 2</figureCitation>
</paragraph>
<paragraph id="8BFE369DFF90FFDDFFD6515CECF6AB17" blockId="1.[106,745,1842,1892]" pageId="1" pageNumber="56">
1909
<emphasis id="B935EA8FFF90FFDDFF76515CEF21AB34" box="[202,333,1842,1863]" italics="true" pageId="1" pageNumber="56">Newton, E. T.</emphasis>
, Hamster remains from the
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Forest Bed. Geological Magazine, 6, London, 110-117.
</paragraph>
<paragraph id="8BFE369DFF90FFDDFF2C51E3EC84ABD7" blockId="1.[106,744,1933,1988]" box="[144,744,1933,1956]" pageId="1" pageNumber="56">
<materialsCitation id="3B293CC0FF90FFDDFF2C51E3EC84ABD7" box="[144,744,1933,1956]" collectionCode="T" country="United Kingdom" location="West Runton" pageId="1" pageNumber="56" specimenCount="1" stateProvince="Norfolk">
L o c u s
<collectionCode id="ED50AE58FF90FFDDFF4651E3EF66ABD7" box="[250,266,1933,1956]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="56">T</collectionCode>
y p i c u s.
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,
<collectingRegion id="4985F87FFF90FFDDFD9451E3ECEDABD7" box="[552,641,1933,1956]" country="United Kingdom" name="Norfolk" pageId="1" pageNumber="56">Norfolk</collectingRegion>
,
<collectingCountry id="F356760DFF90FFDDFD3251E3EC84ABD7" box="[654,744,1933,1956]" name="United Kingdom" pageId="1" pageNumber="56">England</collectingCountry>
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</paragraph>
<paragraph id="8BFE369DFF90FFDDFFD651C3EC18ABB7" blockId="1.[106,744,1933,1988]" box="[106,628,1965,1988]" pageId="1" pageNumber="56">(Cromerian Complex, early Middle Pleistocene).</paragraph>
<paragraph id="8BFE369DFF90FFDDFF2C51B2EFD0A440" blockId="1.[106,745,2012,2099]" pageId="1" pageNumber="56">S t r a t i g r a p h i c a n d g e o g r a p h i c r a n g e. Early to late Middle Pleistocene (Nagyharsányhegyian to Solymárian substages), Europe.</paragraph>
<paragraph id="8BFE369DFF90FFDDFCE652E8EBDFA94A" blockId="1.[820,1459,1158,1337]" pageId="1" pageNumber="56">
S p e c i e s c h a r a c t e r i s t i c s. A relatively large extinct representative of the genus
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<emphasis id="B935EA8FFF90FFDDFB1552CBEB6FA8CF" box="[1193,1283,1189,1212]" italics="true" pageId="1" pageNumber="56">Cricetus</emphasis>
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, smaller than
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<emphasis id="B935EA8FFF90FFDDFA2652CBEBC0A8CF" box="[1434,1452,1189,1212]" italics="true" pageId="1" pageNumber="56">C</emphasis>
.
<emphasis id="B935EA8FFF90FFDDFC8852AAED18A8A8" box="[820,884,1220,1243]" italics="true" pageId="1" pageNumber="56">major</emphasis>
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, but larger than
<taxonomicName id="4C414D1EFF90FFDDFB9F52AAEAA3A8A8" authorityName="SCHAUB" authorityYear="1930" box="[1059,1231,1220,1243]" class="Mammalia" family="Cricetidae" genus="Cricetus" kingdom="Animalia" order="Rodentia" pageId="1" pageNumber="56" phylum="Chordata" rank="species" species="praeglacialis">
<emphasis id="B935EA8FFF90FFDDFB9F52AAEAA3A8A8" box="[1059,1231,1220,1243]" italics="true" pageId="1" pageNumber="56">C. praeglacialis</emphasis>
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and
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<emphasis id="B935EA8FFF90FFDDFABF52AAEB1AA8A8" box="[1283,1398,1220,1243]" italics="true" pageId="1" pageNumber="56">C. cricetus</emphasis>
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, with a cusp-shaped entostyle on M1 and M2, and an ectostyle on m2 and m3 with more frequently doubled mesolophid in stratigraphically later forms (
<bibRefCitation id="EFD04B6CFF90FFDDFBD6534CEAB7A94A" author="Hir, J." box="[1130,1243,1314,1337]" pageId="1" pageNumber="56" pagination="43 - 49" refId="ref3422" refString="Hir, J. (1997 b): A short sketch of the evolution and stratigraphy of the Plio-Pleistocene cricetids (Rodentia, Mammalia) in Hungary. - Folia Historico Naturalia Musei Matraensis, 22: 43 - 49." type="journal article" year="1997">Hír 1997b</bibRefCitation>
, c, 1998; modified).
