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<mods:title id="24D9C5B5FAF717C4429A07B52CDA5C81">Taxonomic and Biogeographical Status of Three Species of the Spider Crabs of the Genus Acanthonyx Latreille, 1828 (Majoidea: Epialtidae) as Determined by DNA Barcoding and Morphological Analyses Along the Western Atlantic</mods:title>
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H. 
<bibRefCitation id="EFA069D8FFE7FF9B6F8EA6C4FD5BFEA0" author="Milne Edwards H." box="[478,738,253,278]" pageId="6" pageNumber="7" pagination="1 - 468" refId="ref13717" refString="Milne Edwards H. 1834. Histoire naturelle des Crustaces, comprenant l'anatomie, la physiologie et la classification de ces animaux, 1: 1 - 468." type="journal article" year="1834">Milne Edwards, 1834</bibRefCitation>
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(
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)
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<taxonomicName id="4C316FAAFFE7FF9B6EFEA761FDE2FEDA" authority="Petiver, 1712" authorityName="Petiver" authorityYear="1712" box="[174,603,344,364]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="subSpecies" species="muricatus" subSpecies="compressum">
<emphasis id="B945C83BFFE7FF9B6EFEA761FE6CFEDA" box="[174,469,344,364]" italics="true" pageId="6" pageNumber="7">Cancer muricatus compressum</emphasis>
<bibRefCitation id="EFA069D8FFE7FF9B6F8BA761FDE2FEDA" author="Petiver J." box="[475,603,344,364]" pageId="6" pageNumber="7" pagination="1 - 3" refId="ref13959" refString="Petiver J. 1712. Pteri-graphia Americana, icones continens... Filicum variarum specierum... nec non Muscos, Lichenes, Fungos, Corallia, Spongias... Crustacea, Testacea, aliaque animalia &amp; c.: 1 - 3, 20 pls. (London)." type="book chapter" year="1712">Petiver, 1712</bibRefCitation>
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: plate 20: Fig. 8.
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<emphasis id="B945C83BFFE7FF9B6EFEA74AFECCFE31" box="[174,373,371,391]" italics="true" pageId="6" pageNumber="7">Acanthonyx petiverii</emphasis>
H. 
<bibRefCitation id="EFA069D8FFE7FF9B6FCCA74AFD1CFE31" author="Milne Edwards H." box="[412,677,371,391]" pageId="6" pageNumber="7" pagination="1 - 468" refId="ref13717" refString="Milne Edwards H. 1834. Histoire naturelle des Crustaces, comprenant l'anatomie, la physiologie et la classification de ces animaux, 1: 1 - 468." type="journal article" year="1834">Milne Edwards, 1834: 343</bibRefCitation>
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, plate 15: Fig.s 6-8; 
<treatmentCitation id="0A903238FFE7FF9B6F1CA7B4FE64FE17" author="Bell T." box="[332,477,397,417]" page="62" pageId="6" pageNumber="7" year="1841">
<bibRefCitation id="EFA069D8FFE7FF9B6F1CA7B4FE64FE17" author="Bell T." box="[332,477,397,417]" pageId="6" pageNumber="7" pagination="39 - 66" refId="ref12408" refString="Bell T. 1841. Some account of the Crustacea of the coasts of South America, with descriptions of new genera and species: founded principally on the collections obtained by Mr. Cuming and Mr. Miller. Trans ZooL Soc 2 (1): 39 - 66." type="journal article" year="1841">Bell, 1841: 62</bibRefCitation>
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; 
<treatmentCitation id="0A903238FFE7FF9B6FB9A7B4FD19FE17" author="Moreira C." box="[489,672,397,417]" page="66" pageId="6" pageNumber="7" year="1901">
<bibRefCitation id="EFA069D8FFE7FF9B6FB9A7B4FD19FE17" author="Moreira C." box="[489,672,397,417]" pageId="6" pageNumber="7" pagination="1 - 151" refId="ref13832" refString="Moreira C. 1901. Contribuicoes para o conhecimento da fauna brazileira. Crustaceos do Brazil. Arq Mus Nac R Janeiro 11: 1 - 151." type="journal article" year="1901">Moreira, 1901: 66</bibRefCitation>
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; 
<treatmentCitation id="0A903238FFE7FF9B6CFCA7B4FEEAFE0A" author="Rathbun MJ" page="142" pageId="6" pageNumber="7" year="1925">
<bibRefCitation id="EFA069D8FFE7FF9B6CFCA7B4FEEAFE0A" author="Rathbun MJ" pageId="6" pageNumber="7" pagination="1 - 613" refId="ref14038" refString="Rathbun MJ. 1925. The spider crabs of America. Bull U S Natl Mus 129: 1 - 613." type="journal article" year="1925">Rathbun, 1925: 142</bibRefCitation>
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, Fig. 52, plate 44, plate 222: 
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-6; 
<treatmentCitation id="0A903238FFE7FF9B6EB3A7FAFE2FFE61" author="Rathbun MJ" box="[227,406,451,471]" page="13" pageId="6" pageNumber="7" year="1933">
<bibRefCitation id="EFA069D8FFE7FF9B6EB3A7FAFE2FFE61" author="Rathbun MJ" box="[227,406,451,471]" pageId="6" pageNumber="7" pagination="1 - 121" refId="ref14060" refString="Rathbun MJ. 1933. Brachyuran crabs of Porto Rico and the Virgin Islands. In: Scientific Survey of Porto Rico and the Virgin Islands. New York Acad Sci 15 (1): 1 - 121." type="journal article" year="1933">Rathbun, 1933: 13</bibRefCitation>
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, Fig. 11; 
<treatmentCitation id="0A903238FFE7FF9B6FBBA7FBFD37FE61" author="Garth JS" box="[491,654,450,471]" page="376" pageId="6" pageNumber="7" year="1946">
<bibRefCitation id="EFA069D8FFE7FF9B6FBBA7FBFD37FE61" author="Garth JS" box="[491,654,450,471]" pageId="6" pageNumber="7" pagination="341 - 601" refId="ref12857" refString="Garth JS. 1946. Littoral brachyuran fauna of the Galapagos Archipelago. Allan Hancock Pac Exped 5 (10): 341 - 601." type="journal article" year="1946">Garth, 1946: 376</bibRefCitation>
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, Fig. 4, plate 63; 
<treatmentCitation id="0A903238FFE7FF9B6F57A7E4FE12FE47" author="Garth JS" box="[263,427,477,497]" page="223" pageId="6" pageNumber="7" year="1958">
<bibRefCitation id="EFA069D8FFE7FF9B6F57A7E4FE12FE47" author="Garth JS" box="[263,427,477,497]" pageId="6" pageNumber="7" pagination="1 - 854" refId="ref12883" refString="Garth JS. 1958. Brachyura of the Pacific coast of America; Oxyrhyncha. Allan Hancock Pac Exped 21 (1 - 2): 1 - 854." type="journal article" year="1958">Garth, 1958: 223</bibRefCitation>
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, plate 25: 
<figureCitation id="130A08ACFFE7FF9B6C5DA7E4FDFDFE47" box="[525,580,477,497]" captionStart="Fig" captionStartId="7.[136,170,1814,1834]" captionTargetBox="[136,1413,500,1788]" captionTargetId="figure-228@7.[136,1413,443,1851]" captionTargetPageId="7" captionText="Fig. 2. Overview of samples of Acanthonyx Latreille, 1828 and some parts of their body. “A. petiverii” H. Milne Edwards, 1834, male CL 13.5 mm (CCDB 1063). (A) dorsal view, (B) ventral view; “A. dissimulatus” Coelho, 1993 male CL 12.2 mm (CCDB 1421) (C) dorsal view, (D) ventral view; (E) bifid rostrum, male CL 24.5 mm (CCDB 3949); (F) dorsal view of male left cheliped, CL 24.5 mm (CCDB 3949); (G) ventral view of female right cheliped, CL 8.3 mm (CCDB 3423); (H) subchela of the propodus of the last three ambulatory pereopods, male CL 24.5 mm (CCDB 3949). Scale bars: 5 mm (A, B, C, D, F); 2 mm (E, G); 0.6 mm (H). The species names in inverted commas refer to the original identification." figureDoi="http://doi.org/10.5281/zenodo.8060307" httpUri="https://zenodo.org/record/8060307/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
; 
<treatmentCitation id="0A903238FFE7FF9B6C1FA7E4FF46FDBA" author="Fausto-Filho J." page="33" pageId="6" pageNumber="7" year="1966">
<bibRefCitation id="EFA069D8FFE7FF9B6C1FA7E4FF46FDBA" author="Fausto-Filho J." pageId="6" pageNumber="7" pagination="31 - 37" refId="ref12759" refString="Fausto-Filho J. 1966. Primeira contribuicao ao inventario dos crustaceos decapodos marinhos do Nordeste brasileiro. Arq Est Biol Mar Univ Fed Ceara 6 (1): 31 - 37." type="journal article" year="1966">Fausto-Filho, 1966: 33</bibRefCitation>
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; 
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; 
<treatmentCitation id="0A903238FFE7FF9B6C9DA7C1FE20FD91" author="Abele LG &amp; Kim W." page="43" pageId="6" pageNumber="7" year="1986">
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, 495 (key); 
<treatmentCitation id="0A903238FFE7FF9B6C5BA42BFE80FDF7" author="Hernandez-Aguilera JL &amp; Toral-Alamzan RE &amp; Ruiz-Nuno JA" page="60" pageId="6" pageNumber="7" year="1997">
<bibRefCitation id="EFA069D8FFE7FF9B6C5BA42BFE80FDF7" author="Hernandez-Aguilera JL &amp; Toral-Alamzan RE &amp; Ruiz-Nuno JA" pageId="6" pageNumber="7" refId="ref13058" refString="Hernandez-Aguilera JL, Toral-Alamzan RE, Ruiz-Nuno JA. 1997. Especies catalogadas de crustaceos estomatopodos y decapodos para el Golfo de Mexico, Rio Bravo, Tamaulipas a Progreso, Yucatan. Secretaria de Marina-Comision Nacional para el Conocimiento y Uso de la Biodiversidad, Mexico." type="book" year="1997">Hernández-Aguilera et al., 1997: 60</bibRefCitation>
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, Fig. d, plate 3; 
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<emphasis id="B945C83BFFE7FF9B6EFEA471FE21FDEA" box="[174,408,584,604]" italics="true" pageId="6" pageNumber="7">Acanthonyx emarginatus</emphasis>
H. 
<bibRefCitation id="EFA069D8FFE7FF9B6FEAA471FD56FDEA" author="Milne Edwards H &amp; Lucas H." box="[442,751,584,604]" pageId="6" pageNumber="7" pagination="1 - 39" refId="ref13746" refString="Milne Edwards H, Lucas H. 1843. Crustaces. In: A. d'Orbigny, Voyage dans l'Amerique meridionale (le Bresil, la republique orientale de l'Uruguay, la republique Argentine, La Patagonie, la republique du Chile, la republique de Bolivia, la republique du Perou), execute pendant les annees 1826, 1827, 1828, 1829, 1830, 1831, 1832 et 1833. Bertrand, Paris 6 (1): 1 - 39." type="journal article" year="1843">Milne Edwards &amp; Lucas, 1843: 9</bibRefCitation>
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<emphasis id="B945C83BFFE7FF9B6EFEA45AFEE6FDC1" box="[174,351,611,631]" italics="true" pageId="6" pageNumber="7">Acanthonyx debilis</emphasis>
<bibRefCitation id="EFA069D8FFE7FF9B6F36A45AFDBEFDC1" author="Dana JD" box="[358,519,611,631]" pageId="6" pageNumber="7" pagination="268 - 274" refId="ref12610" refString="Dana JD. 1851. Conspectus Crustaceorum quae in orbis Terrarum circumnavigatione, Carolo Wilkes e Classe Reipublicae Foederatae Duce, lexit et descripsit J. A. Dana - Pars VI. Am J Sci Arts, ser. 2, 11 (32): 268 - 274." type="journal article" year="1851">Dana, 1851: 272</bibRefCitation>
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<paragraph id="8B8E1429FFE7FF9B6EFEA444FDB2FD27" blockId="6.[174,776,344,871]" box="[174,523,637,657]" pageId="6" pageNumber="7">
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<taxonomicName id="4C316FAAFFE7FF9B6EFEA444FDBEFD27" ID-CoL="58b1bbf0-f5d9-4386-86f3-aaf791db2350" authority="Dana, 1851: 273" authorityName="Dana" authorityPageNumber="273" authorityYear="1851" box="[174,519,637,657]" class="Malacostraca" family="Epialtidae" genus="Peltinia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="scutiformis">
<emphasis id="B945C83BFFE7FF9B6EFEA444FED9FD27" box="[174,352,637,657]" italics="true" pageId="6" pageNumber="7">Peltinia scutiformis</emphasis>
<bibRefCitation id="EFA069D8FFE7FF9B6F36A444FDBEFD27" author="Dana JD" box="[358,519,637,657]" pageId="6" pageNumber="7" pagination="268 - 274" refId="ref12610" refString="Dana JD. 1851. Conspectus Crustaceorum quae in orbis Terrarum circumnavigatione, Carolo Wilkes e Classe Reipublicae Foederatae Duce, lexit et descripsit J. A. Dana - Pars VI. Am J Sci Arts, ser. 2, 11 (32): 268 - 274." type="journal article" year="1851">Dana, 1851: 273</bibRefCitation>
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<paragraph id="8B8E1429FFE7FF9B6EFEA4A1FE15FD71" blockId="6.[174,776,344,871]" pageId="6" pageNumber="7">
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<taxonomicName id="4C316FAAFFE7FF9B6EFEA4A1FE5BFD1A" ID-CoL="64GGR" authority="Dana, 1852" authorityName="Dana" authorityYear="1852" box="[174,482,664,684]" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="B945C83BFFE7FF9B6EFEA4A1FED5FD1A" box="[174,364,664,684]" italics="true" pageId="6" pageNumber="7">Acanthonyx simplex</emphasis>
<bibRefCitation id="EFA069D8FFE7FF9B6F23A4A1FE5BFD1A" author="Dana JD" box="[371,482,664,684]" pageId="6" pageNumber="7" pagination="685" refId="ref12661" refString="Dana JD. 1852. Crustacea, Part 1. In: United States Exploring Expedition during the years 1838, 1839, 1840, 1841, 1842 under the command of Charles Wilkes. U. S. N. Philadelphia 13, p. 685." type="journal article" year="1852">Dana, 1852</bibRefCitation>
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- 
<treatmentCitation id="0A903238FFE7FF9B6FA7A4A1FCBEFD1A" author="Emparanza EJM &amp; Guzman GL &amp; Ng PKL" box="[503,775,664,684]" page="534" pageId="6" pageNumber="7" year="2007">
<bibRefCitation id="EFA069D8FFE7FF9B6FA7A4A1FCBEFD1A" author="Emparanza EJM &amp; Guzman GL &amp; Ng PKL" box="[503,775,664,684]" pageId="6" pageNumber="7" pagination="533 - 543" refId="ref12709" refString="Emparanza EJM, Guzman GL, Ng PKL. 2007. A redescription of Acanthonyx petiverii H. Milne Edwards, 1834, and designation of a neotype for Acanthonyx simplex Dana, 1852 (Brachyura, Majidae). Crustaceana 80 (5): 533 - 543." type="journal article" year="2007">Emparanza et al., 2007: 534</bibRefCitation>
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(
<tableCitation id="C6B32192FFE7FF9B6EB8A48AFE97FD71" box="[232,302,691,711]" captionStart="Table 1" captionStartId="2.[174,241,510,535]" captionTargetBox="[178,1449,610,1925]" captionText="Table 1. Morphological characters used to compare the species Acanthonyx dissimulatus, A. petiverii and A. scutiformis" httpUri="http://table.plazi.org/id/DF4E44A1FFE3FF9F6EFEA7C7FE94FD8C" pageId="6" pageNumber="7" tableUuid="DF4E44A1FFE3FF9F6EFEA7C7FE94FD8C">Table 1</tableCitation>
), 535, 
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.
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</paragraph>
<paragraph id="8B8E1429FFE7FF9B6EFEA4F4FD45FD57" blockId="6.[174,776,344,871]" box="[174,764,717,737]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="AB213307FFE7FF9B6EFEA4F4FD45FD57" box="[174,764,717,737]" pageId="6" pageNumber="7">
<treatmentCitation id="0A903238FFE7FF9B6EFEA4F4FDD8FD57" author="Kinahan JR" box="[174,609,717,737]" page="334" pageId="6" pageNumber="7" year="1857">
<taxonomicName id="4C316FAAFFE7FF9B6EFEA4F4FDD8FD57" ID-CoL="64GFX" authority="Kinahan, 1857: 334" authorityName="Kinahan" authorityPageNumber="334" authorityYear="1857" box="[174,609,717,737]" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="concamerata">
<emphasis id="B945C83BFFE7FF9B6EFEA4F4FE27FD57" box="[174,414,717,737]" italics="true" pageId="6" pageNumber="7">Acanthonyx concamerata</emphasis>
<bibRefCitation id="EFA069D8FFE7FF9B6FF5A4F4FDD8FD57" author="Kinahan JR" box="[421,609,717,737]" pageId="6" pageNumber="7" pagination="328" refId="ref13242" refString="Kinahan JR. 1857. Remarks on Crustacea collected in Peru, the high seas, and South Australia; with descriptions of undescribed species. Journ Roy Dublin Soc 1: 328 352." type="journal article" year="1857">Kinahan, 1857: 334</bibRefCitation>
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, plate 14: 
<figureCitation id="130A08ACFFE7FF9B6C91A4F4FD41FD57" box="[705,760,717,737]" captionStart="Fig" captionStartId="5.[134,168,1005,1025]" captionTargetBox="[86,1464,388,980]" captionTargetId="figure-1@5.[59,1502,280,1032]" captionTargetPageId="5" captionText="Fig. 1. Genetic comparison between species of the genus Acanthonyx Latreille, 1828. Phylogram obtained from Maximum Likelihood analysis of barcoding COI sequences. The numbers are significance values for 1000 bootstraps; values ≤ 50% are not shown. The species names in inverted commas refer to the original identification (*GenBank sequences: JQ305885; JQ305884, EU682455, EU682454, EU682857, EU682856; A: Central and South America; ATL: Atlantic coast; B: North America; BA: Bahia; Br: Brazil; CR: Costa Rica; Mx: Mexico; SP: São Paulo; Pa: Panama; PE: Pernambuco; USA: United States of America; Ve: Venezuela)." figureDoi="http://doi.org/10.5281/zenodo.8060303" httpUri="https://zenodo.org/record/8060303/files/figure.png" pageId="6" pageNumber="7">Fig. 1</figureCitation>
.
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</paragraph>
<paragraph id="8B8E1429FFE7FF9B6EFEA4D1FE84FC87" blockId="6.[174,776,344,871]" pageId="6" pageNumber="7">
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<taxonomicName id="4C316FAAFFE7FF9B6EFEA4D1FE35FD4A" ID-CoL="7QBM5" baseAuthorityName="Dana" baseAuthorityYear="1851" box="[174,396,744,764]" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="scutiformis">
<emphasis id="B945C83BFFE7FF9B6EFEA4D1FE35FD4A" box="[174,396,744,764]" italics="true" pageId="6" pageNumber="7">Acanthonyx scutiformis</emphasis>
</taxonomicName>
- 
<treatmentCitation id="0A903238FFE7FF9B6FF3A4D1FD71FD4A" author="Coelho PA &amp; Torres MFA" box="[419,712,744,764]" page="228" pageId="6" pageNumber="7" year="1993">
<bibRefCitation id="EFA069D8FFE7FF9B6FF3A4D1FD71FD4A" author="Coelho PA &amp; Torres MFA" box="[419,712,744,764]" pageId="6" pageNumber="7" pagination="221 - 241" refId="ref12460" refString="Coelho PA, Torres MFA. 1993. Taxonomia e distribuicao das especies do genero Acanthonyx Latreille, no Brasil (Crustacea: Decapoda: Majidae). Trab Oceanogr Univ Fed Pe 22: 221 - 241." type="journal article" year="1993">Coelho and Torres, 1993: 228</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="0A903238FFE7FF9B6C83A4D1FEFCFCA1" author="Melo GAS" page="171" pageId="6" pageNumber="7" year="1996">
<bibRefCitation id="EFA069D8FFE7FF9B6C83A4D1FEFCFCA1" author="Melo GAS" pageId="6" pageNumber="7" refId="ref13578" refString="Melo GAS. 1996. Manual de identificacao dos Brachyura (caranguejos e siris) do Litoral Brasileiro. Sao Paulo: Editora Pleiade / FAPESP." type="book" year="1996">Melo, 1996: 171</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="0A903238FFE7FF9B6F1EA53AFE55FCA1" author="Melo GAS" box="[334,492,771,791]" page="455" pageId="6" pageNumber="7" year="1998">
<bibRefCitation id="EFA069D8FFE7FF9B6F1EA53AFE55FCA1" author="Melo GAS" box="[334,492,771,791]" pageId="6" pageNumber="7" pagination="455 - 515" refId="ref13605" refString="Melo GAS. 1998. Malacostraca-Eucarida. Brachyura. Oxyrhyncha and Brachyrhyncha. In: Young OS (ed.) Catalogue of Crustacea of Brazil. Rio de Janeiro, Museu Nacional (Serie Livros n ° 6): 455 - 515." type="journal article" year="1998">Melo, 1998: 455</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="0A903238FFE7FF9B6FA4A53AFDC3FCA1" author="Melo GAS" box="[500,634,771,791]" page="4" pageId="6" pageNumber="7" year="2008">
<bibRefCitation id="EFA069D8FFE7FF9B6FA4A53AFDC3FCA1" author="Melo GAS" box="[500,634,771,791]" pageId="6" pageNumber="7" pagination="1 - 22" refId="ref13650" refString="Melo GAS. 2008. The Brachyura (Decapoda) of Ilha Grande Bay, Rio de Janeiro, Brazil. Nauplius 16 (1): 1 - 22." type="journal article" year="2008">Melo, 2008: 4</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="0A903238FFE7FF9B6CD2A53AFE80FC87" author="Teixeira GM &amp; Fransozo V &amp; Cobo VJ &amp; Hiyodo CM" page="89" pageId="6" pageNumber="7" year="2009">
<bibRefCitation id="EFA069D8FFE7FF9B6CD2A53AFE80FC87" author="Teixeira GM &amp; Fransozo V &amp; Cobo VJ &amp; Hiyodo CM" pageId="6" pageNumber="7" pagination="87 - 95" refId="ref14378" refString="Teixeira GM, Fransozo V, Cobo VJ, Hiyodo CM. 2009. Population features of the spider crab Acanthonyx scutiformis (Dana 1851) (Crustacea, Majoidea, Epialtidae) associated with rocky-shore algae from southeastern Brazil. PanamJAS 4 (1): 87 - 95." type="journal article" year="2009">Teixeira et al., 2009: 89</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="8B8E1429FFE7FF9B6EFEA501FEB7FCD1" blockId="6.[174,776,344,871]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="AB213307FFE7FF9B6EFEA501FEB7FCD1" pageId="6" pageNumber="7">
<treatmentCitation id="0A903238FFE7FF9B6EFEA501FDF6FCFA" author="Coelho PA &amp; Torres MFA" box="[174,591,824,844]" page="231" pageId="6" pageNumber="7" year="1993">
<taxonomicName id="4C316FAAFFE7FF9B6EFEA501FDF6FCFA" ID-CoL="7QBLG" authority="Coelho, 1993: 231" authorityName="&quot; Coelho" authorityPageNumber="231" authorityYear="1993" box="[174,591,824,844]" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="dissimulatus">
<emphasis id="B945C83BFFE7FF9B6EFEA501FE21FCFA" box="[174,408,824,844]" italics="true" pageId="6" pageNumber="7">Acanthonyx dissimulatus</emphasis>
<bibRefCitation id="EFA069D8FFE7FF9B6FCEA501FDF6FCFA" author="Coelho PA &amp; Torres MFA" box="[414,591,824,844]" pageId="6" pageNumber="7" pagination="221 - 241" refId="ref12460" refString="Coelho PA, Torres MFA. 1993. Taxonomia e distribuicao das especies do genero Acanthonyx Latreille, no Brasil (Crustacea: Decapoda: Majidae). Trab Oceanogr Univ Fed Pe 22: 221 - 241." type="journal article" year="1993">Coelho, 1993: 231</bibRefCitation>
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, 
<figureCitation id="130A08ACFFE7FF9B6C07A501FD35FCFA" box="[599,652,824,844]" captionStart="Fig" captionStartId="5.[134,168,1005,1025]" captionTargetBox="[86,1464,388,980]" captionTargetId="figure-1@5.[59,1502,280,1032]" captionTargetPageId="5" captionText="Fig. 1. Genetic comparison between species of the genus Acanthonyx Latreille, 1828. Phylogram obtained from Maximum Likelihood analysis of barcoding COI sequences. The numbers are significance values for 1000 bootstraps; values ≤ 50% are not shown. The species names in inverted commas refer to the original identification (*GenBank sequences: JQ305885; JQ305884, EU682455, EU682454, EU682857, EU682856; A: Central and South America; ATL: Atlantic coast; B: North America; BA: Bahia; Br: Brazil; CR: Costa Rica; Mx: Mexico; SP: São Paulo; Pa: Panama; PE: Pernambuco; USA: United States of America; Ve: Venezuela)." figureDoi="http://doi.org/10.5281/zenodo.8060303" httpUri="https://zenodo.org/record/8060303/files/figure.png" pageId="6" pageNumber="7">Fig. 1</figureCitation>
; 
<treatmentCitation id="0A903238FFE7FF9B6CC8A501FEB3FCD1" author="Melo GAS" page="455" pageId="6" pageNumber="7" year="1998">
<bibRefCitation id="EFA069D8FFE7FF9B6CC8A501FEB3FCD1" author="Melo GAS" pageId="6" pageNumber="7" pagination="455 - 515" refId="ref13605" refString="Melo GAS. 1998. Malacostraca-Eucarida. Brachyura. Oxyrhyncha and Brachyrhyncha. In: Young OS (ed.) Catalogue of Crustacea of Brazil. Rio de Janeiro, Museu Nacional (Serie Livros n ° 6): 455 - 515." type="journal article" year="1998">Melo, 1998: 455</bibRefCitation>
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</subSubSection>
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<paragraph id="8B8E1429FFE7FF9B6EBAA5B7FDF5FBF1" blockId="6.[174,776,910,1960]" pageId="6" pageNumber="7">
<emphasis id="B945C83BFFE7FF9B6EBAA5B7FE39FC11" box="[234,384,910,935]" italics="true" pageId="6" pageNumber="7">Type locality</emphasis>
: The Antilles (H. 
<bibRefCitation id="EFA069D8FFE7FF9B6C05A5B7FF57FC71" author="Milne Edwards H." pageId="6" pageNumber="7" pagination="1 - 468" refId="ref13717" refString="Milne Edwards H. 1834. Histoire naturelle des Crustaces, comprenant l'anatomie, la physiologie et la classification de ces animaux, 1: 1 - 468." type="journal article" year="1834">Milne Edwards 1834</bibRefCitation>
; 
<bibRefCitation id="EFA069D8FFE7FF9B6EABA597FE13FC71" author="Rathbun MJ" box="[251,426,942,967]" pageId="6" pageNumber="7" pagination="1 - 613" refId="ref14038" refString="Rathbun MJ. 1925. The spider crabs of America. Bull U S Natl Mus 129: 1 - 613." type="journal article" year="1925">Rathbun 1925</bibRefCitation>
; 
<bibRefCitation id="EFA069D8FFE7FF9B6FE7A597FDFEFC71" author="Garth JS" box="[439,583,942,968]" pageId="6" pageNumber="7" pagination="1 - 854" refId="ref12883" refString="Garth JS. 1958. Brachyura of the Pacific coast of America; Oxyrhyncha. Allan Hancock Pac Exped 21 (1 - 2): 1 - 854." type="journal article" year="1958">Garth 1958</bibRefCitation>
), in the western Atlantic. The 
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of 
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<emphasis id="B945C83BFFE7FF9B6FA1A5F7FDC1FC51" box="[497,632,974,999]" italics="true" pageId="6" pageNumber="7">A. petiverii</emphasis>
</taxonomicName>
(male with 
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) is in the Muséum National d’Histoire Naturelle, Paris, 
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(
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; 
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). Examined (see Discussion).
</paragraph>
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<emphasis id="B945C83BFFE7FF9B6EBAA277FE64FBD1" box="[234,477,1102,1127]" italics="true" pageId="6" pageNumber="7">Material examined</emphasis>
: 
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, 
<collectingRegion id="49F5DACBFFE7FF9B6F68A257FE37FB31" box="[312,398,1134,1159]" country="United States of America" name="Florida" pageId="6" pageNumber="7">Florida</collectingRegion>
, Miami: 
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(CL 
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), 
<date id="FF8F32E9FFE7FF9B6EFEA2B7FEFAFB11" box="[174,323,1166,1191]" pageId="6" pageNumber="7" value="1965-02-22">22/II/1965</date>
, J. Cabrera leg. (
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1152); 
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, Compostela, 
<collectingRegion id="49F5DACBFFE7FF9B6FB6A297FDF0FB71" box="[486,585,1198,1223]" country="Mexico" name="Nayarit" pageId="6" pageNumber="7">Nayarit</collectingRegion>
: 
<specimenCount id="9D37DFA0FFE7FF9B6C0AA297FD74FB71" box="[602,717,1198,1223]" pageId="6" pageNumber="7" type="female">1 female</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE7FF9B6EFEA2F7FEABFB51" box="[174,274,1230,1255]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="6" pageNumber="7" unit="mm" value="7.7">7.7 mm</quantity>
), 
<date id="FF8F32E9FFE7FF9B6F78A2F7FE14FB51" box="[296,429,1230,1255]" pageId="6" pageNumber="7" value="1993-03-11">11/III/1993</date>
, J.L. Villalobos, E. Cadena, M.E. Camacho, F. Álvarez and E. Lira leg. (
<collectionCode id="ED208CECFFE7FF9B6CE7A2D7FCBEFABE" box="[695,775,1262,1288]" pageId="6" pageNumber="7">CNCR</collectionCode>
15302); San Andrés Tuxtla, 
<collectingRegion id="49F5DACBFFE7FF9B6C5EA337FEF8FAF1" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="6" pageNumber="7">Veracruz de Ignacio de la Llave</collectingRegion>
: 
<specimenCount id="9D37DFA0FFE7FF9B6F00A317FE1FFAF1" box="[336,422,1326,1351]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE7FF9B6FB9A317FDF6FAF1" box="[489,591,1326,1351]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="6" pageNumber="7" unit="mm" value="4.2">4.2 mm</quantity>
), 
<date id="FF8F32E9FFE7FF9B6C36A317FCBAFAF1" box="[614,771,1326,1351]" pageId="6" pageNumber="7" value="2007-08-26">26/VIII/2007</date>
, coll. not available (
<collectionCode id="ED208CECFFE7FF9B6FF1A377FE4DFADE" box="[417,500,1358,1384]" pageId="6" pageNumber="7">CNCR</collectionCode>
24952); 
<collectingRegion id="49F5DACBFFE7FF9B6C3BA377FD59FAD1" box="[619,736,1358,1383]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="6" pageNumber="7">Veracruz</collectingRegion>
, El Moro: 
<specimenCount id="9D37DFA0FFE7FF9B6F54A357FEE4FA31" box="[260,349,1390,1415]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE7FF9B6FF2A357FDA2FA31" box="[418,539,1390,1415]" metricMagnitude="-3" metricUnit="m" metricValue="9.35" pageId="6" pageNumber="7" unit="mm" value="9.35">9.35 mm</quantity>
), 
<date id="FF8F32E9FFE7FF9B6C63A357FD7CFA31" box="[563,709,1390,1415]" pageId="6" pageNumber="7" value="1973-06-01">01/VI/1973</date>
, J.A. Rickner leg. (
<collectionCode id="ED208CECFFE7FF9B6F35A3B7FE16FA11" box="[357,431,1422,1447]" country="USA" httpUri="http://grbio.org/cool/ggzt-j23h" name="University of Louisiana at Layafette, Zoological Collection" pageId="6" pageNumber="7">ULLZ</collectionCode>
11303); 
<collectingRegion id="49F5DACBFFE7FF9B6C7EA3B7FD13FA11" box="[558,682,1422,1447]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="6" pageNumber="7">Veracruz</collectingRegion>
, Punta Delgada: 
<specimenCount id="9D37DFA0FFE7FF9B6F71A397FE39FA71" box="[289,384,1454,1479]" pageId="6" pageNumber="7" type="male">3 males</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE7FF9B6FEBA397FDE3FA71" box="[443,602,1454,1479]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" metricValueMax="5.8" metricValueMin="4.8" pageId="6" pageNumber="7" unit="mm" value="5.3" valueMax="5.8" valueMin="4.8">4.8 - 5.8 mm</quantity>
), 
<specimenCount id="9D37DFA0FFE7FF9B6C3DA397FD6DFA71" box="[621,724,1454,1479]" pageId="6" pageNumber="7" type="female">1 female</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE7FF9B6EFEA3F7FEADFA51" box="[174,276,1486,1511]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="6" pageNumber="7" unit="mm" value="4.9">4.9 mm</quantity>
), 
<date id="FF8F32E9FFE7FF9B6F7AA3F7FE1DFA51" box="[298,420,1486,1511]" pageId="6" pageNumber="7" value="2002-01-07">07/I/2002</date>
, D.L. Felder, R. Robles and T. Rodriguez leg. (
<collectionCode id="ED208CECFFE7FF9B6FF0A3D7FE51F9B1" box="[416,488,1518,1543]" country="USA" httpUri="http://grbio.org/cool/ggzt-j23h" name="University of Louisiana at Layafette, Zoological Collection" pageId="6" pageNumber="7">ULLZ</collectionCode>
6163); 
<collectingCountry id="F32654B9FFE7FF9B6C02A3D7FCBBF9BE" box="[594,770,1518,1544]" name="Costa Rica" pageId="6" pageNumber="7">COSTA RICA</collectingCountry>
, Isla Murciélago: 2 ovigerous females (CL 7.0 and 
<quantity id="4CC9B9CCFFE7FF9B6EFEA017FEAEF9F1" box="[174,279,1582,1607]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="6" pageNumber="7" unit="mm" value="8.6">8.6 mm</quantity>
), 
<date id="FF8F32E9FFE7FF9B6F60A017FE03F9F1" box="[304,442,1582,1607]" pageId="6" pageNumber="7" value="1999-05-08">08/V/1999</date>
, coll. not available (UCR 
<date id="FF8F32E9FFE7FF9B6EFEA077FEB5F9D1" box="[174,268,1614,1639]" bridgedPair="-" pageId="6" pageNumber="7" value="2295-02">2295-II</date>
); 
<collectingCountry id="F32654B9FFE7FF9B6F72A077FE20F9D1" box="[290,409,1614,1639]" name="Panama" pageId="6" pageNumber="7">PANAMA</collectingCountry>
, 
<collectingRegion id="49F5DACBFFE7FF9B6FF8A077FDD2F9D1" box="[424,619,1614,1639]" country="Panama" name="Bocas del Toro" pageId="6" pageNumber="7">Bocas del Toro</collectingRegion>
: 
<specimenCount id="9D37DFA0FFE7FF9B6C2AA077FD69F9D1" box="[634,720,1614,1639]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE7FF9B6EFEA057FEAFF931" box="[174,278,1646,1671]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="6" pageNumber="7" unit="mm" value="5.2">5.2 mm</quantity>
), 
<date id="FF8F32E9FFE7FF9B6F7EA057FE69F931" box="[302,464,1646,1671]" pageId="6" pageNumber="7" value="2004-08-09">09/VIII/2004</date>
, D.L. Felder leg. (
<collectionCode id="ED208CECFFE7FF9B6CEFA057FCBFF931" box="[703,774,1646,1671]" country="USA" httpUri="http://grbio.org/cool/ggzt-j23h" name="University of Louisiana at Layafette, Zoological Collection" pageId="6" pageNumber="7">ULLZ</collectionCode>
10755); 
<collectingRegion id="49F5DACBFFE7FF9B6F49A0B7FE64F911" box="[281,477,1678,1703]" country="Panama" name="Bocas del Toro" pageId="6" pageNumber="7">Bocas del Toro</collectingRegion>
, Playa Paunch: 
<specimenCount id="9D37DFA0FFE7FF9B6CE0A0B7FCBCF911" box="[688,773,1678,1703]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE7FF9B6EB2A097FEEBF971" box="[226,338,1710,1735]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="6" pageNumber="7" unit="mm" value="13.5">13.5 mm</quantity>
), 1 ovigerous female (CL 
<quantity id="4CC9B9CCFFE7FF9B6CDEA097FD45F971" box="[654,764,1710,1735]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="6" pageNumber="7" unit="mm" value="11.4">11.4 mm</quantity>
), 
<date id="FF8F32E9FFE7FF9B6EFEA0F7FEF0F951" box="[174,329,1742,1767]" pageId="6" pageNumber="7" value="2011-08-05">05/VIII/2011</date>
