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<document id="214F64DFFC5258C208C8F90148210ADF" ID-CLB-Dataset="21799" ID-DOI="10.5852/ejt.2020.653" ID-GBIF-Dataset="a2776578-f968-442b-8317-9599e5884e3f" ID-ISSN="2118-9773" ID-Zenodo-Dep="3862477" ID-ZooBank="9E6EB3E1-E20F-48CB-BDD4-6859FC472382" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590670836830" checkinUser="valdenar" docAuthor="Wichard, Wilfried, Espeland, Marianne, Müller, Patrick &amp; Wang, Bo" docDate="2020" docId="8005B657FFCFFF8D6C7F457BFF6AFB7B" docLanguage="en" docName="ejt-653_wichard_espeland_mueller.pdf.imf" docOrigin="European Journal of Taxonomy 653" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Bipectinata orientalis Wichard &amp; Espeland &amp; Müller &amp; Wang 2020, gen. et sp. nov." docType="treatment" docUuid="988D3108-67F2-473D-85AB-E21A4E02DA46" docUuidSource="ZooBank" docVersion="6" lastPageNumber="15" masterDocId="7C3CCE2FFFC3FF836E444043FA51FFCF" masterDocTitle="New species of caddisflies with bipectinate antennae from Cretaceous Burmese amber (Insecta, Trichoptera: Odontoceratidae, Calamoceratidae)" masterLastPageNumber="17" masterPageNumber="1" pageNumber="13" updateTime="1698841811242" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="BA26FB2A119D3BD0FF8F156A5867B63D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="51ABABACD5527E58E7F62509DBDEB184">
<mods:title id="174A1916014C96818FCB97FF9C93257C">New species of caddisflies with bipectinate antennae from Cretaceous Burmese amber (Insecta, Trichoptera: Odontoceratidae, Calamoceratidae)</mods:title>
</mods:titleInfo>
<mods:name id="3E7401B368F27076AAC2B92F318F7F77" type="personal">
<mods:role id="FD918321011DC0CA86CA3CB44195B8BC">
<mods:roleTerm id="9AB26733FF66460C394380D1EE247CA0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D10F69253BE6AF93F6AEE2B8E97AF27A">Wichard, Wilfried</mods:namePart>
<mods:nameIdentifier id="719548BC89DDCFD711DE92CF9795A4E9" type="LSID">1E8601B2-A8F8-44A6-B185-4FB0B2E5219E</mods:nameIdentifier>
<mods:affiliation id="600EEF0A2CBF78E812AF6653DF66FC13">Institute of Biology and its Didactics, University of Cologne, Herbert Lewinstrasse 2, 50931 Köln, Germany. &amp; Email: Wichard @ uni-koeln. de &amp; urn: lsid: zoobank. org: author: 1 E 8601 B 2 - A 8 F 8 - 44 A 6 - B 185 - 4 FB 0 B 2 E 5219 E</mods:affiliation>
</mods:name>
<mods:name id="F5B7DED9E88EE1D96FD33E74FEC104DB" type="personal">
<mods:role id="6140A8F56D9EBC2FBA42BF5FC8A3CB3A">
<mods:roleTerm id="D38922F00913309E74443BBC32F2622D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="27A7AD0794BBD2B2613DEBA0FDC9CFF0">Espeland, Marianne</mods:namePart>
<mods:nameIdentifier id="3BF3EA3B9A1C3B35D6A7BFFF8BE87968" type="LSID">00D6F9F9-3902-4A8B-846F-720AB32922A6</mods:nameIdentifier>
<mods:affiliation id="6133EBD57DCE615979E16607B8F55405">Arthropoda Department, Zoological Research Museum Alexander Koenig, Adenauer-Allee 160, 53113 Bonn, Germany. &amp; urn: lsid: zoobank. org: author: 00 D 6 F 9 F 9 - 3902 - 4 A 8 B- 846 F- 720 AB 32922 A 6 &amp; Corresponding author: M. Espeland @ leibniz-zfmk. de</mods:affiliation>
</mods:name>
<mods:name id="1C2CF7E6517BFB00E1B85809977F197E" type="personal">
<mods:role id="F512C0729C2324BAC5E52BD1861AF517">
<mods:roleTerm id="37C317570179523AC1D3A888F7E6A673">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="56B1E92B32781064336B9090A262E216">Müller, Patrick</mods:namePart>
<mods:nameIdentifier id="19E18F94FA86724469C775E97F69CC06" type="LSID">CFA297B2-35D2-4C05-8F60-D73EBF620B5A</mods:nameIdentifier>
<mods:affiliation id="E27A59DE8DBE1A6E2AF0621737A2C552">Friedhofstrasse 9, 66894 Kashofen, Germany. &amp; Email: pat 14789 @ web. de &amp; urn: lsid: zoobank. org: author: CFA 297 B 2 - 35 D 2 - 4 C 05 - 8 F 60 - D 73 EBF 620 B 5 A</mods:affiliation>
<mods:nameIdentifier id="12B680CF7640B2BA9322949416D56956" type="email">pat14789@web.de</mods:nameIdentifier>
</mods:name>
<mods:name id="193B98FA9F9F4C4C732D1D46A0E68405" type="personal">
<mods:role id="B624FD1708C4924C9EE9A28B3B68FE98">
<mods:roleTerm id="CF4A981E95E3F1E70FEA34C2250C3C65">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9095ED29555CDA525BC9AB75215B25C6">Wang, Bo</mods:namePart>
<mods:nameIdentifier id="A45739614BA2FFFDC0CBAA86D47B2B3D" type="LSID">7CBE9A76-16C4-4D6D-9B4D-B2BEC000ADDF</mods:nameIdentifier>
<mods:affiliation id="070C126F837E1E31F6B617FFC7FDDF5A">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China. &amp; Email: bowang @ nigpas. ac. cn &amp; urn: lsid: zoobank. org: author: 7 CBE 9 A 76 - 16 C 4 - 4 D 6 D- 9 B 4 D-B 2 BEC 000 ADDF</mods:affiliation>
<mods:nameIdentifier id="58F6F23FB14F0ABE050A7D6376E26201" type="email">bowang@nigpas.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="976AB713311D0F7A9B0889AF8C62BC09">text</mods:typeOfResource>
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<mods:title id="00C5CA92110445DD23A0C9EEF30694B0">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="848FFEA11654299599DC045B45D38026">
<mods:date id="9F2FA317829BFF37C561752D71A99CFF">2020</mods:date>
<mods:detail id="C288ACAFBDCB1859A01E6844A22B5BEF" type="pubDate">
<mods:number id="FE9CAAA8731EB650037B16DAC92A35D0">2020-05-27</mods:number>
</mods:detail>
<mods:detail id="CEB8A395666ECA08E3A24584D64DB21A" type="volume">
<mods:number id="14A936CA83431F4D9E93C64DC506FB7D">653</mods:number>
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<mods:start id="40C9A858F0B3231BAD564247244C367D">1</mods:start>
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<mods:classification id="FDC139E2E9DE83EE4EE945ACE3F2C111">journal article</mods:classification>
<mods:identifier id="4703BE203A21421379F2AF431F278537" type="CLB-Dataset">21799</mods:identifier>
<mods:identifier id="18C873BEC0A6DA050A40B81B01330587" type="DOI">10.5852/ejt.2020.653</mods:identifier>
<mods:identifier id="BACE9F95A3A382F5706D8599B1D85360" type="GBIF-Dataset">a2776578-f968-442b-8317-9599e5884e3f</mods:identifier>
<mods:identifier id="436AF6B058DA81A6FF747B72E1133DEF" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="B61497335E26AD7C80DCF4260A4EF4CC" type="Zenodo-Dep">3862477</mods:identifier>
<mods:identifier id="15148E3F6F8753715006D72AC9E35E64" type="ZooBank">9E6EB3E1-E20F-48CB-BDD4-6859FC472382</mods:identifier>
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<treatment id="8005B657FFCFFF8D6C7F457BFF6AFB7B" ID-DOI="http://doi.org/10.5281/zenodo.3864338" ID-GBIF-Taxon="164363910" ID-Zenodo-Dep="3864338" LSID="urn:lsid:zoobank.org:act:988D3108-67F2-473D-85AB-E21A4E02DA46" httpUri="http://treatment.plazi.org/id/8005B657FFCFFF8D6C7F457BFF6AFB7B" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13">
<subSubSection id="40B654CAFFCFFF8F6C7F457BF9A9FA9D" box="[571,1016,1335,1362]" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="08130741FFCFFF8F6C7F457BF9A9FA9D" blockId="12.[367,1220,1335,1434]" box="[571,1016,1335,1362]" pageId="12" pageNumber="13">
<heading id="535BB02DFFCFFF8F6C7F457BF9A9FA9D" box="[571,1016,1335,1362]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="13" reason="2">
<taxonomicName id="CFAC7CC2FFCFFF8F6C7F457BF96EFA9D" authority="Wichard &amp; Espeland &amp; Müller &amp; Wang, 2020" authorityName="Wichard &amp; Espeland &amp; Müller &amp; Wang" authorityYear="2020" box="[571,831,1335,1362]" class="Insecta" family="Calamoceratidae" genus="Bipectinata" kingdom="Animalia" order="Trichoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="orientalis" status="gen. et sp. nov.">
<emphasis id="3AD8DB53FFCFFF8F6C7F457BF96EFA9D" bold="true" box="[571,831,1335,1362]" italics="true" pageId="12" pageNumber="13">Bipectinata orientalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="21EB6628FFCFFF8F6D03457BF9A9FA9D" box="[839,1016,1336,1362]" pageId="12" pageNumber="13" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="40B654CAFFCFFF8F6F2B4518F96DFA55" pageId="12" pageNumber="13" type="description">
<paragraph id="08130741FFCFFF8F6F2B4518FE95FAB9" blockId="12.[367,1220,1335,1434]" box="[367,1220,1371,1398]" pageId="12" pageNumber="13">
<uri id="7C3D0B43FFCFFF8F6F2B4518FE95FAB9" box="[367,1220,1371,1398]" pageId="12" pageNumber="13">
urn:lsid:zoobank.org:act:
<uuid id="7C0A3D94FFCFFF8F6CC84518FE95FAB9" box="[652,1220,1371,1398]" pageId="12" pageNumber="13">988D3108-67F2-473D-85AB-E21A4E02DA46</uuid>
</uri>
</paragraph>
<paragraph id="08130741FFCFFF8F6CB3453CF96DFA55" blockId="12.[367,1220,1335,1434]" box="[759,828,1407,1434]" pageId="12" pageNumber="13">
<figureCitation id="90971BC4FFCFFF8F6CB3453CF96DFA55" box="[759,828,1407,1434]" captionStart="Fig" captionStartId="13.[189,232,1929,1955]" captionTargetBox="[221,1369,265,1888]" captionTargetId="figure@13.[221,1369,265,1889]" captionTargetPageId="13" captionText="Fig. 5. Bipectinata orientalis sp. nov. (NIGP172206) A. Holotype male in ventral view. B. Bipectinate antennae with scapus, pedicellus and flagellomeres. C. Drawing of fore- and hindwings. D. Drawing of the male genitalia. E. Male genitalia in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3862493" httpUri="https://zenodo.org/record/3862493/files/figure.png" pageId="12" pageNumber="13">Fig. 5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="40B654CAFFCFFF8F6EF94582F89FF9C5" box="[189,718,1473,1546]" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="08130741FFCFFF8F6EF94582FB67FA14" blockId="12.[189,310,1473,1499]" box="[189,310,1473,1499]" pageId="12" pageNumber="13">
<heading id="535BB02DFFCFFF8F6EF94582FB67FA14" bold="true" box="[189,310,1473,1499]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="3AD8DB53FFCFFF8F6EF94582FB67FA14" bold="true" box="[189,310,1473,1499]" pageId="12" pageNumber="13">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="08130741FFCFFF8F6EF945B3F89FF9C5" blockId="12.[189,718,1520,1546]" box="[189,718,1520,1546]" pageId="12" pageNumber="13">See the corresponding diagnosis of the genus</paragraph>
</subSubSection>
<subSubSection id="40B654CAFFCFFF8F6EF94671FF60F9B4" box="[189,1329,1586,1659]" pageId="12" pageNumber="13" type="etymology">
<paragraph id="08130741FFCFFF8F6EF94671FB15F983" blockId="12.[189,324,1586,1612]" box="[189,324,1586,1612]" pageId="12" pageNumber="13">
<heading id="535BB02DFFCFFF8F6EF94671FB15F983" bold="true" box="[189,324,1586,1612]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="3AD8DB53FFCFFF8F6EF94671FB15F983" bold="true" box="[189,324,1586,1612]" pageId="12" pageNumber="13">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="08130741FFCFFF8F6EF94622FF60F9B4" blockId="12.[189,1329,1633,1659]" box="[189,1329,1633,1659]" pageId="12" pageNumber="13">The species name describes the biogeographic region in which this new fossil species was found.</paragraph>
</subSubSection>
<subSubSection id="40B654CAFFCFFF8F6EF946E1FB30F8FC" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="08130741FFCFFF8F6EF946E1FBFBF973" blockId="12.[189,426,1698,1724]" box="[189,426,1698,1724]" pageId="12" pageNumber="13">
<heading id="535BB02DFFCFFF8F6EF946E1FBFBF973" bold="true" box="[189,426,1698,1724]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="3AD8DB53FFCFFF8F6EF946E1FBFBF973" bold="true" box="[189,426,1698,1724]" pageId="12" pageNumber="13">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="08130741FFCFFF8F6EAE4691FB0DF923" blockId="12.[189,1398,1746,1843]" box="[234,348,1746,1772]" pageId="12" pageNumber="13">
<emphasis id="3AD8DB53FFCFFF8F6EAE4691FB0DF923" bold="true" box="[234,348,1746,1772]" pageId="12" pageNumber="13">
<typeStatus id="D717B9E3FFCFFF8F6EAE4691FB0DF923" box="[234,348,1746,1772]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="08130741FFCFFF8F6EF946B6FB30F8FC" blockId="12.[189,1398,1746,1843]" pageId="12" pageNumber="13">
<materialsCitation id="B8C40D1CFFCFFF8F6EF946B6FB30F8FC" ID-GBIF-Occurrence="2628167301" collectionCode="NIGP" country="Myanmar" location="Kachin State" pageId="12" pageNumber="13" specimenCode="NIGP172206" specimenCount="1" stateProvince="Kachin State" typeStatus="holotype">
<collectingCountry id="70BB47D1FFCFFF8F6EF946B6FB07F8C0" box="[189,342,1781,1807]" name="Myanmar" pageId="12" pageNumber="13">MYANMAR</collectingCountry>
–
<emphasis id="3AD8DB53FFCFFF8F6F3146B6F849F8C0" bold="true" box="[373,536,1781,1807]" pageId="12" pageNumber="13">
<collectingRegion id="CA68C9A3FFCFFF8F6F3146B6F849F8C0" box="[373,536,1781,1807]" country="Myanmar" name="Kachin" pageId="12" pageNumber="13">Kachin State</collectingRegion>
</emphasis>
•
<specimenCount id="1EAACCC8FFCFFF8F6C7646B5F81BF8DF" box="[562,586,1782,1808]" pageId="12" pageNumber="13" type="male">♂</specimenCount>
; exact locality unknown; Mid-Cretaceous Burmese amber inclusion;
<specimenCode id="580AAF3AFFCFFF8F6EF9475AFB0CF8FC" box="[189,349,1817,1843]" pageId="12" pageNumber="13">NIGP172206</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="40B654CAFFCFFF8D6EF94719FF6AFB7B" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13" type="description">
<paragraph id="08130741FFCFFF8F6EF94719FB1FF8BB" blockId="12.[189,334,1882,1908]" box="[189,334,1882,1908]" pageId="12" pageNumber="13">
<heading id="535BB02DFFCFFF8F6EF94719FB1FF8BB" bold="true" box="[189,334,1882,1908]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="3AD8DB53FFCFFF8F6EF94719FB1FF8BB" bold="true" box="[189,334,1882,1908]" pageId="12" pageNumber="13">Description</emphasis>
</heading>
</paragraph>
<paragraph id="08130741FFCFFF8D6EF947C9FB12FE0D" blockId="12.[189,1399,1929,2027]" lastBlockId="14.[189,1399,267,450]" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13">HEAD. Antennae about as long as forewings, partially bipectinate, first consisting of long scapus, short pedicellus and simple flagellomere longer than pedicellus, both together about as long as scapus; followed by 16 successive bipectinate flagellomeres and, finally, 18 simple flagellomeres without rami. Rami originating on both sides at base of flagellomere, basally slender and gradually expanding towards rounded apex. Maxillary palps 5-segmented, labial palps 3-segmented, their terminal segments not flexible or annulate. Head dorsally with broad posterior setose warts, transverse, covered with single, long hairs; scutum densely haired, setose warts not visible, scutellum shows single dome-shaped, large, ovoid wart.</paragraph>
<caption id="5CD357C9FFCEFF8E6EF947CAF94BF825" ID-DOI="http://doi.org/10.5281/zenodo.3862493" ID-Zenodo-Dep="3862493" httpUri="https://zenodo.org/record/3862493/files/figure.png" pageId="13" pageNumber="14" startId="13.[189,232,1929,1955]" targetBox="[221,1369,265,1888]" targetPageId="13">
<paragraph id="08130741FFCEFF8E6EF947CAF94BF825" blockId="13.[189,1399,1929,2026]" pageId="13" pageNumber="14">
<emphasis id="3AD8DB53FFCEFF8E6EF947CAFB5AF86C" bold="true" box="[189,267,1929,1955]" pageId="13" pageNumber="14">Fig. 5.</emphasis>
<taxonomicName id="CFAC7CC2FFCEFF8E6F5747C9F843F86C" authority="Wichard &amp; Espeland &amp; Müller &amp; Wang, 2020" authorityName="Wichard &amp; Espeland &amp; Müller &amp; Wang" authorityYear="2020" box="[275,530,1929,1955]" class="Insecta" family="Calamoceratidae" genus="Bipectinata" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="orientalis" status="sp. nov.">
<emphasis id="3AD8DB53FFCEFF8E6F5747C9F843F86C" box="[275,530,1929,1955]" italics="true" pageId="13" pageNumber="14">Bipectinata orientalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="21EB6628FFCEFF8E6C5E47C9F827F86C" box="[538,630,1930,1956]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
(NIGP172206)
<emphasis id="3AD8DB53FFCEFF8E6D7247CAF91AF86C" bold="true" box="[822,843,1929,1955]" pageId="13" pageNumber="14">A</emphasis>
. Holotype male in ventral view.
<emphasis id="3AD8DB53FFCEFF8E6A8A47CAFEB3F86C" bold="true" box="[1230,1250,1929,1955]" pageId="13" pageNumber="14">B</emphasis>
. Bipectinate antennae with scapus, pedicellus and flagellomeres.
<emphasis id="3AD8DB53FFCEFF8E6D6547EFF967F809" bold="true" box="[801,822,1964,1990]" pageId="13" pageNumber="14">C</emphasis>
. Drawing of fore- and hindwings.
<emphasis id="3AD8DB53FFCEFF8E6A8847EEFEB0F808" bold="true" box="[1228,1249,1965,1991]" pageId="13" pageNumber="14">D</emphasis>
. Drawing of the male genitalia.
<emphasis id="3AD8DB53FFCEFF8E6FD84793FBE1F825" bold="true" box="[412,432,2000,2026]" pageId="13" pageNumber="14">E</emphasis>
. Male genitalia in dorsal view.
</paragraph>
</caption>
<paragraph id="08130741FFCDFF8D6EF941AEFE23FCA9" blockId="14.[189,1399,493,870]" pageId="14" pageNumber="15">
FOREWINGS. Light brown, slightly translucent,
<quantity id="CF54AAA4FFCDFF8D6CA141AEF97DFDC8" box="[741,812,493,519]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="14" pageNumber="15" unit="mm" value="6.0">6 mm</quantity>
long. Radius R1 running straight to apical wing margin. Forks I and II of Rs present, sessile, branching at same level. Nygma at base of fork II. Discoidal cell closed by crossveins, about as long as its stem Rs. Media branching first into M1+2 and M3+4 and then forks III and IV, medial cell absent or open. Cu1 running straight, apically bifurcated into fork V (Cu1a–Cu1b); crossvein m-cu between Cu1a and M4, therefore thyridial cell present, very probably with nygma. Cu2 simple, not forked. Anal veins probably complete. Hindwings (
<figureCitation id="90971BC4FFCDFF8D6A3F42F3FE84FD04" box="[1147,1237,688,715]" captionStart="Fig" captionStartId="13.[189,232,1929,1955]" captionTargetBox="[221,1369,265,1888]" captionTargetId="figure@13.[221,1369,265,1889]" captionTargetPageId="13" captionText="Fig. 5. Bipectinata orientalis sp. nov. (NIGP172206) A. Holotype male in ventral view. B. Bipectinate antennae with scapus, pedicellus and flagellomeres. C. Drawing of fore- and hindwings. D. Drawing of the male genitalia. E. Male genitalia in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3862493" httpUri="https://zenodo.org/record/3862493/files/figure.png" pageId="14" pageNumber="15">Fig. 5C</figureCitation>
) light brown, translucent, smaller than forewings, about
<quantity id="CF54AAA4FFCDFF8D6CF24294F943FD3E" box="[694,786,727,754]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="14" pageNumber="15" unit="mm" value="4.5">4.5 mm</quantity>
long, Radius R1 straight, forks I, II present, fork II with nygma basally; discoidal cell absent or open. M branched in fork III and in M3+4. Cu1 with fork V, Cu2 simple. Crossveins not clearly visible, but indicated in forewings through fine, small hairs, running parallel to each other and forming ‘staggered bridge’ between longitudinal veins.
</paragraph>
<paragraph id="08130741FFCDFF8D6EF943D1FBCFFC64" blockId="14.[189,414,913,940]" box="[189,414,913,940]" pageId="14" pageNumber="15">TIBIAL SPURS. 2/4/4.</paragraph>
<paragraph id="08130741FFCDFF8D6EF94394FF6AFB7B" blockId="14.[189,1399,982,1204]" pageId="14" pageNumber="15">MALE GENITALIA. Visible only in ventral view: Inferior appendages (gonocoxites) 2-segmented consisting of straight digitiform coxopodite and curved rod-shaped harpago, length ratio 3:1; harpago initially thin and gradually enlarging into small club. Pair of long needles situated between inferior appendages, slightly dorsally placed and curved medio-dorsad, sitting on slightly stronger pedestal. In ventral view central phallic apparatus is possibly covered by 2 membranous dorsal scales, probably derived from the Xth segment, and laterally flanked by 2 short parameres [phallic apparatus itself remains unclear].</paragraph>
</subSubSection>
</treatment>
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