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<document id="CE8867FC31650AF9088BC47235ED3AF1" ID-DOI="10.5281/zenodo.198427" ID-GBIF-Dataset="1c32eb21-6978-41da-b589-c98453d4f572" ID-ISSN="1175-5326" ID-Zenodo-Dep="198427" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460297110161" checkinUser="plazi" docAuthor="Short, Andrew E. Z. &amp; García, Mauricio" docDate="2010" docId="03CEA91F8C29B90DA2C062C5FE04F7EB" docLanguage="en" docName="zt02635p031.pdf" docOrigin="Zootaxa 2635" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Oocyclus meridensis Short &amp; Garcia, sp. n." docType="treatment" docVersion="12" lastPageNumber="18" masterDocId="FFF7D1678C27B91CA2576660FFD8F557" masterDocTitle="A review of the Oocyclus Sharp of Venezuela with description of twelve new species (Coleoptera: Hydrophilidae: Laccobiini)" masterLastPageNumber="31" masterPageNumber="1" pageNumber="15" updateTime="1698252212253" updateUser="plazi">
<mods:mods id="4B25915C2E9CC072EC3A8F01DDA09D86" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8B3E85FB7666037EFB4501AC7CB457E9">
<mods:title id="C13EB1E029040AE0CFC6666F8158A878">A review of the Oocyclus Sharp of Venezuela with description of twelve new species (Coleoptera: Hydrophilidae: Laccobiini)</mods:title>
</mods:titleInfo>
<mods:name id="EBAC5086547272D4A28D9622CA412D60" type="personal">
<mods:role id="E8978505BB552634112F4F7AE09E6F61">
<mods:roleTerm id="C0C436CEC8DE3850DA3621B468724EAE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="08ABA028CE419FD6F528570AC48FA669">Short, Andrew E. Z.</mods:namePart>
</mods:name>
<mods:name id="1EAB4BA5D1C3AD6FA965DF788AD3C975" type="personal">
<mods:role id="DEFED6398A6F0AEDEC131874AC5851AC">
<mods:roleTerm id="1F5C2BB19EE60A365A91728C24B018B7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E54C1BACE1010D499C2C622B9A7C1802">García, Mauricio</mods:namePart>
</mods:name>
<mods:typeOfResource id="D39D155F0B12DAF4D314BCE09AD05B9E">text</mods:typeOfResource>
<mods:relatedItem id="BC8490B5770AC63909C2A0EE005029FC" type="host">
<mods:titleInfo id="9F4C2BA362E4A8633755490AA5ED00E1">
<mods:title id="99A6CAC198F5D0448A91BA7EBCA80DF6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="7DFD1C73BE17333628861021FAD757CD">
<mods:date id="D162B63C1AE05EBF711E40DA62B8E255">2010</mods:date>
<mods:detail id="ECEBA40A7033E40128F0A4978D5AB7F5" type="volume">
<mods:number id="DF9CA98E4AF538509D3926FA3E2DE6E7">2635</mods:number>
</mods:detail>
<mods:extent id="7741A5EE55BB5091F3D60B1782C2307F" unit="page">
<mods:start id="87BB826A5AA78F4F5E866011C9765E69">1</mods:start>
<mods:end id="21B5D4BE02CFF798D5F1CA6A292AF022">31</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="EACAE659E3EFF90E562212548B41192C">journal article</mods:classification>
<mods:identifier id="327621BB183A93CE537CFB5F8870D6BA" type="DOI">10.5281/zenodo.198427</mods:identifier>
<mods:identifier id="B077FF886F7C050644574642903039E1" type="GBIF-Dataset">1c32eb21-6978-41da-b589-c98453d4f572</mods:identifier>
<mods:identifier id="126E6D2143FFA76AA8CD61B461DDE186" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="9FDC4EDF20CF34E0EF643B73F5BD058C" type="Zenodo-Dep">198427</mods:identifier>
</mods:mods>
<treatment id="03CEA91F8C29B90DA2C062C5FE04F7EB" ID-DOI="http://doi.org/10.5281/zenodo.6205304" ID-GBIF-Taxon="119399274" ID-Zenodo-Dep="6205304" LSID="urn:lsid:plazi:treatment:03CEA91F8C29B90DA2C062C5FE04F7EB" httpUri="http://treatment.plazi.org/id/03CEA91F8C29B90DA2C062C5FE04F7EB" lastPageId="17" lastPageNumber="18" pageId="14" pageNumber="15">
<subSubSection id="C37D4B828C29B912A2C062C5FE55F1B5" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="8BD818098C29B912A2C062C5FD7BF1E8" blockId="14.[151,675,1189,1250]" box="[151,675,1189,1215]" pageId="14" pageNumber="15">
<heading id="D090AF658C29B912A2C062C5FD7BF1E8" bold="true" box="[151,675,1189,1215]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<emphasis id="B913C41B8C29B912A2C062C5FD7BF1E8" bold="true" box="[151,675,1189,1215]" pageId="14" pageNumber="15">
<taxonomicName id="4C67638A8C29B912A2C062C5FD82F1E8" ID-CoL="49RW9" authority="Short &amp; Garcia" authorityName="Short &amp; Garcia" box="[151,602,1189,1215]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="meridensis" status="sp. nov.">
<emphasis id="B913C41B8C29B912A2C062C5FE57F1E8" bold="true" box="[151,399,1189,1215]" italics="true" pageId="14" pageNumber="15">Oocyclus meridensis</emphasis>
Short &amp; García
</taxonomicName>
<taxonomicNameLabel id="A22079608C29B912A03762C5FD7BF1E8" box="[608,675,1189,1215]" pageId="14" pageNumber="15" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD818098C29B912A2C062A8FE55F1B5" blockId="14.[151,675,1189,1250]" box="[151,397,1224,1250]" pageId="14" pageNumber="15">
(
<figureCitation id="135C048C8C29B912A2C862A8FF28F1B5" box="[159,240,1224,1250]" captionStart="FIGURE 3" captionStartId="7.[151,255,1852,1876]" captionTargetBox="[158,1416,651,1818]" captionTargetId="figure@7.[151,1436,642,1830]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal and lateral habitus of Oocyclus species found in the Venezuelan Andes region. A) O. meridensis sp. n.; B) O. trujillo sp. n.; C) O. andinus sp. n.; D) O. pico sp. n. Scale bar = 2.0 mm." httpUri="https://zenodo.org/record/198430/files/figure.png" pageId="14" pageNumber="15">Figs. 3</figureCitation>
A, 5G, 7, 9A)
</paragraph>
</subSubSection>
<subSubSection id="C37D4B828C29B912A2C06372FB93F390" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="8BD818098C29B912A2C06372FB93F390" blockId="14.[151,1437,1298,1996]" pageId="14" pageNumber="15">
<emphasis id="B913C41B8C29B912A2C06372FC8CF07B" bold="true" box="[151,852,1298,1324]" pageId="14" pageNumber="15">
<typeStatus id="54DCA6AB8C29B912A2C06372FF0DF07B" box="[151,213,1298,1324]" pageId="14" pageNumber="15">Type</typeStatus>
material.
<typeStatus id="54DCA6AB8C29B912A3006372FE16F07B" box="[343,462,1298,1324]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F37058998C29B912A38B6372FD40F07B" box="[476,664,1298,1324]" name="Venezuela" pageId="14" pageNumber="15">VENEZUELA</collectingCountry>
: Mérida State:
</emphasis>
male, “VENZUELA: Mérida State/
<geoCoordinate id="EE537ECE8C29B912A75E6372FA4FF07B" box="[1289,1431,1298,1324]" direction="north" orientation="latitude" pageId="14" pageNumber="15" precision="1" value="8.574817">8°34.489’N</geoCoordinate>
,
<geoCoordinate id="EE537ECE8C29B912A2C06358FEE4F005" box="[151,316,1336,1362]" direction="west" orientation="longitude" pageId="14" pageNumber="15" precision="1" value="-71.33608">71°20.165’W</geoCoordinate>
;
<quantity id="4C9FB5EC8C29B912A31E6358FE7CF005" box="[329,420,1336,1362]" metricMagnitude="3" metricUnit="m" metricValue="1.833" pageId="14" pageNumber="15" unit="m" value="1833.0">1833 m</quantity>
/ ca.
<quantity id="4C9FB5EC8C29B912A3B76358FDC6F005" box="[480,542,1336,1362]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="15" unit="km" value="1.0">1 km</quantity>
E. Jaji;
<date id="FFD93EC98C29B912A0D46358FD2CF005" box="[643,756,1336,1362]" pageId="14" pageNumber="15" value="2006-01-16">16.i.2006</date>
/ stream &amp; wet cement seepage/ AS-06-036; leg. A.E.Z. Short”, “[barcode]/ SM0828517/ KUNHM-ENT”, “
<typeStatus id="54DCA6AB8C29B912A0AA633DFC43F020" box="[765,923,1373,1399]" pageId="14" pageNumber="15" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="4C67638A8C29B912A1FA633DFB9CF021" box="[941,1092,1373,1398]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B913C41B8C29B912A1FA633DFB9CF021" box="[941,1092,1373,1398]" italics="true" pageId="14" pageNumber="15">OOCYCLUS</emphasis>
</taxonomicName>
/
<taxonomicName id="4C67638A8C29B912A603633DFB0CF021" box="[1108,1236,1373,1398]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="meridensis" status="sp. nov.">
<emphasis id="B913C41B8C29B912A603633DFB0CF021" box="[1108,1236,1373,1398]" italics="true" pageId="14" pageNumber="15">meridensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22079608C29B912A68B633DFACAF020" box="[1244,1298,1373,1399]" pageId="14" pageNumber="15" rank="species">sp.n.</taxonomicNameLabel>
/ des.
<bibRefCitation id="EFF665F88C29B912A70B633DFE8CF0CB" author="Short" pageId="14" pageNumber="15" refString="Hall, W. E. &amp; Short, A. E. Z. (2010) A new species of Hydroscapha LeConte from northwestern Venezuela (Coleoptera: Myxophaga: Hydroscaphidae). Zootaxa, 2358, 49 - 56." type="journal article" year="2010">Short &amp; García 2010</bibRefCitation>
” (
<collectionCode id="ED7680CC8C29B912A32663E2FE1FF0CB" LSID="urn:lsid:biocol.org:col:33834" box="[369,455,1410,1436]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="14" pageNumber="15">MIZA</collectionCode>
).
<typeStatus id="54DCA6AB8C29B912A38B63E2FDB9F0CB" box="[476,609,1410,1436]" pageId="14" pageNumber="15" type="paratype">
<emphasis id="B913C41B8C29B912A38B63E2FDB9F0CB" bold="true" box="[476,609,1410,1436]" pageId="14" pageNumber="15">Paratypes</emphasis>
</typeStatus>
(171 exs.):
<emphasis id="B913C41B8C29B912A0AE63E2FB50F0CB" bold="true" box="[761,1160,1410,1436]" pageId="14" pageNumber="15">
<collectingCountry id="F37058998C29B912A0AE63E2FC64F0CB" box="[761,956,1410,1436]" name="Venezuela" pageId="14" pageNumber="15">VENEZUELA</collectingCountry>
: Barinas State:
</emphasis>
<quantity id="4C9FB5EC8C29B912A6C763E2FB07F0CB" box="[1168,1247,1410,1436]" metricMagnitude="4" metricUnit="m" metricValue="2.4" pageId="14" pageNumber="15" unit="km" value="24.0">24 km</quantity>
NW Barinitas,
<date id="FFD93EC98C29B912A2C063C8FEC8F095" box="[151,272,1448,1474]" pageId="14" pageNumber="15" value="1969-02-23">23.ii.1969</date>
, leg. P. &amp; P. Spangler (19 exs.,
<collectionCode id="ED7680CC8C29B912A0D463C8FD05F095" LSID="urn:lsid:biocol.org:col:34871" box="[643,733,1448,1474]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="15">USNM</collectionCode>
);
<emphasis id="B913C41B8C29B912A0A563C7FC46F096" bold="true" box="[754,926,1447,1473]" pageId="14" pageNumber="15">Mérida State:</emphasis>
Mérida, Rio Milla, Mérida Zoo,
<date id="FFD93EC98C29B912A77263C8FA40F095" box="[1317,1432,1448,1474]" pageId="14" pageNumber="15" value="2006-01-14">14.i.2006</date>
, wet rock/seeps, leg. A.E.Z. Short, AS-06-026 (1 ex.); same data but leg. M. García (19 exs.); Along road between Jaji &amp; Mérida, small waterfall and stream pools,
<geoCoordinate id="EE537ECE8C29B912A16D6392FC65F35B" box="[826,957,1522,1548]" direction="north" orientation="latitude" pageId="14" pageNumber="15" precision="1" value="8.58515">8.58515°N</geoCoordinate>
,
<geoCoordinate id="EE537ECE8C29B912A19F6392FB85F35B" box="[968,1117,1522,1548]" direction="west" orientation="longitude" pageId="14" pageNumber="15" precision="1" value="-71.30103">71.30103°W</geoCoordinate>
,
<quantity id="4C9FB5EC8C29B912A63E6392FB1DF35B" box="[1129,1221,1522,1548]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="14" pageNumber="15" unit="m" value="1750.0">1750 m</quantity>
,
<date id="FFD93EC98C29B912A6876392FA99F35B" box="[1232,1345,1522,1548]" pageId="14" pageNumber="15" value="2006-01-16">16.i.2006</date>
, leg. A. Short &amp; P. Torres, AS-06-034, (1 ex.); same data as
<typeStatus id="54DCA6AB8C29B912A1506078FCB7F365" box="[775,879,1560,1586]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
(102 exs.); ca.
<quantity id="4C9FB5EC8C29B912A67F6078FBBEF365" box="[1064,1126,1560,1586]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="15" unit="km" value="2.0">2 km</quantity>
S. La Azulita, Cascada la Palmita,
<date id="FFD93EC98C29B912A354605DFE56F300" box="[259,398,1597,1623]" pageId="14" pageNumber="15" value="2009-07-21">21.vii.2009</date>
, leg. Short et al., VZ09-0721-01A (15 exs.);
<emphasis id="B913C41B8C29B912A1E5605DFBB1F300" bold="true" box="[946,1129,1597,1623]" pageId="14" pageNumber="15">Táchira State:</emphasis>
Chorro el Indio National Park,
<geoCoordinate id="EE537ECE8C29B912A2886002FE55F32B" box="[223,397,1634,1660]" direction="north" orientation="latitude" pageId="14" pageNumber="15" precision="1" value="7.7232804">7°43’23.81”N</geoCoordinate>
, 72°
<date id="FFD93EC98C29B912A3946002FDF7F32B" box="[451,559,1634,1660]" pageId="14" pageNumber="15" value="13-11-31">11.31.13</date>
”W,
<quantity id="4C9FB5EC8C29B912A0326002FD1CF32B" box="[613,708,1634,1660]" metricMagnitude="3" metricUnit="m" metricValue="1.225" pageId="14" pageNumber="15" unit="m" value="1225.0">1225 m</quantity>
,
<date id="FFD93EC98C29B912A0856002FC87F32B" box="[722,863,1634,1660]" pageId="14" pageNumber="15" value="2009-07-16">16.vii.2009</date>
, leg. Short &amp; García, wet rock and rock seeps, VZ09-0716-01A, (11 exs.);
<emphasis id="B913C41B8C29B912A38A60E7FD53F3F6" bold="true" box="[477,651,1671,1697]" pageId="14" pageNumber="15">
<taxonomicName id="4C67638A8C29B912A38A60E7FDE5F3F6" box="[477,573,1671,1697]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="28" phylum="Arthropoda" rank="species" species="trujillo" status="sp. nov.">Trujillo</taxonomicName>
State:
</emphasis>
E. of Pampan,
<geoCoordinate id="EE537ECE8C29B912A16960E8FC12F3F5" box="[830,970,1672,1698]" direction="north" orientation="latitude" pageId="14" pageNumber="15" precision="1" value="9.46325">9°27.795’N</geoCoordinate>
,
<geoCoordinate id="EE537ECE8C29B912A18160E8FBADF3F5" box="[982,1141,1672,1698]" direction="west" orientation="longitude" pageId="14" pageNumber="15" precision="1" value="-70.41893">70°25.136’W</geoCoordinate>
,
<quantity id="4C9FB5EC8C29B912A6D760E8FB03F3F5" box="[1152,1243,1672,1698]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="14" pageNumber="15" unit="m" value="1100.0">1100 m</quantity>
, rock seep along road,
<date id="FFD93EC98C29B912A28D60CDFE92F390" box="[218,330,1709,1735]" pageId="14" pageNumber="15" value="2009-01-21">21.i.2009</date>
, leg. A. Short, M. García, J. Camacho, VZ09-0121-
<date id="FFD93EC98C29B912A1FB60CDFC07F390" box="[940,991,1709,1735]" pageId="14" pageNumber="15" value="2005-10">05X</date>
, (3 exs.).
</paragraph>
</subSubSection>
<subSubSection id="C37D4B828C29B912A29260B2FAAEF2D5" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="8BD818098C29B912A29260B2FAAEF2D5" blockId="14.[151,1437,1298,1996]" pageId="14" pageNumber="15">
<emphasis id="B913C41B8C29B912A29260B2FE93F3BB" bold="true" box="[197,331,1746,1772]" pageId="14" pageNumber="15">Diagnosis.</emphasis>
Larger species (
<figureCitation id="135C048C8C29B912A04E60B2FDBBF3BB" box="[537,611,1746,1772]" captionStart="FIGURE 3" captionStartId="7.[151,255,1852,1876]" captionTargetBox="[158,1416,651,1818]" captionTargetId="figure@7.[151,1436,642,1830]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal and lateral habitus of Oocyclus species found in the Venezuelan Andes region. A) O. meridensis sp. n.; B) O. trujillo sp. n.; C) O. andinus sp. n.; D) O. pico sp. n. Scale bar = 2.0 mm." httpUri="https://zenodo.org/record/198430/files/figure.png" pageId="14" pageNumber="15">Fig. 3</figureCitation>
A). Posterolateral corners of pronotum spinose. Procoxae strongly spinose. Elytra with rows of larger systematic punctures, with first row very distinct, continuous, and bearing erect setae; the lateral rows also very distinct. Elytral suture raised at apex. Elytral margins explanate, without fringe of yellow setae. This species is most similar to
<taxonomicName id="4C67638A8C29B912A1756123FC7EF20B" box="[802,934,1859,1884]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="andinus">
<emphasis id="B913C41B8C29B912A1756123FC7EF20B" box="[802,934,1859,1884]" italics="true" pageId="14" pageNumber="15">
<collectingCountry id="F37058998C29B912A1756123FCE3F20B" box="[802,827,1859,1884]" name="Iceland" pageId="14" pageNumber="15">O</collectingCountry>
. andinus
</emphasis>
</taxonomicName>
and
<taxonomicName id="4C67638A8C29B912A1B66123FBAFF20B" box="[993,1143,1859,1884]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="substillus">
<emphasis id="B913C41B8C29B912A1B66123FBAFF20B" box="[993,1143,1859,1884]" italics="true" pageId="14" pageNumber="15">
<collectingCountry id="F37058998C29B912A1B66123FC22F20B" box="[993,1018,1859,1884]" name="Iceland" pageId="14" pageNumber="15">O</collectingCountry>
. substillus
</emphasis>
</taxonomicName>
, two other large species with spinose posterolateral pronotal corners. Refer to the diagnoses of those species for further discussion.
</paragraph>
</subSubSection>
<subSubSection id="C37D4B828C29B90DA29261EDFBEAF48B" lastPageId="17" lastPageNumber="18" pageId="14" pageNumber="15" type="description">
<paragraph id="8BD818098C29B912A29261EDFF2BF29B" blockId="14.[151,1437,1298,1996]" pageId="14" pageNumber="15">
<emphasis id="B913C41B8C29B912A29261EDFEBBF2F0" bold="true" box="[197,355,1933,1959]" pageId="14" pageNumber="15">Description.</emphasis>
<emphasis id="B913C41B8C29B912A33B61EDFDC6F2F1" box="[364,542,1933,1958]" italics="true" pageId="14" pageNumber="15">Size and form.</emphasis>
Body length =
<quantity id="4C9FB5EC8C29B912A08861EDFCAFF2F0" box="[735,887,1933,1959]" metricMagnitude="-3" metricUnit="m" metricValue="4.85" metricValueMax="5.5" metricValueMin="4.2" pageId="14" pageNumber="15" unit="mm" value="4.85" valueMax="5.5" valueMin="4.2">4.2–5.5 mm</quantity>
. TL/GW = 1.62. Body broadly oval, slightly convex.
</paragraph>
<caption id="DF1848818C28B913A2C061E8FEDEF288" httpUri="https://zenodo.org/record/198432/files/figure.png" pageId="15" pageNumber="16" targetBox="[174,1419,194,1908]" targetPageId="15">
<paragraph id="8BD818098C28B913A2C061E8FEDEF288" blockId="15.[151,1436,1928,2015]" pageId="15" pageNumber="16">
<emphasis id="B913C41B8C28B913A2C061E8FEC1F2F7" bold="true" box="[151,281,1928,1952]" pageId="15" pageNumber="16">FIGURE 5.</emphasis>
Aedeagi of
<taxonomicName id="4C67638A8C28B913A3CB61E9FE27F2F7" box="[412,511,1929,1952]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B913C41B8C28B913A3CB61E9FE27F2F7" box="[412,511,1929,1952]" italics="true" pageId="15" pageNumber="16">Oocyclus</emphasis>
</taxonomicName>
species. A)
<taxonomicName id="4C67638A8C28B913A0D561E9FD33F2F7" box="[642,747,1928,1952]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="pittieri" status="sp. nov.">
<emphasis id="B913C41B8C28B913A0D561E9FD4DF2F7" box="[642,661,1929,1952]" italics="true" pageId="15" pageNumber="16">O</emphasis>
.
<emphasis id="B913C41B8C28B913A0F561E9FD33F2F7" box="[674,747,1929,1952]" italics="true" pageId="15" pageNumber="16">pittieri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22079608C28B913A0A661E8FCF5F2F7" box="[753,813,1928,1952]" pageId="15" pageNumber="16" rank="species">
<emphasis id="B913C41B8C28B913A0A661E8FCF5F2F7" bold="true" box="[753,813,1928,1952]" pageId="15" pageNumber="16">sp. n.</emphasis>
</taxonomicNameLabel>
, B)
<taxonomicName id="4C67638A8C28B913A10B61E9FBEAF2C8" authority="Sharp" authorityName="Sharp" box="[860,1074,1928,1952]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B913C41B8C28B913A10B61E9FCB7F2F7" box="[860,879,1929,1952]" italics="true" pageId="15" pageNumber="16">O</emphasis>
.
<emphasis id="B913C41B8C28B913A12B61E9FC32F2F7" box="[892,1002,1929,1952]" italics="true" pageId="15" pageNumber="16">maculatus</emphasis>
Sharp
</taxonomicName>
; C)
<emphasis id="B913C41B8C28B913A60A61E9FBA8F2F7" box="[1117,1136,1929,1952]" italics="true" pageId="15" pageNumber="16">O</emphasis>
. s
<emphasis id="B913C41B8C28B913A6DF61E9FB3BF2F7" box="[1160,1251,1929,1952]" italics="true" pageId="15" pageNumber="16">ubstillus</emphasis>
Short &amp; Perkins; D)
<taxonomicName id="4C67638A8C28B913A2ED61C9FEFFF297" box="[186,295,1960,1984]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="floccus" status="sp. nov.">
<emphasis id="B913C41B8C28B913A2ED61C9FF15F297" box="[186,205,1961,1984]" italics="true" pageId="15" pageNumber="16">O</emphasis>
.
<emphasis id="B913C41B8C28B913A28C61C9FEFFF297" box="[219,295,1961,1984]" italics="true" pageId="15" pageNumber="16">floccus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22079608C28B913A37A61C8FEB1F297" box="[301,361,1960,1984]" pageId="15" pageNumber="16" rank="species">
<emphasis id="B913C41B8C28B913A37A61C8FEB1F297" bold="true" box="[301,361,1960,1984]" pageId="15" pageNumber="16">sp. n.</emphasis>
</taxonomicNameLabel>
; E)
<taxonomicName id="4C67638A8C28B913A3CF61C9FDDAF297" box="[408,514,1960,1984]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="trujillo" status="sp. nov.">
<emphasis id="B913C41B8C28B913A3CF61C9FE73F297" box="[408,427,1961,1984]" italics="true" pageId="15" pageNumber="16">O</emphasis>
.
<emphasis id="B913C41B8C28B913A3EF61C9FDDAF297" box="[440,514,1961,1984]" italics="true" pageId="15" pageNumber="16">trujillo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22079608C28B913A05E61C8FD9DF297" box="[521,581,1960,1984]" pageId="15" pageNumber="16" rank="species">
<emphasis id="B913C41B8C28B913A05E61C8FD9DF297" bold="true" box="[521,581,1960,1984]" pageId="15" pageNumber="16">sp. n.</emphasis>
</taxonomicNameLabel>
; F)
<taxonomicName id="4C67638A8C28B913A02461C9FD30F297" box="[627,744,1960,1984]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="andinus" status="sp. nov.">
<emphasis id="B913C41B8C28B913A02461C9FD5EF297" box="[627,646,1961,1984]" italics="true" pageId="15" pageNumber="16">O</emphasis>
.
<emphasis id="B913C41B8C28B913A0C361C9FD30F297" box="[660,744,1961,1984]" italics="true" pageId="15" pageNumber="16">andinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22079608C28B913A0B861C8FCF3F297" box="[751,811,1960,1984]" pageId="15" pageNumber="16" rank="species">
<emphasis id="B913C41B8C28B913A0B861C8FCF3F297" bold="true" box="[751,811,1960,1984]" pageId="15" pageNumber="16">sp. n.</emphasis>
</taxonomicNameLabel>
; G)
<taxonomicName id="4C67638A8C28B913A10B61C9FC29F297" box="[860,1009,1960,1984]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="meridensis" status="sp. nov.">
<emphasis id="B913C41B8C28B913A10B61C9FCB7F297" box="[860,879,1961,1984]" italics="true" pageId="15" pageNumber="16">O</emphasis>
.
<emphasis id="B913C41B8C28B913A12A61C9FC29F297" box="[893,1009,1961,1984]" italics="true" pageId="15" pageNumber="16">meridensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22079608C28B913A1AF61C8FBEBF297" box="[1016,1075,1960,1984]" pageId="15" pageNumber="16" rank="species">
<emphasis id="B913C41B8C28B913A1AF61C8FBEBF297" bold="true" box="[1016,1075,1960,1984]" pageId="15" pageNumber="16">sp. n.</emphasis>
</taxonomicNameLabel>
; H)
<taxonomicName id="4C67638A8C28B913A63261C9FB37F297" box="[1125,1263,1960,1984]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="sapphirus" status="sp. nov.">
<emphasis id="B913C41B8C28B913A63261C9FBA0F297" box="[1125,1144,1961,1984]" italics="true" pageId="15" pageNumber="16">O</emphasis>
.
<emphasis id="B913C41B8C28B913A6D161C9FB37F297" box="[1158,1263,1961,1984]" italics="true" pageId="15" pageNumber="16">sapphirus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22079608C28B913A6A161C8FAEAF297" box="[1270,1330,1960,1984]" pageId="15" pageNumber="16" rank="species">
<emphasis id="B913C41B8C28B913A6A161C8FAEAF297" bold="true" box="[1270,1330,1960,1984]" pageId="15" pageNumber="16">sp. n.</emphasis>
</taxonomicNameLabel>
Scale bar = 0.2 mm.
</paragraph>
</caption>
<caption id="DF1848818C37B90CA2C0627DFEB6F103" httpUri="https://zenodo.org/record/198433/files/figure.png" pageId="16" pageNumber="17" targetBox="[155,1436,199,1024]" targetPageId="16">
<paragraph id="8BD818098C37B90CA2C0627DFEB6F103" blockId="16.[151,1436,1053,1108]" pageId="16" pageNumber="17">
<emphasis id="B913C41B8C37B90CA2C0627DFEC2F162" bold="true" box="[151,282,1053,1077]" pageId="16" pageNumber="17">FIGURE 6.</emphasis>
Aedeagi of
<taxonomicName id="4C67638A8C37B90CA3CB627EFE27F162" box="[412,511,1054,1077]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="B913C41B8C37B90CA3CB627EFE27F162" box="[412,511,1054,1077]" italics="true" pageId="16" pageNumber="17">Oocyclus</emphasis>
</taxonomicName>
species. A)
<taxonomicName id="4C67638A8C37B90CA0D3627EFD04F162" box="[644,732,1053,1077]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="petra" status="sp. nov.">
<emphasis id="B913C41B8C37B90CA0D3627EFD4FF162" box="[644,663,1054,1077]" italics="true" pageId="16" pageNumber="17">O</emphasis>
.
<emphasis id="B913C41B8C37B90CA0F3627EFD04F162" box="[676,732,1054,1077]" italics="true" pageId="16" pageNumber="17">petra</emphasis>
</taxonomicName>
<emphasis id="B913C41B8C37B90CA0B3627DFCC7F162" bold="true" box="[740,799,1053,1077]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="A22079608C37B90CA0B3627DFCC7F162" box="[740,799,1053,1077]" pageId="16" pageNumber="17" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, B)
<taxonomicName id="4C67638A8C37B90CA119627EFC6EF162" box="[846,950,1053,1077]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="galbus" status="sp. nov.">
<emphasis id="B913C41B8C37B90CA119627EFCB9F162" box="[846,865,1054,1077]" italics="true" pageId="16" pageNumber="17">O</emphasis>
.
<emphasis id="B913C41B8C37B90CA138627EFC6EF162" box="[879,950,1054,1077]" italics="true" pageId="16" pageNumber="17">galbus</emphasis>
</taxonomicName>
<emphasis id="B913C41B8C37B90CA1EA627DFC21F162" bold="true" box="[957,1017,1053,1077]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="A22079608C37B90CA1EA627DFC21F162" box="[957,1017,1053,1077]" pageId="16" pageNumber="17" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; C)
<taxonomicName id="4C67638A8C37B90CA67E627EFBAEF162" box="[1065,1142,1053,1077]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="pico" status="sp. nov.">
<emphasis id="B913C41B8C37B90CA67E627EFBE4F162" box="[1065,1084,1054,1077]" italics="true" pageId="16" pageNumber="17">O</emphasis>
.
<emphasis id="B913C41B8C37B90CA61E627EFBAEF162" box="[1097,1142,1054,1077]" italics="true" pageId="16" pageNumber="17">pico</emphasis>
</taxonomicName>
<emphasis id="B913C41B8C37B90CA629627DFB62F162" bold="true" box="[1150,1210,1053,1077]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="A22079608C37B90CA629627DFB62F162" box="[1150,1210,1053,1077]" pageId="16" pageNumber="17" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
D)
<taxonomicName id="4C67638A8C37B90CA6B3627EFA80F162" box="[1252,1368,1053,1077]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="bolivari" status="sp. nov.">
<emphasis id="B913C41B8C37B90CA6B3627EFB2FF162" box="[1252,1271,1054,1077]" italics="true" pageId="16" pageNumber="17">O</emphasis>
.
<emphasis id="B913C41B8C37B90CA753627EFA80F162" box="[1284,1368,1054,1077]" italics="true" pageId="16" pageNumber="17">bolivari</emphasis>
</taxonomicName>
<emphasis id="B913C41B8C37B90CA737627DFA44F162" bold="true" box="[1376,1436,1053,1077]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="A22079608C37B90CA737627DFA44F162" box="[1376,1436,1053,1077]" pageId="16" pageNumber="17" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
Scale bar = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BD818098C37B90CA29262E3FCB5F02A" blockId="16.[151,1437,1155,2002]" pageId="16" pageNumber="17">
<emphasis id="B913C41B8C37B90CA29262E3FED6F1CB" box="[197,270,1155,1180]" italics="true" pageId="16" pageNumber="17">Color.</emphasis>
Dorsum of head, pronotum, and elytra black, and usually covered with moderately iridescent green. Anterolateral margins of pronotum with white patch, approximately one-third the length of the margin. Elytra usually with layer of iridescent green maculae which often appear faint and with defuse borders. Maxillary and labial palps yellow. Mentum and stipes dark brown, similar in color to venter of head. Legs, epipleura, lateral margins of prosternum and lateral margins of abdominal ventrites 2–4 medium to light brown, with remainder of thoracic venter dark brown. Abdominal ventrites brown with the posterior margin (sometimes as much as posterior half) of each ventrite light brown to yellow.
</paragraph>
<paragraph id="8BD818098C37B90CA29263E9FE07F3F0" blockId="16.[151,1437,1155,2002]" pageId="16" pageNumber="17">
<emphasis id="B913C41B8C37B90CA29263E9FED5F0F5" box="[197,269,1417,1442]" italics="true" pageId="16" pageNumber="17">Head.</emphasis>
Ground punctation on labrum, clypeus and frons moderately coarse, distance between punctures 1.5–2.0× the width of one puncture. Systematic punctures on labrum consisting of several indistinct punctures, sometimes bearing short setae. Frons with irregular row of systematic punctures mesad of each eye, bearing fine setae. Clypeus with a few very indistinct systematic punctures along anterolateral margins, slightly larger than surrounding punctation. Maxillary palps short, slightly shorter to about as long as width of labrum; segment 2 slightly bulbous, apical segment slightly longer than penultimate. Labial palps two-thirds as long as width of mentum. Mentum quadrate, anterior margin slightly convex but not distinctly depressed, bearing scattered punctures.
</paragraph>
<paragraph id="8BD818098C37B90DA29260D3FEC6F4C5" blockId="16.[151,1437,1155,2002]" lastBlockId="17.[151,1437,152,700]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<emphasis id="B913C41B8C37B90CA29260D3FEFFF39B" box="[197,295,1715,1740]" italics="true" pageId="16" pageNumber="17">Thorax.</emphasis>
Ground punctation on pronotum and elytra composed of moderate fine, evenly distributed punctures. Pronotal systematic punctures with short fine setae, distinctly larger than ground punctures, anterior and posterior series each forming irregular field. Lateral margins of pronotum with sparse row of setiferous punctures; with punctures almost appearing absent, and bearing very fine, short setae which are almost not noticed. Posterolateral corners of pronotum strongly spinose. Sutural punctation represented by sparse, irregular band of small punctures; sutural interval distinctly raised in posterior third. Elytra with five rows of systematic punctures: row 1 represented as grove with dense, continuous row of erect setae on anterior two-thirds, giving a “Mohawk”-like appearance. Rows 2 and 3 each represented by a nearly regular series of coarse punctures usually bearing small recumbent setae. Punctures in rows 4 and 5 similar to 2 and 3 but irregular in distribution. Prosternum with strongly raised median carina; with two short spine-like setae at anterior margin of carina. Elevated process of mesoventrite slightly longer than wide, with 5–7 thickened, coarse spine-like setae. Metaventrite with oval glabrous area posteromedially, ca. slightly longer than wide, length of glabrous area slightly less than one-half length of metaventrite. Procoxae with dense, fine short pubescence and with distinct, very coarse spine-like setae. Protibiae each with 12–15 spine-like setae on dorsal face.
</paragraph>
<paragraph id="8BD818098C36B90DA29267FDFBEAF48B" blockId="17.[151,1437,152,700]" pageId="17" pageNumber="18">
<emphasis id="B913C41B8C36B90DA29267FDFEE5F4E1" box="[197,317,413,438]" italics="true" pageId="17" pageNumber="18">Abdomen.</emphasis>
Ventrites covered with moderately dense setae, the longest setae about as long as the longest setae surrounding the glabrous area of metaventrite. Aedeagus as in
<figureCitation id="135C048C8C36B90DA1E167A2FBCFF48B" box="[950,1047,450,476]" captionStart="FIGURE 5" captionStartId="15.[151,255,1928,1952]" captionTargetBox="[174,1419,194,1908]" captionTargetId="figure@15.[174,1420,194,1909]" captionTargetPageId="15" captionText="FIGURE 5. Aedeagi of Oocyclus species. A) O. pittieri sp. n., B) O. maculatus Sharp; C) O. s ubstillus Short &amp; Perkins; D) O. floccus sp. n.; E) O. trujillo sp. n.; F) O. andinus sp. n.; G) O. meridensis sp. n.; H) O. sapphirus sp. n. Scale bar = 0.2 mm." httpUri="https://zenodo.org/record/198432/files/figure.png" pageId="17" pageNumber="18">Figure 5</figureCitation>
G.
</paragraph>
</subSubSection>
<subSubSection id="C37D4B828C36B90DA2926787FC35F770" pageId="17" pageNumber="18" type="etymology">
<paragraph id="8BD818098C36B90DA2926787FC35F770" blockId="17.[151,1437,152,700]" pageId="17" pageNumber="18">
<emphasis id="B913C41B8C36B90DA2926787FE82F756" bold="true" box="[197,346,487,513]" pageId="17" pageNumber="18">Etymology.</emphasis>
<taxonomicName id="4C67638A8C36B90DA3346788FE35F756" box="[355,493,488,513]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="29" phylum="Arthropoda" rank="species" species="meridensis">
<emphasis id="B913C41B8C36B90DA3346788FE35F756" box="[355,493,488,513]" italics="true" pageId="17" pageNumber="18">Meridensis</emphasis>
</taxonomicName>
, an arbitrary combination of letters similar to the city and state of Mérida, “capital” of the Venezuelan Andes, where this species is known to occur.
</paragraph>
</subSubSection>
<subSubSection id="C37D4B828C36B90DA2926452FB76F71B" box="[197,1198,562,588]" pageId="17" pageNumber="18" type="distribution">
<paragraph id="8BD818098C36B90DA2926452FB76F71B" blockId="17.[151,1437,152,700]" box="[197,1198,562,588]" pageId="17" pageNumber="18">
<emphasis id="B913C41B8C36B90DA2926452FEBDF71B" bold="true" box="[197,357,562,588]" pageId="17" pageNumber="18">Distribution.</emphasis>
This species is known from localities along the Mérida Andes. (
<figureCitation id="135C048C8C36B90DA60E6452FB78F71B" box="[1113,1184,562,588]" captionStart="FIGURE 7" captionStartId="22.[151,255,1419,1443]" captionTargetBox="[151,1431,198,1376]" captionTargetId="figure@22.[151,1436,197,1398]" captionTargetPageId="22" captionText="FIGURE 7. Distribution of Oocyclus species in Venezuela." httpUri="https://zenodo.org/record/198434/files/figure.png" pageId="17" pageNumber="18">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37D4B828C36B90DA2926437FE04F7EB" pageId="17" pageNumber="18" type="discussion">
<paragraph id="8BD818098C36B90DA2926437FE04F7EB" blockId="17.[151,1437,152,700]" pageId="17" pageNumber="18">
<emphasis id="B913C41B8C36B90DA2926437FEE6F726" bold="true" box="[197,318,599,625]" pageId="17" pageNumber="18">Remarks.</emphasis>
This larger species has been found on rock seeps and other hygropetric habitats. The
<typeStatus id="54DCA6AB8C36B90DA7776438FA8BF725" box="[1312,1363,600,626]" pageId="17" pageNumber="18">type</typeStatus>
series (including more than half the specimens) was collected from a large expanse of wet cement covered in mud with a little moss (
<figureCitation id="135C048C8C36B90DA32764C2FE6EF7EB" box="[368,438,674,700]" captionStart="FIGURE 9" captionStartId="24.[151,258,1792,1816]" captionTargetBox="[150,1436,637,1765]" captionTargetId="figure@24.[145,1438,637,1765]" captionTargetPageId="24" captionText="FIGURE 9. Habitat of Oocyclus species. A) Mérida State, E. of Jaji, collecting event AS- 06 - 036, habitat and type locality for O. meridensis. B) Mérida State, Cascada de Bailadores, collecting event VZ 09 - 0718 - 02 B; type locality for O. andinus." httpUri="https://zenodo.org/record/198436/files/figure.png" pageId="17" pageNumber="18">Fig. 9</figureCitation>
A).
</paragraph>
</subSubSection>
</treatment>
</document>
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