474 lines
59 KiB
XML
474 lines
59 KiB
XML
<document id="421250CFBB03EF2690C7F1A2F48E9AC2" ID-CLB-Dataset="163377" ID-DOI="10.11646/zootaxa.5195.3.3" ID-GBIF-Dataset="fd2040c7-2e4e-4c36-87ce-85efe9f45003" ID-ISSN="1175-5326" ID-Zenodo-Dep="7187839" ID-ZooBank="A9963C93-290B-44BA-BBDB-0C4B0299FDB5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1665554369870" checkinUser="plazi" docAuthor="Nakahara, Shinichi, Zilli, Alberto, Calhoun, John V., Espeland, Marianne, Padrón, Pablo Sebastián & Grishin, Nick V." docDate="2022" docId="03B787FAA55C717C4E85FE8ABAC79D3C" docLanguage="en" docName="zootaxa.5195.3.3.pdf" docOrigin="Zootaxa 5195 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Vettius phyllides Rober 1925" docType="treatment" docVersion="7" lastPageNumber="252" masterDocId="FF8EFF82A55671774E12FFE5BE3A9E29" masterDocTitle="Resolving two centuries of mistaken identity: Reinterpretation of Papilio marcus Fabricius, 1787 (Lepidoptera: Nymphalidae, Hesperiidae)" masterLastPageNumber="255" masterPageNumber="241" pageNumber="251" updateTime="1699407275020" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="82E81243DD2885C77D0417A9F9E58220" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="55C4BEE1CE8FC0572BBC075FF1E60325">
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<mods:title id="B623E43D50C24882F6546003F9E7F8DE">Resolving two centuries of mistaken identity: Reinterpretation of Papilio marcus Fabricius, 1787 (Lepidoptera: Nymphalidae, Hesperiidae)</mods:title>
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<mods:name id="3FA1773D6173DD5D8F08B6B75F43AD25" type="personal">
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<mods:roleTerm id="1D02B8D3336D9E5956CC28D8C6D6B063">Author</mods:roleTerm>
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<mods:namePart id="09A9E117EB4A7945008442F20C410012">Nakahara, Shinichi</mods:namePart>
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<mods:affiliation id="B3A4629BE36E1463D6CD9663F43C9A75">Department of Organismic and Evolutionary Biology, Museum of Comparative Zoology, Harvard University, 26 Oxford St. Cambridge, MA 02318, USA</mods:affiliation>
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<mods:name id="A87327E5CE88293705F61EB277398BCD" type="personal">
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<mods:namePart id="B444248D1EC8931433DA99DE5B41FD9E">Zilli, Alberto</mods:namePart>
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<mods:affiliation id="212EE8B69522F2F25E1731BB9BDB8E8F">Insects Division, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
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</mods:name>
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<mods:name id="B385726D9DA11585DAA1A56138055EBE" type="personal">
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<mods:roleTerm id="B1A3B03DC5FC0E50914F4340B85AE186">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D3F2B7864A34D2EF45F96E4EBAA64D4D">Calhoun, John V.</mods:namePart>
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<mods:affiliation id="316043B0CBEA99549C7B0DCECC230AB5">McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Florida, Gainesville, FL 32611, USA</mods:affiliation>
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</mods:name>
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<mods:name id="280A00B93A7912978E8F2E3322C4EB5C" type="personal">
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<mods:roleTerm id="51C024DC8C77FAA500C0C18A262B7353">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="796B8583D804D4B4B35FA68478D29B4F">Espeland, Marianne</mods:namePart>
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<mods:affiliation id="2355F262F86AE2BC3B5AA4D4B29189A6">Leibniz Institute for the Analysis of Biodiversity Change, Zoological Research Museum Alexander Koenig, Adenauerallee 127 53113 Bonn, Germany</mods:affiliation>
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</mods:name>
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<mods:name id="E11ABC3C9920F040C3C93CE3038DE982" type="personal">
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</mods:role>
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<mods:namePart id="1FF462949388254514B08EC63DE5ED22">Padrón, Pablo Sebastián</mods:namePart>
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<mods:affiliation id="592E80EC6D8C5A161713F853F1ECDEA0">Laboratorio de Entomología, Museo de Zoología, Universidad del Azuay, Cuenca, Ecuador</mods:affiliation>
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</mods:name>
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<mods:name id="87C628B6300585869E9580431C8BD33F" type="personal">
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<mods:namePart id="C97BAB4FE3EAAF0ACB13C27A6A4E9024">Grishin, Nick V.</mods:namePart>
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<mods:affiliation id="8CD3F9DD5B4739D052E106B414BB6236">Howard Hughes Medical Institute, Departments of Biophysics and Biochemistry, University of Texas Southwestern Medical Center, 5323 Harry Hines Blvd., Dallas, TX 75390 - 9050, USA</mods:affiliation>
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<mods:typeOfResource id="7DE5E8A766FDB7B30ACDD687EE6CD103">text</mods:typeOfResource>
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<mods:title id="A49B8E812F4DF0E586CC50A6123654AB">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="52387236DC6E4468718F95FE1388E37D">
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<mods:date id="FA45090ABE723594150B6FC5A9D94338">2022</mods:date>
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<mods:detail id="AF242120F9A1BCF6F99607F6F093CC90" type="pubDate">
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<mods:number id="95009FDC6E3F14EAE73BF0CAC96A4A9D">2022-10-12</mods:number>
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</mods:detail>
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<mods:detail id="20F416E193055B5D3800054C67394947" type="volume">
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<mods:number id="3CC07EE737A8E38A48623984A7B8878B">5195</mods:number>
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</mods:detail>
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<mods:detail id="05026337EA226F19DA3738AD92099186" type="issue">
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<mods:number id="D63489F281ED494EE051453E5C4B3D9C">3</mods:number>
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</mods:detail>
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<mods:extent id="E2F7CE72E4662A2ED381CBB1AE9B2747" unit="page">
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<mods:start id="9C6F74686DFFDAC763E27E4876454953">241</mods:start>
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<mods:end id="CBA2B3898C08CF02138D9D4F800C8891">255</mods:end>
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<mods:classification id="59379CEEFB204CAE0A14448C3CBD5752">journal article</mods:classification>
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<mods:identifier id="66F9C47BCEAA42A73A6733DE877F9929" type="CLB-Dataset">163377</mods:identifier>
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<mods:identifier id="8CF949DCDDCBA64AC70F8E3F019D95C7" type="DOI">10.11646/zootaxa.5195.3.3</mods:identifier>
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<mods:identifier id="B75B237A21E717C68E0C4FA3300CD509" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<treatment id="03B787FAA55C717C4E85FE8ABAC79D3C" ID-DOI="http://doi.org/10.5281/zenodo.7190638" ID-GBIF-Taxon="202596683" ID-Zenodo-Dep="7190638" LSID="urn:lsid:plazi:treatment:03B787FAA55C717C4E85FE8ABAC79D3C" httpUri="http://treatment.plazi.org/id/03B787FAA55C717C4E85FE8ABAC79D3C" lastPageId="11" lastPageNumber="252" pageId="10" pageNumber="251">
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<subSubSection id="C3046567A55C717D4E85FE8ABFC29FA1" box="[151,504,366,393]" pageId="10" pageNumber="251" type="nomenclature">
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<paragraph id="8BA136ECA55C717D4E85FE8ABFC29FA1" blockId="10.[151,504,366,393]" box="[151,504,366,393]" pageId="10" pageNumber="251">
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<heading id="D0E98180A55C717D4E85FE8ABFC29FA1" bold="true" box="[151,504,366,393]" fontSize="11" level="1" pageId="10" pageNumber="251" reason="1">
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<taxonomicName id="4C1E4D6FA55C717D4E85FE8ABFC29FA1" ID-CoL="7FR7T" authority="Rober, 1925" authorityName="Rober" authorityYear="1925" box="[151,504,366,393]" class="Insecta" family="Hesperiidae" genus="Vettius" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="phyllides">
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<emphasis id="B96AEAFEA55C717D4E85FE8ABFC29FA1" bold="true" box="[151,504,366,393]" pageId="10" pageNumber="251">
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<emphasis id="B96AEAFEA55C717D4E85FE8ABF669FA0" bold="true" box="[151,348,366,393]" italics="true" pageId="10" pageNumber="251">Vettius phyllides</emphasis>
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<bibRefCitation id="EF8F4B1DA55C717D4F71FE8ABFC29FA1" author="Rober, J. K. M." box="[355,504,366,393]" pageId="10" pageNumber="251" pagination="85 - 100" refId="ref12884" refString="Rober, J. K. M. (1925) Neue su ¨ damerikanische Falter (Lep.). Entomologische Mitteilungen, 14 (1 & 2), 85 - 100 (15 January) & 156 - 162 (10 April)." type="journal article" year="1925">Röber, 1925</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3046567A55C717C4E85FE52BAC79D3C" lastPageId="11" lastPageNumber="252" pageId="10" pageNumber="251" type="discussion">
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<paragraph id="8BA136ECA55C717D4E85FE52BDFD9A1C" blockId="10.[151,1437,439,2013]" pageId="10" pageNumber="251">
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The species-group name
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<taxonomicName id="4C1E4D6FA55C717D4FB8FE52BC359FF8" authorityName="Rober" authorityYear="1925" box="[426,527,439,465]" class="Insecta" family="Hesperiidae" genus="Vettius" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="phyllides">
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<emphasis id="B96AEAFEA55C717D4FB8FE52BC359FF8" box="[426,527,439,465]" italics="true" pageId="10" pageNumber="251">phyllides</emphasis>
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</taxonomicName>
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was established based on a single male and female individuals from
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<collectingRegion id="49DAF80EA55C717D4AE8FE52BBA69FF8" box="[1274,1436,439,465]" country="Brazil" name="Santa Catarina" pageId="10" pageNumber="251">Santa Catarina</collectingRegion>
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[
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<collectingCountry id="F309767CA55C717D4E8CFE3EBED29FDC" box="[158,232,475,501]" name="Brazil" pageId="10" pageNumber="251">Brazil</collectingCountry>
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] (
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<bibRefCitation id="EF8F4B1DA55C717D4EEDFE3EBF829FDC" author="Rober, J. K. M." box="[255,440,474,501]" pageId="10" pageNumber="251" pagination="85 - 100" refId="ref12884" refString="Rober, J. K. M. (1925) Neue su ¨ damerikanische Falter (Lep.). Entomologische Mitteilungen, 14 (1 & 2), 85 - 100 (15 January) & 156 - 162 (10 April)." type="journal article" year="1925">Röber 1925: 99</bibRefCitation>
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). The first half of the original description serves as the diagnosis to distinguish
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<taxonomicName id="4C1E4D6FA55C717D4B40FE3EBEC69C30" authorityName="Rober" authorityYear="1925" class="Insecta" family="Hesperiidae" genus="Vettius" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="phyllides">
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<emphasis id="B96AEAFEA55C717D4B40FE3EBEC69C30" italics="true" pageId="10" pageNumber="251">Vettius phyllides</emphasis>
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</taxonomicName>
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from
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<taxonomicName id="4C1E4D6FA55C717D4F50FE1ABF959C30" baseAuthorityName="Mielke" baseAuthorityYear="2005" box="[322,431,511,537]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="phyllus">
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<emphasis id="B96AEAFEA55C717D4F50FE1ABF959C30" box="[322,431,511,537]" italics="true" pageId="10" pageNumber="251">V. phyllus</emphasis>
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</taxonomicName>
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based on wing pattern differences: “
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<emphasis id="B96AEAFEA55C717D4D5FFE1ABFE19C14" italics="true" pageId="10" pageNumber="251">
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I have one pair of this species from Santa
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<taxonomicName id="4C1E4D6FA55C717D4B3BFE1ABBA69C30" authorityName="Dunker" authorityYear="1882" box="[1321,1436,511,537]" class="Polyplacophora" family="Mopaliidae" genus="Catharina" kingdom="Animalia" order="Chitonida" pageId="10" pageNumber="251" phylum="Mollusca" rank="genus">Catharina</taxonomicName>
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[
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<collectingCountry id="F309767CA55C717D4EB0FDC6BED39C14" box="[162,233,547,573]" name="Brazil" pageId="10" pageNumber="251">Brazil</collectingCountry>
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], and also one pair of
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</emphasis>
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phyllus
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<emphasis id="B96AEAFEA55C717D4C2BFDC6BE979C48" italics="true" pageId="10" pageNumber="251">from the same locality before me, both species are similar sized. The forewings of</emphasis>
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phyllades
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<emphasis id="B96AEAFEA55C717D4F37FDA2BB079C49" box="[293,1341,583,609]" italics="true" pageId="10" pageNumber="251">[sic] are somewhat narrower, and the posterior half of the hindwings is more curved in than in</emphasis>
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phyllus.
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<emphasis id="B96AEAFEA55C717D4E85FD8EBC309CAD" box="[151,522,619,645]" italics="true" pageId="10" pageNumber="251">The hyaline spots are smaller in</emphasis>
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phyllades
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<emphasis id="B96AEAFEA55C717D4C9AFD8EBD2D9CAD" box="[648,791,619,645]" italics="true" pageId="10" pageNumber="251">[sic] than in</emphasis>
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phyllus,
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<emphasis id="B96AEAFEA55C717D4D91FD8EBD9F9CAC" box="[899,933,619,645]" italics="true" pageId="10" pageNumber="251">the</emphasis>
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phyllides
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<emphasis id="B96AEAFEA55C717D4A0CFD8EBA519D3D" italics="true" pageId="10" pageNumber="251">
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female has two post-apical spots. The hind wings show clean, broad white, middle bands (without blue coloration) that start before the M
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<subScript id="179A34A9A55C717D4B30FD44BB109C86" attach="left" box="[1314,1322,673,687]" fontSize="6" pageId="10" pageNumber="251">3</subScript>
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(=hintere Radialis) and reach until the above the 3A (=Innenrandrippe), but is divided by the black 2A (=Submediana); the fringes on the inner margin and in the anal angle are strikingly white and long; the underside of the forewings is almost identical in both species, but the white subapical drawings are more developed in
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</emphasis>
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phyllides,
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<emphasis id="B96AEAFEA55C717D4AF6FD1EBDFD9A1C" italics="true" pageId="10" pageNumber="251">
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which is also the case for the light spot at the inner margin. The underside of the hindwings is white with yellow tint on the anal part; from the wing root stretching along the M
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<subScript id="179A34A9A55C717D4C62FCB0BC429D4A" attach="left" box="[624,632,853,867]" fontSize="6" pageId="10" pageNumber="251">3</subScript>
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(=Mediana?) and the M
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<subScript id="179A34A9A55C717D4D83FCB0BDA39D4A" attach="left" box="[913,921,853,867]" fontSize="6" pageId="10" pageNumber="251">1</subScript>
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(= vordere Radialis), as well as at the end of the discoidal cell, from off the 2A to the outer margin there is one narrow grey band, respectively, which both unite in a large, proximally not clearly differentiated spot, at the outer margin between M
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<subScript id="179A34A9A55C717D4A44FC78BA649D82" attach="left" box="[1110,1118,925,939]" fontSize="6" pageId="10" pageNumber="251">1</subScript>
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and Cu
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<subScript id="179A34A9A55C717D4AA8FC78BAF89D82" attach="left" box="[1210,1218,925,939]" fontSize="6" pageId="10" pageNumber="251">1</subScript>
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; in the anal angle, there, however, is a large, dark green distally and proximately undifferentiated spot set apart from the margin. The color of the upper side of the body is identical to that of the wings, but with white hairs, the collar brown, a large white spot in front of each antenna, palpi white with bluish scales; underside of the thorax and body white, and so are the legs; antennae as the wings, the clubs with yellowish underside.”
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</emphasis>
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</paragraph>
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<paragraph id="8BA136ECA55C717D4ED5FBDABD539BE8" blockId="10.[151,1437,439,2013]" pageId="10" pageNumber="251">
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Subsequently,
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<bibRefCitation id="EF8F4B1DA55C717D4F61FBDABC689A70" author="Evans, W. H." box="[371,594,1086,1113]" pageId="10" pageNumber="251" refId="ref10918" refString="Evans, W. H. (1955) A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. British Museum (Natural History), London, 499 pp." type="book" year="1955">Evans (1955: 182)</bibRefCitation>
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synonymized
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<taxonomicName id="4C1E4D6FA55C717D4D15FBDABDAA9A70" authorityName="Rober" authorityYear="1925" box="[775,912,1087,1113]" class="Insecta" family="Hesperiidae" genus="Vettius" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="phyllides">
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<emphasis id="B96AEAFEA55C717D4D15FBDABDAA9A70" box="[775,912,1087,1113]" italics="true" pageId="10" pageNumber="251">V. phyllides</emphasis>
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</taxonomicName>
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under
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<taxonomicName id="4C1E4D6FA55C717D4DFDFBDABA7B9A70" box="[1007,1089,1087,1113]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="genus">
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<emphasis id="B96AEAFEA55C717D4DFDFBDABA7B9A70" box="[1007,1089,1087,1113]" italics="true" pageId="10" pageNumber="251">Papilio</emphasis>
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</taxonomicName>
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“
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<taxonomicName id="4C1E4D6FA55C717D4A4FFBDABA949A71" authorityName="Fabricius" authorityYear="1787" box="[1117,1198,1087,1112]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="marcus">
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<emphasis id="B96AEAFEA55C717D4A4FFBDABA949A71" box="[1117,1198,1087,1112]" italics="true" pageId="10" pageNumber="251">marcus</emphasis>
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</taxonomicName>
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” without providing justification for this taxonomic rearrangement. He apparently examined specimens from southern
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<collectingCountry id="F309767CA55C717D4ACFFB86BB1E9A54" box="[1245,1316,1123,1149]" name="Brazil" pageId="10" pageNumber="251">Brazil</collectingCountry>
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,
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<collectingCountry id="F309767CA55C717D4B3DFB86BBA29A54" box="[1327,1432,1123,1149]" name="Paraguay" pageId="10" pageNumber="251">Paraguay</collectingCountry>
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, and
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<collectingCountry id="F309767CA55C717D4EDAFB62BF009A88" box="[200,314,1159,1185]" name="Argentina" pageId="10" pageNumber="251">Argentina</collectingCountry>
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, and they enabled him to reach a conclusion regarding the identity of
|
|
<taxonomicName id="4C1E4D6FA55C717D4A7EFB62BAD59A88" authorityName="Rober" authorityYear="1925" box="[1132,1263,1159,1185]" class="Insecta" family="Hesperiidae" genus="Vettius" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="phyllides">
|
|
<emphasis id="B96AEAFEA55C717D4A7EFB62BAD59A88" box="[1132,1263,1159,1185]" italics="true" pageId="10" pageNumber="251">V. phyllides</emphasis>
|
|
</taxonomicName>
|
|
, given its type locality of
|
|
<collectingRegion id="49DAF80EA55C717D4F03FB4EBF829AEC" box="[273,440,1195,1221]" country="Brazil" name="Santa Catarina" pageId="10" pageNumber="251">Santa Catarina</collectingRegion>
|
|
,
|
|
<collectingCountry id="F309767CA55C717D4FD1FB4EBC319AEC" box="[451,523,1195,1221]" name="Brazil" pageId="10" pageNumber="251">Brazil</collectingCountry>
|
|
. This status as a junior synonym of
|
|
<taxonomicName id="4C1E4D6FA55C717D4D8CFB4EBAC59AEC" authority="Fabricius, 1787" authorityName="Fabricius" authorityYear="1787" box="[926,1279,1195,1221]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="marcus">
|
|
<emphasis id="B96AEAFEA55C717D4D8CFB4EBA739AED" box="[926,1097,1195,1221]" italics="true" pageId="10" pageNumber="251">Papilio marcus</emphasis>
|
|
<bibRefCitation id="EF8F4B1DA55C717D4A42FB4EBAC59AEC" author="Fabricius, J. C." box="[1104,1279,1195,1221]" pageId="10" pageNumber="251" refId="ref10967" refString="Fabricius, J. C. (1787) Mantissa insectorvm sistens species nvper detectas adiectis synonymis, observationibvs, descriptionibvs, emendationibvs. Christ [ian] Gottl [ieb] Proft, Hafniae, [i] + 382 pp." type="book" year="1787">Fabricius, 1787</bibRefCitation>
|
|
</taxonomicName>
|
|
was followed by subsequent authors (e.g.,
|
|
<bibRefCitation id="EF8F4B1DA55C717D4FC8FB2ABC4E9AC0" author="Bridges, C. A." box="[474,628,1231,1257]" pageId="10" pageNumber="251" refId="ref10034" refString="Bridges, C. A. (1983) Lepidoptera: Hesperiidae. Notes on species-group names. Author, Urbana, [2] + ii + 129 + 41 + 62 + 30 + 13 pp." type="book" year="1983">Bridges 1983</bibRefCitation>
|
|
,
|
|
<bibRefCitation id="EF8F4B1DA55C717D4C92FB2ABC869AC0" author="Bridges, C. A." box="[640,700,1231,1257]" pageId="10" pageNumber="251" refId="ref10073" refString="Bridges, C. A. (1994) Catalogue of the family-group, genus-group and speciesgroup names of the Hesperioidea (Lepidoptera) of the world. Published by the Author, Urbana, 598 pp." type="book" year="1994">1994</bibRefCitation>
|
|
). In particular,
|
|
<bibRefCitation id="EF8F4B1DA55C717D4D7AFB2ABA799AC0" author="Mielke, O. H. H." box="[872,1091,1231,1257]" pageId="10" pageNumber="251" pagination="131 - 146" refId="ref12245" refString="Mielke, O. H. H. (1989) Sobre os tipos de Hesperiidae descritos por Roeber (Lepidoptera). Revista brasileira de Zoologia, 6 (1), 131 - 146." type="journal article" year="1989">Mielke (1989: 138)</bibRefCitation>
|
|
mentioned the examination of a pair of
|
|
<typeStatus id="54A5884EA55C717D4EECFB16BF5A9B24" box="[254,352,1267,1293]" pageId="10" pageNumber="251" type="syntype">syntypes</typeStatus>
|
|
housed at SMT and designated the male
|
|
<typeStatus id="54A5884EA55C717D4D21FB16BDB09B24" box="[819,906,1267,1293]" pageId="10" pageNumber="251" type="syntype">syntype</typeStatus>
|
|
as a
|
|
<typeStatus id="54A5884EA55C717D4DD7FB16BA169B24" box="[965,1068,1267,1293]" pageId="10" pageNumber="251" type="lectotype">lectotype</typeStatus>
|
|
(
|
|
<figureCitation id="13252A69A55C717D4A2EFB16BAAF9B24" box="[1084,1173,1267,1293]" captionStart="FIGURE 1" captionStartId="4.[152,255,1909,1933]" captionTargetBox="[294,1293,179,1882]" captionTargetId="figure-18@4.[294,1293,179,1882]" captionTargetPageId="4" captionText="FIGURE 1. Pictures of relevant Troyus species:A) sequenced Troyus phyllides from Suriname (molecular voucher: 19022E04; deposited in USNM); B) genitalia of Troyus phyllides from Rondônia, Brazil (dissection vial: SN-20-107; deposited in MGCL); C) Lectotype of Papilio marcus Fabricius, 1787 housed in ZMUC (ZMUC 00029699); D) Lectotype of Vettius phyllides housed at SMT (molecular voucher: 18094F06)." figureDoi="http://doi.org/10.5281/zenodo.7187841" httpUri="https://zenodo.org/record/7187841/files/figure.png" pageId="10" pageNumber="251">Fig. 1D</figureCitation>
|
|
), consequently treating the name as a junior subjective synonym of
|
|
<taxonomicName id="4C1E4D6FA55C717D4C93FAF2BD9F9B18" authority="Fabricius, 1787" authorityName="Fabricius" authorityYear="1787" box="[641,933,1303,1329]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="marcus">
