422 lines
55 KiB
XML
422 lines
55 KiB
XML
<document id="A7F2B00E103F6A614742701C5828D0E7" ID-DOI="10.11646/zootaxa.4802.2.8" ID-ISSN="1175-5326" ID-Zenodo-Dep="3907594" ID-ZooBank="FC760D02-262B-42AB-83D6-1EA9C7C1D986" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1593080422928" checkinUser="plazi" docAuthor="Gorbunov, Oleg G. & Arita, Yutaka" docDate="2020" docId="03B7879FFF8CEE7CF6C3BBE085A27DD3" docLanguage="en" docName="zootaxa.4802.2.8.pdf" docOrigin="Zootaxa 4802 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Toleria vietnamica Gorbunov & Arita 2020, sp. nov." docType="treatment" docVersion="3" lastPageNumber="353" masterDocId="FF8EFFE7FF8DEE78F654BE4986427B0C" masterDocTitle="A new species of the genus Toleria Walker, 1865 [“ 1864 ”] from Vietnam, with a catalogue of Asian Cissuvorini (Lepidoptera: Sesiidae)" masterLastPageNumber="360" masterPageNumber="349" pageNumber="350" updateTime="1700276952666" updateUser="plazi" zenodo-license-document="CLOSED">
|
||
<mods:mods id="F650F052773209BEC1563C1763DB0FEC" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="54CD3FE794E5D160C3B0123A59847BC1">
|
||
<mods:title id="CF09B16BD34A9400AA490C6D00479B58">A new species of the genus Toleria Walker, 1865 [“ 1864 ”] from Vietnam, with a catalogue of Asian Cissuvorini (Lepidoptera: Sesiidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="16848968FFA1AF9A6B5DBAAEAE61487A" type="personal">
|
||
<mods:role id="1B4E565D3BBB29C05275CB74DD726773">
|
||
<mods:roleTerm id="1F4466C53D0D0F2CCB9106C10F17A570">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="9728F555ED1130C952DBEEDA8B6C6389">Gorbunov, Oleg G.</mods:namePart>
|
||
<mods:affiliation id="A0EA630DACC789E8155D0CC25562F9B0">A. N. Severtsov Institute of Ecology and Evolution, Russian Academy of Sciences, Leninsky prospect 33, 119071 Moscow, Russia. gorbunov. oleg @ mail. ru; https: // orcid. org / 0000 - 0001 - 5706 - 621 X</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="92E97DE1333EB46F0FA4C9EF8AF6EA05" type="personal">
|
||
<mods:role id="2C0E22023D85D1187F49BBDA49CE723B">
|
||
<mods:roleTerm id="39087B0E0C6733254CCE71551AB7261C">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="1701C9BAFAD1F019AB4224DBD902E357">Arita, Yutaka</mods:namePart>
|
||
<mods:affiliation id="5893D9BA4DCF2D5B8B9F322D04DA7C10">Ida 2489, Iga-shi, Mie-ken, 518 - 0123 Japan. arita. yutaka @ gmail. com; https: // orcid. org / 0000 - 0003 - 4815 - 5389 Corresponding author</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource id="F5E142EBBD68A6C9A9D1D10471EBEAFE">text</mods:typeOfResource>
|
||
<mods:relatedItem id="DD15281B1BE572B5E9DC61AB4E10C0FD" type="host">
|
||
<mods:titleInfo id="6CA364E5EF3678C373938BB38DD3C158">
|
||
<mods:title id="5D0F0378C67E6A9A7BC48A2EE6F64F46">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="E429624D94064E7AF1F24411A356644E">
|
||
<mods:date id="67E6A9195A24D50D59CD3ABBC0D94492">2020</mods:date>
|
||
<mods:detail id="EB580B8207C2C51BF02BEBAD053D0B32" type="pubDate">
|
||
<mods:number id="7C64DF66779BC0E3DB26FDFBE6EB7655">2020-06-23</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="4D38BBBA23678A508D91770D6E3F0F75" type="volume">
|
||
<mods:number id="D0AEAC44427BB44BB6013DA17CCDE9FD">4802</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="250C05D2B99860CF0A1265373AA2B775" type="issue">
|
||
<mods:number id="54CD8D9777F744A4FE9B174EC388DDA2">2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="797B75D49BCDAABD0608732590C49065" unit="page">
|
||
<mods:start id="98AE2906FEC49AB33E0A67DF85E4AE55">349</mods:start>
|
||
<mods:end id="27B9FC2C993696E68E486039F983443B">360</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="7EAEA7EB8126A01E19D2923B8D262900">journal article</mods:classification>
|
||
<mods:identifier id="27ACD79828344DCE8D75E896299018BC" type="DOI">10.11646/zootaxa.4802.2.8</mods:identifier>
|
||
<mods:identifier id="E3A7347AE21DD34A4539C688FF3212DB" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="722212C6AB79BB917B9C5A7EFC9E849E" type="Zenodo-Dep">3907594</mods:identifier>
|
||
<mods:identifier id="A5834D1D9EBFFE81B563CEDDC70FF66F" type="ZooBank">FC760D02-262B-42AB-83D6-1EA9C7C1D986</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03B7879FFF8CEE7CF6C3BBE085A27DD3" LSID="urn:lsid:plazi:treatment:03B7879FFF8CEE7CF6C3BBE085A27DD3" httpUri="http://treatment.plazi.org/id/03B7879FFF8CEE7CF6C3BBE085A27DD3" lastPageId="4" lastPageNumber="353" pageId="1" pageNumber="350">
|
||
<subSubSection id="C3046502FF8CEE79F6C3BBE0879F7ECF" box="[151,477,1449,1476]" pageId="1" pageNumber="350" type="nomenclature">
|
||
<paragraph id="8BA13689FF8CEE79F6C3BBE0879F7ECF" blockId="1.[151,477,1449,1511]" box="[151,477,1449,1476]" pageId="1" pageNumber="350">
|
||
<heading id="D0E981E5FF8CEE79F6C3BBE0879F7ECF" bold="true" box="[151,477,1449,1476]" fontSize="11" level="1" pageId="1" pageNumber="350" reason="1">
|
||
<emphasis id="B96AEA9BFF8CEE79F6C3BBE0879F7ECF" bold="true" box="[151,477,1449,1476]" pageId="1" pageNumber="350">
|
||
<taxonomicName id="4C1E4D0AFF8CEE79F6C3BBE0873A7ECF" ID-CoL="BDLZM" authority="Gorbunov & Arita, 2020" authorityName="Gorbunov & Arita" authorityYear="2020" box="[151,376,1449,1476]" class="Insecta" family="Sesiidae" genus="Toleria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="350" phylum="Arthropoda" rank="species" species="vietnamica" status="sp. nov.">
|
||
<emphasis id="B96AEA9BFF8CEE79F6C3BBE0873A7ECF" bold="true" box="[151,376,1449,1476]" italics="true" pageId="1" pageNumber="350">Toleria vietnamica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25957E0FF8CEE79F72BBBE0879F7ECF" box="[383,477,1449,1475]" pageId="1" pageNumber="350" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3046502FF8CEE79F6C3BB8487497EEB" box="[151,267,1485,1511]" pageId="1" pageNumber="350" type="description">
|
||
<paragraph id="8BA13689FF8CEE79F6C3BB8487497EEB" blockId="1.[151,477,1449,1511]" box="[151,267,1485,1511]" pageId="1" pageNumber="350">
|
||
(
|
||
<figureCitation id="13252A0CFF8CEE79F6CBBB8487407EEB" box="[159,258,1485,1511]" captionStart-0="FIGURES 1–2" captionStart-1="FIGURE 3" captionStartId-0="2.[151,264,1526,1551]" captionStartId-1="3.[151,250,1983,2008]" captionTargetBox-0="[155,1433,1101,1501]" captionTargetBox-1="[564,1023,190,1950]" captionTargetId-0="figure@2.[155,1433,1101,1501]" captionTargetId-1="figure@3.[555,1032,181,1958]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1–2. Female (holotype) of Toleria vietnamica sp. nov. 1) upside. Sesiidae picture № 0159–2018. Alar expanse 29.0 mm; 2) ditto, underside. Sesiidae picture № 0160–2018." captionText-1="FIGURE 3. Female genitalia of Toleria vietnamica sp. nov. Genital preparation № OG–027-2018. Scale bar: 2.0 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3907596" figureDoi-1="http://doi.org/10.5281/zenodo.3907598" httpUri-0="https://zenodo.org/record/3907596/files/figure.png" httpUri-1="https://zenodo.org/record/3907598/files/figure.png" pageId="1" pageNumber="350">Figs 1–3</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3046502FF8CEE79F6C3B85C85B87D97" pageId="1" pageNumber="350" type="materials_examined">
|
||
<paragraph id="8BA13689FF8CEE79F6C3B85C85B87D97" blockId="1.[151,1437,1557,2015]" pageId="1" pageNumber="350">
|
||
<materialsCitation id="3B763CD4FF8CEE79F6C3B85C85B87D97" collectingDate="2006-05-01" collectionCode="NSMT" collectorName="R. Matsumoto & O. Gorbunov" location="Ban Vai" municipality="Ba Be" pageId="1" pageNumber="350" specimenCount="2" specimenCount-female="2" stateProvince="Bac Kan" typeStatus="holotype">
|
||
<emphasis id="B96AEA9BFF8CEE79F6C3B85C87467D23" bold="true" box="[151,260,1557,1583]" pageId="1" pageNumber="350">
|
||
<typeStatus id="54A5882BFF8CEE79F6C3B85C87467D23" box="[151,260,1557,1583]" pageId="1" pageNumber="350" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<specimenCount id="9D18FD00FF8CEE79F75EB85C875D7D23" box="[266,287,1557,1583]" pageId="1" pageNumber="350" type="female">♀</specimenCount>
|
||
(
|
||
<figureCitation id="13252A0CFF8CEE79F779B85C87CD7D23" box="[301,399,1557,1583]" captionStart-0="FIGURES 1–2" captionStart-1="FIGURE 3" captionStartId-0="2.[151,264,1526,1551]" captionStartId-1="3.[151,250,1983,2008]" captionTargetBox-0="[155,1433,1101,1501]" captionTargetBox-1="[564,1023,190,1950]" captionTargetId-0="figure@2.[155,1433,1101,1501]" captionTargetId-1="figure@3.[555,1032,181,1958]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1–2. Female (holotype) of Toleria vietnamica sp. nov. 1) upside. Sesiidae picture № 0159–2018. Alar expanse 29.0 mm; 2) ditto, underside. Sesiidae picture № 0160–2018." captionText-1="FIGURE 3. Female genitalia of Toleria vietnamica sp. nov. Genital preparation № OG–027-2018. Scale bar: 2.0 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3907596" figureDoi-1="http://doi.org/10.5281/zenodo.3907598" httpUri-0="https://zenodo.org/record/3907596/files/figure.png" httpUri-1="https://zenodo.org/record/3907598/files/figure.png" pageId="1" pageNumber="350">Figs 1–3</figureCitation>
|
||
) with labels: “
|
||
<collectingRegion id="49DAF86BFF8CEE79F464B85C84D17D23" box="[560,659,1557,1583]" country="Vietnam" name="Bac Kan" pageId="1" pageNumber="350">Bac Kan</collectingRegion>
|
||
/
|
||
<collectingMunicipality id="6BC5ACF3FF8CEE79F4F2B85C84AC7D23" box="[678,750,1557,1583]" pageId="1" pageNumber="350">Ba Be</collectingMunicipality>
|
||
,
|
||
<location id="8EC16052FF8CEE79F4ACB85C85117D23" LSID="urn:lsid:plazi:treatment:03B7879FFF8CEE7CF6C3BBE085A27DD3:8EC16052FF8CEE79F4ACB85C85117D23" box="[760,851,1557,1583]" municipality="Ba Be" name="Ban Vai" pageId="1" pageNumber="350" stateProvince="Bac Kan">Ban Vai</location>
|
||
, /
|
||
<date id="FFA01049FF8CEE79F53FB85C858D7D23" box="[875,975,1557,1583]" pageId="1" pageNumber="350" value="2006-05-01">
|
||
<collectingDate id="EFE4E9A1FF8CEE79F53FB85C858D7D23" box="[875,975,1557,1583]" pageId="1" pageNumber="350" value="2006-05-01">1.V.2006</collectingDate>
|
||
</date>
|
||
/
|
||
<collectorName id="26EB535FFF8CEE79F5B0B85C82CB7D23" box="[996,1161,1557,1583]" pageId="1" pageNumber="350">R. Matsumoto</collectorName>
|
||
”; “Yutaka Arita / Collec- tion, NSMT / Donation, 2009 /
|
||
<emphasis id="B96AEA9BFF8CEE79F7ADB87E849B7D42" bold="true" box="[505,729,1591,1614]" pageId="1" pageNumber="350">Ħm DZOi± dzDz</emphasis>
|
||
”; “
|
||
<taxonomicName id="4C1E4D0AFF8CEE79F4AAB870853F7D5F" authorityName="Boisduval" authorityYear="1828" box="[766,893,1593,1619]" class="Insecta" family="Sesiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="350" phylum="Arthropoda" rank="family">SESIIDAE</taxonomicName>
|
||
/ Pictures №№ / 0159-0160–2018 / Photo by
|
||
<collectorName id="26EB535FFF8CEE79F3D5B870874D7D7B" pageId="1" pageNumber="350">O. Gorbunov</collectorName>
|
||
”; “Genitalia examined / by
|
||
<collectorName id="26EB535FFF8CEE79F411B81484987D7B" box="[581,730,1629,1655]" pageId="1" pageNumber="350">O. Gorbunov</collectorName>
|
||
/ Preparation № / OG–027-2018”; “
|
||
<typeStatus id="54A5882BFF8CEE79F224B81483587D7B" box="[1136,1306,1629,1655]" pageId="1" pageNumber="350" type="holotype">HOLOTYPUS</typeStatus>
|
||
<specimenCount id="9D18FD00FF8CEE79F375B81483747D7B" box="[1313,1334,1629,1655]" pageId="1" pageNumber="350" type="female">♀</specimenCount>
|
||
/
|
||
<taxonomicName id="4C1E4D0AFF8CEE79F318B81487507D97" authorityName="Gorbunov & Arita" authorityYear="2020" class="Insecta" family="Sesiidae" genus="Toleria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="350" phylum="Arthropoda" rank="species" species="vietnamica">
|
||
<emphasis id="B96AEA9BFF8CEE79F318B81487507D97" italics="true" pageId="1" pageNumber="350">Toleria vietnamica</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<collectorName id="26EB535FFF8CEE79F77CB8C887FE7D97" box="[296,444,1665,1691]" pageId="1" pageNumber="350">O. Gorbunov</collectorName>
|
||
et Arita, 2020 /
|
||
<collectorName id="26EB535FFF8CEE79F426B8C885447D97" box="[626,774,1665,1691]" pageId="1" pageNumber="350">O. Gorbunov</collectorName>
|
||
des., 2018” (
|
||
<collectionCode id="ED0FAE4CFF8CEE79F5CEB8C885AC7D97" box="[922,1006,1665,1691]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="1" pageNumber="350">NSMT</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3046502FF8CEE7AF693B8ED84087CB3" lastPageId="2" lastPageNumber="351" pageId="1" pageNumber="350" type="description">
|
||
<paragraph id="8BA13689FF8CEE79F693B8ED87107DEF" blockId="1.[151,1437,1557,2015]" pageId="1" pageNumber="350">
|
||
<emphasis id="B96AEA9BFF8CEE79F693B8ED87F57DB3" bold="true" box="[199,439,1700,1727]" pageId="1" pageNumber="350">Description. Female</emphasis>
|
||
(
|
||
<typeStatus id="54A5882BFF8CEE79F791B8EC84687DCC" box="[453,554,1701,1728]" pageId="1" pageNumber="350" type="holotype">holotype</typeStatus>
|
||
) (
|
||
<figureCitation id="13252A0CFF8CEE79F415B8EC84E57DB3" box="[577,679,1701,1727]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1526,1551]" captionTargetBox="[155,1433,1101,1501]" captionTargetId="figure@2.[155,1433,1101,1501]" captionTargetPageId="2" captionText="FIGURES 1–2. Female (holotype) of Toleria vietnamica sp. nov. 1) upside. Sesiidae picture № 0159–2018. Alar expanse 29.0 mm; 2) ditto, underside. Sesiidae picture № 0160–2018." figureDoi="http://doi.org/10.5281/zenodo.3907596" httpUri="https://zenodo.org/record/3907596/files/figure.png" pageId="1" pageNumber="350">Figs 1, 2</figureCitation>
|
||
). Alar expanse 31.0 mm; body length 15.1 mm; forewing 13.8 mm; antenna 6.2 mm.
