119 lines
13 KiB
XML
119 lines
13 KiB
XML
<document id="EBB5C5A10862806EFD5BEFB25C5E1B0E" ID-DOI="10.11646/zootaxa.4048.4.1" ID-GBIF-Dataset="bbb51c92-e184-489d-aba8-18d1dc9a25cd" ID-ISSN="1175-5326" ID-Zenodo-Dep="234023" ID-ZooBank="71E2B8DA-825E-429F-BAA0-F582702B4A80" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461348015003" checkinUser="plazi" docAuthor="López-García, Margarita M., Gasca-Álvarez, Héctor J. & Amat-García, Germán" docDate="2015" docId="990DA529FF9BFF82FF4647F913C2F90A" docLanguage="en" docName="zt04048p492.pdf" docOrigin="Zootaxa 4048 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bothynus" docType="key" docVersion="11" lastPageNumber="456" masterDocId="6534DD51FF9EFF87FFD1425F1659FFE6" masterDocTitle="The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution" masterLastPageNumber="492" masterPageNumber="451" pageNumber="456" updateTime="1698621022009" updateUser="plazi">
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<mods:title id="6AC9E31EF8FDD7775E0F92908F0FEDA4">The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution</mods:title>
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<mods:namePart id="4060C8E0C2AF06D055A62CA52664D885">López-García, Margarita M.</mods:namePart>
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<mods:namePart id="87BC3327855FD63E737A2642A0D55D40">Gasca-Álvarez, Héctor J.</mods:namePart>
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<mods:roleTerm id="B925F2B0BACA62EF9285BC626D5B7796">Author</mods:roleTerm>
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<mods:namePart id="2580DCDDF2AFF26C79E018F586582A30">Amat-García, Germán</mods:namePart>
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<mods:title id="D1726AD63AEAAE14605AC8D25B0AEFDC">Zootaxa</mods:title>
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<mods:date id="1AF09D905774211E8719828B307876C2">2015</mods:date>
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<mods:number id="16669E7628041A6F281B9F2D2FC84E8F">4048</mods:number>
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<mods:classification id="CE068AA6E7B566C6FEF47CE9138530C9">journal article</mods:classification>
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<mods:identifier id="3B193D7273023D0BF010652D714B9A84" type="DOI">10.11646/zootaxa.4048.4.1</mods:identifier>
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<treatment id="990DA529FF9BFF82FF4647F913C2F90A" ID-DOI="http://doi.org/10.5281/zenodo.5699336" ID-GBIF-Taxon="119645638" ID-Zenodo-Dep="5699336" LSID="urn:lsid:plazi:treatment:990DA529FF9BFF82FF4647F913C2F90A" httpUri="http://treatment.plazi.org/id/990DA529FF9BFF82FF4647F913C2F90A" lastPageNumber="456" pageId="5" pageNumber="456">
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<paragraph id="111B143FFF9BFF82FF4647F914A3FA26" blockId="5.[151,762,1446,1472]" box="[151,762,1446,1472]" pageId="5" pageNumber="456">
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<heading id="4A53A353FF9BFF82FF4647F914A3FA26" bold="true" box="[151,762,1446,1472]" fontSize="11" level="1" pageId="5" pageNumber="456" reason="1">
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<emphasis id="23D0C82DFF9BFF82FF4647F914A3FA26" bold="true" box="[151,762,1446,1472]" pageId="5" pageNumber="456">
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Key to the species of adult
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<taxonomicName id="D6A46FBCFF9BFF82FE3547F91401FA26" box="[484,600,1446,1472]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="genus">
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<emphasis id="23D0C82DFF9BFF82FE3547F91401FA26" bold="true" box="[484,600,1446,1472]" italics="true" pageId="5" pageNumber="456">Bothynus</emphasis>
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</taxonomicName>
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of
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<collectingCountry id="69B354AFFF9BFF82FDAE47F914A3FA26" box="[639,762,1446,1472]" name="Colombia" pageId="5" pageNumber="456">Colombia</collectingCountry>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="59BE47B4FF9BFF82FF4647B213C2F90A" pageId="5" pageNumber="456" type="key">
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<keyStep id="AA500C9AFF9BFF82FF4647B213C2F991" pageId="5" pageNumber="456">
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<paragraph id="111B143FFF9BFF82FF4647B213C5F9DA" blockId="5.[151,1437,1517,1772]" pageId="5" pageNumber="456">
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<keyLead id="AA55B70AFF9BFF82FF4647B213C5F9DA" pageId="5" pageNumber="456">
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1 Head without central tubercle. Protibia tridentate (
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<figureCitation id="899F08BAFF9BFF82FD6E47B2155FF9E4" box="[703,774,1517,1538]" captionStart="FIGURES 9 – 16" captionStartId="8.[151,264,1260,1283]" captionTargetBox="[151,1431,487,1233]" captionTargetId="figure@8.[151,1437,483,1239]" captionTargetPageId="8" captionText="FIGURES 9 – 16. Bothynus herbivorus. 9) female, habitus in dorsal view, 10) female, habitus in frontal view, 11) head of male, 12) parameres in frontal view, 13) parameres in lateral view, 14) spiculum gastrale, 15) pygidium of female, 16) locality records in Colombia." httpUri="https://zenodo.org/record/234024/files/figure.png" pageId="5" pageNumber="456">Fig. 10</figureCitation>
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). Elytra strongly punctate with deep punctures arranged in distinctive rows. Apex of metatibia truncate and without spinules. Length
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<quantity id="D65CB9DAFF9BFF82FC97445515EBF9F9" box="[838,946,1546,1567]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" metricValueMax="2.0" metricValueMin="1.4" pageId="5" pageNumber="456" unit="mm" value="17.0" valueMax="20.0" valueMin="14.0">14–20 mm</quantity>
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. Parameres elongate, apices strongly dilated (
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<figureCitation id="899F08BAFF9BFF82FABA44551747F9DA" captionStart="FIGURES 9 – 16" captionStartId="8.[151,264,1260,1283]" captionTargetBox="[151,1431,487,1233]" captionTargetId="figure@8.[151,1437,483,1239]" captionTargetPageId="8" captionText="FIGURES 9 – 16. Bothynus herbivorus. 9) female, habitus in dorsal view, 10) female, habitus in frontal view, 11) head of male, 12) parameres in frontal view, 13) parameres in lateral view, 14) spiculum gastrale, 15) pygidium of female, 16) locality records in Colombia." httpUri="https://zenodo.org/record/234024/files/figure.png" pageId="5" pageNumber="456">Figs. 12–13</figureCitation>
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)...................................................................... ..
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<taxonomicName id="D6A46FBCFF9BFF82FB51447813C5F9DA" authority="Arrow" authorityName="Arrow" box="[1152,1436,1575,1596]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="herbivorus">
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<emphasis id="23D0C82DFF9BFF82FB514478131EF9DA" box="[1152,1351,1575,1596]" italics="true" pageId="5" pageNumber="456">Bothynus herbivorus</emphasis>
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(Arrow)
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="111B143FFF9BFF82FF46441A13C2F991" blockId="5.[151,1437,1517,1772]" pageId="5" pageNumber="456">
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<keyLead id="AA55B70AFF9BFF82FF46441A13C2F991" pageId="5" pageNumber="456">
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- Head with central tubercle. Protibia quadridentate (
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<figureCitation id="899F08BAFF9BFF82FD00441A1540F9BC" box="[721,793,1605,1626]" captionStart="FIGURES 17 – 24" captionStartId="9.[151,264,1195,1218]" captionTargetBox="[163,1435,414,1169]" captionTargetId="figure@9.[151,1436,409,1174]" captionTargetPageId="9" captionText="FIGURES 17 – 24. Bothynus quadridens. 17) habitus in dorsal view, 18) habitus in frontal view, 19) head of male, 20) parameres in frontal view, 21) parameres in lateral view, 22) spiculum gastrale, 23) male inner proclaw, 24) locality records in Colombia." httpUri="https://zenodo.org/record/234025/files/figure.png" pageId="5" pageNumber="456">Fig. 18</figureCitation>
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). Elytra nearly smooth, with very small, inconspicuous punctures. Metatibial apex with irregular margin and spinules. Length>
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<quantity id="D65CB9DAFF9BFF82FCFE443D152AF991" box="[815,883,1634,1655]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="5" pageNumber="456" unit="mm" value="22.0">22 mm</quantity>
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............................................. 2
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="AA500C9AFF9BFF82FF46442013C2F90A" pageId="5" pageNumber="456">
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<paragraph id="111B143FFF9BFF82FF46442013C2F954" blockId="5.[151,1437,1517,1772]" pageId="5" pageNumber="456">
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<keyLead id="AA55B70AFF9BFF82FF46442013C2F954" pageId="5" pageNumber="456">
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<specimenCount id="07A2DFB6FF9BFF82FF4644201745F972" box="[151,284,1663,1684]" pageId="5" pageNumber="456" type="male">2 Males</specimenCount>
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with protarsal inner claw simple (Fig. 7). Parameres elongate, apices strongly dilated, lobe-shaped (Figs. 4–5). Females can be only identified by association with males...................................
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<taxonomicName id="D6A46FBCFF9BFF82FB8144C213C2F954" authority="Burmeister" authorityName="Burmeister" box="[1104,1435,1693,1714]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="complanus">
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<emphasis id="23D0C82DFF9BFF82FB8144C21342F954" box="[1104,1307,1693,1714]" italics="true" pageId="5" pageNumber="456">Bothynus complanus</emphasis>
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(Burmeister)
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="111B143FFF9BFF82FF4644E513C2F90A" blockId="5.[151,1437,1517,1772]" pageId="5" pageNumber="456">
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<keyLead id="AA55B70AFF9BFF82FF4644E513C2F90A" pageId="5" pageNumber="456">
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- Males with protarsal inner claw with median tooth (
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<figureCitation id="899F08BAFF9BFF82FD1C44E5154CF929" box="[717,789,1722,1743]" captionStart="FIGURES 17 – 24" captionStartId="9.[151,264,1195,1218]" captionTargetBox="[163,1435,414,1169]" captionTargetId="figure@9.[151,1436,409,1174]" captionTargetPageId="9" captionText="FIGURES 17 – 24. Bothynus quadridens. 17) habitus in dorsal view, 18) habitus in frontal view, 19) head of male, 20) parameres in frontal view, 21) parameres in lateral view, 22) spiculum gastrale, 23) male inner proclaw, 24) locality records in Colombia." httpUri="https://zenodo.org/record/234025/files/figure.png" pageId="5" pageNumber="456">Fig. 23</figureCitation>
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). Parameres shorter and wider, apices slightly dilated (
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<figureCitation id="899F08BAFF9BFF82FACD44E513D7F929" box="[1308,1422,1722,1743]" captionStart="FIGURES 17 – 24" captionStartId="9.[151,264,1195,1218]" captionTargetBox="[163,1435,414,1169]" captionTargetId="figure@9.[151,1436,409,1174]" captionTargetPageId="9" captionText="FIGURES 17 – 24. Bothynus quadridens. 17) habitus in dorsal view, 18) habitus in frontal view, 19) head of male, 20) parameres in frontal view, 21) parameres in lateral view, 22) spiculum gastrale, 23) male inner proclaw, 24) locality records in Colombia." httpUri="https://zenodo.org/record/234025/files/figure.png" pageId="5" pageNumber="456">Figs. 20–21</figureCitation>
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). Females can be only identified by association with males..........................
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<taxonomicName id="D6A46FBCFF9BFF82FB93448813C2F90A" authority="Taschenberg" authorityName="Taschenberg" box="[1090,1435,1751,1772]" class="Insecta" family="Dynastidae" genus="Bothynus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="456" phylum="Arthropoda" rank="species" species="quadridens">
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<emphasis id="23D0C82DFF9BFF82FB9344881354F90A" box="[1090,1293,1751,1772]" italics="true" pageId="5" pageNumber="456">Bothynus quadridens</emphasis>
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(Taschenberg)
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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</subSubSection>
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</treatment>
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