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<document id="332A8F13C44C46F08017588BF7657F8B" ID-CLB-Dataset="29925" ID-DOI="10.11646/zootaxa.4433.1.6" ID-GBIF-Dataset="1fdf27ec-2318-4726-ac75-a4c5e2121634" ID-ISSN="1175-5326" ID-Zenodo-Dep="1287135" ID-ZooBank="4D532EFF-7063-4772-AF77-58F5A8213B44" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1528732733516" checkinUser="plazi" docAuthor="Osawa, Masayuki &amp; Ng, Peter K. L." docDate="2018" docId="03B787EBFFDB051ED4F80AC1FC09F875" docLanguage="en" docName="zootaxa.4433.1.6.pdf" docOrigin="Zootaxa 4433 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Raphidopus johnsoni Ng &amp; Nakasone 1994" docType="treatment" docVersion="7" lastPageNumber="118" masterDocId="FF8EFF93FFDF0519D46F0A56FFEAFF98" masterDocTitle="A new species of the genus Raphidopus Stimpson, 1858 (Crustacea: Decapoda: Anomura: Porcellanidae) from Peninsular Malaysia, with additional records of R. johnsoni Ng &amp; Nakasone, 1994 from Southeast Asia, and a key to species in the genus" masterLastPageNumber="126" masterPageNumber="111" pageNumber="115" updateTime="1698548299394" updateUser="ExternalLinkService">
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<mods:title id="B8919CC77FB9CAB59DA890B674A642E6">A new species of the genus Raphidopus Stimpson, 1858 (Crustacea: Decapoda: Anomura: Porcellanidae) from Peninsular Malaysia, with additional records of R. johnsoni Ng &amp; Nakasone, 1994 from Southeast Asia, and a key to species in the genus</mods:title>
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<mods:namePart id="F847FA4732995C59B1E6D0F9E04FD6B7">Osawa, Masayuki</mods:namePart>
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<mods:namePart id="556961B11EA1775BD04DB7BD007E13D2">Ng, Peter K. L.</mods:namePart>
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<mods:title id="B67AAB3CE902CB2A8D20810B2DE7C72C">Zootaxa</mods:title>
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<mods:date id="F3877B3A6EF6E2F5E31DBF41E5E0A18E">2018</mods:date>
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<mods:number id="970B0C0E52F1C8A4D9C0A7CE840A1EF1">2018-06-08</mods:number>
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<treatment id="03B787EBFFDB051ED4F80AC1FC09F875" ID-DOI="http://doi.org/10.5281/zenodo.5967589" ID-GBIF-Taxon="145939560" ID-Zenodo-Dep="5967589" LSID="urn:lsid:plazi:treatment:03B787EBFFDB051ED4F80AC1FC09F875" httpUri="http://treatment.plazi.org/id/03B787EBFFDB051ED4F80AC1FC09F875" lastPageId="7" lastPageNumber="118" pageId="4" pageNumber="115">
<subSubSection id="C3046576FFDB051DD4F80AC1FE8BFF4B" pageId="4" pageNumber="115" type="nomenclature">
<paragraph id="8BA136FDFFDB051DD4F80AC1FD45FF2A" blockId="4.[151,687,151,211]" box="[151,687,151,178]" pageId="4" pageNumber="115">
<heading id="D0E98191FFDB051DD4F80AC1FD45FF2A" bold="true" box="[151,687,151,178]" fontSize="11" level="1" pageId="4" pageNumber="115" reason="1">
<taxonomicName id="4C1E4D7EFFDB051DD4F80AC1FD45FF2A" ID-CoL="8fb0d90a-5c10-4380-8b83-481795ba2f33" authority="Ng &amp; Nakasone, 1994" authorityName="Ng &amp; Nakasone" authorityYear="1994" box="[151,687,151,178]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="115" phylum="Arthropoda" rank="species" species="johnsoni">
<emphasis id="B96AEAEFFFDB051DD4F80AC1FD45FF2A" bold="true" box="[151,687,151,178]" pageId="4" pageNumber="115">
<emphasis id="B96AEAEFFFDB051DD4F80AC1FE73FF29" bold="true" box="[151,409,151,177]" italics="true" pageId="4" pageNumber="115">Raphidopus johnsoni</emphasis>
Ng &amp;
<bibRefCitation id="EF8F4B0CFFDB051DD5840ACEFD45FF2A" box="[491,687,152,178]" pageId="4" pageNumber="115" refId="ref6869" refString="Ng, P. K. L. &amp; Nakasone, Y. (1994) On the porcellanid genera Raphidopus Stimpson, 1858, and Pseudoporcellanella SankarankUtty, 1961, With description of a neW mangrove species, Raphidopus Johnsoni from Singapore (Decapoda, AnomUra). Crustaceana, 66, 1 - 21. https: // doi. org / 10.1163 / 156854094 X 00116" type="journal article">Nakasone, 1994</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BA136FDFFDB051DD4F80AECFE8BFF4B" blockId="4.[151,687,151,211]" box="[151,353,186,211]" pageId="4" pageNumber="115">
(
<figureCitation id="13252A78FFDB051DD4F00AECFEE1FF4B" box="[159,267,186,211]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStartId-0="5.[151,250,1751,1773]" captionStartId-1="6.[151,250,1765,1787]" captionStartId-2="7.[151,250,1377,1399]" captionTargetBox-0="[218,1362,206,1697]" captionTargetBox-1="[263,1328,214,1697]" captionTargetBox-2="[173,1420,218,1300]" captionTargetId-0="figure@5.[205,1375,193,1724]" captionTargetId-1="figure@6.[251,1343,193,1738]" captionTargetId-2="figure@7.[150,1436,193,1356]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="FIGURE 3. Raphidopus Johnsoni Ng &amp; Nakasone, 1994, holotype, male (cl 3.8 mm), ZRC 1992.10669, Singapore. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (transverse striae and ridges not draWn); B, rostrUm, dorso-anterior vieW; C, third and foUrth thoracic sternites, ventral vieW; D, basal article of left antennUlar pedUncle, ventral vieW; E, left third maxilliped, lateral vieW (sUrface strUctUre not draWn); F, larger cheliped, merUs, ventral vieW (sUrface strUctUre not draWn); G, larger cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); H, larger cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); I, smaller cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); J, smaller cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); K, left foUrth pereopod, lateral vieW (short striae not draWn); L, same, merUs, lateral perpendicUlar vieW (short striae not draWn). Setae are not draWn on all figUres. Scale bars = 1.0 mm." captionText-1="FIGURE 4. Raphidopus Johnsoni Ng &amp; Nakasone, 1994, male (cl 5.0 mm), CBM-ZC 12485, PhUket. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (marginal setae not draWn on right side); B, right branchial margin of carapace, dorsomesial vieW; C, rostrUm, dorso-anterior vieW; D, third and foUrth thoracic sternites, ventral vieW (setae not draWn on left side); E, telson, external vieW; F, basal article of left antennUlar pedUncle, ventral vieW; G, left ocUlar and antennal pedUncles, dorsolateral vieW; H, left third maxilliped, lateral vieW; I, left second pereopod, lateral vieW; J, left second pereopod, merUs, mesial vieW (short striae not draWn); K, left second pereopod, dactylUs, lateral vieW (setae not draWn); L, left third pereopod, lateral vieW (setae and short striae not draWn); M, left third pereopod, merUs, mesial vieW (short striae not draWn); N, left foUrth pereopod, lateral vieW (setae and short striae not draWn); O, left foUrth pereopod, merUs, mesial vieW (short striae not draWn). Scale bars = 1.0 mm." captionText-2="FIGURE 5. Raphidopus Johnsoni Ng &amp; Nakasone, 1994, male (cl 4.7 mm), CBM-ZC 12485, PhUket. A, larger cheliped, dorsal vieW; B, larger cheliped, merUs and carpUs, ventral vieW; C, larger cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre on palm and dactylUs not draWn); D, smaller cheliped, dorsal vieW (sUrface strUctUre not draWn); E, smaller cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn). Setae are not draWn on all figUres. Scale bars = 1.0 mm." httpUri-0="https://zenodo.org/record/1287141/files/figure.png" httpUri-1="https://zenodo.org/record/1287143/files/figure.png" httpUri-2="https://zenodo.org/record/1287145/files/figure.png" pageId="4" pageNumber="115">Figs. 3–5</figureCitation>
,
<figureCitation id="13252A78FFDB051DD5790AEDFEB3FF4B" box="[278,345,187,211]" captionStart="FIGURE 9" captionStartId="12.[151,250,1662,1684]" captionTargetBox="[151,1436,193,1641]" captionTargetId="figure@12.[151,1436,193,1641]" captionTargetPageId="12" captionText="FIGURE 9. Preserved specimens of three Raphidopus species. Entire animal, dorsal vieW (A–D); carapace, dorsal vieW (E); larger and smaller chelipeds, dorsal vieW (F). A, Raphidopus indicus Henderson, 1893, holotype, male (cl 6.5 mm), NHM 1892.7.15.246; B, Raphidopus Johnsoni Ng &amp; Nakasone, 1994, holotype, male (cl 3.8 mm), ZRC 1992.10669; C, Raphidopus Johnsoni Ng &amp; Nakasone, 1994, female (cl 5.9 mm), CBM-ZC 12485; D, Raphidopus brevipes n. sp., female (cl 5.1 mm), ZRC 2016.528; E, Raphidopus brevipes n. sp., male, holotype, male (cl 5.8 mm), ZRC 1992.10521; F, Raphidopus brevipes n. sp., paratypes, ZRC 1992.10522–10526." httpUri="https://zenodo.org/record/1287153/files/figure.png" pageId="4" pageNumber="115">9B, C</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3046576FFDB051DD4F80B54FE43FECE" pageId="4" pageNumber="115" type="reference_group">
<paragraph id="8BA136FDFFDB051DD4F80B54FC92FE80" blockId="4.[151,1436,258,342]" box="[151,888,258,280]" pageId="4" pageNumber="115">
<treatmentCitationGroup id="AB0E11D3FFDB051DD4F80B54FC92FE80" box="[151,888,258,280]" pageId="4" pageNumber="115">
<emphasis id="B96AEAEFFFDB051DD4F80B54FE96FE80" box="[151,380,258,280]" italics="true" pageId="4" pageNumber="115">
<taxonomicName id="4C1E4D7EFFDB051DD4F80B54FEB4FE80" ID-CoL="445049fb-994e-4886-ba7b-e2521935c222" authorityName="Henderson" authorityYear="1893" box="[151,350,258,280]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="115" phylum="Arthropoda" rank="species" species="indicus">Raphidopus indicus</taxonomicName>
.—
</emphasis>
<treatmentCitation id="0ABF10ECFFDB051DD5130B54FDF0FE80" author="" box="[380,538,258,280]" page="275" pageId="4" pageNumber="115">
<bibRefCitation id="EF8F4B0CFFDB051DD5130B54FE03FE80" box="[380,489,258,280]" pageId="4" pageNumber="115" refId="ref6637" refString="Haig, J. (1981) Porcellanid crabs from the Indo-West Pacific, Part II. Steenstrupia, 7 (12), 269 - 291." type="journal article">Haig 1981</bibRefCitation>
: 275
</treatmentCitation>
(not
<taxonomicName id="4C1E4D7EFFDB051DD6200B55FC87FE80" ID-CoL="445049fb-994e-4886-ba7b-e2521935c222" authority="Henderson, 1893" authorityName="Henderson" authorityYear="1893" box="[591,877,258,280]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="125" phylum="Arthropoda" rank="species" species="indicus">
R. indicUs
<bibRefCitation id="EF8F4B0CFFDB051DD6D30B54FC87FE80" box="[700,877,258,280]" pageId="4" pageNumber="115" refId="ref6699" refString="Henderson, J. R. (1893) A contribUtion to Indian carcinology. Transaction of the Linnean Society, London, Zoology, Series 2, 5, 325 - 458. https: // doi. org / 10.1111 / j. 1096 - 3642.1893. tb 00653. x" type="journal article">Henderson, 1893</bibRefCitation>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA136FDFFDB051DD4F80B77FE43FECE" blockId="4.[151,1436,258,342]" pageId="4" pageNumber="115">
<treatmentCitationGroup id="AB0E11D3FFDB051DD4F80B77FE43FECE" pageId="4" pageNumber="115">
<taxonomicName id="4C1E4D7EFFDB051DD4F80B77FD7DFEAF" ID-CoL="1a4de4bf-a9a2-43d3-9bea-a5b9d186ce4e" authority="Johnsoni Ng &amp; Nakasone, 1994: 9" authorityName="Johnsoni Ng &amp; Nakasone" authorityPageNumber="9" authorityYear="1994" box="[151,663,289,311]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAEFFFDB051DD4F80B77FE99FEAF" box="[151,371,289,311]" italics="true" pageId="4" pageNumber="115">Raphidopus Johnsoni</emphasis>
Ng &amp;
<treatmentCitation id="0ABF10ECFFDB051DD5A30B77FD7DFEAF" author="" box="[460,663,289,311]" page="9" pageId="4" pageNumber="115">
<bibRefCitation id="EF8F4B0CFFDB051DD5A30B77FD92FEAF" box="[460,632,289,311]" pageId="4" pageNumber="115" refId="ref6869" refString="Ng, P. K. L. &amp; Nakasone, Y. (1994) On the porcellanid genera Raphidopus Stimpson, 1858, and Pseudoporcellanella SankarankUtty, 1961, With description of a neW mangrove species, Raphidopus Johnsoni from Singapore (Decapoda, AnomUra). Crustaceana, 66, 1 - 21. https: // doi. org / 10.1163 / 156854094 X 00116" type="journal article">Nakasone, 1994</bibRefCitation>
: 9
</treatmentCitation>
</taxonomicName>
, fig. 4.—
<treatmentCitation id="0ABF10ECFFDB051DD6900B77FB80FEAF" author="" box="[767,1130,289,311]" page="115" pageId="4" pageNumber="115">
<bibRefCitation id="EF8F4B0CFFDB051DD6900B77FBDCFEAF" box="[767,1078,289,311]" pageId="4" pageNumber="115" refId="ref7058" refString="OsaWa, M. &amp; McLaUghlin, P. A. (2010) Annotated checklist of anomUran decapod crUstaceans of the World (exclUsive of the KiWaoidea and families Chirostylidae and Galatheidae of the Galatheoidea) Part II-Porcellanidae. In: LoW, M. E. Y. &amp; Tan, S. H. (Eds.), Checklists of anomuran decapod crustaceans of the world (exclusive of the Kiwaioidea and Families Chirostylidae and Galatheidae of the Galathoidea) and marine lobsters of the world. The Raffles Bulletin of Zoology, Supplement, 23, 109 - 129." type="journal article">OsaWa &amp; McLaUghlin 2010</bibRefCitation>
: 115
</treatmentCitation>
(list).—Ng
<emphasis id="B96AEAEFFFDB051DD09D0B74FAC1FEAF" box="[1266,1323,289,311]" italics="true" pageId="4" pageNumber="115">et al.</emphasis>
2011: 27, UnnUmbered color fig.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3046576FFDB051DD4F80BD4FAF9FD0F" pageId="4" pageNumber="115" type="materials_examined">
<paragraph id="8BA136FDFFDB051DD4F80BD4FDA5FE27" blockId="4.[151,1437,386,1851]" pageId="4" pageNumber="115">
<materialsCitation id="3B763CA0FFDB051DD4F80BD4FDA5FE27" ID-GBIF-Occurrence="1848744201" collectingDate="1989-04-04" collectionCode="ZRC" collectorName="P. K. L. Ng." country="Singapore" location="Lim Chu Kang" pageId="4" pageNumber="115" specimenCode="ZRC 1992.10669" specimenCount="2" typeStatus="holotype">
<emphasis id="B96AEAEFFFDB051DD4F80BD4FEAFFE03" bold="true" box="[151,325,386,411]" pageId="4" pageNumber="115">
<typeStatus id="54A5885FFFDB051DD4F80BD4FF38FE03" box="[151,210,386,411]" pageId="4" pageNumber="115">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54A5885FFFDB051DD5230BD4FE53FE03" box="[332,441,386,411]" pageId="4" pageNumber="115" type="holotype">Holotype</typeStatus>
:
<specimenCode id="DBB89E86FFDB051DD5AB0BD5FD60FE03" box="[452,650,386,411]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="115">ZRC 1992.10669</specimenCode>
, male (cl
<quantity id="4CE69B18FFDB051DD6960BD5FCB9FE03" box="[761,851,387,411]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="4" pageNumber="115" unit="mm" value="3.8">3.8 mm</quantity>
),
<location id="8EC16026FFDB051DD7080BD4FBF8FE03" LSID="urn:lsid:plazi:treatment:03B787EBFFDB051ED4F80AC1FC09F875:8EC16026FFDB051DD7080BD4FBF8FE03" box="[871,1042,386,411]" country="Singapore" name="Lim Chu Kang" pageId="4" pageNumber="115">Lim Chu Kang</location>
,
<collectingCountry id="F309766DFFDB051DD0700BD4FB79FE03" box="[1055,1171,386,411]" name="Singapore" pageId="4" pageNumber="115">Singapore</collectingCountry>
, road end, in mangroVe swamp,
<collectingDate id="EFE4E9D5FFDB051DD49B0BF1FE6CFE27" box="[244,390,422,447]" pageId="4" pageNumber="115" value="1989-04-04">
<date id="FFA0103DFFDB051DD49B0BF1FE6CFE27" box="[244,390,422,447]" pageId="4" pageNumber="115" value="1989-04-04">4 April 1989</date>
</collectingDate>
, coll.
<collectorName id="26EB532BFFDB051DD5A60BF1FDA5FE27" box="[457,591,423,447]" pageId="4" pageNumber="115">P. K. L. Ng.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BA136FDFFDB051DD4A80B9CFAF9FD0F" blockId="4.[151,1437,386,1851]" pageId="4" pageNumber="115">
<materialsCitation id="3B763CA0FFDB051DD4A80B9CFB66FDB3" ID-GBIF-Occurrence="1848744179" collectingDate="2009-12-22" collectorName="T. Komai" country="Thailand" location="Koh Sirae" municipality="South China Sea" pageId="4" pageNumber="115" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Phuket">
<emphasis id="B96AEAEFFFDB051DD4A80B9CFE16FE7B" bold="true" box="[199,508,458,483]" pageId="4" pageNumber="115">Other material examined.</emphasis>
“Indo-Chinese Sea” (= probably
<collectingMunicipality id="6BC5AC87FFDB051DD71A0B9CFBDFFE7B" box="[885,1077,458,483]" pageId="4" pageNumber="115">South China Sea</collectingMunicipality>
): ZMUC-CRU-20517,
<specimenCount id="9D18FD74FFDB051DD1550B9DFA76FE7B" box="[1338,1436,458,483]" pageId="4" pageNumber="115" type="female">1 female</specimenCount>
(cl
<quantity id="4CE69B18FFDB051DD4D30BB9FEFCFD9F" box="[188,278,495,519]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="4" pageNumber="115" unit="mm" value="3.2">3.2 mm</quantity>
), coll. Schmidt.
<collectingCountry id="F309766DFFDB051DD5A20BB8FDDFFD9F" box="[461,565,494,519]" name="Thailand" pageId="4" pageNumber="115">Thailand</collectingCountry>
: CBM-ZC 12485,
<specimenCount id="9D18FD74FFDB051DD7680BB9FCB7FD9F" box="[775,861,494,519]" pageId="4" pageNumber="115" type="male">2 males</specimenCount>
(cl 4.7, 5.0 mm),
<specimenCount id="9D18FD74FFDB051DD0460BB9FB60FD9F" box="[1065,1162,494,519]" pageId="4" pageNumber="115" type="female">1 female</specimenCount>
(cl
<quantity id="4CE69B18FFDB051DD0D80BB8FAFBFD9F" box="[1207,1297,494,519]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="4" pageNumber="115" unit="mm" value="5.9">5.9 mm</quantity>
),
<location id="8EC16026FFDB051DD14B0BB8FA72FD9F" LSID="urn:lsid:plazi:treatment:03B787EBFFDB051ED4F80AC1FC09F875:8EC16026FFDB051DD14B0BB8FA72FD9F" box="[1316,1432,494,519]" country="Thailand" municipality="South China Sea" name="Koh Sirae" pageId="4" pageNumber="115" stateProvince="Phuket">Koh Sirae</location>
,
<collectingRegion id="49DAF81FFFDB051DD4F80844FF02FDB3" box="[151,232,530,555]" country="Thailand" name="Phuket" pageId="4" pageNumber="115">Phuket</collectingRegion>
, intertidal, sand flat, in burrows,
<collectingDate id="EFE4E9D5FFDB051DD60D0845FCD6FDB3" box="[610,828,530,555]" pageId="4" pageNumber="115" value="2009-12-22">
<date id="FFA0103DFFDB051DD60D0845FCD6FDB3" box="[610,828,530,555]" pageId="4" pageNumber="115" value="2009-12-22">22 December 2009</date>
</collectingDate>
, yabby pump, coll.
<collectorName id="26EB532BFFDB051DD0710845FB66FDB3" box="[1054,1164,530,555]" pageId="4" pageNumber="115">T. Komai</collectorName>
</materialsCitation>
;
<materialsCitation id="3B763CA0FFDB051DD0F70845FC16FDD7" ID-GBIF-Occurrence="1848744180" collectingDate="2009-12-22" collectionCode="ZRC" collectorName="T. Komai" country="Singapore" location="Koh Sirae" municipality="South China Sea" pageId="4" pageNumber="115" specimenCode="ZRC 2018.256" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Phuket">
<specimenCode id="DBB89E86FFDB051DD0F70845FAA9FDB3" box="[1176,1347,530,555]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="115">ZRC 2018.256</specimenCode>
,
<specimenCount id="9D18FD74FFDB051DD1200845FA76FDB3" box="[1359,1436,530,555]" pageId="4" pageNumber="115" type="male">1 male</specimenCount>
(cl
<quantity id="4CE69B18FFDB051DD4D30861FEFDFDD7" box="[188,279,567,591]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="4" pageNumber="115" unit="mm" value="3.7">3.7 mm</quantity>
),
<specimenCount id="9D18FD74FFDB051DD5450861FE66FDD7" box="[298,396,566,591]" pageId="4" pageNumber="115" type="female">1 female</specimenCount>
(cl
<quantity id="4CE69B18FFDB051DD5D60861FDFEFDD7" box="[441,532,567,591]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="4" pageNumber="115" unit="mm" value="4.2">4.2 mm</quantity>
), same data as CBM-ZC 12485.
<collectingCountry id="F309766DFFDB051DD7E90860FC16FDD7" box="[902,1020,566,591]" name="Singapore" pageId="4" pageNumber="115">Singapore</collectingCountry>
</materialsCitation>
:
<materialsCitation id="3B763CA0FFDB051DD0680861FAF9FD0F" ID-GBIF-Occurrence="1848744176" collectingDate="2014-04-09" collectionCode="CMBS" collectorName="Surabaya &amp; Andrea." country="Indonesia" county="Seabed" elevation="12" location="Tengah ReserVoir" municipality="West Johor Strait" pageId="4" pageNumber="115" specimenCode="ZRC 2018.247" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="JaVa">
<specimenCode id="DBB89E86FFDB051DD0680861FB5BFDD7" box="[1031,1201,567,591]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="115">ZRC 2018.247</specimenCode>
,
<specimenCount id="9D18FD74FFDB051DD0D30861FAE2FDD7" box="[1212,1288,566,591]" pageId="4" pageNumber="115" type="male">1 male</specimenCount>
(cl
<quantity id="4CE69B18FFDB051DD1590861FA7AFDD7" box="[1334,1424,567,591]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="4" pageNumber="115" unit="mm" value="4.2">4.2 mm</quantity>
), sta. 3821 [SEA-7603 (
<collectionCode id="ED0FAE38FFDB051DD5FF080DFE0AFDEB" box="[400,480,603,627]" pageId="4" pageNumber="115">CMBS</collectionCode>
<collectingCounty id="62C04E71FFDB051DD589080CFDD4FDEB" box="[486,574,602,627]" pageId="4" pageNumber="115">Seabed</collectingCounty>
)],
<collectingMunicipality id="6BC5AC87FFDB051DD638080DFCFCFDEB" box="[599,790,602,627]" pageId="4" pageNumber="115">West Johor Strait</collectingMunicipality>
off
<location id="8EC16026FFDB051DD72C080CFBE1FDEB" LSID="urn:lsid:plazi:treatment:03B787EBFFDB051ED4F80AC1FC09F875:8EC16026FFDB051DD72C080CFBE1FDEB" box="[835,1035,602,627]" country="Indonesia" county="Seabed" municipality="West Johor Strait" name="Tengah ReserVoir" pageId="4" pageNumber="115" stateProvince="JaVa">Tengah ReserVoir</location>
, 11.0–
<elevation id="0033D1CEFFDB051DD03B080CFB48FDEA" box="[1108,1186,602,627]" metricMagnitude="1" metricUnit="m" metricValue="1.15" pageId="4" pageNumber="115" unit="m" value="11.5">
<quantity id="4CE69B18FFDB051DD03B080CFB48FDEA" box="[1108,1186,602,627]" metricMagnitude="1" metricUnit="m" metricValue="1.15" pageId="4" pageNumber="115" unit="m" value="11.5">11.5 m</quantity>
</elevation>
, dredge,
<collectingDate id="EFE4E9D5FFDB051DD168080DFA73FDEB" box="[1287,1433,602,627]" pageId="4" pageNumber="115" value="2014-04-09">
<date id="FFA0103DFFDB051DD168080DFA73FDEB" box="[1287,1433,602,627]" pageId="4" pageNumber="115" value="2014-04-09">9 April 2014</date>
</collectingDate>
.
<collectingCountry id="F309766DFFDB051DD4F80828FEE3FD0F" box="[151,265,638,663]" name="Indonesia" pageId="4" pageNumber="115">Indonesia</collectingCountry>
: ZMUC-CRU-20518,
<specimenCount id="9D18FD74FFDB051DD66B0829FD9AFD0F" box="[516,624,638,663]" pageId="4" pageNumber="115" type="female">3 females</specimenCount>
(cl 3.4–4.0 mm),
<collectorName id="26EB532BFFDB051DD7560828FC4EFD0F" box="[825,932,638,663]" pageId="4" pageNumber="115">Surabaya</collectorName>
,
<collectingRegion id="49DAF81FFFDB051DD7C00829FC0BFD0E" box="[943,993,639,662]" country="Indonesia" name="Jawa Timur" pageId="4" pageNumber="115">JaVa</collectingRegion>
Island, 1870, coll.
<collectorName id="26EB532BFFDB051DD0D60828FAF9FD0F" box="[1209,1299,638,663]" pageId="4" pageNumber="115">Andrea.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3046576FFDB051DD4A808F4FB10F957" pageId="4" pageNumber="115" type="description">
<paragraph id="8BA136FDFFDB051DD4A808F4FE78FCD3" blockId="4.[151,1437,386,1851]" pageId="4" pageNumber="115">
<emphasis id="B96AEAEFFFDB051DD4A808F4FEB3FD23" bold="true" box="[199,345,674,699]" pageId="4" pageNumber="115">Description.</emphasis>
Carapace (
<figureCitation id="13252A78FFDB051DD5B608F4FDD7FD23" box="[473,573,674,699]" captionStart="FIGURE 3" captionStartId="5.[151,250,1751,1773]" captionTargetBox="[218,1362,206,1697]" captionTargetId="figure@5.[205,1375,193,1724]" captionTargetPageId="5" captionText="FIGURE 3. Raphidopus Johnsoni Ng &amp; Nakasone, 1994, holotype, male (cl 3.8 mm), ZRC 1992.10669, Singapore. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (transverse striae and ridges not draWn); B, rostrUm, dorso-anterior vieW; C, third and foUrth thoracic sternites, ventral vieW; D, basal article of left antennUlar pedUncle, ventral vieW; E, left third maxilliped, lateral vieW (sUrface strUctUre not draWn); F, larger cheliped, merUs, ventral vieW (sUrface strUctUre not draWn); G, larger cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); H, larger cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); I, smaller cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); J, smaller cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); K, left foUrth pereopod, lateral vieW (short striae not draWn); L, same, merUs, lateral perpendicUlar vieW (short striae not draWn). Setae are not draWn on all figUres. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287141/files/figure.png" pageId="4" pageNumber="115">Figs. 3A</figureCitation>
,
<figureCitation id="13252A78FFDB051DD62408F5FD84FD23" box="[587,622,675,699]" captionStart="FIGURE 4" captionStartId="6.[151,250,1765,1787]" captionTargetBox="[263,1328,214,1697]" captionTargetId="figure@6.[251,1343,193,1738]" captionTargetPageId="6" captionText="FIGURE 4. Raphidopus Johnsoni Ng &amp; Nakasone, 1994, male (cl 5.0 mm), CBM-ZC 12485, PhUket. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (marginal setae not draWn on right side); B, right branchial margin of carapace, dorsomesial vieW; C, rostrUm, dorso-anterior vieW; D, third and foUrth thoracic sternites, ventral vieW (setae not draWn on left side); E, telson, external vieW; F, basal article of left antennUlar pedUncle, ventral vieW; G, left ocUlar and antennal pedUncles, dorsolateral vieW; H, left third maxilliped, lateral vieW; I, left second pereopod, lateral vieW; J, left second pereopod, merUs, mesial vieW (short striae not draWn); K, left second pereopod, dactylUs, lateral vieW (setae not draWn); L, left third pereopod, lateral vieW (setae and short striae not draWn); M, left third pereopod, merUs, mesial vieW (short striae not draWn); N, left foUrth pereopod, lateral vieW (setae and short striae not draWn); O, left foUrth pereopod, merUs, mesial vieW (short striae not draWn). Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287143/files/figure.png" pageId="4" pageNumber="115">4A</figureCitation>
) 1.4 times as broad as long; dorsal surface with scattered, short or Very short striae; anterior branchial region with small tubercles and short rows of beaded tubercles (
<figureCitation id="13252A78FFDB051DD0A60890FAC9FD47" box="[1225,1315,710,735]" captionStart="FIGURE 4" captionStartId="6.[151,250,1765,1787]" captionTargetBox="[263,1328,214,1697]" captionTargetId="figure@6.[251,1343,193,1738]" captionTargetPageId="6" captionText="FIGURE 4. Raphidopus Johnsoni Ng &amp; Nakasone, 1994, male (cl 5.0 mm), CBM-ZC 12485, PhUket. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (marginal setae not draWn on right side); B, right branchial margin of carapace, dorsomesial vieW; C, rostrUm, dorso-anterior vieW; D, third and foUrth thoracic sternites, ventral vieW (setae not draWn on left side); E, telson, external vieW; F, basal article of left antennUlar pedUncle, ventral vieW; G, left ocUlar and antennal pedUncles, dorsolateral vieW; H, left third maxilliped, lateral vieW; I, left second pereopod, lateral vieW; J, left second pereopod, merUs, mesial vieW (short striae not draWn); K, left second pereopod, dactylUs, lateral vieW (setae not draWn); L, left third pereopod, lateral vieW (setae and short striae not draWn); M, left third pereopod, merUs, mesial vieW (short striae not draWn); N, left foUrth pereopod, lateral vieW (setae and short striae not draWn); O, left foUrth pereopod, merUs, mesial vieW (short striae not draWn). Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287143/files/figure.png" pageId="4" pageNumber="115">Fig. 4B</figureCitation>
); posterior branchial and intestinal regions with long, tuberculate ridges; branchial margins narrowly laminar, each with row of small, rounded and subacute tubercles. Rostrum (
<figureCitation id="13252A78FFDB051DD6B30958FCD4FCBF" box="[732,830,782,807]" captionStart="FIGURE 3" captionStartId="5.[151,250,1751,1773]" captionTargetBox="[218,1362,206,1697]" captionTargetId="figure@5.[205,1375,193,1724]" captionTargetPageId="5" captionText="FIGURE 3. Raphidopus Johnsoni Ng &amp; Nakasone, 1994, holotype, male (cl 3.8 mm), ZRC 1992.10669, Singapore. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (transverse striae and ridges not draWn); B, rostrUm, dorso-anterior vieW; C, third and foUrth thoracic sternites, ventral vieW; D, basal article of left antennUlar pedUncle, ventral vieW; E, left third maxilliped, lateral vieW (sUrface strUctUre not draWn); F, larger cheliped, merUs, ventral vieW (sUrface strUctUre not draWn); G, larger cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); H, larger cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); I, smaller cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); J, smaller cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); K, left foUrth pereopod, lateral vieW (short striae not draWn); L, same, merUs, lateral perpendicUlar vieW (short striae not draWn). Setae are not draWn on all figUres. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287141/files/figure.png" pageId="4" pageNumber="115">Figs. 3B</figureCitation>
,
<figureCitation id="13252A78FFDB051DD7260959FC81FCBF" box="[841,875,783,807]" captionStart="FIGURE 4" captionStartId="6.[151,250,1765,1787]" captionTargetBox="[263,1328,214,1697]" captionTargetId="figure@6.[251,1343,193,1738]" captionTargetPageId="6" captionText="FIGURE 4. Raphidopus Johnsoni Ng &amp; Nakasone, 1994, male (cl 5.0 mm), CBM-ZC 12485, PhUket. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (marginal setae not draWn on right side); B, right branchial margin of carapace, dorsomesial vieW; C, rostrUm, dorso-anterior vieW; D, third and foUrth thoracic sternites, ventral vieW (setae not draWn on left side); E, telson, external vieW; F, basal article of left antennUlar pedUncle, ventral vieW; G, left ocUlar and antennal pedUncles, dorsolateral vieW; H, left third maxilliped, lateral vieW; I, left second pereopod, lateral vieW; J, left second pereopod, merUs, mesial vieW (short striae not draWn); K, left second pereopod, dactylUs, lateral vieW (setae not draWn); L, left third pereopod, lateral vieW (setae and short striae not draWn); M, left third pereopod, merUs, mesial vieW (short striae not draWn); N, left foUrth pereopod, lateral vieW (setae and short striae not draWn); O, left foUrth pereopod, merUs, mesial vieW (short striae not draWn). Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287143/files/figure.png" pageId="4" pageNumber="115">4C</figureCitation>
) trilobate, median lobe slightly exceeding leVel of apices of lateral lobes.
</paragraph>
<paragraph id="8BA136FDFFDB051DD4A80900FB05FCF7" blockId="4.[151,1437,386,1851]" box="[199,1263,854,879]" pageId="4" pageNumber="115">
Third thoracic sternite (
<figureCitation id="13252A78FFDB051DD5A00900FDDBFCF7" box="[463,561,854,879]" captionStart="FIGURE 3" captionStartId="5.[151,250,1751,1773]" captionTargetBox="[218,1362,206,1697]" captionTargetId="figure@5.[205,1375,193,1724]" captionTargetPageId="5" captionText="FIGURE 3. Raphidopus Johnsoni Ng &amp; Nakasone, 1994, holotype, male (cl 3.8 mm), ZRC 1992.10669, Singapore. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (transverse striae and ridges not draWn); B, rostrUm, dorso-anterior vieW; C, third and foUrth thoracic sternites, ventral vieW; D, basal article of left antennUlar pedUncle, ventral vieW; E, left third maxilliped, lateral vieW (sUrface strUctUre not draWn); F, larger cheliped, merUs, ventral vieW (sUrface strUctUre not draWn); G, larger cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); H, larger cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); I, smaller cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); J, smaller cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); K, left foUrth pereopod, lateral vieW (short striae not draWn); L, same, merUs, lateral perpendicUlar vieW (short striae not draWn). Setae are not draWn on all figUres. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287141/files/figure.png" pageId="4" pageNumber="115">Figs. 3C</figureCitation>
,
<figureCitation id="13252A78FFDB051DD6520901FD8AFCF7" box="[573,608,855,879]" captionStart="FIGURE 4" captionStartId="6.[151,250,1765,1787]" captionTargetBox="[263,1328,214,1697]" captionTargetId="figure@6.[251,1343,193,1738]" captionTargetPageId="6" captionText="FIGURE 4. Raphidopus Johnsoni Ng &amp; Nakasone, 1994, male (cl 5.0 mm), CBM-ZC 12485, PhUket. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (marginal setae not draWn on right side); B, right branchial margin of carapace, dorsomesial vieW; C, rostrUm, dorso-anterior vieW; D, third and foUrth thoracic sternites, ventral vieW (setae not draWn on left side); E, telson, external vieW; F, basal article of left antennUlar pedUncle, ventral vieW; G, left ocUlar and antennal pedUncles, dorsolateral vieW; H, left third maxilliped, lateral vieW; I, left second pereopod, lateral vieW; J, left second pereopod, merUs, mesial vieW (short striae not draWn); K, left second pereopod, dactylUs, lateral vieW (setae not draWn); L, left third pereopod, lateral vieW (setae and short striae not draWn); M, left third pereopod, merUs, mesial vieW (short striae not draWn); N, left foUrth pereopod, lateral vieW (setae and short striae not draWn); O, left foUrth pereopod, merUs, mesial vieW (short striae not draWn). Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287143/files/figure.png" pageId="4" pageNumber="115">4D</figureCitation>
) 5.3–7.0 times broader than long measured along midline.
</paragraph>
<paragraph id="8BA136FDFFDB051DD4A8092CFB45FC0B" blockId="4.[151,1437,386,1851]" box="[199,1199,890,915]" pageId="4" pageNumber="115">
Telson (
<figureCitation id="13252A78FFDB051DD54F092CFE9FFC0B" box="[288,373,890,915]" captionStart="FIGURE 4" captionStartId="6.[151,250,1765,1787]" captionTargetBox="[263,1328,214,1697]" captionTargetId="figure@6.[251,1343,193,1738]" captionTargetPageId="6" captionText="FIGURE 4. Raphidopus Johnsoni Ng &amp; Nakasone, 1994, male (cl 5.0 mm), CBM-ZC 12485, PhUket. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (marginal setae not draWn on right side); B, right branchial margin of carapace, dorsomesial vieW; C, rostrUm, dorso-anterior vieW; D, third and foUrth thoracic sternites, ventral vieW (setae not draWn on left side); E, telson, external vieW; F, basal article of left antennUlar pedUncle, ventral vieW; G, left ocUlar and antennal pedUncles, dorsolateral vieW; H, left third maxilliped, lateral vieW; I, left second pereopod, lateral vieW; J, left second pereopod, merUs, mesial vieW (short striae not draWn); K, left second pereopod, dactylUs, lateral vieW (setae not draWn); L, left third pereopod, lateral vieW (setae and short striae not draWn); M, left third pereopod, merUs, mesial vieW (short striae not draWn); N, left foUrth pereopod, lateral vieW (setae and short striae not draWn); O, left foUrth pereopod, merUs, mesial vieW (short striae not draWn). Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287143/files/figure.png" pageId="4" pageNumber="115">Fig. 4E</figureCitation>
) distinctly broader than long, with each posterior plate broader than long.
</paragraph>
<paragraph id="8BA136FDFFDB051DD4A809C9FACDFC2F" blockId="4.[151,1437,386,1851]" box="[199,1319,926,951]" pageId="4" pageNumber="115">
Eyes (
<figureCitation id="13252A78FFDB051DD56309C8FE9AFC2F" box="[268,368,926,951]" captionStart="FIGURE 3" captionStartId="5.[151,250,1751,1773]" captionTargetBox="[218,1362,206,1697]" captionTargetId="figure@5.[205,1375,193,1724]" captionTargetPageId="5" captionText="FIGURE 3. Raphidopus Johnsoni Ng &amp; Nakasone, 1994, holotype, male (cl 3.8 mm), ZRC 1992.10669, Singapore. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (transverse striae and ridges not draWn); B, rostrUm, dorso-anterior vieW; C, third and foUrth thoracic sternites, ventral vieW; D, basal article of left antennUlar pedUncle, ventral vieW; E, left third maxilliped, lateral vieW (sUrface strUctUre not draWn); F, larger cheliped, merUs, ventral vieW (sUrface strUctUre not draWn); G, larger cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); H, larger cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); I, smaller cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); J, smaller cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); K, left foUrth pereopod, lateral vieW (short striae not draWn); L, same, merUs, lateral perpendicUlar vieW (short striae not draWn). Setae are not draWn on all figUres. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287141/files/figure.png" pageId="4" pageNumber="115">Figs. 3A</figureCitation>
,
<figureCitation id="13252A78FFDB051DD51309C9FE2BFC2F" box="[380,449,927,951]" captionStart="FIGURE 4" captionStartId="6.[151,250,1765,1787]" captionTargetBox="[263,1328,214,1697]" captionTargetId="figure@6.[251,1343,193,1738]" captionTargetPageId="6" captionText="FIGURE 4. Raphidopus Johnsoni Ng &amp; Nakasone, 1994, male (cl 5.0 mm), CBM-ZC 12485, PhUket. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (marginal setae not draWn on right side); B, right branchial margin of carapace, dorsomesial vieW; C, rostrUm, dorso-anterior vieW; D, third and foUrth thoracic sternites, ventral vieW (setae not draWn on left side); E, telson, external vieW; F, basal article of left antennUlar pedUncle, ventral vieW; G, left ocUlar and antennal pedUncles, dorsolateral vieW; H, left third maxilliped, lateral vieW; I, left second pereopod, lateral vieW; J, left second pereopod, merUs, mesial vieW (short striae not draWn); K, left second pereopod, dactylUs, lateral vieW (setae not draWn); L, left third pereopod, lateral vieW (setae and short striae not draWn); M, left third pereopod, merUs, mesial vieW (short striae not draWn); N, left foUrth pereopod, lateral vieW (setae and short striae not draWn); O, left foUrth pereopod, merUs, mesial vieW (short striae not draWn). Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287143/files/figure.png" pageId="4" pageNumber="115">4A, G</figureCitation>
) moderately small, reaching to distinctly oVerreaching lateral limits of orbits.
</paragraph>
<paragraph id="8BA136FDFFDB051DD4A80994FAE3FC43" blockId="4.[151,1437,386,1851]" box="[199,1289,962,987]" pageId="4" pageNumber="115">
Basal article of antennular peduncle (
<figureCitation id="13252A78FFDB051DD6060994FD26FC43" box="[617,716,962,987]" captionStart="FIGURE 3" captionStartId="5.[151,250,1751,1773]" captionTargetBox="[218,1362,206,1697]" captionTargetId="figure@5.[205,1375,193,1724]" captionTargetPageId="5" captionText="FIGURE 3. Raphidopus Johnsoni Ng &amp; Nakasone, 1994, holotype, male (cl 3.8 mm), ZRC 1992.10669, Singapore. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (transverse striae and ridges not draWn); B, rostrUm, dorso-anterior vieW; C, third and foUrth thoracic sternites, ventral vieW; D, basal article of left antennUlar pedUncle, ventral vieW; E, left third maxilliped, lateral vieW (sUrface strUctUre not draWn); F, larger cheliped, merUs, ventral vieW (sUrface strUctUre not draWn); G, larger cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); H, larger cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); I, smaller cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); J, smaller cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); K, left foUrth pereopod, lateral vieW (short striae not draWn); L, same, merUs, lateral perpendicUlar vieW (short striae not draWn). Setae are not draWn on all figUres. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287141/files/figure.png" pageId="4" pageNumber="115">Figs. 3D</figureCitation>
,
<figureCitation id="13252A78FFDB051DD6B70995FD12FC43" box="[728,760,963,987]" captionStart="FIGURE 4" captionStartId="6.[151,250,1765,1787]" captionTargetBox="[263,1328,214,1697]" captionTargetId="figure@6.[251,1343,193,1738]" captionTargetPageId="6" captionText="FIGURE 4. Raphidopus Johnsoni Ng &amp; Nakasone, 1994, male (cl 5.0 mm), CBM-ZC 12485, PhUket. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (marginal setae not draWn on right side); B, right branchial margin of carapace, dorsomesial vieW; C, rostrUm, dorso-anterior vieW; D, third and foUrth thoracic sternites, ventral vieW (setae not draWn on left side); E, telson, external vieW; F, basal article of left antennUlar pedUncle, ventral vieW; G, left ocUlar and antennal pedUncles, dorsolateral vieW; H, left third maxilliped, lateral vieW; I, left second pereopod, lateral vieW; J, left second pereopod, merUs, mesial vieW (short striae not draWn); K, left second pereopod, dactylUs, lateral vieW (setae not draWn); L, left third pereopod, lateral vieW (setae and short striae not draWn); M, left third pereopod, merUs, mesial vieW (short striae not draWn); N, left foUrth pereopod, lateral vieW (setae and short striae not draWn); O, left foUrth pereopod, merUs, mesial vieW (short striae not draWn). Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287143/files/figure.png" pageId="4" pageNumber="115">4F</figureCitation>
) as long as broad or slightly longer than broad.
</paragraph>
<paragraph id="8BA136FDFFDB051DD4A809B0FC42FBBB" blockId="4.[151,1437,386,1851]" pageId="4" pageNumber="115">
Antennal peduncle slender (
<figureCitation id="13252A78FFDB051DD66309B0FD81FC67" box="[524,619,998,1023]" captionStart="FIGURE 4" captionStartId="6.[151,250,1765,1787]" captionTargetBox="[263,1328,214,1697]" captionTargetId="figure@6.[251,1343,193,1738]" captionTargetPageId="6" captionText="FIGURE 4. Raphidopus Johnsoni Ng &amp; Nakasone, 1994, male (cl 5.0 mm), CBM-ZC 12485, PhUket. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (marginal setae not draWn on right side); B, right branchial margin of carapace, dorsomesial vieW; C, rostrUm, dorso-anterior vieW; D, third and foUrth thoracic sternites, ventral vieW (setae not draWn on left side); E, telson, external vieW; F, basal article of left antennUlar pedUncle, ventral vieW; G, left ocUlar and antennal pedUncles, dorsolateral vieW; H, left third maxilliped, lateral vieW; I, left second pereopod, lateral vieW; J, left second pereopod, merUs, mesial vieW (short striae not draWn); K, left second pereopod, dactylUs, lateral vieW (setae not draWn); L, left third pereopod, lateral vieW (setae and short striae not draWn); M, left third pereopod, merUs, mesial vieW (short striae not draWn); N, left foUrth pereopod, lateral vieW (setae and short striae not draWn); O, left foUrth pereopod, merUs, mesial vieW (short striae not draWn). Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287143/files/figure.png" pageId="4" pageNumber="115">Fig. 4G</figureCitation>
). First article slightly shorter than combined second to fourth articles. Third article elongate rectangular, about 1.5 times longer than second.
</paragraph>
<paragraph id="8BA136FDFFDB051DD4A80E78FE5DFB17" blockId="4.[151,1437,386,1851]" pageId="4" pageNumber="115">
Third maxilliped (
<figureCitation id="13252A78FFDB051DD5F80E78FE10FBDF" box="[407,506,1070,1095]" captionStart="FIGURE 3" captionStartId="5.[151,250,1751,1773]" captionTargetBox="[218,1362,206,1697]" captionTargetId="figure@5.[205,1375,193,1724]" captionTargetPageId="5" captionText="FIGURE 3. Raphidopus Johnsoni Ng &amp; Nakasone, 1994, holotype, male (cl 3.8 mm), ZRC 1992.10669, Singapore. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (transverse striae and ridges not draWn); B, rostrUm, dorso-anterior vieW; C, third and foUrth thoracic sternites, ventral vieW; D, basal article of left antennUlar pedUncle, ventral vieW; E, left third maxilliped, lateral vieW (sUrface strUctUre not draWn); F, larger cheliped, merUs, ventral vieW (sUrface strUctUre not draWn); G, larger cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); H, larger cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); I, smaller cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); J, smaller cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); K, left foUrth pereopod, lateral vieW (short striae not draWn); L, same, merUs, lateral perpendicUlar vieW (short striae not draWn). Setae are not draWn on all figUres. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287141/files/figure.png" pageId="4" pageNumber="115">Figs. 3E</figureCitation>
,
<figureCitation id="13252A78FFDB051DD6680E79FDC1FBDF" box="[519,555,1071,1095]" captionStart="FIGURE 4" captionStartId="6.[151,250,1765,1787]" captionTargetBox="[263,1328,214,1697]" captionTargetId="figure@6.[251,1343,193,1738]" captionTargetPageId="6" captionText="FIGURE 4. Raphidopus Johnsoni Ng &amp; Nakasone, 1994, male (cl 5.0 mm), CBM-ZC 12485, PhUket. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (marginal setae not draWn on right side); B, right branchial margin of carapace, dorsomesial vieW; C, rostrUm, dorso-anterior vieW; D, third and foUrth thoracic sternites, ventral vieW (setae not draWn on left side); E, telson, external vieW; F, basal article of left antennUlar pedUncle, ventral vieW; G, left ocUlar and antennal pedUncles, dorsolateral vieW; H, left third maxilliped, lateral vieW; I, left second pereopod, lateral vieW; J, left second pereopod, merUs, mesial vieW (short striae not draWn); K, left second pereopod, dactylUs, lateral vieW (setae not draWn); L, left third pereopod, lateral vieW (setae and short striae not draWn); M, left third pereopod, merUs, mesial vieW (short striae not draWn); N, left foUrth pereopod, lateral vieW (setae and short striae not draWn); O, left foUrth pereopod, merUs, mesial vieW (short striae not draWn). Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287143/files/figure.png" pageId="4" pageNumber="115">4H</figureCitation>
) moderately stout, with short ridges on lateral surfaces of ischium to carpus. Merus with rounded lobe proximal to median part of Ventral margin. Propodus comparatiVely stout. Exopod broad, 0.7–0.8 width of ischium.
</paragraph>
<paragraph id="8BA136FDFFDB051DD4A80ECCFC17FA4B" blockId="4.[151,1437,386,1851]" pageId="4" pageNumber="115">
Chelipeds (
<figureCitation id="13252A78FFDB051DD5250ECCFE2CFB2B" box="[330,454,1178,1203]" captionStart="FIGURE 3" captionStartId="5.[151,250,1751,1773]" captionTargetBox="[218,1362,206,1697]" captionTargetId="figure@5.[205,1375,193,1724]" captionTargetPageId="5" captionText="FIGURE 3. Raphidopus Johnsoni Ng &amp; Nakasone, 1994, holotype, male (cl 3.8 mm), ZRC 1992.10669, Singapore. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (transverse striae and ridges not draWn); B, rostrUm, dorso-anterior vieW; C, third and foUrth thoracic sternites, ventral vieW; D, basal article of left antennUlar pedUncle, ventral vieW; E, left third maxilliped, lateral vieW (sUrface strUctUre not draWn); F, larger cheliped, merUs, ventral vieW (sUrface strUctUre not draWn); G, larger cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); H, larger cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); I, smaller cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); J, smaller cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); K, left foUrth pereopod, lateral vieW (short striae not draWn); L, same, merUs, lateral perpendicUlar vieW (short striae not draWn). Setae are not draWn on all figUres. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287141/files/figure.png" pageId="4" pageNumber="115">Figs. 3F–J</figureCitation>
,
<figureCitation id="13252A78FFDB051DD5BA0ECCFDF1FB2B" box="[469,539,1178,1203]" captionStart="FIGURE 5" captionStartId="7.[151,250,1377,1399]" captionTargetBox="[173,1420,218,1300]" captionTargetId="figure@7.[150,1436,193,1356]" captionTargetPageId="7" captionText="FIGURE 5. Raphidopus Johnsoni Ng &amp; Nakasone, 1994, male (cl 4.7 mm), CBM-ZC 12485, PhUket. A, larger cheliped, dorsal vieW; B, larger cheliped, merUs and carpUs, ventral vieW; C, larger cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre on palm and dactylUs not draWn); D, smaller cheliped, dorsal vieW (sUrface strUctUre not draWn); E, smaller cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn). Setae are not draWn on all figUres. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287145/files/figure.png" pageId="4" pageNumber="115">5A–G</figureCitation>
) unequal in size. Merus with strong slender spine on Ventro-anterior margin. Carpus 1.5–2.0 (larger cheliped) and 1.6–1.9 (smaller cheliped) times as long as broad; dorsal surface with short and moderately long rows of small tubercles entirely and with single, flattened tubercles distally, no blunt ridge on longitudinal midline; posterior margin unarmed, rounded entirely (larger) and somewhat bluntly angled on distal part (smaller). Chela 2.5–2.6 (larger) and 2.5–2.8 (smaller) times longer than broad; dorsal surface with coarse, small and relatiVely larger tubercles; cutting edge of fixed finger with row of small and moderately large, rounded or subacute teeth (larger) or with row of small teeth decreasing in size distally (smaller). Dactylus 0.4–0.5 length of chela; cutting edge with some small rounded teeth and distinct, blunt tooth on each of proximal and median parts, proximal tooth larger (larger), and with row of small, rounded teeth (smaller).
</paragraph>
<paragraph id="8BA136FDFFDB051DD4A80F88FE56F9FB" blockId="4.[151,1437,386,1851]" pageId="4" pageNumber="115">
Ambulatory legs (
<figureCitation id="13252A78FFDB051DD5FD0F88FDFEFA6E" box="[402,532,1502,1527]" captionStart="FIGURE 3" captionStartId="5.[151,250,1751,1773]" captionTargetBox="[218,1362,206,1697]" captionTargetId="figure@5.[205,1375,193,1724]" captionTargetPageId="5" captionText="FIGURE 3. Raphidopus Johnsoni Ng &amp; Nakasone, 1994, holotype, male (cl 3.8 mm), ZRC 1992.10669, Singapore. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (transverse striae and ridges not draWn); B, rostrUm, dorso-anterior vieW; C, third and foUrth thoracic sternites, ventral vieW; D, basal article of left antennUlar pedUncle, ventral vieW; E, left third maxilliped, lateral vieW (sUrface strUctUre not draWn); F, larger cheliped, merUs, ventral vieW (sUrface strUctUre not draWn); G, larger cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); H, larger cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); I, smaller cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); J, smaller cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); K, left foUrth pereopod, lateral vieW (short striae not draWn); L, same, merUs, lateral perpendicUlar vieW (short striae not draWn). Setae are not draWn on all figUres. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287141/files/figure.png" pageId="4" pageNumber="115">Figs. 3K, L</figureCitation>
,
<figureCitation id="13252A78FFDB051DD64F0F89FDB1FA6F" box="[544,603,1503,1527]" captionStart="FIGURE 4" captionStartId="6.[151,250,1765,1787]" captionTargetBox="[263,1328,214,1697]" captionTargetId="figure@6.[251,1343,193,1738]" captionTargetPageId="6" captionText="FIGURE 4. Raphidopus Johnsoni Ng &amp; Nakasone, 1994, male (cl 5.0 mm), CBM-ZC 12485, PhUket. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (marginal setae not draWn on right side); B, right branchial margin of carapace, dorsomesial vieW; C, rostrUm, dorso-anterior vieW; D, third and foUrth thoracic sternites, ventral vieW (setae not draWn on left side); E, telson, external vieW; F, basal article of left antennUlar pedUncle, ventral vieW; G, left ocUlar and antennal pedUncles, dorsolateral vieW; H, left third maxilliped, lateral vieW; I, left second pereopod, lateral vieW; J, left second pereopod, merUs, mesial vieW (short striae not draWn); K, left second pereopod, dactylUs, lateral vieW (setae not draWn); L, left third pereopod, lateral vieW (setae and short striae not draWn); M, left third pereopod, merUs, mesial vieW (short striae not draWn); N, left foUrth pereopod, lateral vieW (setae and short striae not draWn); O, left foUrth pereopod, merUs, mesial vieW (short striae not draWn). Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/1287143/files/figure.png" pageId="4" pageNumber="115">
<date id="FFA0103DFFDB051DD64F0F89FDDDFA6F" box="[544,567,1503,1527]" pageId="4" pageNumber="115">4I</date>
–O
</figureCitation>
) comparatiVely slender, lateral surface with short and minute, weak striae. Meri 3.7–3.9, 3.4–3.6, 2.8–3.1 times longer than high on respectiVe second to fourth pereopods. Carpi relatiVely slender. Propodi 3.0–3.5, 2.9–3.3, 2.7–3.0 times longer than high on respectiVe second to fourth pereopods. Dactyli 0.7–0.9 length of propodi.
</paragraph>
<paragraph id="8BA136FDFFDB051DD4A80C38FB10F957" blockId="4.[151,1437,386,1851]" pageId="4" pageNumber="115">
<emphasis id="B96AEAEFFFDB051DD4A80C38FEA2F91F" bold="true" box="[199,328,1646,1671]" pageId="4" pageNumber="115">Coloration</emphasis>
. Ng &amp;
<bibRefCitation id="EF8F4B0CFFDB051DD5C70C38FD86F91F" box="[424,620,1646,1671]" pageId="4" pageNumber="115" refId="ref6869" refString="Ng, P. K. L. &amp; Nakasone, Y. (1994) On the porcellanid genera Raphidopus Stimpson, 1858, and Pseudoporcellanella SankarankUtty, 1961, With description of a neW mangrove species, Raphidopus Johnsoni from Singapore (Decapoda, AnomUra). Crustaceana, 66, 1 - 21. https: // doi. org / 10.1163 / 156854094 X 00116" type="journal article">Nakasone (1994)</bibRefCitation>
noted that “The colour of the hairs of the liVe and freshly preserVed specimen is beige brown, the carapace and appendages being white”. In the preserVed specimens (
<figureCitation id="13252A78FFDB051DD1670CC4FA89F933" box="[1288,1379,1682,1707]" captionStart="FIGURE 9" captionStartId="12.[151,250,1662,1684]" captionTargetBox="[151,1436,193,1641]" captionTargetId="figure@12.[151,1436,193,1641]" captionTargetPageId="12" captionText="FIGURE 9. Preserved specimens of three Raphidopus species. Entire animal, dorsal vieW (A–D); carapace, dorsal vieW (E); larger and smaller chelipeds, dorsal vieW (F). A, Raphidopus indicus Henderson, 1893, holotype, male (cl 6.5 mm), NHM 1892.7.15.246; B, Raphidopus Johnsoni Ng &amp; Nakasone, 1994, holotype, male (cl 3.8 mm), ZRC 1992.10669; C, Raphidopus Johnsoni Ng &amp; Nakasone, 1994, female (cl 5.9 mm), CBM-ZC 12485; D, Raphidopus brevipes n. sp., female (cl 5.1 mm), ZRC 2016.528; E, Raphidopus brevipes n. sp., male, holotype, male (cl 5.8 mm), ZRC 1992.10521; F, Raphidopus brevipes n. sp., paratypes, ZRC 1992.10522–10526." httpUri="https://zenodo.org/record/1287153/files/figure.png" pageId="4" pageNumber="115">Fig. 9C</figureCitation>
), the body and pereopods are generally whitish or pale brown, and the soft plumose setae are dark brown.
</paragraph>
</subSubSection>
<subSubSection id="C3046576FFDB051ED4A80C8CFE34F8A0" lastPageId="7" lastPageNumber="118" pageId="4" pageNumber="115" type="distribution">
<paragraph id="8BA136FDFFDB051DD4A80C8CFC33F8A3" blockId="4.[151,1437,386,1851]" pageId="4" pageNumber="115">
<emphasis id="B96AEAEFFFDB051DD4A80C8CFE8AF96B" bold="true" box="[199,352,1754,1779]" pageId="4" pageNumber="115">Distribution.</emphasis>
PreViously known only from
<collectingCountry id="F309766DFFDB051DD6D20C8CFCC5F96B" box="[701,815,1754,1779]" name="Singapore" pageId="4" pageNumber="115">Singapore</collectingCountry>
(Ng &amp;
<bibRefCitation id="EF8F4B0CFFDB051DD7E20C8CFBABF96B" box="[909,1089,1754,1779]" pageId="4" pageNumber="115" refId="ref6869" refString="Ng, P. K. L. &amp; Nakasone, Y. (1994) On the porcellanid genera Raphidopus Stimpson, 1858, and Pseudoporcellanella SankarankUtty, 1961, With description of a neW mangrove species, Raphidopus Johnsoni from Singapore (Decapoda, AnomUra). Crustaceana, 66, 1 - 21. https: // doi. org / 10.1163 / 156854094 X 00116" type="journal article">Nakasone 1994</bibRefCitation>
). The present specimens from
<collectingRegion id="49DAF81FFFDB051DD4F80CA8FF0FF88F" box="[151,229,1790,1815]" country="Thailand" name="Phuket" pageId="4" pageNumber="115">Phuket</collectingRegion>
(
<collectingCountry id="F309766DFFDB051DD49B0CA8FEB1F88F" box="[244,347,1790,1815]" name="Thailand" pageId="4" pageNumber="115">Thailand</collectingCountry>
) (western coast of Peninsular
<collectingCountry id="F309766DFFDB051DD6C00CA8FCFDF88F" box="[687,791,1790,1815]" name="Malaysia" pageId="4" pageNumber="115">Malaysia</collectingCountry>
) and
<collectingRegion id="49DAF81FFFDB051DD7380CA9FC63F88E" box="[855,905,1791,1814]" country="Indonesia" name="Jawa Timur" pageId="4" pageNumber="115">JaVa</collectingRegion>
Island (
<collectingCountry id="F309766DFFDB051DD78B0CA8FBBDF88F" box="[996,1111,1790,1815]" name="Indonesia" pageId="4" pageNumber="115">Indonesia</collectingCountry>
), are additional records of
<taxonomicName id="4C1E4D7EFFDB051DD1EB0CA9FF1DF8A3" authorityName="Ng &amp; Nakasone" authorityYear="1994" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="115" phylum="Arthropoda" rank="species" species="johnsoni">
<emphasis id="B96AEAEFFFDB051DD1EB0CA9FF1DF8A3" italics="true" pageId="4" pageNumber="115">R. johnsoni</emphasis>
</taxonomicName>
and extend the known range of the species north and southwards.
</paragraph>
<caption id="DF616675FFDA051CD4F80C81FEDBF87A" httpUri="https://zenodo.org/record/1287141/files/figure.png" pageId="5" pageNumber="116" startId="5.[151,250,1751,1773]" targetBox="[218,1362,206,1697]" targetPageId="5">
<paragraph id="8BA136FDFFDA051CD4F80C81FEDBF87A" blockId="5.[151,1436,1751,2018]" pageId="5" pageNumber="116">
<emphasis id="B96AEAEFFFDA051CD4F80C81FEFDF974" bold="true" box="[151,279,1751,1773]" pageId="5" pageNumber="116">FIGURE 3.</emphasis>
<taxonomicName id="4C1E4D7EFFDA051CD54C0C81FD17F975" authority="Johnsoni Ng &amp; Nakasone, 1994" authorityName="Johnsoni Ng &amp; Nakasone" authorityYear="1994" box="[291,765,1751,1773]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAEFFFDA051CD54C0C81FE14F975" box="[291,510,1751,1773]" italics="true" pageId="5" pageNumber="116">Raphidopus Johnsoni</emphasis>
Ng &amp; Nakasone, 1994
</taxonomicName>
, holotype, male (cl 3.8 mm), ZRC 1992.10669, Singapore. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (transverse striae and ridges not draWn); B, rostrUm, dorso-anterior vieW; C, third and foUrth thoracic sternites, ventral vieW; D, basal article of left antennUlar pedUncle, ventral vieW; E, left third maxilliped, lateral vieW (sUrface strUctUre not draWn); F, larger cheliped, merUs, ventral vieW (sUrface strUctUre not draWn); G, larger cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); H, larger cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); I, smaller cheliped, carpUs, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); J, smaller cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn); K, left foUrth pereopod, lateral vieW (short striae not draWn); L, same, merUs, lateral perpendicUlar vieW (short striae not draWn). Setae are not draWn on all figUres. Scale bars = 1.0 mm.
</paragraph>
</caption>
<caption id="DF616675FFD9051FD4F80CB3FE84F869" httpUri="https://zenodo.org/record/1287143/files/figure.png" pageId="6" pageNumber="117" startId="6.[151,250,1765,1787]" targetBox="[263,1328,214,1697]" targetPageId="6">
<paragraph id="8BA136FDFFD9051FD4F80CB3FE84F869" blockId="6.[151,1436,1765,2033]" pageId="6" pageNumber="117">
<emphasis id="B96AEAEFFFD9051FD4F80CB3FEF9F962" bold="true" box="[151,275,1765,1787]" pageId="6" pageNumber="117">FIGURE 4.</emphasis>
<taxonomicName id="4C1E4D7EFFD9051FD5750CB3FD0AF963" authority="Johnsoni Ng &amp; Nakasone, 1994" authorityName="Johnsoni Ng &amp; Nakasone" authorityYear="1994" box="[282,736,1765,1787]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAEFFFD9051FD5750CB3FE1BF963" box="[282,497,1765,1787]" italics="true" pageId="6" pageNumber="117">Raphidopus Johnsoni</emphasis>
Ng &amp; Nakasone, 1994
</taxonomicName>
, male (cl 5.0 mm), CBM-ZC 12485, PhUket. A, carapace and ocUlar and antennal pedUncles, dorsal vieW (marginal setae not draWn on right side); B, right branchial margin of carapace, dorsomesial vieW; C, rostrUm, dorso-anterior vieW; D, third and foUrth thoracic sternites, ventral vieW (setae not draWn on left side); E, telson, external vieW; F, basal article of left antennUlar pedUncle, ventral vieW; G, left ocUlar and antennal pedUncles, dorsolateral vieW; H, left third maxilliped, lateral vieW; I, left second pereopod, lateral vieW; J, left second pereopod, merUs, mesial vieW (short striae not draWn); K, left second pereopod, dactylUs, lateral vieW (setae not draWn); L, left third pereopod, lateral vieW (setae and short striae not draWn); M, left third pereopod, merUs, mesial vieW (short striae not draWn); N, left foUrth pereopod, lateral vieW (setae and short striae not draWn); O, left foUrth pereopod, merUs, mesial vieW (short striae not draWn). Scale bars = 1.0 mm.
</paragraph>
</caption>
<caption id="DF616675FFD8051ED4F80F37FAE7FA4B" httpUri="https://zenodo.org/record/1287145/files/figure.png" pageId="7" pageNumber="118" startId="7.[151,250,1377,1399]" targetBox="[173,1420,218,1300]" targetPageId="7">
<paragraph id="8BA136FDFFD8051ED4F80F37FAE7FA4B" blockId="7.[151,1436,1377,1491]" pageId="7" pageNumber="118">
<emphasis id="B96AEAEFFFD8051ED4F80F37FEF8FAEF" bold="true" box="[151,274,1377,1399]" pageId="7" pageNumber="118">FIGURE 5.</emphasis>
<taxonomicName id="4C1E4D7EFFD8051ED5770F37FD3CFAEF" authority="Johnsoni Ng &amp; Nakasone, 1994" authorityName="Johnsoni Ng &amp; Nakasone" authorityYear="1994" box="[280,726,1377,1399]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="118" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAEFFFD8051ED5770F37FE07FAEF" box="[280,493,1377,1399]" italics="true" pageId="7" pageNumber="118">Raphidopus Johnsoni</emphasis>
Ng &amp; Nakasone, 1994
</taxonomicName>
, male (cl 4.7 mm), CBM-ZC 12485, PhUket. A, larger cheliped, dorsal vieW; B, larger cheliped, merUs and carpUs, ventral vieW; C, larger cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre on palm and dactylUs not draWn); D, smaller cheliped, dorsal vieW (sUrface strUctUre not draWn); E, smaller cheliped, chela, dorsal perpendicUlar vieW (sUrface strUctUre not draWn). Setae are not draWn on all figUres. Scale bars = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BA136FDFFD8051ED4A80C56FE34F8A0" blockId="7.[151,1437,1536,2029]" pageId="7" pageNumber="118">
<emphasis id="B96AEAEFFFD8051ED4A80C56FEC0F981" bold="true" box="[199,298,1536,1561]" pageId="7" pageNumber="118">Habitat.</emphasis>
The specimens examined from Phuket were collected from burrows on the tidal sand flat using a stainless-steel suction pump (yabby pump; cf.
<bibRefCitation id="EF8F4B0CFFD8051ED6CC0C72FC8FF9A5" box="[675,869,1572,1597]" pageId="7" pageNumber="118" refId="ref6566" refString="DWorschak, P. C. (2015) Methods collecting Axiidea and Gebiidea (Decapoda): a revieW. Annalen des Naturhistorischen Museums in Wien, 117 B, 5 - 21." type="journal article">Dworschak 2015</bibRefCitation>
;
<bibRefCitation id="EF8F4B0CFFD8051ED71E0C72FBA5F9A5" box="[881,1103,1572,1597]" pageId="7" pageNumber="118" refId="ref6450" refString="Ahyong, S. T., CaldWell, R. L. &amp; Erdmann, M. V. (2017) Collecting and processing stomatopods. Journal of Crustacean Biology, 37, 109 - 114. https: // doi. org / 10.1093 / jcbiol / rUW 005" type="journal article">
Ahyong
<emphasis id="B96AEAEFFFD8051ED7BC0C73FBE6F9A4" box="[979,1036,1572,1596]" italics="true" pageId="7" pageNumber="118">et al.</emphasis>
2017
</bibRefCitation>
), but the hosts in the burrows could not be collected. Ng &amp;
<bibRefCitation id="EF8F4B0CFFD8051ED5890C1EFD4DF9F8" box="[486,679,1608,1633]" pageId="7" pageNumber="118" refId="ref6869" refString="Ng, P. K. L. &amp; Nakasone, Y. (1994) On the porcellanid genera Raphidopus Stimpson, 1858, and Pseudoporcellanella SankarankUtty, 1961, With description of a neW mangrove species, Raphidopus Johnsoni from Singapore (Decapoda, AnomUra). Crustaceana, 66, 1 - 21. https: // doi. org / 10.1163 / 156854094 X 00116" type="journal article">Nakasone (1994)</bibRefCitation>
mentioned that the holotype of
<taxonomicName id="4C1E4D7EFFD8051ED07F0C1FFB7AF9F8" authorityName="Ng &amp; Nakasone" authorityYear="1994" box="[1040,1168,1608,1632]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="118" phylum="Arthropoda" rank="species" species="johnsoni">
<emphasis id="B96AEAEFFFD8051ED07F0C1FFB7AF9F8" box="[1040,1168,1608,1632]" italics="true" pageId="7" pageNumber="118">R. johnsoni</emphasis>
</taxonomicName>
was obtained from soft mangroVe mud in the littoral zone and it seemed to be commensal with tubicolous worms that were relatiVely common in the mangroVes of the collection site. Although the substratum from which
<taxonomicName id="4C1E4D7EFFD8051ED00C0CC7FB08F930" authorityName="Ng &amp; Nakasone" authorityYear="1994" box="[1123,1250,1680,1704]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="118" phylum="Arthropoda" rank="species" species="johnsoni">
<emphasis id="B96AEAEFFFD8051ED00C0CC7FB08F930" box="[1123,1250,1680,1704]" italics="true" pageId="7" pageNumber="118">R. johnsoni</emphasis>
</taxonomicName>
was collected is different between Singapore and Phuket, we expect the species is probably commensal with polychaetes such as those in the family
<taxonomicName id="4C1E4D7EFFD8051ED5EE0C8EFDE4F969" box="[385,526,1752,1777]" class="Polychaeta" family="Terebellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Terebellida" pageId="7" pageNumber="118" phylum="Annelida" rank="family">Terebellidae</taxonomicName>
, as presumed by Ng &amp;
<bibRefCitation id="EF8F4B0CFFD8051ED75F0C8EFC1DF968" box="[816,1015,1752,1777]" pageId="7" pageNumber="118" refId="ref6869" refString="Ng, P. K. L. &amp; Nakasone, Y. (1994) On the porcellanid genera Raphidopus Stimpson, 1858, and Pseudoporcellanella SankarankUtty, 1961, With description of a neW mangrove species, Raphidopus Johnsoni from Singapore (Decapoda, AnomUra). Crustaceana, 66, 1 - 21. https: // doi. org / 10.1163 / 156854094 X 00116" type="journal article">Nakasone (1994)</bibRefCitation>
, as is also the case for
<taxonomicName id="4C1E4D7EFFD8051ED1790C8EFF1FF88C" authority="Osawa &amp; Ng, 2018" authorityName="Osawa &amp; Ng" authorityYear="2018" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="118" phylum="Arthropoda" rank="species" species="brevipes" status="sp. nov.">
<emphasis id="B96AEAEFFFD8051ED1790C8EFF1FF88C" italics="true" pageId="7" pageNumber="118">Raphidopus brevipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2595794FFD8051ED56F0CAAFEAEF88D" box="[256,324,1788,1813]" pageId="7" pageNumber="118" rank="species">
<emphasis id="B96AEAEFFFD8051ED56F0CAAFEAEF88D" bold="true" box="[256,324,1788,1813]" pageId="7" pageNumber="118">n. sp.</emphasis>
</taxonomicNameLabel>
shown below. AVailable specimens indicate that
<taxonomicName id="4C1E4D7EFFD8051ED7E30CABFBFBF88C" authorityName="Ng &amp; Nakasone" authorityYear="1994" box="[908,1041,1788,1812]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="118" phylum="Arthropoda" rank="species" species="johnsoni">
<emphasis id="B96AEAEFFFD8051ED7E30CABFBFBF88C" box="[908,1041,1788,1812]" italics="true" pageId="7" pageNumber="118">R. johnsoni</emphasis>
</taxonomicName>
occurs in the intertidal region to subtidal depth of about
<quantity id="4CE69B18FFD8051ED5F00D76FE30F8A0" box="[415,474,1824,1848]" metricMagnitude="1" metricUnit="m" metricValue="1.1" pageId="7" pageNumber="118" unit="m" value="11.0">11 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3046576FFD8051ED4A80D12FC09F875" pageId="7" pageNumber="118" type="discussion">
<paragraph id="8BA136FDFFD8051ED4A80D12FE3DF83D" blockId="7.[151,1437,1536,2029]" pageId="7" pageNumber="118">
<emphasis id="B96AEAEFFFD8051ED4A80D12FED1F8C5" bold="true" box="[199,315,1860,1885]" pageId="7" pageNumber="118">Remarks.</emphasis>
Re-examination of the specimens reported as
<taxonomicName id="4C1E4D7EFFD8051ED72D0D13FC58F8C4" authorityName="Henderson" authorityYear="1893" box="[834,946,1860,1884]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="118" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B96AEAEFFFD8051ED72D0D13FC58F8C4" box="[834,946,1860,1884]" italics="true" pageId="7" pageNumber="118">R. indicus</emphasis>
</taxonomicName>
by
<bibRefCitation id="EF8F4B0CFFD8051ED7B30D12FB8EF8C5" box="[988,1124,1860,1885]" pageId="7" pageNumber="118" refId="ref6637" refString="Haig, J. (1981) Porcellanid crabs from the Indo-West Pacific, Part II. Steenstrupia, 7 (12), 269 - 291." type="journal article">Haig (1981)</bibRefCitation>
has shown that they belong to
<taxonomicName id="4C1E4D7EFFD8051ED4DC0D3FFED8F818" authorityName="Ng &amp; Nakasone" authorityYear="1994" box="[179,306,1896,1920]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="118" phylum="Arthropoda" rank="species" species="johnsoni">
<emphasis id="B96AEAEFFFD8051ED4DC0D3FFED8F818" box="[179,306,1896,1920]" italics="true" pageId="7" pageNumber="118">R. johnsoni</emphasis>
</taxonomicName>
instead. Distinctions among
<taxonomicName id="4C1E4D7EFFD8051ED6190D3FFD0CF818" authorityName="Henderson" authorityYear="1893" box="[630,742,1896,1920]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="118" phylum="Arthropoda" rank="species" species="indicus">
<emphasis id="B96AEAEFFFD8051ED6190D3FFD0CF818" box="[630,742,1896,1920]" italics="true" pageId="7" pageNumber="118">R. indicus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1E4D7EFFD8051ED69B0D3FFC98F818" authorityName="Ng &amp; Nakasone" authorityYear="1994" box="[756,882,1896,1920]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="118" phylum="Arthropoda" rank="species" species="johnsoni">
<emphasis id="B96AEAEFFFD8051ED69B0D3FFC98F818" box="[756,882,1896,1920]" italics="true" pageId="7" pageNumber="118">R. johnsoni</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C1E4D7EFFD8051ED7C00D3FFBC6F818" authority="Osawa &amp; Ng, 2018" authorityName="Osawa &amp; Ng" authorityYear="2018" box="[943,1068,1896,1920]" class="Malacostraca" family="Porcellanidae" genus="Raphidopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="118" phylum="Arthropoda" rank="species" species="brevipes" status="sp. nov.">
<emphasis id="B96AEAEFFFD8051ED7C00D3FFBC6F818" box="[943,1068,1896,1920]" italics="true" pageId="7" pageNumber="118">R. brevipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2595794FFD8051ED05C0D3EFB9BF819" box="[1075,1137,1896,1921]" pageId="7" pageNumber="118" rank="species">
<emphasis id="B96AEAEFFFD8051ED05C0D3EFB9BF819" bold="true" box="[1075,1137,1896,1921]" pageId="7" pageNumber="118">n. sp.</emphasis>
</taxonomicNameLabel>
are discussed below in the Remarks of the new species.
</paragraph>
<paragraph id="8BA136FDFFD8051ED4A80DE7FC09F875" blockId="7.[151,1437,1536,2029]" pageId="7" pageNumber="118">
Four of the fiVe Phuket specimens (cl
<quantity id="4CE69B18FFD8051ED6140DE6FCEFF850" box="[635,773,1968,1993]" metricMagnitude="-3" metricUnit="m" metricValue="5.05" metricValueMax="5.9" metricValueMin="4.2" pageId="7" pageNumber="118" unit="mm" value="5.05" valueMax="5.9" valueMin="4.2">4.2–5.9 mm</quantity>
) are larger than the holotype from Singapore (cl
<quantity id="4CE69B18FFD8051ED15A0DE6FA7AF850" box="[1333,1424,1968,1992]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="7" pageNumber="118" unit="mm" value="3.8">3.8 mm</quantity>
), and the aVailable material shows that females reach larger sizes than males.
</paragraph>
</subSubSection>
</treatment>
</document>
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