|
|
<emphasis id="B96AEAFEA55C717D4C93FAF2BCD79B19" box="[641,749,1303,1328]" italics="true" pageId="10" pageNumber="251">P. marcus</emphasis>
|
|
<bibRefCitation id="EF8F4B1DA55C717D4CE6FAF2BD9F9B18" author="Fabricius, J. C." box="[756,933,1303,1329]" pageId="10" pageNumber="251" refId="ref10967" refString="Fabricius, J. C. (1787) Mantissa insectorvm sistens species nvper detectas adiectis synonymis, observationibvs, descriptionibvs, emendationibvs. Christ [ian] Gottl [ieb] Proft, Hafniae, [i] + 382 pp." type="book" year="1787">Fabricius, 1787</bibRefCitation>
|
|
</taxonomicName>
|
|
. The information regarding the
|
|
<typeStatus id="54A5884EA55C717D4B15FAF2BB549B18" box="[1287,1390,1303,1329]" pageId="10" pageNumber="251" type="lectotype">lectotype</typeStatus>
|
|
and
|
|
<typeStatus id="54A5884EA55C717D4E85FADEBF149B7C" box="[151,302,1339,1365]" pageId="10" pageNumber="251" type="paralectotype">paralectotype</typeStatus>
|
|
of
|
|
<taxonomicName id="4C1E4D6FA55C717D4F40FADEBFEB9B7C" authorityName="Rober" authorityYear="1925" box="[338,465,1339,1365]" class="Insecta" family="Hesperiidae" genus="Vettius" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="phyllides">
|
|
<emphasis id="B96AEAFEA55C717D4F40FADEBFEB9B7C" box="[338,465,1339,1365]" italics="true" pageId="10" pageNumber="251">V. phyllides</emphasis>
|
|
</taxonomicName>
|
|
, including associated labels, is also reflected in
|
|
<bibRefCitation id="EF8F4B1DA55C717D4DF5FADEBAD39B7C" author="Nekrutenko, Y. P." box="[999,1257,1339,1365]" pageId="10" pageNumber="251" pagination="79 - 109" refId="ref12594" refString="Nekrutenko, Y. P. (2003) Catalogue of the type specimens of Hesperiidae and Pieridae deposited at the Museum fur Tierkunde Dresden, with additions to the catalogues of Nymphalidae, Riodinidae and Lycaenidae (Insecta: Lepidoptera). Entomologische Abhandlungen, 60, 79 - 109." type="journal article" year="2003">Nekrutenko (2003: 87)</bibRefCitation>
|
|
. There exists no revisionary work on relevant groups to date, and the status of
|
|
<taxonomicName id="4C1E4D6FA55C717D4D54FABABDFC9B50" authorityName="Rober" authorityYear="1925" box="[838,966,1375,1401]" class="Insecta" family="Hesperiidae" genus="Vettius" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="phyllides">
|
|
<emphasis id="B96AEAFEA55C717D4D54FABABDFC9B50" box="[838,966,1375,1401]" italics="true" pageId="10" pageNumber="251">V. phyllides</emphasis>
|
|
</taxonomicName>
|
|
as a junior subjective synonym of
|
|
<taxonomicName id="4C1E4D6FA55C717D4B58FABABED29BB5" authorityName="Fabricius" authorityYear="1787" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="marcus">
|
|
<emphasis id="B96AEAFEA55C717D4B58FABABED29BB5" italics="true" pageId="10" pageNumber="251">Papilio marcus</emphasis>
|
|
</taxonomicName>
|
|
has been retained in all the literature regarding
|
|
<taxonomicName id="4C1E4D6FA55C717D4D14FA66BDB69BB4" box="[774,908,1411,1437]" class="Insecta" family="Hesperiidae" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="family">Hesperiidae</taxonomicName>
|
|
classification (
|
|
<bibRefCitation id="EF8F4B1DA55C717D4A25FA66BAEB9BB4" author="Mielke, O. H. H." box="[1079,1233,1411,1437]" pageId="10" pageNumber="251" pagination="3 - 11" refId="ref12282" refString="Mielke, O. H. H. (2004) Hesperiidae. In: Lamas, G. (Ed.), Checklist. Part 4 a. Hesperioidea - Papilionoidea. In: Heppner, J. B. (Ed.), Atlas of Neotropical Lepidoptera. Vol. 5 A. Association of Tropical Lepidoptera, Gainsville, pp. 3 - 11 + 25 - 86." type="book chapter" year="2004">Mielke, 2004</bibRefCitation>
|
|
,
|
|
<bibRefCitation id="EF8F4B1DA55C717D4ACFFA67BB209BB4" author="Mielke, O. H. H." box="[1245,1306,1410,1437]" pageId="10" pageNumber="251" refId="ref12356" refString="Mielke, O. H. H. (2005) Catalogue of the American Hesperioidea: Hesperiidae (Lepidoptera). Vols. 1 - 6. Sociedade Brasileira de Zoologia, Curitiba, 1536 pp." type="book" year="2005">2005</bibRefCitation>
|
|
). The name was therefore never treated as valid subsequent to
|
|
<bibRefCitation id="EF8F4B1DA55C717D4CDEFA42BD5C9BE8" author="Evans, W. H." box="[716,870,1446,1473]" pageId="10" pageNumber="251" refId="ref10918" refString="Evans, W. H. (1955) A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. British Museum (Natural History), London, 499 pp." type="book" year="1955">Evans (1955)</bibRefCitation>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="8BA136ECA55C717D4ED5FA2EBB2E992C" blockId="10.[151,1437,439,2013]" pageId="10" pageNumber="251">
|
|
Phenotypic inspection and genomic sequencing of the
|
|
<taxonomicName id="4C1E4D6FA55C717D4D24FA2EBD839BCC" authorityName="Rober" authorityYear="1925" box="[822,953,1483,1509]" class="Insecta" family="Hesperiidae" genus="Vettius" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="phyllides">
|
|
<emphasis id="B96AEAFEA55C717D4D24FA2EBD839BCC" box="[822,953,1483,1509]" italics="true" pageId="10" pageNumber="251">V. phyllides</emphasis>
|
|
</taxonomicName>
|
|
<typeStatus id="54A5884EA55C717D4DD0FA2EBA139BCC" box="[962,1065,1483,1509]" pageId="10" pageNumber="251" type="lectotype">lectotype</typeStatus>
|
|
reveals that it is not conspecific with
|
|
<taxonomicName id="4C1E4D6FA55C717D4EC1FA0ABF789820" baseAuthorityName="Mielke" baseAuthorityYear="2005" box="[211,322,1519,1545]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="phyllus">
|
|
<emphasis id="B96AEAFEA55C717D4EC1FA0ABF789820" box="[211,322,1519,1545]" italics="true" pageId="10" pageNumber="251">V. phyllus</emphasis>
|
|
</taxonomicName>
|
|
, and instead belongs to the genus
|
|
<taxonomicName id="4C1E4D6FA55C717D4CC4FA0ABD1B9821" authorityName="Warren & Turland" authorityYear="2012" box="[726,801,1519,1544]" class="Insecta" family="Hesperiidae" genus="Troyus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96AEAFEA55C717D4CC4FA0ABD1B9821" box="[726,801,1519,1544]" italics="true" pageId="10" pageNumber="251">Troyus</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<figureCitation id="13252A69A55C717D4D21FA0ABD409820" box="[819,890,1519,1545]" captionStart="FIGURE 3" captionStartId="5.[152,255,1729,1753]" captionTargetBox="[157,1426,1319,1702]" captionTargetId="figure-54@5.[151,1436,1314,1706]" captionTargetPageId="5" captionText="FIGURE 3. Phylogenetic tree of Vettius and Troyus representatives constructed from protein-coding regions of the Z chromosome." figureDoi="http://doi.org/10.5281/zenodo.7187845" httpUri="https://zenodo.org/record/7187845/files/figure.png" pageId="10" pageNumber="251">Fig. 3</figureCitation>
|
|
, red and magenta). The two genera
|
|
<taxonomicName id="4C1E4D6FA55C717D4B0DFA0ABB509821" authorityName="Godman" authorityYear="1901" box="[1311,1386,1519,1544]" class="Insecta" family="Hesperiidae" genus="Vettius" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96AEAFEA55C717D4B0DFA0ABB509821" box="[1311,1386,1519,1544]" italics="true" pageId="10" pageNumber="251">Vettius</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C1E4D6FA55C717D4E85F9F6BED89805" authorityName="Warren & Turland" authorityYear="2012" box="[151,226,1555,1580]" class="Insecta" family="Hesperiidae" genus="Troyus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96AEAFEA55C717D4E85F9F6BED89805" box="[151,226,1555,1580]" italics="true" pageId="10" pageNumber="251">Troyus</emphasis>
|
|
</taxonomicName>
|
|
are not sisters, each being recovered as sister to other genera,
|
|
<taxonomicName id="4C1E4D6FA55C717D4DA6F9F6BAFA9804" authority="Godman, [1900]" authorityName="Godman" authorityYear="1900" box="[948,1216,1555,1581]" class="Insecta" family="Hesperiidae" genus="Thoon" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96AEAFEA55C717D4DA6F9F6BDC69804" box="[948,1020,1555,1581]" italics="true" pageId="10" pageNumber="251">Thoon</emphasis>
|
|
Godman, [1900]
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="4C1E4D6FA55C717D4AE8F9F6BEE89878" authority="Evans, 1955" authorityName="Evans" authorityYear="1955" class="Insecta" family="Hesperiidae" genus="Monca" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96AEAFEA55C717D4AE8F9F6BB729805" box="[1274,1352,1555,1580]" italics="true" pageId="10" pageNumber="251">Monca</emphasis>
|
|
<bibRefCitation id="EF8F4B1DA55C717D4B43F9F6BEE89878" author="Evans, W. H." pageId="10" pageNumber="251" refId="ref10918" refString="Evans, W. H. (1955) A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. British Museum (Natural History), London, 499 pp." type="book" year="1955">Evans, 1955</bibRefCitation>
|
|
</taxonomicName>
|
|
, respectively, than to each other (
|
|
<figureCitation id="13252A69A55C717D4C5BF9D2BCB49878" box="[585,654,1591,1617]" captionStart="FIGURE 3" captionStartId="5.[152,255,1729,1753]" captionTargetBox="[157,1426,1319,1702]" captionTargetId="figure-54@5.[151,1436,1314,1706]" captionTargetPageId="5" captionText="FIGURE 3. Phylogenetic tree of Vettius and Troyus representatives constructed from protein-coding regions of the Z chromosome." figureDoi="http://doi.org/10.5281/zenodo.7187845" httpUri="https://zenodo.org/record/7187845/files/figure.png" pageId="10" pageNumber="251">Fig. 3</figureCitation>
|
|
). Because
|
|
<taxonomicName id="4C1E4D6FA55C717D4D1AF9D2BD4E9879" authorityName="Fabricius" authorityYear="1787" box="[776,884,1591,1616]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="marcus">
|
|
<emphasis id="B96AEAFEA55C717D4D1AF9D2BD4E9879" box="[776,884,1591,1616]" italics="true" pageId="10" pageNumber="251">P. marcus</emphasis>
|
|
</taxonomicName>
|
|
is a junior subjective synonym of
|
|
<taxonomicName id="4C1E4D6FA55C717D4AEBF9D2BB5C9878" baseAuthorityName="Mielke" baseAuthorityYear="2005" box="[1273,1382,1591,1617]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="phyllus">
|
|
<emphasis id="B96AEAFEA55C717D4AEBF9D2BB5C9878" box="[1273,1382,1591,1617]" italics="true" pageId="10" pageNumber="251">V. phyllus</emphasis>
|
|
</taxonomicName>
|
|
, and
|
|
<taxonomicName id="4C1E4D6FA55C717D4E85F9BEBF2C985C" authorityName="Rober" authorityYear="1925" box="[151,278,1627,1653]" class="Insecta" family="Hesperiidae" genus="Vettius" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="phyllides">
|
|
<emphasis id="B96AEAFEA55C717D4E85F9BEBF2C985C" box="[151,278,1627,1653]" italics="true" pageId="10" pageNumber="251">V. phyllides</emphasis>
|
|
</taxonomicName>
|
|
is not even congeneric with
|
|
<taxonomicName id="4C1E4D6FA55C717D4C43F9BEBC86985C" baseAuthorityName="Mielke" baseAuthorityYear="2005" box="[593,700,1627,1653]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="phyllus">
|
|
<emphasis id="B96AEAFEA55C717D4C43F9BEBC86985C" box="[593,700,1627,1653]" italics="true" pageId="10" pageNumber="251">V. phyllus</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C1E4D6FA55C717D4CDBF9BEBD72985C" authorityName="Rober" authorityYear="1925" box="[713,840,1627,1653]" class="Insecta" family="Hesperiidae" genus="Vettius" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="phyllides">
|
|
<emphasis id="B96AEAFEA55C717D4CDBF9BEBD72985C" box="[713,840,1627,1653]" italics="true" pageId="10" pageNumber="251">V. phyllides</emphasis>
|
|
</taxonomicName>
|
|
cannot remain as a synonym of
|
|
<taxonomicName id="4C1E4D6FA55C717D4AB9F9BEBB2C985D" authorityName="Fabricius" authorityYear="1787" box="[1195,1302,1627,1652]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="marcus">
|
|
<emphasis id="B96AEAFEA55C717D4AB9F9BEBB2C985D" box="[1195,1302,1627,1652]" italics="true" pageId="10" pageNumber="251">P. marcus</emphasis>
|
|
</taxonomicName>
|
|
. Reviewing all available names in
|
|
<taxonomicName id="4C1E4D6FA55C717D4F8EF99ABFDD98B1" authorityName="Warren & Turland" authorityYear="2012" box="[412,487,1663,1688]" class="Insecta" family="Hesperiidae" genus="Troyus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96AEAFEA55C717D4F8EF99ABFDD98B1" box="[412,487,1663,1688]" italics="true" pageId="10" pageNumber="251">Troyus</emphasis>
|
|
</taxonomicName>
|
|
, we were not able to find one that could be applied to
|
|
<taxonomicName id="4C1E4D6FA55C717D4A78F99ABAD798B0" authorityName="Rober" authorityYear="1925" box="[1130,1261,1663,1689]" class="Insecta" family="Hesperiidae" genus="Vettius" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="phyllides">
|
|
<emphasis id="B96AEAFEA55C717D4A78F99ABAD798B0" box="[1130,1261,1663,1689]" italics="true" pageId="10" pageNumber="251">V. phyllides</emphasis>
|
|
</taxonomicName>
|
|
. Therefore, we propose that
|
|
<taxonomicName id="4C1E4D6FA55C717D4F22F946BFD19894" box="[304,491,1699,1725]" class="Insecta" family="Hesperiidae" genus="Troyus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="phyllides">
|
|
<emphasis id="B96AEAFEA55C717D4F22F946BFD19894" box="[304,491,1699,1725]" italics="true" pageId="10" pageNumber="251">Troyus phyllides</emphasis>
|
|
</taxonomicName>
|
|
is a species-level taxon (
|
|
<emphasis id="B96AEAFEA55C717D4D0BF947BDDB9894" bold="true" box="[793,993,1698,1725]" pageId="10" pageNumber="251">reinstated status</emphasis>
|
|
,
|
|
<emphasis id="B96AEAFEA55C717D4DE1F946BAF99895" bold="true" box="[1011,1219,1698,1725]" pageId="10" pageNumber="251">new combination</emphasis>
|
|
). This species has been frequently misidentified as
|
|
<taxonomicName id="4C1E4D6FA55C717D4C1AF922BC9198C9" box="[520,683,1735,1760]" class="Insecta" family="Hesperiidae" genus="Troyus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="marcus">
|
|
<emphasis id="B96AEAFEA55C717D4C1AF922BC9198C9" box="[520,683,1735,1760]" italics="true" pageId="10" pageNumber="251">Troyus marcus</emphasis>
|
|
</taxonomicName>
|
|
: one such specimen from
|
|
<collectingCountry id="F309767CA55C717D4DC6F922BA0598C8" box="[980,1087,1735,1761]" name="Suriname" pageId="10" pageNumber="251">Suriname</collectingCountry>
|
|
is included in the tree (
|
|
<figureCitation id="13252A69A55C717D4B5BF922BBB498C8" box="[1353,1422,1735,1761]" captionStart="FIGURE 3" captionStartId="5.[152,255,1729,1753]" captionTargetBox="[157,1426,1319,1702]" captionTargetId="figure-54@5.[151,1436,1314,1706]" captionTargetPageId="5" captionText="FIGURE 3. Phylogenetic tree of Vettius and Troyus representatives constructed from protein-coding regions of the Z chromosome." figureDoi="http://doi.org/10.5281/zenodo.7187845" httpUri="https://zenodo.org/record/7187845/files/figure.png" pageId="10" pageNumber="251">Fig. 3</figureCitation>
|
|
). Finally, the phylogenetic tree reveals that
|
|
<taxonomicName id="4C1E4D6FA55C717D4C7EF90EBD23992C" baseAuthorityName="Plotz" baseAuthorityYear="1882" box="[620,793,1771,1797]" class="Insecta" family="Hesperiidae" genus="Troyus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="aurelius">
|
|
<emphasis id="B96AEAFEA55C717D4C7EF90EBD23992C" box="[620,793,1771,1797]" italics="true" pageId="10" pageNumber="251">Troyus aurelius</emphasis>
|
|
</taxonomicName>
|
|
is sister to
|
|
<taxonomicName id="4C1E4D6FA55C717D4D8AF90EBA23992C" box="[920,1049,1771,1797]" class="Insecta" family="Hesperiidae" genus="Troyus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="phyllides">
|
|
<emphasis id="B96AEAFEA55C717D4D8AF90EBA23992C" box="[920,1049,1771,1797]" italics="true" pageId="10" pageNumber="251">T. phyllides</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<figureCitation id="13252A69A55C717D4A3AF90EBA56992C" box="[1064,1132,1771,1797]" captionStart="FIGURE 3" captionStartId="5.[152,255,1729,1753]" captionTargetBox="[157,1426,1319,1702]" captionTargetId="figure-54@5.[151,1436,1314,1706]" captionTargetPageId="5" captionText="FIGURE 3. Phylogenetic tree of Vettius and Troyus representatives constructed from protein-coding regions of the Z chromosome." figureDoi="http://doi.org/10.5281/zenodo.7187845" httpUri="https://zenodo.org/record/7187845/files/figure.png" pageId="10" pageNumber="251">Fig. 3</figureCitation>
|
|
, red and blue).
|
|
</paragraph>
|
|
<paragraph id="8BA136ECA55C717D4ED5F8EABDAD9964" blockId="10.[151,1437,439,2013]" pageId="10" pageNumber="251">
|
|
In conclusion, given the circumstances surrounding the species-group names associated with
|
|
<taxonomicName id="4C1E4D6FA55C717D4AE2F8EABF729964" authority="Fabricius, 1787" authorityName="Fabricius" authorityYear="1787" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="marcus">
|
|
<emphasis id="B96AEAFEA55C717D4AE2F8EABBA69901" box="[1264,1436,1807,1833]" italics="true" pageId="10" pageNumber="251">Papilio marcus</emphasis>
|
|
<bibRefCitation id="EF8F4B1DA55C717D4E85F8D6BF729964" author="Fabricius, J. C." box="[151,328,1843,1869]" pageId="10" pageNumber="251" refId="ref10967" refString="Fabricius, J. C. (1787) Mantissa insectorvm sistens species nvper detectas adiectis synonymis, observationibvs, descriptionibvs, emendationibvs. Christ [ian] Gottl [ieb] Proft, Hafniae, [i] + 382 pp." type="book" year="1787">Fabricius, 1787</bibRefCitation>
|
|
</taxonomicName>
|
|
, we propose the following three taxonomic changes:
|
|
</paragraph>
|
|
<paragraph id="8BA136ECA55C717C4ED5F8B2BAD19EFC" blockId="10.[151,1437,439,2013]" lastBlockId="11.[151,1437,151,789]" lastPageId="11" lastPageNumber="252" pageId="10" pageNumber="251">
|
|
(1) Treatment of
|
|
<taxonomicName id="4C1E4D6FA55C717D4F9EF8B2BCCB9958" authority="Fabricius, 1787" authorityName="Fabricius" authorityYear="1787" box="[396,753,1879,1905]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="marcus">
|
|
<emphasis id="B96AEAFEA55C717D4F9EF8B2BC039959" box="[396,569,1879,1905]" italics="true" pageId="10" pageNumber="251">Papilio marcus</emphasis>
|
|
<bibRefCitation id="EF8F4B1DA55C717D4C52F8B2BCCB9958" author="Fabricius, J. C." box="[576,753,1879,1905]" pageId="10" pageNumber="251" refId="ref10967" refString="Fabricius, J. C. (1787) Mantissa insectorvm sistens species nvper detectas adiectis synonymis, observationibvs, descriptionibvs, emendationibvs. Christ [ian] Gottl [ieb] Proft, Hafniae, [i] + 382 pp." type="book" year="1787">Fabricius, 1787</bibRefCitation>
|
|
</taxonomicName>
|
|
as a junior subjective synonym of
|
|
<taxonomicName id="4C1E4D6FA55C717D4A98F8B2BBA69958" authority="Cramer, 1777" authorityName="Cramer" authorityYear="1777" box="[1162,1436,1879,1905]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="phyllus">
|
|
<emphasis id="B96AEAFEA55C717D4A98F8B2BAC09958" box="[1162,1274,1879,1905]" italics="true" pageId="10" pageNumber="251">P. phyllus</emphasis>
|
|
<bibRefCitation id="EF8F4B1DA55C717D4B12F8B2BBA69958" author="Cramer, P." box="[1280,1436,1879,1905]" pageId="10" pageNumber="251" pagination="1 - 151" refId="ref10471" refString="Cramer, P. (1777) De uitlandsche Kapellen voorkomende in de drie Waereld-Deelen Asia, Africa en America. Papillons exotiques des trois parties du monde l'Asie, l'Afrique et l'Ameirique. 2 (9 / 16). S. J. Baalde, Amsteldam, Barthelemy Wild and J. Van Schoonhoven & Comp, Utrecht, 1 - 151." type="book chapter" year="1777">Cramer, 1777</bibRefCitation>
|
|
</taxonomicName>
|
|
(
|
|
<emphasis id="B96AEAFEA55C717D4EB3F89EBF7F99BC" bold="true" box="[161,325,1915,1941]" pageId="10" pageNumber="251">new synonym</emphasis>
|
|
). By virtue of article 23.9.5 of
|
|
<bibRefCitation id="EF8F4B1DA55C717D4CA6F89EBD7699BC" author="ICZN & International Commission on Zoological Nomenclature" box="[692,844,1915,1941]" pageId="10" pageNumber="251" refId="ref11719" refString="ICZN [International Commission on Zoological Nomenclature] (1999) International Code of Zoological Nomenclature. 4 th Edition. International Trust for Zoological Nomenclature, London, 306 pp." type="book" year="1999">ICZN (1999)</bibRefCitation>
|
|
, the current placement of
|
|
<taxonomicName id="4C1E4D6FA55C717D4A6BF89EBEF59990" authority="Fabricius, 1787" authorityName="Fabricius" authorityYear="1787" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="marcus">
|
|
<emphasis id="B96AEAFEA55C717D4A6BF89EBB1C99BD" box="[1145,1318,1915,1941]" italics="true" pageId="10" pageNumber="251">Papilio marcus</emphasis>
|
|
<bibRefCitation id="EF8F4B1DA55C717D4B3FF89EBEF59990" author="Fabricius, J. C." pageId="10" pageNumber="251" refId="ref10967" refString="Fabricius, J. C. (1787) Mantissa insectorvm sistens species nvper detectas adiectis synonymis, observationibvs, descriptionibvs, emendationibvs. Christ [ian] Gottl [ieb] Proft, Hafniae, [i] + 382 pp." type="book" year="1787">
|
|
Fabricius,
|
|
<quantity id="4CE69B09A55C717D4E85F87ABEF59990" box="[151,207,1951,1977]" metricMagnitude="1" metricUnit="m" metricValue="4.53898" pageId="10" pageNumber="251" unit="in" value="1787.0">1787</quantity>
|
|
</bibRefCitation>
|
|
</taxonomicName>
|
|
in
|
|
<taxonomicName id="4C1E4D6FA55C717D4EE4F87ABFC89990" authority="Godman, 1901" authorityName="Godman" authorityYear="1901" box="[246,498,1951,1977]" class="Insecta" family="Hesperiidae" genus="Vettius" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96AEAFEA55C717D4EE4F87ABF7B9991" box="[246,321,1951,1976]" italics="true" pageId="10" pageNumber="251">Vettius</emphasis>
|
|
Godman, 1901
|
|
</taxonomicName>
|
|
was already preventing its automatic replacement as a junior primary homonym of
|
|
<taxonomicName id="4C1E4D6FA55C717D4EA4F826BC3799F4" authority="Schaller, 1785" authorityName="Schaller" authorityYear="1785" box="[182,525,1986,2013]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="marcus">
|
|
<emphasis id="B96AEAFEA55C717D4EA4F826BF5899F5" box="[182,354,1987,2013]" italics="true" pageId="10" pageNumber="251">Papilio marcus</emphasis>
|
|
Schaller, 1785
|
|
</taxonomicName>
|
|
(now in
|
|
<taxonomicName id="4C1E4D6FA55C717D4C64F826BDB699F4" authority="Fabricius, 1807" authorityName="Fabricius" authorityYear="1807" box="[630,908,1987,2013]" class="Insecta" family="Nymphalidae" genus="Morpho" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96AEAFEA55C717D4C64F826BCEA99F4" box="[630,720,1987,2013]" italics="true" pageId="10" pageNumber="251">Morpho</emphasis>
|
|
Fabricius, 1807
|
|
</taxonomicName>
|
|
), so pending referral of the case to the ICZN, prevailing usage had to be maintained. Removing
|
|
<taxonomicName id="4C1E4D6FA55D717C4CADFF72BD109E99" authorityName="Schaller" authorityYear="1785" box="[703,810,151,176]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="252" phylum="Arthropoda" rank="species" species="marcus">
|
|
<emphasis id="B96AEAFEA55D717C4CADFF72BD109E99" box="[703,810,151,176]" italics="true" pageId="11" pageNumber="252">P. marcus</emphasis>
|
|
</taxonomicName>
|
|
from use as a valid taxon following the above synonymy also eliminates referral to the ICZN as Article 23.9.5 applies only to species-group names “in use”.
|
|
</paragraph>
|
|
<paragraph id="8BA136ECA55D717C4ED5FF3ABDB79CE4" blockId="11.[151,1437,151,789]" pageId="11" pageNumber="252">
|
|
(2) Removal of the name
|
|
<taxonomicName id="4C1E4D6FA55D717C4FF9FF3ABC999ED0" authorityName="Rober" authorityYear="1925" box="[491,675,223,249]" class="Insecta" family="Hesperiidae" genus="Vettius" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="252" phylum="Arthropoda" rank="species" species="phyllides">
|
|
<emphasis id="B96AEAFEA55D717C4FF9FF3ABC999ED0" box="[491,675,223,249]" italics="true" pageId="11" pageNumber="252">Vettius phyllides</emphasis>
|
|
</taxonomicName>
|
|
from the synonymy of
|
|
<taxonomicName id="4C1E4D6FA55D717C4DA2FF3ABA279ED1" authorityName="Schaller" authorityYear="1785" box="[944,1053,223,248]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="252" phylum="Arthropoda" rank="species" species="marcus">
|
|
<emphasis id="B96AEAFEA55D717C4DA2FF3ABA279ED1" box="[944,1053,223,248]" italics="true" pageId="11" pageNumber="252">P. marcus</emphasis>
|
|
</taxonomicName>
|
|
and consideration of this specific epithet as a name to represent a concept regarded by the name
|
|
<taxonomicName id="4C1E4D6FA55D717C4D79FEE6BD879F34" box="[875,957,259,285]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="252" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96AEAFEA55D717C4D79FEE6BD879F34" box="[875,957,259,285]" italics="true" pageId="11" pageNumber="252">Papilio</emphasis>
|
|
</taxonomicName>
|
|
“
|
|
<taxonomicName id="4C1E4D6FA55D717C4DC1FEE6BA1E9F35" authorityName="Schaller" authorityYear="1785" box="[979,1060,259,284]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="252" phylum="Arthropoda" rank="species" species="marcus">
|
|
<emphasis id="B96AEAFEA55D717C4DC1FEE6BA1E9F35" box="[979,1060,259,284]" italics="true" pageId="11" pageNumber="252">marcus</emphasis>
|
|
</taxonomicName>
|
|
” for nearly 200 years, following Article 23.3.5 of
|
|
<bibRefCitation id="EF8F4B1DA55D717C4F41FEC2BFDF9F68" author="ICZN & International Commission on Zoological Nomenclature" box="[339,485,295,321]" pageId="11" pageNumber="252" refId="ref11719" refString="ICZN [International Commission on Zoological Nomenclature] (1999) International Code of Zoological Nomenclature. 4 th Edition. International Trust for Zoological Nomenclature, London, 306 pp." type="book" year="1999">ICZN (1999)</bibRefCitation>
|
|
(
|
|
<emphasis id="B96AEAFEA55D717C4FE6FEC3BC8D9F68" bold="true" box="[500,695,294,321]" pageId="11" pageNumber="252">reinstated status</emphasis>
|
|
). The rationale of applying
|
|
<taxonomicName id="4C1E4D6FA55D717C4DF8FEC2BA759F68" authorityName="Rober" authorityYear="1925" box="[1002,1103,295,321]" class="Insecta" family="Hesperiidae" genus="Vettius" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="252" phylum="Arthropoda" rank="species" species="phyllides">
|
|
<emphasis id="B96AEAFEA55D717C4DF8FEC2BA759F68" box="[1002,1103,295,321]" italics="true" pageId="11" pageNumber="252">phyllides</emphasis>
|
|
</taxonomicName>
|
|
to
|
|
<emphasis id="B96AEAFEA55D717C4A62FEC2BABC9F69" box="[1136,1158,295,320]" italics="true" pageId="11" pageNumber="252">P.</emphasis>
|
|
“
|
|
<taxonomicName id="4C1E4D6FA55D717C4A85FEC2BAD29F69" authorityName="Schaller" authorityYear="1785" box="[1175,1256,295,320]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="252" phylum="Arthropoda" rank="species" species="marcus">
|
|
<emphasis id="B96AEAFEA55D717C4A85FEC2BAD29F69" box="[1175,1256,295,320]" italics="true" pageId="11" pageNumber="252">marcus</emphasis>
|
|
</taxonomicName>
|
|
” is based on the fact that these two specific epithets have been associated with each other since
|
|
<bibRefCitation id="EF8F4B1DA55D717C4A0AFEAEBA8B9F4C" author="Evans, W. H." box="[1048,1201,330,357]" pageId="11" pageNumber="252" refId="ref10918" refString="Evans, W. H. (1955) A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. British Museum (Natural History), London, 499 pp." type="book" year="1955">Evans (1955)</bibRefCitation>
|
|
, and continued to be considered as conspecific in the recent comprehensive catalogue of
|
|
<taxonomicName id="4C1E4D6FA55D717C4D98FE8ABA2A9FA0" box="[906,1040,367,393]" class="Insecta" family="Hesperiidae" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="252" phylum="Arthropoda" rank="family">Hesperiidae</taxonomicName>
|
|
(
|
|
<bibRefCitation id="EF8F4B1DA55D717C4A0FFE8ABA8A9FA0" author="Mielke, O. H. H." box="[1053,1200,367,393]" pageId="11" pageNumber="252" pagination="3 - 11" refId="ref12282" refString="Mielke, O. H. H. (2004) Hesperiidae. In: Lamas, G. (Ed.), Checklist. Part 4 a. Hesperioidea - Papilionoidea. In: Heppner, J. B. (Ed.), Atlas of Neotropical Lepidoptera. Vol. 5 A. Association of Tropical Lepidoptera, Gainsville, pp. 3 - 11 + 25 - 86." type="book chapter" year="2004">Mielke 2004</bibRefCitation>
|
|
,
|
|
<bibRefCitation id="EF8F4B1DA55D717C4AA8FE8BBACD9FA0" author="Mielke, O. H. H." box="[1210,1271,366,393]" pageId="11" pageNumber="252" refId="ref12356" refString="Mielke, O. H. H. (2005) Catalogue of the American Hesperioidea: Hesperiidae (Lepidoptera). Vols. 1 - 6. Sociedade Brasileira de Zoologia, Curitiba, 1536 pp." type="book" year="2005">2005</bibRefCitation>
|
|
). Furthermore, the genomic sequencing data supports the
|
|
<typeStatus id="54A5884EA55D717C4C6EFE76BCD99F84" box="[636,739,403,429]" pageId="11" pageNumber="252" type="lectotype">lectotype</typeStatus>
|
|
of
|
|
<taxonomicName id="4C1E4D6FA55D717C4D1EFE76BDB49F84" authorityName="Rober" authorityYear="1925" box="[780,910,403,429]" class="Insecta" family="Hesperiidae" genus="Vettius" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="252" phylum="Arthropoda" rank="species" species="phyllides">
|
|
<emphasis id="B96AEAFEA55D717C4D1EFE76BDB49F84" box="[780,910,403,429]" italics="true" pageId="11" pageNumber="252">V. phyllides</emphasis>
|
|
</taxonomicName>
|
|
as conspecific with specimens that have been misidentified as
|
|
<taxonomicName id="4C1E4D6FA55D717C4F45FE52BCBC9FF8" authority="Fabricius, 1787" authorityName="Fabricius" authorityYear="1787" box="[343,646,439,465]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="252" phylum="Arthropoda" rank="species" species="marcus">
|
|
<emphasis id="B96AEAFEA55D717C4F45FE52BFFD9FF9" box="[343,455,439,464]" italics="true" pageId="11" pageNumber="252">P. marcus</emphasis>
|
|
<bibRefCitation id="EF8F4B1DA55D717C4FC0FE52BCBC9FF8" author="Fabricius, J. C." box="[466,646,439,465]" pageId="11" pageNumber="252" refId="ref10967" refString="Fabricius, J. C. (1787) Mantissa insectorvm sistens species nvper detectas adiectis synonymis, observationibvs, descriptionibvs, emendationibvs. Christ [ian] Gottl [ieb] Proft, Hafniae, [i] + 382 pp." type="book" year="1787">Fabricius, 1787</bibRefCitation>
|
|
</taxonomicName>
|
|
. On the other hand, as explained above,
|
|
<taxonomicName id="4C1E4D6FA55D717C4A74FE52BAFB9FF8" baseAuthorityName="Plotz" baseAuthorityYear="1882" box="[1126,1217,439,465]" class="Insecta" family="Hesperiidae" genus="Troyus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="252" phylum="Arthropoda" rank="species" species="aurelius">
|
|
<emphasis id="B96AEAFEA55D717C4A74FE52BAFB9FF8" box="[1126,1217,439,465]" italics="true" pageId="11" pageNumber="252">aurelius</emphasis>
|
|
</taxonomicName>
|
|
and other relevant species-group names currently associated with the genus
|
|
<taxonomicName id="4C1E4D6FA55D717C4D3BFE3EBD4E9FDD" authorityName="Warren & Turland" authorityYear="2012" box="[809,884,475,500]" class="Insecta" family="Hesperiidae" genus="Troyus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="252" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96AEAFEA55D717C4D3BFE3EBD4E9FDD" box="[809,884,475,500]" italics="true" pageId="11" pageNumber="252">Troyus</emphasis>
|
|
</taxonomicName>
|
|
are no longer associated with
|
|
<taxonomicName id="4C1E4D6FA55D717C4AC4FE3EBB129FDC" box="[1238,1320,475,501]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="252" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B96AEAFEA55D717C4AC4FE3EBB129FDC" box="[1238,1320,475,501]" italics="true" pageId="11" pageNumber="252">Papilio</emphasis>
|
|
</taxonomicName>
|
|
“
|
|
<taxonomicName id="4C1E4D6FA55D717C4B2CFE3EBBB59FDD" authorityName="Fabricius" authorityYear="1787" box="[1342,1423,475,500]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="252" phylum="Arthropoda" rank="species" species="marcus">
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<emphasis id="B96AEAFEA55D717C4B2CFE3EBBB59FDD" box="[1342,1423,475,500]" italics="true" pageId="11" pageNumber="252">marcus</emphasis>
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</taxonomicName>
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” and, importantly, type material is not available for these taxa to settle their nomenclature. In addition, genetic data exists to support the specific status of these names, which are currently in use (Zhang
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<emphasis id="B96AEAFEA55D717C4A7CFDC6BA9A9C14" box="[1134,1184,547,573]" italics="true" pageId="11" pageNumber="252">et al</emphasis>
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. 2019). We argue that the application of “in use” species-group names to different taxa would cause confusion, and we avoid this situation by proposing a replacement name based on a taxon which has been considered to be a junior subjective synonym. Name-bearing types exist for both
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<taxonomicName id="4C1E4D6FA55D717C4C32FD6ABDBB9C80" authority="Fabricius, 1787" authorityName="Fabricius" authorityYear="1787" box="[544,897,655,681]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="252" phylum="Arthropoda" rank="species" species="marcus">
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<emphasis id="B96AEAFEA55D717C4C32FD6ABCF19C81" box="[544,715,655,681]" italics="true" pageId="11" pageNumber="252">Papilio marcus</emphasis>
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<bibRefCitation id="EF8F4B1DA55D717C4CC1FD6ABDBB9C80" author="Fabricius, J. C." box="[723,897,655,681]" pageId="11" pageNumber="252" refId="ref10967" refString="Fabricius, J. C. (1787) Mantissa insectorvm sistens species nvper detectas adiectis synonymis, observationibvs, descriptionibvs, emendationibvs. Christ [ian] Gottl [ieb] Proft, Hafniae, [i] + 382 pp." type="book" year="1787">Fabricius, 1787</bibRefCitation>
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</taxonomicName>
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and
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<taxonomicName id="4C1E4D6FA55D717C4DABFD6ABB3F9C80" authority="Rober, 1925" authorityName="Rober" authorityYear="1925" box="[953,1285,654,681]" class="Insecta" family="Hesperiidae" genus="Vettius" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="252" phylum="Arthropoda" rank="species" species="phyllides">
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<emphasis id="B96AEAFEA55D717C4DABFD6ABA4A9C80" box="[953,1136,655,681]" italics="true" pageId="11" pageNumber="252">Vettius phyllides</emphasis>
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|
<bibRefCitation id="EF8F4B1DA55D717C4A6AFD6ABB3F9C80" author="Rober, J. K. M." box="[1144,1285,654,681]" pageId="11" pageNumber="252" pagination="85 - 100" refId="ref12884" refString="Rober, J. K. M. (1925) Neue su ¨ damerikanische Falter (Lep.). Entomologische Mitteilungen, 14 (1 & 2), 85 - 100 (15 January) & 156 - 162 (10 April)." type="journal article" year="1925">Röber, 1925</bibRefCitation>
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</taxonomicName>
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, allowing the removal of any ambiguity regarding the application of these names.
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</paragraph>
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<paragraph id="8BA136ECA55D717C4ED5FD32BAC79D3C" blockId="11.[151,1437,151,789]" pageId="11" pageNumber="252">
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(3) Proposal of the combination
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<taxonomicName id="4C1E4D6FA55D717C4C29FD32BCCE9CD8" box="[571,756,727,753]" class="Insecta" family="Hesperiidae" genus="Troyus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="252" phylum="Arthropoda" rank="species" species="phyllides">
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<emphasis id="B96AEAFEA55D717C4C29FD32BCCE9CD8" box="[571,756,727,753]" italics="true" pageId="11" pageNumber="252">Troyus phyllides</emphasis>
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</taxonomicName>
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|
(
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<emphasis id="B96AEAFEA55D717C4D14FD32BDEE9CD9" bold="true" box="[774,980,726,753]" pageId="11" pageNumber="252">new combination</emphasis>
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|
) as a name to refer to a skipper species known as
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<taxonomicName id="4C1E4D6FA55D717C4F1BFD1EBF919D3D" authorityName="Fabricius" authorityYear="1787" box="[265,427,763,788]" class="Insecta" family="Hesperiidae" genus="Vettius" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="252" phylum="Arthropoda" rank="species" species="marcus">
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<emphasis id="B96AEAFEA55D717C4F1BFD1EBF919D3D" box="[265,427,763,788]" italics="true" pageId="11" pageNumber="252">Vettius marcus</emphasis>
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</taxonomicName>
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|
for over 100 years until its recent taxonomic change in Zhang
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|
<emphasis id="B96AEAFEA55D717C4A7EFD1EBAA79D3C" box="[1132,1181,763,789]" italics="true" pageId="11" pageNumber="252">et al</emphasis>
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. (2019).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |