333 lines
36 KiB
XML
333 lines
36 KiB
XML
<document id="AE95C04D290AD497A06E147AB04C2FEF" ID-DOI="10.11646/zootaxa.4048.4.1" ID-GBIF-Dataset="bbb51c92-e184-489d-aba8-18d1dc9a25cd" ID-ISSN="1175-5326" ID-Zenodo-Dep="234023" ID-ZooBank="71E2B8DA-825E-429F-BAA0-F582702B4A80" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461348015003" checkinUser="plazi" docAuthor="López-García, Margarita M., Gasca-Álvarez, Héctor J. & Amat-García, Germán" docDate="2015" docId="990DA529FF92FF89FF46443617D6FEA6" docLanguage="en" docName="zt04048p492.pdf" docOrigin="Zootaxa 4048 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Euetheola humilis Burmeister 1847" docType="treatment" docVersion="11" lastPageNumber="465" masterDocId="6534DD51FF9EFF87FFD1425F1659FFE6" masterDocTitle="The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution" masterLastPageNumber="492" masterPageNumber="451" pageNumber="463" updateTime="1698621022009" updateUser="plazi">
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<mods:title id="9CF4A600AE72F9EB5776A4875DBD91FC">The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution</mods:title>
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<mods:namePart id="A1E75F7D24B9F12618ED69D622E5F2DF">López-García, Margarita M.</mods:namePart>
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<mods:namePart id="6DC9770076693CB6171B5DDEEED23B59">Gasca-Álvarez, Héctor J.</mods:namePart>
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<mods:namePart id="B55A853EA2B91E5D909904F5AFF7D653">Amat-García, Germán</mods:namePart>
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<mods:title id="CBE9FE8AAF9FB32E722645F6A90C5CF2">Zootaxa</mods:title>
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<mods:date id="23A4E979840CDB253C8435E340C972AA">2015</mods:date>
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<mods:number id="8BBA63FA21486E731BAC4B85A62A8EA7">4048</mods:number>
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<treatment id="990DA529FF92FF89FF46443617D6FEA6" ID-DOI="http://doi.org/10.5281/zenodo.5699354" ID-GBIF-Taxon="119645650" ID-Zenodo-Dep="5699354" LSID="urn:lsid:plazi:treatment:990DA529FF92FF89FF46443617D6FEA6" httpUri="http://treatment.plazi.org/id/990DA529FF92FF89FF46443617D6FEA6" lastPageId="14" lastPageNumber="465" pageId="12" pageNumber="463">
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<paragraph id="111B143FFF92FF8BFF464436143DF965" blockId="12.[151,612,1641,1700]" box="[151,612,1641,1667]" pageId="12" pageNumber="463">
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<heading id="4A53A353FF92FF8BFF464436143DF965" bold="true" box="[151,612,1641,1667]" fontSize="11" level="1" pageId="12" pageNumber="463" reason="1">
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<taxonomicName id="D6A46FBCFF92FF8BFF464436143DF965" authority="Burmeister, 1847" authorityName="Burmeister" authorityYear="1847" box="[151,612,1641,1667]" class="Insecta" family="Dynastidae" genus="Euetheola" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="humilis">
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<emphasis id="23D0C82DFF92FF8BFF464436143DF965" bold="true" box="[151,612,1641,1667]" pageId="12" pageNumber="463">
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<emphasis id="23D0C82DFF92FF8BFF464436172CF965" bold="true" box="[151,373,1641,1667]" italics="true" pageId="12" pageNumber="463">Euetheola humilis</emphasis>
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(
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<bibRefCitation id="753569CEFF92FF8BFE5544361405F965" author="Burmeister" box="[388,604,1641,1667]" pageId="12" pageNumber="463" refString="Burmeister, H. (1847) Handbuch der Entomologie. Coleoptera Lamellicornia, Xylophila et Pectinicornia, Band 5. Enslin, Berlin, Germany." type="book" year="1847">Burmeister, 1847</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="111B143FFF92FF8BFF4644D41777F942" blockId="12.[151,612,1641,1700]" box="[151,302,1675,1700]" pageId="12" pageNumber="463">
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(
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<figureCitation id="899F08BAFF92FF8BFF4E44D4177CF942" box="[159,293,1675,1700]" captionStart="FIGURES 33 – 39" captionStartId="13.[151,264,1391,1414]" captionTargetBox="[163,1428,594,1361]" captionTargetId="figure@13.[150,1436,589,1371]" captionTargetPageId="13" captionText="FIGURES 33 – 39. Euetheola humilis. 33) habitus in dorsal view, 34) habitus in frontal view, 35) parameres in frontal view, 36) parameres in lateral view, 37) spiculum gastrale, 38) head in dorsal view, 39) locality records in Colombia." httpUri="https://zenodo.org/record/234027/files/figure.png" pageId="12" pageNumber="463">
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Figs. 33
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<emphasis id="23D0C82DFF92FF8BFF2B44D41751F942" bold="true" box="[250,264,1675,1700]" pageId="12" pageNumber="463">–</emphasis>
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39
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</figureCitation>
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||
)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="59BE47B4FF92FF8BFF46448B146EF8A3" pageId="12" pageNumber="463" type="reference_group">
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<paragraph id="111B143FFF92FF8BFF46448B1455F8EE" blockId="12.[151,598,1748,1862]" pageId="12" pageNumber="463">
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<treatmentCitationGroup id="31B43311FF92FF8BFF46448B1455F8EE" pageId="12" pageNumber="463">
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<taxonomicName id="D6A46FBCFF92FF8BFF46448B140BF90C" authority="Burmeister, 1847: 93" authorityName="Burmeister" authorityPageNumber="93" authorityYear="1847" box="[151,594,1748,1770]" class="Insecta" family="Dynastidae" genus="Heteronychus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="humilis">
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<emphasis id="23D0C82DFF92FF8BFF46448B172DF90C" box="[151,372,1748,1770]" italics="true" pageId="12" pageNumber="463">Heteronychus humilis</emphasis>
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<treatmentCitation id="9005322EFF92FF8BFEAB448B140BF90C" author="Burmeister" box="[378,594,1748,1770]" page="93" pageId="12" pageNumber="463" year="1847">
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<bibRefCitation id="753569CEFF92FF8BFEAB448B1475F90C" author="Burmeister" box="[378,556,1748,1770]" pageId="12" pageNumber="463" refString="Burmeister, H. (1847) Handbuch der Entomologie. Coleoptera Lamellicornia, Xylophila et Pectinicornia, Band 5. Enslin, Berlin, Germany." type="book" year="1847">Burmeister, 1847</bibRefCitation>
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: 93
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</treatmentCitation>
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||
</taxonomicName>
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.
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<taxonomicName id="D6A46FBCFF92FF8BFF4644AC1451F8EE" authority="LeConte, 1856: 21" authorityName="LeConte" authorityPageNumber="21" authorityYear="1856" box="[151,520,1778,1800]" class="Insecta" family="Scarabaeidae" genus="Ligyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="rugiceps">
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<emphasis id="23D0C82DFF92FF8BFF4644AC171BF8EE" box="[151,322,1779,1800]" italics="true" pageId="12" pageNumber="463">Ligyrus rugiceps</emphasis>
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<treatmentCitation id="9005322EFF92FF8BFE9944AD1451F8EE" author="LeConte" box="[328,520,1778,1800]" page="21" pageId="12" pageNumber="463" year="1856">
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<bibRefCitation id="753569CEFF92FF8BFE9944AD17BBF8EE" author="LeConte" box="[328,482,1778,1800]" pageId="12" pageNumber="463" refString="LeConte, J. L. (1856) Notice of three genera of Scarabaeidae found in the United States. Proceedings of the Academy of Natural Sciences of Philadelphia, 8, 19 - 25." type="journal article" year="1856">LeConte, 1856</bibRefCitation>
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: 21
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</treatmentCitation>
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</taxonomicName>
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||
.
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||
</treatmentCitationGroup>
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</paragraph>
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||
<paragraph id="111B143FFF92FF8BFF46454D1454F8C1" blockId="12.[151,598,1748,1862]" box="[151,525,1809,1831]" pageId="12" pageNumber="463">
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<treatmentCitationGroup id="31B43311FF92FF8BFF46454D1454F8C1" box="[151,525,1809,1831]" pageId="12" pageNumber="463">
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<taxonomicName id="D6A46FBCFF92FF8BFF46454D1450F8C1" authority="Casey, 1915: 172" authorityName="Casey" authorityPageNumber="172" authorityYear="1915" box="[151,521,1809,1831]" class="Insecta" family="Dynastidae" genus="Dyscinetus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="parvus">
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<emphasis id="23D0C82DFF92FF8BFF46454D1708F8C1" box="[151,337,1810,1831]" italics="true" pageId="12" pageNumber="463">Dyscinetus parvus</emphasis>
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<treatmentCitation id="9005322EFF92FF8BFE86454E1450F8C1" author="Casey" box="[343,521,1809,1831]" page="172" pageId="12" pageNumber="463" year="1915">
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<bibRefCitation id="753569CEFF92FF8BFE86454E178FF8C1" author="Casey" box="[343,470,1809,1831]" pageId="12" pageNumber="463" refString="Casey, T. L. (1915) A review of the American species of Rutelinae, Dynastinae and Cetoniinae. Memoirs on the Coleoptera, 6, 1 - 394." type="journal article" year="1915">Casey, 1915</bibRefCitation>
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: 172
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||
</treatmentCitation>
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||
</taxonomicName>
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||
.
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||
</treatmentCitationGroup>
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||
</paragraph>
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<paragraph id="111B143FFF92FF8BFF46456E146EF8A3" blockId="12.[151,598,1748,1862]" box="[151,567,1839,1862]" pageId="12" pageNumber="463">
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||
<treatmentCitationGroup id="31B43311FF92FF8BFF46456E146EF8A3" box="[151,567,1839,1862]" pageId="12" pageNumber="463">
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||
<taxonomicName id="D6A46FBCFF92FF8BFF46456E146DF8A3" authority="Casey, 1915: 188" authorityName="Casey" authorityPageNumber="188" authorityYear="1915" box="[151,564,1839,1862]" class="Insecta" family="Dynastidae" genus="Dyscinetus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="hondurana">
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<emphasis id="23D0C82DFF92FF8BFF46456E1722F8A3" box="[151,379,1839,1862]" italics="true" pageId="12" pageNumber="463">Dyscinetus hondurana</emphasis>
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<treatmentCitation id="9005322EFF92FF8BFE534570146DF8A3" author="Casey" box="[386,564,1839,1861]" page="188" pageId="12" pageNumber="463" year="1915">
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<bibRefCitation id="753569CEFF92FF8BFE5345701458F8A3" author="Casey" box="[386,513,1839,1861]" pageId="12" pageNumber="463" refString="Casey, T. L. (1915) A review of the American species of Rutelinae, Dynastinae and Cetoniinae. Memoirs on the Coleoptera, 6, 1 - 394." type="journal article" year="1915">Casey, 1915</bibRefCitation>
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: 188
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</treatmentCitation>
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||
</taxonomicName>
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||
.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="59BE47B4FF92FF8AFF46452D157AFDFE" lastPageId="13" lastPageNumber="464" pageId="12" pageNumber="463" type="description">
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<paragraph id="111B143FFF92FF8AFF46452D157AFDFE" blockId="12.[151,1437,1906,2003]" lastBlockId="13.[151,1437,151,536]" lastPageId="13" lastPageNumber="464" pageId="12" pageNumber="463">
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<emphasis id="23D0C82DFF92FF8BFF46452D1770F86D" bold="true" box="[151,297,1906,1931]" pageId="12" pageNumber="463">Description.</emphasis>
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Habitus as in
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<figureCitation id="899F08BAFF92FF8BFE19452D1443F86D" box="[456,538,1906,1931]" captionStart="FIGURES 33 – 39" captionStartId="13.[151,264,1391,1414]" captionTargetBox="[163,1428,594,1361]" captionTargetId="figure@13.[150,1436,589,1371]" captionTargetPageId="13" captionText="FIGURES 33 – 39. Euetheola humilis. 33) habitus in dorsal view, 34) habitus in frontal view, 35) parameres in frontal view, 36) parameres in lateral view, 37) spiculum gastrale, 38) head in dorsal view, 39) locality records in Colombia." httpUri="https://zenodo.org/record/234027/files/figure.png" pageId="12" pageNumber="463">Fig. 33</figureCitation>
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. Length 13.0
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<emphasis id="23D0C82DFF92FF8BFD7D452D14E3F86D" bold="true" box="[684,698,1906,1931]" pageId="12" pageNumber="463">–</emphasis>
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<quantity id="D65CB9DAFF92FF8BFD6B452D1547F86C" box="[698,798,1906,1931]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="12" pageNumber="463" unit="mm" value="13.5">13.5 mm</quantity>
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(♂), 13.0
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||
<emphasis id="23D0C82DFF92FF8BFC5A452D15C0F86D" bold="true" box="[907,921,1906,1931]" pageId="12" pageNumber="463">–</emphasis>
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||
14.0 mm (♀). Width 6.0
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||
<emphasis id="23D0C82DFF92FF8BFB79452D12EFF86D" bold="true" box="[1192,1206,1906,1931]" pageId="12" pageNumber="463">–</emphasis>
|
||
<quantity id="D65CB9DAFF92FF8BFB67452D1355F86C" box="[1206,1292,1906,1931]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="12" pageNumber="463" unit="mm" value="6.5">6.5 mm</quantity>
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||
(♂), 6.0
|
||
<emphasis id="23D0C82DFF92FF8BFABA452D1320F86D" bold="true" box="[1387,1401,1906,1931]" pageId="12" pageNumber="463">–</emphasis>
|
||
7.0 mm (♀). Color black to reddish brown (
|
||
<figureCitation id="899F08BAFF92FF8BFD4F45C914A5F849" box="[670,764,1942,1967]" captionStart="FIGURES 33 – 39" captionStartId="13.[151,264,1391,1414]" captionTargetBox="[163,1428,594,1361]" captionTargetId="figure@13.[150,1436,589,1371]" captionTargetPageId="13" captionText="FIGURES 33 – 39. Euetheola humilis. 33) habitus in dorsal view, 34) habitus in frontal view, 35) parameres in frontal view, 36) parameres in lateral view, 37) spiculum gastrale, 38) head in dorsal view, 39) locality records in Colombia." httpUri="https://zenodo.org/record/234027/files/figure.png" pageId="12" pageNumber="463">Fig. 33</figureCitation>
|
||
).
|
||
<emphasis id="23D0C82DFF92FF8BFCCB45C91538F849" bold="true" box="[794,865,1942,1967]" pageId="12" pageNumber="463">
|
||
<emphasis id="23D0C82DFF92FF8BFCCB45C91501F849" bold="true" box="[794,856,1942,1967]" italics="true" pageId="12" pageNumber="463">Head</emphasis>
|
||
:
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||
</emphasis>
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||
Frons transversely rugopunctate and grooved. Frontoclypeal suture present; carinate, elevated, and interrupted at middle (
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||
<figureCitation id="899F08BAFF92FF8BFC3E45E5121DF835" box="[1007,1092,1978,2003]" captionStart="FIGURES 33 – 39" captionStartId="13.[151,264,1391,1414]" captionTargetBox="[163,1428,594,1361]" captionTargetId="figure@13.[150,1436,589,1371]" captionTargetPageId="13" captionText="FIGURES 33 – 39. Euetheola humilis. 33) habitus in dorsal view, 34) habitus in frontal view, 35) parameres in frontal view, 36) parameres in lateral view, 37) spiculum gastrale, 38) head in dorsal view, 39) locality records in Colombia." httpUri="https://zenodo.org/record/234027/files/figure.png" pageId="12" pageNumber="463">Fig. 38</figureCitation>
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||
). Clypeus subtrapezoidal with apex truncate; surface transversely grooved and rugose; apex with 2 small, reflexed teeth. Interocular distance equals 4.0
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<emphasis id="23D0C82DFF93FF8AFEDC42E31742FF33" bold="true" box="[269,283,188,213]" pageId="13" pageNumber="464">–</emphasis>
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||
4.8 times the transverse ocular diameter. Mandibles strongly sinuate, with a sharp tooth and basal rounded lobe. Antennal club subequal in length to antennomeres 2
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||
<emphasis id="23D0C82DFF93FF8AFC7C428015E2FF1E" bold="true" box="[941,955,223,248]" pageId="13" pageNumber="464">–</emphasis>
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||
7.
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||
<emphasis id="23D0C82DFF93FF8AFC0D42BF1203FF1E" bold="true" box="[988,1114,223,248]" pageId="13" pageNumber="464">
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||
<emphasis id="23D0C82DFF93FF8AFC0D42BF1209FF11" bold="true" box="[988,1104,224,247]" italics="true" pageId="13" pageNumber="464">Pronotum</emphasis>
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||
:
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</emphasis>
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Surface with sparse, small punctures, most of them not larger than those on vertex.
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<taxonomicName id="D6A46FBCFF93FF8AFCC8435C15ACFEFB" authority="Surface" authorityName="Surface" box="[793,1013,259,285]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Corynexochida" pageId="13" pageNumber="464" phylum="Arthropoda" rank="genus">
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<emphasis id="23D0C82DFF93FF8AFCC8435C15CEFEFB" bold="true" box="[793,919,259,285]" pageId="13" pageNumber="464">
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||
<emphasis id="23D0C82DFF93FF8AFCC8435C15D7FEFA" bold="true" box="[793,910,259,284]" italics="true" pageId="13" pageNumber="464">Scutellum</emphasis>
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||
:
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</emphasis>
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||
Surface
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</taxonomicName>
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with a few minute, sparse punctures.
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<emphasis id="23D0C82DFF93FF8AFF46437816BEFEA6" bold="true" box="[151,231,295,320]" pageId="13" pageNumber="464">
|
||
<emphasis id="23D0C82DFF93FF8AFF4643781686FEA6" bold="true" box="[151,223,295,320]" italics="true" pageId="13" pageNumber="464">Elytra</emphasis>
|
||
:
|
||
</emphasis>
|
||
Surface with 3 double rows of punctures, each small, deep and round (
|
||
<figureCitation id="899F08BAFF93FF8AFB9F437812FEFEA6" box="[1102,1191,295,320]" captionStart="FIGURES 33 – 39" captionStartId="13.[151,264,1391,1414]" captionTargetBox="[163,1428,594,1361]" captionTargetId="figure@13.[150,1436,589,1371]" captionTargetPageId="13" captionText="FIGURES 33 – 39. Euetheola humilis. 33) habitus in dorsal view, 34) habitus in frontal view, 35) parameres in frontal view, 36) parameres in lateral view, 37) spiculum gastrale, 38) head in dorsal view, 39) locality records in Colombia." httpUri="https://zenodo.org/record/234027/files/figure.png" pageId="13" pageNumber="464">Fig. 33</figureCitation>
|
||
). Lateral margin not thickened in females. Sutural stria complete, with contiguous punctures.
|
||
<taxonomicName id="D6A46FBCFF93FF8AFC2143141291FE83" authority="Surface" authorityName="Surface" box="[1008,1224,331,357]" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="13" pageNumber="464" phylum="Chordata" rank="genus">
|
||
<emphasis id="23D0C82DFF93FF8AFC214314123EFE83" bold="true" box="[1008,1127,331,357]" pageId="13" pageNumber="464">
|
||
<emphasis id="23D0C82DFF93FF8AFC2143141207FE82" bold="true" box="[1008,1118,331,356]" italics="true" pageId="13" pageNumber="464">Pygidium</emphasis>
|
||
:
|
||
</emphasis>
|
||
Surface
|
||
</taxonomicName>
|
||
densely punctate, punctures larger and denser on the basal half.
|
||
<emphasis id="23D0C82DFF93FF8AFD75432F14BAFE6E" bold="true" box="[676,739,367,392]" pageId="13" pageNumber="464">
|
||
<emphasis id="23D0C82DFF93FF8AFD75432F1483FE61" bold="true" box="[676,730,368,391]" italics="true" pageId="13" pageNumber="464">Legs</emphasis>
|
||
:
|
||
</emphasis>
|
||
Protibia tridentate, basal tooth slightly removed from others. Protarsus simple (
|
||
<figureCitation id="899F08BAFF93FF8AFEB843CB179BFE4A" box="[361,450,404,429]" captionStart="FIGURES 33 – 39" captionStartId="13.[151,264,1391,1414]" captionTargetBox="[163,1428,594,1361]" captionTargetId="figure@13.[150,1436,589,1371]" captionTargetPageId="13" captionText="FIGURES 33 – 39. Euetheola humilis. 33) habitus in dorsal view, 34) habitus in frontal view, 35) parameres in frontal view, 36) parameres in lateral view, 37) spiculum gastrale, 38) head in dorsal view, 39) locality records in Colombia." httpUri="https://zenodo.org/record/234027/files/figure.png" pageId="13" pageNumber="464">Fig. 34</figureCitation>
|
||
). Metatibia with apex slightly crenulate and with 9 spinules of nearly equal length.
|
||
<emphasis id="23D0C82DFF93FF8AFF4643E716B3FE36" bold="true" box="[151,234,439,464]" pageId="13" pageNumber="464">
|
||
<emphasis id="23D0C82DFF93FF8AFF4643E716BBFE29" bold="true" box="[151,226,440,463]" italics="true" pageId="13" pageNumber="464">Venter</emphasis>
|
||
:
|
||
</emphasis>
|
||
Prosternal process long, apex truncate and subquadrate.
|
||
<emphasis id="23D0C82DFF93FF8AFC7B43E71276FE36" bold="true" box="[938,1071,439,464]" pageId="13" pageNumber="464">
|
||
<emphasis id="23D0C82DFF93FF8AFC7B43E7127FFE29" bold="true" box="[938,1062,440,463]" italics="true" pageId="13" pageNumber="464">Parameres</emphasis>
|
||
:
|
||
</emphasis>
|
||
Basal half broad, apical half narrowed, with 2 curve and acute lateral projections, apex rounded (
|
||
<figureCitation id="899F08BAFF93FF8AFC4343831242FE13" box="[914,1051,476,501]" captionStart="FIGURES 33 – 39" captionStartId="13.[151,264,1391,1414]" captionTargetBox="[163,1428,594,1361]" captionTargetId="figure@13.[150,1436,589,1371]" captionTargetPageId="13" captionText="FIGURES 33 – 39. Euetheola humilis. 33) habitus in dorsal view, 34) habitus in frontal view, 35) parameres in frontal view, 36) parameres in lateral view, 37) spiculum gastrale, 38) head in dorsal view, 39) locality records in Colombia." httpUri="https://zenodo.org/record/234027/files/figure.png" pageId="13" pageNumber="464">
|
||
Figs. 35
|
||
<emphasis id="23D0C82DFF93FF8AFC3C438315A2FE13" bold="true" box="[1005,1019,476,501]" pageId="13" pageNumber="464">–</emphasis>
|
||
36
|
||
</figureCitation>
|
||
).
|
||
<emphasis id="23D0C82DFF93FF8AFBFF43841359FE12" bold="true" box="[1070,1280,475,500]" italics="true" pageId="13" pageNumber="464">Spiculum gastrale</emphasis>
|
||
: Base as long as lateral branches, apex broad, slightly truncate (
|
||
<figureCitation id="899F08BAFF93FF8AFD1343A0154CFDFE" box="[706,789,511,536]" captionStart="FIGURES 33 – 39" captionStartId="13.[151,264,1391,1414]" captionTargetBox="[163,1428,594,1361]" captionTargetId="figure@13.[150,1436,589,1371]" captionTargetPageId="13" captionText="FIGURES 33 – 39. Euetheola humilis. 33) habitus in dorsal view, 34) habitus in frontal view, 35) parameres in frontal view, 36) parameres in lateral view, 37) spiculum gastrale, 38) head in dorsal view, 39) locality records in Colombia." httpUri="https://zenodo.org/record/234027/files/figure.png" pageId="13" pageNumber="464">Fig. 37</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="45DB44B7FF93FF8AFF4647301289FA43" httpUri="https://zenodo.org/record/234027/files/figure.png" pageId="13" pageNumber="464" targetBox="[163,1428,594,1361]" targetPageId="13">
|
||
<paragraph id="111B143FFF93FF8AFF4647301289FA43" blockId="13.[151,1435,1391,1445]" pageId="13" pageNumber="464">
|
||
<emphasis id="23D0C82DFF93FF8AFF464730170AFA63" bold="true" box="[151,339,1391,1414]" pageId="13" pageNumber="464">FIGURES 33–39.</emphasis>
|
||
<taxonomicName id="D6A46FBCFF93FF8AFE88472F1449FA60" box="[345,528,1392,1414]" class="Insecta" family="Dynastidae" genus="Euetheola" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="464" phylum="Arthropoda" rank="species" species="humilis">
|
||
<emphasis id="23D0C82DFF93FF8AFE88472F1449FA60" box="[345,528,1392,1414]" italics="true" pageId="13" pageNumber="464">Euetheola humilis</emphasis>
|
||
</taxonomicName>
|
||
. 33) habitus in dorsal view, 34) habitus in frontal view, 35) parameres in frontal view, 36) parameres in lateral view, 37) spiculum gastrale, 38) head in dorsal view, 39) locality records in Colombia.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="59BE47B4FF93FF8AFF16478E14F9F890" pageId="13" pageNumber="464" type="diagnosis">
|
||
<paragraph id="111B143FFF93FF8AFF16478E12D3F9D7" blockId="13.[151,1436,1489,2018]" pageId="13" pageNumber="464">
|
||
<emphasis id="23D0C82DFF93FF8AFF16478E1718FA0C" bold="true" box="[199,321,1489,1514]" pageId="13" pageNumber="464">Diagnosis.</emphasis>
|
||
<taxonomicName id="D6A46FBCFF93FF8AFE9A478E1440FA0C" box="[331,537,1489,1514]" class="Insecta" family="Dynastidae" genus="Euetheola" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="464" phylum="Arthropoda" rank="species" species="humilis">
|
||
<emphasis id="23D0C82DFF93FF8AFE9A478E1440FA0C" box="[331,537,1489,1514]" italics="true" pageId="13" pageNumber="464">Euetheola humilis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="D6A46FBCFF93FF8AFD87478C14BBFA0C" box="[598,738,1489,1514]" class="Insecta" family="Dynastidae" genus="Euetheola" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="464" phylum="Arthropoda" rank="species" species="bidentata">
|
||
<emphasis id="23D0C82DFF93FF8AFD87478C143EFA0C" box="[598,615,1491,1514]" italics="true" pageId="13" pageNumber="464">E</emphasis>
|
||
.
|
||
<emphasis id="23D0C82DFF93FF8AFDA9478E14BBFA0C" box="[632,738,1489,1514]" italics="true" pageId="13" pageNumber="464">bidentata</emphasis>
|
||
</taxonomicName>
|
||
are similar species, but the former has sinuate mandibles, a cariniform frontoclypeal suture, and smaller, sparser pronotal punctation than in
|
||
<taxonomicName id="D6A46FBCFF93FF8AFBC847A912F8F9E8" box="[1049,1185,1525,1550]" class="Insecta" family="Dynastidae" genus="Euetheola" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="464" phylum="Arthropoda" rank="species" species="bidentata">
|
||
<emphasis id="23D0C82DFF93FF8AFBC847A912F8F9E8" box="[1049,1185,1525,1550]" italics="true" pageId="13" pageNumber="464">E. bidentata</emphasis>
|
||
</taxonomicName>
|
||
. The male protarsi are not enlarged, and females do not have elytral margin thickened as they are in
|
||
<taxonomicName id="D6A46FBCFF93FF8AFC2B444412DAF9D4" box="[1018,1155,1561,1586]" class="Insecta" family="Dynastidae" genus="Euetheola" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="464" phylum="Arthropoda" rank="species" species="bidentata">
|
||
<emphasis id="23D0C82DFF93FF8AFC2B444412DAF9D4" box="[1018,1155,1561,1586]" italics="true" pageId="13" pageNumber="464">E. bidentata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="111B143FFF93FF8AFF16446213D0F8C8" blockId="13.[151,1436,1489,2018]" pageId="13" pageNumber="464">
|
||
<emphasis id="23D0C82DFF93FF8AFF16446217C9F9B0" bold="true" box="[199,400,1597,1622]" pageId="13" pageNumber="464">Locality records.</emphasis>
|
||
(
|
||
<figureCitation id="899F08BAFF93FF8AFE4E446217A8F9B0" box="[415,497,1597,1622]" captionStart="FIGURES 33 – 39" captionStartId="13.[151,264,1391,1414]" captionTargetBox="[163,1428,594,1361]" captionTargetId="figure@13.[150,1436,589,1371]" captionTargetPageId="13" captionText="FIGURES 33 – 39. Euetheola humilis. 33) habitus in dorsal view, 34) habitus in frontal view, 35) parameres in frontal view, 36) parameres in lateral view, 37) spiculum gastrale, 38) head in dorsal view, 39) locality records in Colombia." httpUri="https://zenodo.org/record/234027/files/figure.png" pageId="13" pageNumber="464">Fig. 39</figureCitation>
|
||
)
|
||
<specimenCount id="07A2DFB6FF93FF8AFDD1446114C3F9B0" box="[512,666,1597,1622]" pageId="13" pageNumber="464" type="generic">34 specimens</specimenCount>
|
||
,
|
||
<specimenCount id="07A2DFB6FF93FF8AFD7444611481F9B0" box="[677,728,1598,1622]" pageId="13" pageNumber="464" type="generic">12♂</specimenCount>
|
||
, 22♀. Specimens were seen from ANDES, CTI, ICN, MEFLG, MPUJ, UPN.
|
||
<emphasis id="23D0C82DFF93FF8AFEE2443E17E4F99C" bold="true" box="[307,445,1633,1658]" pageId="13" pageNumber="464">Arauca (3):</emphasis>
|
||
Arauca, Alred. río Arauca (3).
|
||
<emphasis id="23D0C82DFF93FF8AFCCE443E15E6F99C" bold="true" box="[799,959,1633,1658]" pageId="13" pageNumber="464">Atlántico (3):</emphasis>
|
||
Barranquilla (1). Juan de Acosta, Reserva Campesina La Montaña (1). Repelón, Reserva Bijibana (1).
|
||
<emphasis id="23D0C82DFF93FF8AFCEB44DA159DF978" bold="true" box="[826,964,1669,1694]" pageId="13" pageNumber="464">Bolívar (6):</emphasis>
|
||
Mompox (5). Turbaco (1).
|
||
<emphasis id="23D0C82DFF93FF8AFB2B44DA13C5F978" bold="true" box="[1274,1436,1669,1694]" pageId="13" pageNumber="464">Casanare (7):</emphasis>
|
||
Hato Corozal, Caserío indígena Mochuelo, Selva de Galería (1).
|
||
<collectingCountry id="69B354AFFF93FF8AFC6544F6124FF924" box="[948,1046,1705,1730]" name="Trinidad and Tobago" pageId="13" pageNumber="464">Trinidad</collectingCountry>
|
||
, Fundación RN La Palmita (6).
|
||
<emphasis id="23D0C82DFF93FF8AFF4644921717F900" bold="true" box="[151,334,1741,1766]" pageId="13" pageNumber="464">Magdalena (4):</emphasis>
|
||
Sitio Nuevo, Los Cocos,
|
||
<collectingCountry id="69B354AFFF93FF8AFDA5449214C5F900" box="[628,668,1741,1766]" name="Iceland" pageId="13" pageNumber="464">Isla</collectingCountry>
|
||
de Salamanca (4).
|
||
<emphasis id="23D0C82DFF93FF8AFCA6449215BFF900" bold="true" box="[887,998,1741,1766]" pageId="13" pageNumber="464">Meta (6):</emphasis>
|
||
Los Medios, Hda. La Cabaña (1). San Martín, RN El Caduceo (4). Villavicencio, Buena Vista (1).
|
||
<emphasis id="23D0C82DFF93FF8AFC9744AE15E4F8EC" bold="true" box="[838,957,1777,1802]" pageId="13" pageNumber="464">Sucre (1):</emphasis>
|
||
San Marcos, Hda. Cocodrilia (1).
|
||
<emphasis id="23D0C82DFF93FF8AFA9844AE1698F8C8" bold="true" pageId="13" pageNumber="464">Tolima (1):</emphasis>
|
||
Ibagué (1).
|
||
<emphasis id="23D0C82DFF93FF8AFE9B454A146DF8C8" bold="true" box="[330,564,1813,1838]" pageId="13" pageNumber="464">Valle del Cauca (1):</emphasis>
|
||
Bugalagrande (1).
|
||
<emphasis id="23D0C82DFF93FF8AFCDD454A125BF8C8" bold="true" box="[780,1026,1813,1838]" pageId="13" pageNumber="464">Ambiguous data (2):</emphasis>
|
||
<collectingCountry id="69B354AFFF93FF8AFBD8454A122EF8C8" box="[1033,1143,1813,1838]" name="Colombia" pageId="13" pageNumber="464">Colombia</collectingCountry>
|
||
(1). Costa Atlántica (1).
|
||
</paragraph>
|
||
<paragraph id="111B143FFF93FF8AFF16456614F9F890" blockId="13.[151,1436,1489,2018]" pageId="13" pageNumber="464">
|
||
<emphasis id="23D0C82DFF93FF8AFF1645661781F8B4" bold="true" box="[199,472,1849,1874]" pageId="13" pageNumber="464">Temporal distribution.</emphasis>
|
||
February (3), March (4), April (6), May (1), June (4), July (1), August (1), October (1), November (4), December (5). No data (6).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="59BE47B4FF93FF89FF1645DE17D6FEA6" lastPageId="14" lastPageNumber="465" pageId="13" pageNumber="464" type="distribution">
|
||
<paragraph id="111B143FFF93FF8AFF1645DE157BF804" blockId="13.[151,1436,1489,2018]" pageId="13" pageNumber="464">
|
||
<emphasis id="23D0C82DFF93FF8AFF1645DE1739F87C" bold="true" box="[199,352,1921,1946]" pageId="13" pageNumber="464">Distribution.</emphasis>
|
||
Southern
|
||
<collectingCountry id="69B354AFFF93FF8AFE3045DE154BF87C" box="[481,786,1921,1946]" name="United States of America" pageId="13" pageNumber="464">United States of America</collectingCountry>
|
||
to
|
||
<collectingCountry id="69B354AFFF93FF8AFC9545DE15EDF87C" box="[836,948,1921,1946]" name="Argentina" pageId="13" pageNumber="464">Argentina</collectingCountry>
|
||
(
|
||
<bibRefCitation id="753569CEFF93FF8AFC1845DE1237F87C" author="Endrodi" box="[969,1134,1921,1946]" pageId="13" pageNumber="464" refString="Endrodi, S. (1969) Monographie der Dynastinae. 4. Tribus: Pentodontini (Coleoptera, Lamellicornia). Entomologische Abhandlungen Museum fur Tierkunde, Dresden, 37, 1 - 145." type="journal article" year="1969">Endrödi 1969</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="753569CEFF93FF8AFB5045DE13CEF87C" author="Ratcliffe" box="[1153,1431,1921,1946]" pageId="13" pageNumber="464" refString="Ratcliffe, B. C. & Cave, R. D. (2006) The Dynastinae Scarab Beetles of Honduras, Nicaragua and El Salvador. Bulletin of the University of Nebraska State Museum, 21, 1 - 424." type="journal article" year="2006">Ratcliffe & Cave 2006</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="753569CEFF93FF8AFF4645FA17D6F858" author="Ratcliffe" box="[151,399,1957,1982]" pageId="13" pageNumber="464" refString="Ratcliffe, B. C., Cave, R. D. & Cano, E. B. (2013) The dynastine scarab beetles of Mexico, Guatemala, and Belize (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum, 27, 1 - 666." type="journal article" year="2013">
|
||
Ratcliffe
|
||
<emphasis id="23D0C82DFF93FF8AFED745F81767F858" box="[262,318,1957,1982]" italics="true" pageId="13" pageNumber="464">et al</emphasis>
|
||
. 2013
|
||
</bibRefCitation>
|
||
). In
|
||
<collectingCountry id="69B354AFFF93FF8AFE1C45FA1467F858" box="[461,574,1957,1982]" name="Colombia" pageId="13" pageNumber="464">Colombia</collectingCountry>
|
||
, besides the eight departments recorded here, the species was found in Antioquia and Córdoba by
|
||
<bibRefCitation id="753569CEFF93FF8AFE1945961547F804" box="[456,798,1993,2018]" pageId="13" pageNumber="464" refString="Restrepo-Giraldo, H., Moron, M. A., Vallejo, L. F. & Pardo, L. C. (2003) Catalogo de Coleoptera Melolonthidae (Scarabaeidae- Pleuristicti) de Colombia. Folia Entomologica Mexicana, 2 (42), 239 - 263." type="journal article">
|
||
Restrepo-Giraldo
|
||
<emphasis id="23D0C82DFF93FF8AFD404594149BF804" box="[657,706,1993,2018]" italics="true" pageId="13" pageNumber="464">et al</emphasis>
|
||
. (2003)
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="111B143FFF90FF89FF1642C817D6FEA6" blockId="14.[151,1436,151,320]" pageId="14" pageNumber="465">
|
||
<emphasis id="23D0C82DFF90FF89FF1642C817DCFF56" bold="true" box="[199,389,151,176]" pageId="14" pageNumber="465">Natural history.</emphasis>
|
||
Several specimens were collected with light traps, between
|
||
<quantity id="D65CB9DAFF90FF89FBFB42C712D6FF49" box="[1066,1167,151,176]" metricMagnitude="2" metricUnit="m" metricValue="3.02" metricValueMax="6.0" metricValueMin="0.04" pageId="14" pageNumber="465" unit="m" value="302.0" valueMax="600.0" valueMin="4.0">
|
||
4
|
||
<emphasis id="23D0C82DFF90FF89FBE942C8121FFF56" bold="true" box="[1080,1094,151,176]" pageId="14" pageNumber="465">–</emphasis>
|
||
<specimenCount id="07A2DFB6FF90FF89FB9442C812D6FF49" box="[1093,1167,151,176]" pageId="14" pageNumber="465" type="generic">600 m</specimenCount>
|
||
</quantity>
|
||
. Some individuals were found in dry forest and gallery forest. Adults are attracted to lights. They feed on stems of grasses, and larvae are found in the soil feeding on roots (
|
||
<bibRefCitation id="753569CEFF90FF89FDC742801575FF1E" author="Ratcliffe" box="[534,812,223,248]" pageId="14" pageNumber="465" refString="Ratcliffe, B. C. & Moron, M. A. (1997) Dynastinae. In: Moron, M. A., Ratcliffe, B. C. & Deloya, C. (Eds.) Atlas de los escarabajos de Mexico. Coleoptera: Lamellicornia. Vol. I Familia Melolonthidae. Subfamilias Rutelinae, Dynastinae, Cetoniinae, Trichinae, Valginae y Melolonthinae. Comision Nacional para el Conocimiento y Uso de la Biodiversidad (CONABIO), Mexico City, Mexico." type="book" year="1997">Ratcliffe & Morón 1997</bibRefCitation>
|
||
). This species can become a pest of rice, sugarcane, and tobacco (
|
||
<bibRefCitation id="753569CEFF90FF89FF2F435B1784FEFA" author="Scavo" box="[254,477,260,285]" pageId="14" pageNumber="465" refString="Scavo, M. & Joly, L. J. (1998) Las especies venezolanas del genero Euetheola (Bates, 1888) (Coleoptera, Scarabaeidae, Dynastinae). Boletin Entomologico Venezolano, 13 (12), 173 - 186." type="journal article" year="1998">Scavo & Joly 1998</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="753569CEFF90FF89FE20435B14B1FEFB" box="[497,744,260,285]" pageId="14" pageNumber="465" refString="Bernardi, O., Garcia, M. S., Da Cunha, U. S., Back, E. C. U., Glodoaldo, A. R. & Finkenauer, E. (2006) Ocorrencia de Euetheola humilis (Burmeister) (Coleoptera: Scarabaeidae) em Eucalyptus saligna Smith (Myrtaceae), no Rio Grande do Sul. Neotropical Entomology, 37, 100 - 103. http: // dx. doi. org / 10.1590 / S 1519 - 566 X 2008000100017" type="journal article">
|
||
Bernardi
|
||
<emphasis id="23D0C82DFF90FF89FD8D435A14CCFEFB" box="[604,661,260,285]" italics="true" pageId="14" pageNumber="465">et al.</emphasis>
|
||
(2006)
|
||
</bibRefCitation>
|
||
reported adults attacking plants of
|
||
<taxonomicName id="D6A46FBCFF90FF89FBAB435B1779FEA6" authority="Smith" authorityName="Smith" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" pageId="14" pageNumber="465" phylum="Tracheophyta" rank="species" species="saligna">
|
||
<emphasis id="23D0C82DFF90FF89FBAB435B1309FEFB" box="[1146,1360,260,285]" italics="true" pageId="14" pageNumber="465">Eucalyptus saligna</emphasis>
|
||
Smith (Myrtaceae)
|
||
</taxonomicName>
|
||
in
|
||
<collectingCountry id="69B354AFFF90FF89FE92437817D2FEA6" box="[323,395,295,320]" name="Brazil" pageId="14" pageNumber="465">Brazil</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |