treatments-xml/data/03/E3/87/03E387A1FFC4FFD2FF1BFE1CD004FF72.xml
2024-06-21 12:22:17 +02:00

452 lines
52 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="94AA65D793E5C852E027ACB07390CB3D" ID-DOI="10.11646/zootaxa.3936.4.7" ID-GBIF-Dataset="eae1b1d0-0ba9-4a39-b10c-8a8f5ba02085" ID-ISSN="1175-5326" ID-Zenodo-Dep="244127" ID-ZooBank="9F14082D-3680-4D75-B3F3-398F912ECCDD" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461017551092" checkinUser="plazi" docAuthor="Batuwita, Sudesh, Maduwage, Kalana &amp; Sudasinghe, Hiranya" docDate="2015" docId="03E387A1FFC4FFD2FF1BFE1CD004FF72" docLanguage="en" docName="zt03936p583.pdf" docOrigin="Zootaxa 3936 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pethia melanomaculata Deraniyagala 1956" docType="treatment" docVersion="8" lastPageNumber="580" masterDocId="FFDAFFD9FFC5FFD7FF8CFFC7D431FF85" masterDocTitle="Redescription of Pethia melanomaculata (Teleostei: Cyprinidae) from Sri Lanka" masterLastPageNumber="583" masterPageNumber="575" pageNumber="576" updateTime="1698569171914" updateUser="plazi">
<mods:mods id="E6D200950E2CD93BBC95A4E659456D5E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9C716466743D12A396F1FB7B7D5A56BE">
<mods:title id="B821D1CD9A4B33B342F23AEDE0613C1C">Redescription of Pethia melanomaculata (Teleostei: Cyprinidae) from Sri Lanka</mods:title>
</mods:titleInfo>
<mods:name id="AA130B4BAAF23273BE45654F89F7D609" type="personal">
<mods:role id="0B79E44A44C0EC0EEFA5C5E505DF3900">
<mods:roleTerm id="ABAA19930BC9DB1A8EFFE626A5CC60E9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A55A5A7DA4C0F4BCA9909D551ADFEE16">Batuwita, Sudesh</mods:namePart>
</mods:name>
<mods:name id="D59E6C3C845839F3CC651841833F0213" type="personal">
<mods:role id="D99C5AE2DADB3E671EA724ED7969E2CB">
<mods:roleTerm id="AB18BE06293DE2869CA004180BFF8AE7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0F423126BA1EC2C4336280F1C648966B">Maduwage, Kalana</mods:namePart>
</mods:name>
<mods:name id="4E71422B4E726A46B46F15761AA60AEA" type="personal">
<mods:role id="16BC42B2CAD52B6977E7703E91AD52E2">
<mods:roleTerm id="FE8A9AFA97B59CC30B03C6B119A9640D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D79C7797B3D91DAD77E7FCF7B3B170C0">Sudasinghe, Hiranya</mods:namePart>
</mods:name>
<mods:typeOfResource id="E61EED542DFD716D8B1BD30E84A4B4AD">text</mods:typeOfResource>
<mods:relatedItem id="1AC5F8DDD7727485FA2DF4DE01A2E583" type="host">
<mods:titleInfo id="A20B1214870C2BD2764C014288EC9064">
<mods:title id="7609653B30CFF16784E35A433929351F">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="2388264F6C114F656F7FD71A42884095">
<mods:date id="9E3BDD2FBF162D31BB893EA23E431FD5">2015</mods:date>
<mods:detail id="D352FB862C594FC961A1C74C526C85BE" type="volume">
<mods:number id="A4B9877B0AFDD21A6BA1B7DE090EA543">3936</mods:number>
</mods:detail>
<mods:detail id="1ACF28AF291E269CB791AA61CB129290" type="issue">
<mods:number id="E5636F05940467985D15E127E06935BD">4</mods:number>
</mods:detail>
<mods:extent id="B825BD2F276437CA9A3093CAB06939C7" unit="page">
<mods:start id="AA0EDB1E3239395D3B43187DBE6A2AEE">575</mods:start>
<mods:end id="823FD6AA14E62D6329B9541D880C87CC">583</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="C8688329B997224A2BCBE9DBFBA6E0B0">journal article</mods:classification>
<mods:identifier id="81CC9AC22D24B3F3E73C396AB6DB4114" type="DOI">10.11646/zootaxa.3936.4.7</mods:identifier>
<mods:identifier id="BF4C3EAC6BF04469E80859059E4E073A" type="GBIF-Dataset">eae1b1d0-0ba9-4a39-b10c-8a8f5ba02085</mods:identifier>
<mods:identifier id="843155B5AB74405FC358DEC2A8E2FE4B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="188E12BA0679F6F06E437AFBEA1F0783" type="Zenodo-Dep">244127</mods:identifier>
<mods:identifier id="8D70BEE17738484B87241BD4008DDF3A" type="ZooBank">9F14082D-3680-4D75-B3F3-398F912ECCDD</mods:identifier>
</mods:mods>
<treatment id="03E387A1FFC4FFD2FF1BFE1CD004FF72" ID-DOI="http://doi.org/10.5281/zenodo.5693636" ID-GBIF-Taxon="119601828" ID-Zenodo-Dep="5693636" LSID="urn:lsid:plazi:treatment:03E387A1FFC4FFD2FF1BFE1CD004FF72" httpUri="http://treatment.plazi.org/id/03E387A1FFC4FFD2FF1BFE1CD004FF72" lastPageId="5" lastPageNumber="580" pageId="1" pageNumber="576">
<subSubSection id="C350653CFFC4FFD6FF1BFE1CD542FD92" pageId="1" pageNumber="576" type="nomenclature">
<paragraph id="8BF536B7FFC4FFD6FF1BFE1CD6F0FE73" blockId="1.[151,705,475,535]" box="[151,705,475,502]" pageId="1" pageNumber="576">
<heading id="D0BD81DBFFC4FFD6FF1BFE1CD6F0FE73" bold="true" box="[151,705,475,502]" fontSize="11" level="1" pageId="1" pageNumber="576" reason="1">
<taxonomicName id="4C4A4D34FFC4FFD6FF1BFE1CD6F0FE73" ID-CoL="4FDCV" authority="Deraniyagala, 1956" authorityName="Deraniyagala" authorityYear="1956" box="[151,705,475,502]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="576" phylum="Chordata" rank="species" species="melanomaculata">
<emphasis id="B93EEAA5FFC4FFD6FF1BFE1CD6F0FE73" bold="true" box="[151,705,475,502]" pageId="1" pageNumber="576">
<emphasis id="B93EEAA5FFC4FFD6FF1BFE1CD587FE70" bold="true" box="[151,438,475,501]" italics="true" pageId="1" pageNumber="576">Pethia melanomaculata</emphasis>
(
<bibRefCitation id="EFDB4B46FFC4FFD6FE49FE1BD688FE73" author="Deraniyagala" box="[453,697,476,502]" pageId="1" pageNumber="576" refString="Deraniyagala, P. E. P. (1956) Two new subspecies and one new species of cyprinoid fishes from Ceylon. Proceedings of the 12 th Annual Sessions of the Ceylon Association for the Advancement of Science, 1, 34 - 35." type="journal article" year="1956">Deraniyagala, 1956</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BF536B7FFC4FFD6FF1BFE39D542FD92" blockId="1.[151,705,475,535]" box="[151,371,510,535]" pageId="1" pageNumber="576">
(
<figureCitation id="13712A32FFC4FFD6FF12FE39D52EFD92" box="[158,287,510,535]" captionStart="FIGURE 1. A" captionStartId="2.[151,250,1896,1918]" captionTargetBox="[200,1380,231,1850]" captionTargetId="figure@2.[200,1381,231,1852]" captionTargetPageId="2" captionText="FIGURE 1. A, Pethia melanomaculata, colour in life (not preserved); B, copy of iconotype of Cyprinus ticto Hamilton, 1822 (Courtesy Linnean Society of London); C, illustration of Puntius ticto, from Day (1878, pl. 144, fig. 7); D, distribution of Pethia species in Sri Lanka: ● Pethia melanomaculata (location of holotype in red); other known locations (black); {■}, Pethia bandula (purple); P. cumingii (yellow); P. nigrofasciata (green); P. reval (blue)." httpUri="https://zenodo.org/record/244128/files/figure.png" pageId="1" pageNumber="576">Figures 1A</figureCitation>
,