</paragraph>
<paragraph id="8BFE369DFF90FFDDFCE6533EEA40A9B7" blockId="1.[820,1459,1360,1476]" pageId="1" pageNumber="56">
M a t e r i a l. A right fragmented mandible with dentition (incisor, isolated m1, and m2 m
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the rock) and a left calcaneus. The bones are a light colour (brown-white) and teeth crowns are white.
</paragraph>
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D e s c r i p t i o n. The anterior part of the mandible fragment includes a slightly damaged incisor. The mental foramen is large, situated below the diastema, between the incisor and m1. The rest of the mandible ramus is located below the molars in the rock. The m1 is damaged in the posterior part on both lateral sides. This tooth is isolated but in anatomical order with m2 and m3, belonging to the posterior part of the mandible. The double-cuspid anteroconid of m1 is more or less isolated, slightly worn on the lingual conelet tip and damaged on the buccal conelet. The anterolophulid can be considered more likely to be reduced rather than connected to the buccal conelet of the anteroconid. Other cuspids (metaconid, protoconid, entoconid, hypoconid) are moderately worn. The mesolophid is missing. Since the posterior part of the crown is damaged, the morphology of the posterolophulid cannot be classified. The remaining part, however, suggests that it was probably simple rather than ramified. Based on that, this molar would more likely represent morphotype F1a than E1a as defined by
<bibRefCitation id="EFD04B6CFF93FFDEFF1F56F6EF76ACDC" author="Hir, J." box="[163,282,152,175]" pageId="2" pageNumber="57" pagination="51 - 79" refId="ref3377" refString="Hir, J. (1997 a): A Comparative study on the dental morphology of the Early Pleistocene Cricetus praeglacialis SCHAUB, 1930 and recent Hungarian Cricetus cricetus L. - Folia Historico Naturalia Musei Matraensis, 22: 51 - 79." type="journal article" year="1997">Hír (1997a</bibRefCitation>
, c, 1998). The two morphotypes differ in the level of anterolophulid reduction (reduced in the F1a morphotype and non-reduced, connected to the anteroconid buccal conelet in the E1a morphotype).
