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<document id="8453AB2B71566DA8F305AAAB1A7D3A9C" ID-CLB-Dataset="68941" ID-DOI="10.5281/zenodo.5184182" ID-GBIF-Dataset="dcb37aba-9f9f-402a-91db-c1c6e310d61c" ID-ISSN="0253-116X" ID-Zenodo-Dep="5184182" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="juliana" checkinTime="1628735848729" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2015" docId="03915A153A17FF81FF0AEC3DFC41FC0F" docLanguage="en" docName="LinzerbiolBeitr.47.2.1133-1178.pdf" docOrigin="Linzer biologische Beiträge 47 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Lithocharis sordidula HERMAN 2003" docType="treatment" docVersion="10" lastPageNumber="1157" masterDocId="FFA8226D3A00FF99FFB8EF41FFB6FFD8" masterDocTitle="A revison of the Lithocharis species of the Palaearctic, Oriental and Australian regions (Coleoptera: Staphylinidae: Paederinae: Medonina)" masterLastPageNumber="1178" masterPageNumber="1133" pageNumber="1156" updateTime="1699050958296" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="C113E8C6AFA44FC94135FC80371DC244">A revison of the Lithocharis species of the Palaearctic, Oriental and Australian regions (Coleoptera: Staphylinidae: Paederinae: Medonina)</mods:title>
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<treatment id="03915A153A17FF81FF0AEC3DFC41FC0F" ID-DOI="http://doi.org/10.5281/zenodo.6631520" ID-GBIF-Taxon="195832413" ID-Zenodo-Dep="6631520" LSID="urn:lsid:plazi:treatment:03915A153A17FF81FF0AEC3DFC41FC0F" httpUri="http://treatment.plazi.org/id/03915A153A17FF81FF0AEC3DFC41FC0F" lastPageId="24" lastPageNumber="1157" pageId="23" pageNumber="1156">
<subSubSection id="C322B8883A17FF8EFF0AEC3DFD0AFC4A" box="[178,700,892,914]" pageId="23" pageNumber="1156" type="nomenclature">
<paragraph id="8B87EB033A17FF8EFF0AEC3DFD0AFC4A" blockId="23.[178,700,892,914]" box="[178,700,892,914]" pageId="23" pageNumber="1156">
<heading id="D0CF5C6F3A17FF8EFF0AEC3DFD0AFC4A" box="[178,700,892,914]" fontSize="8" level="2" pageId="23" pageNumber="1156" reason="2">
<taxonomicName id="4C3890803A17FF8EFF0AEC3DFD81FC4A" ID-CoL="72KYX" authority="HERMAN, 2003" authorityName="HERMAN" authorityYear="2003" box="[178,567,892,914]" class="Insecta" family="Staphylinidae" genus="Lithocharis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="1156" phylum="Arthropoda" rank="species" species="sordidula">
<emphasis id="B94C37113A17FF8EFF0AEC3DFE4FFC49" bold="true" box="[178,505,892,914]" pageId="23" pageNumber="1156">
<emphasis id="B94C37113A17FF8EFF0AEC3DFE25FC4A" bold="true" box="[178,403,892,914]" italics="true" pageId="23" pageNumber="1156">Lithocharis sordidula</emphasis>
<bibRefCitation id="EFA996F23A17FF8EFE22EC3CFE4FFC49" author="HERMAN L" box="[410,505,893,914]" pageId="23" pageNumber="1156" pagination="1 - 28" refId="ref28393" refString="HERMAN L. H. (2003): Nomenclatural changes in the Paederinae (Coleoptera: Staphylinidae). - American Museum Novitates 3416: 1 - 28." type="journal article" year="2003">HERMAN</bibRefCitation>
</emphasis>
,
<emphasis id="B94C37113A17FF8EFDBCEC3CFD81FC4A" bold="true" box="[516,567,893,914]" pageId="23" pageNumber="1156">2003</emphasis>
</taxonomicName>
<emphasis id="B94C37113A17FF8EFD85EC3DFD0AFC4A" bold="true" box="[573,700,892,914]" pageId="23" pageNumber="1156">
(
<figureCitation id="1303F7863A17FF8EFDFCEC3DFD00FC4A" box="[580,694,892,914]" captionStart="Figs 60-66" captionStartId="22.[178,218,1224,1246]" captionTargetBox="[186,1081,212,1202]" captionTargetId="figure-1@22.[178,1091,212,1215]" captionTargetPageId="22" captionText="Figs 60-66: Lithocharis sordidula HERMAN (64-65: lectotype): (60) habitus; (61) forebody; (62) male sternite VII; (63) male sternite VIII; (64-65) aedeagus in lateral and in ventral view; (66) apical portion of aedeagus in ventral view. Scale bars: 60-61: 1.0 mm; 62-66: 0.2 mm. A d d i t i o n a l m a t e r i a l e x a m i n e d India: 1♂, Assam, Bhalukpong, 27°02'N, 92°35'E, 150 m, 26.V.-3.VI.2006, leg. Pacholátko (cAss); 1♂, 2♀♀, Assam, Umrongso env., 25°27'N, 92°43'E, 700 m, 3.-8.VI.2002, leg. Trýzna (cSch, cAss); 1♀, Meghalaya, 1 km E Tura, 25°30'N, 90°14'E, 500-600 m, 2.-5.V.2002, leg. Trýzna &amp; Benda (cSch). China: 1♂, Guangxi, Bama, cow dung, I.1988 (BMNH). Thailand: 2♂♂, 2♀♀, Chumphon prov., Pha To env., 9°48'N, 98°47'E, 27.III.-14.IV.1996, leg. Majer (NHMB, cAss); 1♂, Chumphon prov., Pha To env. 9°48'N, 98°47'E, 1.-21.III.1996, K. Majer leg. (NHMB); 1♂, Chumphon, Pha To env., 9°48'N, 98°47'E, III-IV.1996, at light, leg. Prudek (cSch); 1♂, 1♀, Ranong prov., Ranong: Hot Springs, 9°56'N, 98°40'E, 23.-25.II.1996, leg. Majer&quot; (NHMB, cAss). Laos: 8 exs., 240 km N Vientiane, 10 km N Luang Prabang, Mekong river, XI.1992, leg. Somsy (cSch, cAss). Singapore: 1♂, &quot;B.T.Ra.&quot; 4 m, I.1922, leg. Saunders (BMNH); 1 ex., &quot;Scott land&quot;, 5.VIII.1922, leg. Saunders (BMNH).Indonesia: 6 exs., Sulawesi Tengah, near Morowali, Ranu River area, lowland rain forest, at light, I-IV.1980, leg. Brendell (BMNH, cAss)." figureDoi="http://doi.org/10.5281/zenodo.5184202" httpUri="https://zenodo.org/record/5184202/files/figure.png" pageId="23" pageNumber="1156">Figs 60-66</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C322B8883A17FF8EFF0AECE2FD6FFC0A" pageId="23" pageNumber="1156" type="reference_group">
<paragraph id="8B87EB033A17FF8EFF0AECE2FD05FC61" blockId="23.[178,1085,930,1072]" box="[178,691,930,953]" pageId="23" pageNumber="1156">
<treatmentCitationGroup id="AB28CC2D3A17FF8EFF0AECE2FD05FC61" box="[178,691,930,953]" pageId="23" pageNumber="1156">
<treatmentCitation id="0A99CD123A17FF8EFF0AECE2FD82FC60" author="CAMERON M" box="[178,564,930,952]" page="245" pageId="23" pageNumber="1156" year="1928">
<taxonomicName id="4C3890803A17FF8EFF0AECE2FD82FC60" ID-CoL="8Y62V" authority="CAMERON, 1928: 245" authorityName="Cameron" authorityPageNumber="245" authorityYear="1928" box="[178,564,930,952]" class="Insecta" family="Staphylinidae" genus="Lithocharis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="1156" phylum="Arthropoda" rank="species" species="sordida">
<emphasis id="B94C37113A17FF8EFF0AECE2FED3FC60" box="[178,357,931,952]" italics="true" pageId="23" pageNumber="1156">Lithocharis sordida</emphasis>
<bibRefCitation id="EFA996F23A17FF8EFED3ECE2FD82FC60" author="CAMERON M" box="[363,564,930,952]" pageId="23" pageNumber="1156" pagination="244 - 247" refId="ref27756" refString="CAMERON M. (1928): On the Oriental species of the Lithocharis ochracea group of Staphylinidae (Col.). - The Entomologist's Monthly Magazine 64: 244 - 247." type="journal article" year="1928">CAMERON, 1928: 245</bibRefCitation>
</taxonomicName>
</treatmentCitation>
; preoccupied.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B87EB033A17FF8EFF0AECFCFD6FFC0A" blockId="23.[178,1085,930,1072]" box="[178,729,957,979]" pageId="23" pageNumber="1156">
<treatmentCitationGroup id="AB28CC2D3A17FF8EFF0AECFCFD6FFC0A" box="[178,729,957,979]" pageId="23" pageNumber="1156">
<treatmentCitation id="0A99CD123A17FF8EFF0AECFCFD94FC0B" author="HERMAN L" box="[178,546,957,979]" page="6" pageId="23" pageNumber="1156" year="2003">
<taxonomicName id="4C3890803A17FF8EFF0AECFCFD94FC0B" ID-CoL="72KYX" authority="HERMAN, 2003: 6" authorityName="HERMAN" authorityPageNumber="6" authorityYear="2003" box="[178,546,957,979]" class="Insecta" family="Staphylinidae" genus="Lithocharis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="1156" phylum="Arthropoda" rank="species" species="sordidula">
<emphasis id="B94C37113A17FF8EFF0AECFCFEC1FC0A" box="[178,375,957,978]" italics="true" pageId="23" pageNumber="1156">Lithocharis sordidula</emphasis>
<bibRefCitation id="EFA996F23A17FF8EFEC5ECFCFD94FC0B" author="HERMAN L" box="[381,546,957,979]" pageId="23" pageNumber="1156" pagination="1 - 28" refId="ref28393" refString="HERMAN L. H. (2003): Nomenclatural changes in the Paederinae (Coleoptera: Staphylinidae). - American Museum Novitates 3416: 1 - 28." type="journal article" year="2003">HERMAN, 2003: 6</bibRefCitation>
</taxonomicName>
</treatmentCitation>
; replacement name.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C322B8883A17FF8EFF0AEC97FD90FBE8" pageId="23" pageNumber="1156" type="materials_examined">
<paragraph id="8B87EB033A17FF8EFF0AEC97FD90FBE8" blockId="23.[178,1085,930,1072]" pageId="23" pageNumber="1156">
Type material examined:
<materialsCitation id="3B50E15E3A17FF8EFDFDEC97FD90FBE8" ID-GBIF-Occurrence="3805307497" pageId="23" pageNumber="1156" typeStatus="lectotype">
<typeStatus id="548355A13A17FF8EFDFDEC97FD15FC33" box="[581,675,982,1003]" pageId="23" pageNumber="1156" type="lectotype">Lectotype</typeStatus>
♂, present designation: &quot;Bentong,
<collectingRegion id="49FC25E13A17FF8EFC4CEC97FB8FFC34" box="[1012,1081,982,1004]" country="Malaysia" name="Pahang" pageId="23" pageNumber="1156">Pahang</collectingRegion>
, F.M.S., Dr. Cameron / Dung /
<taxonomicName id="4C3890803A17FF8EFE61ECACFDD8FBDA" authority="Cam" authorityName="Cameron" authorityYear="1928" box="[473,622,1004,1026]" class="Insecta" family="Staphylinidae" genus="Lithocharis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="1156" phylum="Arthropoda" rank="species" species="sordida">L. sordida Cam</taxonomicName>
, Type / M. Cameron. Bequest. B.M. 1955-147. / Type / Lectotypus ♂
<taxonomicName id="4C3890803A17FF8EFEC3EB42FD38FBC0" authority="Cameron" authorityName="Cameron" authorityYear="1928" box="[379,654,1027,1048]" class="Insecta" family="Staphylinidae" genus="Lithocharis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="1156" phylum="Arthropoda" rank="species" species="sordida">
<emphasis id="B94C37113A17FF8EFEC3EB42FD86FBC0" box="[379,560,1027,1048]" italics="true" pageId="23" pageNumber="1156">Lithocharis sordida</emphasis>
Cameron
</taxonomicName>
, desig. V.
<bibRefCitation id="EFA996F23A17FF8EFD4AEB42FCD3FBC1" author="ASSING V" box="[754,869,1027,1049]" pageId="23" pageNumber="1156" pagination="191 - 206" refId="ref27538" refString="ASSING V. (2015): Two new genera of Medonina from China and Japan (Coleoptera: Staphylinidae: Paederinae). - Linzer Biologische Beitrage 47 (1): 191 - 206." type="journal article" year="2015">Assing 2015</bibRefCitation>
/
<taxonomicName id="4C3890803A17FF8EFCC2EB42FF48FBF7" authority="Herman" authorityName="HERMAN" authorityYear="2003" class="Insecta" family="Staphylinidae" genus="Lithocharis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="1156" phylum="Arthropoda" rank="species" species="sordidula">Lithocharis sordidula Herman</taxonomicName>
, det. V.
<bibRefCitation id="EFA996F23A17FF8EFEF2EB5BFE08FBE8" author="ASSING V" box="[330,446,1050,1072]" pageId="23" pageNumber="1156" pagination="191 - 206" refId="ref27538" refString="ASSING V. (2015): Two new genera of Medonina from China and Japan (Coleoptera: Staphylinidae: Paederinae). - Linzer Biologische Beitrage 47 (1): 191 - 206." type="journal article" year="2015">Assing 2015</bibRefCitation>
&quot; (BMNH).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C322B8883A17FF8EFF0AEB7FFEEBFB3E" pageId="23" pageNumber="1156" type="discussion">
<paragraph id="8B87EB033A17FF8EFF0AEB7FFEEBFB3E" blockId="23.[178,1087,1086,1333]" pageId="23" pageNumber="1156">
Comment: The original description of
<taxonomicName id="4C3890803A17FF8EFDCAEB7EFD6CFB8B" authorityName="Cameron" authorityYear="1928" box="[626,730,1086,1108]" class="Insecta" family="Staphylinidae" genus="Lithocharis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="1156" phylum="Arthropoda" rank="species" species="sordida">
<emphasis id="B94C37113A17FF8EFDCAEB7EFD6CFB8B" box="[626,730,1086,1108]" italics="true" pageId="23" pageNumber="1156">L. sordida</emphasis>
</taxonomicName>
is based on an unspecified number of
<typeStatus id="548355A13A17FF8EFF69EB1DFE9CFBA9" box="[209,298,1116,1137]" pageId="23" pageNumber="1156" type="syntype">syntypes</typeStatus>
from &quot;Malay Peninsula,
<collectingRegion id="49FC25E13A17FF8EFD8DEB1AFD34FBA9" box="[565,642,1115,1137]" country="Malaysia" name="Pahang" pageId="23" pageNumber="1156">Pahang</collectingRegion>
&quot; (
<bibRefCitation id="EFA996F23A17FF8EFD23EB1AFCF5FBA9" author="CAMERON M" box="[667,835,1115,1137]" pageId="23" pageNumber="1156" pagination="244 - 247" refId="ref27756" refString="CAMERON M. (1928): On the Oriental species of the Lithocharis ochracea group of Staphylinidae (Col.). - The Entomologist's Monthly Magazine 64: 244 - 247." type="journal article" year="1928">CAMERON 1928</bibRefCitation>
). One slightly damaged male
<typeStatus id="548355A13A17FF8EFF56EB38FE8BFB56" box="[238,317,1145,1166]" pageId="23" pageNumber="1156" type="syntype">syntype</typeStatus>
(left elytron missing; male sternite VIII incomplete) was located in the Cameron collection at the BMNH; it is designated as the
<typeStatus id="548355A13A17FF8EFC9AEBD7FC34FB74" box="[802,898,1174,1196]" pageId="23" pageNumber="1156" type="lectotype">lectotype</typeStatus>
. The preoccupied name
<taxonomicName id="4C3890803A17FF8EFF48EBF2FE0CFB10" authorityName="Cameron" authorityYear="1928" box="[240,442,1203,1224]" class="Insecta" family="Staphylinidae" genus="Lithocharis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="1156" phylum="Arthropoda" rank="species" species="sordida">
<emphasis id="B94C37113A17FF8EFF48EBF2FE0CFB10" box="[240,442,1203,1224]" italics="true" pageId="23" pageNumber="1156">Lithocharis sordida</emphasis>
</taxonomicName>
was recently replaced with the nomen novum
<taxonomicName id="4C3890803A17FF8EFC26EBF5FBAAFB10" authorityName="HERMAN" authorityYear="2003" box="[926,1052,1203,1225]" class="Insecta" family="Staphylinidae" genus="Lithocharis" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="1156" phylum="Arthropoda" rank="species" species="sordidula">
<emphasis id="B94C37113A17FF8EFC26EBF5FBAAFB10" box="[926,1052,1203,1225]" italics="true" pageId="23" pageNumber="1156">L. sordidula</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFA996F23A17FF8EFF0AEB90FEEFFB3E" author="HERMAN L" box="[178,345,1232,1254]" pageId="23" pageNumber="1156" pagination="1 - 28" refId="ref28393" refString="HERMAN L. H. (2003): Nomenclatural changes in the Paederinae (Coleoptera: Staphylinidae). - American Museum Novitates 3416: 1 - 28." type="journal article" year="2003">HERMAN (2003)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C322B8883A17FF8EFF0AEBB0FEDDFAED" pageId="23" pageNumber="1156" type="materials_examined">
<paragraph id="8B87EB033A17FF8EFF0AEBB0FEDDFAED" blockId="23.[178,1087,1086,1333]" pageId="23" pageNumber="1156">
Additional material examined:
<materialsCitation id="3B50E15E3A17FF8EFD1EEBB0FD00FAC6" ID-GBIF-Occurrence="3805307358" collectingDate="1996-03-01" collectingDateMax="1996-03-21" collectingDateMin="1996-03-01" collectionCode="NHMB" collectorName="Majer" country="Thailand" latitude="9.8" location="Pha To" longLatPrecision="1299" longitude="98.78333" pageId="23" pageNumber="1156" specimenCount="1" stateProvince="Chumphon prov.">
<collectingCountry id="F32FAB933A17FF8EFD1EEBB0FD4AFADF" box="[678,764,1265,1287]" name="Thailand" pageId="23" pageNumber="1156">Thailand</collectingCountry>
: 1♂,
<collectingRegion id="49FC25E13A17FF8EFC8AEBB0FC7CFADE" box="[818,970,1265,1287]" country="Thailand" name="Chumphon" pageId="23" pageNumber="1156">Chumphon prov.</collectingRegion>
,
<location id="8EE7BDD83A17FF8EFC6AEBB0FBA5FADE" LSID="urn:lsid:plazi:treatment:03915A153A17FF81FF0AEC3DFC41FC0F:8EE7BDD83A17FF8EFC6AEBB0FBA5FADE" box="[978,1043,1265,1287]" country="Thailand" latitude="9.8" longLatPrecision="1299" longitude="98.78333" name="Pha To" pageId="23" pageNumber="1156" stateProvince="Chumphon prov.">Pha To</location>
env.
<geoCoordinate id="EE0C8DC43A17FF8EFF67EA49FEA8FAC5" box="[223,286,1288,1309]" degrees="9" direction="north" minutes="48" orientation="latitude" pageId="23" pageNumber="1156" precision="925" value="9.8">9°48'N</geoCoordinate>
,
<geoCoordinate id="EE0C8DC43A17FF8EFE91EA49FEC6FAC5" box="[297,368,1288,1309]" degrees="98" direction="east" minutes="47" orientation="longitude" pageId="23" pageNumber="1156" precision="925" value="98.78333">98°47'E</geoCoordinate>
,
<date id="FF86CDC33A17FF8EFEC3EA49FE43FAC6" box="[379,501,1288,1310]" pageId="23" pageNumber="1156" value="1996-03-01" valueMax="1996-03-21" valueMin="1996-03-01">
<collectingDate id="EFC2342B3A17FF8EFEC3EA49FE43FAC6" box="[379,501,1288,1310]" pageId="23" pageNumber="1156" value="1996-03-01" valueMax="1996-03-21" valueMin="1996-03-01">1.-21.III.1996</collectingDate>
</date>
, leg.
<collectorName id="26CD8ED53A17FF8EFD9FEA49FDEDFAC6" box="[551,603,1288,1310]" pageId="23" pageNumber="1156">Majer</collectorName>
(
<collectionCode id="ED2973C63A17FF8EFDD1EA49FD06FAC6" box="[617,688,1288,1310]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="23" pageNumber="1156">NHMB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B50E15E3A17FF8EFD79EA49FED1FAED" ID-GBIF-Occurrence="3805307464" collectingDate="1996-03-27" collectingDateMax="1996-04-14" collectingDateMin="1996-03-27" collectionCode="NHMB, cAss" collectorName="Majer" country="Thailand" latitude="9.8" location="Pha To" longLatPrecision="1299" longitude="98.78333" pageId="23" pageNumber="1156" specimenCount="2" specimenCount-female="1" stateProvince="Chumphon prov.">
<specimenCount id="9D3E208A3A17FF8EFD79EA49FD69FAC5" box="[705,735,1288,1309]" pageId="23" pageNumber="1156">1♂</specimenCount>
,
<specimenCount id="9D3E208A3A17FF8EFD53EA49FCBFFAFA" box="[747,777,1288,1314]" pageId="23" pageNumber="1156" type="female">1♀</specimenCount>
, same data, but
<date id="FF86CDC33A17FF8EFC26EA49FB8BFAC6" box="[926,1085,1288,1310]" pageId="23" pageNumber="1156" value="1996-03-27" valueMax="1996-04-14" valueMin="1996-03-27">
<collectingDate id="EFC2342B3A17FF8EFC26EA49FB8BFAC6" box="[926,1085,1288,1310]" pageId="23" pageNumber="1156" value="1996-03-27" valueMax="1996-04-14" valueMin="1996-03-27">27.III.-14.IV.1996</collectingDate>
</date>
(
<collectionCode id="ED2973C63A17FF8EFF5DEA5EFE9DFAEC" box="[229,299,1311,1332]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="23" pageNumber="1156">NHMB</collectionCode>
,
<collectionCode id="ED2973C63A17FF8EFE8BEA5EFED7FAED" box="[307,353,1311,1333]" pageId="23" pageNumber="1156">cAss</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C322B8883A17FF81FF0AEA02FE47FD18" lastPageId="24" lastPageNumber="1157" pageId="23" pageNumber="1156" type="description">
<paragraph id="8B87EB033A17FF8EFF0AEA02FCE6FA69" blockId="23.[178,1085,1347,1457]" pageId="23" pageNumber="1156">
Redescription: Body length
<quantity id="4CC046E63A17FF8EFDFEEA02FD73FA81" box="[582,709,1347,1369]" metricMagnitude="-3" metricUnit="m" metricValue="3.5999999999999996" metricValueMax="3.8" metricValueMin="3.4" pageId="23" pageNumber="1156" unit="mm" value="3.5999999999999996" valueMax="3.8" valueMin="3.4">3.4-3.8 mm</quantity>
; length of forebody
<quantity id="4CC046E63A17FF8EFC16EA02FB8FFA81" box="[942,1081,1347,1369]" metricMagnitude="-3" metricUnit="m" metricValue="1.975" metricValueMax="2.1" metricValueMin="1.85" pageId="23" pageNumber="1156" unit="mm" value="1.975" valueMax="2.1" valueMin="1.85">1.85-2.1 mm</quantity>
. Habitus as in