, F.L. Mantelatto leg. (
<collectionCode id="ED208CECFFE7FF9B6C0CA0F7FD14F95E" box="[604,685,1742,1768]" country="Brazil" httpUri="http://grbio.org/cool/4wb1-ykjs" name="Crustacean Collection of the Department of Biology" pageId="6" pageNumber="7">CCDB</collectionCode>
1063); 
<collectingCountry id="F32654B9FFE7FF9B6EFEA0D7FEEBF8B1" box="[174,338,1774,1799]" name="Venezuela" pageId="6" pageNumber="7">VENEZUELA</collectingCountry>
, Isla Margarita, Boca Chica: 
<specimenCount id="9D37DFA0FFE7FF9B6CE4A0D7FCBEF8B1" box="[692,775,1774,1799]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE7FF9B6EB1A137FEFBF891" box="[225,322,1806,1831]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="6" pageNumber="7" unit="mm" value="8.4">8.4 mm</quantity>
), 
<specimenCount id="9D37DFA0FFE7FF9B6F06A137FE07F891" box="[342,446,1806,1831]" pageId="6" pageNumber="7" type="female">1 female</specimenCount>
(CL 10.0 mm), 
<date id="FF8F32E9FFE7FF9B6C2EA137FCBAF891" box="[638,771,1806,1831]" pageId="6" pageNumber="7" value="2010-11-03">03/XI/2010</date>
, R. Lopez leg. (
<collectionCode id="ED208CECFFE7FF9B6F32A117FE08F8FE" box="[354,433,1838,1864]" country="Brazil" httpUri="http://grbio.org/cool/4wb1-ykjs" name="Crustacean Collection of the Department of Biology" pageId="6" pageNumber="7">CCDB</collectionCode>
2428); La Restinga: 
<specimenCount id="9D37DFA0FFE7FF9B6CE4A117FCBEF8F1" box="[692,775,1838,1863]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE7FF9B6EBAA177FED0F8D1" box="[234,361,1870,1895]" metricMagnitude="-2" metricUnit="m" metricValue="1.24" pageId="6" pageNumber="7" unit="mm" value="12.4">12.4 mm</quantity>
), 
<specimenCount id="9D37DFA0FFE7FF9B6FD3A177FE43F8D1" box="[387,506,1870,1895]" pageId="6" pageNumber="7" type="female">1 female</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE7FF9B6C15A177FD7DF8D1" box="[581,708,1870,1895]" metricMagnitude="-2" metricUnit="m" metricValue="1.64" pageId="6" pageNumber="7" unit="mm" value="16.4">16.4 mm</quantity>
), 
<date id="FF8F32E9FFE7FF9B6C8EA177FEA4F831" pageId="6" pageNumber="7" value="2011-08-25">25/ VIII/2011</date>
, J. Bolaños leg. (
<collectionCode id="ED208CECFFE7FF9B6FA1A157FDF9F83E" box="[497,576,1902,1928]" country="Brazil" httpUri="http://grbio.org/cool/4wb1-ykjs" name="Crustacean Collection of the Department of Biology" pageId="6" pageNumber="7">CCDB</collectionCode>
3633); 
<collectingCountry id="F32654B9FFE7FF9B6CF1A157FCBDF831" box="[673,772,1902,1927]" name="Brazil" pageId="6" pageNumber="7">BRAZIL</collectingCountry>
, 
<collectingRegion id="49F5DACBFFE7FF9B6EFEA1B7FF42F81E" box="[174,251,1934,1960]" country="Brazil" name="Ceara" pageId="6" pageNumber="7">Ceará</collectingRegion>
, Aracati, Retiro Grande Beach: 
<specimenCount id="9D37DFA0FFE7FF9B6C2FA1B7FD6BF811" box="[639,722,1934,1959]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE7FF9B6D02A6C4FC7AFEA0" box="[850,963,253,278]" metricMagnitude="-2" metricUnit="m" metricValue="1.47" pageId="6" pageNumber="7" unit="mm" value="14.7">14.7 mm</quantity>
), 1 ovigerous female (CL 
<quantity id="4CC9B9CCFFE7FF9B6B50A6C4FAC8FEA0" box="[1280,1393,253,278]" metricMagnitude="-2" metricUnit="m" metricValue="1.38" pageId="6" pageNumber="7" unit="mm" value="13.8">13.8 mm</quantity>
), 
<date id="FF8F32E9FFE7FF9B6BD6A6C4FC11FE80" pageId="6" pageNumber="7" value="1964-01-17">17/ I/1964</date>
, A.L. Castro leg. (
<collectionCode id="ED208CECFFE7FF9B6AC9A724FB53FE80" box="[1177,1258,285,310]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="6" pageNumber="7">MNRJ</collectionCode>
4468); 
<collectingRegion id="49F5DACBFFE7FF9B6B05A724FA1EFE81" box="[1365,1447,285,311]" country="Brazil" name="Ceara" pageId="6" pageNumber="7">Ceará</collectingRegion>
, Mucuripe: 
<specimenCount id="9D37DFA0FFE7FF9B6D86A704FBFAFEE0" box="[982,1091,317,342]" pageId="6" pageNumber="7" type="female">1 female</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE7FF9B6AD3A704FB5EFEE0" box="[1155,1255,317,342]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="6" pageNumber="7" unit="mm" value="9.9">9.9 mm</quantity>
), 
<specimenCount id="9D37DFA0FFE7FF9B6AAEA704FACCFEE0" box="[1278,1397,317,342]" pageId="6" pageNumber="7" type="juvenile">1 juvenile</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE7FF9B6D02A764FC7CFEC0" box="[850,965,349,374]" metricMagnitude="-3" metricUnit="m" metricValue="6.54" pageId="6" pageNumber="7" unit="mm" value="6.54">6.54 mm</quantity>
), 
<date id="FF8F32E9FFE7FF9B6D8BA764FBDCFEC0" box="[987,1125,349,374]" pageId="6" pageNumber="7" value="1965-04-23">23/IV/1965</date>
, coll. not available (
<collectionCode id="ED208CECFFE7FF9B6B0DA764FA12FEC0" box="[1373,1451,349,374]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="6" pageNumber="7">MNRJ</collectionCode>
4462); 
<collectingRegion id="49F5DACBFFE7FF9B6DE3A744FBE7FE20" box="[947,1118,381,406]" country="Brazil" name="Pernambuco" pageId="6" pageNumber="7">Pernambuco</collectingRegion>
, Recife, Piedade Beach: 
<specimenCount id="9D37DFA0FFE7FF9B6D02A7A4FC01FE00" box="[850,952,413,438]" pageId="6" pageNumber="7" type="male">2 males</specimenCount>
(CL 10.3 and 
<quantity id="4CC9B9CCFFE7FF9B6A27A7A4FB54FE00" box="[1143,1261,413,438]" metricMagnitude="-2" metricUnit="m" metricValue="1.92" pageId="6" pageNumber="7" unit="mm" value="19.2">19.2 mm</quantity>
), 
<specimenCount id="9D37DFA0FFE7FF9B6B55A7A4FACDFE00" box="[1285,1396,413,438]" pageId="6" pageNumber="7" type="female">1 female</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE7FF9B6D02A784FC78FE60" box="[850,961,445,470]" metricMagnitude="-3" metricUnit="m" metricValue="8.52" pageId="6" pageNumber="7" unit="mm" value="8.52">8.52 mm</quantity>
), 1 ovigerous female (CL 
<quantity id="4CC9B9CCFFE7FF9B6AA5A784FACAFE60" box="[1269,1395,445,470]" metricMagnitude="-2" metricUnit="m" metricValue="1.506" pageId="6" pageNumber="7" unit="mm" value="15.06">15.06 mm</quantity>
), 
<date id="FF8F32E9FFE7FF9B6BD6A784FC0CFE40" pageId="6" pageNumber="7" value="1986-06-24">24/ VI/1986</date>
, A.L. Castro, P. Coelho and G. Melo leg. (
<collectionCode id="ED208CECFFE7FF9B6D09A7C4FC1EFDA0" box="[857,935,509,534]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="6" pageNumber="7">MNRJ</collectionCode>
4478); 
<collectingRegion id="49F5DACBFFE7FF9B6A59A7C4FB11FDA0" box="[1033,1192,509,534]" country="Brazil" name="Pernambuco" pageId="6" pageNumber="7">Pernambuco</collectingRegion>
, Recife, Boa Viagem Beach: 
<specimenCount id="9D37DFA0FFE7FF9B6DFFA424FBB7FD80" box="[943,1038,541,566]" pageId="6" pageNumber="7" type="male">4 males</specimenCount>
(CL 7.0 - 
<quantity id="4CC9B9CCFFE7FF9B6ADCA424FB42FD80" box="[1164,1275,541,566]" metricMagnitude="-2" metricUnit="m" metricValue="1.76" pageId="6" pageNumber="7" unit="mm" value="17.6">17.6 mm</quantity>
), 
<specimenCount id="9D37DFA0FFE7FF9B6B40A424FAC1FD80" box="[1296,1400,541,566]" pageId="6" pageNumber="7" type="female">1 female</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE7FF9B6D02A404FC02FDE0" box="[850,955,573,598]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" pageId="6" pageNumber="7" unit="mm" value="9.7">9.7 mm</quantity>
), 
<specimenCount id="9D37DFA0FFE7FF9B6D83A404FBE8FDE0" box="[979,1105,573,598]" pageId="6" pageNumber="7" type="juvenile">1 juvenile</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE7FF9B6AC6A404FB46FDE0" box="[1174,1279,573,598]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="6" pageNumber="7" unit="mm" value="5.5">5.5 mm</quantity>
), 
<date id="FF8F32E9FFE7FF9B6B47A404FA1EFDE0" box="[1303,1447,573,598]" pageId="6" pageNumber="7" value="2012-04-06">06/IV/2012</date>
, F.L. Mantelatto leg. (
<collectionCode id="ED208CECFFE7FF9B6A04A464FB1DFDC1" box="[1108,1188,605,631]" country="Brazil" httpUri="http://grbio.org/cool/4wb1-ykjs" name="Crustacean Collection of the Department of Biology" pageId="6" pageNumber="7">CCDB</collectionCode>
3814); 
<collectingRegion id="49F5DACBFFE7FF9B6B58A464FA11FDC0" box="[1288,1448,605,630]" country="Brazil" name="Pernambuco" pageId="6" pageNumber="7">Pernambuco</collectingRegion>
, Ipojuca, Serrambi county, Serrambi Beach: 
<specimenCount id="9D37DFA0FFE7FF9B6B0AA444FA15FD20" box="[1370,1452,637,662]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE7FF9B6DD5A4A4FC5FFD00" box="[901,998,669,694]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="6" pageNumber="7" unit="mm" value="8.3">8.3 mm</quantity>
), 
<specimenCount id="9D37DFA0FFE7FF9B6DA9A4A4FBD2FD00" box="[1017,1131,669,694]" pageId="6" pageNumber="7" type="juvenile">1 juvenile</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE7FF9B6AF8A4A4FAB1FD00" box="[1192,1288,669,694]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="6" pageNumber="7" unit="mm" value="5.3">5.3 mm</quantity>
), 
<date id="FF8F32E9FFE7FF9B6B4CA4A4FA11FD00" box="[1308,1448,669,694]" pageId="6" pageNumber="7" value="2012-12-25">25/XII/2012</date>
, F.L. Mantelatto and F.B. Mantelatto leg. (
<collectionCode id="ED208CECFFE7FF9B6B0BA484FA12FD61" box="[1371,1451,701,727]" country="Brazil" httpUri="http://grbio.org/cool/4wb1-ykjs" name="Crustacean Collection of the Department of Biology" pageId="6" pageNumber="7">CCDB</collectionCode>
4481); 
<collectingRegion id="49F5DACBFFE7FF9B6DF7A4E4FBB0FD41" box="[935,1033,733,759]" country="Brazil" name="Paraiba" pageId="6" pageNumber="7">Paraíba</collectingRegion>
, Jacunã, Tambaba Beach: 
<specimenCount id="9D37DFA0FFE7FF9B6B1DA4E4FA12FD40" box="[1357,1451,733,758]" pageId="6" pageNumber="7" type="male">2 males</specimenCount>
(CL 6.9 and 
<quantity id="4CC9B9CCFFE7FF9B6DA0A4C4FBDAFCA0" box="[1008,1123,765,790]" metricMagnitude="-2" metricUnit="m" metricValue="1.78" pageId="6" pageNumber="7" unit="mm" value="17.8">17.8 mm</quantity>
), 
<specimenCount id="9D37DFA0FFE7FF9B6A28A4C4FB5DFCA0" box="[1144,1252,765,790]" pageId="6" pageNumber="7" type="female">1 female</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE7FF9B6B73A4C4FA3EFCA0" box="[1315,1415,765,790]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="6" pageNumber="7" unit="mm" value="9.1">9.1 mm</quantity>
), 1 ovigerous female (CL 14.0 mm), 
<date id="FF8F32E9FFE7FF9B6ABEA524FAD4FC80" box="[1262,1389,797,822]" pageId="6" pageNumber="7" value="1995-02-23">23/II/1995</date>
, P.S. Young and C.S. Serejo leg. (
<collectionCode id="ED208CECFFE7FF9B6ACFA504FB52FCE0" box="[1183,1259,829,854]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="6" pageNumber="7">MNRJ</collectionCode>
6487); 
<collectingRegion id="49F5DACBFFE7FF9B6B14A504FA11FCE0" box="[1348,1448,829,854]" country="Brazil" name="Alagoas" pageId="6" pageNumber="7">Alagoas</collectingRegion>
, Marechal Deodoro, Francês Beach: 1 ovigerous female (CL 
<quantity id="4CC9B9CCFFE7FF9B6DB2A544FBEBFC20" box="[994,1106,893,918]" metricMagnitude="-2" metricUnit="m" metricValue="1.8699999999999999" pageId="6" pageNumber="7" unit="mm" value="18.7">18.7 mm</quantity>
), 
<date id="FF8F32E9FFE7FF9B6A37A544FB5AFC20" box="[1127,1251,893,918]" pageId="6" pageNumber="7" value="1995-02-16">16/II/1995</date>
, P.S. Young and C.S. Serejo leg. (
<collectionCode id="ED208CECFFE7FF9B6A72A5A4FBD6FC00" box="[1058,1135,925,950]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="6" pageNumber="7">MNRJ</collectionCode>
6630); 
<collectingRegion id="49F5DACBFFE7FF9B6A9CA5A4FA88FC00" box="[1228,1329,925,950]" country="Brazil" name="Alagoas" pageId="6" pageNumber="7">Alagoas</collectingRegion>
, Marechal Deodoro, Torto Beach: 
<specimenCount id="9D37DFA0FFE7FF9B6A36A584FB75FC60" box="[1126,1228,957,982]" pageId="6" pageNumber="7" type="female">1 female</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE7FF9B6B56A584FACDFC60" box="[1286,1396,957,982]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="6" pageNumber="7" unit="mm" value="10.7">10.7 mm</quantity>
), 
<date id="FF8F32E9FFE7FF9B6BD6A584FC15FC40" pageId="6" pageNumber="7" value="1995-02-16">16/ II/1995</date>
, P.S Young and C.S. Serejo leg. (
<collectionCode id="ED208CECFFE7FF9B6B0CA5E4FA12FC40" box="[1372,1451,989,1014]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="6" pageNumber="7">MNRJ</collectionCode>
6671); 
<collectingRegion id="49F5DACBFFE7FF9B6DFCA5C4FC4EFBA0" box="[940,1015,1021,1046]" country="Brazil" name="Bahia" pageId="6" pageNumber="7">Bahia</collectingRegion>
, Lauro de Freitas, Ipitanga Beach: 
<specimenCount id="9D37DFA0FFE7FF9B6D02A224FC03FB80" box="[850,954,1053,1078]" pageId="6" pageNumber="7" type="female">1 female</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE7FF9B6DA5A224FBDAFB80" box="[1013,1123,1053,1078]" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" pageId="6" pageNumber="7" unit="mm" value="13.2">13.2 mm</quantity>
), 
<date id="FF8F32E9FFE7FF9B6A27A224FAB8FB80" box="[1143,1281,1053,1078]" pageId="6" pageNumber="7" value="2011-12-22">22/XII/2011</date>
, F.L. Carvalho and E.A. Souza-Carvalho leg. (
<collectionCode id="ED208CECFFE7FF9B6AA7A204FAF2FBE1" box="[1271,1355,1085,1111]" country="Brazil" httpUri="http://grbio.org/cool/4wb1-ykjs" name="Crustacean Collection of the Department of Biology" pageId="6" pageNumber="7">CCDB</collectionCode>
3789); 
<collectingRegion id="49F5DACBFFE7FF9B6D02A264FC25FBC0" box="[850,924,1117,1142]" country="Brazil" name="Bahia" pageId="6" pageNumber="7">Bahia</collectingRegion>
, Itacaré, Ribeira Beach: 
<specimenCount id="9D37DFA0FFE7FF9B6A81A264FA8AFBC0" box="[1233,1331,1117,1142]" pageId="6" pageNumber="7" type="male">3 males</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE7FF9B6B22A264FC07FB20" metricMagnitude="-3" metricUnit="m" metricValue="7.55" metricValueMax="11.4" metricValueMin="3.7" pageId="6" pageNumber="7" unit="mm" value="7.55" valueMax="11.4" valueMin="3.7">3.7 - 11.4 mm</quantity>
), 
<specimenCount id="9D37DFA0FFE7FF9B6D81A244FBFDFB20" box="[977,1092,1149,1174]" pageId="6" pageNumber="7" type="female">4 females</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE7FF9B6A2EA244FAA2FB20" box="[1150,1307,1149,1174]" metricMagnitude="-3" metricUnit="m" metricValue="6.199999999999999" metricValueMax="7.8" metricValueMin="4.6" pageId="6" pageNumber="7" unit="mm" value="6.199999999999999" valueMax="7.8" valueMin="4.6">4.6 - 7.8 mm</quantity>
), 
<specimenCount id="9D37DFA0FFE7FF9B6B7DA244FA15FB20" box="[1325,1452,1149,1174]" pageId="6" pageNumber="7" type="juvenile">9 juveniles</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE7FF9B6DDAA2A4FB80FB00" box="[906,1081,1181,1206]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.7" metricValueMin="2.3" pageId="6" pageNumber="7" unit="mm" value="3.0" valueMax="3.7" valueMin="2.3">2.3 - 3.7 mm</quantity>
), 
<date id="FF8F32E9FFE7FF9B6A01A2A4FB6FFB00" box="[1105,1238,1181,1206]" pageId="6" pageNumber="7" value="1994-02-22">22/II/1994</date>
, P.S. Young and M.M. Britto-Pereira leg. (
<collectionCode id="ED208CECFFE7FF9B6ACCA284FB57FB60" box="[1180,1262,1213,1238]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="6" pageNumber="7">MNRJ</collectionCode>
4507); 
<collectingRegion id="49F5DACBFFE7FF9B6B09A284FA1EFB60" box="[1369,1447,1213,1238]" country="Brazil" name="Bahia" pageId="6" pageNumber="7">Bahia</collectingRegion>
, Ilhéus, Morro de 
<collectingRegion id="49F5DACBFFE7FF9B6A7DA2E4FB76FB40" box="[1069,1231,1245,1270]" country="Brazil" name="Pernambuco" pageId="6" pageNumber="7">Pernambuco</collectingRegion>
, Badusca Beach: 
<specimenCount id="9D37DFA0FFE7FF9B6D02A2C4FC0CFAA0" box="[850,949,1277,1302]" pageId="6" pageNumber="7" type="male">4 males</specimenCount>
(CL 4.2 - 6.0 mm), 
<specimenCount id="9D37DFA0FFE7FF9B6AE6A2C4FA88FAA0" box="[1206,1329,1277,1302]" pageId="6" pageNumber="7" type="female">6 females</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE7FF9B6B22A2C4FC7AFA80" metricMagnitude="-3" metricUnit="m" metricValue="8.7" metricValueMax="13.8" metricValueMin="3.6" pageId="6" pageNumber="7" unit="mm" value="8.7" valueMax="13.8" valueMin="3.6">3.6 - 13.8 mm</quantity>
), 
<date id="FF8F32E9FFE7FF9B6D88A324FBF6FA80" box="[984,1103,1309,1334]" pageId="6" pageNumber="7" value="2010-11-06">6/XI/2010</date>
, F.L. Mantelatto, F. Carvalho and L. Pileggi leg. (
<collectionCode id="ED208CECFFE7FF9B6A1AA304FB23FAE1" box="[1098,1178,1341,1367]" country="Brazil" httpUri="http://grbio.org/cool/4wb1-ykjs" name="Crustacean Collection of the Department of Biology" pageId="6" pageNumber="7">CCDB</collectionCode>
2427); 
<collectingRegion id="49F5DACBFFE7FF9B6AAFA304FAF3FAE0" box="[1279,1354,1341,1366]" country="Brazil" name="Bahia" pageId="6" pageNumber="7">Bahia</collectingRegion>
, Ilhéus, Morro de 
<collectingRegion id="49F5DACBFFE7FF9B6D9CA364FBD3FAC0" box="[972,1130,1373,1398]" country="Brazil" name="Pernambuco" pageId="6" pageNumber="7">Pernambuco</collectingRegion>
, Badusca Beach: 
<specimenCount id="9D37DFA0FFE7FF9B6B1AA364FA12FAC0" box="[1354,1451,1373,1398]" pageId="6" pageNumber="7" type="male">4 males</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE7FF9B6DD6A344FB8FFA20" box="[902,1078,1405,1430]" metricMagnitude="-3" metricUnit="m" metricValue="9.45" metricValueMax="11.5" metricValueMin="7.4" pageId="6" pageNumber="7" unit="mm" value="9.45" valueMax="11.5" valueMin="7.4">7.4 - 11.5 mm</quantity>
), 
<specimenCount id="9D37DFA0FFE7FF9B6A1AA344FB79FA20" box="[1098,1216,1405,1430]" pageId="6" pageNumber="7" type="female">3 females</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE7FF9B6AAEA344FA18FA20" box="[1278,1441,1405,1430]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" metricValueMax="9.9" metricValueMin="6.7" pageId="6" pageNumber="7" unit="mm" value="8.3" valueMax="9.9" valueMin="6.7">6.7 - 9.9 mm</quantity>
), 2 ovigerous females (CL 9.9 and 
<quantity id="4CC9B9CCFFE7FF9B6B5DA3A4FA3CFA00" box="[1293,1413,1437,1462]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="6" pageNumber="7" unit="mm" value="13.1">13.1 mm</quantity>
), 
<specimenCount id="9D37DFA0FFE7FF9B6BCCA3A4FC0EFA60" pageId="6" pageNumber="7" type="juvenile">1 juvenile</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE7FF9B6DAFA384FBD3FA60" box="[1023,1130,1469,1494]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="6" pageNumber="7" unit="mm" value="3.9">3.9 mm</quantity>
), 
<date id="FF8F32E9FFE7FF9B6AD3A384FAB8FA60" box="[1155,1281,1469,1494]" pageId="6" pageNumber="7" value="2011-01-21">21/I/2011</date>
, F. Carvalho and E.A. Souza-Carvalho leg. (
<collectionCode id="ED208CECFFE7FF9B6AA7A3E4FAF2FA41" box="[1271,1355,1501,1527]" country="Brazil" httpUri="http://grbio.org/cool/4wb1-ykjs" name="Crustacean Collection of the Department of Biology" pageId="6" pageNumber="7">CCDB</collectionCode>
3423); 
<collectingRegion id="49F5DACBFFE7FF9B6D02A3C4FBB8F9A1" box="[850,1025,1533,1559]" country="Brazil" name="Espirito Santo" pageId="6" pageNumber="7">Espírito Santo</collectingRegion>
, Guarapari, Castanheiros Beach: 1 ovigerous female (CL 
<quantity id="4CC9B9CCFFE7FF9B6A35A024FB6EF980" box="[1125,1239,1565,1590]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="6" pageNumber="7" unit="mm" value="10.1">10.1 mm</quantity>
), 
<date id="FF8F32E9FFE7FF9B6ABDA024FAD7F980" box="[1261,1390,1565,1590]" pageId="6" pageNumber="7" value="1993-10-30">30/X/1993</date>
, P.S. Young leg. (
<collectionCode id="ED208CECFFE7FF9B6DB6A004FB8DF9E0" box="[998,1076,1597,1622]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="6" pageNumber="7">MNRJ</collectionCode>
4547); 
<collectingRegion id="49F5DACBFFE7FF9B6AC6A004FAF0F9E1" box="[1174,1353,1597,1623]" country="Brazil" name="Espirito Santo" pageId="6" pageNumber="7">Espírito Santo</collectingRegion>
, Vitória, Boi Island: 
<specimenCount id="9D37DFA0FFE7FF9B6D87A064FB9EF9C0" box="[983,1063,1629,1654]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE7FF9B6A31A064FB77F9C0" box="[1121,1230,1629,1654]" metricMagnitude="-2" metricUnit="m" metricValue="2.07" pageId="6" pageNumber="7" unit="mm" value="20.7">20.7 mm</quantity>
), 
<date id="FF8F32E9FFE7FF9B6AB0A064FAE3F9C0" box="[1248,1370,1629,1654]" pageId="6" pageNumber="7" value="1987-02-11">11/II/1987</date>
, Flávio and Iara leg. (
<collectionCode id="ED208CECFFE7FF9B6DA7A044FBFDF920" box="[1015,1092,1661,1686]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="6" pageNumber="7">MNRJ</collectionCode>
4664); 
<collectingRegion id="49F5DACBFFE7FF9B6AF1A044FAF4F921" box="[1185,1357,1661,1687]" country="Brazil" name="Espirito Santo" pageId="6" pageNumber="7">Espírito Santo</collectingRegion>
, Vitória, Tubarão Harbor: 
<specimenCount id="9D37DFA0FFE7FF9B6A4BA0A4FBD2F900" box="[1051,1131,1693,1718]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE7FF9B6AF3A0A4FAA9F900" box="[1187,1296,1693,1718]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" pageId="6" pageNumber="7" unit="mm" value="18.5">18.5 mm</quantity>
), 1 ovigerous female (CL 
<quantity id="4CC9B9CCFFE7FF9B6DA7A084FBCEF960" box="[1015,1143,1725,1750]" metricMagnitude="-2" metricUnit="m" metricValue="1.66" pageId="6" pageNumber="7" unit="mm" value="16.6">16.6 mm</quantity>
), 
<date id="FF8F32E9FFE7FF9B6AC1A084FA92F960" box="[1169,1323,1725,1750]" pageId="6" pageNumber="7" value="1987-11-10">10/XI/1987</date>
, coll. not available (
<collectionCode id="ED208CECFFE7FF9B6D9AA0E4FBAEF940" box="[970,1047,1757,1782]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="6" pageNumber="7">MNRJ</collectionCode>
4663); 
<collectingRegion id="49F5DACBFFE7FF9B6A22A0E4FA9DF940" box="[1138,1316,1757,1782]" country="Brazil" name="Rio de Janeiro" pageId="6" pageNumber="7">Rio de Janeiro</collectingRegion>
, Cabo Frio, Peró Beach: 
<specimenCount id="9D37DFA0FFE7FF9B6DA4A0C4FBF1F8A0" box="[1012,1096,1789,1814]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE7FF9B6AD7A0C4FB53F8A0" box="[1159,1258,1789,1814]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="6" pageNumber="7" unit="mm" value="9.4">9.4 mm</quantity>
), 
<specimenCount id="9D37DFA0FFE7FF9B6AAFA0C4FACEF8A0" box="[1279,1399,1789,1814]" pageId="6" pageNumber="7" type="female">2 females</specimenCount>
(CL 6.3 and 7.9), 
<date id="FF8F32E9FFE7FF9B6DA8A124FB3CF880" box="[1016,1157,1821,1846]" pageId="6" pageNumber="7" value="1965-07-28">28/VII/1965</date>
, A.L. Castro leg. (
<collectionCode id="ED208CECFFE7FF9B6B0EA124FA12F880" box="[1374,1451,1821,1846]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="6" pageNumber="7">MNRJ</collectionCode>
4661); 
<collectingRegion id="49F5DACBFFE7FF9B6DF5A104FBEDF8E0" box="[933,1108,1853,1878]" country="Brazil" name="Rio de Janeiro" pageId="6" pageNumber="7">Rio de Janeiro</collectingRegion>
, Niterói, Itaipú Beach: 
<specimenCount id="9D37DFA0FFE7FF9B6B0BA104FA15F8E0" box="[1371,1452,1853,1878]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE7FF9B6DDBA164FBBAF8C0" box="[907,1027,1885,1910]" metricMagnitude="-2" metricUnit="m" metricValue="1.96" pageId="6" pageNumber="7" unit="mm" value="19.6">19.6 mm</quantity>
), 
<date id="FF8F32E9FFE7FF9B6A4BA164FB3AF8C0" box="[1051,1155,1885,1910]" bridgedPair="/" pageId="6" pageNumber="7" value="1963-04">IV/1963</date>
, J. Becker leg. (
<collectionCode id="ED208CECFFE7FF9B6B0AA164FA13F8C0" box="[1370,1450,1885,1910]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="6" pageNumber="7">MNRJ</collectionCode>
4471); 
<collectingRegion id="49F5DACBFFE7FF9B6DF5A144FBEDF820" box="[933,1108,1917,1942]" country="Brazil" name="Rio de Janeiro" pageId="6" pageNumber="7">Rio de Janeiro</collectingRegion>
, Niterói, Itaipú Beach: 
<specimenCount id="9D37DFA0FFE7FF9B6B0BA144FA15F820" box="[1371,1452,1917,1942]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE7FF9B6DD4A1A4FC48F800" box="[900,1009,1949,1974]" metricMagnitude="-2" metricUnit="m" metricValue="1.44" pageId="6" pageNumber="7" unit="mm" value="14.4">14.4 mm</quantity>
), 
<specimenCount id="9D37DFA0FFE7FF9B6A53A1A4FBD3F800" box="[1027,1130,1949,1974]" pageId="6" pageNumber="7" type="female">1 female</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE7FF9B6AF2A1A4FAB8F800" box="[1186,1281,1949,1974]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="6" pageNumber="7" unit="mm" value="9.6">9.6 mm</quantity>
), 
<date id="FF8F32E9FFE7FF9B6B43A1A4FA30F800" box="[1299,1417,1949,1974]" pageId="6" pageNumber="7" value="1992-04-03">3/IV/1992</date>
, A. Macedo leg. (
<collectionCode id="ED208CECFFE6FF9A6F7AA6C4FECEFEA0" box="[298,375,253,278]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="7" pageNumber="8">MNRJ</collectionCode>
4515); 
<collectingRegion id="49F5DACBFFE6FF9A6F84A6C4FD3EFEA0" box="[468,647,253,278]" country="Brazil" name="Rio de Janeiro" pageId="7" pageNumber="8">Rio de Janeiro</collectingRegion>
, 
<collectingRegion id="49F5DACBFFE6FF9A6CC3A6C4FF5DFE80" country="Brazil" name="Rio de Janeiro" pageId="7" pageNumber="8">Rio de Janeiro</collectingRegion>
, Barra de Guaratiba: 
<specimenCount id="9D37DFA0FFE6FF9A6FB0A724FD89FE80" box="[480,560,285,310]" pageId="7" pageNumber="8" type="male">1 male</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE6FF9A6C39A724FD6FFE80" box="[617,726,285,310]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="7" pageNumber="8" unit="mm" value="13.1">13.1 mm</quantity>
), 
<date id="FF8F32E9FFE6FF9A6ED8A704FEA8FEE0" box="[136,273,317,342]" pageId="7" pageNumber="8" value="1953-04-30">30/IV/1953</date>
, N. Santos leg. (
<collectionCode id="ED208CECFFE6FF9A6FB0A704FD97FEE0" box="[480,558,317,342]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="7" pageNumber="8">MNRJ</collectionCode>
4467); 
<collectingRegion id="49F5DACBFFE6FF9A6CC0A704FF51FEC0" country="Brazil" name="Rio de Janeiro" pageId="7" pageNumber="8">Rio de Janeiro</collectingRegion>
, 
<collectingRegion id="49F5DACBFFE6FF9A6EA7A764FE0DFEC0" box="[247,436,349,374]" country="Brazil" name="Rio de Janeiro" pageId="7" pageNumber="8">Rio de Janeiro</collectingRegion>
, Flamengo Bay: 
<specimenCount id="9D37DFA0FFE6FF9A6CDBA764FD59FEC0" box="[651,736,349,374]" pageId="7" pageNumber="8" type="male">1 male</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE6FF9A6EEDA744FE89FE20" box="[189,304,381,406]" metricMagnitude="-2" metricUnit="m" metricValue="1.7899999999999998" pageId="7" pageNumber="8" unit="mm" value="17.9">17.9 mm</quantity>
), 
<date id="FF8F32E9FFE6FF9A6F15A744FE64FE20" box="[325,477,381,406]" pageId="7" pageNumber="8" value="1982-08-06">06/VIII/1982</date>
, R. Leite leg. (
<collectionCode id="ED208CECFFE6FF9A6CC3A744FD58FE20" box="[659,737,381,406]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="7" pageNumber="8">MNRJ</collectionCode>
4470); 
<collectingRegion id="49F5DACBFFE6FF9A6EBBA7A4FEC5FE00" box="[235,380,413,439]" country="Brazil" name="Sao Paulo" pageId="7" pageNumber="8">São Paulo</collectingRegion>
, Ubatuba, Grande Beach: 
<specimenCount id="9D37DFA0FFE6FF9A6D7DA6C4FCC7FEA0" box="[813,894,253,278]" pageId="7" pageNumber="8" type="male">1 male</specimenCount>
(
<quantity id="4CC9B9CCFFE6FF9A6DDDA6C4FC47FEA0" box="[909,1022,253,278]" metricMagnitude="-2" metricUnit="m" metricValue="2.26" pageId="7" pageNumber="8" unit="mm" value="22.6">22.6 mm</quantity>
), 
<date id="FF8F32E9FFE6FF9A6A41A6C4FB2FFEA0" box="[1041,1174,253,278]" pageId="7" pageNumber="8" value="1987-11-04">04/XI/1987</date>
, F.L. Mantelatto leg. (
<collectionCode id="ED208CECFFE6FF9A6D63A724FC38FE81" box="[819,897,285,311]" country="Brazil" httpUri="http://grbio.org/cool/4wb1-ykjs" name="Crustacean Collection of the Department of Biology" pageId="7" pageNumber="8">CCDB</collectionCode>
0067); 
<collectingRegion id="49F5DACBFFE6FF9A6D8EA724FBE7FE80" box="[990,1118,285,311]" country="Brazil" name="Sao Paulo" pageId="7" pageNumber="8">São Paulo</collectingRegion>
, Ubatuba, Itaguá Beach: 
<specimenCount id="9D37DFA0FFE6FF9A6D7DA704FC35FEE0" box="[813,908,317,342]" pageId="7" pageNumber="8" type="male">2 males</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE6FF9A6D99A704FB81FEE0" box="[969,1080,317,342]" metricMagnitude="-2" metricUnit="m" metricValue="2.13" pageId="7" pageNumber="8" unit="mm" value="21.3">21.3 mm</quantity>
), 2 ovigerous females (15.7 and 
<quantity id="4CC9B9CCFFE6FF9A6D30A764FC74FEC0" box="[864,973,349,374]" metricMagnitude="-2" metricUnit="m" metricValue="1.61" pageId="7" pageNumber="8" unit="mm" value="16.1">16.1 mm</quantity>
), 
<date id="FF8F32E9FFE6FF9A6D8FA764FB86FEC0" box="[991,1087,349,374]" bridgedPair="/" pageId="7" pageNumber="8" value="1999-06">VI/1999</date>
, F.L. Mantelatto leg. (
<collectionCode id="ED208CECFFE6FF9A6B67A764FA3CFEC1" box="[1335,1413,349,375]" country="Brazil" httpUri="http://grbio.org/cool/4wb1-ykjs" name="Crustacean Collection of the Department of Biology" pageId="7" pageNumber="8">CCDB</collectionCode>
0046); 
<collectingRegion id="49F5DACBFFE6FF9A6D2FA744FC47FE20" box="[895,1022,381,407]" country="Brazil" name="Sao Paulo" pageId="7" pageNumber="8">São Paulo</collectingRegion>
, Ubatuba, Grande Beach: 
<specimenCount id="9D37DFA0FFE6FF9A6B66A744FA3FFE20" box="[1334,1414,381,406]" pageId="7" pageNumber="8" type="male">1 male</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE6FF9A6D31A7A4FC68FE00" box="[865,977,413,438]" metricMagnitude="-2" metricUnit="m" metricValue="1.76" pageId="7" pageNumber="8" unit="mm" value="17.6">17.6 mm</quantity>
), 1 ovigerous female (CL 
<quantity id="4CC9B9CCFFE6FF9A6B5BA7A4FAC2FE00" box="[1291,1403,413,438]" metricMagnitude="-2" metricUnit="m" metricValue="1.52" pageId="7" pageNumber="8" unit="mm" value="15.2">15.2 mm</quantity>
), 
<date id="FF8F32E9FFE9FF956EFEA6C4FEB2FEA0" box="[174,267,253,278]" bridgedPair="/" pageId="8" pageNumber="9" value="2002-10">X/2002</date>
, F.L. Mantelatto leg. (
<collectionCode id="ED208CECFFE9FF956C4FA6C4FDC9FEA1" box="[543,624,253,279]" country="Brazil" httpUri="http://grbio.org/cool/4wb1-ykjs" name="Crustacean Collection of the Department of Biology" pageId="8" pageNumber="9">CCDB</collectionCode>
0760); 
<collectingRegion id="49F5DACBFFE9FF956C86A6C4FF47FE80" country="Brazil" name="Sao Paulo" pageId="8" pageNumber="9">São Paulo</collectingRegion>
, Ubatuba, Ubatuba Bay: 
<specimenCount id="9D37DFA0FFE9FF956C08A724FD75FE80" box="[600,716,285,310]" pageId="8" pageNumber="9" type="female">1 female</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE9FF956EFEA704FEA7FEE0" box="[174,286,317,342]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="8" pageNumber="9" unit="mm" value="12.3">12.3 mm</quantity>
), 
<date id="FF8F32E9FFE9FF956F63A704FE0BFEE0" box="[307,434,317,342]" pageId="8" pageNumber="9" value="2010-05-13">13/V/2010</date>
, F.L. Mantelatto leg. (
<collectionCode id="ED208CECFFE9FF956CE8A704FCBFFEE1" box="[696,774,317,343]" country="Brazil" httpUri="http://grbio.org/cool/4wb1-ykjs" name="Crustacean Collection of the Department of Biology" pageId="8" pageNumber="9">CCDB</collectionCode>
2436); 
<collectingRegion id="49F5DACBFFE9FF956F50A764FEC6FEC0" box="[256,383,349,375]" country="Brazil" name="Sao Paulo" pageId="8" pageNumber="9">São Paulo</collectingRegion>
, Ubatuba, Grande Beach: 
<specimenCount id="9D37DFA0FFE9FF956CE7A764FCBEFEC0" box="[695,775,349,374]" pageId="8" pageNumber="9" type="male">1 male</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE9FF956EB0A744FEF6FE20" box="[224,335,381,406]" metricMagnitude="-2" metricUnit="m" metricValue="2.48" pageId="8" pageNumber="9" unit="mm" value="24.8">24.8 mm</quantity>
), 
<date id="FF8F32E9FFE9FF956F32A744FE56FE20" box="[354,495,381,406]" pageId="8" pageNumber="9" value="2012-12-08">08/XII/2012</date>
, I.C. Leone leg. (
<collectionCode id="ED208CECFFE9FF956CE9A744FCBEFE21" box="[697,775,381,407]" country="Brazil" httpUri="http://grbio.org/cool/4wb1-ykjs" name="Crustacean Collection of the Department of Biology" pageId="8" pageNumber="9">CCDB</collectionCode>
3949).
</paragraph>
<caption id="DF4E44A1FFE6FF9A6ED8A12FFDACF803" ID-DOI="http://doi.org/10.5281/zenodo.8060307" ID-Zenodo-Dep="8060307" httpUri="https://zenodo.org/record/8060307/files/figure.png" pageId="7" pageNumber="8" startId="7.[136,170,1814,1834]" targetBox="[136,1413,500,1788]" targetPageId="7" targetType="figure">
<paragraph id="8B8E1429FFE6FF9A6ED8A12FFDACF803" blockId="7.[136,1414,1813,1973]" pageId="7" pageNumber="8">
<emphasis id="B945C83BFFE6FF9A6ED8A12FFF7EF89C" bold="true" box="[136,199,1814,1834]" pageId="7" pageNumber="8">Fig. 2.</emphasis>
Overview of samples of 
<taxonomicName id="4C316FAAFFE6FF9A6F90A12FFD07F89C" authority="Latreille, 1828" authorityName="Latreille" authorityYear="1828" box="[448,702,1814,1834]" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B945C83BFFE6FF9A6F90A12FFD96F89C" box="[448,559,1814,1834]" italics="true" pageId="7" pageNumber="8">Acanthonyx</emphasis>
Latreille, 1828
</taxonomicName>
and some parts of their body. “ 
<taxonomicName id="4C316FAAFFE6FF9A6DBFA12FFAF4F89C" authority="&quot; H. Milne Edwards, 1834" authorityName="&quot; H. Milne Edwards" authorityYear="1834" box="[1007,1357,1814,1834]" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="petiverii">
<emphasis id="B945C83BFFE6FF9A6DBFA12FFBEDF89C" box="[1007,1108,1814,1834]" italics="true" pageId="7" pageNumber="8">A. petiverii</emphasis>
” H. 
<bibRefCitation id="EFA069D8FFE6FF9A6AD0A12FFAF4F89C" author="Milne Edwards H." box="[1152,1357,1814,1834]" pageId="7" pageNumber="8" pagination="1 - 468" refId="ref13717" refString="Milne Edwards H. 1834. Histoire naturelle des Crustaces, comprenant l'anatomie, la physiologie et la classification de ces animaux, 1: 1 - 468." type="journal article" year="1834">Milne Edwards, 1834</bibRefCitation>
</taxonomicName>
, male CL 13.5 mm (CCDB 1063). (A) dorsal view, (B) ventral view; “ 
<taxonomicName id="4C316FAAFFE6FF9A6C9FA10BFC52F8F0" authority="&quot; Coelho, 1993" authorityName="&quot; Coelho" authorityYear="1993" box="[719,1003,1841,1862]" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="dissimulatus">
<emphasis id="B945C83BFFE6FF9A6C9FA10BFCE7F8F0" box="[719,862,1842,1862]" italics="true" pageId="7" pageNumber="8">A. dissimulatus</emphasis>
” 
<bibRefCitation id="EFA069D8FFE6FF9A6D3CA108FC52F8F0" author="Coelho PA &amp; Torres MFA" box="[876,1003,1841,1862]" pageId="7" pageNumber="8" pagination="221 - 241" refId="ref12460" refString="Coelho PA, Torres MFA. 1993. Taxonomia e distribuicao das especies do genero Acanthonyx Latreille, no Brasil (Crustacea: Decapoda: Majidae). Trab Oceanogr Univ Fed Pe 22: 221 - 241." type="journal article" year="1993">Coelho, 1993</bibRefCitation>
</taxonomicName>
male CL 12.2 mm (CCDB 1421) (C) dorsal view, (D) ventral view; (E) bifid rostrum, male CL 24.5 mm (CCDB 3949); (F) dorsal view of male left cheliped, CL 24.5 mm (CCDB 3949); (G) ventral view of female right cheliped, CL 8.3 mm (CCDB 3423); (H) subchela of the propodus of the last three ambulatory pereopods, male CL 24.5 mm (CCDB 3949). Scale bars: 5 mm (A, B, C, D, F); 2 mm (E, G); 0.6 mm (H). The species names in inverted commas refer to the original identification.
</paragraph>
</caption>
<paragraph id="8B8E1429FFE9FF956EBAA784FD87FA00" blockId="8.[174,776,253,1974]" pageId="8" pageNumber="9">
Additional material initially identified as 
<taxonomicName id="4C316FAAFFE9FF956CBDA784FEF8FE40" authorityName="&quot; Coelho" authorityYear="1993" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="dissimulatus">
<emphasis id="B945C83BFFE9FF956CBDA784FEF8FE40" italics="true" pageId="8" pageNumber="9">A. dissimulatus</emphasis>
</taxonomicName>
. 
<collectingCountry id="F32654B9FFE9FF956F01A7E4FE06FE41" box="[337,447,477,503]" name="Mexico" pageId="8" pageNumber="9">MEXICO</collectingCountry>
, 
<collectingRegion id="49F5DACBFFE9FF956F9BA7E4FDCCFE40" box="[459,629,477,503]" country="Mexico" name="Quintana Roo" pageId="8" pageNumber="9">Quintana Roo</collectingRegion>
, La Mancha Rodes: 
<specimenCount id="9D37DFA0FFE9FF956F5CA7C4FEE6FDA0" box="[268,351,509,534]" pageId="8" pageNumber="9" type="male">1 male</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE9FF956FCCA7C4FDB5FDA0" box="[412,524,509,534]" metricMagnitude="-2" metricUnit="m" metricValue="1.59" pageId="8" pageNumber="9" unit="mm" value="15.9">15.9 mm</quantity>
), 1 ovigerous female (CL 
<quantity id="4CC9B9CCFFE9FF956EB8A424FEEAFD80" box="[232,339,541,566]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="8" pageNumber="9" unit="mm" value="8.2">8.2 mm</quantity>
), 
<date id="FF8F32E9FFE9FF956F3BA424FDB0FD80" box="[363,521,541,566]" pageId="8" pageNumber="9" value="2002-07-02">02/VII/2002</date>
, coll. not available (
<collectionCode id="ED208CECFFE9FF956EE5A404FEB1FDE1" box="[181,264,573,599]" country="Brazil" httpUri="http://grbio.org/cool/4wb1-ykjs" name="Crustacean Collection of the Department of Biology" pageId="8" pageNumber="9">CCDB</collectionCode>
2430); 
<collectingCountry id="F32654B9FFE9FF956F22A404FE62FDE0" box="[370,475,573,598]" name="Brazil" pageId="8" pageNumber="9">BRAZIL</collectingCountry>
, 
<collectingRegion id="49F5DACBFFE9FF956FBAA404FCBBFDE0" box="[490,770,573,599]" country="Brazil" name="Rio Grande do Norte" pageId="8" pageNumber="9">Rio Grande do Norte</collectingRegion>
, Potiguar Basin: 1 ovigerous female (CL 
<quantity id="4CC9B9CCFFE9FF956CCAA464FD45FDC0" box="[666,764,605,630]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="8" pageNumber="9" unit="mm" value="9.9">9.9 mm</quantity>
), 
<date id="FF8F32E9FFE9FF956EFEA444FE8EFD20" box="[174,311,637,662]" pageId="8" pageNumber="9" value="2003-11-23">23/XI/2003</date>
, coll. not available (
<collectionCode id="ED208CECFFE9FF956C7CA444FD19FD21" box="[556,672,637,663]" pageId="8" pageNumber="9">DOUFPE</collectionCode>
13837); 
<collectingRegion id="49F5DACBFFE9FF956EFEA4A4FE12FD00" box="[174,427,669,695]" country="Brazil" name="Rio Grande do Norte" pageId="8" pageNumber="9">Rio Grande do Norte</collectingRegion>
, Potiguar Basin: 
<specimenCount id="9D37DFA0FFE9FF956C26A4A4FD6DFD00" box="[630,724,669,694]" pageId="8" pageNumber="9" type="male">2 males</specimenCount>
(CL 7.1 and 
<quantity id="4CC9B9CCFFE9FF956F45A484FE3EFD60" box="[277,391,701,726]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="8" pageNumber="9" unit="mm" value="10.7">10.7 mm</quantity>
), 
<date id="FF8F32E9FFE9FF956FCCA484FD9AFD60" box="[412,547,701,726]" pageId="8" pageNumber="9" value="2003-11-23">23/XI/2003</date>
, coll. not available (
<collectionCode id="ED208CECFFE9FF956EE4A4E4FE9FFD41" box="[180,294,733,759]" pageId="8" pageNumber="9">DOUFPE</collectionCode>
13906); 
<collectingRegion id="49F5DACBFFE9FF956FC4A4E4FD2EFD40" box="[404,663,733,759]" country="Brazil" name="Rio Grande do Norte" pageId="8" pageNumber="9">Rio Grande do Norte</collectingRegion>
, Potiguar Basin: 
<specimenCount id="9D37DFA0FFE9FF956F52A4C4FEEDFCA0" box="[258,340,765,790]" pageId="8" pageNumber="9" type="male">1 male</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE9FF956FC2A4C4FDBBFCA0" box="[402,514,765,790]" metricMagnitude="-2" metricUnit="m" metricValue="1.54" pageId="8" pageNumber="9" unit="mm" value="15.4">15.4 mm</quantity>
), 
<date id="FF8F32E9FFE9FF956C46A4C4FD22FCA0" box="[534,667,765,790]" pageId="8" pageNumber="9" value="2003-11-21">21/XI/2003</date>
, coll. not available (
<collectionCode id="ED208CECFFE9FF956F76A524FE21FC81" box="[294,408,797,823]" pageId="8" pageNumber="9">DOUFPE</collectionCode>
13920); 
<collectingRegion id="49F5DACBFFE9FF956C55A524FCBAFC80" box="[517,771,797,823]" country="Brazil" name="Rio Grande do Norte" pageId="8" pageNumber="9">Rio Grande do Norte</collectingRegion>
, Potiguar Basin: 
<specimenCount id="9D37DFA0FFE9FF956F3BA504FE6BFCE0" box="[363,466,829,854]" pageId="8" pageNumber="9" type="female">1 female</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE9FF956C5BA504FDD3FCE0" box="[523,618,829,854]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="8" pageNumber="9" unit="mm" value="8.6">8.6 mm</quantity>
), 2 ovigerous females (CL 10.5 and 
<quantity id="4CC9B9CCFFE9FF956FEFA564FD96FCC0" box="[447,559,861,886]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="8" pageNumber="9" unit="mm" value="13.5">13.5 mm</quantity>
), 
<date id="FF8F32E9FFE9FF956C14A564FD73FCC0" box="[580,714,861,886]" pageId="8" pageNumber="9" value="2003-11-21">21/XI/2003</date>
, coll. not available (
<collectionCode id="ED208CECFFE9FF956F38A544FE58FC21" box="[360,481,893,919]" pageId="8" pageNumber="9">DOUFPE</collectionCode>
13927); 
<collectingRegion id="49F5DACBFFE9FF956C0CA544FCBAFC20" box="[604,771,893,918]" country="Brazil" name="Pernambuco" pageId="8" pageNumber="9">Pernambuco</collectingRegion>
, Santo Aleixo Island: 
<specimenCount id="9D37DFA0FFE9FF956FE5A5A4FD9BFC00" box="[437,546,925,950]" pageId="8" pageNumber="9" type="female">1 female</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE9FF956C35A5A4FD73FC00" box="[613,714,925,950]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="8" pageNumber="9" unit="mm" value="8.2">8.2 mm</quantity>
), 
<date id="FF8F32E9FFE9FF956CB0A5A4FEB4FC60" pageId="8" pageNumber="9" value="2007-02-06">06/ II/2007</date>
, coll. not available (
<collectionCode id="ED208CECFFE9FF956C4DA584FD2EFC61" box="[541,663,957,983]" pageId="8" pageNumber="9">DOUFPE</collectionCode>
13523); 
<collectingRegion id="49F5DACBFFE9FF956EFEA5E4FEEDFC40" box="[174,340,989,1014]" country="Brazil" name="Pernambuco" pageId="8" pageNumber="9">Pernambuco</collectingRegion>
, Santo Aleixo Island: 1 ovigerous female (CL 
<quantity id="4CC9B9CCFFE9FF956F18A5C4FE07FBA0" box="[328,446,1021,1046]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="8" pageNumber="9" unit="mm" value="11.5">11.5 mm</quantity>
), 
<specimenCount id="9D37DFA0FFE9FF956F85A5C4FDD9FBA0" box="[469,608,1021,1046]" pageId="8" pageNumber="9" type="juvenile">2 juveniles</specimenCount>
(CL 2.9 and 
<quantity id="4CC9B9CCFFE9FF956EFEA224FEB6FB80" box="[174,271,1053,1078]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="8" pageNumber="9" unit="mm" value="3.6">3.6 mm</quantity>
), 
<date id="FF8F32E9FFE9FF956F75A224FE1BFB80" box="[293,418,1053,1078]" pageId="8" pageNumber="9" value="2007-02-06">06/II/2007</date>
, coll. not available (
<collectionCode id="ED208CECFFE9FF956CC4A224FCBEFB81" box="[660,775,1053,1079]" pageId="8" pageNumber="9">DOUFPE</collectionCode>
13524); 
<collectingRegion id="49F5DACBFFE9FF956F4AA204FEDEFBE0" box="[282,359,1085,1110]" country="Brazil" name="Bahia" pageId="8" pageNumber="9">Bahia</collectingRegion>
, Corumbau, Itacolomis: 
<specimenCount id="9D37DFA0FFE9FF956CF1A204FCBFFBE0" box="[673,774,1085,1110]" pageId="8" pageNumber="9" type="male">3 males</specimenCount>
(CL 4.9 - 8.0 mm), 
<date id="FF8F32E9FFE9FF956FFAA264FD80FBC0" box="[426,569,1117,1142]" pageId="8" pageNumber="9" value="2000-02-16">16/ II/2000</date>
, P.C. Paiva leg. (
<collectionCode id="ED208CECFFE9FF956EE4A244FEBBFB20" box="[180,258,1149,1174]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="8" pageNumber="9">MNRJ</collectionCode>
16748); 
<collectingRegion id="49F5DACBFFE9FF956F24A244FD89FB20" box="[372,560,1149,1174]" country="Brazil" name="Rio de Janeiro" pageId="8" pageNumber="9">Rio de Janeiro</collectingRegion>
, Arraial do Cabo, Anjos Beach: 
<specimenCount id="9D37DFA0FFE9FF956F0CA2A4FE09FB00" box="[348,432,1181,1206]" pageId="8" pageNumber="9" type="male">1 male</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE9FF956FA0A2A4FDDBFB00" box="[496,610,1181,1206]" metricMagnitude="-2" metricUnit="m" metricValue="2.43" pageId="8" pageNumber="9" unit="mm" value="24.3">24.3 mm</quantity>
), 1 ovigerous female (CL 
<quantity id="4CC9B9CCFFE9FF956F6CA284FE15FB60" box="[316,428,1213,1238]" metricMagnitude="-2" metricUnit="m" metricValue="2.02" pageId="8" pageNumber="9" unit="mm" value="20.2">20.2 mm</quantity>
), 
<date id="FF8F32E9FFE9FF956F91A284FDF1FB60" box="[449,584,1213,1238]" pageId="8" pageNumber="9" value="2003-09-06">06/IX/2003</date>
, C.E.L. Ferreira leg. (
<collectionCode id="ED208CECFFE9FF956EBBA2E4FE80FB40" box="[235,313,1245,1270]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="8" pageNumber="9">MNRJ</collectionCode>
19254); 
<collectingRegion id="49F5DACBFFE9FF956FFCA2E4FD8BFB40" box="[428,562,1245,1271]" country="Brazil" name="Sao Paulo" pageId="8" pageNumber="9">São Paulo</collectingRegion>
, Ubatuba, Itaguá Beach: 
<specimenCount id="9D37DFA0FFE9FF956F5DA2C4FED7FAA0" box="[269,366,1277,1302]" pageId="8" pageNumber="9" type="male">2 males</specimenCount>
(CL 8.8 and 
<quantity id="4CC9B9CCFFE9FF956C45A2C4FD3EFAA0" box="[533,647,1277,1302]" metricMagnitude="-2" metricUnit="m" metricValue="1.8699999999999999" pageId="8" pageNumber="9" unit="mm" value="18.7">18.7 mm</quantity>
), 
<specimenCount id="9D37DFA0FFE9FF956CCCA2C4FCBFFAA0" box="[668,774,1277,1302]" pageId="8" pageNumber="9" type="female">1 female</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE9FF956EB4A324FEFEFA80" box="[228,327,1309,1334]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="8" pageNumber="9" unit="mm" value="8.5">8.5 mm</quantity>
), 2 ovigerous females (CL 17.2 and 
<quantity id="4CC9B9CCFFE9FF956EFEA304FEA6FAE0" box="[174,287,1341,1366]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" pageId="8" pageNumber="9" unit="mm" value="18.5">18.5 mm</quantity>
), 
<specimenCount id="9D37DFA0FFE9FF956F64A304FE10FAE0" box="[308,425,1341,1366]" pageId="8" pageNumber="9" type="juvenile">1 juvenile</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE9FF956FB8A304FDF3FAE0" box="[488,586,1341,1366]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="8" pageNumber="9" unit="mm" value="9.3">9.3 mm</quantity>
), 
<date id="FF8F32E9FFE9FF956C30A304FD70FAE0" box="[608,713,1341,1366]" bridgedPair="/" pageId="8" pageNumber="9" value="1995-12">XII/1995</date>
, coll. not available (
<collectionCode id="ED208CECFFE9FF956F0EA364FE17FAC1" box="[350,430,1373,1399]" country="Brazil" httpUri="http://grbio.org/cool/4wb1-ykjs" name="Crustacean Collection of the Department of Biology" pageId="8" pageNumber="9">CCDB</collectionCode>
103); 
<collectingRegion id="49F5DACBFFE9FF956C53A364FD30FAC0" box="[515,649,1373,1399]" country="Brazil" name="Sao Paulo" pageId="8" pageNumber="9">São Paulo</collectingRegion>
, Ubatuba, Grande Beach: 
<specimenCount id="9D37DFA0FFE9FF956F3FA344FE7BFA20" box="[367,450,1405,1430]" pageId="8" pageNumber="9" type="male">1 male</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE9FF956C51A344FDD6FA20" box="[513,623,1405,1430]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="8" pageNumber="9" unit="mm" value="11.7">11.7 mm</quantity>
), 
<date id="FF8F32E9FFE9FF956CD5A344FCBAFA20" box="[645,771,1405,1430]" pageId="8" pageNumber="9" value="2004-05-04">04/V/2004</date>
, F.L. Mantelatto leg. (
<collectionCode id="ED208CECFFE9FF956FCDA3A4FE52FA01" box="[413,491,1437,1463]" country="Brazil" httpUri="http://grbio.org/cool/4wb1-ykjs" name="Crustacean Collection of the Department of Biology" pageId="8" pageNumber="9">CCDB</collectionCode>
1421).
</paragraph>
<paragraph id="8B8E1429FFE9FF956EBAA384FF43F9C0" blockId="8.[174,776,253,1974]" pageId="8" pageNumber="9">
Additional material initially identified as 
<taxonomicName id="4C316FAAFFE9FF956CBDA384FE8EFA40" baseAuthorityName="Dana" baseAuthorityYear="1851" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="scutiformis">
<emphasis id="B945C83BFFE9FF956CBDA384FE8EFA40" italics="true" pageId="8" pageNumber="9">A. scutiformis</emphasis>
</taxonomicName>
. 
<collectingCountry id="F32654B9FFE9FF956F1BA3E4FE0BFA40" box="[331,434,1501,1526]" name="Brazil" pageId="8" pageNumber="9">BRAZIL</collectingCountry>
, 
<collectingRegion id="49F5DACBFFE9FF956F91A3E4FD3AFA40" box="[449,643,1501,1526]" country="Brazil" name="Rio de Janeiro" pageId="8" pageNumber="9">Rio de Janeiro</collectingRegion>
, Angra do Reis: 
<specimenCount id="9D37DFA0FFE9FF956EA7A3C4FEF5F9A0" box="[247,332,1533,1558]" pageId="8" pageNumber="9" type="male">1 male</specimenCount>
(CL 
<quantity id="4CC9B9CCFFE9FF956FDCA3C4FE46F9A0" box="[396,511,1533,1558]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="8" pageNumber="9" unit="mm" value="13.1">13.1 mm</quantity>
), 
<date id="FF8F32E9FFE9FF956C45A3C4FD2EF9A0" box="[533,663,1533,1558]" pageId="8" pageNumber="9" value="1966-05-22">22/V/1966</date>
, coll. not available (
<collectionCode id="ED208CECFFE9FF956F79A024FE30F981" box="[297,393,1565,1591]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="8" pageNumber="9">MZUSP</collectionCode>
2781); 1 ovigerous female (CL 
<quantity id="4CC9B9CCFFE9FF956EFEA004FEA6F9E0" box="[174,287,1597,1622]" metricMagnitude="-2" metricUnit="m" metricValue="1.56" pageId="8" pageNumber="9" unit="mm" value="15.6">15.6 mm</quantity>
), 
<date id="FF8F32E9FFE9FF956F64A004FE0DF9E0" box="[308,436,1597,1622]" pageId="8" pageNumber="9" value="1966-05-21">21/V/1966</date>
, coll. not available (
<collectionCode id="ED208CECFFE9FF956CF6A004FCBFF9E1" box="[678,774,1597,1623]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="8" pageNumber="9">MZUSP</collectionCode>
2782).
</paragraph>
<paragraph id="8B8E1429FFE9FF956EBAA044FDF1F8A0" blockId="8.[174,776,253,1974]" pageId="8" pageNumber="9">
<emphasis id="B945C83BFFE9FF956EBAA044FED1F920" box="[234,360,1661,1686]" italics="true" pageId="8" pageNumber="9">Diagnosis</emphasis>
: Rostrum short, deflexed, bifid; gastric region protuberant; hepatic region with lateral lobes curved forwards and upwards; propodus of chelipeds gaping in adult male; ambulatory pereopods subchelate.
</paragraph>
<paragraph id="8B8E1429FFE9FF946EBAA124FD5BFBA0" blockId="8.[174,776,253,1974]" lastBlockId="9.[136,738,253,1974]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<emphasis id="B945C83BFFE9FF956EBAA124FE38F880" box="[234,385,1821,1846]" italics="true" pageId="8" pageNumber="9">Description</emphasis>
: Carapace elongate (almost elliptical), subpentagonal, smooth; tubercles in gastric, sometimes absent; cardiac and intestinal region, with no apparent pattern; lateral margins nearly parallel or parallel, with 3 setiferous teeth, 1 on margin of hepatic region, 2 on branchial region, sometimes absent; antennae visible on either side of rostrum. Rostrum short, deflexed, bifid; extremity with minute spine and 1 tuft of setae, setae filling entire sinus of rostrum; long and fine setae distributed laterally on dorsal margin; row of hooked setae on each side of dorsal surface adjacent to preorbital tooth. Orbits absent; orbital region with lateral angles obtuse; preorbital lobes not pointed, elevated, curved forwards and inwards. Eyes visible from above carapace, small, mobile, with short and thick setae on dorsal surface, sometimes absent. Preorbital teeth elevated, not pointed, presence of setae with variable length and thickness along lateral margin. Postorbital teeth absent. Hepatic region usually with almost rectangular angles in dorsal view, lateral margins curved forwards and upwards, with setae of variable length and thickness along margins. Gastric region slightly elevated, with 3 small setiferous tubercles distributed in a triangle, 2 on protogastric and 1 on mesogastric region. Cardiac region not evident; obsolete tubercle, with tuft of setae of variable length and thickness. Branchial region with 2 small teeth not pointed, with short and long setae. Intestinal region smooth or 2 lateral obsolete tubercles, with 3 tufts of setae, a central and 2 lateral tufts. Basal article of antenna with base wider than extremity; following 2 articles subcylindrical, attaining end of rostrum; distal end of second article with long setae; third article with tufts of short and long setae on inner margin; flagellum slender, with setae, sometimes absent. Chelipeds less strong, short, smaller than first pair of ambulatory pereopods in females and males up to 
<quantity id="4CC9B9CCFFE9FF956DCDA364FC5AFAC0" box="[925,995,1373,1398]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="9" unit="mm" value="8.0">8 mm</quantity>
of carapace length; chelipeds short, considerably enlarged in adult males (carapace length more than 
<quantity id="4CC9B9CCFFE9FF956A62A3A4FB37FA00" box="[1074,1166,1437,1462]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="8" pageNumber="9" unit="mm" value="12.0">12 mm</quantity>
). Distal end of ischium with 1-5 thin setae on ventral surface. Merus with 3 setiferous lobes at the distal end (1 central and 2 lateral lobes); 1-2 spines on the proximal dorsal surface; a row of 1-3 thin setae on ventral surface. Carpus with setiferous external crest, sometimes smooth; 3 setiferous tubercles (1 proximal, 1 distal, 1 central); row of 7 thin setae along inner margin. Propodus enlarged in lateral margins and compressed in upper margin, less enlarged and compressed in females; fingers gaping extremity in males, almost entirely closed in females; fixed finger smooth in larger males, dentate on outer surface and smooth on inner surface in other males and females, with tufts of setae near distal end. Dactylus dentate on outer surface and smooth on inner surface, with tufts of setae near distal end. Ambulatory pereopods subchelate in larger specimens, not subchelate in smaller specimens; posteriorly decreasing in size. Merus with 3 lobes in distal end of first and second pereopods, 1 or 2 long and stout setae in the central lobe; 1or 2 setae on dorsal surface and sometimes on ventral surface. Merus of third and fourth pairs with less pronounced distal lobes, with setae on extremity, sometimes smooth. Carpus of first, second and third pairs with a setae on distal inner margin, 1 or 2 on median of dorsal surface, 1 setae on distal outer margin; fourth pair smooth; subtriangular in last 2 ambulatory pereopods. Propodus of ambulatory pereopods compressed, subtriangular, with rounded end where dactylus articulas, forming structure with dactylus; distal margin lined with setae. Subchela more pronounced from fourth to first pair. Dactylus with 2 rows of minute spines intercalated with fine setae on ventral surface; end smooth, not pointed. Anterior thoracic sternum smooth. Fourth and fifth abdominal somites fused in both genders, triangular abdomen in males, almost elliptical in females, rounded in ovigerous females; telson triangular; extremity of male first pleopod with subtriangular lobe with minute spines.
</paragraph>
<paragraph id="8B8E1429FFE8FF946E95A224FD6EF900" blockId="9.[136,738,253,1974]" pageId="9" pageNumber="10">
<emphasis id="B945C83BFFE8FF946E95A224FE40FB80" box="[197,505,1053,1078]" italics="true" pageId="9" pageNumber="10">Description of juveniles</emphasis>
: Carapace transparent, lobes and spines less pronounced than adults; rostrum not deflexed, bifid and more divergent than in adults, forming an obtuse angle and a pronounced cavity; preorbital lobes not curved forwards and inwards, with long and thick setae between preorbital lobes and between preorbital and hepatic lobes; hepatic region smooth, without rectangular angles in dorsal view, lateral margins not curved forwards and upwards; gastric region without tubercles and setae. Chelipeds not strong, articles with setae distributed arbitrarily; carpus without external crest or less pronounced, without tubercles and sometimes only with setae (row with up to 6 thin setae on inner margin); propodus not enlarged and compressed, fingers entirely closed. Ambulatory pereopods subcylindrical, with setae distributed arbitrarily; propodus with almost rounded end where dactylus articulas, but not subchelate. Abdomen almost triangular, but not so marked and thin as in males.
</paragraph>
<paragraph id="8B8E1429FFE8FF946E95A084FAD3FE00" blockId="9.[136,738,253,1974]" lastBlockId="9.[813,1415,253,1974]" pageId="9" pageNumber="10">
<emphasis id="B945C83BFFE8FF946E95A084FE86F960" box="[197,319,1725,1750]" italics="true" pageId="9" pageNumber="10">Coloration</emphasis>
: It is variable and depends entirely upon the color of the surrounding macroalgae (
<bibRefCitation id="EFA069D8FFE8FF946EDFA0C4FE71F8A0" author="Coelho PA &amp; Torres MFA" box="[143,456,1789,1815]" pageId="9" pageNumber="10" pagination="221 - 241" refId="ref12460" refString="Coelho PA, Torres MFA. 1993. Taxonomia e distribuicao das especies do genero Acanthonyx Latreille, no Brasil (Crustacea: Decapoda: Majidae). Trab Oceanogr Univ Fed Pe 22: 221 - 241." type="journal article" year="1993">Coelho and Torres 1993</bibRefCitation>
). Co-ordinated with its body ornaments, its color allows the crab to be confused with algae and difficult to recognize (
<bibRefCitation id="EFA069D8FFE8FF946EDEA164FEC4F8C0" author="Teixeira GM &amp; Fransozo V &amp; Cobo VJ &amp; Hiyodo CM" box="[142,381,1885,1910]" pageId="9" pageNumber="10" pagination="87 - 95" refId="ref14378" refString="Teixeira GM, Fransozo V, Cobo VJ, Hiyodo CM. 2009. Population features of the spider crab Acanthonyx scutiformis (Dana 1851) (Crustacea, Majoidea, Epialtidae) associated with rocky-shore algae from southeastern Brazil. PanamJAS 4 (1): 87 - 95." type="journal article" year="2009">Teixeira et al. 2009</bibRefCitation>
). Crabs are typically brown or dark green, with bits of algae, grains of sand and/or debris attached to the body. Fresh specimens can present a dark heart-shaped stain in the gastric region and two round spots in the protogastric region around the area of the tubercles. The tips of the cheliped fingers are white and red, and the hepatic region is orange. There are two darker bands in the region between the preorbital teeth.
</paragraph>
<paragraph id="8B8E1429FFE8FF946D39A784FBD2FC20" blockId="9.[813,1415,253,1974]" pageId="9" pageNumber="10">
<emphasis id="B945C83BFFE8FF946D39A784FC60FE60" box="[873,985,445,470]" italics="true" pageId="9" pageNumber="10">Remarks</emphasis>
: The species in 
<collectingCountry id="F32654B9FFE8FF946AE4A784FB42FE60" box="[1204,1275,445,470]" name="Brazil" pageId="9" pageNumber="10">Brazil</collectingCountry>
cannot be effectively separated with the current accepted suite of morphological characters (H. 
<bibRefCitation id="EFA069D8FFE8FF946B6EA7C4FC62FD80" author="Milne Edwards H." pageId="9" pageNumber="10" pagination="1 - 468" refId="ref13717" refString="Milne Edwards H. 1834. Histoire naturelle des Crustaces, comprenant l'anatomie, la physiologie et la classification de ces animaux, 1: 1 - 468." type="journal article" year="1834">Milne Edwards 1834</bibRefCitation>
; 
<bibRefCitation id="EFA069D8FFE8FF946DB7A424FBD7FD80" author="Dana JD" box="[999,1134,541,566]" pageId="9" pageNumber="10" pagination="685" refId="ref12661" refString="Dana JD. 1852. Crustacea, Part 1. In: United States Exploring Expedition during the years 1838, 1839, 1840, 1841, 1842 under the command of Charles Wilkes. U. S. N. Philadelphia 13, p. 685." type="journal article" year="1852">Dana 1852</bibRefCitation>
; 
<bibRefCitation id="EFA069D8FFE8FF946A2AA424FA9FFD80" author="Rathbun MJ" box="[1146,1318,541,566]" pageId="9" pageNumber="10" pagination="1 - 613" refId="ref14038" refString="Rathbun MJ. 1925. The spider crabs of America. Bull U S Natl Mus 129: 1 - 613." type="journal article" year="1925">Rathbun 1925</bibRefCitation>
, 
<bibRefCitation id="EFA069D8FFE8FF946B61A424FC45FDE0" author="Coelho PA &amp; Torres MFA" pageId="9" pageNumber="10" pagination="221 - 241" refId="ref12460" refString="Coelho PA, Torres MFA. 1993. Taxonomia e distribuicao das especies do genero Acanthonyx Latreille, no Brasil (Crustacea: Decapoda: Majidae). Trab Oceanogr Univ Fed Pe 22: 221 - 241." type="journal article" year="1993">Coelho and Torres 1993</bibRefCitation>
; 
<bibRefCitation id="EFA069D8FFE8FF946A59A404FB2BFDE0" author="Melo GAS" box="[1033,1170,573,598]" pageId="9" pageNumber="10" refId="ref13578" refString="Melo GAS. 1996. Manual de identificacao dos Brachyura (caranguejos e siris) do Litoral Brasileiro. Sao Paulo: Editora Pleiade / FAPESP." type="book" year="1996">Melo 1996</bibRefCitation>
), and with the male first pleopods proving not to be useful. The male first pleopod of 
<taxonomicName id="4C316FAAFFE8FF946DBDA444FB58FD20" authorityName="&quot; H. Milne Edwards" authorityYear="1834" box="[1005,1249,637,662]" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="petiverii">
<emphasis id="B945C83BFFE8FF946DBDA444FB58FD20" box="[1005,1249,637,662]" italics="true" pageId="9" pageNumber="10">Acanthonyx petiverii</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFA069D8FFE8FF946AA2A444FA3BFD20" author="Garth JS" box="[1266,1410,637,663]" pageId="9" pageNumber="10" pagination="1 - 854" refId="ref12883" refString="Garth JS. 1958. Brachyura of the Pacific coast of America; Oxyrhyncha. Allan Hancock Pac Exped 21 (1 - 2): 1 - 854." type="journal article" year="1958">Garth 1958</bibRefCitation>
: plate O, Fig. 3) is similar to that of other species, such as 
<taxonomicName id="4C316FAAFFE8FF946DC5A484FC42FD60" authorityName="Manning and Holthuis" authorityYear="1981" box="[917,1019,701,726]" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="B945C83BFFE8FF946DC5A484FC42FD60" box="[917,1019,701,726]" italics="true" pageId="9" pageNumber="10">A. minor</emphasis>
</taxonomicName>
(cf. 
<bibRefCitation id="EFA069D8FFE8FF946A63A484FA3BFD60" author="Manning RB &amp; Holthuis LB" box="[1075,1410,701,726]" pageId="9" pageNumber="10" pagination="1 - 379" refId="ref13277" refString="Manning RB, Holthuis LB. 1981. West African brachyuran crabs (Crustacea: Decapoda). Smithson Contrib Zool 306: 1 - 379." type="journal article" year="1981">Manning and Holthuis 1981</bibRefCitation>
: Fig. 64h), and clearly differs from 
<taxonomicName id="4C316FAAFFE8FF946A93A4E4FA3FFD40" authorityName="Manning and Holthuis" authorityYear="1981" box="[1219,1414,733,758]" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="depressifrons">
<emphasis id="B945C83BFFE8FF946A93A4E4FA3FFD40" box="[1219,1414,733,758]" italics="true" pageId="9" pageNumber="10">A. depressifrons</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C316FAAFFE8FF946D35A4C4FC55FCA0" authorityName="Wu, Yu &amp; Ng" authorityYear="1999" box="[869,1004,765,790]" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="formosa">
<emphasis id="B945C83BFFE8FF946D35A4C4FC55FCA0" box="[869,1004,765,790]" italics="true" pageId="9" pageNumber="10">A. formosa</emphasis>
</taxonomicName>
(cf. 
<bibRefCitation id="EFA069D8FFE8FF946A78A4C4FA38FCA0" author="Manning RB &amp; Holthuis LB" box="[1064,1409,765,790]" pageId="9" pageNumber="10" pagination="1 - 379" refId="ref13277" refString="Manning RB, Holthuis LB. 1981. West African brachyuran crabs (Crustacea: Decapoda). Smithson Contrib Zool 306: 1 - 379." type="journal article" year="1981">Manning and Holthuis 1981</bibRefCitation>
: Fig. 62f, 
<bibRefCitation id="EFA069D8FFE8FF946DCFA524FBE6FC80" author="Wu S-H &amp; Yu H-P &amp; Ng PKL" box="[927,1119,797,822]" pageId="9" pageNumber="10" pagination="193 - 202" refId="ref14700" refString="Wu S-H, Yu H-P, Ng PKL. 1999. Acanthonyx formosa, a new species of spider crab (Decapoda, Brachyura, Majidae) from seaweed beds in Taiwan. Crustaceana 72 (2): 193 - 202." type="journal article" year="1999">Wu et al. 1999</bibRefCitation>
: Fig. 3d), which means that this character can be used to recognize some species within the genus but is not informative for the species here analyzed.
</paragraph>
<paragraph id="8B8E1429FFE8FF946D39A5A4FB32F9A0" blockId="9.[813,1415,253,1974]" pageId="9" pageNumber="10">
The original description of 
<taxonomicName id="4C316FAAFFE8FF946A99A5A4FBFFFC60" authority="(H. Milne Edwards 1834)" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1834" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="petiverii">
<emphasis id="B945C83BFFE8FF946A99A5A4FAEDFC00" box="[1225,1364,925,950]" italics="true" pageId="9" pageNumber="10">A. petiverii</emphasis>
(H. 
<bibRefCitation id="EFA069D8FFE8FF946D7DA584FB86FC60" author="Milne Edwards H." box="[813,1087,957,982]" pageId="9" pageNumber="10" pagination="1 - 468" refId="ref13717" refString="Milne Edwards H. 1834. Histoire naturelle des Crustaces, comprenant l'anatomie, la physiologie et la classification de ces animaux, 1: 1 - 468." type="journal article" year="1834">Milne Edwards 1834</bibRefCitation>
)
</taxonomicName>
was short and general, addressing only features of three morphological characters (orbits, carapace and legs). We redescribed 
<taxonomicName id="4C316FAAFFE8FF946D81A224FBE4FB80" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1834" box="[977,1117,1053,1078]" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="petiverii">
<emphasis id="B945C83BFFE8FF946D81A224FBE4FB80" box="[977,1117,1053,1078]" italics="true" pageId="9" pageNumber="10">A. petiverii</emphasis>
</taxonomicName>
with more details and characters based on samples from distinct localities of its distribution. The specimens of 
<taxonomicName id="4C316FAAFFE8FF946B3CA264FC2BFB20" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1834" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="petiverii">
<emphasis id="B945C83BFFE8FF946B3CA264FC2BFB20" italics="true" pageId="9" pageNumber="10">A. petiverii</emphasis>
</taxonomicName>
studied herein agree with the original description (H. 
<bibRefCitation id="EFA069D8FFE8FF946DA6A2A4FABBFB00" author="Milne Edwards H." box="[1014,1282,1181,1206]" pageId="9" pageNumber="10" pagination="1 - 468" refId="ref13717" refString="Milne Edwards H. 1834. Histoire naturelle des Crustaces, comprenant l'anatomie, la physiologie et la classification de ces animaux, 1: 1 - 468." type="journal article" year="1834">Milne Edwards 1834</bibRefCitation>
) in lateral margin with three teeth, the anterior one bigger than the others; they were similar to those described by 
<bibRefCitation id="EFA069D8FFE8FF946D82A2C4FBD7FAA0" author="Garth JS" box="[978,1134,1277,1303]" pageId="9" pageNumber="10" pagination="1 - 854" refId="ref12883" refString="Garth JS. 1958. Brachyura of the Pacific coast of America; Oxyrhyncha. Allan Hancock Pac Exped 21 (1 - 2): 1 - 854." type="journal article" year="1958">Garth (1958)</bibRefCitation>
and 
<bibRefCitation id="EFA069D8FFE8FF946AFEA2C4FA3BFAA0" author="Hendrickx ME" box="[1198,1410,1277,1302]" pageId="9" pageNumber="10" refId="ref13013" refString="Hendrickx ME. 1999. Los cangrejos braquiuros (Crustacea: Brachyura: Majoidea y Parthenopoidea) del Pacifico Mexicano. Comision Nacional para el conocimiento y uso de la biodiversidad, Instituto de Ciencias del Mar y Limnologia Universidad Nacional Autonoma de Mexico." type="book" year="1999">Hendrickx (1999)</bibRefCitation>
, except for the size variation (the smallest specimen had a CL of 
<quantity id="4CC9B9CCFFE8FF946DEEA304FBA2FAE0" box="[958,1051,1341,1366]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="9" pageNumber="10" unit="mm" value="2.9">2.9 mm</quantity>
, and the largest specimen had a CL of 
<quantity id="4CC9B9CCFFE8FF946DC2A364FBB9FAC0" box="[914,1024,1373,1398]" metricMagnitude="-2" metricUnit="m" metricValue="2.48" pageId="9" pageNumber="10" unit="mm" value="24.8">24.8 mm</quantity>
) observed in the present study, which was less than that reported in the original description (approximately 20.0 mm) and greater than that reported by 
<bibRefCitation id="EFA069D8FFE8FF946A14A384FB5FFA60" author="Garth JS" box="[1092,1254,1469,1495]" pageId="9" pageNumber="10" pagination="1 - 854" refId="ref12883" refString="Garth JS. 1958. Brachyura of the Pacific coast of America; Oxyrhyncha. Allan Hancock Pac Exped 21 (1 - 2): 1 - 854." type="journal article" year="1958">Garth (1958)</bibRefCitation>
(CL ranging from 
<quantity id="4CC9B9CCFFE8FF946D3EA3E4FB89FA40" box="[878,1072,1501,1526]" metricMagnitude="-2" metricUnit="m" metricValue="1.955" metricValueMax="3.46" metricValueMin="0.45" pageId="9" pageNumber="10" unit="mm" value="19.55" valueMax="34.6" valueMin="4.5">4.5 to 34.6 mm</quantity>
) and 
<bibRefCitation id="EFA069D8FFE8FF946A2BA3E4FAE9FA40" author="Hendrickx ME" box="[1147,1360,1501,1526]" pageId="9" pageNumber="10" refId="ref13013" refString="Hendrickx ME. 1999. Los cangrejos braquiuros (Crustacea: Brachyura: Majoidea y Parthenopoidea) del Pacifico Mexicano. Comision Nacional para el conocimiento y uso de la biodiversidad, Instituto de Ciencias del Mar y Limnologia Universidad Nacional Autonoma de Mexico." type="book" year="1999">Hendrickx (1999)</bibRefCitation>
(CL ranging from 
<quantity id="4CC9B9CCFFE8FF946D9AA3C4FBC6F9A0" box="[970,1151,1533,1558]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" metricValueMax="11.8" metricValueMin="3.6" pageId="9" pageNumber="10" unit="mm" value="7.7" valueMax="11.8" valueMin="3.6">3.6 to 11.8 mm</quantity>
).
</paragraph>
<paragraph id="8B8E1429FFE8FF976D39A024FE34FEE0" blockId="9.[813,1415,253,1974]" lastBlockId="10.[174,775,253,1974]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
Similar to the present study, 
<bibRefCitation id="EFA069D8FFE8FF946AEAA024FAEBF980" author="Garth JS" box="[1210,1362,1565,1591]" pageId="9" pageNumber="10" pagination="1 - 854" refId="ref12883" refString="Garth JS. 1958. Brachyura of the Pacific coast of America; Oxyrhyncha. Allan Hancock Pac Exped 21 (1 - 2): 1 - 854." type="journal article" year="1958">Garth (1958)</bibRefCitation>
and 
<bibRefCitation id="EFA069D8FFE8FF946D7DA004FBD9F9E0" author="Emparanza EJM &amp; Guzman GL &amp; Ng PKL" box="[813,1120,1597,1622]" pageId="9" pageNumber="10" pagination="533 - 543" refId="ref12709" refString="Emparanza EJM, Guzman GL, Ng PKL. 2007. A redescription of Acanthonyx petiverii H. Milne Edwards, 1834, and designation of a neotype for Acanthonyx simplex Dana, 1852 (Brachyura, Majidae). Crustaceana 80 (5): 533 - 543." type="journal article" year="2007">Emparanza et al. (2007)</bibRefCitation>
observed considerable variation in the presence or absence of tufts of setae and tubercles on the carapace and in the angle of the hepatic lobe, whether acute or obtuse. Both variations in the carapace as well as its color appear to be related to the color of the host algae (
<emphasis id="B945C83BFFE8FF946D66A0C4FCDBF8A0" box="[822,866,1789,1814]" italics="true" pageId="9" pageNumber="10">e.g.</emphasis>
green, brown, or red) in the region where the crabs live (
<bibRefCitation id="EFA069D8FFE8FF946DFBA124FB78F880" author="Emparanza EJM &amp; Guzman GL &amp; Ng PKL" box="[939,1217,1821,1846]" pageId="9" pageNumber="10" pagination="533 - 543" refId="ref12709" refString="Emparanza EJM, Guzman GL, Ng PKL. 2007. A redescription of Acanthonyx petiverii H. Milne Edwards, 1834, and designation of a neotype for Acanthonyx simplex Dana, 1852 (Brachyura, Majidae). Crustaceana 80 (5): 533 - 543." type="journal article" year="2007">Emparanza et al. 2007</bibRefCitation>
). The specimens analyzed here have their colors related to the color 
<typeStatus id="548AAA8BFFE8FF946D27A164FC14F8C0" box="[887,941,1885,1910]" pageId="9" pageNumber="10">type</typeStatus>
of algae that they were taken from, brown specimens occuring on 
<taxonomicName id="4C316FAAFFE8FF946ACCA144FAE9F820" box="[1180,1360,1917,1943]" class="Phaeophyceae" family="Sargassaceae" genus="Sargassum" kingdom="Chromista" order="Fucales" pageId="9" pageNumber="10" phylum="Ochrophyta" rank="species" species="undetermined">
<emphasis id="B945C83BFFE8FF946ACCA144FA9AF821" box="[1180,1315,1917,1943]" italics="true" pageId="9" pageNumber="10">Sargassum</emphasis>
sp.
</taxonomicName>
and 
<taxonomicName id="4C316FAAFFE8FF946D7DA1A4FC7BF800" box="[813,962,1949,1974]" class="Florideophyceae" family="Cystocloniaceae" genus="Hypnea" kingdom="Plantae" order="Gigartinales" pageId="9" pageNumber="10" phylum="Rhodophyta" rank="species" species="undetermined">
<emphasis id="B945C83BFFE8FF946D7DA1A4FC34F800" box="[813,909,1949,1974]" italics="true" pageId="9" pageNumber="10">Hypnea</emphasis>
sp.
</taxonomicName>
; green specimens living on 
<taxonomicName id="4C316FAAFFE8FF976B7EA1A4FF6BFEA0" class="Phaeophyceae" family="Dictyotaceae" genus="Padina" kingdom="Chromista" lastPageId="10" lastPageNumber="11" order="Dictyotales" pageId="9" pageNumber="10" phylum="Ochrophyta" rank="species" species="undetermined">
<emphasis id="B945C83BFFE8FF946B7EA1A4FA3CF800" box="[1326,1413,1949,1974]" italics="true" pageId="9" pageNumber="10">Padina</emphasis>
sp.
</taxonomicName>
and 
<taxonomicName id="4C316FAAFFEBFF976F41A6C4FECDFEA0" box="[273,372,253,278]" class="Ulvophyceae" family="Ulvaceae" genus="Ulva" kingdom="Plantae" order="Ulvales" pageId="10" pageNumber="11" phylum="Chlorophyta" rank="species" species="undetermined">
<emphasis id="B945C83BFFEBFF976F41A6C4FEFFFEA0" box="[273,326,253,278]" italics="true" pageId="10" pageNumber="11">Ulva</emphasis>
sp.
</taxonomicName>
Finally, carapace shape variation seems to be morphological not a consequence of the ornamentation.
</paragraph>
<paragraph id="8B8E1429FFEBFF976EBAA764FD9DFDC0" blockId="10.[174,775,253,1974]" pageId="10" pageNumber="11">
Ambulatory legs of specimens analyzed herein were less cheliform in females and in young males compared with adult males, and this had already been noted was an age-related character (
<bibRefCitation id="EFA069D8FFEBFF976F6AA7E4FE44FE40" author="Rathbun MJ" box="[314,509,477,502]" pageId="10" pageNumber="11" pagination="1 - 613" refId="ref14038" refString="Rathbun MJ. 1925. The spider crabs of America. Bull U S Natl Mus 129: 1 - 613." type="journal article" year="1925">Rathbun 1925</bibRefCitation>
). 
<bibRefCitation id="EFA069D8FFEBFF976C44A7E4FF46FDA0" author="Coelho PA &amp; Torres MFA" pageId="10" pageNumber="11" pagination="221 - 241" refId="ref12460" refString="Coelho PA, Torres MFA. 1993. Taxonomia e distribuicao das especies do genero Acanthonyx Latreille, no Brasil (Crustacea: Decapoda: Majidae). Trab Oceanogr Univ Fed Pe 22: 221 - 241." type="journal article" year="1993">Coelho and Torres (1993)</bibRefCitation>
described 
<taxonomicName id="4C316FAAFFEBFF976FC0A7C4FDEBFDA0" authorityName="&quot; Coelho" authorityYear="1993" box="[400,594,509,534]" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="dissimulatus">
<emphasis id="B945C83BFFEBFF976FC0A7C4FDEBFDA0" box="[400,594,509,534]" italics="true" pageId="10" pageNumber="11">A. dissimulatus</emphasis>
</taxonomicName>
and reviewed 
<taxonomicName id="4C316FAAFFEBFF976EFEA424FEE4FD80" baseAuthorityName="Dana" baseAuthorityYear="1851" box="[174,349,541,566]" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="scutiformis">
<emphasis id="B945C83BFFEBFF976EFEA424FEE4FD80" box="[174,349,541,566]" italics="true" pageId="10" pageNumber="11">A. scutiformis</emphasis>
</taxonomicName>
, but they noted that adults of 
<taxonomicName id="4C316FAAFFEBFF976CBDA424FEB2FDE0" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1834" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="petiverii">
<emphasis id="B945C83BFFEBFF976CBDA424FEB2FDE0" italics="true" pageId="10" pageNumber="11">A. petiverii</emphasis>
</taxonomicName>
were similar to juveniles and adults of 
<taxonomicName id="4C316FAAFFEBFF976CBEA404FE94FDC0" baseAuthorityName="Dana" baseAuthorityYear="1851" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="scutiformis">
<emphasis id="B945C83BFFEBFF976CBEA404FE94FDC0" italics="true" pageId="10" pageNumber="11">A. scutiformis</emphasis>
</taxonomicName>
and 
<emphasis id="B945C83BFFEBFF976F39A464FD9DFDC0" box="[361,548,605,630]" italics="true" pageId="10" pageNumber="11">
<taxonomicName id="4C316FAAFFEBFF976F39A464FD99FDC0" authorityName="&quot; Coelho" authorityYear="1993" box="[361,544,605,630]" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="dissimulatus">A. dissimulatus</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="8B8E1429FFEBFF976EBAA444FD5AFAC0" blockId="10.[174,775,253,1974]" pageId="10" pageNumber="11">
The type locality of 
<taxonomicName id="4C316FAAFFEBFF976F8BA444FDE5FD20" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1834" box="[475,604,637,662]" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="petiverii">
<emphasis id="B945C83BFFEBFF976F8BA444FDE5FD20" box="[475,604,637,662]" italics="true" pageId="10" pageNumber="11">A. petiverii</emphasis>
</taxonomicName>
is the Antilles (H. 
<bibRefCitation id="EFA069D8FFEBFF976E8FA4A4FE53FD00" author="Milne Edwards H." box="[223,490,669,694]" pageId="10" pageNumber="11" pagination="1 - 468" refId="ref13717" refString="Milne Edwards H. 1834. Histoire naturelle des Crustaces, comprenant l'anatomie, la physiologie et la classification de ces animaux, 1: 1 - 468." type="journal article" year="1834">Milne Edwards 1834</bibRefCitation>
; 
<bibRefCitation id="EFA069D8FFEBFF976FA9A4A4FD0BFD00" author="Rathbun MJ" box="[505,690,669,694]" pageId="10" pageNumber="11" pagination="1 - 613" refId="ref14038" refString="Rathbun MJ. 1925. The spider crabs of America. Bull U S Natl Mus 129: 1 - 613." type="journal article" year="1925">Rathbun 1925</bibRefCitation>
; 
<bibRefCitation id="EFA069D8FFEBFF976C91A4A4FF48FD60" author="Garth JS" pageId="10" pageNumber="11" pagination="1 - 854" refId="ref12883" refString="Garth JS. 1958. Brachyura of the Pacific coast of America; Oxyrhyncha. Allan Hancock Pac Exped 21 (1 - 2): 1 - 854." type="journal article" year="1958">Garth 1958</bibRefCitation>
) but we did not analyze material of this region. Then, we had observed specimens from 
<collectingCountry id="F32654B9FFEBFF976EFEA4C4FEF8FCA0" box="[174,321,765,791]" name="Costa Rica" pageId="10" pageNumber="11">Costa Rica</collectingCountry>
, 
<collectingCountry id="F32654B9FFEBFF976F00A4C4FE00FCA0" box="[336,441,765,790]" name="Panama" pageId="10" pageNumber="11">Panama</collectingCountry>
and 
<collectingCountry id="F32654B9FFEBFF976FAFA4C4FD3DFCA0" box="[511,644,765,790]" name="Venezuela" pageId="10" pageNumber="11">Venezuela</collectingCountry>
aiming to cover some Central America regions, places that are close to the type locality. We examined a possible specimen that correspond to the 
<typeStatus id="548AAA8BFFEBFF976CF2A564FCBEFCC0" box="[674,775,861,886]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
in the Muséum National d’Histoire Naturelle via recent photos made by the curator, but doubts raised about its origin due to lack of information available in the original label, which lead us uncomfortable and with uncertainties during our analysis. The type locality of 
<taxonomicName id="4C316FAAFFEBFF976C43A224FD73FB80" authorityName="&quot; Coelho" authorityYear="1993" box="[531,714,1053,1078]" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="dissimulatus">
<emphasis id="B945C83BFFEBFF976C43A224FD73FB80" box="[531,714,1053,1078]" italics="true" pageId="10" pageNumber="11">A. dissimulatus</emphasis>
</taxonomicName>
is in 
<collectingRegion id="49F5DACBFFEBFF976EFEA204FEE7FBE1" box="[174,350,1085,1111]" country="Brazil" name="Paraiba" pageId="10" pageNumber="11">Paraíba State</collectingRegion>
(
<collectingCountry id="F32654B9FFEBFF976F20A204FE79FBE0" box="[368,448,1085,1110]" name="Brazil" pageId="10" pageNumber="11">Brazil</collectingCountry>
), and the 
<typeStatus id="548AAA8BFFEBFF976C13A204FD17FBE0" box="[579,686,1085,1110]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
(male, 
<collectionCode id="ED208CECFFEBFF976EFEA264FEABFBC1" box="[174,274,1117,1143]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="10" pageNumber="11">MZUSP</collectionCode>
6596) was supposedly deposited in Museu de Zoologia da Universidade de 
<collectingRegion id="49F5DACBFFEBFF976CDBA244FCBEFB20" box="[651,775,1149,1175]" country="Brazil" name="Sao Paulo" pageId="10" pageNumber="11">São Paulo</collectingRegion>
(
<bibRefCitation id="EFA069D8FFEBFF976EE4A2A4FE5CFB00" author="Coelho PA &amp; Torres MFA" box="[180,485,1181,1207]" pageId="10" pageNumber="11" pagination="221 - 241" refId="ref12460" refString="Coelho PA, Torres MFA. 1993. Taxonomia e distribuicao das especies do genero Acanthonyx Latreille, no Brasil (Crustacea: Decapoda: Majidae). Trab Oceanogr Univ Fed Pe 22: 221 - 241." type="journal article" year="1993">Coelho and Torres 1993</bibRefCitation>
). The type locality of 
<taxonomicName id="4C316FAAFFEBFF976CBDA2A4FE94FB60" baseAuthorityName="Dana" baseAuthorityYear="1851" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="scutiformis">
<emphasis id="B945C83BFFEBFF976CBDA2A4FE94FB60" italics="true" pageId="10" pageNumber="11">A. scutiformis</emphasis>
</taxonomicName>
is 
<collectingRegion id="49F5DACBFFEBFF976F02A284FDF0FB61" box="[338,585,1213,1239]" country="Brazil" name="Rio de Janeiro" pageId="10" pageNumber="11">Rio de Janeiro State</collectingRegion>
(
<collectingCountry id="F32654B9FFEBFF976C09A284FD1AFB60" box="[601,675,1213,1238]" name="Brazil" pageId="10" pageNumber="11">Brazil</collectingCountry>
), and its 
<typeStatus id="548AAA8BFFEBFF976EFEA2E4FEADFB40" box="[174,276,1245,1270]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
is noted in the literature as non-existent (
<bibRefCitation id="EFA069D8FFEBFF976EE5A2C4FEF0FAA0" author="Dana JD" box="[181,329,1277,1302]" pageId="10" pageNumber="11" pagination="268 - 274" refId="ref12610" refString="Dana JD. 1851. Conspectus Crustaceorum quae in orbis Terrarum circumnavigatione, Carolo Wilkes e Classe Reipublicae Foederatae Duce, lexit et descripsit J. A. Dana - Pars VI. Am J Sci Arts, ser. 2, 11 (32): 268 - 274." type="journal article" year="1851">Dana 1851</bibRefCitation>
; 
<bibRefCitation id="EFA069D8FFEBFF976F09A2C4FDB1FAA0" author="Moreira C." box="[345,520,1277,1302]" pageId="10" pageNumber="11" pagination="1 - 151" refId="ref13832" refString="Moreira C. 1901. Contribuicoes para o conhecimento da fauna brazileira. Crustaceos do Brazil. Arq Mus Nac R Janeiro 11: 1 - 151." type="journal article" year="1901">Moreira 1901</bibRefCitation>
; 
<bibRefCitation id="EFA069D8FFEBFF976C48A2C4FF4BFA80" author="Coelho PA &amp; Torres MFA" pageId="10" pageNumber="11" pagination="221 - 241" refId="ref12460" refString="Coelho PA, Torres MFA. 1993. Taxonomia e distribuicao das especies do genero Acanthonyx Latreille, no Brasil (Crustacea: Decapoda: Majidae). Trab Oceanogr Univ Fed Pe 22: 221 - 241." type="journal article" year="1993">Coelho and Torres 1993</bibRefCitation>
; 
<bibRefCitation id="EFA069D8FFEBFF976F52A324FE2AFA80" author="Melo GAS" box="[258,403,1309,1334]" pageId="10" pageNumber="11" refId="ref13578" refString="Melo GAS. 1996. Manual de identificacao dos Brachyura (caranguejos e siris) do Litoral Brasileiro. Sao Paulo: Editora Pleiade / FAPESP." type="book" year="1996">Melo 1996</bibRefCitation>
). At this stage, we analyzed the original description but not the specimens, although we analyzed material from this locality.
</paragraph>
<paragraph id="8B8E1429FFEBFF976EBAA344FE82F800" blockId="10.[174,775,253,1974]" pageId="10" pageNumber="11">
The genus 
<taxonomicName id="4C316FAAFFEBFF976F3FA344FE43FA20" authorityName="Latreille" authorityYear="1828" box="[367,506,1405,1430]" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B945C83BFFEBFF976F3FA344FE43FA20" box="[367,506,1405,1430]" italics="true" pageId="10" pageNumber="11">Acanthonyx</emphasis>
</taxonomicName>
has a wide distribution with 17 valid species, but its systematics and taxonomy are little studied. 
<taxonomicName id="4C316FAAFFEBFF976C57A384FD46FA60" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1834" box="[519,767,1469,1494]" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="petiverii">
<emphasis id="B945C83BFFEBFF976C57A384FD46FA60" box="[519,767,1469,1494]" italics="true" pageId="10" pageNumber="11">Acanthonyx petiverii</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C316FAAFFEBFF976EFEA3E4FED3FA40" authorityName="&quot; Coelho" authorityYear="1993" box="[174,362,1501,1526]" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="dissimulatus">
<emphasis id="B945C83BFFEBFF976EFEA3E4FED3FA40" box="[174,362,1501,1526]" italics="true" pageId="10" pageNumber="11">A. dissimulatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C316FAAFFEBFF976FFEA3E4FDEFFA40" baseAuthorityName="Dana" baseAuthorityYear="1851" box="[430,598,1501,1526]" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="scutiformis">
<emphasis id="B945C83BFFEBFF976FFEA3E4FDEFFA40" box="[430,598,1501,1526]" italics="true" pageId="10" pageNumber="11">A. scutiformis</emphasis>
</taxonomicName>
are American species of this genus; however, we did not find a consistent morphological feature to differentiate these species from each other. Thus, we chose s o m e a v a i l a b l e c h a r a c t e r a n d p e r f o r m e d a morphological comparison based on species from distinct geographical distributions described and well supported in the literature (
<tableCitation id="C6B32192FFEBFF976C22A084FD6CF960" box="[626,725,1725,1750]" captionStart="Table 4" captionStartId="11.[136,204,253,278]" captionTargetPageId="11" captionText="Table 4. Morphological characters of some species of Acanthonyx Latreille, 1828. (CL: carapace length)" httpUri="http://table.plazi.org/id/DF4E44A1FFEAFF966ED8A6C4FAE1FEA0" pageId="10" pageNumber="11" tableUuid="DF4E44A1FFEAFF966ED8A6C4FAE1FEA0">Table 4</tableCitation>
): 
<taxonomicName id="4C316FAAFFEBFF976CBDA084FD74F940" authority="Manning and Holthuis, 1981" authorityName="Manning and Holthuis" authorityYear="1981" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="depressifrons">
<emphasis id="B945C83BFFEBFF976CBDA084FEE1F940" italics="true" pageId="10" pageNumber="11">A. depressifrons</emphasis>
<bibRefCitation id="EFA069D8FFEBFF976F34A0E4FD74F940" author="Manning RB &amp; Holthuis LB" box="[356,717,1757,1782]" pageId="10" pageNumber="11" pagination="1 - 379" refId="ref13277" refString="Manning RB, Holthuis LB. 1981. West African brachyuran crabs (Crustacea: Decapoda). Smithson Contrib Zool 306: 1 - 379." type="journal article" year="1981">Manning and Holthuis, 1981</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C316FAAFFEBFF976EFEA0C4FDC4F8A0" authority="Manning and Holthuis, 1981" authorityName="Manning and Holthuis" authorityYear="1981" box="[174,637,1789,1814]" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="B945C83BFFEBFF976EFEA0C4FEAFF8A0" box="[174,278,1789,1814]" italics="true" pageId="10" pageNumber="11">A. minor</emphasis>
<bibRefCitation id="EFA069D8FFEBFF976F71A0C4FDC4F8A0" author="Manning RB &amp; Holthuis LB" box="[289,637,1789,1814]" pageId="10" pageNumber="11" pagination="1 - 379" refId="ref13277" refString="Manning RB, Holthuis LB. 1981. West African brachyuran crabs (Crustacea: Decapoda). Smithson Contrib Zool 306: 1 - 379." type="journal article" year="1981">Manning and Holthuis, 1981</bibRefCitation>
</taxonomicName>
from West Africa (
<bibRefCitation id="EFA069D8FFEBFF976F50A124FDF7F880" author="Manning RB &amp; Holthuis LB" box="[256,590,1821,1846]" pageId="10" pageNumber="11" pagination="1 - 379" refId="ref13277" refString="Manning RB, Holthuis LB. 1981. West African brachyuran crabs (Crustacea: Decapoda). Smithson Contrib Zool 306: 1 - 379." type="journal article" year="1981">Manning and Holthuis 1981</bibRefCitation>
); 
<taxonomicName id="4C316FAAFFEBFF976C31A124FE0DF8E0" authority="A. Milne-Edwards, 1862" authorityName="A. Milne-Edwards" authorityYear="1862" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="limbatus">
<emphasis id="B945C83BFFEBFF976C31A124FD5CF880" box="[609,741,1821,1846]" italics="true" pageId="10" pageNumber="11">A. limbatus</emphasis>
A. Milne-Edwards, 1862
</taxonomicName>
and 
<taxonomicName id="4C316FAAFFEBFF976FA6A104FE30F8C0" authority="Griffin and Tranter, 1986" authorityName="Griffin and Tranter" authorityYear="1986" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="euryseroche">
<emphasis id="B945C83BFFEBFF976FA6A104FD0AF8E0" box="[502,691,1853,1878]" italics="true" pageId="10" pageNumber="11">A. euryseroche</emphasis>
<bibRefCitation id="EFA069D8FFEBFF976CEDA104FE30F8C0" author="Griffin DJG &amp; Tranter HA" pageId="10" pageNumber="11" pagination="1 - 335" refId="ref12913" refString="Griffin DJG, Tranter HA. 1986. The Decapoda Brachyura of the Siboga expedition. Part VIII. Majidae. Siboga Expeditie 39 (C 4): 1 - 335." type="journal article" year="1986">Griffin and Tranter, 1986</bibRefCitation>
</taxonomicName>
from Indo-West Pacific (
<bibRefCitation id="EFA069D8FFEBFF976CECA164FEC3F820" author="Griffin DJG &amp; Tranter HA" pageId="10" pageNumber="11" pagination="1 - 335" refId="ref12913" refString="Griffin DJG, Tranter HA. 1986. The Decapoda Brachyura of the Siboga expedition. Part VIII. Majidae. Siboga Expeditie 39 (C 4): 1 - 335." type="journal article" year="1986">Griffin and Tranter 1986</bibRefCitation>
); and 
<taxonomicName id="4C316FAAFFEBFF976FEEA144FD84F820" authorityName="Wu, Yu &amp; Ng" authorityYear="1999" box="[446,573,1917,1942]" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="formosa">
<emphasis id="B945C83BFFEBFF976FEEA144FD84F820" box="[446,573,1917,1942]" italics="true" pageId="10" pageNumber="11">A. formosa</emphasis>
</taxonomicName>
from 
<collectingCountry id="F32654B9FFEBFF976C2EA144FD69F820" box="[638,720,1917,1942]" name="Taiwan" pageId="10" pageNumber="11">Taiwan</collectingCountry>
(
<bibRefCitation id="EFA069D8FFEBFF976C8DA144FE89F800" author="Wu S-H &amp; Yu H-P &amp; Ng PKL" pageId="10" pageNumber="11" pagination="193 - 202" refId="ref14700" refString="Wu S-H, Yu H-P, Ng PKL. 1999. Acanthonyx formosa, a new species of spider crab (Decapoda, Brachyura, Majidae) from seaweed beds in Taiwan. Crustaceana 72 (2): 193 - 202." type="journal article" year="1999">Wu et al. 1999</bibRefCitation>
).
</paragraph>
<paragraph id="8B8E1429FFEBFF976DDFA6C4FC27FE60" blockId="10.[850,1452,253,950]" pageId="10" pageNumber="11">
We can infer that those combined character that fail to separate 
<taxonomicName id="4C316FAAFFEBFF976A6BA724FB0FFE80" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1834" box="[1083,1206,285,310]" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="petiverii">
<emphasis id="B945C83BFFEBFF976A6BA724FB0FFE80" box="[1083,1206,285,310]" italics="true" pageId="10" pageNumber="11">A. petiverii</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C316FAAFFEBFF976A95A724FAC1FE80" authorityName="&quot; Coelho" authorityYear="1993" box="[1221,1400,285,310]" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="dissimulatus">
<emphasis id="B945C83BFFEBFF976A95A724FAC1FE80" box="[1221,1400,285,310]" italics="true" pageId="10" pageNumber="11">A. dissimulatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C316FAAFFEBFF976D02A704FC4FFEE0" baseAuthorityName="Dana" baseAuthorityYear="1851" box="[850,1014,317,342]" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="scutiformis">
<emphasis id="B945C83BFFEBFF976D02A704FC4FFEE0" box="[850,1014,317,342]" italics="true" pageId="10" pageNumber="11">A. scutiformis</emphasis>
</taxonomicName>
, can be used to distinguish species from other geographic region, 
<emphasis id="B945C83BFFEBFF976A84A764FABBFEC0" box="[1236,1282,349,374]" italics="true" pageId="10" pageNumber="11">e.g.</emphasis>
shape of the rostrum plus male first pleopod can distinguish 
<taxonomicName id="4C316FAAFFEBFF976BC2A744FC4BFE00" authorityName="Manning and Holthuis" authorityYear="1981" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="depressifrons">
<emphasis id="B945C83BFFEBFF976BC2A744FC4BFE00" italics="true" pageId="10" pageNumber="11">A. depressifrons</emphasis>
</taxonomicName>
from 
<taxonomicName id="4C316FAAFFEBFF976A66A7A4FC23FE60" authority="(Manning and Holthuis 1981)" baseAuthorityName="Manning and Holthuis" baseAuthorityYear="1981" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="B945C83BFFEBFF976A66A7A4FB23FE00" box="[1078,1178,413,438]" italics="true" pageId="10" pageNumber="11">A. minor</emphasis>
(
<bibRefCitation id="EFA069D8FFEBFF976AF7A7A4FC2AFE60" author="Manning RB &amp; Holthuis LB" pageId="10" pageNumber="11" pagination="1 - 379" refId="ref13277" refString="Manning RB, Holthuis LB. 1981. West African brachyuran crabs (Crustacea: Decapoda). Smithson Contrib Zool 306: 1 - 379." type="journal article" year="1981">Manning and Holthuis 1981</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8B8E1429FFEBFF976DDFA7E4FAFEFC00" blockId="10.[850,1452,253,950]" pageId="10" pageNumber="11">
The variation observed in taxonomic relevant characters among specimens of 
<taxonomicName id="4C316FAAFFEBFF976B41A7C4FA15FDA0" authorityName="Latreille" authorityYear="1828" box="[1297,1452,509,534]" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B945C83BFFEBFF976B41A7C4FA15FDA0" box="[1297,1452,509,534]" italics="true" pageId="10" pageNumber="11">Acanthonyx</emphasis>
</taxonomicName>
from 
<collectingCountry id="F32654B9FFEBFF976DC7A424FC5DFD80" box="[919,996,541,566]" name="Brazil" pageId="10" pageNumber="11">Brazil</collectingCountry>
, allow us to question their validity. However, our molecular results suggested a clear division in two groups. Therefore, for specimens from Caribbean and 
<collectingCountry id="F32654B9FFEBFF976A05A444FB23FD20" box="[1109,1178,637,662]" name="Brazil" pageId="10" pageNumber="11">Brazil</collectingCountry>
(clade A) we suggest the synonymization of 
<taxonomicName id="4C316FAAFFEBFF976AD0A4A4FAF1FD00" authorityName="&quot; Coelho" authorityYear="1993" box="[1152,1352,669,694]" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="dissimulatus">
<emphasis id="B945C83BFFEBFF976AD0A4A4FAF1FD00" box="[1152,1352,669,694]" italics="true" pageId="10" pageNumber="11">A. dissimulatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C316FAAFFEBFF976BC1A4A4FC68FD60" baseAuthorityName="Dana" baseAuthorityYear="1851" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="scutiformis">
<emphasis id="B945C83BFFEBFF976BC1A4A4FC68FD60" italics="true" pageId="10" pageNumber="11">A. scutiformis</emphasis>
</taxonomicName>
with 
<taxonomicName id="4C316FAAFFEBFF976A44A484FB2AFD60" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1834" box="[1044,1171,701,726]" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="petiverii">
<emphasis id="B945C83BFFEBFF976A44A484FB2AFD60" box="[1044,1171,701,726]" italics="true" pageId="10" pageNumber="11">A. petiverii</emphasis>
</taxonomicName>
. We validate the name of the last one because its 
<typeStatus id="548AAA8BFFEBFF976AFAA4E4FB67FD40" box="[1194,1246,733,758]" pageId="10" pageNumber="11">type</typeStatus>
locality is in the Antilles, and it is a senior subjective synonym of 
<taxonomicName id="4C316FAAFFEBFF976BC2A4C4FC5FFC80" authorityName="&quot; Coelho" authorityYear="1993" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="dissimulatus">
<emphasis id="B945C83BFFEBFF976BC2A4C4FC5FFC80" italics="true" pageId="10" pageNumber="11">A. dissimulatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C316FAAFFEBFF976A78A524FB75FC80" baseAuthorityName="Dana" baseAuthorityYear="1851" box="[1064,1228,797,822]" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="scutiformis">
<emphasis id="B945C83BFFEBFF976A78A524FB75FC80" box="[1064,1228,797,822]" italics="true" pageId="10" pageNumber="11">A. scutiformis</emphasis>
</taxonomicName>
. Additionally, from North America (clade B), we suggest that there is a different species, here named as 
<taxonomicName id="4C316FAAFFEBFF976AA5A564FA39FCC0" authority="Mantelatto, 2016" authorityName="Mantelatto" authorityYear="2016" box="[1269,1408,861,886]" class="Malacostraca" family="Epialtidae" genus="Acanthonyx" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="B945C83BFFEBFF976AA5A564FA39FCC0" box="[1269,1408,861,886]" italics="true" pageId="10" pageNumber="11">Acanthonyx</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2767540FFEBFF976BD8A564FC3DFC20" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
because we cannot find any clear character to diagnose and chose an appropriate name.
</paragraph>
</subSubSection>
</treatment>
</document>