<figureCitation id="13712A32FFC4FFD6FEA7FE38D50AFD92" box="[299,315,511,535]" captionStart="FIGURE 2. A" captionStartId="3.[151,250,1924,1946]" captionTargetBox="[206,1375,209,1879]" captionTargetId="figure@3.[195,1391,193,1896]" captionTargetPageId="3" captionText="FIGURE 2. A, left dentary (buccal view; CP, coronoid process; AA, anguloarticular); B, left premaxilla (dorsal view); C, left maxilla (buccal view; PP, palatine process); D, left infraorbitals 3 + 4 (lateral view); E, 5 th ceratobranchial: of Pethia melanomaculata (WHT 11066, 32.8 mm SL), P. punctata (WHT 11078, 34.9 mm SL) and P. ticto (WHT 11054, 37.0 mm SL). Scale bar: 1 mm." httpUri="https://zenodo.org/record/244129/files/figure.png" pageId="1" pageNumber="576">2</figureCitation>
,
<figureCitation id="13712A32FFC4FFD6FECBFE38D565FD92" box="[327,340,511,535]" captionStart="FIGURE 3" captionStartId="4.[151,250,627,649]" captionTargetBox="[176,1397,198,599]" captionTargetId="figure@4.[167,1419,193,605]" captionTargetPageId="4" captionText="FIGURE 3. Position of caudal spot in A, Pethia melanomaculata; B, P. punctata; and C, P. ticto. Scale bar: 1 mm." httpUri="https://zenodo.org/record/244130/files/figure.png" pageId="1" pageNumber="576">3</figureCitation>
A)
</paragraph>
</subSubSection>
<subSubSection id="C350653CFFC4FFD6FF1BFD8FD7A7FD52" pageId="1" pageNumber="576" type="reference_group">
<paragraph id="8BF536B7FFC4FFD6FF1BFD8FD70FFDD9" blockId="1.[151,1194,582,727]" box="[151,830,582,605]" pageId="1" pageNumber="576">
<treatmentCitationGroup id="AB5A1199FFC4FFD6FF1BFD8FD70FFDD9" box="[151,830,582,605]" pageId="1" pageNumber="576">
<taxonomicName id="4C4A4D34FFC4FFD6FF1BFD8FD6F7FDD9" ID-CoL="4QJ68" authority="Deraniyagala, 1956: 34" authorityName="Deraniyagala" authorityPageNumber="34" authorityYear="1956" box="[151,710,582,605]" class="Actinopterygii" family="Cyprinidae" genus="Puntius" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="576" phylum="Chordata" rank="subSpecies" species="ticto" subSpecies="melanomaculatus">
<emphasis id="B93EEAA5FFC4FFD6FF1BFD8FD5E0FDD9" box="[151,465,582,605]" italics="true" pageId="1" pageNumber="576">Puntius ticto melanomaculatus</emphasis>
<treatmentCitation id="0AEB10A6FFC4FFD6FE5BFD81D6F7FDD9" author="Deraniyagala" box="[471,710,582,604]" page="34" pageId="1" pageNumber="576" year="1956">
<bibRefCitation id="EFDB4B46FFC4FFD6FE5BFD81D691FDD9" author="Deraniyagala" box="[471,672,582,604]" pageId="1" pageNumber="576" refString="Deraniyagala, P. E. P. (1956) Two new subspecies and one new species of cyprinoid fishes from Ceylon. Proceedings of the 12 th Annual Sessions of the Ceylon Association for the Advancement of Science, 1, 34 - 35." type="journal article" year="1956">Deraniyagala, 1956</bibRefCitation>
: 34
</treatmentCitation>
</taxonomicName>
; 1958: 129.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF536B7FFC4FFD6FF1BFDA2D7A6FDFE" blockId="1.[151,1194,582,727]" box="[151,919,613,635]" pageId="1" pageNumber="576">
<treatmentCitationGroup id="AB5A1199FFC4FFD6FF1BFDA2D7A6FDFE" box="[151,919,613,635]" pageId="1" pageNumber="576">
<taxonomicName id="4C4A4D34FFC4FFD6FF1BFDA2D522FDFE" ID-CoL="KNY4" box="[151,275,613,635]" class="Actinopterygii" family="Cyprinidae" genus="Barbus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="576" phylum="Chordata" rank="species" species="ticto">
<emphasis id="B93EEAA5FFC4FFD6FF1BFDA2D522FDFE" box="[151,275,613,635]" italics="true" pageId="1" pageNumber="576">Barbus ticto</emphasis>
</taxonomicName>
(not
<bibRefCitation id="EFDB4B46FFC4FFD6FEC5FDA2D5DDFDFE" author="Hamilton" box="[329,492,613,635]" pageId="1" pageNumber="576" refString="Hamilton, F. (1822) An Account of the Fishes of River Ganges and its Branches. George Ramsay and Co., London, vii + 405 pp., 39 pls." type="book" year="1822">Hamilton, 1822</bibRefCitation>
):
<treatmentCitation id="0AEB10A6FFC4FFD6FE71FDA1D6ADFDFE" author="Day" box="[509,668,613,635]" page="576" pageId="1" pageNumber="576" year="1878">
<bibRefCitation id="EFDB4B46FFC4FFD6FE71FDA1D658FDFE" author="Day" box="[509,617,613,635]" pageId="1" pageNumber="576" refString="Day, F. (1878) Fishes of India. Part 4. Quaritch, London, pp. i - xx, 553 - 778, pls. 134 - 195." type="book chapter" year="1878">Day, 1878</bibRefCitation>
: 576
</treatmentCitation>
;
<treatmentCitation id="0AEB10A6FFC4FFD6FD2AFDA2D7A2FDFE" author="Senanayake" box="[678,915,613,635]" page="118" pageId="1" pageNumber="576" year="1980">
<bibRefCitation id="EFDB4B46FFC4FFD6FD2AFDA2D750FDFE" author="Senanayake" box="[678,865,613,635]" pageId="1" pageNumber="576" refString="Senanayake, F. R. (1980) The biogeography and ecology of the inland fishes of Sri Lanka. Unpublished Ph. D. dissertation, Department of Wildlife and Fisheries Biology, University of California, Davis. [unkown total page number]" type="book" year="1980">Senanayake, 1980</bibRefCitation>
: 118
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF536B7FFC4FFD6FF1BFD42D7A7FD52" blockId="1.[151,1194,582,727]" pageId="1" pageNumber="576">
<treatmentCitationGroup id="AB5A1199FFC4FFD6FF1BFD42D7A7FD52" pageId="1" pageNumber="576">
<taxonomicName id="4C4A4D34FFC4FFD6FF1BFD42D529FD1F" ID-CoL="6WPFT" box="[151,280,645,666]" class="Actinopterygii" family="Cyprinidae" genus="Puntius" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="576" phylum="Chordata" rank="species" species="ticto">
<emphasis id="B93EEAA5FFC4FFD6FF1BFD42D529FD1F" box="[151,280,645,666]" italics="true" pageId="1" pageNumber="576">Puntius ticto</emphasis>
</taxonomicName>
(not
<bibRefCitation id="EFDB4B46FFC4FFD6FEC1FD43D5C1FD1F" author="Hamilton" box="[333,496,644,666]" pageId="1" pageNumber="576" refString="Hamilton, F. (1822) An Account of the Fishes of River Ganges and its Branches. George Ramsay and Co., London, vii + 405 pp., 39 pls." type="book" year="1822">Hamilton, 1822</bibRefCitation>
):
<treatmentCitation id="0AEB10A6FFC4FFD6FD8EFD43D6C0FD1F" author="Deraniyagala" box="[514,753,644,666]" page="35" pageId="1" pageNumber="576" year="1952">
<bibRefCitation id="EFDB4B46FFC4FFD6FD8EFD43D6FAFD1F" author="Deraniyagala" box="[514,715,644,666]" pageId="1" pageNumber="576" refString="Deraniyagala, P. E. P. (1952) A coloured atlas of some vertebrates from Ceylon. Vol. 1. Fishes. National Museums of Ceylon, Colombo, 149 pp., 34 pls." type="book" year="1952">Deraniyagala, 1952</bibRefCitation>
: 35
</treatmentCitation>
;
<treatmentCitation id="0AEB10A6FFC4FFD6FD70FD42D798FD1F" author="Munro" box="[764,937,644,666]" page="46" pageId="1" pageNumber="576" year="1955">
<bibRefCitation id="EFDB4B46FFC4FFD6FD70FD42D7B2FD1F" author="Munro" box="[764,899,644,666]" pageId="1" pageNumber="576" refString="Munro, I. S. R. (1955) The Marine and Freshwater Fishes of Ceylon. Department of External Affairs, Canberra, 349 pp." type="book" year="1955">Munro, 1955</bibRefCitation>
: 46
</treatmentCitation>
; Pethiyagoda, 1991: 118.
<taxonomicName id="4C4A4D34FFC4FFD6FF1BFD64D64EFD3D" ID-CoL="4QJ68" authority="Deraniyagala, 1956" authorityName="Deraniyagala" authorityYear="1956" box="[151,639,674,696]" class="Actinopterygii" family="Cyprinidae" genus="Puntius" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="576" phylum="Chordata" rank="species" species="melanomaculatus">
<emphasis id="B93EEAA5FFC4FFD6FF1BFD64D5AFFD3D" box="[151,414,674,696]" italics="true" pageId="1" pageNumber="576">Puntius melanomaculatus</emphasis>
(
<bibRefCitation id="EFDB4B46FFC4FFD6FE26FD65D647FD3D" author="Deraniyagala" box="[426,630,674,696]" pageId="1" pageNumber="576" refString="Deraniyagala, P. E. P. (1956) Two new subspecies and one new species of cyprinoid fishes from Ceylon. Proceedings of the 12 th Annual Sessions of the Ceylon Association for the Advancement of Science, 1, 34 - 35." type="journal article" year="1956">Deraniyagala, 1956</bibRefCitation>
)
</taxonomicName>
:
<treatmentCitation id="0AEB10A6FFC4FFD6FD04FD65D7EEFD3D" author="Meegaskumbura" box="[648,991,674,696]" page="148" pageId="1" pageNumber="576" year="2008">
<bibRefCitation id="EFDB4B46FFC4FFD6FD04FD65D79DFD3D" author="Meegaskumbura" box="[648,940,674,696]" pageId="1" pageNumber="576" refString="Meegaskumbura, M., Silva, A., Maduwage, K. &amp; Pethiyagoda, R. (2008) Puntius reval, a new barb from Sri Lanka (Teleostei: Cyprinidae). Ichthyological Exploration of Freshwaters, 19, 141 - 152." type="journal article" year="2008">
Meegaskumbura
<emphasis id="B93EEAA5FFC4FFD6FCBBFD64D75BFD3D" box="[823,874,674,696]" italics="true" pageId="1" pageNumber="576">et al.</emphasis>
, 2008
</bibRefCitation>
: 148
</treatmentCitation>
.
<taxonomicName id="4C4A4D34FFC4FFD6FF1BFD06D658FD52" ID-CoL="4FDCV" authority="Deraniyagala, 1956" authorityName="Deraniyagala" authorityYear="1956" box="[151,617,705,727]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="576" phylum="Chordata" rank="species" species="melanomaculata">
<emphasis id="B93EEAA5FFC4FFD6FF1BFD06D5B9FD52" box="[151,392,705,727]" italics="true" pageId="1" pageNumber="576">Pethia melanomaculata</emphasis>
(
<bibRefCitation id="EFDB4B46FFC4FFD6FE19FD06D650FD52" author="Deraniyagala" box="[405,609,705,727]" pageId="1" pageNumber="576" refString="Deraniyagala, P. E. P. (1956) Two new subspecies and one new species of cyprinoid fishes from Ceylon. Proceedings of the 12 th Annual Sessions of the Ceylon Association for the Advancement of Science, 1, 34 - 35." type="journal article" year="1956">Deraniyagala, 1956</bibRefCitation>
)
</taxonomicName>
:
<treatmentCitation id="0AEB10A6FFC4FFD6FDFEFD06D7A0FD52" author="Pethiyagoda" box="[626,913,705,727]" page="71" pageId="1" pageNumber="576" year="2012">
<bibRefCitation id="EFDB4B46FFC4FFD6FDFEFD06D75DFD52" author="Pethiyagoda" box="[626,876,705,727]" pageId="1" pageNumber="576" refString="Pethiyagoda, R., Meegaskumbura, M. &amp; Maduwage, K. (2012) A synopsis of the South Asian fishes referred to Puntius (Pisces: Cyprinidae). Ichthyological Exploration of Freshwaters, 23, 69 - 95." type="journal article" year="2012">
Pethiyagoda
<emphasis id="B93EEAA5FFC4FFD6FD7AFD05D718FD52" box="[758,809,705,727]" italics="true" pageId="1" pageNumber="576">et al.</emphasis>
, 2012
</bibRefCitation>
: 71
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C350653CFFC4FFD6FF1BFCC4D629FBB8" pageId="1" pageNumber="576" type="diagnosis">
<paragraph id="8BF536B7FFC4FFD6FF1BFCC4D629FBB8" blockId="1.[151,1437,771,2021]" pageId="1" pageNumber="576">
<emphasis id="B93EEAA5FFC4FFD6FF1BFCC4D520FC99" bold="true" box="[151,273,771,796]" pageId="1" pageNumber="576">Diagnosis.</emphasis>
<taxonomicName id="4C4A4D34FFC4FFD6FE94FCC3D615FC99" box="[280,548,772,796]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="576" phylum="Chordata" rank="species" species="melanomaculata">
<emphasis id="B93EEAA5FFC4FFD6FE94FCC3D615FC99" box="[280,548,772,796]" italics="true" pageId="1" pageNumber="576">Pethia melanomaculata</emphasis>
</taxonomicName>
is distinguished from all the other congeners in
<collectingCountry id="F35D7627FFC4FFD6FBCFFCC3D080FC98" box="[1091,1201,772,797]" name="Sri Lanka" pageId="1" pageNumber="576">Sri Lanka</collectingCountry>
and the peninsula of
<collectingCountry id="F35D7627FFC4FFD6FF1BFCE0D4E0FCC5" box="[151,209,807,832]" name="India" pageId="1" pageNumber="576">India</collectingCountry>
by the combination of the following characters: barbels absent; lateral line incomplete, with 710 perforated scales; 2023 scales in lateral series; 3 unbranched dorsal-fin rays, the last one with 811 strong serrae; ½4/1/3½ scales in transverse line on body; ½2/1/2½ scales in transverse line on caudal peduncle; predorsal scales 89; body depth 32.441.5% SL; head length 26.129.2% SL; snout length 25.335.6% HL; eye diameter 24.431.9% HL; a small black humeral spot on the 3rd or 4th lateral-line scales; a black spot on caudal peduncle, on scales 1618 of the lateral-line series; no prominent projection at postero-lateral border of premaxilla; superior angle of palatine process of maxilla blunt; dorsal border of anguloarticular curved, with a deep concavity; lateral border of 5th ceratobranchial with a deep notch.
</paragraph>
</subSubSection>
<subSubSection id="C350653CFFC4FFD3FF4BFB80D599F86B" lastPageId="4" lastPageNumber="579" pageId="1" pageNumber="576" type="description">
<paragraph id="8BF536B7FFC4FFD6FF4BFB80D0F1FB75" blockId="1.[151,1437,771,2021]" pageId="1" pageNumber="576">
<emphasis id="B93EEAA5FFC4FFD6FF4BFB80D568FBE5" bold="true" box="[199,345,1095,1120]" pageId="1" pageNumber="576">Description.</emphasis>
See
<figureCitation id="13712A32FFC4FFD6FE2CFB80D5CFFBE5" box="[416,510,1095,1120]" captionStart="FIGURE 1. A" captionStartId="2.[151,250,1896,1918]" captionTargetBox="[200,1380,231,1850]" captionTargetId="figure@2.[200,1381,231,1852]" captionTargetPageId="2" captionText="FIGURE 1. A, Pethia melanomaculata, colour in life (not preserved); B, copy of iconotype of Cyprinus ticto Hamilton, 1822 (Courtesy Linnean Society of London); C, illustration of Puntius ticto, from Day (1878, pl. 144, fig. 7); D, distribution of Pethia species in Sri Lanka: ● Pethia melanomaculata (location of holotype in red); other known locations (black); {■}, Pethia bandula (purple); P. cumingii (yellow); P. nigrofasciata (green); P. reval (blue)." httpUri="https://zenodo.org/record/244128/files/figure.png" pageId="1" pageNumber="576">Fig. 1A</figureCitation>
for general appearance and
<tableCitation id="C6C8030CFFC4FFD6FCE9FB80D7F0FBE5" box="[869,961,1095,1120]" captionStart="TABLE 1" captionStartId="4.[151,239,689,712]" captionTargetBox="[159,1246,771,1833]" captionTargetPageId="4" captionText="TABLE 1. Morphometric data for Pethia melanomaculata, holotype, NMSL FF 728 / A; and WHT 7719 and WHT 7748 (n = 13)." httpUri="http://table.plazi.org/id/DF35663FFFC1FFD3FF1BFD76D4D6FD6D" pageId="1" pageNumber="576" tableUuid="DF35663FFFC1FFD3FF1BFD76D4D6FD6D">Table 1</tableCitation>
for morphometric data. Body laterally compressed. Dorsal profile of head gently notched above posterior orbital margin. Predorsal profile convex, rising gradually to dorsal-fin origin. Postdorsal profile sloping down to hypural notch. Ventral profile convex up to pelvic-fin base, then almost flat to anal-fin origin, rising markedly between anal-fin origin and insertion, then approximately level to hypural notch. Caudal peduncle short, its depth 1.01.4 times its length.
</paragraph>
<paragraph id="8BF536B7FFC4FFD6FF4BFB3BD6FDFAD8" blockId="1.[151,1437,771,2021]" pageId="1" pageNumber="576">Head small, laterally compressed. Snout blunt, rounded, subequal to eye diameter. Mouth small, subterminal, angle of gape not reaching vertical from anterior margin of eye. Barbels absent. Eyes large, dorsolaterally oriented, closer to tip of snout than to margin of operculum.</paragraph>
<paragraph id="8BF536B7FFC4FFD6FF4BFAA0D04EF9B0" blockId="1.[151,1437,771,2021]" pageId="1" pageNumber="576">Dorsal-fin with its origin posterior to pelvic-fin origin, approximately equidistant from tip of snout and caudalfin base; its distal margin concave; 3 unbranched and 8 branched rays (18), last one strong with 8 (8), 9 (4), 10 (4) or 11 (1) serrae posteriorly. Pectoral fin with 1 unbranched and 11 (2) or 12 (15) branched rays, its tip reaching pelvic-fin base when adpressed. Pelvic fin with 1 unbranched and 8 (16) branched rays, its distal margin rounded, not reaching anal-fin origin when adpressed. Anal fin with 3 unbranched and 5 (17) branched rays, its distal margin concave. Caudal fin forked, with i,8,8,i (1), i,8,9,i (5), i,9,8,i (4), i,9,9,i (7) principal rays.</paragraph>
<paragraph id="8BF536B7FFC4FFD6FF4BF9F8D6F1F925" blockId="1.[151,1437,771,2021]" pageId="1" pageNumber="576">Lateral line incomplete, with 710 pored scales; 20 (4), 21 (9), 22 (3) or 23 (2) scales in lateral series. Scales in transverse line on body ½4/1/3½ (18), predorsal scales 8 (16) or 9 (1), prepelvic scales 9 (3) or 10 (2), scales in transverse line on caudal peduncle ½2/1/2½ (18).</paragraph>
<paragraph id="8BF536B7FFC4FFD6FF4BF96BD149F8D0" blockId="1.[151,1437,771,2021]" pageId="1" pageNumber="576">Precaudal vertebrae 16; caudal vertebrae 14. Supraneurals 4. No prominent projection at postero-lateral border of premaxilla. Superior angle of palatine process of maxilla blunt. Lengths of anterior and superior articular borders of infraorbital 3+4 less than half of its width. Tip of coronoid process of dentary blunt. Dorsal border of anguloarticular curved, with deep concavity. Lateral arm of 5th ceratobranchial narrow, pointed, with a small foramen at its base. Fifth ceratobranchial with a deep notch on its lateral border and 5+3+2 small, conical teeth.</paragraph>
<paragraph id="8BF536B7FFC4FFD6FF4BF898D6B4F860" blockId="1.[151,1437,771,2021]" pageId="1" pageNumber="576">
<emphasis id="B93EEAA5FFC4FFD6FF4BF898D57EF8FD" bold="true" box="[199,335,1887,1912]" pageId="1" pageNumber="576">Coloration.</emphasis>
In preservative, body overall pale brown. A small black humeral spot above pectoral fin, level with mid eye, confined to 3rd or 4th lateral-line scale. Black spot on caudal peduncle larger than humeral spot, somewhat smaller than eye diameter, originating posterior to anal-fin insertion, on scales 1618 of lateral series. All fins hyaline, without any distinct marks.
</paragraph>
<caption id="DF35663FFFC7FFD5FF1BF8AFD78AF85F" httpUri="https://zenodo.org/record/244128/files/figure.png" pageId="2" pageNumber="577" targetBox="[200,1380,231,1850]" targetPageId="2">
<paragraph id="8BF536B7FFC7FFD5FF1BF8AFD78AF85F" blockId="2.[151,1436,1896,2010]" pageId="2" pageNumber="577">
<emphasis id="B93EEAA5FFC7FFD5FF1BF8AFD503F8F8" bold="true" box="[151,306,1896,1918]" pageId="2" pageNumber="577">FIGURE 1. A,</emphasis>
<taxonomicName id="4C4A4D34FFC7FFD5FEB6F8AFD61DF8FB" box="[314,556,1896,1918]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="577" phylum="Chordata" rank="species" species="melanomaculata">
<emphasis id="B93EEAA5FFC7FFD5FEB6F8AFD61DF8FB" box="[314,556,1896,1918]" italics="true" pageId="2" pageNumber="577">Pethia melanomaculata</emphasis>
</taxonomicName>
, colour in life (not preserved);
<emphasis id="B93EEAA5FFC7FFD5FCE0F8AFD7B3F8F8" bold="true" box="[876,898,1896,1917]" pageId="2" pageNumber="577">B,</emphasis>
copy of iconotype of
<taxonomicName id="4C4A4D34FFC7FFD5FBE9F8AED1ADF8FB" authority="Hamilton, 1822" authorityName="Hamilton" authorityYear="1822" box="[1125,1436,1896,1918]" class="Actinopterygii" family="Cyprinidae" genus="Cyprinus" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="577" phylum="Chordata" rank="species" species="ticto">
<emphasis id="B93EEAA5FFC7FFD5FBE9F8AED0C4F8FB" box="[1125,1269,1897,1918]" italics="true" pageId="2" pageNumber="577">Cyprinus ticto</emphasis>
Hamilton, 1822
</taxonomicName>
(Courtesy Linnean Society of London);
<emphasis id="B93EEAA5FFC7FFD5FDB7F841D665F819" bold="true" box="[571,596,1926,1948]" pageId="2" pageNumber="577">C,</emphasis>
illustration of
<taxonomicName id="4C4A4D34FFC7FFD5FD7EF84FD744F818" box="[754,885,1928,1949]" class="Actinopterygii" family="Cyprinidae" genus="Puntius" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="577" phylum="Chordata" rank="species" species="ticto">
<emphasis id="B93EEAA5FFC7FFD5FD7EF84FD744F818" box="[754,885,1928,1949]" italics="true" pageId="2" pageNumber="577">Puntius ticto</emphasis>
</taxonomicName>
, from Day (1878, pl. 144, fig. 7); D, distribution of
<taxonomicName id="4C4A4D34FFC7FFD5FF1BF862D4E8F83E" box="[151,217,1957,1979]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="577" phylum="Chordata" rank="genus">
<emphasis id="B93EEAA5FFC7FFD5FF1BF862D4E8F83E" box="[151,217,1957,1979]" italics="true" pageId="2" pageNumber="577">Pethia</emphasis>
</taxonomicName>
species in Sri Lanka: ●
<taxonomicName id="4C4A4D34FFC7FFD5FE41F862D68FF83E" box="[461,702,1957,1979]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="577" phylum="Chordata" rank="species" species="melanomaculata">
<emphasis id="B93EEAA5FFC7FFD5FE41F862D68FF83E" box="[461,702,1957,1979]" italics="true" pageId="2" pageNumber="577">Pethia melanomaculata</emphasis>
</taxonomicName>
(location of holotype in red); other known locations (black); {■},
<taxonomicName id="4C4A4D34FFC7FFD5FAD6F862D4DBF85F" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="577" phylum="Chordata" rank="species" species="bandula">
<emphasis id="B93EEAA5FFC7FFD5FAD6F862D4DBF85F" italics="true" pageId="2" pageNumber="577">Pethia bandula</emphasis>
</taxonomicName>
(purple);
<taxonomicName id="4C4A4D34FFC7FFD5FEC3F802D5F0F85F" box="[335,449,1989,2010]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="577" phylum="Chordata" rank="species" species="cumingii">
<emphasis id="B93EEAA5FFC7FFD5FEC3F802D5F0F85F" box="[335,449,1989,2010]" italics="true" pageId="2" pageNumber="577">P. cumingii</emphasis>
</taxonomicName>
(yellow);
<taxonomicName id="4C4A4D34FFC7FFD5FDA6F802D6F8F85F" box="[554,713,1988,2010]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="577" phylum="Chordata" rank="species" species="nigrofasciata">
<emphasis id="B93EEAA5FFC7FFD5FDA6F802D6F8F85F" box="[554,713,1988,2010]" italics="true" pageId="2" pageNumber="577">P. nigrofasciata</emphasis>
</taxonomicName>
(green);
<taxonomicName id="4C4A4D34FFC7FFD5FCA9F802D740F85F" box="[805,881,1988,2010]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="577" phylum="Chordata" rank="species" species="reval">
<emphasis id="B93EEAA5FFC7FFD5FCA9F802D740F85F" box="[805,881,1988,2010]" italics="true" pageId="2" pageNumber="577">P. reval</emphasis>
</taxonomicName>
(blue).
</paragraph>
</caption>
<caption id="DF35663FFFC6FFD4FF1BF843D572F873" httpUri="https://zenodo.org/record/244129/files/figure.png" pageId="3" pageNumber="578" targetBox="[206,1375,209,1879]" targetPageId="3">
<paragraph id="8BF536B7FFC6FFD4FF1BF843D572F873" blockId="3.[151,1436,1924,2038]" pageId="3" pageNumber="578">
<emphasis id="B93EEAA5FFC6FFD4FF1BF843D503F81C" bold="true" box="[151,306,1924,1946]" pageId="3" pageNumber="578">FIGURE 2. A,</emphasis>
left dentary (buccal view; CP, coronoid process; AA, anguloarticular);
<emphasis id="B93EEAA5FFC6FFD4FB80F843D013F81C" bold="true" box="[1036,1058,1924,1945]" pageId="3" pageNumber="578">B,</emphasis>
left premaxilla (dorsal view);
<emphasis id="B93EEAA5FFC6FFD4FAD7F843D145F81F" bold="true" box="[1371,1396,1924,1946]" pageId="3" pageNumber="578">C,</emphasis>
left maxilla (buccal view; PP, palatine process);
<emphasis id="B93EEAA5FFC6FFD4FD0CF864D6A9F83D" bold="true" box="[640,664,1955,1976]" pageId="3" pageNumber="578">D,</emphasis>
left infraorbitals 3+4 (lateral view);
<emphasis id="B93EEAA5FFC6FFD4FBB9F864D07DF83D" bold="true" box="[1077,1100,1955,1976]" pageId="3" pageNumber="578">E,</emphasis>
5th ceratobranchial: of
<taxonomicName id="4C4A4D34FFC6FFD4FAD6F865D571F852" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="578" phylum="Chordata" rank="species" species="melanomaculata">
<emphasis id="B93EEAA5FFC6FFD4FAD6F865D571F852" italics="true" pageId="3" pageNumber="578">Pethia melanomaculata</emphasis>
</taxonomicName>
(WHT 11066, 32.8 mm SL),
<taxonomicName id="4C4A4D34FFC6FFD4FDE0F805D6ECF852" box="[620,733,1986,2007]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="578" phylum="Chordata" rank="species" species="punctata">
<emphasis id="B93EEAA5FFC6FFD4FDE0F805D6ECF852" box="[620,733,1986,2007]" italics="true" pageId="3" pageNumber="578">P. punctata</emphasis>
</taxonomicName>
(WHT 11078, 34.9 mm SL) and
<taxonomicName id="4C4A4D34FFC6FFD4FBA3F805D044F852" box="[1071,1141,1986,2007]" class="Actinopterygii" family="Cyprinidae" genus="Puntius" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="578" phylum="Chordata" rank="species" species="ticto">
<emphasis id="B93EEAA5FFC6FFD4FBA3F805D044F852" box="[1071,1141,1986,2007]" italics="true" pageId="3" pageNumber="578">P. ticto</emphasis>
</taxonomicName>
(WHT 11054, 37.0 mm SL). Scale bar: 1 mm.
</paragraph>
</caption>
<caption id="DF35663FFFC1FFD3FF1BFDB4D112FD0C" box="[151,1315,627,649]" httpUri="https://zenodo.org/record/244130/files/figure.png" pageId="4" pageNumber="579" targetBox="[176,1397,198,599]" targetPageId="4">
<paragraph id="8BF536B7FFC1FFD3FF1BFDB4D112FD0C" blockId="4.[151,1315,627,649]" box="[151,1315,627,649]" pageId="4" pageNumber="579">
<emphasis id="B93EEAA5FFC1FFD3FF1BFDB4D523FD0D" bold="true" box="[151,274,627,649]" pageId="4" pageNumber="579">FIGURE 3.</emphasis>
Position of caudal spot in
<emphasis id="B93EEAA5FFC1FFD3FDACFDB4D609FD0D" bold="true" box="[544,568,627,648]" pageId="4" pageNumber="579">A,</emphasis>
<taxonomicName id="4C4A4D34FFC1FFD3FDB3FDB4D700FD0C" box="[575,817,627,649]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="579" phylum="Chordata" rank="species" species="melanomaculata">
<emphasis id="B93EEAA5FFC1FFD3FDB3FDB4D700FD0C" box="[575,817,627,649]" italics="true" pageId="4" pageNumber="579">Pethia melanomaculata</emphasis>
</taxonomicName>
;
<emphasis id="B93EEAA5FFC1FFD3FCB2FDB4D765FD0D" bold="true" box="[830,852,627,648]" pageId="4" pageNumber="579">B,</emphasis>
<taxonomicName id="4C4A4D34FFC1FFD3FCD0FDB3D7FDFD0C" box="[860,972,628,649]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="579" phylum="Chordata" rank="species" species="punctata">
<emphasis id="B93EEAA5FFC1FFD3FCD0FDB3D7FDFD0C" box="[860,972,628,649]" italics="true" pageId="4" pageNumber="579">P. punctata</emphasis>
</taxonomicName>
; and
<emphasis id="B93EEAA5FFC1FFD3FB89FDB4D02FFD0C" bold="true" box="[1029,1054,627,649]" pageId="4" pageNumber="579">C,</emphasis>
<emphasis id="B93EEAA5FFC1FFD3FBA8FDB3D040FD0C" box="[1060,1137,628,649]" italics="true" pageId="4" pageNumber="579">
<taxonomicName id="4C4A4D34FFC1FFD3FBA8FDB3D05CFD0C" box="[1060,1133,628,649]" class="Actinopterygii" family="Cyprinidae" genus="Puntius" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="579" phylum="Chordata" rank="species" species="ticto">P. ticto</taxonomicName>
.
</emphasis>
Scale bar: 1 mm.
</paragraph>
</caption>
<caption id="DF35663FFFC1FFD3FF1BFD76D4D6FD6D" ID-Table-UUID="DF35663FFFC1FFD3FF1BFD76D4D6FD6D" httpUri="http://table.plazi.org/id/DF35663FFFC1FFD3FF1BFD76D4D6FD6D" pageId="4" pageNumber="579" targetBox="[159,1246,771,1833]" targetIsTable="true" targetPageId="4">
<paragraph id="8BF536B7FFC1FFD3FF1BFD76D4D6FD6D" blockId="4.[151,1436,688,744]" pageId="4" pageNumber="579">
<emphasis id="B93EEAA5FFC1FFD3FF1BFD76D53BFD4D" bold="true" box="[151,266,689,712]" pageId="4" pageNumber="579">TABLE 1.</emphasis>
Morphometric data for
<taxonomicName id="4C4A4D34FFC1FFD3FD87FD77D73BFD4D" box="[523,778,688,712]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="579" phylum="Chordata" rank="species" species="melanomaculata">
<emphasis id="B93EEAA5FFC1FFD3FD87FD77D73BFD4D" box="[523,778,688,712]" italics="true" pageId="4" pageNumber="579">Pethia melanomaculata</emphasis>
</taxonomicName>
, holotype, NMSL FF 728/A; and WHT 7719 and WHT 7748 (n=13).
</paragraph>
</caption>
<paragraph id="8BF536B7FFC1FFD3FDA6FCC4D0EFF8AC" pageId="4" pageNumber="579">
<table id="F94AC417FFC10028FF13FCC4D0EFF8AC" box="[159,1246,771,1833]" gridcols="3" gridrows="27" pageId="4" pageNumber="579">
<tr id="357A34F5FFC10028FF13FCC4D0EFFC9C" box="[159,1246,771,793]" gridrow="0" pageId="4" pageNumber="579" rowspan-0="1" rowspan-2="1">
<th id="76AB5D89FFC10028FDA6FCC4D7B6FC9C" box="[554,903,771,793]" gridcol="1" gridrow="0" pageId="4" pageNumber="579">
<taxonomicName id="4C4A4D34FFC1FFD3FDA6FCC4D73EFC9C" box="[554,783,771,793]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="579" phylum="Chordata" rank="species" species="melanomaculata">
<emphasis id="B93EEAA5FFC1FFD3FDA6FCC4D73EFC9C" box="[554,783,771,793]" italics="true" pageId="4" pageNumber="579">Pethia melanomaculata</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="357A34F5FFC10028FF13FCEBD0EFFCC4" box="[159,1246,812,833]" gridrow="1" pageId="4" pageNumber="579" rowspan-0="1">
<td id="76AB5D89FFC10028FDA6FCEBD7B6FCC4" box="[554,903,812,833]" gridcol="1" gridrow="1" pageId="4" pageNumber="579">holotype mean (s.d)</td>
<td id="76AB5D89FFC10028FBF2FCEBD0EFFCC4" box="[1150,1246,812,833]" gridcol="2" gridrow="1" pageId="4" pageNumber="579">range</td>
</tr>
<tr id="357A34F5FFC10028FF13FC93D0EFFCEC" box="[159,1246,852,873]" gridrow="2" pageId="4" pageNumber="579">
<th id="76AB5D89FFC10028FF13FC93D5E8FCEC" box="[159,473,852,873]" gridcol="0" gridrow="2" pageId="4" pageNumber="579">Standard length (mm)</th>
<td id="76AB5D89FFC10028FDA6FC93D7B6FCEC" box="[554,903,852,873]" gridcol="1" gridrow="2" pageId="4" pageNumber="579">37.4 33.8 (2.3)</td>
<td id="76AB5D89FFC10028FBF2FC93D0EFFCEC" box="[1150,1246,852,873]" gridcol="2" gridrow="2" pageId="4" pageNumber="579">31.037.4</td>
</tr>
<tr id="357A34F5FFC10028FF13FCBBD0EFFC14" box="[159,1246,892,913]" gridrow="3" pageId="4" pageNumber="579" rowspan-1="1" rowspan-2="1">
<th id="76AB5D89FFC10028FF13FCBBD5E8FC14" box="[159,473,892,913]" gridcol="0" gridrow="3" pageId="4" pageNumber="579">In percent of standard length</th>
</tr>
<tr id="357A34F5FFC10028FF13FC63D0EFFC3C" box="[159,1246,932,953]" gridrow="4" pageId="4" pageNumber="579">
<th id="76AB5D89FFC10028FF13FC63D5E8FC3C" box="[159,473,932,953]" gridcol="0" gridrow="4" pageId="4" pageNumber="579">Total length</th>
<td id="76AB5D89FFC10028FDA6FC63D7B6FC3C" box="[554,903,932,953]" gridcol="1" gridrow="4" pageId="4" pageNumber="579"> 132 (2.3)</td>
<td id="76AB5D89FFC10028FBF2FC63D0EFFC3C" box="[1150,1246,932,953]" gridcol="2" gridrow="4" pageId="4" pageNumber="579">128136</td>
</tr>
<tr id="357A34F5FFC10028FF13FC0BD0EFFC64" box="[159,1246,972,993]" gridrow="5" pageId="4" pageNumber="579">
<th id="76AB5D89FFC10028FF13FC0BD5E8FC64" box="[159,473,972,993]" gridcol="0" gridrow="5" pageId="4" pageNumber="579">Predorsal length</th>
<td id="76AB5D89FFC10028FDA6FC0BD7B6FC64" box="[554,903,972,993]" gridcol="1" gridrow="5" pageId="4" pageNumber="579">45.5 55.0 (3.0)</td>
<td id="76AB5D89FFC10028FBF2FC0BD0EFFC64" box="[1150,1246,972,993]" gridcol="2" gridrow="5" pageId="4" pageNumber="579">45.557.4</td>
</tr>
<tr id="357A34F5FFC10028FF13FC33D0EFFB8C" box="[159,1246,1012,1033]" gridrow="6" pageId="4" pageNumber="579">
<th id="76AB5D89FFC10028FF13FC33D5E8FB8C" box="[159,473,1012,1033]" gridcol="0" gridrow="6" pageId="4" pageNumber="579">Postdorsal length</th>
<td id="76AB5D89FFC10028FDA6FC33D7B6FB8C" box="[554,903,1012,1033]" gridcol="1" gridrow="6" pageId="4" pageNumber="579">50.8 54.5 (2.4)</td>
<td id="76AB5D89FFC10028FBF2FC33D0EFFB8C" box="[1150,1246,1012,1033]" gridcol="2" gridrow="6" pageId="4" pageNumber="579">50.858.0</td>
</tr>
<tr id="357A34F5FFC10028FF13FBDBD0EFFBB4" box="[159,1246,1052,1073]" gridrow="7" pageId="4" pageNumber="579">
<th id="76AB5D89FFC10028FF13FBDBD5E8FBB4" box="[159,473,1052,1073]" gridcol="0" gridrow="7" pageId="4" pageNumber="579">Preanal length</th>
<td id="76AB5D89FFC10028FDA6FBDBD7B6FBB4" box="[554,903,1052,1073]" gridcol="1" gridrow="7" pageId="4" pageNumber="579">65.5 69.9 (2.0)</td>
<td id="76AB5D89FFC10028FBF2FBDBD0EFFBB4" box="[1150,1246,1052,1073]" gridcol="2" gridrow="7" pageId="4" pageNumber="579">65.572.6</td>
</tr>
<tr id="357A34F5FFC10028FF13FB83D0EFFBDC" box="[159,1246,1092,1113]" gridrow="8" pageId="4" pageNumber="579">
<th id="76AB5D89FFC10028FF13FB83D5E8FBDC" box="[159,473,1092,1113]" gridcol="0" gridrow="8" pageId="4" pageNumber="579">Prepelvic length</th>
<td id="76AB5D89FFC10028FDA6FB83D7B6FBDC" box="[554,903,1092,1113]" gridcol="1" gridrow="8" pageId="4" pageNumber="579">46.8 47.5 (1.6)</td>
<td id="76AB5D89FFC10028FBF2FB83D0EFFBDC" box="[1150,1246,1092,1113]" gridcol="2" gridrow="8" pageId="4" pageNumber="579">44.550.3</td>
</tr>
<tr id="357A34F5FFC10028FF13FBABD0EFFB04" box="[159,1246,1132,1153]" gridrow="9" pageId="4" pageNumber="579">
<th id="76AB5D89FFC10028FF13FBABD5E8FB04" box="[159,473,1132,1153]" gridcol="0" gridrow="9" pageId="4" pageNumber="579">Body depth</th>
<td id="76AB5D89FFC10028FDA6FBABD7B6FB04" box="[554,903,1132,1153]" gridcol="1" gridrow="9" pageId="4" pageNumber="579">32.4 38.3 (2.3)</td>
<td id="76AB5D89FFC10028FBF2FBABD0EFFB04" box="[1150,1246,1132,1153]" gridcol="2" gridrow="9" pageId="4" pageNumber="579">32.441.5</td>
</tr>
<tr id="357A34F5FFC10028FF13FB53D0EFFB2C" box="[159,1246,1172,1193]" gridrow="10" pageId="4" pageNumber="579">
<th id="76AB5D89FFC10028FF13FB53D5E8FB2C" box="[159,473,1172,1193]" gridcol="0" gridrow="10" pageId="4" pageNumber="579">Head length</th>
<td id="76AB5D89FFC10028FDA6FB53D7B6FB2C" box="[554,903,1172,1193]" gridcol="1" gridrow="10" pageId="4" pageNumber="579">28.1 27.8 (0.9)</td>
<td id="76AB5D89FFC10028FBF2FB53D0EFFB2C" box="[1150,1246,1172,1193]" gridcol="2" gridrow="10" pageId="4" pageNumber="579">26.129.2</td>
</tr>
<tr id="357A34F5FFC10028FF13FB7BD0EFFB54" box="[159,1246,1212,1233]" gridrow="11" pageId="4" pageNumber="579">
<th id="76AB5D89FFC10028FF13FB7BD5E8FB54" box="[159,473,1212,1233]" gridcol="0" gridrow="11" pageId="4" pageNumber="579">Dorsal-fin base length</th>
<td id="76AB5D89FFC10028FDA6FB7BD7B6FB54" box="[554,903,1212,1233]" gridcol="1" gridrow="11" pageId="4" pageNumber="579">15.5 17.1 (0.8)</td>
<td id="76AB5D89FFC10028FBF2FB7BD0EFFB54" box="[1150,1246,1212,1233]" gridcol="2" gridrow="11" pageId="4" pageNumber="579">15.518.1</td>
</tr>
<tr id="357A34F5FFC10028FF13FB23D0EFFB7C" box="[159,1246,1252,1273]" gridrow="12" pageId="4" pageNumber="579">
<th id="76AB5D89FFC10028FF13FB23D5E8FB7C" box="[159,473,1252,1273]" gridcol="0" gridrow="12" pageId="4" pageNumber="579">Dorsal-fin height</th>
<td id="76AB5D89FFC10028FDA6FB23D7B6FB7C" box="[554,903,1252,1273]" gridcol="1" gridrow="12" pageId="4" pageNumber="579">23.0 25.3 (1.5)</td>
<td id="76AB5D89FFC10028FBF2FB23D0EFFB7C" box="[1150,1246,1252,1273]" gridcol="2" gridrow="12" pageId="4" pageNumber="579">23.028.4</td>
</tr>
<tr id="357A34F5FFC10028FF13FACBD0EFFAA4" box="[159,1246,1292,1313]" gridrow="13" pageId="4" pageNumber="579">
<th id="76AB5D89FFC10028FF13FACBD5E8FAA4" box="[159,473,1292,1313]" gridcol="0" gridrow="13" pageId="4" pageNumber="579">Anal-fin base length</th>
<td id="76AB5D89FFC10028FDA6FACBD7B6FAA4" box="[554,903,1292,1313]" gridcol="1" gridrow="13" pageId="4" pageNumber="579">10.2 11.8 (1.2)</td>
<td id="76AB5D89FFC10028FBF2FACBD0EFFAA4" box="[1150,1246,1292,1313]" gridcol="2" gridrow="13" pageId="4" pageNumber="579">9.714.0</td>
</tr>
<tr id="357A34F5FFC10028FF13FAF3D0EFFACC" box="[159,1246,1332,1353]" gridrow="14" pageId="4" pageNumber="579">
<th id="76AB5D89FFC10028FF13FAF3D5E8FACC" box="[159,473,1332,1353]" gridcol="0" gridrow="14" pageId="4" pageNumber="579">Anal-fin depth</th>
<td id="76AB5D89FFC10028FDA6FAF3D7B6FACC" box="[554,903,1332,1353]" gridcol="1" gridrow="14" pageId="4" pageNumber="579">17.1 17.0 (1.0)</td>
<td id="76AB5D89FFC10028FBF2FAF3D0EFFACC" box="[1150,1246,1332,1353]" gridcol="2" gridrow="14" pageId="4" pageNumber="579">15.718.9</td>
</tr>
<tr id="357A34F5FFC10028FF13FA9BD0EFFAF4" box="[159,1246,1372,1393]" gridrow="15" pageId="4" pageNumber="579">
<th id="76AB5D89FFC10028FF13FA9BD5E8FAF4" box="[159,473,1372,1393]" gridcol="0" gridrow="15" pageId="4" pageNumber="579">Pelvic-fin length</th>
<td id="76AB5D89FFC10028FDA6FA9BD7B6FAF4" box="[554,903,1372,1393]" gridcol="1" gridrow="15" pageId="4" pageNumber="579">18.7 21.6 (1.4)</td>
<td id="76AB5D89FFC10028FBF2FA9BD0EFFAF4" box="[1150,1246,1372,1393]" gridcol="2" gridrow="15" pageId="4" pageNumber="579">18.723.4</td>
</tr>
<tr id="357A34F5FFC10028FF13FA43D0EFFA1C" box="[159,1246,1412,1433]" gridrow="16" pageId="4" pageNumber="579">
<th id="76AB5D89FFC10028FF13FA43D5E8FA1C" box="[159,473,1412,1433]" gridcol="0" gridrow="16" pageId="4" pageNumber="579">Pectoral-fin length</th>
<td id="76AB5D89FFC10028FDA6FA43D7B6FA1C" box="[554,903,1412,1433]" gridcol="1" gridrow="16" pageId="4" pageNumber="579">20.1 20.9 (1.5)</td>
<td id="76AB5D89FFC10028FBF2FA43D0EFFA1C" box="[1150,1246,1412,1433]" gridcol="2" gridrow="16" pageId="4" pageNumber="579">16.822.4</td>
</tr>
<tr id="357A34F5FFC10028FF13FA6BD0EFFA44" box="[159,1246,1452,1473]" gridrow="17" pageId="4" pageNumber="579">
<th id="76AB5D89FFC10028FF13FA6BD5E8FA44" box="[159,473,1452,1473]" gridcol="0" gridrow="17" pageId="4" pageNumber="579">Caudal-peduncle length</th>
<td id="76AB5D89FFC10028FDA6FA6BD7B6FA44" box="[554,903,1452,1473]" gridcol="1" gridrow="17" pageId="4" pageNumber="579">18.7 19.0 (1.9)</td>
<td id="76AB5D89FFC10028FBF2FA6BD0EFFA44" box="[1150,1246,1452,1473]" gridcol="2" gridrow="17" pageId="4" pageNumber="579">16.322.5</td>
</tr>
<tr id="357A34F5FFC10028FF13FA13D0EFFA6C" box="[159,1246,1492,1513]" gridrow="18" pageId="4" pageNumber="579">
<th id="76AB5D89FFC10028FF13FA13D5E8FA6C" box="[159,473,1492,1513]" gridcol="0" gridrow="18" pageId="4" pageNumber="579">Caudal-peduncle depth</th>
<td id="76AB5D89FFC10028FDA6FA13D7B6FA6C" box="[554,903,1492,1513]" gridcol="1" gridrow="18" pageId="4" pageNumber="579">13.1 15.3 (0.9)</td>
<td id="76AB5D89FFC10028FBF2FA13D0EFFA6C" box="[1150,1246,1492,1513]" gridcol="2" gridrow="18" pageId="4" pageNumber="579">13.117.0</td>
</tr>
<tr id="357A34F5FFC10028FF13FA3BD0EFF994" box="[159,1246,1532,1553]" gridrow="19" pageId="4" pageNumber="579">
<th id="76AB5D89FFC10028FF13FA3BD5E8F994" box="[159,473,1532,1553]" gridcol="0" gridrow="19" pageId="4" pageNumber="579">Length of upper caudal-fin lobe</th>
<td id="76AB5D89FFC10028FDA6FA3BD7B6F994" box="[554,903,1532,1553]" gridcol="1" gridrow="19" pageId="4" pageNumber="579"> 32.4 (5.2)</td>
<td id="76AB5D89FFC10028FBF2FA3BD0EFF994" box="[1150,1246,1532,1553]" gridcol="2" gridrow="19" pageId="4" pageNumber="579">27.246.8</td>
</tr>
<tr id="357A34F5FFC10028FF13F9E3D0EFF9BC" box="[159,1246,1572,1593]" gridrow="20" pageId="4" pageNumber="579">
<th id="76AB5D89FFC10028FF13F9E3D5E8F9BC" box="[159,473,1572,1593]" gridcol="0" gridrow="20" pageId="4" pageNumber="579">Length of lower caudal-fin lobe</th>
<td id="76AB5D89FFC10028FDA6F9E3D7B6F9BC" box="[554,903,1572,1593]" gridcol="1" gridrow="20" pageId="4" pageNumber="579"> 33.0 (1.9)</td>
<td id="76AB5D89FFC10028FBF2F9E3D0EFF9BC" box="[1150,1246,1572,1593]" gridcol="2" gridrow="20" pageId="4" pageNumber="579">29.236.7</td>
</tr>
<tr id="357A34F5FFC10028FF13F98BD0EFF9E4" box="[159,1246,1612,1633]" gridrow="21" pageId="4" pageNumber="579">
<th id="76AB5D89FFC10028FF13F98BD5E8F9E4" box="[159,473,1612,1633]" gridcol="0" gridrow="21" pageId="4" pageNumber="579">Length of medial caudal-fin rays</th>
<td id="76AB5D89FFC10028FDA6F98BD7B6F9E4" box="[554,903,1612,1633]" gridcol="1" gridrow="21" pageId="4" pageNumber="579">10.4 14.7 (1.8)</td>
<td id="76AB5D89FFC10028FBF2F98BD0EFF9E4" box="[1150,1246,1612,1633]" gridcol="2" gridrow="21" pageId="4" pageNumber="579">10.416.5</td>
</tr>
<tr id="357A34F5FFC10028FF13F9B3D0EFF90C" box="[159,1246,1652,1673]" gridrow="22" pageId="4" pageNumber="579" rowspan-1="1" rowspan-2="1">
<th id="76AB5D89FFC10028FF13F9B3D5E8F90C" box="[159,473,1652,1673]" gridcol="0" gridrow="22" pageId="4" pageNumber="579">In percent of head length</th>
</tr>
<tr id="357A34F5FFC10028FF13F95BD0EFF934" box="[159,1246,1692,1713]" gridrow="23" pageId="4" pageNumber="579">
<th id="76AB5D89FFC10028FF13F95BD5E8F934" box="[159,473,1692,1713]" gridcol="0" gridrow="23" pageId="4" pageNumber="579">Snout length</th>
<td id="76AB5D89FFC10028FDA6F95BD7B6F934" box="[554,903,1692,1713]" gridcol="1" gridrow="23" pageId="4" pageNumber="579">27.6 29.3 (3.1)</td>
<td id="76AB5D89FFC10028FBF2F95BD0EFF934" box="[1150,1246,1692,1713]" gridcol="2" gridrow="23" pageId="4" pageNumber="579">25.335.6</td>
</tr>
<tr id="357A34F5FFC10028FF13F903D0EFF95C" box="[159,1246,1732,1753]" gridrow="24" pageId="4" pageNumber="579">
<th id="76AB5D89FFC10028FF13F903D5E8F95C" box="[159,473,1732,1753]" gridcol="0" gridrow="24" pageId="4" pageNumber="579">Orbital diameter</th>
<td id="76AB5D89FFC10028FDA6F903D7B6F95C" box="[554,903,1732,1753]" gridcol="1" gridrow="24" pageId="4" pageNumber="579">30.5 29.9 (1.9)</td>
<td id="76AB5D89FFC10028FBF2F903D0EFF95C" box="[1150,1246,1732,1753]" gridcol="2" gridrow="24" pageId="4" pageNumber="579">24.431.9</td>
</tr>
<tr id="357A34F5FFC10028FF13F92BD0EFF884" box="[159,1246,1772,1793]" gridrow="25" pageId="4" pageNumber="579">
<th id="76AB5D89FFC10028FF13F92BD5E8F884" box="[159,473,1772,1793]" gridcol="0" gridrow="25" pageId="4" pageNumber="579">Interorbital width</th>
<td id="76AB5D89FFC10028FDA6F92BD7B6F884" box="[554,903,1772,1793]" gridcol="1" gridrow="25" pageId="4" pageNumber="579">33.3 37.6 (2.0)</td>
<td id="76AB5D89FFC10028FBF2F92BD0EFF884" box="[1150,1246,1772,1793]" gridcol="2" gridrow="25" pageId="4" pageNumber="579">33.339.6</td>
</tr>
<tr id="357A34F5FFC10028FF13F8D3D0EFF8AC" box="[159,1246,1812,1833]" gridrow="26" pageId="4" pageNumber="579">
<th id="76AB5D89FFC10028FF13F8D3D5E8F8AC" box="[159,473,1812,1833]" gridcol="0" gridrow="26" pageId="4" pageNumber="579">Internarial width</th>
<td id="76AB5D89FFC10028FDA6F8D3D7B6F8AC" box="[554,903,1812,1833]" gridcol="1" gridrow="26" pageId="4" pageNumber="579">19.0 18.8 (2.9)</td>
<td id="76AB5D89FFC10028FBF2F8D3D0EFF8AC" box="[1150,1246,1812,1833]" gridcol="2" gridrow="26" pageId="4" pageNumber="579">12.822.0</td>
</tr>
</table>
</paragraph>
<paragraph id="8BF536B7FFC1FFD3FF4BF8A9D599F86B" blockId="4.[151,1437,1901,2030]" pageId="4" pageNumber="579">In life, dorsally light golden brown. Sides of body iridescent silver with scattered melanophores on distal margin of anterior mid-body scales. Position of humeral and caudal spots same as in preserved specimens. Ventrally white. Prominent markings on dorsal fin absent. Fins hyaline to light yellow. Breeding males with a reddish body coloration.</paragraph>
</subSubSection>
<subSubSection id="C350653CFFC0FFD2FF4BFF50D004FF72" pageId="5" pageNumber="580" type="distribution">
<paragraph id="8BF536B7FFC0FFD2FF4BFF50D004FF72" blockId="5.[151,1436,151,248]" pageId="5" pageNumber="580">
<emphasis id="B93EEAA5FFC0FFD2FF4BFF50D551FF35" bold="true" box="[199,352,151,176]" pageId="5" pageNumber="580">Distribution.</emphasis>
<taxonomicName id="4C4A4D34FFC0FFD2FEEAFF5FD640FF35" box="[358,625,152,176]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="580" phylum="Chordata" rank="species" species="melanomaculata">
<emphasis id="B93EEAA5FFC0FFD2FEEAFF5FD640FF35" box="[358,625,152,176]" italics="true" pageId="5" pageNumber="580">Pethia melanomaculata</emphasis>
</taxonomicName>
is restricted to the northern and eastern dry zone (annual rainfall &lt;
<quantity id="4CB29B52FFC0FFD2FAD1FF50D4F5FF51" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="580" unit="mm" value="1500.0">1,500 mm</quantity>
) of
<collectingCountry id="F35D7627FFC0FFD2FF7AFF7BD557FF50" box="[246,358,188,213]" name="Sri Lanka" pageId="5" pageNumber="580">Sri Lanka</collectingCountry>
where it is widely distributed in tanks and small rivers, primarily within the Mahaweli River basin (
<figureCitation id="13712A32FFC0FFD2FF6DFF18D511FF7D" box="[225,288,223,248]" captionStart="FIGURE 1. A" captionStartId="2.[151,250,1896,1918]" captionTargetBox="[200,1380,231,1850]" captionTargetId="figure@2.[200,1381,231,1852]" captionTargetPageId="2" captionText="FIGURE 1. A, Pethia melanomaculata, colour in life (not preserved); B, copy of iconotype of Cyprinus ticto Hamilton, 1822 (Courtesy Linnean Society of London); C, illustration of Puntius ticto, from Day (1878, pl. 144, fig. 7); D, distribution of Pethia species in Sri Lanka: ● Pethia melanomaculata (location of holotype in red); other known locations (black); {■}, Pethia bandula (purple); P. cumingii (yellow); P. nigrofasciata (green); P. reval (blue)." httpUri="https://zenodo.org/record/244128/files/figure.png" pageId="5" pageNumber="580">Fig. 1</figureCitation>
D). It is the only species of
<taxonomicName id="4C4A4D34FFC0FFD2FDD5FF27D693FF7D" box="[601,674,224,248]" class="Actinopterygii" family="Cyprinidae" genus="Pethia" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="580" phylum="Chordata" rank="genus">
<emphasis id="B93EEAA5FFC0FFD2FDD5FF27D693FF7D" box="[601,674,224,248]" italics="true" pageId="5" pageNumber="580">Pethia</emphasis>
</taxonomicName>
that occurs in the islands dry zone.
</paragraph>
</subSubSection>
</treatment>
</document>