<figureCitation id="1303F7863A17FF8EFEF6EA21FE28FAAE" box="[334,414,1376,1398]" captionStart="Figs 60-66" captionStartId="22.[178,218,1224,1246]" captionTargetBox="[186,1081,212,1202]" captionTargetId="figure-1@22.[178,1091,212,1215]" captionTargetPageId="22" captionText="Figs 60-66: Lithocharis sordidula HERMAN (64-65: lectotype): (60) habitus; (61) forebody; (62) male sternite VII; (63) male sternite VIII; (64-65) aedeagus in lateral and in ventral view; (66) apical portion of aedeagus in ventral view. Scale bars: 60-61: 1.0 mm; 62-66: 0.2 mm. A d d i t i o n a l m a t e r i a l e x a m i n e d India: 1♂, Assam, Bhalukpong, 27°02'N, 92°35'E, 150 m, 26.V.-3.VI.2006, leg. Pacholátko (cAss); 1♂, 2♀♀, Assam, Umrongso env., 25°27'N, 92°43'E, 700 m, 3.-8.VI.2002, leg. Trýzna (cSch, cAss); 1♀, Meghalaya, 1 km E Tura, 25°30'N, 90°14'E, 500-600 m, 2.-5.V.2002, leg. Trýzna &amp; Benda (cSch). China: 1♂, Guangxi, Bama, cow dung, I.1988 (BMNH). Thailand: 2♂♂, 2♀♀, Chumphon prov., Pha To env., 9°48'N, 98°47'E, 27.III.-14.IV.1996, leg. Majer (NHMB, cAss); 1♂, Chumphon prov., Pha To env. 9°48'N, 98°47'E, 1.-21.III.1996, K. Majer leg. (NHMB); 1♂, Chumphon, Pha To env., 9°48'N, 98°47'E, III-IV.1996, at light, leg. Prudek (cSch); 1♂, 1♀, Ranong prov., Ranong: Hot Springs, 9°56'N, 98°40'E, 23.-25.II.1996, leg. Majer&quot; (NHMB, cAss). Laos: 8 exs., 240 km N Vientiane, 10 km N Luang Prabang, Mekong river, XI.1992, leg. Somsy (cSch, cAss). Singapore: 1♂, &quot;B.T.Ra.&quot; 4 m, I.1922, leg. Saunders (BMNH); 1 ex., &quot;Scott land&quot;, 5.VIII.1922, leg. Saunders (BMNH).Indonesia: 6 exs., Sulawesi Tengah, near Morowali, Ranu River area, lowland rain forest, at light, I-IV.1980, leg. Brendell (BMNH, cAss)." figureDoi="http://doi.org/10.5281/zenodo.5184202" httpUri="https://zenodo.org/record/5184202/files/figure.png" pageId="23" pageNumber="1156">Fig. 60</figureCitation>
. Coloration: head blackish; pronotum and elytra pale reddishbrown; abdomen brown, with the apex and the posterior margins of the segments paler; legs yellowish; antennae brown with antennomeres I-II yellowish.
</paragraph>
<paragraph id="8B87EB033A17FF8EFF0AEA81FE1EF9F6" blockId="23.[178,1085,1472,1582]" pageId="23" pageNumber="1156">
Head (
<figureCitation id="1303F7863A17FF8EFF4EEA81FEF6FA0E" box="[246,320,1472,1494]" captionStart="Figs 60-66" captionStartId="22.[178,218,1224,1246]" captionTargetBox="[186,1081,212,1202]" captionTargetId="figure-1@22.[178,1091,212,1215]" captionTargetPageId="22" captionText="Figs 60-66: Lithocharis sordidula HERMAN (64-65: lectotype): (60) habitus; (61) forebody; (62) male sternite VII; (63) male sternite VIII; (64-65) aedeagus in lateral and in ventral view; (66) apical portion of aedeagus in ventral view. Scale bars: 60-61: 1.0 mm; 62-66: 0.2 mm. A d d i t i o n a l m a t e r i a l e x a m i n e d India: 1♂, Assam, Bhalukpong, 27°02'N, 92°35'E, 150 m, 26.V.-3.VI.2006, leg. Pacholátko (cAss); 1♂, 2♀♀, Assam, Umrongso env., 25°27'N, 92°43'E, 700 m, 3.-8.VI.2002, leg. Trýzna (cSch, cAss); 1♀, Meghalaya, 1 km E Tura, 25°30'N, 90°14'E, 500-600 m, 2.-5.V.2002, leg. Trýzna &amp; Benda (cSch). China: 1♂, Guangxi, Bama, cow dung, I.1988 (BMNH). Thailand: 2♂♂, 2♀♀, Chumphon prov., Pha To env., 9°48'N, 98°47'E, 27.III.-14.IV.1996, leg. Majer (NHMB, cAss); 1♂, Chumphon prov., Pha To env. 9°48'N, 98°47'E, 1.-21.III.1996, K. Majer leg. (NHMB); 1♂, Chumphon, Pha To env., 9°48'N, 98°47'E, III-IV.1996, at light, leg. Prudek (cSch); 1♂, 1♀, Ranong prov., Ranong: Hot Springs, 9°56'N, 98°40'E, 23.-25.II.1996, leg. Majer&quot; (NHMB, cAss). Laos: 8 exs., 240 km N Vientiane, 10 km N Luang Prabang, Mekong river, XI.1992, leg. Somsy (cSch, cAss). Singapore: 1♂, &quot;B.T.Ra.&quot; 4 m, I.1922, leg. Saunders (BMNH); 1 ex., &quot;Scott land&quot;, 5.VIII.1922, leg. Saunders (BMNH).Indonesia: 6 exs., Sulawesi Tengah, near Morowali, Ranu River area, lowland rain forest, at light, I-IV.1980, leg. Brendell (BMNH, cAss)." figureDoi="http://doi.org/10.5281/zenodo.5184202" httpUri="https://zenodo.org/record/5184202/files/figure.png" pageId="23" pageNumber="1156">Fig. 61</figureCitation>
) as long as broad or weakly oblong; punctation dense and very fine; interstices with shallow microsculpture. Eyes large, distinctly longer than postocular portion in dorsal view. Antennae approximately
<quantity id="4CC046E63A17FF8EFDE1EABAFD1EF9C8" box="[601,680,1531,1552]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="23" pageNumber="1156" unit="mm" value="1.2">1.2 mm</quantity>
long; preapical antennomeres approximately as long as broad.
</paragraph>
<paragraph id="8B87EB033A18FF81FF0AEF9AFD71FEF3" blockId="24.[178,1085,219,299]" pageId="24" pageNumber="1157">
Pronotum (
<figureCitation id="1303F7863A18FF81FE91EF9AFECEFF29" box="[297,376,219,241]" captionStart="Figs 60-66" captionStartId="22.[178,218,1224,1246]" captionTargetBox="[186,1081,212,1202]" captionTargetId="figure-1@22.[178,1091,212,1215]" captionTargetPageId="22" captionText="Figs 60-66: Lithocharis sordidula HERMAN (64-65: lectotype): (60) habitus; (61) forebody; (62) male sternite VII; (63) male sternite VIII; (64-65) aedeagus in lateral and in ventral view; (66) apical portion of aedeagus in ventral view. Scale bars: 60-61: 1.0 mm; 62-66: 0.2 mm. A d d i t i o n a l m a t e r i a l e x a m i n e d India: 1♂, Assam, Bhalukpong, 27°02'N, 92°35'E, 150 m, 26.V.-3.VI.2006, leg. Pacholátko (cAss); 1♂, 2♀♀, Assam, Umrongso env., 25°27'N, 92°43'E, 700 m, 3.-8.VI.2002, leg. Trýzna (cSch, cAss); 1♀, Meghalaya, 1 km E Tura, 25°30'N, 90°14'E, 500-600 m, 2.-5.V.2002, leg. Trýzna &amp; Benda (cSch). China: 1♂, Guangxi, Bama, cow dung, I.1988 (BMNH). Thailand: 2♂♂, 2♀♀, Chumphon prov., Pha To env., 9°48'N, 98°47'E, 27.III.-14.IV.1996, leg. Majer (NHMB, cAss); 1♂, Chumphon prov., Pha To env. 9°48'N, 98°47'E, 1.-21.III.1996, K. Majer leg. (NHMB); 1♂, Chumphon, Pha To env., 9°48'N, 98°47'E, III-IV.1996, at light, leg. Prudek (cSch); 1♂, 1♀, Ranong prov., Ranong: Hot Springs, 9°56'N, 98°40'E, 23.-25.II.1996, leg. Majer&quot; (NHMB, cAss). Laos: 8 exs., 240 km N Vientiane, 10 km N Luang Prabang, Mekong river, XI.1992, leg. Somsy (cSch, cAss). Singapore: 1♂, &quot;B.T.Ra.&quot; 4 m, I.1922, leg. Saunders (BMNH); 1 ex., &quot;Scott land&quot;, 5.VIII.1922, leg. Saunders (BMNH).Indonesia: 6 exs., Sulawesi Tengah, near Morowali, Ranu River area, lowland rain forest, at light, I-IV.1980, leg. Brendell (BMNH, cAss)." figureDoi="http://doi.org/10.5281/zenodo.5184202" httpUri="https://zenodo.org/record/5184202/files/figure.png" pageId="24" pageNumber="1157">Fig. 61</figureCitation>
) approximately as long as broad and as broad as head; posterior angles rounded, weakly marked; punctation similar to that of head; midline with distinct impunctate band; interstices without microsculpture.
</paragraph>
<paragraph id="8B87EB033A18FF81FF0AEE7AFE56FE70" blockId="24.[178,1085,315,424]" pageId="24" pageNumber="1157">
Elytra (
<figureCitation id="1303F7863A18FF81FEB8EE7AFEFDFE89" box="[256,331,315,337]" captionStart="Figs 60-66" captionStartId="22.[178,218,1224,1246]" captionTargetBox="[186,1081,212,1202]" captionTargetId="figure-1@22.[178,1091,212,1215]" captionTargetPageId="22" captionText="Figs 60-66: Lithocharis sordidula HERMAN (64-65: lectotype): (60) habitus; (61) forebody; (62) male sternite VII; (63) male sternite VIII; (64-65) aedeagus in lateral and in ventral view; (66) apical portion of aedeagus in ventral view. Scale bars: 60-61: 1.0 mm; 62-66: 0.2 mm. A d d i t i o n a l m a t e r i a l e x a m i n e d India: 1♂, Assam, Bhalukpong, 27°02'N, 92°35'E, 150 m, 26.V.-3.VI.2006, leg. Pacholátko (cAss); 1♂, 2♀♀, Assam, Umrongso env., 25°27'N, 92°43'E, 700 m, 3.-8.VI.2002, leg. Trýzna (cSch, cAss); 1♀, Meghalaya, 1 km E Tura, 25°30'N, 90°14'E, 500-600 m, 2.-5.V.2002, leg. Trýzna &amp; Benda (cSch). China: 1♂, Guangxi, Bama, cow dung, I.1988 (BMNH). Thailand: 2♂♂, 2♀♀, Chumphon prov., Pha To env., 9°48'N, 98°47'E, 27.III.-14.IV.1996, leg. Majer (NHMB, cAss); 1♂, Chumphon prov., Pha To env. 9°48'N, 98°47'E, 1.-21.III.1996, K. Majer leg. (NHMB); 1♂, Chumphon, Pha To env., 9°48'N, 98°47'E, III-IV.1996, at light, leg. Prudek (cSch); 1♂, 1♀, Ranong prov., Ranong: Hot Springs, 9°56'N, 98°40'E, 23.-25.II.1996, leg. Majer&quot; (NHMB, cAss). Laos: 8 exs., 240 km N Vientiane, 10 km N Luang Prabang, Mekong river, XI.1992, leg. Somsy (cSch, cAss). Singapore: 1♂, &quot;B.T.Ra.&quot; 4 m, I.1922, leg. Saunders (BMNH); 1 ex., &quot;Scott land&quot;, 5.VIII.1922, leg. Saunders (BMNH).Indonesia: 6 exs., Sulawesi Tengah, near Morowali, Ranu River area, lowland rain forest, at light, I-IV.1980, leg. Brendell (BMNH, cAss)." figureDoi="http://doi.org/10.5281/zenodo.5184202" httpUri="https://zenodo.org/record/5184202/files/figure.png" pageId="24" pageNumber="1157">Fig. 61</figureCitation>
) as long as pronotum or slightly longer; punctation very dense and fine; interstices without microsculpture. Hind wings present. Protarsomeres I-IV moderately dilated, slightly more so in male than in female. Metatarsomere I nearly as long as the combined length of II and III.
</paragraph>
<paragraph id="8B87EB033A18FF81FF0AEEF9FCCEFE33" blockId="24.[178,1085,440,491]" pageId="24" pageNumber="1157">Abdomen narrower than elytra; punctation fine and very dense; interstices with distinct microreticulation; posterior margin of tergite VII with palisade fringe.</paragraph>
<paragraph id="8B87EB033A18FF81FF0AEEBBFE47FD18" blockId="24.[178,1085,506,704]" pageId="24" pageNumber="1157">
♂: sternite VII (
<figureCitation id="1303F7863A18FF81FEEFEEBBFE17FDC8" box="[343,417,506,528]" captionStart="Figs 60-66" captionStartId="22.[178,218,1224,1246]" captionTargetBox="[186,1081,212,1202]" captionTargetId="figure-1@22.[178,1091,212,1215]" captionTargetPageId="22" captionText="Figs 60-66: Lithocharis sordidula HERMAN (64-65: lectotype): (60) habitus; (61) forebody; (62) male sternite VII; (63) male sternite VIII; (64-65) aedeagus in lateral and in ventral view; (66) apical portion of aedeagus in ventral view. Scale bars: 60-61: 1.0 mm; 62-66: 0.2 mm. A d d i t i o n a l m a t e r i a l e x a m i n e d India: 1♂, Assam, Bhalukpong, 27°02'N, 92°35'E, 150 m, 26.V.-3.VI.2006, leg. Pacholátko (cAss); 1♂, 2♀♀, Assam, Umrongso env., 25°27'N, 92°43'E, 700 m, 3.-8.VI.2002, leg. Trýzna (cSch, cAss); 1♀, Meghalaya, 1 km E Tura, 25°30'N, 90°14'E, 500-600 m, 2.-5.V.2002, leg. Trýzna &amp; Benda (cSch). China: 1♂, Guangxi, Bama, cow dung, I.1988 (BMNH). Thailand: 2♂♂, 2♀♀, Chumphon prov., Pha To env., 9°48'N, 98°47'E, 27.III.-14.IV.1996, leg. Majer (NHMB, cAss); 1♂, Chumphon prov., Pha To env. 9°48'N, 98°47'E, 1.-21.III.1996, K. Majer leg. (NHMB); 1♂, Chumphon, Pha To env., 9°48'N, 98°47'E, III-IV.1996, at light, leg. Prudek (cSch); 1♂, 1♀, Ranong prov., Ranong: Hot Springs, 9°56'N, 98°40'E, 23.-25.II.1996, leg. Majer&quot; (NHMB, cAss). Laos: 8 exs., 240 km N Vientiane, 10 km N Luang Prabang, Mekong river, XI.1992, leg. Somsy (cSch, cAss). Singapore: 1♂, &quot;B.T.Ra.&quot; 4 m, I.1922, leg. Saunders (BMNH); 1 ex., &quot;Scott land&quot;, 5.VIII.1922, leg. Saunders (BMNH).Indonesia: 6 exs., Sulawesi Tengah, near Morowali, Ranu River area, lowland rain forest, at light, I-IV.1980, leg. Brendell (BMNH, cAss)." figureDoi="http://doi.org/10.5281/zenodo.5184202" httpUri="https://zenodo.org/record/5184202/files/figure.png" pageId="24" pageNumber="1157">Fig. 62</figureCitation>
) moderately transverse, posterior margin nearly truncate and with comb of approximately 20 moderately long palisade setae; sternite VIII (
<figureCitation id="1303F7863A18FF81FC5CED59FB83FDF5" box="[996,1077,535,558]" captionStart="Figs 60-66" captionStartId="22.[178,218,1224,1246]" captionTargetBox="[186,1081,212,1202]" captionTargetId="figure-1@22.[178,1091,212,1215]" captionTargetPageId="22" captionText="Figs 60-66: Lithocharis sordidula HERMAN (64-65: lectotype): (60) habitus; (61) forebody; (62) male sternite VII; (63) male sternite VIII; (64-65) aedeagus in lateral and in ventral view; (66) apical portion of aedeagus in ventral view. Scale bars: 60-61: 1.0 mm; 62-66: 0.2 mm. A d d i t i o n a l m a t e r i a l e x a m i n e d India: 1♂, Assam, Bhalukpong, 27°02'N, 92°35'E, 150 m, 26.V.-3.VI.2006, leg. Pacholátko (cAss); 1♂, 2♀♀, Assam, Umrongso env., 25°27'N, 92°43'E, 700 m, 3.-8.VI.2002, leg. Trýzna (cSch, cAss); 1♀, Meghalaya, 1 km E Tura, 25°30'N, 90°14'E, 500-600 m, 2.-5.V.2002, leg. Trýzna &amp; Benda (cSch). China: 1♂, Guangxi, Bama, cow dung, I.1988 (BMNH). Thailand: 2♂♂, 2♀♀, Chumphon prov., Pha To env., 9°48'N, 98°47'E, 27.III.-14.IV.1996, leg. Majer (NHMB, cAss); 1♂, Chumphon prov., Pha To env. 9°48'N, 98°47'E, 1.-21.III.1996, K. Majer leg. (NHMB); 1♂, Chumphon, Pha To env., 9°48'N, 98°47'E, III-IV.1996, at light, leg. Prudek (cSch); 1♂, 1♀, Ranong prov., Ranong: Hot Springs, 9°56'N, 98°40'E, 23.-25.II.1996, leg. Majer&quot; (NHMB, cAss). Laos: 8 exs., 240 km N Vientiane, 10 km N Luang Prabang, Mekong river, XI.1992, leg. Somsy (cSch, cAss). Singapore: 1♂, &quot;B.T.Ra.&quot; 4 m, I.1922, leg. Saunders (BMNH); 1 ex., &quot;Scott land&quot;, 5.VIII.1922, leg. Saunders (BMNH).Indonesia: 6 exs., Sulawesi Tengah, near Morowali, Ranu River area, lowland rain forest, at light, I-IV.1980, leg. Brendell (BMNH, cAss)." figureDoi="http://doi.org/10.5281/zenodo.5184202" httpUri="https://zenodo.org/record/5184202/files/figure.png" pageId="24" pageNumber="1157">Fig. 63</figureCitation>
) approximately as long as broad and with posterior excision of somewhat variable shape (sometimes more broadly V-shaped); aedeagus (
<figureCitation id="1303F7863A18FF81FD26ED13FCB8FDB0" box="[670,782,594,616]" captionStart="Figs 60-66" captionStartId="22.[178,218,1224,1246]" captionTargetBox="[186,1081,212,1202]" captionTargetId="figure-1@22.[178,1091,212,1215]" captionTargetPageId="22" captionText="Figs 60-66: Lithocharis sordidula HERMAN (64-65: lectotype): (60) habitus; (61) forebody; (62) male sternite VII; (63) male sternite VIII; (64-65) aedeagus in lateral and in ventral view; (66) apical portion of aedeagus in ventral view. Scale bars: 60-61: 1.0 mm; 62-66: 0.2 mm. A d d i t i o n a l m a t e r i a l e x a m i n e d India: 1♂, Assam, Bhalukpong, 27°02'N, 92°35'E, 150 m, 26.V.-3.VI.2006, leg. Pacholátko (cAss); 1♂, 2♀♀, Assam, Umrongso env., 25°27'N, 92°43'E, 700 m, 3.-8.VI.2002, leg. Trýzna (cSch, cAss); 1♀, Meghalaya, 1 km E Tura, 25°30'N, 90°14'E, 500-600 m, 2.-5.V.2002, leg. Trýzna &amp; Benda (cSch). China: 1♂, Guangxi, Bama, cow dung, I.1988 (BMNH). Thailand: 2♂♂, 2♀♀, Chumphon prov., Pha To env., 9°48'N, 98°47'E, 27.III.-14.IV.1996, leg. Majer (NHMB, cAss); 1♂, Chumphon prov., Pha To env. 9°48'N, 98°47'E, 1.-21.III.1996, K. Majer leg. (NHMB); 1♂, Chumphon, Pha To env., 9°48'N, 98°47'E, III-IV.1996, at light, leg. Prudek (cSch); 1♂, 1♀, Ranong prov., Ranong: Hot Springs, 9°56'N, 98°40'E, 23.-25.II.1996, leg. Majer&quot; (NHMB, cAss). Laos: 8 exs., 240 km N Vientiane, 10 km N Luang Prabang, Mekong river, XI.1992, leg. Somsy (cSch, cAss). Singapore: 1♂, &quot;B.T.Ra.&quot; 4 m, I.1922, leg. Saunders (BMNH); 1 ex., &quot;Scott land&quot;, 5.VIII.1922, leg. Saunders (BMNH).Indonesia: 6 exs., Sulawesi Tengah, near Morowali, Ranu River area, lowland rain forest, at light, I-IV.1980, leg. Brendell (BMNH, cAss)." figureDoi="http://doi.org/10.5281/zenodo.5184202" httpUri="https://zenodo.org/record/5184202/files/figure.png" pageId="24" pageNumber="1157">Figs 64-66</figureCitation>
) approximately
<quantity id="4CC046E63A18FF81FC0DED13FBB4FDB0" box="[949,1026,594,616]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="24" pageNumber="1157" unit="mm" value="0.7">0.7 mm</quantity>
long; ventral process with pair of ventral teeth in the middle (best visible in lateral view); dorso-lateral apophyses very long, nearly reaching apex of ventral process; internal structures of distinctive shapes.
</paragraph>
</subSubSection>
<subSubSection id="C322B8883A18FF81FF0AED8EFDF2FCA0" pageId="24" pageNumber="1157" type="discussion">
<paragraph id="8B87EB033A18FF81FF0AED8EFDF2FCA0" blockId="24.[178,1085,719,888]" pageId="24" pageNumber="1157">
Comparative notes: This species is characterized particularly by the slender head and pronotum, by rather small body size, by the shapes and chaetotaxy of the male sternites VII and VIII, and by the morphology of the aedeagus (ventral process with pair of teeth, dorso-lateral apophyses very long, shapes of internal structures). For characters distinguishing it from the similarly small and syntopic
<taxonomicName id="4C3890803A18FF81FC9BEC04FC3CFC82" authorityName="Assing" authorityYear="2015" box="[803,906,837,858]" class="Insecta" family="Staphylinidae" genus="Lithocharis" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="1157" phylum="Arthropoda" rank="species" species="inermis">
<emphasis id="B94C37113A18FF81FC9BEC04FC3CFC82" box="[803,906,837,858]" italics="true" pageId="24" pageNumber="1157">L. inermis</emphasis>
</taxonomicName>
see the comparative notes in the section on that species.
</paragraph>
</subSubSection>
<subSubSection id="C322B8883A18FF81FF0AECC6FC41FC0F" pageId="24" pageNumber="1157" type="distribution">
<paragraph id="8B87EB033A18FF81FF0AECC6FC41FC0F" blockId="24.[178,1085,903,983]" pageId="24" pageNumber="1157">
Distribution and natural history: Confirmed records are known only from two localities, one in
<collectingRegion id="49FC25E13A18FF81FDACECE5FD77FC62" box="[532,705,932,954]" country="Malaysia" name="Pahang" pageId="24" pageNumber="1157">Pahang province</collectingRegion>
in Peninsular
<collectingCountry id="F32FAB933A18FF81FCDAECE5FC09FC62" box="[866,959,932,954]" name="Malaysia" pageId="24" pageNumber="1157">Malaysia</collectingCountry>
and one in
<collectingRegion id="49FC25E13A18FF81FF0AEC80FE37FC0F" box="[178,385,961,983]" country="Thailand" name="Chumphon" pageId="24" pageNumber="1157">Chumphon province</collectingRegion>
in southern
<collectingCountry id="F32FAB933A18FF81FE47EC80FDEAFC0F" box="[511,604,961,983]" name="Thailand" pageId="24" pageNumber="1157">Thailand</collectingCountry>
. The
<typeStatus id="548355A13A18FF81FD2BEC80FD46FC0F" box="[659,752,961,983]" pageId="24" pageNumber="1157" type="lectotype">lectotype</typeStatus>
was collected from dung.
</paragraph>
</subSubSection>
</treatment>
</document>
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