</paragraph>
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Table 1. Size comparison of m1 from Za Hájovnou Cave with that of various Quaternary species of
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<emphasis id="B935EA8FFF93FFDEFB7256D9EB4DACBE" bold="true" box="[1230,1313,183,205]" italics="true" pageId="2" pageNumber="57">Cricetus</emphasis>
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(L = length, WA = anterior width, WP = posterior width). Measurements in mm. Data source: Fahlbusch 1986,
<bibRefCitation id="EFD04B6CFF93FFDEFB37569EEB69AD76" author="Fejfar, O." box="[1163,1285,240,261]" pageId="2" pageNumber="57" pagination="277 - 296" refId="ref3191" refString="Fejfar, O. (1970): Die plio-pleistozanen Wirbeltierfaunen von Hajnacka und Ivanovce (Slowakei, CSSR) VI. Cricetidae (Rodentia, Mammalia) [Plio-Pleistocene Vertebrate Fauna of Hajnacka and Ivanovce (Slovakia, CSSR) VI. Cricetidae (Rodentia, Mammalia)]. - Mitteilungen der Bayerischen Staatssammlung fur Palaontologie und Histor. Geologie., 10: 277 - 296." type="journal article" year="1970">Fejfar 1970</bibRefCitation>
,
<bibRefCitation id="EFD04B6CFF93FFDEFAAD569EEB17AD76" author="Hir, J." box="[1297,1403,240,261]" pageId="2" pageNumber="57" pagination="51 - 79" refId="ref3377" refString="Hir, J. (1997 a): A Comparative study on the dental morphology of the Early Pleistocene Cricetus praeglacialis SCHAUB, 1930 and recent Hungarian Cricetus cricetus L. - Folia Historico Naturalia Musei Matraensis, 22: 51 - 79." type="journal article" year="1997">Hír 1997a</bibRefCitation>
,
<bibRefCitation id="EFD04B6CFF93FFDEFA34569EEBA9AD76" author="Hir, J." box="[1416,1477,240,261]" pageId="2" pageNumber="57" pagination="23 - 42" refId="ref3464" refString="Hir, J. (1997 c): Cricetus runtonensis solymarensis ssp. n. (Mammalia, Rodentia) from the Late Middle Pleistocene fauna of Solymar. - Annales Historico-Naturales Musei Nationalis Hungarici, 89: 23 - 42." type="journal article" year="1997">1997c</bibRefCitation>
,
<bibRefCitation id="EFD04B6CFF93FFDEFCF75763ED10AD51" author="Hir, J." box="[843,892,269,290]" pageId="2" pageNumber="57" pagination="57 - 89" refId="ref3507" refString="Hir, J. (1998): Cricetids (Rodentia, Mammalia) of the Early Pleistocene vertebrate fauna of Somssich-hegy 2 (Southern Hungary, Villany Mountains). - Annales Historico-Naturales Musei Nationalis Hungarici, 90: 57 - 89." type="journal article" year="1998">1998</bibRefCitation>
,
<bibRefCitation id="EFD04B6CFF93FFDEFC3B5763EA6FAD51" author="Pradel, A." box="[903,1027,269,290]" pageId="2" pageNumber="57" pagination="235 - 296" refId="ref4164" refString="Pradel, A. (1988): Fossil hamsters (Cricetinae, Rodentia) from the Pliocene and Quaternary of Poland. - Acta zoologica cracoviensia, 31 (6): 235 - 296." type="journal article" year="1988">Pradel 1988</bibRefCitation>
.
</paragraph>
</caption>
<paragraph id="8BFE369DFF93FFDEFF1A5779EF18ADFF" blockId="2.[129,768,152,833]" pageId="2" pageNumber="57">The lower molars (m2 and m3) are mostly covered by rock, and only the lingual cuspids (larger metaconid and lower entoconid) can be recognized. The antero-buccal part of m2 crown is absent.</paragraph>
<paragraph id="8BFE369DFF93FFDEFF1A57FAECEBAF32" blockId="2.[129,768,152,833]" pageId="2" pageNumber="57">
The left calcaneus is completely preserved, probably belonging to the same individual as the fragmented mandible with teeth. The calcaneus is 9.0 mm long and
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wide from the peroneal process to the medial tip of the sustentaculum. The calcaneal tubercle is long, transversely compressed with a slightly expanded proximal head. The calcaneoastragalar facet is convex and smoothly rounded. The sustentacular facet is flat and faces more or less dorsally. The narrow and relatively poorly defined sulcus calcaneus separates the sustentacular facet from the calcaneoastragalar facet. The sustentaculum talus is prominent, projecting medially. The distinct, shelf-shaped peroneal process is more expanded proximodistally, situated almost at the same level as the sustentaculum talus. The anterior plantar tubercle is faint, positioned more medially.
</paragraph>
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