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<document id="243776DA3EDFDF8A8E06DC79A31BE710" ID-DOI="10.1649/0010-065X-77.3.462" ID-ISSN="1938-4394" ID-Zenodo-Dep="13243309" ID-ZooBank="urn:lsid:zoobank.org:pub:86518A71-4AA0-46CF-955B-729FB6E5B07D" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722968277150" checkinUser="felipe" docAuthor="Ito, Naoya &amp; Powell, Gareth S." docDate="2023" docId="03C3CB76FF89FFA997AF0745FC64F9F8" docLanguage="en" docName="TheColeopBull.77.3.462-468.pdf" docOrigin="The Coleopterists Bulletin 77 (3)" docSource="http://dx.doi.org/10.1649/0010-065x-77.3.462" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Urophorus (Anophorus) cordiformis Ito and Powell 2023, new species" docType="treatment" docUuid="8993FF7B-216C-4DE6-AA91-076967301516" docUuidSource="ZooBank" docVersion="2" lastPageNumber="468" masterDocId="FFFAB30EFF8BFFAE976C023CFFCBFF99" masterDocTitle="A New Species of Urophorus (Anophorus) Kirejtshuk, 1990 (Coleoptera: Nitidulidae) from Ishigaki-Jima Island, Southern Ryukyu, Southwestern Japan" masterLastPageNumber="468" masterPageNumber="462" pageNumber="463" updateTime="1722971589281" updateUser="ExternalLinkService">
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<mods:title id="6E527B092036535627EC6E793A1F2AE0">A New Species of Urophorus (Anophorus) Kirejtshuk, 1990 (Coleoptera: Nitidulidae) from Ishigaki-Jima Island, Southern Ryukyu, Southwestern Japan</mods:title>
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<mods:name id="76A0FCAAB5B5B9D5FAC456CDA1B6AE25" type="personal">
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<mods:namePart id="5DC6E58113A1118D6FBC7DB8989D5770">Ito, Naoya</mods:namePart>
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<mods:namePart id="F07232F32103E32DBC17E289E7AF2B1C">Powell, Gareth S.</mods:namePart>
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<mods:title id="DC236B70676BEE3E49D0C228865C6972">The Coleopterists Bulletin</mods:title>
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<mods:date id="896C93296257E566DAFF5AF787719B27">2023</mods:date>
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<mods:number id="973534BB1BE8874F113A2B05625FF438">2023-09-29</mods:number>
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<subSubSection id="C37029EBFF89FFAC97AF0745FE1EF981" pageId="2" pageNumber="463" type="nomenclature">
<paragraph id="8BD57A60FF89FFAC97AF0745FE1EF981" blockId="2.[180,652,1401,1560]" pageId="2" pageNumber="463">
<heading id="D09DCD0CFF89FFAC97AF0745FDC8FA34" centered="true" fontSize="9" level="2" pageId="2" pageNumber="463" reason="2">
<emphasis id="B91EA672FF89FFAC97AF0745FDC8FA34" bold="true" pageId="2" pageNumber="463">
<taxonomicName id="4C6A01E3FF89FFAC97AF0745FE49FA35" authority="Ito and Powell" authorityName="Ito and Powell" authorityYear="2023" class="Insecta" family="Nitidulidae" genus="Urophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="463" phylum="Arthropoda" rank="species" species="cordiformis" status="sp. nov." subGenus="Anophorus">
<emphasis id="B91EA672FF89FFAC97AF0745FEE4FA16" bold="true" box="[195,303,1401,1423]" italics="true" pageId="2" pageNumber="463">Urophorus</emphasis>
(
<emphasis id="B91EA672FF89FFAC96510745FE67FA16" bold="true" box="[317,428,1401,1423]" italics="true" pageId="2" pageNumber="463">Anophorus</emphasis>
)
<emphasis id="B91EA672FF89FFAC96D60745FDE6FA16" bold="true" box="[442,557,1401,1423]" italics="true" pageId="2" pageNumber="463">cordiformis</emphasis>
Ito and Powell
</taxonomicName>
,
<taxonomicNameLabel id="A22D1B09FF89FFAC96E107ABFDC8FA34" box="[397,515,1431,1453]" pageId="2" pageNumber="463" rank="species">new species</taxonomicNameLabel>
</emphasis>
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zoobank.org/
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urn:lsid:zoobank.org:act:
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(Japanese name: Kaku-mune-kata-beni-deokisui) (
<figureCitation id="135166E5FF89FFAC961E043EFE06F981" box="[370,461,1538,1560]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStartId-0="3.[176,209,1668,1687]" captionStartId-1="4.[197,228,1392,1411]" captionStartId-2="5.[176,207,711,730]" captionStartId-3="5.[176,207,1535,1554]" captionStartId-4="6.[197,228,1231,1250]" captionTargetBox-0="[159,1145,213,1634]" captionTargetBox-1="[179,1180,197,1366]" captionTargetBox-2="[178,1164,202,691]" captionTargetBox-3="[151,1003,805,1518]" captionTargetBox-4="[179,1164,205,1211]" captionTargetId-0="figure-8@3.[151,1171,194,1651]" captionTargetId-1="figure-83@4.[173,1193,193,1375]" captionTargetId-2="figure-69@5.[151,1171,193,693]" captionTargetId-3="figure-97@5.[151,1171,805,1518]" captionTargetId-4="figure-251@6.[173,1193,193,1213]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="5" captionTargetPageId-4="6" captionText-0="Fig. 1. Urophorus (Anophorus) cordiformis, new species. AB) Male, dorsal and ventral view, respectively, CD) Female, dorsal and ventral view, respectively." captionText-1="Fig. 2. Urophorus (Anophorus) cordiformis, new species. A) Labrum, dorsal view, B) Labial palpi with mentum, ventral view, C) Right maxilla, ventral view, D) Right mandible, dorsal view, E) Head capsule, dorsal view, F) Right antenna, dorsal view, G) Abdominal sternite VI of male, ventral view, H) Abdominal sternite VII of male, ventral view, I) Spiculum gastrale, ventral view, J) Anal sclerite, ventral view, K) Abdominal sternite VI of female, ventral view, L) Abdominal sternite VII of female, ventral view." captionText-2="Fig. 3. Genitalia of Urophorus (Anophorus) cordiformis, new species. AC) Male genitalia, ventral, lateral, and dorsal views, respectively, D) Ovipositor, ventral view." captionText-3="Fig. 4. The mating posture of Urophorus (Anophorus) cordiformis, new species under laboratory conditions." captionText-4="Fig. 5. Geographic location of Ishigaki-jima Island." figureDoi-0="http://doi.org/10.5281/zenodo.13243313" figureDoi-1="http://doi.org/10.5281/zenodo.13243315" figureDoi-2="http://doi.org/10.5281/zenodo.13243317" figureDoi-3="http://doi.org/10.5281/zenodo.13243321" figureDoi-4="http://doi.org/10.5281/zenodo.13243323" httpUri-0="https://zenodo.org/record/13243313/files/figure.png" httpUri-1="https://zenodo.org/record/13243315/files/figure.png" httpUri-2="https://zenodo.org/record/13243317/files/figure.png" httpUri-3="https://zenodo.org/record/13243321/files/figure.png" httpUri-4="https://zenodo.org/record/13243323/files/figure.png" pageId="2" pageNumber="463">Figs. 15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37029EBFF89FFAC97A90411FC22FD8E" pageId="2" pageNumber="463" type="materials_examined">
<paragraph id="8BD57A60FF89FFAC97A90411FC22FD8E" blockId="2.[173,660,1580,1709]" lastBlockId="2.[706,1193,194,1709]" pageId="2" pageNumber="463">
<emphasis id="B91EA672FF89FFAC97A90411FEBAF9DB" bold="true" box="[197,369,1580,1602]" pageId="2" pageNumber="463">Type Specimens.</emphasis>
<materialsCitation id="3B02703DFF89FFAC96140411FD5BF935" collectingDate="2022-06-27" collectingDateMax="2022-06-29" collectingDateMin="2022-06-27" collectionCode="BLKU" country="Japan" location="Okinawa Pref." pageId="2" pageNumber="463" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa" typeStatus="holotype">
<typeStatus id="54D1C4C2FF89FFAC96140411FE1EF9DA" box="[376,469,1581,1603]" pageId="2" pageNumber="463" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D6CB1E9FF89FFAC96B30411FE38F9DB" box="[479,499,1581,1602]" count="1" pageId="2" pageNumber="463" type="male"></specimenCount>
,
<collectingCountry id="F37D3AF0FF89FFAC96920411FDF3F9DB" box="[510,568,1581,1602]" name="Japan" pageId="2" pageNumber="463">Japan</collectingCountry>
: “[
<collectingCountry id="F37D3AF0FF89FFAC953E0411FD5BF9DB" box="[594,656,1581,1602]" name="Japan" pageId="2" pageNumber="463">Japan</collectingCountry>
: Ishigaki-jima Is.]/ Hirakubo, Ishigaki City,/
<collectingRegion id="49AEB482FF89FFAC952C047BFF17F9E1" country="Japan" name="Okinawa" pageId="2" pageNumber="463">Okinawa Pref.</collectingRegion>
,/
<date id="FFD45CA0FF89FFAC9787045EFEBCF9EE" box="[235,375,1634,1655]" pageId="2" pageNumber="463" value="2022-06-27" valueMax="2022-06-29" valueMin="2022-06-27">
<collectingDate id="EF90A548FF89FFAC9787045EFEBCF9EE" box="[235,375,1634,1655]" pageId="2" pageNumber="463" value="2022-06-27" valueMax="2022-06-29" valueMin="2022-06-27">2729.VI.2022</collectingDate>
</date>
,/ Naoya ITO leg./
<collectingMethod id="522B0277FF89FFAC9548045EFD58F9EE" box="[548,659,1634,1656]" pageId="2" pageNumber="463">by fruit trap</collectingMethod>
(pineapple)//
<typeStatus id="54D1C4C2FF89FFAC964B0441FE6AF90B" box="[295,417,1661,1682]" pageId="2" pageNumber="463" type="holotype">HOLOTYPE</typeStatus>
//
<taxonomicName id="4C6A01E3FF89FFAC96D80441FED7F935" authorityName="Ito &amp; Powell" authorityYear="2023" class="Insecta" family="Nitidulidae" genus="Urophorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="463" phylum="Arthropoda" rank="species" species="cordiformis" status="sp. nov." subGenus="Anophorus">
<emphasis id="B91EA672FF89FFAC96D80441FDD2F90B" box="[436,537,1661,1682]" italics="true" pageId="2" pageNumber="463">Urophorus</emphasis>
(
<emphasis id="B91EA672FF89FFAC95490441FD40F90B" box="[549,651,1661,1682]" italics="true" pageId="2" pageNumber="463">Anophorus</emphasis>
)
<emphasis id="B91EA672FF89FFAC97C104ABFED7F935" box="[173,284,1687,1708]" italics="true" pageId="2" pageNumber="463">cordiformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D1B09FF89FFAC964E04A4FEA0F935" box="[290,363,1687,1709]" pageId="2" pageNumber="463" rank="species">sp. nov.</taxonomicNameLabel>
/ Ito et Powell, 2023” (
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)
</materialsCitation>
.
<emphasis id="B91EA672FF89FFAC95AF02FEFCE5FF4E" bold="true" box="[707,814,194,215]" pageId="2" pageNumber="463">
<typeStatus id="54D1C4C2FF89FFAC95AF02FEFCE5FF4E" box="[707,814,194,215]" pageId="2" pageNumber="463" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="9D6CB1E9FF89FFAC945302FEFCBCFF4E" box="[831,887,194,216]" count="15" pageId="2" pageNumber="463" type="male">15 ♂</specimenCount>
,
<specimenCount id="9D6CB1E9FF89FFAC94E802FEFC67FF4E" box="[900,940,194,215]" count="8" pageId="2" pageNumber="463" type="female">8 ♀</specimenCount>
): [Ishigaki-jima Is.]
<materialsCitation id="3B02703DFF89FFAC931002FEFCDDFE95" collectingDate="2013-10-16" collectionCode="BLKU" collectorName="Hirae &amp; N. Ito" country="Japan" pageId="2" pageNumber="463" specimenCount="2" specimenCount-male="2" stateProvince="Okinawa" typeStatus="paratype">
<specimenCount id="9D6CB1E9FF89FFAC931002FEFB6FFF4E" box="[1148,1188,194,215]" count="2" pageId="2" pageNumber="463" type="male">2 ♂</specimenCount>
,
<collectorName id="269F1FB6FF89FFAC95AF02E1FD31FF6A" box="[707,762,221,243]" pageId="2" pageNumber="463">Hirae</collectorName>
,
<date id="FFD45CA0FF89FFAC946F02E0FCA3FF6B" box="[771,872,220,242]" pageId="2" pageNumber="463" value="2013-10-16">
<collectingDate id="EF90A548FF89FFAC946F02E0FCA3FF6B" box="[771,872,220,242]" pageId="2" pageNumber="463" value="2013-10-16">16.X.2013</collectingDate>
</date>
,
<collectorName id="269F1FB6FF89FFAC941E02E1FC61FF6B" box="[882,938,221,242]" pageId="2" pageNumber="463">N. Ito</collectorName>
leg.,
<collectingMethod id="522B0277FF89FFAC94B202E1FB9BFF6B" box="[990,1104,221,243]" pageId="2" pageNumber="463">by fruit trap</collectingMethod>
(banana) (
<collectionCode id="ED7BE2A5FF89FFAC95A502CBFCC5FE95" box="[713,782,247,268]" pageId="2" pageNumber="463">BLKU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B02703DFF89FFAC947202CBFBCCFEBE" collectingDate="2014-03-24" collectingDateMax="2014-03-28" collectingDateMin="2014-03-24" collectionCode="BLKU" collectorName="N. Ito" country="Japan" pageId="2" pageNumber="463" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Okinawa" typeStatus="paratype">
<specimenCount id="9D6CB1E9FF89FFAC947202CBFC89FE94" box="[798,834,247,269]" count="1" pageId="2" pageNumber="463" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D6CB1E9FF89FFAC942702CBFCA5FE94" box="[843,878,247,269]" count="1" pageId="2" pageNumber="463" type="female">1 ♀</specimenCount>
, ditto,
<date id="FFD45CA0FF89FFAC94C202CBFBF6FE95" box="[942,1085,247,268]" pageId="2" pageNumber="463" value="2014-03-24" valueMax="2014-03-28" valueMin="2014-03-24">
<collectingDate id="EF90A548FF89FFAC94C202CBFBF6FE95" box="[942,1085,247,268]" pageId="2" pageNumber="463" value="2014-03-24" valueMax="2014-03-28" valueMin="2014-03-24">2428.III.2014</collectingDate>
</date>
,
<collectorName id="269F1FB6FF89FFAC932902C4FBB0FE94" box="[1093,1147,248,269]" pageId="2" pageNumber="463">N. Ito</collectorName>
leg.,
<collectingMethod id="522B0277FF89FFAC95AF032EFCFCFEBE" box="[707,823,274,296]" pageId="2" pageNumber="463">by fruit trap</collectingMethod>
(pineapple) (
<collectionCode id="ED7BE2A5FF89FFAC94D6032EFBCAFEBE" box="[954,1025,274,295]" pageId="2" pageNumber="463">BLKU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B02703DFF89FFAC937E032EFCB3FEC5" collectingDate="2014-03-27" collectingDateMax="2014-03-28" collectingDateMin="2014-03-24" collectionCode="BLKU" collectorName="N. Ito" country="Japan" pageId="2" pageNumber="463" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Okinawa" typeStatus="paratype">
<specimenCount id="9D6CB1E9FF89FFAC937E032EFBF2FEBE" box="[1042,1081,274,295]" count="1" pageId="2" pageNumber="463" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D6CB1E9FF89FFAC9328032EFBA2FEBE" box="[1092,1129,274,295]" count="1" pageId="2" pageNumber="463" type="female">1 ♀</specimenCount>
, ditto,
<date id="FFD45CA0FF89FFAC95AF0311FCFEFEDB" box="[707,821,301,322]" pageId="2" pageNumber="463" value="2014-03-27">
<collectingDate id="EF90A548FF89FFAC95AF0311FCFEFEDB" box="[707,821,301,322]" pageId="2" pageNumber="463" value="2014-03-27">27.III.2014</collectingDate>
</date>
,
<collectorName id="269F1FB6FF89FFAC94290311FC4FFEDB" box="[837,900,301,322]" pageId="2" pageNumber="463">N. Ito</collectorName>
leg., by tree sap (
<taxonomicName id="4C6A01E3FF89FFAC93220311FCDEFEC5" class="Magnoliopsida" family="Oleaceae" genus="Fraxinus" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="463" phylum="Tracheophyta" rank="species" species="griffithii">
<emphasis id="B91EA672FF89FFAC93220311FCDEFEC5" italics="true" pageId="2" pageNumber="463">Fraxinus griffithii</emphasis>
</taxonomicName>
) (
<collectionCode id="ED7BE2A5FF89FFAC9446037BFCBAFEC5" box="[810,881,327,348]" pageId="2" pageNumber="463">BLKU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B02703DFF89FFAC94EF037BFBB3FEEE" collectingDate="2014-03-28" collectingDateMax="2014-03-28" collectingDateMin="2014-03-24" collectionCode="BLKU" collectorName="N. Ito" country="Japan" pageId="2" pageNumber="463" specimenCount="1" specimenCount-female="1" stateProvince="Okinawa" typeStatus="paratype">
<specimenCount id="9D6CB1E9FF89FFAC94EF037BFC6CFEC4" box="[899,935,327,349]" count="1" pageId="2" pageNumber="463" type="female">1 ♀</specimenCount>
, ditto,
<date id="FFD45CA0FF89FFAC9481037BFBAAFEC5" box="[1005,1121,327,348]" pageId="2" pageNumber="463" value="2014-03-28">
<collectingDate id="EF90A548FF89FFAC9481037BFBAAFEC5" box="[1005,1121,327,348]" pageId="2" pageNumber="463" value="2014-03-28">28.III.2014</collectingDate>
</date>
,
<collectorName id="269F1FB6FF89FFAC93010374FB63FEC4" box="[1133,1192,328,349]" pageId="2" pageNumber="463">N. Ito</collectorName>
leg., by tree sap (
<taxonomicName id="4C6A01E3FF89FFAC9403035FFBD1FEEE" box="[879,1050,354,376]" class="Magnoliopsida" family="Oleaceae" genus="Fraxinus" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="463" phylum="Tracheophyta" rank="species" species="griffithii">
<emphasis id="B91EA672FF89FFAC9403035FFBD1FEEE" box="[879,1050,354,376]" italics="true" pageId="2" pageNumber="463">Fraxinus griffithii</emphasis>
</taxonomicName>
) (
<collectionCode id="ED7BE2A5FF89FFAC9341035EFBBAFEEE" box="[1069,1137,354,375]" pageId="2" pageNumber="463">BLKU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B02703DFF89FFAC93EC035EFCDDFE35" collectingDate="2021-10-30" collectingDateMax="2014-03-28" collectingDateMin="2014-03-24" collectionCode="BLKU" collectorName="N. Ito" country="Japan" pageId="2" pageNumber="463" specimenCount="5" specimenCount-male="5" stateProvince="Okinawa" typeStatus="paratype">
<specimenCount id="9D6CB1E9FF89FFAC93EC035EFB6EFEEE" box="[1152,1189,354,376]" count="5" pageId="2" pageNumber="463" type="male">5 ♂</specimenCount>
, ditto,
<date id="FFD45CA0FF89FFAC95950341FC96FE0B" box="[761,861,381,402]" pageId="2" pageNumber="463" value="2021-10-30">
<collectingDate id="EF90A548FF89FFAC95950341FC96FE0B" box="[761,861,381,402]" pageId="2" pageNumber="463" value="2021-10-30">30.X.2021</collectingDate>
</date>
,
<collectorName id="269F1FB6FF89FFAC94040341FC6BFE0B" box="[872,928,381,402]" pageId="2" pageNumber="463">N. Ito</collectorName>
leg.,
<collectingMethod id="522B0277FF89FFAC94B90341FB86FE0B" box="[981,1101,381,403]" pageId="2" pageNumber="463">by fruit trap</collectingMethod>
(banana) (
<collectionCode id="ED7BE2A5FF89FFAC95A503ABFCC5FE35" box="[713,782,407,428]" pageId="2" pageNumber="463">BLKU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B02703DFF89FFAC947203ABFCC7FE7B" collectingDate="2022-06-23" collectingDateMax="2022-06-27" collectingDateMin="2022-06-23" collectionCode="BYU, BLKU" collectorName="N. Ito" country="Japan" pageId="2" pageNumber="463" specimenCount="10" specimenCount-female="4" specimenCount-male="6" stateProvince="Okinawa" typeStatus="paratype">
<specimenCount id="9D6CB1E9FF89FFAC947203ABFC8AFE34" box="[798,833,407,429]" count="3" pageId="2" pageNumber="463" type="male">3 ♂</specimenCount>
,
<specimenCount id="9D6CB1E9FF89FFAC942703ABFCA6FE34" box="[843,877,407,429]" count="2" pageId="2" pageNumber="463" type="female">2 ♀</specimenCount>
, ditto,
<date id="FFD45CA0FF89FFAC94C103ABFBF6FE35" box="[941,1085,407,428]" pageId="2" pageNumber="463" value="2022-06-23" valueMax="2022-06-27" valueMin="2022-06-23">
<collectingDate id="EF90A548FF89FFAC94C103ABFBF6FE35" box="[941,1085,407,428]" pageId="2" pageNumber="463" value="2022-06-23" valueMax="2022-06-27" valueMin="2022-06-23">2327.VI.2022</collectingDate>
</date>
,
<collectorName id="269F1FB6FF89FFAC932903A4FBB0FE34" box="[1093,1147,408,429]" pageId="2" pageNumber="463">N. Ito</collectorName>
leg.,
<collectingMethod id="522B0277FF89FFAC95AF038EFCF1FE5E" box="[707,826,434,456]" pageId="2" pageNumber="463">by fruit trap</collectingMethod>
(banana) (
<specimenCount id="9D6CB1E9FF89FFAC94C4038EFC05FE5E" box="[936,974,434,455]" count="1" pageId="2" pageNumber="463" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D6CB1E9FF89FFAC94B6038EFBCBFE5E" box="[986,1024,434,455]" count="1" pageId="2" pageNumber="463" type="female">1 ♀</specimenCount>
:
<collectionCode id="ED7BE2A5FF89FFAC9360038EFBF4FE5E" box="[1036,1087,434,455]" country="USA" httpUri="http://grbio.org/cool/0am8-rdhc" name="Monte L. Bean Life Science Museum" pageId="2" pageNumber="463" type="University or college">BYU</collectionCode>
;
<specimenCount id="9D6CB1E9FF89FFAC9327038EFBBAFE5E" box="[1099,1137,434,455]" count="2" pageId="2" pageNumber="463" type="male">2 ♂</specimenCount>
,
<specimenCount id="9D6CB1E9FF89FFAC9311038EFB68FE5E" box="[1149,1187,434,455]" count="1" pageId="2" pageNumber="463" type="female">1 ♀</specimenCount>
:
<collectionCode id="ED7BE2A5FF89FFAC95AF03F1FCCDFE7B" box="[707,774,461,482]" pageId="2" pageNumber="463">BLKU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B02703DFF89FFAC947903F1FC2EFD8E" collectingDate="2022-06-27" collectingDateMax="2022-06-29" collectingDateMin="2022-06-27" collectionCode="FSCA, BLKU, KUM" collectorName="Hirakubo &amp; N. Ito" country="Japan" pageId="2" pageNumber="463" specimenCount="12" specimenCount-female="6" specimenCount-male="6" stateProvince="Okinawa" typeStatus="paratype">
<specimenCount id="9D6CB1E9FF89FFAC947903F1FCF2FE7B" box="[789,825,461,482]" count="3" pageId="2" pageNumber="463" type="male">3 ♂</specimenCount>
,
<specimenCount id="9D6CB1E9FF89FFAC942F03F1FCADFE7B" box="[835,870,461,482]" count="3" pageId="2" pageNumber="463" type="female">3 ♀</specimenCount>
,
<collectorName id="269F1FB6FF89FFAC941D03F1FC01FE7A" box="[881,970,461,483]" pageId="2" pageNumber="463">Hirakubo</collectorName>
,
<date id="FFD45CA0FF89FFAC94BE03F1FBADFE7B" box="[978,1126,461,482]" pageId="2" pageNumber="463" value="2022-06-27" valueMax="2022-06-29" valueMin="2022-06-27">
<collectingDate id="EF90A548FF89FFAC94BE03F1FBADFE7B" box="[978,1126,461,482]" pageId="2" pageNumber="463" value="2022-06-27" valueMax="2022-06-29" valueMin="2022-06-27">2729.VI.2022</collectingDate>
</date>
,
<collectorName id="269F1FB6FF89FFAC931C03F1FB63FE7B" box="[1136,1192,461,482]" pageId="2" pageNumber="463">N. Ito</collectorName>
leg.,
<collectingMethod id="522B0277FF89FFAC959C03DBFC95FE64" box="[752,862,487,509]" pageId="2" pageNumber="463">by fruit trap</collectingMethod>
(pineapple) (
<specimenCount id="9D6CB1E9FF89FFAC94B703DBFC35FE64" box="[987,1022,487,509]" count="1" pageId="2" pageNumber="463" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D6CB1E9FF89FFAC936403DBFBE6FE64" box="[1032,1069,487,509]" count="1" pageId="2" pageNumber="463" type="female">1 ♀</specimenCount>
:
<collectionCode id="ED7BE2A5FF89FFAC935403DBFBBEFE65" box="[1080,1141,487,508]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="2" pageNumber="463" type="Museum">FSCA</collectionCode>
;
<specimenCount id="9D6CB1E9FF89FFAC931203DBFB6FFE64" box="[1150,1188,487,509]" count="1" pageId="2" pageNumber="463" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D6CB1E9FF89FFAC95AF003EFD22FD8E" box="[707,745,514,535]" count="1" pageId="2" pageNumber="463" type="female">1 ♀</specimenCount>
:
<collectionCode id="ED7BE2A5FF89FFAC959F003EFCFCFD8E" box="[755,823,514,535]" pageId="2" pageNumber="463">BLKU</collectionCode>
;
<specimenCount id="9D6CB1E9FF89FFAC942D003EFCACFD8E" box="[833,871,514,535]" count="1" pageId="2" pageNumber="463" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D6CB1E9FF89FFAC941D003EFC5CFD8E" box="[881,919,514,535]" count="1" pageId="2" pageNumber="463" type="female">1 ♀</specimenCount>
:
<collectionCode id="ED7BE2A5FF89FFAC94CE003EFC15FD8E" box="[930,990,514,535]" pageId="2" pageNumber="463">KUM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37029EBFF89FFAC95B70021FB8CFD04" pageId="2" pageNumber="463" type="etymology">
<paragraph id="8BD57A60FF89FFAC95B70021FB8CFD04" blockId="2.[706,1193,194,1709]" pageId="2" pageNumber="463">
<emphasis id="B91EA672FF89FFAC95B70021FC85FDAB" bold="true" box="[731,846,541,562]" pageId="2" pageNumber="463">Etymology.</emphasis>
The specific epithet of the new species is formed from the Latin “
<emphasis id="B91EA672FF89FFAC936D000BFBF7FDD5" box="[1025,1084,567,588]" italics="true" pageId="2" pageNumber="463">cordis</emphasis>
” (heart) + “
<emphasis id="B91EA672FF89FFAC95A1006EFCC1FDFE" box="[717,778,594,615]" italics="true" pageId="2" pageNumber="463">formis</emphasis>
” (form), which is derived from the arrangement of minute projections on the male abdominal sternite VII forming a weak heart shape.
</paragraph>
</subSubSection>
<subSubSection id="C37029EBFF89FFAC95B7009EFBACFCA4" pageId="2" pageNumber="463" type="diagnosis">
<paragraph id="8BD57A60FF89FFAC95B7009EFBACFCA4" blockId="2.[706,1193,194,1709]" pageId="2" pageNumber="463">
<emphasis id="B91EA672FF89FFAC95B7009EFC88FD21" bold="true" box="[731,835,674,696]" pageId="2" pageNumber="463">Diagnosis.</emphasis>
This species differs from its conge- ners by the following characteristics: abdominal sternite VII of male with minute projections in a heart-shaped arrangement; abdominal sternite VII of female pentagonal; lateral lobe of tegmen not narrowed from distal half towards the apex.
</paragraph>
</subSubSection>
<subSubSection id="C37029EBFF89FFA995B7017EFC64F9F8" lastPageId="7" lastPageNumber="468" pageId="2" pageNumber="463" type="description">
<paragraph id="8BD57A60FF89FFAC95B7017EFC68FB8B" blockId="2.[706,1193,194,1709]" pageId="2" pageNumber="463">
<emphasis id="B91EA672FF89FFAC95B7017EFC9CFCC1" bold="true" box="[731,855,834,856]" pageId="2" pageNumber="463">Description.</emphasis>
Male. Measurements and proportions (
<emphasis id="B91EA672FF89FFAC95930161FCC0FCEB" box="[767,779,861,882]" italics="true" pageId="2" pageNumber="463">n</emphasis>
= 13): BL
<quantity id="4C92D785FF89FFAC941F0160FC35FCEB" box="[883,1022,860,882]" metricMagnitude="-3" metricUnit="m" metricValue="2.475" metricValueMax="2.7" metricValueMin="2.25" pageId="2" pageNumber="463" unit="mm" value="2.475" valueMax="2.7" valueMin="2.25">2.252.70 mm</quantity>
(mean
<quantity id="4C92D785FF89FFAC93290161FB55FCEB" box="[1093,1182,861,882]" metricMagnitude="-3" metricUnit="m" metricValue="2.43" pageId="2" pageNumber="463" unit="mm" value="2.43">2.43 mm</quantity>
), BW
<quantity id="4C92D785FF89FFAC9587014BFCA7FC14" box="[747,876,887,909]" metricMagnitude="-3" metricUnit="m" metricValue="1.625" metricValueMax="1.8" metricValueMin="1.45" pageId="2" pageNumber="463" unit="mm" value="1.625" valueMax="1.8" valueMin="1.45">1.451.80 mm</quantity>
(mean
<quantity id="4C92D785FF89FFAC94C7014BFC36FC15" box="[939,1021,887,909]" metricMagnitude="-3" metricUnit="m" metricValue="1.58" pageId="2" pageNumber="463" unit="mm" value="1.58">1.58 mm</quantity>
), PL
<quantity id="4C92D785FF89FFAC934B014BFB62FC14" box="[1063,1193,887,909]" metricMagnitude="-3" metricUnit="m" metricValue="1.005" metricValueMax="1.09" metricValueMin="0.92" pageId="2" pageNumber="463" unit="mm" value="1.005" valueMax="1.09" valueMin="0.92">0.921.09 mm</quantity>
(mean 1.00 mm), PW
<quantity id="4C92D785FF89FFAC94AD01AEFB91FC3E" box="[961,1114,914,936]" metricMagnitude="-3" metricUnit="m" metricValue="1.525" metricValueMax="1.67" metricValueMin="1.38" pageId="2" pageNumber="463" unit="mm" value="1.525" valueMax="1.67" valueMin="1.38">1.381.67 mm</quantity>
(mean
<quantity id="4C92D785FF89FFAC95AF0190FCD5FC5B" box="[707,798,940,962]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="463" unit="mm" value="1.5">1.50 mm</quantity>
), EL
<quantity id="4C92D785FF89FFAC94230190FC13FC5B" box="[847,984,940,962]" metricMagnitude="-3" metricUnit="m" metricValue="1.465" metricValueMax="1.6" metricValueMin="1.33" pageId="2" pageNumber="463" unit="mm" value="1.465" valueMax="1.6" valueMin="1.33">1.331.60 mm</quantity>
(mean
<quantity id="4C92D785FF89FFAC93710191FBBFFC5B" box="[1053,1140,941,962]" metricMagnitude="-3" metricUnit="m" metricValue="1.43" pageId="2" pageNumber="463" unit="mm" value="1.43">1.43 mm</quantity>
); EW
<quantity id="4C92D785FF89FFAC95AF01FBFC9FFC44" box="[707,852,967,989]" metricMagnitude="-3" metricUnit="m" metricValue="1.625" metricValueMax="1.8" metricValueMin="1.45" pageId="2" pageNumber="463" unit="mm" value="1.625" valueMax="1.8" valueMin="1.45">1.451.80 mm</quantity>
(mean 1.58); BW/BL 0.640.67 (mean 0.65), PW/PL 1.461.53 (mean 1.50), EW/EL 1.081.13 (mean 1.10).
</paragraph>
<paragraph id="8BD57A60FF89FFAC95B7062BFB41FB2A" blockId="2.[706,1193,194,1709]" pageId="2" pageNumber="463">
Body (
<figureCitation id="135166E5FF89FFAC9477062BFC99FBB5" box="[795,850,1047,1069]" captionStart="Fig" captionStartId="3.[176,209,1668,1687]" captionTargetBox="[159,1145,213,1634]" captionTargetId="figure-8@3.[151,1171,194,1651]" captionTargetPageId="3" captionText="Fig. 1. Urophorus (Anophorus) cordiformis, new species. AB) Male, dorsal and ventral view, respectively, CD) Female, dorsal and ventral view, respectively." figureDoi="http://doi.org/10.5281/zenodo.13243313" httpUri="https://zenodo.org/record/13243313/files/figure.png" pageId="2" pageNumber="463">Fig. 1</figureCitation>
) oblong; moderately convex dorso- ventrally; dorsum and venter shining; dorsum and venter feebly covered with recumbent, short, yel- lowish pubescence. Body coloration dark brown; antennae, mandibles, lateral margin of pronotum, humeral area of elytra, and legs reddish-brown.
</paragraph>
<paragraph id="8BD57A60FF89FFAC95B7068BFC1EF9DA" blockId="2.[706,1193,194,1709]" pageId="2" pageNumber="463">
Head (
<figureCitation id="135166E5FF89FFAC9476068BFCA9FB55" box="[794,866,1207,1229]" captionStart="Fig" captionStartId="4.[197,228,1392,1411]" captionTargetBox="[179,1180,197,1366]" captionTargetId="figure-83@4.[173,1193,193,1375]" captionTargetPageId="4" captionText="Fig. 2. Urophorus (Anophorus) cordiformis, new species. A) Labrum, dorsal view, B) Labial palpi with mentum, ventral view, C) Right maxilla, ventral view, D) Right mandible, dorsal view, E) Head capsule, dorsal view, F) Right antenna, dorsal view, G) Abdominal sternite VI of male, ventral view, H) Abdominal sternite VII of male, ventral view, I) Spiculum gastrale, ventral view, J) Anal sclerite, ventral view, K) Abdominal sternite VI of female, ventral view, L) Abdominal sternite VII of female, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13243315" httpUri="https://zenodo.org/record/13243315/files/figure.png" pageId="2" pageNumber="463">Fig. 2E</figureCitation>
) punctate; punctures on disc large and dense, on occipital region larger and deeper than that of the disc. Labrum (
<figureCitation id="135166E5FF89FFAC94D706D1FBC3FA9B" box="[955,1032,1261,1283]" captionStart="Fig" captionStartId="4.[197,228,1392,1411]" captionTargetBox="[179,1180,197,1366]" captionTargetId="figure-83@4.[173,1193,193,1375]" captionTargetPageId="4" captionText="Fig. 2. Urophorus (Anophorus) cordiformis, new species. A) Labrum, dorsal view, B) Labial palpi with mentum, ventral view, C) Right maxilla, ventral view, D) Right mandible, dorsal view, E) Head capsule, dorsal view, F) Right antenna, dorsal view, G) Abdominal sternite VI of male, ventral view, H) Abdominal sternite VII of male, ventral view, I) Spiculum gastrale, ventral view, J) Anal sclerite, ventral view, K) Abdominal sternite VI of female, ventral view, L) Abdominal sternite VII of female, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13243315" httpUri="https://zenodo.org/record/13243315/files/figure.png" pageId="2" pageNumber="463">Fig. 2A</figureCitation>
) deeply notched in middle. Antenna (
<figureCitation id="135166E5FF89FFAC94EB073BFC05FA85" box="[903,974,1287,1309]" captionStart="Fig" captionStartId="4.[197,228,1392,1411]" captionTargetBox="[179,1180,197,1366]" captionTargetId="figure-83@4.[173,1193,193,1375]" captionTargetPageId="4" captionText="Fig. 2. Urophorus (Anophorus) cordiformis, new species. A) Labrum, dorsal view, B) Labial palpi with mentum, ventral view, C) Right maxilla, ventral view, D) Right mandible, dorsal view, E) Head capsule, dorsal view, F) Right antenna, dorsal view, G) Abdominal sternite VI of male, ventral view, H) Abdominal sternite VII of male, ventral view, I) Spiculum gastrale, ventral view, J) Anal sclerite, ventral view, K) Abdominal sternite VI of female, ventral view, L) Abdominal sternite VII of female, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13243315" httpUri="https://zenodo.org/record/13243315/files/figure.png" pageId="2" pageNumber="463">Fig. 2F</figureCitation>
) with three antennom- eres in club oblong; antennal club approximately 1.2 times as long as wide; antennomere X widest, antennomere IX distinctly narrower than X; antennomere XI obtusely pointed apically. Mandible (
<figureCitation id="135166E5FF89FFAC95A607B1FCD2FA3B" box="[714,793,1421,1443]" captionStart="Fig" captionStartId="4.[197,228,1392,1411]" captionTargetBox="[179,1180,197,1366]" captionTargetId="figure-83@4.[173,1193,193,1375]" captionTargetPageId="4" captionText="Fig. 2. Urophorus (Anophorus) cordiformis, new species. A) Labrum, dorsal view, B) Labial palpi with mentum, ventral view, C) Right maxilla, ventral view, D) Right mandible, dorsal view, E) Head capsule, dorsal view, F) Right antenna, dorsal view, G) Abdominal sternite VI of male, ventral view, H) Abdominal sternite VII of male, ventral view, I) Spiculum gastrale, ventral view, J) Anal sclerite, ventral view, K) Abdominal sternite VI of female, ventral view, L) Abdominal sternite VII of female, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13243315" httpUri="https://zenodo.org/record/13243315/files/figure.png" pageId="2" pageNumber="463">Fig. 2D</figureCitation>
) with an inner tooth. Labial palpi with mentum as figured (
<figureCitation id="135166E5FF89FFAC94EB079BFC18FA25" box="[903,979,1447,1469]" captionStart="Fig" captionStartId="4.[197,228,1392,1411]" captionTargetBox="[179,1180,197,1366]" captionTargetId="figure-83@4.[173,1193,193,1375]" captionTargetPageId="4" captionText="Fig. 2. Urophorus (Anophorus) cordiformis, new species. A) Labrum, dorsal view, B) Labial palpi with mentum, ventral view, C) Right maxilla, ventral view, D) Right mandible, dorsal view, E) Head capsule, dorsal view, F) Right antenna, dorsal view, G) Abdominal sternite VI of male, ventral view, H) Abdominal sternite VII of male, ventral view, I) Spiculum gastrale, ventral view, J) Anal sclerite, ventral view, K) Abdominal sternite VI of female, ventral view, L) Abdominal sternite VII of female, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13243315" httpUri="https://zenodo.org/record/13243315/files/figure.png" pageId="2" pageNumber="463">Fig. 2B</figureCitation>
); terminal segment of labial palpomere widened to truncate apex, and longer than width; mentum with large punctures, and widest near mid-length. Maxilla as shown in
<figureCitation id="135166E5FF89FFAC95AF042EFCDBF9BE" box="[707,784,1554,1576]" captionStart="Fig" captionStartId="4.[197,228,1392,1411]" captionTargetBox="[179,1180,197,1366]" captionTargetId="figure-83@4.[173,1193,193,1375]" captionTargetPageId="4" captionText="Fig. 2. Urophorus (Anophorus) cordiformis, new species. A) Labrum, dorsal view, B) Labial palpi with mentum, ventral view, C) Right maxilla, ventral view, D) Right mandible, dorsal view, E) Head capsule, dorsal view, F) Right antenna, dorsal view, G) Abdominal sternite VI of male, ventral view, H) Abdominal sternite VII of male, ventral view, I) Spiculum gastrale, ventral view, J) Anal sclerite, ventral view, K) Abdominal sternite VI of female, ventral view, L) Abdominal sternite VII of female, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13243315" httpUri="https://zenodo.org/record/13243315/files/figure.png" pageId="2" pageNumber="463">Fig. 2C</figureCitation>
; last maxillary palpomere elongated, narrowed near basal two-thirds.
</paragraph>
<paragraph id="8BD57A60FF89FFAB95B70474FCB5F913" blockId="2.[706,1193,194,1709]" lastBlockId="5.[152,1171,1598,1727]" lastPageId="5" lastPageNumber="466" pageId="2" pageNumber="463">Pronotum convex, transverse, approximately 1.5 times as wide as long; disc moderately punctate, punctures separated by about their diameter, progres- sively larger towards the lateral sides; interspace 464 between punctures smooth; lateral margins widest near basal one-fourth, strongly narrowed anteriorly and weakly narrowed posteriorly; anterior corner subrounded, and posterior corner right angle; anterior margin slightly emarginated, posterior margin sinuate; dorsolateral fovea on pronotum indistinct. Prosternum with large, dense punctures separated by less than their diameter, interspaces smooth; prosternal process strongly expanded behind procoxae, disc without punctures, glabrous. Mesoventrite with punctures that larger than on prosternum, interspaces between punctures with microreticulation. 466 Metaventrite densely punctate, disc flattened and large as on pronotal disc; interspaces between punc- sparsely punctate. tures smooth; apical margins broadly rounded, hind Elytra 1.081.13 times wider than long; lateral sutural angles obtuse.</paragraph>
<caption id="DF152AE8FF88FFAD97DC04B8FD9DF936" ID-DOI="http://doi.org/10.5281/zenodo.13243313" ID-Zenodo-Dep="13243313" httpUri="https://zenodo.org/record/13243313/files/figure.png" pageId="3" pageNumber="464" startId="3.[176,209,1668,1687]" targetBox="[159,1145,213,1634]" targetPageId="3" targetType="figure">
<paragraph id="8BD57A60FF88FFAD97DC04B8FB58F90E" blockId="3.[152,1171,1668,1711]" box="[176,1171,1668,1687]" pageId="3" pageNumber="464">
<emphasis id="B91EA672FF88FFAD97DC04B8FF27F90E" bold="true" box="[176,236,1668,1687]" pageId="3" pageNumber="464">Fig. 1.</emphasis>
<taxonomicName id="4C6A01E3FF88FFAD966E04B8FD81F90E" authorityName="Ito &amp; Powell" authorityYear="2023" box="[258,586,1668,1687]" class="Insecta" family="Nitidulidae" genus="Urophorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="464" phylum="Arthropoda" rank="species" species="cordiformis" status="sp. nov." subGenus="Anophorus">
<emphasis id="B91EA672FF88FFAD966E04B8FEA8F90E" box="[258,355,1668,1687]" italics="true" pageId="3" pageNumber="464">Urophorus</emphasis>
(
<emphasis id="B91EA672FF88FFAD961E04B8FE1FF90E" box="[370,468,1668,1687]" italics="true" pageId="3" pageNumber="464">Anophorus</emphasis>
)
<emphasis id="B91EA672FF88FFAD968E04B8FD81F90E" box="[482,586,1668,1687]" italics="true" pageId="3" pageNumber="464">cordiformis</emphasis>
</taxonomicName>
,
<emphasis id="B91EA672FF88FFAD953A04B8FD0EF90E" bold="true" box="[598,709,1668,1687]" pageId="3" pageNumber="464">
<taxonomicNameLabel id="A22D1B09FF88FFAD953A04B8FD0EF90E" box="[598,709,1668,1687]" pageId="3" pageNumber="464" rank="species">new species</taxonomicNameLabel>
</emphasis>
. AB) Male, dorsal and ventral view, respectively,
</paragraph>
<paragraph id="8BD57A60FF88FFAD97F404A0FD9DF936" blockId="3.[152,1171,1668,1711]" box="[152,598,1692,1711]" pageId="3" pageNumber="464">CD) Female, dorsal and ventral view, respectively.</paragraph>
</caption>
<caption id="DF152AE8FF8FFFAA97A9074CFD96FA7A" ID-DOI="http://doi.org/10.5281/zenodo.13243315" ID-Zenodo-Dep="13243315" httpUri="https://zenodo.org/record/13243315/files/figure.png" pageId="4" pageNumber="465" startId="4.[197,228,1392,1411]" targetBox="[179,1180,197,1366]" targetPageId="4" targetType="figure">
<paragraph id="8BD57A60FF8FFFAA97A9074CFD96FA7A" blockId="4.[173,1193,1392,1508]" pageId="4" pageNumber="465">
<emphasis id="B91EA672FF8FFFAA97A9074CFF35FA1A" bold="true" box="[197,254,1392,1411]" pageId="4" pageNumber="465">Fig. 2.</emphasis>
<taxonomicName id="4C6A01E3FF8FFFAA9678074CFD87FA1A" authorityName="Ito &amp; Powell" authorityYear="2023" box="[276,588,1392,1411]" class="Insecta" family="Nitidulidae" genus="Urophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="465" phylum="Arthropoda" rank="species" species="cordiformis" status="sp. nov." subGenus="Anophorus">
<emphasis id="B91EA672FF8FFFAA9678074CFEBAFA1A" box="[276,369,1392,1411]" italics="true" pageId="4" pageNumber="465">Urophorus</emphasis>
(
<emphasis id="B91EA672FF8FFFAA9612074CFE17FA1A" box="[382,476,1392,1411]" italics="true" pageId="4" pageNumber="465">Anophorus</emphasis>
)
<emphasis id="B91EA672FF8FFFAA9685074CFD87FA1A" box="[489,588,1392,1411]" italics="true" pageId="4" pageNumber="465">cordiformis</emphasis>
</taxonomicName>
,
<emphasis id="B91EA672FF8FFFAA9534074CFD09FA1A" bold="true" box="[600,706,1392,1411]" pageId="4" pageNumber="465">
<taxonomicNameLabel id="A22D1B09FF8FFFAA9534074CFD09FA1A" box="[600,706,1392,1411]" pageId="4" pageNumber="465" rank="species">new species</taxonomicNameLabel>
</emphasis>
. A) Labrum, dorsal view, B) Labial palpi with mentum, ventral view, C) Right maxilla, ventral view, D) Right mandible, dorsal view, E) Head capsule, dorsal view, F) Right antenna, dorsal view, G) Abdominal sternite VI of male, ventral view, H) Abdominal sternite VII of male, ventral view, I) Spiculum gastrale, ventral view, J) Anal sclerite, ventral view, K) Abdominal sternite VI of female, ventral view, L) Abdominal sternite VII of female, ventral view.
</paragraph>
</caption>
<caption id="DF152AE8FF8EFFAB97DC00FBFDA4FD6B" ID-DOI="http://doi.org/10.5281/zenodo.13243317" ID-Zenodo-Dep="13243317" httpUri="https://zenodo.org/record/13243317/files/figure.png" pageId="5" pageNumber="466" startId="5.[176,207,711,730]" targetBox="[178,1164,202,691]" targetPageId="5" targetType="figure">
<paragraph id="8BD57A60FF8EFFAB97DC00FBFDA4FD6B" blockId="5.[152,1171,711,754]" pageId="5" pageNumber="466">
<emphasis id="B91EA672FF8EFFAB97DC00FBFF21FD43" bold="true" box="[176,234,711,730]" pageId="5" pageNumber="466">Fig. 3.</emphasis>
Genitalia of
<taxonomicName id="4C6A01E3FF8EFFAB960200FBFD63FD43" authorityName="Ito &amp; Powell" authorityYear="2023" box="[366,680,711,730]" class="Insecta" family="Nitidulidae" genus="Urophorus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="cordiformis" status="sp. nov." subGenus="Anophorus">
<emphasis id="B91EA672FF8EFFAB960200FBFE00FD43" box="[366,459,711,730]" italics="true" pageId="5" pageNumber="466">Urophorus</emphasis>
(
<emphasis id="B91EA672FF8EFFAB96B500FBFDFCFD43" box="[473,567,711,730]" italics="true" pageId="5" pageNumber="466">Anophorus</emphasis>
)
<emphasis id="B91EA672FF8EFFAB952900FBFD63FD43" box="[581,680,711,730]" italics="true" pageId="5" pageNumber="466">cordiformis</emphasis>
</taxonomicName>
,
<emphasis id="B91EA672FF8EFFAB95D800FBFCD4FD43" bold="true" box="[692,799,711,730]" pageId="5" pageNumber="466">
<taxonomicNameLabel id="A22D1B09FF8EFFAB95D800FBFCD4FD43" box="[692,799,711,730]" pageId="5" pageNumber="466" rank="species">new species</taxonomicNameLabel>
</emphasis>
. AC) Male genitalia, ventral, lateral, and dorsal views, respectively, D) Ovipositor, ventral view.
</paragraph>
</caption>
<caption id="DF152AE8FF8EFFAB97DC07C3FBA9F98B" ID-DOI="http://doi.org/10.5281/zenodo.13243321" ID-Zenodo-Dep="13243321" box="[176,1122,1535,1555]" httpUri="https://zenodo.org/record/13243321/files/figure.png" pageId="5" pageNumber="466" startId="5.[176,207,1535,1554]" targetBox="[151,1003,805,1518]" targetPageId="5" targetType="figure">
<paragraph id="8BD57A60FF8EFFAB97DC07C3FBA9F98B" blockId="5.[176,1122,1535,1555]" box="[176,1122,1535,1555]" pageId="5" pageNumber="466">
<emphasis id="B91EA672FF8EFFAB97DC07C3FF23F98B" bold="true" box="[176,232,1535,1554]" pageId="5" pageNumber="466">Fig. 4.</emphasis>
The mating posture of
<taxonomicName id="4C6A01E3FF8EFFAB96AC07C3FD3DF98B" authorityName="Ito &amp; Powell" authorityYear="2023" box="[448,758,1535,1554]" class="Insecta" family="Nitidulidae" genus="Urophorus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="466" phylum="Arthropoda" rank="species" species="cordiformis" status="sp. nov." subGenus="Anophorus">
<emphasis id="B91EA672FF8EFFAB96AC07C3FDD6F98B" box="[448,541,1535,1554]" italics="true" pageId="5" pageNumber="466">Urophorus</emphasis>
(
<emphasis id="B91EA672FF8EFFAB954507C3FD4CF98B" box="[553,647,1535,1554]" italics="true" pageId="5" pageNumber="466">Anophorus</emphasis>
)
<emphasis id="B91EA672FF8EFFAB95FF07C3FD3DF98B" box="[659,758,1535,1554]" italics="true" pageId="5" pageNumber="466">cordiformis</emphasis>
</taxonomicName>
,
<emphasis id="B91EA672FF8EFFAB946D07C3FCA1F98B" bold="true" box="[769,874,1535,1554]" pageId="5" pageNumber="466">
<taxonomicNameLabel id="A22D1B09FF8EFFAB946D07C3FCA1F98B" box="[769,874,1535,1554]" pageId="5" pageNumber="466" rank="species">new species</taxonomicNameLabel>
</emphasis>
under laboratory conditions.
</paragraph>
</caption>
<paragraph id="8BD57A60FF8EFFA897F404B2FE9DF9B9" blockId="5.[152,1171,1598,1727]" lastBlockId="6.[173,659,1307,1729]" lastPageId="6" lastPageNumber="467" pageId="5" pageNumber="466">
margins subparallel-sided, widest near mid-length, Abdomen with three terminal tergites exposed; wider than pronotal width; punctures on disc as punctures of dorsum and venter finer than those of elytra; setae of dorsum and venter shorter than those of head, pronotum, and elytra; posterior margin of abdominal sternite VI slightly emarginated, possessing a pair of small projections near the middle, with tuft of setae apically about as long as on elytra (
<figureCitation id="135166E5FF8DFFA897D8079CFEC9FA2F" box="[180,258,1440,1462]" captionStart="Fig" captionStartId="4.[197,228,1392,1411]" captionTargetBox="[179,1180,197,1366]" captionTargetId="figure-83@4.[173,1193,193,1375]" captionTargetPageId="4" captionText="Fig. 2. Urophorus (Anophorus) cordiformis, new species. A) Labrum, dorsal view, B) Labial palpi with mentum, ventral view, C) Right maxilla, ventral view, D) Right mandible, dorsal view, E) Head capsule, dorsal view, F) Right antenna, dorsal view, G) Abdominal sternite VI of male, ventral view, H) Abdominal sternite VII of male, ventral view, I) Spiculum gastrale, ventral view, J) Anal sclerite, ventral view, K) Abdominal sternite VI of female, ventral view, L) Abdominal sternite VII of female, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13243315" httpUri="https://zenodo.org/record/13243315/files/figure.png" pageId="6" pageNumber="467">Fig. 2G</figureCitation>
); abdominal sternite VII deeply depressed with densely arranged, minute projections; minute projections of abdominal sternite VII absent along the median line, arrangement somewhat heart shaped (
<figureCitation id="135166E5FF8DFFA897910437FE80F9B9" box="[253,331,1547,1569]" captionStart="Fig" captionStartId="4.[197,228,1392,1411]" captionTargetBox="[179,1180,197,1366]" captionTargetId="figure-83@4.[173,1193,193,1375]" captionTargetPageId="4" captionText="Fig. 2. Urophorus (Anophorus) cordiformis, new species. A) Labrum, dorsal view, B) Labial palpi with mentum, ventral view, C) Right maxilla, ventral view, D) Right mandible, dorsal view, E) Head capsule, dorsal view, F) Right antenna, dorsal view, G) Abdominal sternite VI of male, ventral view, H) Abdominal sternite VII of male, ventral view, I) Spiculum gastrale, ventral view, J) Anal sclerite, ventral view, K) Abdominal sternite VI of female, ventral view, L) Abdominal sternite VII of female, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13243315" httpUri="https://zenodo.org/record/13243315/files/figure.png" pageId="6" pageNumber="467">Fig. 2H</figureCitation>
).
</paragraph>
<caption id="DF152AE8FF8DFFA897A906F3FD44FB7B" ID-DOI="http://doi.org/10.5281/zenodo.13243323" ID-Zenodo-Dep="13243323" box="[197,655,1231,1250]" httpUri="https://zenodo.org/record/13243323/files/figure.png" pageId="6" pageNumber="467" startId="6.[197,228,1231,1250]" targetBox="[179,1164,205,1211]" targetPageId="6" targetType="figure">
<paragraph id="8BD57A60FF8DFFA897A906F3FD44FB7B" blockId="6.[197,655,1231,1250]" box="[197,655,1231,1250]" pageId="6" pageNumber="467">
<emphasis id="B91EA672FF8DFFA897A906F3FF36FB7B" bold="true" box="[197,253,1231,1250]" pageId="6" pageNumber="467">Fig. 5.</emphasis>
Geographic location of Ishigaki-jima Island.
</paragraph>
</caption>
<paragraph id="8BD57A60FF8DFFA897A9041AFB62FAD5" blockId="6.[173,659,1307,1729]" lastBlockId="6.[707,1193,1307,1729]" pageId="6" pageNumber="467">
Male genitalia (
<figureCitation id="135166E5FF8DFFA89637041AFE00F9A2" box="[347,459,1574,1596]" captionStart="Fig" captionStartId="5.[176,207,711,730]" captionTargetBox="[178,1164,202,691]" captionTargetId="figure-69@5.[151,1171,193,693]" captionTargetPageId="5" captionText="Fig. 3. Genitalia of Urophorus (Anophorus) cordiformis, new species. AC) Male genitalia, ventral, lateral, and dorsal views, respectively, D) Ovipositor, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13243317" httpUri="https://zenodo.org/record/13243317/files/figure.png" pageId="6" pageNumber="467">Figs. 3AC</figureCitation>
) as figured; each lateral lobe of tegmen subparallel-sided, apical margin truncated in dorsal view, narrowing to acute apical point in lateral view, apical region with dense, long setae; inner region of each lobe depressed in dorsal view; lateral lobes strongly curved, inner margin in distal half dentate in lateral view; tegminal strut gently widened distally, as long as tegminal length.
</paragraph>
<paragraph id="8BD57A60FF8DFFA895B7076CFCF6F9B9" blockId="6.[707,1193,1307,1729]" pageId="6" pageNumber="467">
Female. Measurements and proportions (
<emphasis id="B91EA672FF8DFFA8930B076DFBB8FAFF" box="[1127,1139,1361,1382]" italics="true" pageId="6" pageNumber="467">n</emphasis>
= 6): BL
<quantity id="4C92D785FF8DFFA895870757FCB0FA18" box="[747,891,1387,1409]" metricMagnitude="-3" metricUnit="m" metricValue="2.7249999999999996" metricValueMax="3.01" metricValueMin="2.44" pageId="6" pageNumber="467" unit="mm" value="2.7249999999999996" valueMax="3.01" valueMin="2.44">2.443.01 mm</quantity>
(mean
<quantity id="4C92D785FF8DFFA894A10757FBE0FA19" box="[973,1067,1387,1409]" metricMagnitude="-3" metricUnit="m" metricValue="2.68" pageId="6" pageNumber="467" unit="mm" value="2.68">2.68 mm</quantity>
); BW
<quantity id="4C92D785FF8DFFA8931E0757FCDDFA02" metricMagnitude="-3" metricUnit="m" metricValue="1.79" metricValueMax="1.94" metricValueMin="1.64" pageId="6" pageNumber="467" unit="mm" value="1.79" valueMax="1.94" valueMin="1.64">1.64 1.94 mm</quantity>
(mean
<quantity id="4C92D785FF8DFFA8943607BAFC7AFA02" box="[858,945,1414,1436]" metricMagnitude="-3" metricUnit="m" metricValue="1.76" pageId="6" pageNumber="467" unit="mm" value="1.76">1.76 mm</quantity>
); PL
<quantity id="4C92D785FF8DFFA8948C07BAFBA3FA02" box="[992,1128,1414,1435]" metricMagnitude="-3" metricUnit="m" metricValue="1.095" metricValueMax="1.2" metricValueMin="0.99" pageId="6" pageNumber="467" unit="mm" value="1.095" valueMax="1.2" valueMin="0.99">0.991.20 mm</quantity>
(mean
<quantity id="4C92D785FF8DFFA895AF079DFCD0FA2F" box="[707,795,1441,1462]" metricMagnitude="-3" metricUnit="m" metricValue="1.09" pageId="6" pageNumber="467" unit="mm" value="1.09">1.09 mm</quantity>
); PW
<quantity id="4C92D785FF8DFFA8943F079CFC16FA2F" box="[851,989,1440,1462]" metricMagnitude="-3" metricUnit="m" metricValue="1.675" metricValueMax="1.82" metricValueMin="1.53" pageId="6" pageNumber="467" unit="mm" value="1.675" valueMax="1.82" valueMin="1.53">1.531.82 mm</quantity>
(mean
<quantity id="4C92D785FF8DFFA8934F079CFBB0FA2F" box="[1059,1147,1440,1462]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" pageId="6" pageNumber="467" unit="mm" value="1.65">1.65 mm</quantity>
); EL
<quantity id="4C92D785FF8DFFA895AF0787FC80FA48" box="[707,843,1467,1489]" metricMagnitude="-3" metricUnit="m" metricValue="1.625" metricValueMax="1.8" metricValueMin="1.45" pageId="6" pageNumber="467" unit="mm" value="1.625" valueMax="1.8" valueMin="1.45">1.451.80 mm</quantity>
(mean
<quantity id="4C92D785FF8DFFA894FC0787FC2CFA49" box="[912,999,1467,1489]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="467" unit="mm" value="1.6">1.60 mm</quantity>
); EW
<quantity id="4C92D785FF8DFFA8934C0787FB63FA48" box="[1056,1192,1467,1489]" metricMagnitude="-3" metricUnit="m" metricValue="1.79" metricValueMax="1.94" metricValueMin="1.64" pageId="6" pageNumber="467" unit="mm" value="1.79" valueMax="1.94" valueMin="1.64">1.641.94 mm</quantity>
(mean
<quantity id="4C92D785FF8DFFA8946A07EAFCABFA72" box="[774,864,1494,1515]" metricMagnitude="-3" metricUnit="m" metricValue="1.74" pageId="6" pageNumber="467" unit="mm" value="1.74">1.74 mm</quantity>
); BW/BL 0.650.67 (mean 0.66); PW/PL 1.511.55 (mean 1.52); EW/EL 1.081.13 (mean 1.10).
</paragraph>
<paragraph id="8BD57A60FF8DFFA995B7041AFDB6FEC4" blockId="6.[707,1193,1307,1729]" lastBlockId="7.[152,1173,194,1634]" lastPageId="7" lastPageNumber="468" pageId="6" pageNumber="467">
External features similar to those of males except for the following. Abdominal sternite VI (
<figureCitation id="135166E5FF8DFFA8933C047CFB56F9CF" box="[1104,1181,1600,1622]" captionStart="Fig" captionStartId="4.[197,228,1392,1411]" captionTargetBox="[179,1180,197,1366]" captionTargetId="figure-83@4.[173,1193,193,1375]" captionTargetPageId="4" captionText="Fig. 2. Urophorus (Anophorus) cordiformis, new species. A) Labrum, dorsal view, B) Labial palpi with mentum, ventral view, C) Right maxilla, ventral view, D) Right mandible, dorsal view, E) Head capsule, dorsal view, F) Right antenna, dorsal view, G) Abdominal sternite VI of male, ventral view, H) Abdominal sternite VII of male, ventral view, I) Spiculum gastrale, ventral view, J) Anal sclerite, ventral view, K) Abdominal sternite VI of female, ventral view, L) Abdominal sternite VII of female, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13243315" httpUri="https://zenodo.org/record/13243315/files/figure.png" pageId="6" pageNumber="467">Fig. 2K</figureCitation>
); posterior margin straight and without a pair of small projections. Abdominal sternite VII (
<figureCitation id="135166E5FF8DFFA8934B044AFBBBF912" box="[1063,1136,1654,1676]" captionStart="Fig" captionStartId="4.[197,228,1392,1411]" captionTargetBox="[179,1180,197,1366]" captionTargetId="figure-83@4.[173,1193,193,1375]" captionTargetPageId="4" captionText="Fig. 2. Urophorus (Anophorus) cordiformis, new species. A) Labrum, dorsal view, B) Labial palpi with mentum, ventral view, C) Right maxilla, ventral view, D) Right mandible, dorsal view, E) Head capsule, dorsal view, F) Right antenna, dorsal view, G) Abdominal sternite VI of male, ventral view, H) Abdominal sternite VII of male, ventral view, I) Spiculum gastrale, ventral view, J) Anal sclerite, ventral view, K) Abdominal sternite VI of female, ventral view, L) Abdominal sternite VII of female, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13243315" httpUri="https://zenodo.org/record/13243315/files/figure.png" pageId="6" pageNumber="467">Fig. 2L</figureCitation>
) pentagonal. Pygidium triangular with apex rounded; punctures on subapical region smaller than on disc. 468 Ovipositor (
<figureCitation id="135166E5FF8CFFA9966702FEFE9EFF4E" box="[267,341,194,216]" captionStart="Fig" captionStartId="5.[176,207,711,730]" captionTargetBox="[178,1164,202,691]" captionTargetId="figure-69@5.[151,1171,193,693]" captionTargetPageId="5" captionText="Fig. 3. Genitalia of Urophorus (Anophorus) cordiformis, new species. AC) Male genitalia, ventral, lateral, and dorsal views, respectively, D) Ovipositor, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13243317" httpUri="https://zenodo.org/record/13243317/files/figure.png" pageId="7" pageNumber="468">Fig. 3D</figureCitation>
) with styli near apex; each sty- Distribution:
<collectingCountry id="F37D3AF0FF8CFFA9941D02FEFC61FF4E" box="[881,938,194,215]" name="Japan" pageId="7" pageNumber="468">Japan</collectingCountry>
(the Ryukyu Islands: lus with 3 short setae at apex. Okinawa-jima, Ishigaki-jima, Iriomote-jima,
<emphasis id="B91EA672FF8CFFA997DC02CBFEF8FE94" bold="true" box="[176,307,247,269]" pageId="7" pageNumber="468">Distribution.</emphasis>
<collectingCountry id="F37D3AF0FF8CFFA9965502CBFEB8FE95" box="[313,371,247,268]" name="Japan" pageId="7" pageNumber="468">Japan</collectingCountry>
: Ishigaki-jima Is. (
<figureCitation id="135166E5FF8CFFA9954B02CBFDA9FE94" box="[551,610,247,269]" captionStart="Fig" captionStartId="6.[197,228,1231,1250]" captionTargetBox="[179,1164,205,1211]" captionTargetId="figure-251@6.[173,1193,193,1213]" captionTargetPageId="6" captionText="Fig. 5. Geographic location of Ishigaki-jima Island." figureDoi="http://doi.org/10.5281/zenodo.13243323" httpUri="https://zenodo.org/record/13243323/files/figure.png" pageId="7" pageNumber="468">Fig. 5</figureCitation>
). and Minamidaito-jima Is.),
<collectingCountry id="F37D3AF0FF8CFFA994B002CBFBDEFE94" box="[988,1045,247,269]" name="Nepal" pageId="7" pageNumber="468">Nepal</collectingCountry>
,
<collectingCountry id="F37D3AF0FF8CFFA9937102CBFB84FE94" box="[1053,1103,247,269]" name="India" pageId="7" pageNumber="468">India</collectingCountry>
,
<collectingCountry id="F37D3AF0FF8CFFA9933A02CBFB5BFE94" box="[1110,1168,247,269]" name="China" pageId="7" pageNumber="468">China</collectingCountry>
,
<emphasis id="B91EA672FF8CFFA997DC032EFED9FEBE" bold="true" box="[176,274,274,295]" pageId="7" pageNumber="468">Remarks.</emphasis>
<taxonomicName id="4C6A01E3FF8CFFA99674032EFEB4FEBE" authorityName="Murray" authorityYear="1864" box="[280,383,274,295]" class="Insecta" family="Nitidulidae" genus="Urophorus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="468" phylum="Arthropoda" rank="genus">
<emphasis id="B91EA672FF8CFFA99674032EFEB4FEBE" box="[280,383,274,295]" italics="true" pageId="7" pageNumber="468">Urophorus</emphasis>
</taxonomicName>
<taxonomicName id="4C6A01E3FF8CFFA996E9032EFDC6FEBE" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[389,525,274,295]" class="Insecta" family="Nitidulidae" genus="Urophorus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="468" phylum="Arthropoda" rank="species" species="humeralis" subGenus="Anophorus">
(
<emphasis id="B91EA672FF8CFFA996E1032EFE50FEBE" box="[397,411,274,295]" italics="true" pageId="7" pageNumber="468">A</emphasis>
.)
<emphasis id="B91EA672FF8CFFA996C3032EFDC6FEBE" box="[431,525,274,295]" italics="true" pageId="7" pageNumber="468">humeralis</emphasis>
</taxonomicName>
and
<emphasis id="B91EA672FF8CFFA99550032EFD86FEBE" box="[572,589,274,295]" italics="true" pageId="7" pageNumber="468">U</emphasis>
. (
<emphasis id="B91EA672FF8CFFA9950D032EFDA4FEBE" box="[609,623,274,295]" italics="true" pageId="7" pageNumber="468">A</emphasis>
.)
<collectingCountry id="F37D3AF0FF8CFFA9958E032EFCECFEB1" box="[738,807,274,296]" name="Taiwan" pageId="7" pageNumber="468">Taiwan</collectingCountry>
....................
<emphasis id="B91EA672FF8CFFA994CC032EFB58FEBE" bold="true" box="[928,1171,273,295]" pageId="7" pageNumber="468">
<emphasis id="B91EA672FF8CFFA994CC032EFC7AFEBE" bold="true" box="[928,945,274,295]" italics="true" pageId="7" pageNumber="468">U</emphasis>
.
<taxonomicName id="4C6A01E3FF8CFFA994D6032DFB58FEBE" authority="Murray" authorityName="Murray" box="[954,1171,273,295]" class="Insecta" family="Nitidulidae" genus="Urophorus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="468" phylum="Arthropoda" rank="species" species="foveicollis" subGenus="Anophorus">
(
<emphasis id="B91EA672FF8CFFA994AE032EFC1AFEBE" bold="true" box="[962,977,274,295]" italics="true" pageId="7" pageNumber="468">A</emphasis>
.)
<emphasis id="B91EA672FF8CFFA9948E032DFB8AFEBE" bold="true" box="[994,1089,273,295]" italics="true" pageId="7" pageNumber="468">foveicollis</emphasis>
Murray
</taxonomicName>
</emphasis>
<taxonomicName id="4C6A01E3FF8CFFA997F40310FF33FED8" box="[152,248,300,321]" class="Insecta" family="Nitidulidae" genus="Urophorus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="468" phylum="Arthropoda" rank="species" species="foveicollis">
<emphasis id="B91EA672FF8CFFA997F40310FF33FED8" box="[152,248,300,321]" italics="true" pageId="7" pageNumber="468">foveicollis</emphasis>
</taxonomicName>
are commonly found around the agricul- tural environments,
<emphasis id="B91EA672FF8CFFA996230374FEBBFEC4" box="[335,368,328,349]" italics="true" pageId="7" pageNumber="468">e.g.</emphasis>
, pineapple fields on Ishigaki-
</paragraph>
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<emphasis id="B91EA672FF8CFFA9945D036EFBCFFEF0" bold="true" box="[817,1028,338,362]" pageId="7" pageNumber="468">ACKNOWLEDGMENTS</emphasis>
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jima
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. In contrast, this new species has been collected only from fruit traps set in forests or tree This study was partly supported by the Collaborative sap of
<taxonomicName id="4C6A01E3FF8CFFA997B003A4FDC1FE34" authority="C. B. Clarke" authorityName="C. B. Clarke" box="[220,522,407,429]" class="Magnoliopsida" family="Oleaceae" genus="Fraxinus" kingdom="Plantae" order="Lamiales" pageId="7" pageNumber="468" phylum="Tracheophyta" rank="species" species="griffithii">
<emphasis id="B91EA672FF8CFFA997B003A4FE4CFE35" box="[220,391,407,429]" italics="true" pageId="7" pageNumber="468">Fraxinus griffithii</emphasis>
C. B. Clarke
</taxonomicName>
(
<taxonomicName id="4C6A01E3FF8CFFA9957403ABFDB8FE34" authorityName="von Hoffmannsegg &amp; Link" authorityYear="1809" box="[536,627,407,429]" class="Magnoliopsida" family="Oleaceae" kingdom="Plantae" order="Lamiales" pageId="7" pageNumber="468" phylum="Tracheophyta" rank="family">Oleaceae</taxonomicName>
), Research of Tropical Biosphere Research Center, suggesting that its preference of habitat is likely to University of the Ryukyus. GSP thanks the Florida differ from that of the other two species. Department of Agriculture and Consumer Services, The mating posture of this species was observed Division of Plant Industry, for their support. Lastly, under laboratory conditions. In the open, after gen- we thank the reviewers and editorial staff for ital contact, the male detaches all legs from the their recommendations that greatly improved the dorsum of the female, leaving only the abdomen manuscript. connected between the sexes (
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). Even in closed spaces, the sexes always maintained only the abdomen connection in an inverted position. The struc-
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<emphasis id="B91EA672FF8CFFA994AB0041FBC0FD0A" bold="true" box="[967,1035,637,661]" pageId="7" pageNumber="468">CITED</emphasis>
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ture of the male genitalia and the abdominal sternite
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<emphasis id="B91EA672FF8CFFA995C1009AFC9BFD20" bold="true" box="[685,848,678,697]" pageId="7" pageNumber="468">Audisio, P. 1993.</emphasis>
<taxonomicName id="4C6A01E3FF8CFFA99437009AFC72FD20" authorityName="Linnaeus" authorityYear="1758" box="[859,953,678,697]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="468" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
Nitidulidae-Kateretidae.
</subScript>
VII may be related to this unique mating posture. Fauna dItalia 32. Edizioni Calderini, Bologna,
</paragraph>
<paragraph id="8BD57A60FF8CFFA9958100EAFCE4FD73" blockId="7.[152,1173,194,1634]" box="[749,815,726,746]" pageId="7" pageNumber="468">971 pp.</paragraph>
<paragraph id="8BD57A60FF8CFFA995C100D2FB58FC80" blockId="7.[152,1173,194,1634]" pageId="7" pageNumber="468">
<emphasis id="B91EA672FF8CFFA995C100D2FB99FC98" bold="true" box="[685,1106,750,769]" pageId="7" pageNumber="468">Dasgupta, J., T. K. Pal, and G. S. Powell. 2021.</emphasis>
Taxon-
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</emphasis>
<taxonomicName id="4C6A01E3FF8CFFA9965E013CFDCBFC8F" box="[306,512,768,792]" class="Insecta" family="Nitidulidae" genus="Urophorus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="468" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="B91EA672FF8CFFA9965E013CFE61FC8F" bold="true" box="[306,426,768,791]" italics="true" pageId="7" pageNumber="468">UROPHORUS</emphasis>
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omy of
<taxonomicName id="4C6A01E3FF8CFFA9945B013AFC7BFC80" authorityName="Erichson" authorityYear="1843" box="[823,944,774,793]" class="Insecta" family="Nitidulidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="468" phylum="Arthropoda" rank="subFamily" subFamily="Carpophilinae">Carpophilinae</taxonomicName>
(
<taxonomicName id="4C6A01E3FF8CFFA994D2013AFBEBFC80" authorityName="Linnaeus" authorityYear="1758" box="[958,1056,774,793]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="468" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
:
<taxonomicName id="4C6A01E3FF8CFFA99340013AFB46FC80" authorityName="Latreille" authorityYear="1802" box="[1068,1165,774,793]" class="Insecta" family="Nitidulidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="468" phylum="Arthropoda" rank="family">Nitidulidae</taxonomicName>
)
</paragraph>
<paragraph id="8BD57A60FF8CFFA995810122FC1AFCD0" blockId="7.[152,1173,194,1634]" pageId="7" pageNumber="468">
from
<collectingRegion id="49AEB482FF8CFFA994710122FC94FCA8" box="[797,863,798,817]" country="India" name="Tripura" pageId="7" pageNumber="468">Tripura</collectingRegion>
,
<collectingCountry id="F37D3AF0FF8CFFA994050122FC5EFCA8" box="[873,917,798,817]" name="India" pageId="7" pageNumber="468">India</collectingCountry>
with a new species. Annales 1. Posterior corner of pronotum right angled Zoologici 71(3): 627649.
</paragraph>
<paragraph id="8BD57A60FF8CFFA997A1017BFC09FC30" blockId="7.[152,1173,194,1634]" pageId="7" pageNumber="468">
(
<figureCitation id="135166E5FF8CFFA997B8017BFEC1FCC5" box="[212,266,839,861]" captionStart="Fig" captionStartId="3.[176,209,1668,1687]" captionTargetBox="[159,1145,213,1634]" captionTargetId="figure-8@3.[151,1171,194,1651]" captionTargetPageId="3" captionText="Fig. 1. Urophorus (Anophorus) cordiformis, new species. AB) Male, dorsal and ventral view, respectively, CD) Female, dorsal and ventral view, respectively." figureDoi="http://doi.org/10.5281/zenodo.13243313" httpUri="https://zenodo.org/record/13243313/files/figure.png" pageId="7" pageNumber="468">Fig. 1</figureCitation>
). Antennomere IX distinctly narrower
<emphasis id="B91EA672FF8CFFA995C10172FC3CFCF8" bold="true" box="[685,1015,846,865]" pageId="7" pageNumber="468">Ewing, C. P., and A. R. Cline. 2005.</emphasis>
Key to adventive than antennomere X (
<figureCitation id="135166E5FF8CFFA996FD015EFE1DFCEE" box="[401,470,866,888]" captionStart="Fig" captionStartId="4.[197,228,1392,1411]" captionTargetBox="[179,1180,197,1366]" captionTargetId="figure-83@4.[173,1193,193,1375]" captionTargetPageId="4" captionText="Fig. 2. Urophorus (Anophorus) cordiformis, new species. A) Labrum, dorsal view, B) Labial palpi with mentum, ventral view, C) Right maxilla, ventral view, D) Right mandible, dorsal view, E) Head capsule, dorsal view, F) Right antenna, dorsal view, G) Abdominal sternite VI of male, ventral view, H) Abdominal sternite VII of male, ventral view, I) Spiculum gastrale, ventral view, J) Anal sclerite, ventral view, K) Abdominal sternite VI of female, ventral view, L) Abdominal sternite VII of female, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13243315" httpUri="https://zenodo.org/record/13243315/files/figure.png" pageId="7" pageNumber="468">Fig. 2F</figureCitation>
). Posterior margin sap beetles (
<taxonomicName id="4C6A01E3FF8CFFA99422015AFC60FCE0" authorityName="Linnaeus" authorityYear="1758" box="[846,939,870,889]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="468" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
:
<taxonomicName id="4C6A01E3FF8CFFA994DD015AFBC6FCE0" authorityName="Latreille" authorityYear="1802" box="[945,1037,870,889]" class="Insecta" family="Nitidulidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="468" phylum="Arthropoda" rank="family">Nitidulidae</taxonomicName>
) in
<collectingRegion id="49AEB482FF8CFFA99347015AFBA3FCE0" box="[1067,1128,870,889]" country="United States of America" name="Hawaii" pageId="7" pageNumber="468">Hawaii</collectingRegion>
, with of abdominal sternite VI slightly emarginate notes on records and habits. The Coleopterists and possessing a pair of small projections near Bulletin 59(2): 167183.
</paragraph>
<paragraph id="8BD57A60FF8CFFA997A1018EFB58FC58" blockId="7.[152,1173,194,1634]" box="[205,1171,942,968]" pageId="7" pageNumber="468">
middle (
<figureCitation id="135166E5FF8CFFA99674018EFEAAFC5E" box="[280,353,946,968]" captionStart="Fig" captionStartId="4.[197,228,1392,1411]" captionTargetBox="[179,1180,197,1366]" captionTargetId="figure-83@4.[173,1193,193,1375]" captionTargetPageId="4" captionText="Fig. 2. Urophorus (Anophorus) cordiformis, new species. A) Labrum, dorsal view, B) Labial palpi with mentum, ventral view, C) Right maxilla, ventral view, D) Right mandible, dorsal view, E) Head capsule, dorsal view, F) Right antenna, dorsal view, G) Abdominal sternite VI of male, ventral view, H) Abdominal sternite VII of male, ventral view, I) Spiculum gastrale, ventral view, J) Anal sclerite, ventral view, K) Abdominal sternite VI of female, ventral view, L) Abdominal sternite VII of female, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13243315" httpUri="https://zenodo.org/record/13243315/files/figure.png" pageId="7" pageNumber="468">Fig. 2G</figureCitation>
). Minute projections of abdom-
<emphasis id="B91EA672FF8CFFA995C10192FCAEFC58" bold="true" box="[685,869,942,961]" pageId="7" pageNumber="468">Hisamatsu, S. 1985.</emphasis>
Family
<taxonomicName id="4C6A01E3FF8CFFA994DD0192FBDAFC58" authorityName="Latreille" authorityYear="1802" box="[945,1041,942,961]" class="Insecta" family="Nitidulidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="468" phylum="Arthropoda" rank="family">Nitidulidae</taxonomicName>
[pp. 174197,
</paragraph>
<paragraph id="8BD57A60FF8CFFA997A101F0FB5FFC40" blockId="7.[152,1173,194,1634]" box="[205,1172,966,994]" pageId="7" pageNumber="468">
inal sternite VII of male absent along median pls. 2831].
<emphasis id="B91EA672FF8CFFA9943B01FBFCA2FC40" box="[855,873,967,985]" italics="true" pageId="7" pageNumber="468">In</emphasis>
: The
<taxonomicName id="4C6A01E3FF8CFFA994F501FAFC3CFC40" authorityName="Linnaeus" authorityYear="1758" box="[921,1015,966,985]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="468" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
of
<collectingCountry id="F37D3AF0FF8CFFA9937F01FBFB8FFC43" box="[1043,1092,967,986]" name="Japan" pageId="7" pageNumber="468">Japan</collectingCountry>
in Color,
</paragraph>
<paragraph id="8BD57A60FF8CFFA997A101DBFB5FFC68" blockId="7.[152,1173,194,1634]" box="[205,1172,990,1021]" pageId="7" pageNumber="468">line, forming a heart-shaped arrangement Vol. 3 (Y. Kurosawa, S. Hisamatsu, and H. Sasaji,</paragraph>
<paragraph id="8BD57A60FF8CFFA9958101CAFC19FB90" blockId="7.[152,1173,194,1634]" box="[749,978,1014,1033]" pageId="7" pageNumber="468">
editors). Hoikusha,
<collectingRegion id="49AEB482FF8CFFA994FA01CAFC05FB90" box="[918,974,1014,1033]" country="Japan" name="Osaka" pageId="7" pageNumber="468">Osaka</collectingRegion>
.
</paragraph>
<paragraph id="8BD57A60FF8CFFA997A1063EFB58FBB8" blockId="7.[152,1173,194,1634]" box="[205,1171,1026,1057]" pageId="7" pageNumber="468">
(
<figureCitation id="135166E5FF8CFFA997B8063EFEE8FB8E" box="[212,291,1026,1048]" captionStart="Fig" captionStartId="4.[197,228,1392,1411]" captionTargetBox="[179,1180,197,1366]" captionTargetId="figure-83@4.[173,1193,193,1375]" captionTargetPageId="4" captionText="Fig. 2. Urophorus (Anophorus) cordiformis, new species. A) Labrum, dorsal view, B) Labial palpi with mentum, ventral view, C) Right maxilla, ventral view, D) Right mandible, dorsal view, E) Head capsule, dorsal view, F) Right antenna, dorsal view, G) Abdominal sternite VI of male, ventral view, H) Abdominal sternite VII of male, ventral view, I) Spiculum gastrale, ventral view, J) Anal sclerite, ventral view, K) Abdominal sternite VI of female, ventral view, L) Abdominal sternite VII of female, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13243315" httpUri="https://zenodo.org/record/13243315/files/figure.png" pageId="7" pageNumber="468">Fig. 2H</figureCitation>
). Abdominal sternite VII of female
<emphasis id="B91EA672FF8CFFA995C10631FC2FFBB8" bold="true" box="[685,996,1037,1057]" pageId="7" pageNumber="468">Jelínek, J., and P. Audisio. 2007.</emphasis>
Family
<taxonomicName id="4C6A01E3FF8CFFA9935F0632FB58FBB8" authorityName="Latreille" authorityYear="1802" box="[1075,1171,1038,1057]" class="Insecta" family="Nitidulidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="468" phylum="Arthropoda" rank="family">Nitidulidae</taxonomicName>
</paragraph>
<paragraph id="8BD57A60FF8CFFA997A10620FB59FBA0" blockId="7.[152,1173,194,1634]" box="[205,1170,1052,1082]" pageId="7" pageNumber="468">
pentagonal (
<figureCitation id="135166E5FF8CFFA9962C0620FE43FBA8" box="[320,392,1052,1074]" captionStart="Fig" captionStartId="4.[197,228,1392,1411]" captionTargetBox="[179,1180,197,1366]" captionTargetId="figure-83@4.[173,1193,193,1375]" captionTargetPageId="4" captionText="Fig. 2. Urophorus (Anophorus) cordiformis, new species. A) Labrum, dorsal view, B) Labial palpi with mentum, ventral view, C) Right maxilla, ventral view, D) Right mandible, dorsal view, E) Head capsule, dorsal view, F) Right antenna, dorsal view, G) Abdominal sternite VI of male, ventral view, H) Abdominal sternite VII of male, ventral view, I) Spiculum gastrale, ventral view, J) Anal sclerite, ventral view, K) Abdominal sternite VI of female, ventral view, L) Abdominal sternite VII of female, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13243315" httpUri="https://zenodo.org/record/13243315/files/figure.png" pageId="7" pageNumber="468">Fig. 2L</figureCitation>
). Male genitalia as figured [pp. 459491].
<emphasis id="B91EA672FF8CFFA994E9061BFC5CFBA0" box="[901,919,1063,1081]" italics="true" pageId="7" pageNumber="468">In</emphasis>
: Catalogue of Palaearctic
</paragraph>
<paragraph id="8BD57A60FF8CFFA997A1060BFB59FBCB" blockId="7.[152,1173,194,1634]" box="[205,1170,1079,1106]" pageId="7" pageNumber="468">
(
<figureCitation id="135166E5FF8CFFA997B8060BFE8FFBD5" box="[212,324,1079,1101]" captionStart="Fig" captionStartId="5.[176,207,711,730]" captionTargetBox="[178,1164,202,691]" captionTargetId="figure-69@5.[151,1171,193,693]" captionTargetPageId="5" captionText="Fig. 3. Genitalia of Urophorus (Anophorus) cordiformis, new species. AC) Male genitalia, ventral, lateral, and dorsal views, respectively, D) Ovipositor, ventral view." figureDoi="http://doi.org/10.5281/zenodo.13243317" httpUri="https://zenodo.org/record/13243317/files/figure.png" pageId="7" pageNumber="468">Figs. 3AC</figureCitation>
). .................................................
<taxonomicName id="4C6A01E3FF8CFFA995810602FC9AFBC8" authorityName="Linnaeus" authorityYear="1758" box="[749,849,1086,1105]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="468" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
, 4. Elateroidea Derodontoidea
</paragraph>
<paragraph id="8BD57A60FF8CFFA997A1066EFB5FFB18" blockId="7.[152,1173,194,1634]" pageId="7" pageNumber="468">
<emphasis id="B91EA672FF8CFFA997A1066EFDCDFBFE" bold="true" box="[205,518,1105,1127]" pageId="7" pageNumber="468">
<emphasis id="B91EA672FF8CFFA997A1066EFF15FBFE" bold="true" box="[205,222,1106,1127]" italics="true" pageId="7" pageNumber="468">U</emphasis>
.
<taxonomicName id="4C6A01E3FF8CFFA9978B066DFDCDFBFE" authority="Ito and Powell" authorityName="Ito and Powell" authorityYear="2023" box="[231,518,1105,1127]" class="Insecta" family="Nitidulidae" genus="Urophorus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="468" phylum="Arthropoda" rank="species" species="cordiformis" status="sp. nov." subGenus="Anophorus">
(
<emphasis id="B91EA672FF8CFFA99783066EFF35FBFE" bold="true" box="[239,254,1106,1127]" italics="true" pageId="7" pageNumber="468">A</emphasis>
.)
<emphasis id="B91EA672FF8CFFA99663066DFEB0FBFE" bold="true" box="[271,379,1105,1127]" italics="true" pageId="7" pageNumber="468">cordiformis</emphasis>
Ito and Powell
</taxonomicName>
</emphasis>
,
<emphasis id="B91EA672FF8CFFA99563066EFDB6FBFE" bold="true" box="[527,637,1105,1127]" pageId="7" pageNumber="468">
<taxonomicNameLabel id="A22D1B09FF8CFFA99563066EFDB6FBFE" box="[527,637,1105,1127]" pageId="7" pageNumber="468" rank="species">new species</taxonomicNameLabel>
</emphasis>
Bostrichoidea Lymexyloidea Cleroidea 1. Posterior corner of pronotum an obtuse angle. Cucujoidea (I. Löbl and A. Smetana, editors).
</paragraph>
<paragraph id="8BD57A60FF8CFFA997A106BBFBC7FB03" blockId="7.[152,1173,194,1634]" box="[205,1036,1158,1181]" pageId="7" pageNumber="468">Antennomere IX as wide as or wider than an- Apollo Books, Stenstrup, 935 pp.</paragraph>
<paragraph id="8BD57A60FF8CFFA997A1069EFB5EFB50" blockId="7.[152,1173,194,1634]" pageId="7" pageNumber="468">
tennomere X. Posterior margin of abdominal
<emphasis id="B91EA672FF8CFFA995C106A2FC43FB28" bold="true" box="[685,904,1182,1201]" pageId="7" pageNumber="468">Kirejtshuk, A. G. 1990.</emphasis>
New taxa of nitidulid beetles sternite VI distinctly emarginate and without (
<taxonomicName id="4C6A01E3FF8CFFA9959F068AFC84FB50" authorityName="Linnaeus" authorityYear="1758" box="[755,847,1206,1225]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="468" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
,
<taxonomicName id="4C6A01E3FF8CFFA99439068AFC7AFB50" authorityName="Latreille" authorityYear="1802" box="[853,945,1206,1225]" class="Insecta" family="Nitidulidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="468" phylum="Arthropoda" rank="family">Nitidulidae</taxonomicName>
) of the Eastern Hemisphere.
</paragraph>
<paragraph id="8BD57A60FF8CFFA997A106EBFB5FFB60" blockId="7.[152,1173,194,1634]" pageId="7" pageNumber="468">a pair of small projections near the middle. Part 4. Proceedings of the Zoological Institute of the USSR Academy of Sciences [in Russian] 211:</paragraph>
<paragraph id="8BD57A60FF8CFFA997A106CEFCFFFA88" blockId="7.[152,1173,194,1634]" box="[205,820,1266,1297]" pageId="7" pageNumber="468">Minute projections of abdominal sternite VII 84103.</paragraph>
<paragraph id="8BD57A60FF8CFFA997A10730FB58FAB3" blockId="7.[152,1173,194,1634]" box="[205,1171,1292,1322]" pageId="7" pageNumber="468">
of male in a circular arrangement.Abdominal
<emphasis id="B91EA672FF8CFFA995C1072AFC20FAB0" bold="true" box="[685,1003,1302,1321]" pageId="7" pageNumber="468">Komagata, S., and K. Ogawa. 2022.</emphasis>
Digital transforma-
</paragraph>
<paragraph id="8BD57A60FF8CFFA997A1071BFB58FAC3" blockId="7.[152,1173,194,1634]" pageId="7" pageNumber="468">sternite VII of female triangular. Male genita- tion for entomology: Scientific drawing of insect lia with lateral lobe narrowed from distal half morphology using digital devices. Japanese Jour-</paragraph>
<paragraph id="8BD57A60FF8CFFA997A10760FB59FA13" blockId="7.[152,1173,194,1634]" pageId="7" pageNumber="468">
towards apex .................................................2 nal of Entomology (New Series) 25(3): 128143. 2. Body narrower, with long, dense setae.
<emphasis id="B91EA672FF8CFFA995C1074AFCA6FA10" bold="true" box="[685,877,1398,1417]" pageId="7" pageNumber="468">Maruyama, M. 2004.</emphasis>
A permanent slide pinned under a
</paragraph>
<paragraph id="8BD57A60FF8CFFA997A107AEFBC1FA38" blockId="7.[152,1173,194,1634]" box="[205,1034,1422,1448]" pageId="7" pageNumber="468">
Dorsolateral fovea on pronotum indistinct. specimen. Elytra,
<collectingRegion id="49AEB482FF8CFFA994EB07B2FC76FA38" box="[903,957,1422,1441]" country="Japan" name="Tokyo" pageId="7" pageNumber="468">Tokyo</collectingRegion>
32: 276.
</paragraph>
<paragraph id="8BD57A60FF8CFFA997A10790FB58FA20" blockId="7.[152,1173,194,1634]" box="[205,1171,1446,1474]" pageId="7" pageNumber="468">
Pygidium of female without longitudinal
<emphasis id="B91EA672FF8CFFA995C1079AFB58FA20" bold="true" box="[685,1171,1446,1465]" pageId="7" pageNumber="468">Powell, G. S., A. R. Cline, A. G. Duffy, and J. M. Zaspel.</emphasis>
</paragraph>
<paragraph id="8BD57A60FF8CFFA997A107FBFB58FA70" blockId="7.[152,1173,194,1634]" pageId="7" pageNumber="468">
carina. Distribution:
<collectingCountry id="F37D3AF0FF8CFFA996E607FBFE75FA45" box="[394,446,1479,1500]" name="Japan" pageId="7" pageNumber="468">Japan</collectingCountry>
(
<collectingRegion id="49AEB482FF8CFFA996A607FBFDE3FA44" box="[458,552,1479,1501]" country="Japan" name="Hokkaido" pageId="7" pageNumber="468">Hokkaido</collectingRegion>
, Honshu,
<emphasis id="B91EA672FF8CFFA995810782FCD7FA48" bold="true" box="[749,796,1470,1489]" pageId="7" pageNumber="468">2020.</emphasis>
Phylogeny and reclassification of
<taxonomicName id="4C6A01E3FF8CFFA993350782FCE4FA70" authorityName="Erichson" authorityYear="1843" class="Insecta" family="Nitidulidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="468" phylum="Arthropoda" rank="subFamily" subFamily="Carpophilinae">Carpophilinae</taxonomicName>
(
<taxonomicName id="4C6A01E3FF8CFFA9942D07EAFC54FA70" authorityName="Linnaeus" authorityYear="1758" box="[833,927,1494,1513]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="468" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
:
<taxonomicName id="4C6A01E3FF8CFFA994C207EAFBC6FA70" authorityName="Latreille" authorityYear="1802" box="[942,1037,1494,1513]" class="Insecta" family="Nitidulidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="468" phylum="Arthropoda" rank="family">Nitidulidae</taxonomicName>
), with insights
</paragraph>
<paragraph id="8BD57A60FF8CFFA997A107DEFB58F998" blockId="7.[152,1173,194,1634]" box="[205,1171,1506,1537]" pageId="7" pageNumber="468">Shikoku, Kyushu, the Ryukyu Archipelago), into the origins of anthophily. Zoological Journal of</paragraph>
<paragraph id="8BD57A60FF8CFFA997A107C0FDB5F9B5" blockId="7.[152,1173,194,1634]" pageId="7" pageNumber="468">
cosmopolitan .....
<emphasis id="B91EA672FF8CFFA9961D07C1FDB6F98B" bold="true" box="[369,637,1532,1554]" pageId="7" pageNumber="468">
<emphasis id="B91EA672FF8CFFA9961D07C1FE4AF98B" bold="true" box="[369,385,1533,1554]" italics="true" pageId="7" pageNumber="468">U</emphasis>
.
<taxonomicName id="4C6A01E3FF8CFFA996E707C0FDB6F98B" authority="(Fabricius)" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[395,637,1532,1554]" class="Insecta" family="Nitidulidae" genus="Urophorus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="468" phylum="Arthropoda" rank="species" species="humeralis" subGenus="Anophorus">
(
<emphasis id="B91EA672FF8CFFA996FF07C0FE69F988" bold="true" box="[403,418,1532,1553]" italics="true" pageId="7" pageNumber="468">A</emphasis>
.)
<emphasis id="B91EA672FF8CFFA996DE07C0FDDBF98B" bold="true" box="[434,528,1532,1554]" italics="true" pageId="7" pageNumber="468">humeralis</emphasis>
(Fabricius)
</taxonomicName>
</emphasis>
the Linnean Society 189(4): 13591369. 2. Body broader, with short, sparse setae.
</paragraph>
<paragraph id="8BD57A60FF8CFFA997A1040EFB58F9D3" blockId="7.[152,1173,194,1634]" box="[205,1171,1586,1610]" pageId="7" pageNumber="468">
Dorsolateral fovea on pronotum distinct. (Received
<date id="FFD45CA0FF8CFFA99466040AFCBEF9D0" box="[778,885,1590,1610]" pageId="7" pageNumber="468" value="2023-06-01">1 June 2023</date>
; accepted
<date id="FFD45CA0FF8CFFA994BC040AFB89F9D0" box="[976,1090,1590,1609]" pageId="7" pageNumber="468" value="2023-07-30">30 July 2023</date>
. Publica-
</paragraph>
<paragraph id="8BD57A60FF8CFFA997A10470FC64F9F8" blockId="7.[152,1173,194,1634]" box="[205,943,1612,1634]" pageId="7" pageNumber="468">
Pygidium of female with longitudinal carina. tion date
<date id="FFD45CA0FF8CFFA995910472FC6DF9F8" box="[765,934,1614,1633]" pageId="7" pageNumber="468" value="2023-09-29">29 September 2023</date>
.)
</paragraph>
</subSubSection>
</treatment>
</document>