|
||
</paragraph>
|
||
<paragraph id="8BA13689FF8CEE79F693B8A482717C43" blockId="1.[151,1437,1557,2015]" pageId="1" pageNumber="350">Head with antenna brown with a dense admixture of light brown scales dorsally on basal half; scapus yellow to pale yellow; frons gray with purple sheen and a very narrow pale yellow lateral margin; vertex and labial palpus entirely yellow to dark yellow; occipital fringe white dorsally and yellow laterally.</paragraph>
|
||
<paragraph id="8BA13689FF8CEE7AF693B91083DE7BBF" blockId="1.[151,1437,1557,2015]" lastBlockId="2.[151,1437,153,1079]" lastPageId="2" lastPageNumber="351" pageId="1" pageNumber="350">Thorax with patagia dark yellow to yellow-orange and with an admixture of dark brown scales with greenishpurple sheen dorsally; tegula dark brown with bronze-purple sheen, densely covered with light brown scales mask- ing background colouration and with a large yellow-orange spot at base of forewing; mesothorax dark brown with bronze-purple sheen and a dense admixture of dark yellow to yellow-orange scales posteriorly; metathorax dark brown with bronze-purple sheen, densely covered with chestnut-brown elongate scales; thorax laterally dark gray</paragraph>
|
||
<paragraph id="8BA13689FF8FEE7AF6C3BEF483DE7BDB" blockId="2.[151,1437,153,1079]" box="[151,1436,189,215]" pageId="2" pageNumber="351">with purple-violet sheen and an admixture of dark yellow and light brown scales anteriorly; posteriorly both met-</paragraph>
|
||
<paragraph id="8BA13689FF8FEE7AF6C3BEA883DE7BF7" blockId="2.[151,1437,153,1079]" box="[151,1436,225,251]" pageId="2" pageNumber="351">epimeron and metameron dark gray-brown with purple-violet sheen, covered with long light brown hairs. Legs with</paragraph>
|
||
<paragraph id="8BA13689FF8FEE7AF6C3BF4C83DE7A13" blockId="2.[151,1437,153,1079]" box="[151,1436,261,287]" pageId="2" pageNumber="351">neck plate light brown; fore coxa dark yellow to yellow-orange with a small dark gray spot with bronze sheen dis-</paragraph>
|
||
<paragraph id="8BA13689FF8FEE7AF6C3BF6083DE7A4F" blockId="2.[151,1437,153,1079]" box="[151,1436,297,323]" pageId="2" pageNumber="351">tally; fore femur dark yellow to yellow-orange; both fore tibia and fore tarsus light brown dorsally and dark yellow</paragraph>
|
||
<paragraph id="8BA13689FF8FEE7AF6C3BF0483DE7A6B" blockId="2.[151,1437,153,1079]" box="[151,1436,333,359]" pageId="2" pageNumber="351">ventrally; mid coxa dark gray with bronze sheen and a few light brown scales internally; mid femur externally dark</paragraph>
|
||
<paragraph id="8BA13689FF8FEE7AF6C3BF3883DE7A87" blockId="2.[151,1437,153,1079]" box="[151,1436,369,395]" pageId="2" pageNumber="351">yellow to yellow-orange and with a small black spot with bronze sheen distally and with a raw of light brown hairs</paragraph>
|
||
<paragraph id="8BA13689FF8FEE7AF6C3BFDC83DE7AA3" blockId="2.[151,1437,153,1079]" box="[151,1436,405,431]" pageId="2" pageNumber="351">at posterior margin, internally dark yellow; mid tibia light brown with an admixture of dark yellow scales medio-</paragraph>
|
||
<paragraph id="8BA13689FF8FEE7AF6C3BFF083DE7ADF" blockId="2.[151,1437,153,1079]" box="[151,1436,441,467]" pageId="2" pageNumber="351">internally; spurs dark yellow; mid tarsus dark yellow, basal tarsomere dorsally light brown; hind coxa mixed with</paragraph>
|
||
<paragraph id="8BA13689FF8FEE7AF6C3BF9483DF7AFB" blockId="2.[151,1437,153,1079]" box="[151,1437,477,503]" pageId="2" pageNumber="351">dark gray, dark yellow and light brown scales; hind femur externally dark yellow to yellow-orange and with a small</paragraph>
|
||
<paragraph id="8BA13689FF8FEE7AF6C3BC4883DE7917" blockId="2.[151,1437,153,1079]" box="[151,1436,513,539]" pageId="2" pageNumber="351">black spot with bronze sheen distally and with a raw of light brown hairs at posterior margin, internally dark yellow;</paragraph>
|
||
<paragraph id="8BA13689FF8FEE7AF6C3BC6C83DE7933" blockId="2.[151,1437,153,1079]" box="[151,1436,549,575]" pageId="2" pageNumber="351">hind tibia light brown, but slightly darker exterior-basally and somewhat lighter internally; spurs dark yellow; hind</paragraph>
|
||
<paragraph id="8BA13689FF8FEE7AF6C3BC0083DE796F" blockId="2.[151,1437,153,1079]" box="[151,1436,585,611]" pageId="2" pageNumber="351">tarsus light brown dorsally and dark yellow internally. Forewing dorsally with basal part black with dark blue sheen;</paragraph>
|
||
<paragraph id="8BA13689FF8FEE7AF6C3BC2483DE798B" blockId="2.[151,1437,153,1079]" box="[151,1436,621,647]" pageId="2" pageNumber="351">costal margin dark brown to black with dark blue sheen and a dense admixture of chestnut brown scales basally;</paragraph>
|
||
<paragraph id="8BA13689FF8FEE7AF6C3BCD883DE79A7" blockId="2.[151,1437,153,1079]" box="[151,1436,657,683]" pageId="2" pageNumber="351">CuA-stem, anal margin and veins distally of discal spot dark brown to black with dark blue sheen; discal spot nar-</paragraph>
|
||
<paragraph id="8BA13689FF8FEE7AF6C3BCFC83DE79C3" blockId="2.[151,1437,153,1079]" box="[151,1436,693,719]" pageId="2" pageNumber="351">row, S-shaped, black with dark blue-violet sheen; transparent areas well-developed; apical area undeveloped, but</paragraph>
|
||
<paragraph id="8BA13689FF8FEE7AF6C3BC9083DE781B" blockId="2.[151,1437,153,1079]" pageId="2" pageNumber="351">
|
||
surface of cells between veins R
|
||
<subScript id="179A34CCFF8FEE7AF7AABCA3844479F4" attach="left" box="[510,518,746,760]" fontSize="6" pageId="2" pageNumber="351">3</subScript>
|
||
–CuA
|
||
<subScript id="179A34CCFF8FEE7AF41DBCA3841379F4" attach="left" box="[585,593,746,760]" fontSize="6" pageId="2" pageNumber="351">1</subScript>
|
||
divided by an extremely narrow scaled line to middle from termen to discal spot; cilia dark gray-brown with bronze sheen; ventrally veins and discal spot dark brown with bronze sheen and
|
||
</paragraph>
|
||
<paragraph id="8BA13689FF8FEE7AF6C3BD6883DE7837" blockId="2.[151,1437,153,1079]" box="[151,1436,801,827]" pageId="2" pageNumber="351">a dense admixture of dark yellow scales on costal and anal margin and CuA-stem basally. Hindwing transparent;</paragraph>
|
||
<paragraph id="8BA13689FF8FEE7AF6C3BD0C83DE7853" blockId="2.[151,1437,153,1079]" box="[151,1436,837,863]" pageId="2" pageNumber="351">dorsally veins veins, discal spot, outer margin and cilia dark brown to black with bronze sheen; discal spot narrow,</paragraph>
|
||
<paragraph id="8BA13689FF8FEE7AF6C3BD20833778AB" blockId="2.[151,1437,153,1079]" pageId="2" pageNumber="351">
|
||
triangular, reaching to vein M
|
||
<subScript id="179A34CCFF8FEE7AF7B8BD3387B67884" attach="left" box="[492,500,890,904]" fontSize="6" pageId="2" pageNumber="351">2</subScript>
|
||
; outer margin extremely narrow, about 0.2 times as broad as cilia; ventrally veins dark brown with bronze sheen and an admixture of dark yellow scales on CuA-stem and veins CuA
|
||
<subScript id="179A34CCFF8FEE7AF2A6BDD782B878A0" attach="left" box="[1266,1274,926,940]" fontSize="6" pageId="2" pageNumber="351">1</subScript>
|
||
and CuA
|
||
<subScript id="179A34CCFF8FEE7AF332BDD7832C78A0" attach="left" box="[1382,1390,926,940]" fontSize="6" pageId="2" pageNumber="351">2</subScript>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA13689FF8FEE7AF693BDF883DE78C7" blockId="2.[151,1437,153,1079]" box="[199,1436,945,971]" pageId="2" pageNumber="351">Abdomen black with dark blue sheen; dorsally tergites 1 and 2 each densely covered with chestnut-brown elon-</paragraph>
|
||
<paragraph id="8BA13689FF8FEE7AF6C3BD9C83DF78E3" blockId="2.[151,1437,153,1079]" box="[151,1437,981,1007]" pageId="2" pageNumber="351">gate scales; tergite 1 with a large dark yellow to yellow-orange spot lateral-distally; tergite 2 with a narrow dark yel-</paragraph>
|
||
<paragraph id="8BA13689FF8FEE7AF6C3BDB083DF7F1F" blockId="2.[151,1437,153,1079]" box="[151,1437,1017,1043]" pageId="2" pageNumber="351">low to yellow-orange stripe distally; ventrally sternites 1+2 and 3 each with a broad dark yellow to yellow-orange</paragraph>
|
||
<paragraph id="8BA13689FF8FEE7AF6C3BA5485687F3B" blockId="2.[151,1437,153,1079]" box="[151,810,1053,1079]" pageId="2" pageNumber="351">stripe distally; anal tuft small dark gray with bronze sheen.</paragraph>
|
||
<caption id="DF616601FF8FEE7AF6C3BBBF84937D3F" ID-DOI="http://doi.org/10.5281/zenodo.3907596" ID-Zenodo-Dep="3907596" httpUri="https://zenodo.org/record/3907596/files/figure.png" pageId="2" pageNumber="351" startId="2.[151,264,1526,1551]" targetBox="[155,1433,1101,1501]" targetPageId="2">
|
||
<paragraph id="8BA13689FF8FEE7AF6C3BBBF84937D3F" blockId="2.[151,1436,1526,1587]" pageId="2" pageNumber="351">
|
||
<emphasis id="B96AEA9BFF8FEE7AF6C3BBBF87787D03" bold="true" box="[151,314,1526,1551]" pageId="2" pageNumber="351">FIGURES 1–2.</emphasis>
|
||
Female (holotype) of
|
||
<taxonomicName id="4C1E4D0AFF8FEE7AF449BBBE84987D02" authority="Gorbunov & Arita, 2020" authorityName="Gorbunov & Arita" authorityYear="2020" box="[541,730,1527,1551]" class="Insecta" family="Sesiidae" genus="Toleria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="351" phylum="Arthropoda" rank="species" species="vietnamica" status="sp. nov.">
|
||
<emphasis id="B96AEA9BFF8FEE7AF449BBBE84987D02" box="[541,730,1527,1551]" italics="true" pageId="2" pageNumber="351">Toleria vietnamica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96AEA9BFF8FEE7AF4B5BBBE85737D03" bold="true" box="[737,817,1527,1551]" pageId="2" pageNumber="351">
|
||
<taxonomicNameLabel id="A25957E0FF8FEE7AF4B5BBBE85737D03" box="[737,817,1527,1551]" pageId="2" pageNumber="351" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
1) upside.
|
||
<taxonomicName id="4C1E4D0AFF8FEE7AF5F6BBBF85B47D03" authorityName="Boisduval" authorityYear="1828" box="[930,1014,1526,1551]" class="Insecta" family="Sesiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="351" phylum="Arthropoda" rank="family">Sesiidae</taxonomicName>
|
||
picture № 0159–2018. Alar expanse 29.0 mm; 2) ditto, underside.
|
||
<taxonomicName id="4C1E4D0AFF8FEE7AF7C6B85387A47D3F" authorityName="Boisduval" authorityYear="1828" box="[402,486,1562,1587]" class="Insecta" family="Sesiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="351" phylum="Arthropoda" rank="family">Sesiidae</taxonomicName>
|
||
picture № 0160–2018.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA13689FF8FEE7AF693B82885217C7B" blockId="2.[151,1437,1632,1983]" pageId="2" pageNumber="351">
|
||
<emphasis id="B96AEA9BFF8FEE7AF693B82887CF7D77" bold="true" box="[199,397,1632,1659]" pageId="2" pageNumber="351">Female genitalia</emphasis>
|
||
(
|
||
<typeStatus id="54A5882BFF8FEE7AF7C9B82884407D70" box="[413,514,1633,1660]" pageId="2" pageNumber="351" type="holotype">holotype</typeStatus>
|
||
) (Genital preparation № OG–027-2018) (
|
||
<figureCitation id="13252A0CFF8FEE7AF589B82882607D77" box="[989,1058,1633,1659]" captionStart="FIGURE 3" captionStartId="3.[151,250,1983,2008]" captionTargetBox="[564,1023,190,1950]" captionTargetId="figure@3.[555,1032,181,1958]" captionTargetPageId="3" captionText="FIGURE 3. Female genitalia of Toleria vietnamica sp. nov. Genital preparation № OG–027-2018. Scale bar: 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3907598" httpUri="https://zenodo.org/record/3907598/files/figure.png" pageId="2" pageNumber="351">Fig. 3</figureCitation>
|
||
). Papillae anales well-sclerotized with rounded membranous areas at base of setae, covered with long and short setae; tergite 8 relatively broad, cov- ered with sparse short setae; posterior apophysis slightly longer than anterior apophysis; lamella postvaginalis pres- ent, H-shaped, with narrow, pointed “wings”; lamella antevaginalis absent; ostium bursae opening near posterior margin of sternite 7, relatively broad, membranous, tube-shaped; antrum short, with a narrow well-sclerotized semi- circle posteriorly and with a narrow well-sclerotized plate at base of ductus seminalis; ductus bursae long, relatively thin, membranous; corpus bursae large, ovoid, membranous, with numerous transverse folds; signum present, oval, with a short fold, consists of many well-sclerotized, small teeth.
