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<document id="03C035797369D60518409764221CBD75" ID-CLB-Dataset="3307" ID-DOI="10.11646/zootaxa.5071.1.5" ID-GBIF-Dataset="f5a054cf-41c7-40cd-bac9-63e1ea8d1b8e" ID-ISSN="1175-5326" ID-Zenodo-Dep="5723327" ID-ZooBank="C7DFE939-E932-4570-8496-7A67A51D4D7A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1637742853538" checkinUser="plazi" docAuthor="Suksai, Benjamart, Gustafson, Grey T., Sites, Robert W. &amp; Sangpradub, Narumon" docDate="2021" docId="03DC697BFF98FFCF77CAFC87FA37F8F7" docLanguage="en" docName="zootaxa.5071.1.5.pdf" docOrigin="Zootaxa 5071 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gyrinidae" docType="key" docVersion="9" lastPageNumber="114" masterDocId="FFE51103FF89FFDE775DFFE0FFABFFCF" masterDocTitle="The Gyrinidae (Coleoptera) fauna of Thailand: Key to tribes and genera, with new records and keys to species of Dineutini and Gyrinini" masterLastPageNumber="117" masterPageNumber="97" pageNumber="114" updateTime="1699469841920" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="6FE9A4E9FFB24008D2EAD6B302366147">The Gyrinidae (Coleoptera) fauna of Thailand: Key to tribes and genera, with new records and keys to species of Dineutini and Gyrinini</mods:title>
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<mods:affiliation id="D729C276DCA7F9EF5D7BA5AF150A80F5">Department of Ecology and Evolutionary Biology, University of Kansas, Lawrence, Kansas 66045, USA. &amp; Department of Biological Sciences, Northern Arizona University, Flagstaff, Arizona, 86011, USA.</mods:affiliation>
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<mods:namePart id="9A3929204BDF1365A99BD71086332B63">Sites, Robert W.</mods:namePart>
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<mods:affiliation id="871954479E8C2348C3C89A79FB36C0C8">Enns Entomology Museum, Division of Plant Science, University of Missouri-Columbia, Missouri 65211, USA. SitesR @ missouri. edu; https: // orcid. org / 0000 - 0002 - 3895 - 813 X</mods:affiliation>
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Key to the tribes and genera of adult
<taxonomicName id="4C75A3EEFF98FFCF7520FC86FD57FC4F" ID-CoL="625FK" ID-ENA="107778" box="[637,764,870,896]" class="Insecta" family="Gyrinidae" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="114" phylum="Arthropoda" rank="family">Gyrinidae</taxonomicName>
in Thailand,
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<paragraph id="8BCAD86DFF98FFCF74F6FC86FEA7FC6B" blockId="17.[151,1436,870,932]" pageId="17" pageNumber="114">
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<emphasis id="B901047FFF98FFCF74F6FC86FEA7FC6B" bold="true" pageId="17" pageNumber="114">
including species of adult
<taxonomicName id="4C75A3EEFF98FFCF73AAFC86FAC8FC4F" authorityName="Latreille" authorityYear="1810" box="[1271,1379,870,896]" class="Insecta" family="Gyrinidae" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="114" phylum="Arthropoda" rank="tribe" tribe="Gyrinini">Gyrinini</taxonomicName>
and
<taxonomicName id="4C75A3EEFF98FFCF77CAFC6AFEA7FC6B" authorityName="Desmarest" authorityYear="1851" box="[151,268,906,932]" class="Insecta" family="Gyrinidae" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="114" phylum="Arthropoda" rank="tribe" tribe="Dineutini">Dineutini</taxonomicName>
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1 Dorsal surface of pronotum and elytron with pubescence; abdominal sternite VIII narrow and long.......
<taxonomicName id="4C75A3EEFF98FFCF73A3FC2FFA2FFC2A" box="[1278,1412,975,997]" class="Insecta" family="Gyrinidae" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="114" phylum="Arthropoda" rank="tribe" tribe="Orectochilini">
<emphasis id="B901047FFF98FFCF73A3FC2FFA2FFC2A" bold="true" box="[1278,1412,975,997]" pageId="17" pageNumber="114">Orectochilini</emphasis>
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, 2
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<paragraph id="8BCAD86DFF98FFCF77CAFC0DFA37FBCD" blockId="17.[151,1437,975,1848]" box="[151,1436,1005,1027]" pageId="17" pageNumber="114">
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- Dorsal surface of pronotum and elytron completely glabrous (
<figureCitation id="134EC4E8FF98FFCF7469FC0DFC03FBCC" box="[820,936,1005,1027]" captionStart="FIGURE 1" captionStartId="3.[151,250,1894,1918]" captionTargetBox="[201,1387,181,1871]" captionTargetId="figure-18@3.[201,1387,181,1871]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Gyrinini and Dineutini species: (A) Gyrinus sericeolimbatus, dorsal view, female; (B) G. smaragdinus, dorsal view, male; (CD) Dineutus australis, dorsal view: (C) male; (D) female; (E) D. sitesi, dorsal view, male; (F) D. spinosus, dorsal view, male; (G) D. unidentatus, dorsal view, male; (H) Porrorhynchus marginatus, dorsal view, male; (I) D. sitesi, ventral view, male; (J) D. spinosus, ventral view, male. Scale bars = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.5723329" httpUri="https://zenodo.org/record/5723329/files/figure.png" pageId="17" pageNumber="114">Figs. 1AH</figureCitation>
); abdominal sternite VIII broad and short....... 3
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<paragraph id="8BCAD86DFF98FFCF77CAFBEAFA37FBF2" blockId="17.[151,1437,975,1848]" pageId="17" pageNumber="114">
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2 Antenna with six flagellomeres; surface of pronotum and elytron with glabrous regions medially and pubescence marginally...............................................................................................
<taxonomicName id="4C75A3EEFF98FFCF7206FBC8FA37FBF2" authorityName="Aube" authorityYear="1838" box="[1371,1436,1064,1085]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="114" phylum="Arthropoda" rank="genus">
<emphasis id="B901047FFF98FFCF7206FBC8FA37FBF2" bold="true" box="[1371,1436,1064,1085]" italics="true" pageId="17" pageNumber="114">Patrus</emphasis>
</taxonomicName>
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<paragraph id="8BCAD86DFF98FFCF77CAFBA5FA36FBB7" blockId="17.[151,1437,975,1848]" pageId="17" pageNumber="114">
<keyLead id="30847B58FF98FFCF77CAFBA5FA36FBB7" pageId="17" pageNumber="114">
- Antenna with nine flagellomeres; surface of pronotum and elytron without glabrous regions, entirely covered with pubescence....................................................................................
<taxonomicName id="4C75A3EEFF98FFCF7243FB82FA36FBB7" box="[1310,1437,1122,1144]" class="Insecta" family="Gyrinidae" genus="Orectochilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="114" phylum="Arthropoda" rank="genus">
<emphasis id="B901047FFF98FFCF7243FB82FA36FBB7" bold="true" box="[1310,1437,1122,1144]" italics="true" pageId="17" pageNumber="114">Orectochilus</emphasis>
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<keyStep id="3081C0C8FF98FFCF77CAFB60FA37FB22" pageId="17" pageNumber="114">
<paragraph id="8BCAD86DFF98FFCF77CAFB60FA37FB7D" blockId="17.[151,1437,975,1848]" pageId="17" pageNumber="114">
<keyLead id="30847B58FF98FFCF77CAFB60FA37FB7D" pageId="17" pageNumber="114">
3 Size smaller, length &lt;6.0 mm; pronotum with median transverse depression; scutellum visible; elytron with 11 longitudinal striae (
<figureCitation id="134EC4E8FF98FFCF767EFB7DFE3EFB7D" box="[291,405,1181,1203]" captionStart="FIGURE 1" captionStartId="3.[151,250,1894,1918]" captionTargetBox="[201,1387,181,1871]" captionTargetId="figure-18@3.[201,1387,181,1871]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Gyrinini and Dineutini species: (A) Gyrinus sericeolimbatus, dorsal view, female; (B) G. smaragdinus, dorsal view, male; (CD) Dineutus australis, dorsal view: (C) male; (D) female; (E) D. sitesi, dorsal view, male; (F) D. spinosus, dorsal view, male; (G) D. unidentatus, dorsal view, male; (H) Porrorhynchus marginatus, dorsal view, male; (I) D. sitesi, ventral view, male; (J) D. spinosus, ventral view, male. Scale bars = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.5723329" httpUri="https://zenodo.org/record/5723329/files/figure.png" pageId="17" pageNumber="114">Figs. 1AB</figureCitation>
,
<figureCitation id="134EC4E8FF98FFCF76C2FB7DFE72FB7C" box="[415,473,1181,1203]" captionStart="FIGURE 2" captionStartId="4.[151,250,1087,1111]" captionTargetBox="[267,1320,181,1063]" captionTargetId="figure-22@4.[267,1320,181,1063]" captionTargetPageId="4" captionText="FIGURE 2. Gyrinus smaragdinus: (A) female pronotum, median transverse depression (arrow), scale bar = 300 μm; (B) male elytra, spaced puncture (arrow), scale bar = 1 mm; (C) male elytral apex, scale bar = 300 μm." figureDoi="http://doi.org/10.5281/zenodo.5723331" httpUri="https://zenodo.org/record/5723331/files/figure.png" pageId="17" pageNumber="114">2AB</figureCitation>
)..............................................................
<taxonomicName id="4C75A3EEFF98FFCF738DFB7DFA83FB7C" authorityName="Latreille" authorityYear="1810" box="[1232,1320,1181,1203]" class="Insecta" family="Gyrinidae" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="114" phylum="Arthropoda" rank="tribe" tribe="Gyrinini">
<emphasis id="B901047FFF98FFCF738DFB7DFA83FB7C" bold="true" box="[1232,1320,1181,1203]" pageId="17" pageNumber="114">Gyrinini</emphasis>
</taxonomicName>
,
<taxonomicName id="4C75A3EEFF98FFCF7269FB7DFA2FFB7D" ID-CoL="4S46" box="[1332,1412,1181,1202]" class="Insecta" family="Gyrinidae" genus="Gyrinus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="114" phylum="Arthropoda" rank="genus">
<emphasis id="B901047FFF98FFCF7269FB7DFA2FFB7D" bold="true" box="[1332,1412,1181,1202]" italics="true" pageId="17" pageNumber="114">Gyrinus</emphasis>
</taxonomicName>
, 4
</keyLead>
</paragraph>
<paragraph id="8BCAD86DFF98FFCF77CAFB5BFA37FB22" blockId="17.[151,1437,975,1848]" pageId="17" pageNumber="114">
<keyLead id="30847B58FF98FFCF77CAFB5BFA37FB22" pageId="17" pageNumber="114">
- Size larger, length 6.017.0 mm; pronotum without median transverse depression; scutellum not visible (
<figureCitation id="134EC4E8FF98FFCF7389FB5AFAECFB1F" box="[1236,1351,1210,1232]" captionStart="FIGURE 1" captionStartId="3.[151,250,1894,1918]" captionTargetBox="[201,1387,181,1871]" captionTargetId="figure-18@3.[201,1387,181,1871]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Gyrinini and Dineutini species: (A) Gyrinus sericeolimbatus, dorsal view, female; (B) G. smaragdinus, dorsal view, male; (CD) Dineutus australis, dorsal view: (C) male; (D) female; (E) D. sitesi, dorsal view, male; (F) D. spinosus, dorsal view, male; (G) D. unidentatus, dorsal view, male; (H) Porrorhynchus marginatus, dorsal view, male; (I) D. sitesi, ventral view, male; (J) D. spinosus, ventral view, male. Scale bars = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.5723329" httpUri="https://zenodo.org/record/5723329/files/figure.png" pageId="17" pageNumber="114">Figs. 1CH</figureCitation>
); elytron without evident longitudinal striae..............................................................
<taxonomicName id="4C75A3EEFF98FFCF7279FB37FA2FFB22" authorityName="Desmarest" authorityYear="1851" box="[1316,1412,1239,1261]" class="Insecta" family="Gyrinidae" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="114" phylum="Arthropoda" rank="tribe" tribe="Dineutini">
<emphasis id="B901047FFF98FFCF7279FB37FA2FFB22" bold="true" box="[1316,1412,1239,1261]" pageId="17" pageNumber="114">Dineutini</emphasis>
</taxonomicName>
, 5
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<keyStep id="3081C0C8FF98FFCF77CAFB15FA37FAAD" pageId="17" pageNumber="114">
<paragraph id="8BCAD86DFF98FFCF77CAFB15FA36FAE7" blockId="17.[151,1437,975,1848]" pageId="17" pageNumber="114">
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4 Dorsally metallic in appearance lateral sides of elytra with bronzy sheen (
<figureCitation id="134EC4E8FF98FFCF74F9FB15FC5BFAC4" box="[932,1008,1269,1291]" captionStart="FIGURE 1" captionStartId="3.[151,250,1894,1918]" captionTargetBox="[201,1387,181,1871]" captionTargetId="figure-18@3.[201,1387,181,1871]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Gyrinini and Dineutini species: (A) Gyrinus sericeolimbatus, dorsal view, female; (B) G. smaragdinus, dorsal view, male; (CD) Dineutus australis, dorsal view: (C) male; (D) female; (E) D. sitesi, dorsal view, male; (F) D. spinosus, dorsal view, male; (G) D. unidentatus, dorsal view, male; (H) Porrorhynchus marginatus, dorsal view, male; (I) D. sitesi, ventral view, male; (J) D. spinosus, ventral view, male. Scale bars = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.5723329" httpUri="https://zenodo.org/record/5723329/files/figure.png" pageId="17" pageNumber="114">Fig. 1A</figureCitation>
), intervals VIX not sulcate............................................................................................
<taxonomicName id="4C75A3EEFF98FFCF73F1FAF2FA36FAE7" authorityName="Regimbart" authorityYear="1883" box="[1196,1437,1298,1320]" class="Insecta" family="Gyrinidae" genus="Gyrinus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="114" phylum="Arthropoda" rank="species" species="sericeolimbatus">
<emphasis id="B901047FFF98FFCF73F1FAF2FA36FAE7" bold="true" box="[1196,1437,1298,1320]" italics="true" pageId="17" pageNumber="114">Gyrinus sericeolimbatus</emphasis>
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<paragraph id="8BCAD86DFF98FFCF77CAFAD0FA37FAAD" blockId="17.[151,1437,975,1848]" pageId="17" pageNumber="114">
<keyLead id="30847B58FF98FFCF77CAFAD0FA37FAAD" pageId="17" pageNumber="114">
- Dorsally iridescent in appearance (
<figureCitation id="134EC4E8FF98FFCF756CFAD0FDD7FA89" box="[561,636,1328,1350]" captionStart="FIGURE 1" captionStartId="3.[151,250,1894,1918]" captionTargetBox="[201,1387,181,1871]" captionTargetId="figure-18@3.[201,1387,181,1871]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Gyrinini and Dineutini species: (A) Gyrinus sericeolimbatus, dorsal view, female; (B) G. smaragdinus, dorsal view, male; (CD) Dineutus australis, dorsal view: (C) male; (D) female; (E) D. sitesi, dorsal view, male; (F) D. spinosus, dorsal view, male; (G) D. unidentatus, dorsal view, male; (H) Porrorhynchus marginatus, dorsal view, male; (I) D. sitesi, ventral view, male; (J) D. spinosus, ventral view, male. Scale bars = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.5723329" httpUri="https://zenodo.org/record/5723329/files/figure.png" pageId="17" pageNumber="114">Fig. 1B</figureCitation>
); lateral sides of elytra without bronzy sheen, intervals VIX strongly sulcate (
<figureCitation id="134EC4E8FF98FFCF7228FAD0FF55FAAC" captionStart="FIGURE 2" captionStartId="4.[151,250,1087,1111]" captionTargetBox="[267,1320,181,1063]" captionTargetId="figure-22@4.[267,1320,181,1063]" captionTargetPageId="4" captionText="FIGURE 2. Gyrinus smaragdinus: (A) female pronotum, median transverse depression (arrow), scale bar = 300 μm; (B) male elytra, spaced puncture (arrow), scale bar = 1 mm; (C) male elytral apex, scale bar = 300 μm." figureDoi="http://doi.org/10.5281/zenodo.5723331" httpUri="https://zenodo.org/record/5723331/files/figure.png" pageId="17" pageNumber="114">Fig. 2B</figureCitation>
)...............................................................................
<taxonomicName id="4C75A3EEFF98FFCF739BFAADFA37FAAD" box="[1222,1436,1356,1378]" class="Insecta" family="Gyrinidae" genus="Gyrinus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="114" phylum="Arthropoda" rank="species" species="smaragdinus">
<emphasis id="B901047FFF98FFCF739BFAADFA37FAAD" bold="true" box="[1222,1436,1356,1378]" italics="true" pageId="17" pageNumber="114">Gyrinus smaragdinus</emphasis>
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<keyStep id="3081C0C8FF98FFCF77CAFA8AFA37FA53" pageId="17" pageNumber="114">
<paragraph id="8BCAD86DFF98FFCF77CAFA8AFA37FA4F" blockId="17.[151,1437,975,1848]" box="[151,1436,1386,1408]" pageId="17" pageNumber="114">
<keyLead id="30847B58FF98FFCF77CAFA8AFA37FA4F" box="[151,1436,1386,1408]" pageId="17" pageNumber="114">
5 Labrum short and broad, not triangular in form (
<figureCitation id="134EC4E8FF98FFCF75EFFA8AFC8FFA4F" box="[690,804,1386,1408]" captionStart="FIGURE 1" captionStartId="3.[151,250,1894,1918]" captionTargetBox="[201,1387,181,1871]" captionTargetId="figure-18@3.[201,1387,181,1871]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Gyrinini and Dineutini species: (A) Gyrinus sericeolimbatus, dorsal view, female; (B) G. smaragdinus, dorsal view, male; (CD) Dineutus australis, dorsal view: (C) male; (D) female; (E) D. sitesi, dorsal view, male; (F) D. spinosus, dorsal view, male; (G) D. unidentatus, dorsal view, male; (H) Porrorhynchus marginatus, dorsal view, male; (I) D. sitesi, ventral view, male; (J) D. spinosus, ventral view, male. Scale bars = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.5723329" httpUri="https://zenodo.org/record/5723329/files/figure.png" pageId="17" pageNumber="114">Figs. 1CG</figureCitation>
); apex of protibia without spine................
<taxonomicName id="4C75A3EEFF98FFCF7277FA8AFA2EFAB0" ID-CoL="45QT" authorityName="MacLeay" authorityYear="1825" box="[1322,1413,1386,1407]" class="Insecta" family="Gyrinidae" genus="Dineutus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="114" phylum="Arthropoda" rank="genus">
<emphasis id="B901047FFF98FFCF7277FA8AFA2EFAB0" bold="true" box="[1322,1413,1386,1407]" italics="true" pageId="17" pageNumber="114">Dineutus</emphasis>
</taxonomicName>
, 6
</keyLead>
</paragraph>
<paragraph id="8BCAD86DFF98FFCF77CAFA68FA37FA53" blockId="17.[151,1437,975,1848]" box="[151,1436,1415,1438]" pageId="17" pageNumber="114">
<keyLead id="30847B58FF98FFCF77CAFA68FA37FA53" box="[151,1436,1415,1438]" pageId="17" pageNumber="114">
- Labrum long and triangular in form (
<figureCitation id="134EC4E8FF98FFCF7518FA68FD3AFA51" box="[581,657,1416,1438]" captionStart="FIGURE 1" captionStartId="3.[151,250,1894,1918]" captionTargetBox="[201,1387,181,1871]" captionTargetId="figure-18@3.[201,1387,181,1871]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Gyrinini and Dineutini species: (A) Gyrinus sericeolimbatus, dorsal view, female; (B) G. smaragdinus, dorsal view, male; (CD) Dineutus australis, dorsal view: (C) male; (D) female; (E) D. sitesi, dorsal view, male; (F) D. spinosus, dorsal view, male; (G) D. unidentatus, dorsal view, male; (H) Porrorhynchus marginatus, dorsal view, male; (I) D. sitesi, ventral view, male; (J) D. spinosus, ventral view, male. Scale bars = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.5723329" httpUri="https://zenodo.org/record/5723329/files/figure.png" pageId="17" pageNumber="114">Fig. 1H</figureCitation>
); apex of protibia with spine (
<figureCitation id="134EC4E8FF98FFCF74EFFA68FC56FA51" box="[946,1021,1416,1438]" captionStart="FIGURE 1" captionStartId="3.[151,250,1894,1918]" captionTargetBox="[201,1387,181,1871]" captionTargetId="figure-18@3.[201,1387,181,1871]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Gyrinini and Dineutini species: (A) Gyrinus sericeolimbatus, dorsal view, female; (B) G. smaragdinus, dorsal view, male; (CD) Dineutus australis, dorsal view: (C) male; (D) female; (E) D. sitesi, dorsal view, male; (F) D. spinosus, dorsal view, male; (G) D. unidentatus, dorsal view, male; (H) Porrorhynchus marginatus, dorsal view, male; (I) D. sitesi, ventral view, male; (J) D. spinosus, ventral view, male. Scale bars = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.5723329" httpUri="https://zenodo.org/record/5723329/files/figure.png" pageId="17" pageNumber="114">Fig. 1H</figureCitation>
).........
<taxonomicName id="4C75A3EEFF98FFCF73D3FA67FA37FA53" authorityName="Laporte" authorityYear="1835" box="[1166,1436,1415,1437]" class="Insecta" family="Gyrinidae" genus="Porrorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="114" phylum="Arthropoda" rank="species" species="marginatus">
<emphasis id="B901047FFF98FFCF73D3FA67FA37FA53" bold="true" box="[1166,1436,1415,1437]" italics="true" pageId="17" pageNumber="114">Porrorhynchus marginatus</emphasis>
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<keyStep id="3081C0C8FF98FFCF77CAFA45FA37F9DD" pageId="17" pageNumber="114">
<paragraph id="8BCAD86DFF98FFCF77CAFA45FA37FA17" blockId="17.[151,1437,975,1848]" pageId="17" pageNumber="114">
<keyLead id="30847B58FF98FFCF77CAFA45FA37FA17" pageId="17" pageNumber="114">
6 Size smaller, length 6.0
<quantity id="4C8D7588FF98FFCF7690FA45FDB3FA75" box="[461,536,1445,1467]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="17" pageNumber="114" unit="mm" value="8.2">8.2 mm</quantity>
; profemur of males without sub-apicoventral tooth; protibia wedge-shaped; elytral apices variously shaped........................................................................................ 7
</keyLead>
</paragraph>
<paragraph id="8BCAD86DFF98FFCF77CAFA00FA37F9DD" blockId="17.[151,1437,975,1848]" pageId="17" pageNumber="114">
<keyLead id="30847B58FF98FFCF77CAFA00FA37F9DD" pageId="17" pageNumber="114">
- Size larger, length 13.417.0 mm; profemur of males with sub-apicoventral tooth; protibia club-shaped; elytral apices rounded (
<figureCitation id="134EC4E8FF98FFCF77B4FA1DFE90F9DD" box="[233,315,1533,1555]" captionStart="FIGURE 1" captionStartId="3.[151,250,1894,1918]" captionTargetBox="[201,1387,181,1871]" captionTargetId="figure-18@3.[201,1387,181,1871]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Gyrinini and Dineutini species: (A) Gyrinus sericeolimbatus, dorsal view, female; (B) G. smaragdinus, dorsal view, male; (CD) Dineutus australis, dorsal view: (C) male; (D) female; (E) D. sitesi, dorsal view, male; (F) D. spinosus, dorsal view, male; (G) D. unidentatus, dorsal view, male; (H) Porrorhynchus marginatus, dorsal view, male; (I) D. sitesi, ventral view, male; (J) D. spinosus, ventral view, male. Scale bars = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.5723329" httpUri="https://zenodo.org/record/5723329/files/figure.png" pageId="17" pageNumber="114">Figs. 1E</figureCitation>
,
<figureCitation id="134EC4E8FF98FFCF761BFA1DFECBF9DC" box="[326,352,1533,1555]" captionStart="FIGURE 5" captionStartId="8.[151,250,1718,1742]" captionTargetBox="[194,1392,374,1695]" captionTargetId="figure-103@8.[165,1422,345,1695]" captionTargetPageId="8" captionText="FIGURE 5. Elytral apex, aedeagus and gonocoxae of species of Dineutus: (AE) D. australis: (AB) elytral apex, scale bar = 2 mm: (A) male; (B) female; (CD) aedeagus, scale bar = 0.5 mm: (C) dorsal view; (D) lateral view; (E) gonocoxae, ventral view, scale bar = 0.5 mm; (FI) D. sitesi: (F) male elytral apex, scale bar = 2 mm; (GH) aedeagus, scale bar = 1 mm: (G) dorsal view; (H) lateral view; (I) gonocoxae, ventral view, scale bar = 1 mm; (JM) D. spinosus: (J) male elytral apex, scale bar = 2 mm, (KL) aedeagus, scale bar = 0.5 mm: (K) dorsal view; (L) lateral view; (M) gonocoxae, ventral view, scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5723337" httpUri="https://zenodo.org/record/5723337/files/figure.png" pageId="17" pageNumber="114">5F</figureCitation>
)..................................................................................
<taxonomicName id="4C75A3EEFF98FFCF7213FA1DFA37F9DD" box="[1358,1436,1533,1554]" class="Insecta" family="Gyrinidae" genus="Dineutus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="114" phylum="Arthropoda" rank="species" species="sitesi">
<emphasis id="B901047FFF98FFCF7213FA1DFA37F9DD" bold="true" box="[1358,1436,1533,1554]" italics="true" pageId="17" pageNumber="114">D. sitesi</emphasis>
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<keyStep id="3081C0C8FF98FFCF77CAF9FBFA37F948" pageId="17" pageNumber="114">
<paragraph id="8BCAD86DFF98FFCF77CAF9FBFA37F982" blockId="17.[151,1437,975,1848]" pageId="17" pageNumber="114">
<keyLead id="30847B58FF98FFCF77CAF9FBFA37F982" pageId="17" pageNumber="114">7 Apex of flagellomere VI darkly colored; elytral apices without parasutural spines; lateral margin of pronotum and elytron black.............................................................................................. 8</keyLead>
</paragraph>
<paragraph id="8BCAD86DFF98FFCF77CAF9B5FA37F948" blockId="17.[151,1437,975,1848]" pageId="17" pageNumber="114">
<keyLead id="30847B58FF98FFCF77CAF9B5FA37F948" pageId="17" pageNumber="114">
- Apex of flagellomere VI lightly colored; elytral apices with parasutural spines (
<figureCitation id="134EC4E8FF98FFCF74BAF9B5FB9CF9A5" box="[999,1079,1621,1643]" captionStart="FIGURE 1" captionStartId="3.[151,250,1894,1918]" captionTargetBox="[201,1387,181,1871]" captionTargetId="figure-18@3.[201,1387,181,1871]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Gyrinini and Dineutini species: (A) Gyrinus sericeolimbatus, dorsal view, female; (B) G. smaragdinus, dorsal view, male; (CD) Dineutus australis, dorsal view: (C) male; (D) female; (E) D. sitesi, dorsal view, male; (F) D. spinosus, dorsal view, male; (G) D. unidentatus, dorsal view, male; (H) Porrorhynchus marginatus, dorsal view, male; (I) D. sitesi, ventral view, male; (J) D. spinosus, ventral view, male. Scale bars = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.5723329" httpUri="https://zenodo.org/record/5723329/files/figure.png" pageId="17" pageNumber="114">Figs. 1F</figureCitation>
,
<figureCitation id="134EC4E8FF98FFCF731FF9B5FBF1F9A4" box="[1090,1114,1621,1643]" captionStart="FIGURE 5" captionStartId="8.[151,250,1718,1742]" captionTargetBox="[194,1392,374,1695]" captionTargetId="figure-103@8.[165,1422,345,1695]" captionTargetPageId="8" captionText="FIGURE 5. Elytral apex, aedeagus and gonocoxae of species of Dineutus: (AE) D. australis: (AB) elytral apex, scale bar = 2 mm: (A) male; (B) female; (CD) aedeagus, scale bar = 0.5 mm: (C) dorsal view; (D) lateral view; (E) gonocoxae, ventral view, scale bar = 0.5 mm; (FI) D. sitesi: (F) male elytral apex, scale bar = 2 mm; (GH) aedeagus, scale bar = 1 mm: (G) dorsal view; (H) lateral view; (I) gonocoxae, ventral view, scale bar = 1 mm; (JM) D. spinosus: (J) male elytral apex, scale bar = 2 mm, (KL) aedeagus, scale bar = 0.5 mm: (K) dorsal view; (L) lateral view; (M) gonocoxae, ventral view, scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5723337" httpUri="https://zenodo.org/record/5723337/files/figure.png" pageId="17" pageNumber="114">5J</figureCitation>
); lateral margin of pronotum and elytron orange or yellow......................................................................
<taxonomicName id="4C75A3EEFF98FFCF7277F993FA37F948" box="[1322,1436,1650,1672]" class="Insecta" family="Gyrinidae" genus="Dineutus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="114" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B901047FFF98FFCF7277F993FA37F948" bold="true" box="[1322,1436,1650,1672]" italics="true" pageId="17" pageNumber="114">D. spinosus</emphasis>
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<keyStep id="3081C0C8FF98FFCF77CAF970FA37F8F7" pageId="17" pageNumber="114">
<paragraph id="8BCAD86DFF98FFCF77CAF970FA37F92F" blockId="17.[151,1437,975,1848]" pageId="17" pageNumber="114">
<keyLead id="30847B58FF98FFCF77CAF970FA37F92F" pageId="17" pageNumber="114">
8 Size smaller, length 6.0
<quantity id="4C8D7588FF98FFCF7693F970FDB1F96A" box="[462,538,1680,1702]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="17" pageNumber="114" unit="mm" value="6.9">6.9 mm</quantity>
; body form narrowly oval; elytral apices with epipleural spines (
<figureCitation id="134EC4E8FF98FFCF73DEF970FB73F96A" box="[1155,1240,1680,1702]" captionStart="FIGURE 1" captionStartId="3.[151,250,1894,1918]" captionTargetBox="[201,1387,181,1871]" captionTargetId="figure-18@3.[201,1387,181,1871]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Gyrinini and Dineutini species: (A) Gyrinus sericeolimbatus, dorsal view, female; (B) G. smaragdinus, dorsal view, male; (CD) Dineutus australis, dorsal view: (C) male; (D) female; (E) D. sitesi, dorsal view, male; (F) D. spinosus, dorsal view, male; (G) D. unidentatus, dorsal view, male; (H) Porrorhynchus marginatus, dorsal view, male; (I) D. sitesi, ventral view, male; (J) D. spinosus, ventral view, male. Scale bars = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.5723329" httpUri="https://zenodo.org/record/5723329/files/figure.png" pageId="17" pageNumber="114">Figs. 1G</figureCitation>
,
<figureCitation id="134EC4E8FF98FFCF73BFF970FAABF969" box="[1250,1280,1680,1702]" captionStart="FIGURE 6" captionStartId="12.[151,250,1219,1243]" captionTargetBox="[177,1411,192,1185]" captionTargetId="figure-22@12.[159,1428,181,1195]" captionTargetPageId="12" captionText="FIGURE 6. Elytral apex, aedeagus and gonocoxae: (AD) D. unidentatus: (A) male elytral apex, scale bar = 2 mm; (BC) aedeagus, scale bar = 0.5 mm: (B) dorsal view; (C) lateral view; (D) gonocoxae, ventral view, scale bar = 0.5 mm; (EG) P. marginatus: (EF) aedeagus, scale bars = 1 mm: (E) dorsal view; (F) lateral view; (G) gonocoxae, ventral view, scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5723339" httpUri="https://zenodo.org/record/5723339/files/figure.png" pageId="17" pageNumber="114">6A</figureCitation>
) evident in both sexes; elytral apex truncate and straight (
<figureCitation id="134EC4E8FF98FFCF7530F94DFD68F90D" box="[621,707,1709,1731]" captionStart="FIGURE 1" captionStartId="3.[151,250,1894,1918]" captionTargetBox="[201,1387,181,1871]" captionTargetId="figure-18@3.[201,1387,181,1871]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Gyrinini and Dineutini species: (A) Gyrinus sericeolimbatus, dorsal view, female; (B) G. smaragdinus, dorsal view, male; (CD) Dineutus australis, dorsal view: (C) male; (D) female; (E) D. sitesi, dorsal view, male; (F) D. spinosus, dorsal view, male; (G) D. unidentatus, dorsal view, male; (H) Porrorhynchus marginatus, dorsal view, male; (I) D. sitesi, ventral view, male; (J) D. spinosus, ventral view, male. Scale bars = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.5723329" httpUri="https://zenodo.org/record/5723329/files/figure.png" pageId="17" pageNumber="114">Figs. 1G</figureCitation>
,
<figureCitation id="134EC4E8FF98FFCF7590F94DFD45F90C" box="[717,750,1709,1731]" captionStart="FIGURE 6" captionStartId="12.[151,250,1219,1243]" captionTargetBox="[177,1411,192,1185]" captionTargetId="figure-22@12.[159,1428,181,1195]" captionTargetPageId="12" captionText="FIGURE 6. Elytral apex, aedeagus and gonocoxae: (AD) D. unidentatus: (A) male elytral apex, scale bar = 2 mm; (BC) aedeagus, scale bar = 0.5 mm: (B) dorsal view; (C) lateral view; (D) gonocoxae, ventral view, scale bar = 0.5 mm; (EG) P. marginatus: (EF) aedeagus, scale bars = 1 mm: (E) dorsal view; (F) lateral view; (G) gonocoxae, ventral view, scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5723339" httpUri="https://zenodo.org/record/5723339/files/figure.png" pageId="17" pageNumber="114">6A</figureCitation>
); protrochanter of male apically without patch of thick setae on ventral surface (
<figureCitation id="134EC4E8FF98FFCF7669F92AFE2BF92F" box="[308,384,1738,1760]" captionStart="FIGURE 4" captionStartId="7.[151,250,1575,1599]" captionTargetBox="[157,1429,181,1551]" captionTargetId="figure-18@7.[157,1429,181,1551]" captionTargetPageId="7" captionText="FIGURE 4. SEM photographs of protochanters in species of Dineutus and Porrorhynchus (males): (A) D. australis, patch of thick setae (arrow), scale bar = 100 μm; (B) D. sitesi, protrochanter without setae (arrow), scale bar = 1 mm; (C) D. spinosus, setae (arrow), scale bar = 100 μm; (D) D. unidentatus, setae (arrow), scale bar = 100 μm; (EF) P. marginatus, cluster setae (square and arrow): (E) scale bar = 500 μm; (F) scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5723335" httpUri="https://zenodo.org/record/5723335/files/figure.png" pageId="17" pageNumber="114">Fig. 4D</figureCitation>
).........................................................................
<taxonomicName id="4C75A3EEFF98FFCF7257F92BFA37F92F" box="[1290,1436,1738,1760]" class="Insecta" family="Gyrinidae" genus="Dineutus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="114" phylum="Arthropoda" rank="species" species="unidentatus">
<emphasis id="B901047FFF98FFCF7257F92BFA37F92F" bold="true" box="[1290,1436,1738,1760]" italics="true" pageId="17" pageNumber="114">D. unidentatus</emphasis>
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<paragraph id="8BCAD86DFF98FFCF77CAF908FA37F8F7" blockId="17.[151,1437,975,1848]" pageId="17" pageNumber="114">
<keyLead id="30847B58FF98FFCF77CAF908FA37F8F7" pageId="17" pageNumber="114">
- Size larger, length
<quantity id="4C8D7588FF98FFCF76C8F908FDA1F932" box="[405,522,1768,1790]" metricMagnitude="-3" metricUnit="m" metricValue="7.8999999999999995" metricValueMax="8.2" metricValueMin="7.6" pageId="17" pageNumber="114" unit="mm" value="7.8999999999999995" valueMax="8.2" valueMin="7.6">7.68.2 mm</quantity>
; body form oval; elytral apices of male only with epipleural spines (
<figureCitation id="134EC4E8FF98FFCF73C9F908FB4CF932" box="[1172,1255,1768,1790]" captionStart="FIGURE 1" captionStartId="3.[151,250,1894,1918]" captionTargetBox="[201,1387,181,1871]" captionTargetId="figure-18@3.[201,1387,181,1871]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Gyrinini and Dineutini species: (A) Gyrinus sericeolimbatus, dorsal view, female; (B) G. smaragdinus, dorsal view, male; (CD) Dineutus australis, dorsal view: (C) male; (D) female; (E) D. sitesi, dorsal view, male; (F) D. spinosus, dorsal view, male; (G) D. unidentatus, dorsal view, male; (H) Porrorhynchus marginatus, dorsal view, male; (I) D. sitesi, ventral view, male; (J) D. spinosus, ventral view, male. Scale bars = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.5723329" httpUri="https://zenodo.org/record/5723329/files/figure.png" pageId="17" pageNumber="114">Figs. 1C</figureCitation>
,
<figureCitation id="134EC4E8FF98FFCF73AFF907FABAF932" box="[1266,1297,1767,1789]" captionStart="FIGURE 5" captionStartId="8.[151,250,1718,1742]" captionTargetBox="[194,1392,374,1695]" captionTargetId="figure-103@8.[165,1422,345,1695]" captionTargetPageId="8" captionText="FIGURE 5. Elytral apex, aedeagus and gonocoxae of species of Dineutus: (AE) D. australis: (AB) elytral apex, scale bar = 2 mm: (A) male; (B) female; (CD) aedeagus, scale bar = 0.5 mm: (C) dorsal view; (D) lateral view; (E) gonocoxae, ventral view, scale bar = 0.5 mm; (FI) D. sitesi: (F) male elytral apex, scale bar = 2 mm; (GH) aedeagus, scale bar = 1 mm: (G) dorsal view; (H) lateral view; (I) gonocoxae, ventral view, scale bar = 1 mm; (JM) D. spinosus: (J) male elytral apex, scale bar = 2 mm, (KL) aedeagus, scale bar = 0.5 mm: (K) dorsal view; (L) lateral view; (M) gonocoxae, ventral view, scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5723337" httpUri="https://zenodo.org/record/5723337/files/figure.png" pageId="17" pageNumber="114">5A</figureCitation>
), female without epipleural spines (
<figureCitation id="134EC4E8FF98FFCF76EAF8E5FDA6F8D5" box="[439,525,1797,1819]" captionStart="FIGURE 1" captionStartId="3.[151,250,1894,1918]" captionTargetBox="[201,1387,181,1871]" captionTargetId="figure-18@3.[201,1387,181,1871]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Gyrinini and Dineutini species: (A) Gyrinus sericeolimbatus, dorsal view, female; (B) G. smaragdinus, dorsal view, male; (CD) Dineutus australis, dorsal view: (C) male; (D) female; (E) D. sitesi, dorsal view, male; (F) D. spinosus, dorsal view, male; (G) D. unidentatus, dorsal view, male; (H) Porrorhynchus marginatus, dorsal view, male; (I) D. sitesi, ventral view, male; (J) D. spinosus, ventral view, male. Scale bars = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.5723329" httpUri="https://zenodo.org/record/5723329/files/figure.png" pageId="17" pageNumber="114">Figs. 1D</figureCitation>
,
<figureCitation id="134EC4E8FF98FFCF7545F8E5FD9DF8D4" box="[536,566,1797,1819]" captionStart="FIGURE 5" captionStartId="8.[151,250,1718,1742]" captionTargetBox="[194,1392,374,1695]" captionTargetId="figure-103@8.[165,1422,345,1695]" captionTargetPageId="8" captionText="FIGURE 5. Elytral apex, aedeagus and gonocoxae of species of Dineutus: (AE) D. australis: (AB) elytral apex, scale bar = 2 mm: (A) male; (B) female; (CD) aedeagus, scale bar = 0.5 mm: (C) dorsal view; (D) lateral view; (E) gonocoxae, ventral view, scale bar = 0.5 mm; (FI) D. sitesi: (F) male elytral apex, scale bar = 2 mm; (GH) aedeagus, scale bar = 1 mm: (G) dorsal view; (H) lateral view; (I) gonocoxae, ventral view, scale bar = 1 mm; (JM) D. spinosus: (J) male elytral apex, scale bar = 2 mm, (KL) aedeagus, scale bar = 0.5 mm: (K) dorsal view; (L) lateral view; (M) gonocoxae, ventral view, scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5723337" httpUri="https://zenodo.org/record/5723337/files/figure.png" pageId="17" pageNumber="114">5B</figureCitation>
); elytral apex obliquely truncate (
<figureCitation id="134EC4E8FF98FFCF7421F8E5FC45F8D5" box="[892,1006,1797,1819]" captionStart="FIGURE 1" captionStartId="3.[151,250,1894,1918]" captionTargetBox="[201,1387,181,1871]" captionTargetId="figure-18@3.[201,1387,181,1871]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of Gyrinini and Dineutini species: (A) Gyrinus sericeolimbatus, dorsal view, female; (B) G. smaragdinus, dorsal view, male; (CD) Dineutus australis, dorsal view: (C) male; (D) female; (E) D. sitesi, dorsal view, male; (F) D. spinosus, dorsal view, male; (G) D. unidentatus, dorsal view, male; (H) Porrorhynchus marginatus, dorsal view, male; (I) D. sitesi, ventral view, male; (J) D. spinosus, ventral view, male. Scale bars = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.5723329" httpUri="https://zenodo.org/record/5723329/files/figure.png" pageId="17" pageNumber="114">Figs. 1CD</figureCitation>
,
<figureCitation id="134EC4E8FF98FFCF74A4F8E5FB9DF8D4" box="[1017,1078,1797,1819]" captionStart="FIGURE 5" captionStartId="8.[151,250,1718,1742]" captionTargetBox="[194,1392,374,1695]" captionTargetId="figure-103@8.[165,1422,345,1695]" captionTargetPageId="8" captionText="FIGURE 5. Elytral apex, aedeagus and gonocoxae of species of Dineutus: (AE) D. australis: (AB) elytral apex, scale bar = 2 mm: (A) male; (B) female; (CD) aedeagus, scale bar = 0.5 mm: (C) dorsal view; (D) lateral view; (E) gonocoxae, ventral view, scale bar = 0.5 mm; (FI) D. sitesi: (F) male elytral apex, scale bar = 2 mm; (GH) aedeagus, scale bar = 1 mm: (G) dorsal view; (H) lateral view; (I) gonocoxae, ventral view, scale bar = 1 mm; (JM) D. spinosus: (J) male elytral apex, scale bar = 2 mm, (KL) aedeagus, scale bar = 0.5 mm: (K) dorsal view; (L) lateral view; (M) gonocoxae, ventral view, scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5723337" httpUri="https://zenodo.org/record/5723337/files/figure.png" pageId="17" pageNumber="114">5AB</figureCitation>
); protrochanter of male apically with patch of thick setae on ventral surface (
<figureCitation id="134EC4E8FF98FFCF7506F8C2FD0DF8F7" box="[603,678,1826,1848]" captionStart="FIGURE 4" captionStartId="7.[151,250,1575,1599]" captionTargetBox="[157,1429,181,1551]" captionTargetId="figure-18@7.[157,1429,181,1551]" captionTargetPageId="7" captionText="FIGURE 4. SEM photographs of protochanters in species of Dineutus and Porrorhynchus (males): (A) D. australis, patch of thick setae (arrow), scale bar = 100 μm; (B) D. sitesi, protrochanter without setae (arrow), scale bar = 1 mm; (C) D. spinosus, setae (arrow), scale bar = 100 μm; (D) D. unidentatus, setae (arrow), scale bar = 100 μm; (EF) P. marginatus, cluster setae (square and arrow): (E) scale bar = 500 μm; (F) scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.5723335" httpUri="https://zenodo.org/record/5723335/files/figure.png" pageId="17" pageNumber="114">Fig. 4A</figureCitation>
)....................................................
<taxonomicName id="4C75A3EEFF98FFCF7277F8C3FA37F8F7" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[1322,1436,1826,1848]" class="Insecta" family="Gyrinidae" genus="Dineutus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="114" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="B901047FFF98FFCF7277F8C3FA37F8F7" bold="true" box="[1322,1436,1826,1848]" italics="true" pageId="17" pageNumber="114">D. australis</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>