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<document id="7362E4FFA46EADB3D28747B7525DD72F" ID-DOI="10.5281/zenodo.194461" ID-GBIF-Dataset="273cd7a4-269e-4278-ac95-58a213cc027b" ID-ISSN="1175-5326" ID-Zenodo-Dep="194461" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460213889124" checkinUser="plazi" docAuthor="Liu, Nan, Zhao, Yunyun &amp; Ren, Dong" docDate="2010" docId="03BAD533FFD6381CFF5EFACD6992FEF1" docLanguage="en" docName="zt02420p045.pdf" docOrigin="Zootaxa 2420" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Itaphlebia exquisita" docType="treatment" docVersion="8" lastPageNumber="41" masterDocId="FF83AD4BFFD43818FFC9FF906C0EFF9D" masterDocTitle="Two new fossil species of Itaphlebia (Mecoptera: Nannochoristidae) from Jiulongshan Formation, Inner Mongolia, China" masterLastPageNumber="45" masterPageNumber="37" pageNumber="39" updateTime="1698236826099" updateUser="plazi">
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<mods:title id="DB4ECC677F1955720428C060DF4670CC">Two new fossil species of Itaphlebia (Mecoptera: Nannochoristidae) from Jiulongshan Formation, Inner Mongolia, China</mods:title>
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<mods:name id="A2578DDCFC8EC7AF0DDC350481B798B7" type="personal">
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<mods:namePart id="DAAFCF311EE40CA0972B70E11B9AE118">Liu, Nan</mods:namePart>
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<mods:namePart id="5011AD675A3928F1B0FDD924DCBB049F">Zhao, Yunyun</mods:namePart>
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<mods:namePart id="90213FF5D97646B6921E23F42B057259">Ren, Dong</mods:namePart>
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<mods:typeOfResource id="5B5F14A917BE056BDCD60E460ACEBD48">text</mods:typeOfResource>
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<mods:title id="D3FDEBF4FC28CD48FAA58EA7DCCC429A">Zootaxa</mods:title>
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<mods:date id="AF141A939894E85BCC0D7F2BB34E4ABF">2010</mods:date>
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<mods:number id="7E9305FFCEF5DFAA2B3C6F8FC0E527D6">2420</mods:number>
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<mods:identifier id="90927DDAEC428D2D6054049CDE893FD6" type="DOI">10.5281/zenodo.194461</mods:identifier>
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<mods:identifier id="77C1CF0D0777D10012F8F7DC1022D140" type="Zenodo-Dep">194461</mods:identifier>
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<treatment id="03BAD533FFD6381CFF5EFACD6992FEF1" ID-DOI="http://doi.org/10.5281/zenodo.6200710" ID-GBIF-Taxon="119383867" ID-Zenodo-Dep="6200710" LSID="urn:lsid:plazi:treatment:03BAD533FFD6381CFF5EFACD6992FEF1" httpUri="http://treatment.plazi.org/id/03BAD533FFD6381CFF5EFACD6992FEF1" lastPageId="4" lastPageNumber="41" pageId="2" pageNumber="39">
<subSubSection id="C30937AEFFD6381AFF5EFACD6DFDFA07" pageId="2" pageNumber="39" type="nomenclature">
<paragraph id="8BAC6425FFD6381AFF5EFACD6DE5FAEB" blockId="2.[151,499,1372,1434]" box="[151,491,1372,1399]" pageId="2" pageNumber="39">
<heading id="D0E4D349FFD6381AFF5EFACD6DE5FAEB" bold="true" box="[151,491,1372,1399]" fontSize="11" level="1" pageId="2" pageNumber="39" reason="1">
<emphasis id="B967B837FFD6381AFF5EFACD6DE5FAEB" bold="true" box="[151,491,1372,1399]" pageId="2" pageNumber="39">
<emphasis id="B967B837FFD6381AFF5EFACD6D96FAEA" bold="true" box="[151,408,1373,1399]" italics="true" pageId="2" pageNumber="39">
<taxonomicName id="4C131FA6FFD6381AFF5EFACD6D88FAEA" box="[151,390,1373,1399]" class="Insecta" family="Nannochoristidae" genus="Itaphlebia" kingdom="Animalia" order="Mecoptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="exquisita">Itaphlebia exquisita</taxonomicName>
s
</emphasis>
p. nov.
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAC6425FFD6381AFF5EFA106DFDFA07" blockId="2.[151,499,1372,1434]" box="[151,499,1408,1434]" pageId="2" pageNumber="39">
(
<figureCitation id="132878A0FFD6381AFF56FA106D0CFA07" box="[159,258,1408,1434]" captionStart="FIGURE 2" captionStartId="3.[151,255,1661,1685]" captionTargetBox="[305,1285,371,1624]" captionTargetId="figure@3.[295,1292,355,1637]" captionTargetPageId="3" captionText="FIGURE 2. Itaphlebia exquisita sp. nov. A. Holotype, part CNU-MEC-NN 2009145 - 1; B. Holotype, counterpart CNU- MEC-NN 2009145 - 2; C. Middle leg; D. Paratype, CNU-MEC-NN 2009236; E. Portion of wings of paratype." httpUri="https://zenodo.org/record/194463/files/figure.png" pageId="2" pageNumber="39">Figure 2</figureCitation>
AE,
<figureCitation id="132878A0FFD6381AFE86FA106DBCFA07" box="[335,434,1408,1434]" captionStart="FIGURE 3" captionStartId="4.[151,255,1613,1637]" captionTargetBox="[320,1278,440,1555]" captionTargetId="figure@4.[303,1284,421,1588]" captionTargetPageId="4" captionText="FIGURE 3. Itaphlebia exquisita sp. nov. Line drawings of holotype. A. Ventral view of body with wings; B. Dorsal view of body with wings. C. Left forewing; D. Left hind wing; E. Right forewing; F. Right hind wing; G. Left middle leg." httpUri="https://zenodo.org/record/194464/files/figure.png" pageId="2" pageNumber="39">Figure 3</figureCitation>
FL)
</paragraph>
</subSubSection>
<subSubSection id="C30937AEFFD6381AFF5EFA5A6DA5F9C9" pageId="2" pageNumber="39" type="materials_examined">
<paragraph id="8BAC6425FFD6381AFF5EFA5A6E9AF997" blockId="2.[151,1437,1482,2012]" pageId="2" pageNumber="39">
<emphasis id="B967B837FFD6381AFF5EFA5A6D0AFA79" bold="true" box="[151,260,1482,1508]" pageId="2" pageNumber="39">Material</emphasis>
.
<typeStatus id="54A8DA87FFD6381AFEDBFA5A6D71FA79" box="[274,383,1482,1508]" pageId="2" pageNumber="39" type="holotype">Holotype</typeStatus>
female, CNU-MEC-NN2009145-1 and CNU-MEC-NN2009145-2, positive and negative.
<typeStatus id="54A8DA87FFD6381AFF5EFA606D0EF997" box="[151,256,1520,1546]" pageId="2" pageNumber="39" type="paratype">Paratype</typeStatus>
, female, CNU-MEC-NN2009236.
</paragraph>
<paragraph id="8BAC6425FFD6381AFF0DF9846DA5F9C9" blockId="2.[151,1437,1482,2012]" pageId="2" pageNumber="39">
<emphasis id="B967B837FFD6381AFF0DF9846DC6F9B3" bold="true" box="[196,456,1556,1582]" pageId="2" pageNumber="39">Horizon and locality</emphasis>
. Jiulongshan Formation, Middle Jurassic, Daohugou Village, Ningcheng County, Inner
<collectingCountry id="F30424B5FFD6381AFF15F9AA6D5FF9C9" box="[220,337,1594,1620]" name="Mongolia" pageId="2" pageNumber="39">Mongolia</collectingCountry>
,
<collectingCountry id="F30424B5FFD6381AFE95F9AA6DA8F9C9" box="[348,422,1594,1620]" name="China" pageId="2" pageNumber="39">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C30937AEFFD6381AFF0DF9CF6CCCF93F" pageId="2" pageNumber="39" type="diagnosis">
<paragraph id="8BAC6425FFD6381AFF0DF9CF6CCCF93F" blockId="2.[151,1437,1482,2012]" pageId="2" pageNumber="39">
<emphasis id="B967B837FFD6381AFF0DF9CF6D30F9E4" bold="true" box="[196,318,1631,1657]" pageId="2" pageNumber="39">Diagnosis</emphasis>
. Costal area slightly broad. M with 5 branches. Cu1+M forked beyond the cross-vein cu1+mcu2.
</paragraph>
</subSubSection>
<subSubSection id="C30937AEFFD6381BFF0DF9206953F840" lastPageId="3" lastPageNumber="40" pageId="2" pageNumber="39" type="description">
<paragraph id="8BAC6425FFD6381AFF0DF9206825F888" blockId="2.[151,1437,1482,2012]" pageId="2" pageNumber="39">
<emphasis id="B967B837FFD6381AFF0DF9206D59F957" bold="true" box="[196,343,1712,1738]" pageId="2" pageNumber="39">Description</emphasis>
. Antenna filiform, with 16 segments visible. Legs slender. Tibia with 2 tibial spurs at apex longer than femur. Tarsus with 5 tarsomeres; basitarsus approximately equal to half tarsus. Two claws at end of pretarsus. Surface of all legs covered with setae. Setae irregularly arranged.
</paragraph>
<paragraph id="8BAC6425FFD6381BFF0DF8B06D50FEB6" blockId="2.[151,1437,1482,2012]" lastBlockId="3.[151,1436,152,299]" lastPageId="3" lastPageNumber="40" pageId="2" pageNumber="39">In forewings, front edge of C somewhat convex. Costal area slightly broad. A distinct pterostigma present. Sc with 2 branches. Sc1 ending at C less than half length of forewing, and Sc2 terminating at C near pterostigmal area. One short cross-vein sc2-r1 before pterostigma. Cross-vein r1-r2 under pterostigma. Crossvein r3-r4 at same level of cross-vein r1-r2. Cross-vein r5-m1+2 approximately at middle of wing length. Crossvein r5-m1 before cross-vein r4-r5. Rs with 4 branches. Conspicuous thyridium at the fork of M. M with 5 branches. M1 with 2 branches (M1a, M1b). M3+4 divided beyond the fork of M1+2. Cross-vein m1+2-m3 between M1+2 and M3. Cross-vein m-cu not straight, somewhat S-shaped. Cu1 coalesced with M for a short distance and separated from M beyond cross-vein cu1+m-cu2. 1A and 2A almost parallel, one cross-vein a1-a2 between them. 3A absent.</paragraph>
<caption id="DF6C34ADFFD7381BFF5EF9ED6914F929" httpUri="https://zenodo.org/record/194463/files/figure.png" pageId="3" pageNumber="40" targetBox="[305,1285,371,1624]" targetPageId="3">
<paragraph id="8BAC6425FFD7381BFF5EF9ED6914F929" blockId="3.[151,1436,1661,1716]" pageId="3" pageNumber="40">
<emphasis id="B967B837FFD7381BFF5EF9ED6D17F908" bold="true" box="[151,281,1661,1685]" pageId="3" pageNumber="40">FIGURE 2.</emphasis>
<taxonomicName id="4C131FA6FFD7381BFEE8F9ED6DF8F909" box="[289,502,1661,1684]" class="Insecta" family="Nannochoristidae" genus="Itaphlebia" kingdom="Animalia" order="Mecoptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="exquisita" status="sp. nov.">
<emphasis id="B967B837FFD7381BFEE8F9ED6DF8F909" box="[289,502,1661,1684]" italics="true" pageId="3" pageNumber="40">Itaphlebia exquisita</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A254054CFFD7381BFE34F9ED6E5DF908" box="[509,595,1661,1685]" pageId="3" pageNumber="40" rank="species">
<emphasis id="B967B837FFD7381BFE34F9ED6E5DF908" bold="true" box="[509,595,1661,1685]" pageId="3" pageNumber="40">sp. nov.</emphasis>
</taxonomicNameLabel>
A. Holotype, part CNU-MEC-NN2009145-1; B. Holotype, counterpart CNU- MEC-NN2009145-2; C. Middle leg; D. Paratype, CNU-MEC-NN2009236; E. Portion of wings of paratype.
</paragraph>
</caption>
<paragraph id="8BAC6425FFD7381BFF0DF8926F22F8F0" blockId="3.[151,1436,1794,2013]" pageId="3" pageNumber="40">
<collectingCountry id="F30424B5FFD7381BFF0DF8926D0FF881" box="[196,257,1794,1820]" name="India" pageId="3" pageNumber="40">Hind</collectingCountry>
wings shorter than forewings. Sc simple, terminating at C much earlier before pterostigma. R1 curved under pterostigma and connected with C by a cross-vein c-r1. M1 divided into 2 branches in right hind wing, but not forked in left hind wing; Anal vein absent.
</paragraph>
<paragraph id="8BAC6425FFD7381BFF0DF8E86953F840" blockId="3.[151,1436,1794,2013]" pageId="3" pageNumber="40">
Abdomen of female with 11 segments, segments 911 distinctly smaller than segment 8. A pair of cerci at the end of abdomen, each one with 3 segments, basal segments fused the eleventh abdomen segment. Body length
<quantity id="4CEBC9C0FFD7381BFEE7F8536D85F840" box="[302,395,1987,2013]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="3" pageNumber="40" unit="mm" value="9.3">9.3 mm</quantity>
; forewing length
<quantity id="4CEBC9C0FFD7381BFD90F8536ECAF840" box="[601,708,1987,2013]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" pageId="3" pageNumber="40" unit="mm" value="10.2">10.2 mm</quantity>
, width
<quantity id="4CEBC9C0FFD7381BFCD0F8536F79F840" box="[793,887,1987,2013]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="40" unit="mm" value="3.5">3.5 mm</quantity>
; hind wing length
<quantity id="4CEBC9C0FFD7381BFB99F85368A3F840" box="[1104,1197,1987,2013]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="3" pageNumber="40" unit="mm" value="8.7">8.7 mm</quantity>
, width
<quantity id="4CEBC9C0FFD7381BFACAF8536957F840" box="[1283,1369,1987,2013]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="3" pageNumber="40" unit="mm" value="3.3">3.3mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C30937AEFFD0381CFF0FFF076D03FF4A" pageId="4" pageNumber="41" type="etymology">
<paragraph id="8BAC6425FFD0381CFF0FFF076D03FF4A" blockId="4.[151,1437,151,364]" pageId="4" pageNumber="41">
<emphasis id="B967B837FFD0381CFF0FFF076D5DFF2C" bold="true" box="[198,339,151,177]" pageId="4" pageNumber="41">Etymology.</emphasis>
The specific epithet derives from Latin word
<emphasis id="B967B837FFD0381CFC43FF086FF1FF2C" box="[906,1023,152,177]" italics="true" pageId="4" pageNumber="41">exquisitus</emphasis>
(exquisite), for its well-preserved condition.
</paragraph>
</subSubSection>
<subSubSection id="C30937AEFFD0381CFF0DFF726992FEF1" pageId="4" pageNumber="41" type="discussion">
<paragraph id="8BAC6425FFD0381CFF0DFF726992FEF1" blockId="4.[151,1437,151,364]" pageId="4" pageNumber="41">
<emphasis id="B967B837FFD0381CFF0DFF726D34FF61" bold="true" box="[196,314,226,252]" pageId="4" pageNumber="41">Remarks</emphasis>
.
<taxonomicName id="4C131FA6FFD0381CFE82FF726DC3FF66" box="[331,461,226,251]" class="Insecta" family="Nannochoristidae" genus="Itaphlebia" kingdom="Animalia" order="Mecoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="exquisita" status="sp. nov.">
<emphasis id="B967B837FFD0381CFE82FF726DC3FF66" box="[331,461,226,251]" italics="true" pageId="4" pageNumber="41">I.exquisita</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A254054CFFD0381CFE1CFF726E37FF61" box="[469,569,226,252]" pageId="4" pageNumber="41" rank="species">
<emphasis id="B967B837FFD0381CFE1CFF726E37FF61" bold="true" box="[469,569,226,252]" pageId="4" pageNumber="41">sp. nov.</emphasis>
</taxonomicNameLabel>
resembles
<taxonomicName id="4C131FA6FFD0381CFD0CFF726F29FF66" box="[709,807,226,251]" class="Insecta" family="Nannochoristidae" genus="Itaphlebia" kingdom="Animalia" order="Mecoptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="multa">
<emphasis id="B967B837FFD0381CFD0CFF726F29FF66" box="[709,807,226,251]" italics="true" pageId="4" pageNumber="41">I. multa</emphasis>
</taxonomicName>
in venation pattern: a distinct pterostigma, similar pattern of longitudinal veins, Rs 4-branched; and the same number and location of cross-veins. However, it differs from the latter by having M 5-branched in the forewings and right hind wing, although the left hind wing has M 4-branched. A 5-branched M, also seen in the
<typeStatus id="54A8DA87FFD0381CFC8BFEC26FA4FEF1" box="[834,938,338,364]" pageId="4" pageNumber="41" type="paratype">paratype</typeStatus>
, appears to be a relatively stable character.
</paragraph>
</subSubSection>
</treatment>
</document>