|
||
</paragraph>
|
||
<paragraph id="8BA13689FF8FEE7AF693B9C887CA7C97" blockId="2.[151,1437,1632,1983]" box="[199,392,1921,1947]" pageId="2" pageNumber="351">
|
||
<emphasis id="B96AEA9BFF8FEE7AF693B9C887497C97" bold="true" box="[199,267,1921,1947]" pageId="2" pageNumber="351">Male.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
<paragraph id="8BA13689FF8FEE7AF693B9ED84087CB3" blockId="2.[151,1437,1632,1983]" box="[199,586,1956,1983]" pageId="2" pageNumber="351">
|
||
<emphasis id="B96AEA9BFF8FEE7AF693B9ED878F7CB3" bold="true" box="[199,461,1956,1983]" pageId="2" pageNumber="351">Individual variability.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF616601FF8EEE7BF6C3B9F683047CDB" ID-DOI="http://doi.org/10.5281/zenodo.3907598" ID-Zenodo-Dep="3907598" box="[151,1350,1983,2008]" httpUri="https://zenodo.org/record/3907598/files/figure.png" pageId="3" pageNumber="352" startId="3.[151,250,1983,2008]" targetBox="[564,1023,190,1950]" targetPageId="3">
|
||
<paragraph id="8BA13689FF8EEE7BF6C3B9F683047CDB" blockId="3.[151,1350,1983,2008]" box="[151,1350,1983,2008]" pageId="3" pageNumber="352">
|
||
<emphasis id="B96AEA9BFF8EEE7BF6C3B9F687517CD4" bold="true" box="[151,275,1983,2008]" pageId="3" pageNumber="352">FIGURE 3.</emphasis>
|
||
Female genitalia of
|
||
<taxonomicName id="4C1E4D0AFF8EEE7BF7B7B98984E37CDB" authority="Gorbunov & Arita, 2020" authorityName="Gorbunov & Arita" authorityYear="2020" box="[483,673,1984,2008]" class="Insecta" family="Sesiidae" genus="Toleria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="352" phylum="Arthropoda" rank="species" species="vietnamica" status="sp. nov.">
|
||
<emphasis id="B96AEA9BFF8EEE7BF7B7B98984E37CDB" box="[483,673,1984,2008]" italics="true" pageId="3" pageNumber="352">Toleria vietnamica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96AEA9BFF8EEE7BF4F3B98984BA7CD4" bold="true" box="[679,760,1984,2008]" pageId="3" pageNumber="352">
|
||
<taxonomicNameLabel id="A25957E0FF8EEE7BF4F3B98984BA7CD4" box="[679,760,1984,2008]" pageId="3" pageNumber="352" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Genital preparation № OG–027-2018. Scale bar: 2.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3046502FF89EE7CF693BED187E57D9B" pageId="4" pageNumber="353" type="diagnosis">
|
||
<paragraph id="8BA13689FF89EE7CF693BED1833F78AB" blockId="4.[151,1437,152,1759]" pageId="4" pageNumber="353">
|
||
<emphasis id="B96AEA9BFF89EE7CF693BED187927BBF" bold="true" box="[199,464,152,179]" pageId="4" pageNumber="353">Differential diagnosis.</emphasis>
|
||
Superficially this new species seems to be the closest to
|
||
<taxonomicName id="4C1E4D0AFF89EE7CF23EBED086997BDB" authority="(Bryk, 1947)" baseAuthorityName="Bryk" baseAuthorityYear="1947" class="Insecta" family="Sesiidae" genus="Toleria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="353" phylum="Arthropoda" rank="species" species="colochelyna">
|
||
<emphasis id="B96AEA9BFF89EE7CF23EBED083087BBF" box="[1130,1354,153,179]" italics="true" pageId="4" pageNumber="353">Toleria colochelyna</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8F4B78FF89EE7CF308BED086917BDB" author="Bryk, F." pageId="4" pageNumber="353" pagination="96 - 109" refId="ref7735" refString="Bryk, F. (1947) Neue ostasiatische Aegeriiden (Lep.). Opuscula Entomologica, 12, 96 - 109." type="journal article" year="1947">Bryk, 1947</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, but it can be distinguished by the colouration of the scapus (scapus yellow dorsally and light brown ventrally in
|
||
<taxonomicName id="4C1E4D0AFF89EE7CF6E0BEAB87147BF7" baseAuthorityName="Bryk" baseAuthorityYear="1947" box="[180,342,225,251]" class="Insecta" family="Sesiidae" genus="Toleria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="353" phylum="Arthropoda" rank="species" species="colochelyna">
|
||
<emphasis id="B96AEA9BFF89EE7CF6E0BEAB87147BF7" box="[180,342,225,251]" italics="true" pageId="4" pageNumber="353">T. colochelyna</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B96AEA9BFF89EE7CF730BEAB87397BF7" box="[356,379,226,251]" italics="true" pageId="4" pageNumber="353">vs</emphasis>
|
||
. scapus entirely yellow to pale yellow in
|
||
<taxonomicName id="4C1E4D0AFF89EE7CF518BEAB85A17BF7" authority="Gorbunov & Arita, 2020" authorityName="Gorbunov & Arita" authorityYear="2020" box="[844,995,226,251]" class="Insecta" family="Sesiidae" genus="Toleria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="353" phylum="Arthropoda" rank="species" species="vietnamica" status="sp. nov.">
|
||
<emphasis id="B96AEA9BFF89EE7CF518BEAB85A17BF7" box="[844,995,226,251]" italics="true" pageId="4" pageNumber="353">T. vietnamica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96AEA9BFF89EE7CF5BDBEA882017BF7" bold="true" box="[1001,1091,225,251]" pageId="4" pageNumber="353">
|
||
<taxonomicNameLabel id="A25957E0FF89EE7CF5BDBEA882017BF7" box="[1001,1091,225,251]" pageId="4" pageNumber="353" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
), occipital fringe (entirely yellow in
|
||
<taxonomicName id="4C1E4D0AFF89EE7CF6B7BF4F87C67A13" baseAuthorityName="Bryk" baseAuthorityYear="1947" box="[227,388,261,287]" class="Insecta" family="Sesiidae" genus="Toleria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="353" phylum="Arthropoda" rank="species" species="colochelyna">
|
||
<emphasis id="B96AEA9BFF89EE7CF6B7BF4F87C67A13" box="[227,388,261,287]" italics="true" pageId="4" pageNumber="353">T. colochelyna</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B96AEA9BFF89EE7CF7C5BF4F87EA7A13" box="[401,424,262,287]" italics="true" pageId="4" pageNumber="353">vs</emphasis>
|
||
. white dorsally and yellow laterally
|
||
<taxonomicName id="4C1E4D0AFF89EE7CF56FBF4F85937A13" authority="Gorbunov & Arita, 2020" authorityName="Gorbunov & Arita" authorityYear="2020" box="[827,977,262,287]" class="Insecta" family="Sesiidae" genus="Toleria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="353" phylum="Arthropoda" rank="species" species="vietnamica" status="sp. nov.">
|
||
<emphasis id="B96AEA9BFF89EE7CF56FBF4F85937A13" box="[827,977,262,287]" italics="true" pageId="4" pageNumber="353">T. vietnamica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96AEA9BFF89EE7CF582BF4C826D7A13" bold="true" box="[982,1071,261,287]" pageId="4" pageNumber="353">
|
||
<taxonomicNameLabel id="A25957E0FF89EE7CF582BF4C826D7A13" box="[982,1071,261,287]" pageId="4" pageNumber="353" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
), tegula (brown to dark brown in the species compared, vs. tegula dark brown densely covered with light brown scales masking background coloura- tion and with a large yellow-orange spot at base of forewing in
|
||
<taxonomicName id="4C1E4D0AFF89EE7CF501BF0785A97A6B" authority="Gorbunov & Arita, 2020" authorityName="Gorbunov & Arita" authorityYear="2020" box="[853,1003,334,359]" class="Insecta" family="Sesiidae" genus="Toleria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="353" phylum="Arthropoda" rank="species" species="vietnamica" status="sp. nov.">
|
||
<emphasis id="B96AEA9BFF89EE7CF501BF0785A97A6B" box="[853,1003,334,359]" italics="true" pageId="4" pageNumber="353">T. vietnamica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96AEA9BFF89EE7CF5A5BF0482087A6B" bold="true" box="[1009,1098,333,359]" pageId="4" pageNumber="353">
|
||
<taxonomicNameLabel id="A25957E0FF89EE7CF5A5BF0482087A6B" box="[1009,1098,333,359]" pageId="4" pageNumber="353" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
), thorax laterally (grey-brown with bronze sheen in
|
||
<taxonomicName id="4C1E4D0AFF89EE7CF7D9BF3B84737A87" baseAuthorityName="Bryk" baseAuthorityYear="1947" box="[397,561,369,395]" class="Insecta" family="Sesiidae" genus="Toleria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="353" phylum="Arthropoda" rank="species" species="colochelyna">
|
||
<emphasis id="B96AEA9BFF89EE7CF7D9BF3B84737A87" box="[397,561,369,395]" italics="true" pageId="4" pageNumber="353">T. colochelyna</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B96AEA9BFF89EE7CF414BF3B84157A87" box="[576,599,370,395]" italics="true" pageId="4" pageNumber="353">vs</emphasis>
|
||
. dark gray with purple-violet sheen and an admixture of dark yellow and light brown scales anteriorly in
|
||
<taxonomicName id="4C1E4D0AFF89EE7CF7A8BFDF84D17AA3" authority="Gorbunov & Arita, 2020" authorityName="Gorbunov & Arita" authorityYear="2020" box="[508,659,406,431]" class="Insecta" family="Sesiidae" genus="Toleria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="353" phylum="Arthropoda" rank="species" species="vietnamica" status="sp. nov.">
|
||
<emphasis id="B96AEA9BFF89EE7CF7A8BFDF84D17AA3" box="[508,659,406,431]" italics="true" pageId="4" pageNumber="353">T. vietnamica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96AEA9BFF89EE7CF4CEBFDC84B67AA3" bold="true" box="[666,756,405,431]" pageId="4" pageNumber="353">
|
||
<taxonomicNameLabel id="A25957E0FF89EE7CF4CEBFDC84B67AA3" box="[666,756,405,431]" pageId="4" pageNumber="353" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
), fore tibia (entirely yellow-orange in the species compared,
|
||
<emphasis id="B96AEA9BFF89EE7CF6C3BFF386EC7ADF" box="[151,174,442,467]" italics="true" pageId="4" pageNumber="353">vs</emphasis>
|
||
. light brown dorsally and dark yellow ventrally in
|
||
<taxonomicName id="4C1E4D0AFF89EE7CF4A3BFF385CD7ADF" authority="Gorbunov & Arita, 2020" authorityName="Gorbunov & Arita" authorityYear="2020" box="[759,911,442,467]" class="Insecta" family="Sesiidae" genus="Toleria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="353" phylum="Arthropoda" rank="species" species="vietnamica" status="sp. nov.">
|
||
<emphasis id="B96AEA9BFF89EE7CF4A3BFF385CD7ADF" box="[759,911,442,467]" italics="true" pageId="4" pageNumber="353">T. vietnamica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96AEA9BFF89EE7CF5C3BFF085B07ADF" bold="true" box="[919,1010,441,467]" pageId="4" pageNumber="353">
|
||
<taxonomicNameLabel id="A25957E0FF89EE7CF5C3BFF085B07ADF" box="[919,1010,441,467]" pageId="4" pageNumber="353" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
), hind tibia (externally orange with a small brown spot basally and yellow internally in
|
||
<taxonomicName id="4C1E4D0AFF89EE7CF4E9BF97851C7AFB" baseAuthorityName="Bryk" baseAuthorityYear="1947" box="[701,862,477,503]" class="Insecta" family="Sesiidae" genus="Toleria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="353" phylum="Arthropoda" rank="species" species="colochelyna">
|
||
<emphasis id="B96AEA9BFF89EE7CF4E9BF97851C7AFB" box="[701,862,477,503]" italics="true" pageId="4" pageNumber="353">T. colochelyna</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B96AEA9BFF89EE7CF53FBF9785C07AFB" box="[875,898,478,503]" italics="true" pageId="4" pageNumber="353">vs</emphasis>
|
||
. light brown, but slightly darker exterior-basally and somewhat lighter internally in
|
||
<taxonomicName id="4C1E4D0AFF89EE7CF475BC4B84FA7917" authority="Gorbunov & Arita, 2020" authorityName="Gorbunov & Arita" authorityYear="2020" box="[545,696,514,539]" class="Insecta" family="Sesiidae" genus="Toleria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="353" phylum="Arthropoda" rank="species" species="vietnamica" status="sp. nov.">
|
||
<emphasis id="B96AEA9BFF89EE7CF475BC4B84FA7917" box="[545,696,514,539]" italics="true" pageId="4" pageNumber="353">T. vietnamica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96AEA9BFF89EE7CF4EBBC48855B7917" bold="true" box="[703,793,513,539]" pageId="4" pageNumber="353">
|
||
<taxonomicNameLabel id="A25957E0FF89EE7CF4EBBC48855B7917" box="[703,793,513,539]" pageId="4" pageNumber="353" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
), and abdomen (dorsally tergites 1–4 each with a narrow yellow stripe posteriorly; tergite 1 with a large yellow spot laterally; tergites 2–3 each with admixture of yellow, orange and light brown scales medially; tergites 4–7 each densely covered with yellow and dirty orange scales, masking background coloration; ventrally sternites 1+2–4 each dark brown anteriorly and yellow to yellow-orange posteriorly; remaining sternites yellow-orange to light brown; anal tuft yellow-orange to orange in the species compared,
|
||
<emphasis id="B96AEA9BFF89EE7CF740BCFF876979C3" box="[276,299,694,719]" italics="true" pageId="4" pageNumber="353">vs</emphasis>
|
||
. dorsally tergites 1 and 2 each densely covered with chestnut-brown elongate scales; tergite 1 with a large dark yellow to yellow-orange spot lateral-distally; tergite 2 with a narrow dark yellow to yellow-orange stripe distally; ventrally sternites 1+2 and 3 each with a broad dark yellow to yellow-orange stripe distally; anal tuft dark gray with bronze sheen in
|
||
<taxonomicName id="4C1E4D0AFF89EE7CF7E9BD6B84117837" authority="Gorbunov & Arita, 2020" authorityName="Gorbunov & Arita" authorityYear="2020" box="[445,595,802,827]" class="Insecta" family="Sesiidae" genus="Toleria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="353" phylum="Arthropoda" rank="species" species="vietnamica" status="sp. nov.">
|
||
<emphasis id="B96AEA9BFF89EE7CF7E9BD6B84117837" box="[445,595,802,827]" italics="true" pageId="4" pageNumber="353">T. vietnamica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96AEA9BFF89EE7CF40EBD6884F17837" bold="true" box="[602,691,801,827]" pageId="4" pageNumber="353">
|
||
<taxonomicNameLabel id="A25957E0FF89EE7CF40EBD6884F17837" box="[602,691,801,827]" pageId="4" pageNumber="353" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
). In addition, these two species have differences in the structure of the female genitalia: papillae anales visibly broader and posterior apophysis relatively shorter in
|
||
<taxonomicName id="4C1E4D0AFF89EE7CF288BD0F83317853" authority="Gorbunov & Arita, 2020" authorityName="Gorbunov & Arita" authorityYear="2020" box="[1244,1395,838,863]" class="Insecta" family="Sesiidae" genus="Toleria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="353" phylum="Arthropoda" rank="species" species="vietnamica" status="sp. nov.">
|
||
<emphasis id="B96AEA9BFF89EE7CF288BD0F83317853" box="[1244,1395,838,863]" italics="true" pageId="4" pageNumber="353">T. vietnamica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96AEA9BFF89EE7CF32FBD0C868A788F" bold="true" pageId="4" pageNumber="353">
|
||
<taxonomicNameLabel id="A25957E0FF89EE7CF32FBD0C868A788F" pageId="4" pageNumber="353" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; tergite 8 and lamella postvaginalis more narrow in the new species; signum relatively larger, but with more short fold medially in
|
||
<taxonomicName id="4C1E4D0AFF89EE7CF7C4BDC7846578AB" authority="Gorbunov & Arita, 2020" authorityName="Gorbunov & Arita" authorityYear="2020" box="[400,551,910,935]" class="Insecta" family="Sesiidae" genus="Toleria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="353" phylum="Arthropoda" rank="species" species="vietnamica" status="sp. nov.">
|
||
<emphasis id="B96AEA9BFF89EE7CF7C4BDC7846578AB" box="[400,551,910,935]" italics="true" pageId="4" pageNumber="353">T. vietnamica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96AEA9BFF89EE7CF479BDC484C578AB" bold="true" box="[557,647,909,935]" pageId="4" pageNumber="353">
|
||
<taxonomicNameLabel id="A25957E0FF89EE7CF479BDC484C578AB" box="[557,647,909,935]" pageId="4" pageNumber="353" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(compare
|
||
<figureCitation id="13252A0CFF89EE7CF4ABBDC4850678AB" box="[767,836,909,935]" captionStart="FIGURE 3" captionStartId="3.[151,250,1983,2008]" captionTargetBox="[564,1023,190,1950]" captionTargetId="figure@3.[555,1032,181,1958]" captionTargetPageId="3" captionText="FIGURE 3. Female genitalia of Toleria vietnamica sp. nov. Genital preparation № OG–027-2018. Scale bar: 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3907598" httpUri="https://zenodo.org/record/3907598/files/figure.png" pageId="4" pageNumber="353">Figs 3</figureCitation>
|
||
and 4 with fig.
|
||
<quantity id="4CE69B6CFF89EE7CF5A1BDC4826C78AB" box="[1013,1070,909,935]" metricMagnitude="-1" metricUnit="m" metricValue="9.652" pageId="4" pageNumber="353" unit="in" value="38.0">38 in</quantity>
|
||
<bibRefCitation id="EF8F4B78FF89EE7CF261BDC4833278AB" author="Gorbunov, O. G. & Arita, Y." box="[1077,1392,909,935]" pageId="4" pageNumber="353" pagination="9 - 51" refId="ref8034" refString="Gorbunov, O. G. & Arita, Y. (2001) A revision of Felix Bryk's clearwing moth types (Lepidoptera, Sesiidae), at the Naturhisto- riska Riksmuseet in Stockholm, Sweden. In: Gorbunov, O. G. (Ed.), Melittia, a Lepidopterological Almanac, 1, pp. 9 - 51." type="journal article" year="2001">Gorbunov & Arita 2001: 40</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA13689FF89EE7CF693BDF8875A7E92" blockId="4.[151,1437,152,1759]" pageId="4" pageNumber="353">
|
||
From
|
||
<taxonomicName id="4C1E4D0AFF89EE7CF759BDFB840378C0" authority="(Strand, 1916)" baseAuthorityName="Strand" baseAuthorityYear="1916" box="[269,577,945,972]" class="Insecta" family="Sesiidae" genus="Toleria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="353" phylum="Arthropoda" rank="species" species="aegerides">
|
||
<emphasis id="B96AEA9BFF89EE7CF759BDFB87D078C7" box="[269,402,945,971]" italics="true" pageId="4" pageNumber="353">T aegerides</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8F4B78FF89EE7CF7F6BDF8847B78C0" author="Strand, E." box="[418,569,945,972]" pageId="4" pageNumber="353" pagination="34 - 49" refId="ref9427" refString="Strand, E. (1916) H. Sauter's Formosa-Ausbeute: Noctuidae p. p. (Agaristinae, Macrobrochis), Aganaidae, Saturniidae, Uraniidae, Cossidae, Callidulidae und Aegeriidae. Archiv fur Naturgeschichte, Abteilung A, 81 (8), 34 - 49." type="journal article" year="1916">Strand, 1916</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, this new species clearly differs by the colouration of the antenna (antenna yellow-orange dorsally and light brown ventrally in
|
||
<emphasis id="B96AEA9BFF89EE7CF4B4BD9F85C478E3" box="[736,902,981,1007]" italics="true" pageId="4" pageNumber="353">
|
||
<taxonomicName id="4C1E4D0AFF89EE7CF4B4BD9F852A78E3" baseAuthorityName="Strand" baseAuthorityYear="1916" box="[736,872,981,1007]" class="Insecta" family="Sesiidae" genus="Toleria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="353" phylum="Arthropoda" rank="species" species="aegerides">T. aegerides</taxonomicName>
|
||
vs
|
||
</emphasis>
|
||
. antenna brown with a dense admixture of light brown scales dorsally on basal half in
|
||
<taxonomicName id="4C1E4D0AFF89EE7CF417BDB384987F1F" authority="Gorbunov & Arita, 2020" authorityName="Gorbunov & Arita" authorityYear="2020" box="[579,730,1018,1043]" class="Insecta" family="Sesiidae" genus="Toleria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="353" phylum="Arthropoda" rank="species" species="vietnamica" status="sp. nov.">
|
||
<emphasis id="B96AEA9BFF89EE7CF417BDB384987F1F" box="[579,730,1018,1043]" italics="true" pageId="4" pageNumber="353">T. vietnamica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96AEA9BFF89EE7CF4B5BDB085787F1F" bold="true" box="[737,826,1017,1043]" pageId="4" pageNumber="353">
|
||
<taxonomicNameLabel id="A25957E0FF89EE7CF4B5BDB085787F1F" box="[737,826,1017,1043]" pageId="4" pageNumber="353" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
), tegula (anteriorly dark brown with bronze sheen and an admixture of individual light brown scales, posteriorly pale yellow to yellow in the species compared
|
||
<emphasis id="B96AEA9BFF89EE7CF366BA57830B7F3B" box="[1330,1353,1054,1079]" italics="true" pageId="4" pageNumber="353">vs</emphasis>
|
||
. tegula dark brown with bronze-purple sheen, densely covered with light brown scales masking background colouration and with a large yellow-orange spot at base of forewing in
|
||
<taxonomicName id="4C1E4D0AFF89EE7CF510BA2F859F7F73" authority="Gorbunov & Arita, 2020" authorityName="Gorbunov & Arita" authorityYear="2020" box="[836,989,1126,1151]" class="Insecta" family="Sesiidae" genus="Toleria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="353" phylum="Arthropoda" rank="species" species="vietnamica" status="sp. nov.">
|
||
<emphasis id="B96AEA9BFF89EE7CF510BA2F859F7F73" box="[836,989,1126,1151]" italics="true" pageId="4" pageNumber="353">T. vietnamica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96AEA9BFF89EE7CF5B2BA2C82007F73" bold="true" box="[998,1090,1125,1151]" pageId="4" pageNumber="353">
|
||
<taxonomicNameLabel id="A25957E0FF89EE7CF5B2BA2C82007F73" box="[998,1090,1125,1151]" pageId="4" pageNumber="353" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
), and abdomen (dorsally dark brown with bronze sheen and an admixture of individual light brown scales; tergite 1 with a narrow yellow stripe distally; tergites 3 and 4 each densely mixed with yellow scales; tergites 5 and 6 each mixed with yellow scales medio-distally; anal tuft light brown-yellow medially and yellow laterally in
|
||
<emphasis id="B96AEA9BFF89EE7CF5A9BA9882E77FE6" box="[1021,1189,1232,1258]" italics="true" pageId="4" pageNumber="353">
|
||
<taxonomicName id="4C1E4D0AFF89EE7CF5A9BA9882C47FE6" baseAuthorityName="Strand" baseAuthorityYear="1916" box="[1021,1158,1232,1258]" class="Insecta" family="Sesiidae" genus="Toleria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="353" phylum="Arthropoda" rank="species" species="aegerides">T. aegerides</taxonomicName>
|
||
vs
|
||
</emphasis>
|
||
. black with dark blue sheen; dorsally tergites 1 and 2 each densely covered with chestnut-brown elongate scales; tergite 1 with a large dark yellow to yellow-orange spot lateral-distally; tergite 2 with a narrow dark yellow to yellow-orange stripe distally; anal tuft dark gray with bronze sheen in
|
||
<taxonomicName id="4C1E4D0AFF89EE7CF40FBB7584B37E59" authority="Gorbunov & Arita, 2020" authorityName="Gorbunov & Arita" authorityYear="2020" box="[603,753,1340,1365]" class="Insecta" family="Sesiidae" genus="Toleria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="353" phylum="Arthropoda" rank="species" species="vietnamica" status="sp. nov.">
|
||
<emphasis id="B96AEA9BFF89EE7CF40FBB7584B37E59" box="[603,753,1340,1365]" italics="true" pageId="4" pageNumber="353">T. vietnamica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96AEA9BFF89EE7CF4ACBB7585137E5A" bold="true" box="[760,849,1340,1366]" pageId="4" pageNumber="353">
|
||
<taxonomicNameLabel id="A25957E0FF89EE7CF4ACBB7585137E5A" box="[760,849,1340,1366]" pageId="4" pageNumber="353" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
). Besides this, these species can be distinguished by the shape of the corpus bursae and signum of the female genitalia (compere
|
||
<figureCitation id="13252A0CFF89EE7CF5B1BB1682647E75" box="[997,1062,1375,1401]" captionStart="FIGURE 3" captionStartId="3.[151,250,1983,2008]" captionTargetBox="[564,1023,190,1950]" captionTargetId="figure@3.[555,1032,181,1958]" captionTargetPageId="3" captionText="FIGURE 3. Female genitalia of Toleria vietnamica sp. nov. Genital preparation № OG–027-2018. Scale bar: 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3907598" httpUri="https://zenodo.org/record/3907598/files/figure.png" pageId="4" pageNumber="353">Fig. 3</figureCitation>
|
||
with fig.
|
||
<quantity id="4CE69B6CFF89EE7CF2C5BB1682887E75" box="[1169,1226,1375,1401]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="4" pageNumber="353" unit="in" value="18.0">18 in</quantity>
|
||
<bibRefCitation id="EF8F4B78FF89EE7CF29ABB1687497E92" author="Arita, Y. & Gorbunov, O. G." pageId="4" pageNumber="353" pagination="141 - 165" refId="ref7472" refString="Arita, Y. & Gorbunov, O. G. (1998) A revision of Embrik Strand's clearwing moth types (Lepidoptera: Sesiidae) from Taiwan. Chinese Journal of Entomology, 18, 141 - 165." type="journal article" year="1998">Arita & Gorbunov 1998: 161</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA13689FF89EE7CF693BBEE83D57D20" blockId="4.[151,1437,152,1759]" pageId="4" pageNumber="353">
|
||
From
|
||
<taxonomicName id="4C1E4D0AFF89EE7CF759BBEE87C17ECC" authorityName="Walker" authorityYear="1864" box="[269,387,1447,1472]" class="Insecta" family="Sesiidae" genus="Toleria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="353" phylum="Arthropoda" rank="species" species="sinensis">
|
||
<emphasis id="B96AEA9BFF89EE7CF759BBEE87C17ECC" box="[269,387,1447,1472]" italics="true" pageId="4" pageNumber="353">T. sinensis</emphasis>
|
||
</taxonomicName>
|
||
(Walker, 1865 [“1864”],
|
||
<taxonomicName id="4C1E4D0AFF89EE7CF4FCBBEE85027ECC" authority="Gorbunov & Arita, 2020" authorityName="Gorbunov & Arita" authorityYear="2020" box="[680,832,1447,1472]" class="Insecta" family="Sesiidae" genus="Toleria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="353" phylum="Arthropoda" rank="species" species="vietnamica" status="sp. nov.">
|
||
<emphasis id="B96AEA9BFF89EE7CF4FCBBEE85027ECC" box="[680,832,1447,1472]" italics="true" pageId="4" pageNumber="353">T. vietnamica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96AEA9BFF89EE7CF51DBBEE85E67ECD" bold="true" box="[841,932,1447,1473]" pageId="4" pageNumber="353">
|
||
<taxonomicNameLabel id="A25957E0FF89EE7CF51DBBEE85E67ECD" box="[841,932,1447,1473]" pageId="4" pageNumber="353" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is clearly separable by the coloration of the abdomen (dorsally tergites 4–6 each yellow with a narrow black stripe anteriorly in
|
||
<taxonomicName id="4C1E4D0AFF89EE7CF267BB8282E47EE8" authorityName="Walker" authorityYear="1864" box="[1075,1190,1483,1508]" class="Insecta" family="Sesiidae" genus="Toleria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="353" phylum="Arthropoda" rank="species" species="sinensis">
|
||
<emphasis id="B96AEA9BFF89EE7CF267BB8282E47EE8" box="[1075,1190,1483,1508]" italics="true" pageId="4" pageNumber="353">T. sinensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B96AEA9BFF89EE7CF2E6BB82828B7EE8" box="[1202,1225,1483,1508]" italics="true" pageId="4" pageNumber="353">vs</emphasis>
|
||
. dorsally tergites 1 and 2 each densely covered with chestnut-brown elongate scales; tergite 1 with a large dark yellow to yellow-orange spot lateral-distally; tergite 2 with a narrow dark yellow to yellow-orange stripe distally in
|
||
<taxonomicName id="4C1E4D0AFF89EE7CF2C4B85A83647D20" authority="Gorbunov & Arita, 2020" authorityName="Gorbunov & Arita" authorityYear="2020" box="[1168,1318,1555,1580]" class="Insecta" family="Sesiidae" genus="Toleria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="353" phylum="Arthropoda" rank="species" species="vietnamica" status="sp. nov.">
|
||
<emphasis id="B96AEA9BFF89EE7CF2C4B85A83647D20" box="[1168,1318,1555,1580]" italics="true" pageId="4" pageNumber="353">T. vietnamica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96AEA9BFF89EE7CF379B85B83C57D20" bold="true" box="[1325,1415,1554,1580]" pageId="4" pageNumber="353">
|
||
<taxonomicNameLabel id="A25957E0FF89EE7CF379B85B83C57D20" box="[1325,1415,1554,1580]" pageId="4" pageNumber="353" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA13689FF89EE7CF693B87C86AD7D78" blockId="4.[151,1437,152,1759]" pageId="4" pageNumber="353">
|
||
<emphasis id="B96AEA9BFF89EE7CF693B87C87097D5C" bold="true" box="[199,331,1589,1616]" pageId="4" pageNumber="353">Bionomics.</emphasis>
|
||
The larval host plant of this new species is unknown. The
|
||
<typeStatus id="54A5882BFF89EE7CF58AB87C82027D43" box="[990,1088,1589,1615]" pageId="4" pageNumber="353" type="holotype">holotype</typeStatus>
|
||
was collected in the beginning of May.
|
||
</paragraph>
|
||
<paragraph id="8BA13689FF89EE7CF693B83587E57D9B" blockId="4.[151,1437,152,1759]" box="[199,423,1660,1687]" pageId="4" pageNumber="353">
|
||
<emphasis id="B96AEA9BFF89EE7CF693B83587687D9B" bold="true" box="[199,298,1660,1687]" pageId="4" pageNumber="353">Habitat.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3046502FF89EE7CF693B8E9828A7DB7" box="[199,1224,1696,1723]" pageId="4" pageNumber="353" type="distribution">
|
||
<paragraph id="8BA13689FF89EE7CF693B8E9828A7DB7" blockId="4.[151,1437,152,1759]" box="[199,1224,1696,1723]" pageId="4" pageNumber="353">
|
||
<emphasis id="B96AEA9BFF89EE7CF693B8E987237DB7" bold="true" box="[199,353,1696,1723]" pageId="4" pageNumber="353">Distribution.</emphasis>
|
||
Known from Ba Bể National Park, Bẳc Kan Province in North
|
||
<collectingCountry id="F3097619FF89EE7CF27FB8E882C87DB7" box="[1067,1162,1697,1723]" name="Vietnam" pageId="4" pageNumber="353">Vietnam</collectingCountry>
|
||
only.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3046502FF89EE7CF693B88C85A27DD3" box="[199,992,1733,1759]" pageId="4" pageNumber="353" type="etymology">
|
||
<paragraph id="8BA13689FF89EE7CF693B88C85A27DD3" blockId="4.[151,1437,152,1759]" box="[199,992,1733,1759]" pageId="4" pageNumber="353">
|
||
<emphasis id="B96AEA9BFF89EE7CF693B88C870C7DD3" bold="true" box="[199,334,1733,1759]" pageId="4" pageNumber="353">Etymology.</emphasis>
|
||
This new species is named after
|
||
<collectingCountry id="F3097619FF89EE7CF494B88C85607DD3" box="[704,802,1733,1759]" name="Vietnam" pageId="4" pageNumber="353">Vietnam</collectingCountry>
|
||
, the
|
||
<emphasis id="B96AEA9BFF89EE7CF502B88F859B7DD3" box="[854,985,1734,1759]" italics="true" pageId="4" pageNumber="353">terra typica</emphasis>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |